Bulletin of the Kagoshima University Museum No. 14
ISSN-L 2188-9074

AN ANNOTATED CHECKLIST OF MARINE AND FRESHWATER FISHES OFKIKAI ISLAND IN THE AMAMI ISLANDS, KAGOSHIMA,
SOUTHERN JAPAN, WITH 259 NEW RECORDS

KYOJI FUJIWARA AND HIROYUKI MOTOMURA

December 2020

The Kagoshima University Museum

Cover photograph: A school of Lutjanus kasmira (Lutjanidae) off Kikai Island. Photo by J. Yoda
Back cover photograph: A school of Chromis viridis (Pomacentridae) off Kikai Island. Photo by M. Matsuoka

Bulletin of the Kagoshima University Museum No.14

ISSN-L 2188-9074

An annotated checklist of marine and freshwater fishes of
Kikai Island in the Amami Islands, Kagoshima,southern Japan, with 259 new records

Kyoji Fujiwara1, 3 and Hiroyuki Motomura2
1

The United Graduate School of Agricultural Sciences, Kagoshima University,
1–21–24 Korimoto, Kagoshima 890–0065, Japan

2

The Kagoshima University Museum, 1–21–30 Korimoto, Kagoshima 890–0065, Japan
3

Corresponding author, e-mail: kyojifujiwara627@yahoo.co.jp

Abstract An annotated checklist of marine and freshwater fishes of Kikai
Island, Amami Islands, Kagoshima Prefecture, southern Japan, was compiled
from specimen and literature surveys. A total of 530 species (255 genera and85 families), including 259 species recorded reliably for the first time on thebasis of collected specimens and/or underwater photographs, are listed withliterature citations, institution registration numbers, size, island localities,and (in some cases) remarks. Two of the newly recorded species from Kikai
Island [Valenciennea sexguttata (Gobiidae) and Paramonacanthus oblongus(Monacanthidae)] also represent the first records from the Amami Islands.
Preliminary comparisons of the island ichthyofauna with those of 16 otherlocalities in Japan show the former to be similar to that of Tokunoshima islandin the Amami Islands.

Fujiwara, K. and H. Motomura. 2020. An annotated checklist of marine and freshwater fishes of Kikai Islandin the Amami Islands, Kagoshima, southern Japan, with 259 new records. Bulletin of the Kagoshima
University Museum, 14: 1–73.
Published on 1 December 2020

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K. Fujiwara and H. Motomura

Introduction
Kikai Island, a boot-shaped island of area ca. 57 km2 and a highest point of 214 m above sea level, is thefifth largest island of the Amami Islands group, located approximately midway down the Ryukyu Archipelago, southern Japan (Fig. 1). The coastline extends for ca. 49 km, with well-developed rocky reefs.
Since comprehensive ichthyofaunal surveys of Kikai Island had not been previously undertaken, althougha five-person mid-level survey had been conducted in May 2015, a major “Kikai Expedition” to study theisland’s ichthyofauna took place in November 2019, involving 17 personnel from the Kagoshima University
Museum and Kindai University. This was followed by a second mid-level expedition to the island in October
2020. In total, ca. 2,900 marine and freshwater fish specimens were collected during these expeditions. In addition, 96 specimens, collected from Kikai Island in May 1954, July 1960, and July/August 1976, and foundin the Faculty of Fisheries, Kagoshima University, were registered into the Kagoshima University Museum
fish collection and incorporated into this study.
Examination of the above-mentioned specimens, published literature, and unpublished underwater photographs taken at Kikai Island confirmed 530 species of marine and freshwater fishes, including 259 newrecords, from the island. All species are listed below. The ichthyofauna of Kikai Island is briefly comparedwith that of other islands in the Ryukyu, Osumi, and Ogasawara islands, and several localities along the Pacific coast of southern Japan.
Material and methods
Sampling methods were described and illustrated in Motomura & Matsuura (2010, 2014). Curatorial procedures for newly-collected specimens followed Motomura & Ishikawa (2013). Standard and total lengths
are abbreviated as SL and TL, respectively. The systematic arrangement of families generally follows Nelson(2006), with scientific names generally following Fricke et al. (2020) and Motomura (2020), with some modifications following recent taxonomic studies (both published and unpublished). Within families, species arearranged by alphabetical order of scientific name. Standard Japanese names transliterated using the Hepburn
system follow Motomura (2020). Each species record was compiled from voucher specimens, underwaterphotographs, and/or published literature records. Registration number, number of specimens registered (ifmore than one), mm (SL or TL), and locality on Kikai Island (Fig. 2) are included in each species record.
Because Kikai Island has only one town (Kikai), the localities of marine and freshwater fishes are indicatedonly by the local area name. All freshwater fishes were collected from small ponds and/or streams. Locality
is restricted to “Kikai Island” for fish(es) collected well off the island or if the precise locality on the island is
unknown.
Institutional codes used in this paper are as follows: BSKU (Laboratory of Marine Biology, Faculty of
Science, Kochi University, Kochi, Japan), KAUM (Kagoshima University Museum, Kagoshima, Japan),
KPM (Kanagawa Prefectural Museum of Natural History, Kanagawa, Japan), KMNH (Kitakyushu Museumof Natural History and Human History, Fukuoka, Japan), KUN (Kindai University, Nara, Japan), SMBL
(Kyoto University Seto Marine Biological Laboratory, Wakayama, Japan), and YCM (Yokosuka City Museum, Kanagawa, Japan).
Remarks
The freshwater fish diversity of Kikai Island is poor, being restricted to small ponds and streams. All
freshwater fish species on the island are introduced, except for Misgurnus anguillicaudatus (Cantor, 1842)(Cobitidae) (currently considered to be an undescribed species, Misgurnus sp. IR) and Oryzias latipes(Temminck & Schlegel, 1846) (Adrianichthyidae) [e.g., two poeciliids, Gambusia affinis affinis (Baird &Girard, 1853) and Poecilia reticulata Peters, 1859, and the cichlid Oreochromis niloticus (Linnaeus, 1758)](Nakajima et al., 2020; this study). Although Hayashi (1995) recorded three freshwater fishes [Carassius sp.
(Cyprinidae), M. anguillicaudatus, and O. latipes], Kano et al. (2016) considered all three to have become
extinct on the island. However, Nakajima et al. (2020) recently confirmed the existence of Carassius sp. (asMarine and freshwater fishes of Kikai Island

3

C. langsdorfii) on Kikai Island, in addition to recording Carassius auratus (Linnaeus, 1758), believing bothto have likely been introduced. During the present study, several individuals of O. latipes were collected, but
M. anguillicaudatus (= Misgurnus sp. IR) was not found.
In addition to the above-mentioned species, 12 fish species were confirmed from freshwater area on Kikai
Island, as follows: Anguilla japonica Temminck & Schlegel, 1846, Anguilla marmorata Quoy & Gaimard,1824 (Anguillidae), Kuhlia mugil (Forster, 1801), Kuhlia rupestris (Lacepède, 1802) (Kuhliidae), Eleotris
acanthopoma Bleeker, 1853, Eleotris fusca (Bloch & Schneider, 1801), Eleotris melanosoma Bleeker, 1853
(Eleotridae), Oligolepis acutipennis (Valenciennes, 1837), Rhinogobius brunneus (Temminck & Schlegel,1845), Rhinogobius similis Gill, 1859, Stenogobius sp., and Trypauchenopsis intermedia Volz, 1903 (Gobiidae).
Marine fishes from Kikai Island have been incompletely listed by Kamohara (1957) (129 species), Kamohara & Yamakawa (1965, 1968a, b) (59, 25, and 12 species, respectively), and Yamakawa (1969, 1971)(4 and 3 species respectively), identifying a total of 185 species, based mostly on unregistered specimens.
However, although Zenion japonicum Kamohara, 1934 (Zeniidae), Hippichthys (Parasyngnathus) penicillus (Cantor, 1849) (as Syngnathus argyrostictus) (Syngnathidae), Epinephelus trimaculatus (Valenciennes,
1828) (as Epinephelus fario), Epinephelus quoyanus (Valenciennes, 1830) (as Epinephelus merra megachir)(Serranidae), Branchiostegus albus Dooley, 1978 (as Branchiostegus argentatus) (Branchiostegidae), Dentex
hypselosomus Bleeker, 1854 (as Taius tumifrons) (Sparidae), Aspasma ubauo Fujiwara & Motomura, 2019
(as Aspasma minima), and Aspasmichthys ciconiae (Jordan & Fowler, 1902) (Gobiesocidae) were recorded
from Kikai Island by Kamohara (1957) and Kamohara & Yamaksawa (1965), those species do not usuallyoccur in the Ryukyu Islands and the above records have been omitted here.
Six nominal species have been described on the basis of a primary type specimen from Kikai Island [Aporops japonicus Kamohara, 1957 (Serranidae), Pseudochromis kikaii Aoyagi, 1941, Pseudochromis luteus
Aoyagi, 1943 (Pseudochromidae), Cirrhitoidea unimacula Kamohara, 1957 (Cirrhitidae), Salarias kikaiensis
Aoyagi, 1954 (Blenniidae), and Mars nigrocellatus Yanagisawa, 1978 (Gobiidae)]. Currently, three of the six
are regarded as valid (Pseudochromis luteus, Amblycirrhitus unimaculus, and Cryptocentrus nigrocellatus).
The number of species in the top 10 most speciose families occurring at Kikai Island are shown in Table
1. Comparable data for 16 localities in southern Japan, including the Sagami Sea, Osezaki (Suruga Bay),
Kushimoto (Kii Peninsula), Kashiwa Island (Shikoku), Iwo and Take islands (Osumi Islands), Yaku Island(Osumi Islands), Amami-oshima Island (Amami Islands), Tokunoshima Island (Amami Islands), Okinoerabu
Island (Amami Islands), Yoron Island (Amami Islands), Okinawa Island (Okinawa Islands), Ie Island (Okinawa Islands), the Miyako Islands, Ishigaki Island (Yaeyama Islands), Iriomote Island (Yaeyama Islands),
and the Ogasawara Islands, were included in Motomura (2016: tables 1, 2), Nakae et al. (2018: table 2),
Mochida & Motomura (2018: table 1), and Motomura
Table 1. Number of species in the top 10 most speciose families
at Kikai Island.
& Uehara (2020: table 1). The three most speciose
families at all localities [except the Sagami Sea, OsFamily
No. of species% in total
Gobiidae63
11.9
ezaki (Suruga Bay), and the Ogasawara Islands], are
Pomacentridae
48
9.1
Gobiidae, Pomacentridae, and Labridae, with SerraniLabridae
45
8.5
dae replacing Pomacentridae and occupying third most
Blennidae
33
6.2
speciose position at the above exceptions (Motomura,
Apogonidae
23
4.32016). Although within the Amami Islands group,
Serranidae
19
3.6
Kikai Island is geographically located close to AmaScorpaenidae
18
3.4
mi-oshima Island, the ichthyofauna of Kikai Island
Lutjanidae
17
3.2
is closer to those of Tokunoshima, Okinoerabu, andChaetodontidae
16
3.0
Acanthuridae
16
3.0
Yoron islands than Amami-oshima Island (Table 1;
Other families
232
43.8
Motomura, 2016; Mochida & Motomura, 2018; MotoTotal
530
100.0
mura & Uehara, 2020). This is because Amami-oshima

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K. Fujiwara and H. Motomura

Island is the largest island (12.5 times larger than Kikai Island) in the Amami Islands and has various environments such as mangroves and large rivers compared with other islands only with mostly coral or rocky
reefs. The top 10 most speciose families, and percentage proportion of species in each family to total numberof species at Kikai Island are very similar to those of Tokunoshima Island (Table 1; Mochida & Motomura,
2018: table 1), the only difference between Kikai and Tokunoshima islands involving the family Lutjanidae(17 species), which is ranked as 10th most speciose family at Kikai Island, but is not so well represented at
Tokunoshima Island [being replaced by the families Syngnathidae (13 species) and Balistidae (13 species)](Mochida & Motomura, 2018: table 1). At Kikai Island, Syngnathidae (9 species) and Balistidae (10 species)
are ranked as 16th and 15th most speciose families, respectively. The ichthyofaunal similarity between Kikai and Tokunoshima islands is likely due to their geographical close distance (Kikai Island is the closet to
Tokunoshima Island in above-mentioned islands, except for Amami-oshima Island).
List of fishes from Kikai Island
ORDER ELOPIFORMESFAMILY MEGALOPIDAE
Megalops cyprinoides (Broussonet, 1782)[Jpn name: Isegoi]
KAUM–I. 134577, 32.4 mm SL, Shitooke.
ORDER ANGUILLIFORMESFAMILY ANGUILLIDAE
Anguilla japonica Temminck & Schlegel, 1846
[Jpn name: Nihon-unagi]
KAUM–I. 7633, 87.2 mm TL, KAUM–I. 8148, 48.0 mm TL, Urahara; KAUM–I. 134971, 166.6 mm TL,
Wan.
Anguilla marmorata Quoy & Gaimard, 1824
[Jpn name: Ounagi]
KAUM–I. 8407, 113.4 mm TL, Urahara; KAUM–I. 134585, 94.4 mm TL, KAUM–I. 134586, 67.0 mm
TL, Shitooke; KAUM–I. 134952, 489.4 mm TL, Shiomichi; KAUM–I. 134960, 51.2 mm TL, Shiramizu;
KAUM–I. 136794, 417.0 mm TL, Isaneku.
FAMILY MORINGUIDAE
Moringua microchir Bleeker, 1853
[Jpn name: Hariganeumihebi]
Yamakawa (1971, as Moringua abbreviatus): 1 unregistered specimen, 267 mm TL, Kikai Island.
FAMILY CHLOPSIDAE
Kaupichthys atronasus Schultz, 1953
[Jpn name: Kurohigeiwaanago]
Hatooka (2013a): Kikai Island.
Remarks: In Japanese waters, this species has been recorded only from Kikai Island (Hatooka, 2013a).
FAMILY MURAENIDAE
Echidna nebulosa (Ahl, 1789)[Jpn name: Kumoutsubo]Kamohara (1957): 3 unregistered specimens, 160–173 mm TL, Wan; 1 unregistered specimen, 257 mm
TL, Somachi. Kamohara & Yamakawa (1965): 4 unregistered specimens, 136–300 mm TL, Tekuzuku. This
study: KAUM–I. 4565, 95.9 mm TL, KAUM–I. 4566, 97.4 mm TL, Kikai Island; KAUM–I. 135130, 269.9mm TL, Araki; KAUM–I. 135138, 250.9 mm TL, Shiomichi.

Marine and freshwater fishes of Kikai Island

5

Echidna polyzona (Richardson, 1845)[Jpn name: Shimaarashiutsubo]
KAUM–I. 134794, 136.8 mm TL, Keraji.
Enchelycore lichenosa (Jordan & Snyder, 1901)[Jpn name: Kokeutsubo]Kamohara & Yamakawa (1965, as Aemasia lichenosa): 1 unregistered specimen, 190 mm TL, Somachi.
Enchelycore pardalis (Temminck & Schlegel, 1846)[Jpn name: Torautsubo]Kamohara & Yamakawa (1965, as Muraena pardalis): 1 unregistered specimen, 135 mm TL, Somachi.
Gymnomuraena zebra (Shaw, 1797)[Jpn name: Zeburautsubo]Kamohara (1957, as Arndha zebra): 2 unregistered specimens, 146–150 mm TL, Onotsu; 1 unregisteredspecimen, 188 mm TL, Somachi. Kamohara & Yamakawa (1965, as Echidna zebra): 1 unregistered specimen, 170 mm TL, Araki.
Gymnothorax enigmaticus McCosker & Randall, 1982
[Jpn name: Shimautsubo]
KAUM–I. 4567, 95.7 mm TL, Kikai Island.
Gymnothorax fimbriatus (Bennett, 1832)[Jpn name: Herigoishiutsubo]Kamohara & Yamakawa (1965): 2 unregistered specimens, 182–267 mm TL, Tekuzuku. This study: KAUM–
I. 2382, 104.1 mm TL, Kikai Island; KAUM–I. 134836, 476.0 mm TL, KAUM–I. 136714, 189.4 mm TL,
KAUM–I. 138112, 348.3 mm TL, KAUM–I. 138113, 211.3 mm TL, KAUM–I. 138114, 199.3 mm TL,
KAUM–I. 138115, 168.1 mm TL, Shiomichi.
Gymnothorax flavimarginatus (Rüppell, 1830)[Jpn name: Gomautsubo]Kamohara (1957): 4 unregistered specimens, 160–365 mm TL, Onotsu; 2 unregistered specimens, 275–280
mm TL, Somachi. Kamohara & Yamakawa (1968a): 1 unregistered specimen, ca. 340+ mm TL, Kikai Island.
This study: KAUM–I. 1983, 87.1 mm TL, KAUM–I. 14743, 114.0 mm TL, Kikai Island; KAUM–I. 134999,
174.7 mm TL, Nakaguma; KUN-P 55739, 240.4 mm TL, Aden.
Gymnothorax meleagris (Shaw, 1795)[Jpn name: Wakautsubo]Kamohara & Yamakawa (1965, as Gymnothorax leucostictus): 1 unregistered specimen, 118 mm TL, Somachi. Hibino (2018a): KAUM–I. 72514, 474.8 mm TL, Akaren. Hibino (2019a): KAUM–I. 72514, 474.8 mm
TL, Akaren. This study: KAUM–I. 72514, 474.8 mm TL, Akaren; KAUM–I. 136795, 356.4 mm TL, Somachi.
Gymnothorax pictus (Ahl, 1789)[Jpn name: Aseutsubo]Kamohara (1957): 1 unregistered specimen, 760 mm TL, Wan; 1 unregistered specimen, 740 mm TL, Onotsu. This study: KAUM–I. 134645, 725.9 mm TL, Akaren; KAUM–I. 136796, 923.0 mm TL, Somachi;
KAUM–I. 146849, 773.0 mm TL, Sakamine; KAUM–I. 147643, 303.2 mm TL, Ikeji.
Gymnothorax rueppellii (McClelland, 1844)[Jpn name: Kurakakeutsubo]

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K. Fujiwara and H. Motomura

Kamohara (1957, as Gymnothorax petelli): 1 unregistered specimen, 72 mm TL, Somachi. Kamohara & Yamakawa (1965, as Gymnothorax petelli): 1 unregistered specimen, 100 mm TL, Araki; 3 unregistered specimens, 97–508 mm TL, Somachi.
Gymnothorax thyrsoideus (Richardson, 1845)[Jpn name: Sabiutsubo]Kamohara & Yamakawa (1965): 1 unregistered specimen, 75 mm TL, Tekuzuku; 1 unregistered specimen,
240 mm TL, Somachi. This study: KAUM–I. 14742, 159.9 mm TL, Kikai Island; KAUM–I. 135137, 299.4mm TL, Shiomichi.
Gymnothorax undulatus (Lacepède, 1803)[Jpn name: Namiutsubo]Kamohara (1957): 3 unregistered specimens, 205–530 mm TL, Onotsu. Kamohara & Yamakawa (1965): 1
unregistered specimen, 195 mm TL, Araki.
Uropterygius micropterus (Bleeker, 1852)[Jpn name: Amikikaiutsubo]
KAUM–I. 135136, 242.9 mm TL, KAUM–I. 135139, 36.8 mm TL, Shiomichi.
FAMILY OPHICHTHIDAE
Leiuranus semicinctus (Lay & Bennett, 1839)[Jpn name: Soraumihebi]
Kamohara (1957): 1 unregistered specimen, 190 mm TL, Wan; 2 unregistered specimens, 198–360 mm TL,
Somachi; 1 unregistered specimen, 408 mm TL (as Sphagebranchus cinctus), Somachi. Kamohara & Yamakawa (1965): 2 unregistered specimens, 190–240 mm TL, Somachi. Hatooka (2013b): Kikai Island.
Myrichthys colubrinus (Boddaert, 1781)[Jpn name: Shimaumihebi]
KAUM–I. 147626, 325.2 mm TL, Kamikatetsu.
Myrichthys maculosus (Cuvier, 1816)[Jpn name: Moyomongaradoshi]
Kamohara (1957): 4 unregistered specimens, 125–545 mm TL, Wan; 3 unregistered specimens, 235–410 mm
TL, Somachi. Kamohara & Yamakawa (1965): 1 unregistered specimen, 275 mm TL, Somachi. Hata et al.
(2015a): KAUM–I. 1982, 169.8 mm TL, Kikai Island. Hibino (2018b): KAUM–I. 1982, 169.8 mm TL, Kikai Island. Hibino (2019b): KAUM–I. 1982, 169.8 mm TL, Kikai Island. This study: KAUM–I. 1982, 169.8mm TL, Kikai Island; KAUM–I. 134705, 460.3 mm TL, KUN-P 55685, 376.0 mm TL, Shiomichi.
Scolecenchelys gymnota (Bleeker, 1857)[Jpn name: Mimizuanago]
Kamohara (1957, as Muraenichthys gymnotus): 1 unregistered specimen, 230 mm TL, Wan. Kamohara &Yamakawa (1965, as Muraenichthys gymnotus): 1 unregistered specimen, 122 mm TL, Somachi.
Scolecenchelys laticaudata (Ogilby, 1897)[Jpn name: Futomimizuanago]
Hibino et al. (2013): BSKU 5498, 223 mm TL, Wan; KAUM–I. 2381, 152.1 mm TL, Kikai Island. Hibino& Kimura (2015): BSKU 5498, 223 mm TL, Wan; KAUM–I. 2381, 152.1 mm TL, Kikai Island. Hibino(2018b): KAUM–I. 2381, 152.1 mm TL, Kikai Island. Hibino (2019b): KAUM–I. 2381, 152.1 mm TL, Kikai Island. This study: KAUM–I. 2381, 152.1 mm TL, Kikai Island.

Marine and freshwater fishes of Kikai Island

7

FAMILY CONGRIDAE
Conger cinereus Rüppell, 1871
[Jpn name: Kirianago]
Kamohara & Yamakawa (1965): 2 unregistered specimens, 386–435 mm TL, Somachi. This study: KAUM–
I. 135025, 744.0 mm TL, Shiomichi.
Conger jordani Kanazawa, 1958
[Jpn name: Kuroanago]
KUN-P 55736, 68.4 mm TL, Aden.
Conger myriaster (Brevoort, 1856)[Jpn name: Maanago]
Kamohara & Yamakawa (1968a): 1 unregistered specimen, 117 mm TL, Kikai Island. Hatooka (2013c): Kikai Island.
ORDER CLUPEIFORMESFAMILY ENGRAULIDAE
Encrasicholina sp. (unidentified)[Jpn name: N/A]
KAUM–I. 146639, 12.8 mm SL, Kamikatetsu.
ORDER CYPRINIFORMESFAMILY CYPRINIDAE
Carassius auratus (Linnaeus, 1758)[Jpn name: Kingyo]
Nakajima et al. (2020): KMNH VR 100259, 61.5 mm SL, Urahara.
Carassius sp.
[Jpn name: Ginbuna]Hayashi (1955): YCM-P 31033, 3 specimens, 40.5–51.3 mm SL, Araki. Matsunuma (2018a): KPM-NI
21069, 40.5 mm SL, KPM-NI 21070, 41.7 mm SL, Kikai Island. Matsunuma (2019a): KPM-NI 21070, 41.7mm SL, Kikai Island. Nakajima et al. (2020, as Carassius langsdorfii): KMNH VR 100258, 65.1 mm SL,
Urahara.
FAMILY COBITIDAE
Misgurnus sp. IR[Jpn name: Shinobidojo]
Hayashi (1955, as Misgurnus anguillicaudatus): YCM-P 31002, 78.5 mm SL, Araki.
ORDER SILURIFORMESFAMILY PLOTOSIDAE
Plotosus japonicus Yoshino & Kishimoto, 2008
[Jpn name: Gonzui]
KAUM–I. 72336, 17.6 mm SL, KAUM–I. 72337, 17.8 mm SL, Onotsu; KAUM–I. 134650, 69.1 mm SL,
KAUM–I. 134651, 73.4 mm SL, KAUM–I. 134652, 83.9 mm SL, Nakasato.
ORDER AULOPIFORMESFAMILY SYNODONTIDAE
Saurida gracilis (Quoy & Gaimard, 1824)[Jpn name: Madaraeso]

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K. Fujiwara and H. Motomura

Kamohara (1957): 1 unregistered specimen, 207 mm TL, Somachi; 2 unregistered specimens, 90–119 mm
TL, Wan. Kamohara & Yamakawa (1968a): 2 unregistered specimens, 127–180 mm TL, Kikai Island. This
study: KAUM–I. 134528, 107.0 mm SL, KAUM–I. 134529, 89.0 mm SL, KAUM–I. 134718, 103.7 mm SL,
Akaren; KAUM–I. 134677, 127.6 mm SL, KAUM–I. 134687, 44.7 mm SL, Shiomichi; KAUM–I. 146714,
69.8 mm SL, Kamikatetsu; KUN-P 55681, 97.6 mm SL, Onotsu.
Synodus dermatogenys Fowler, 1912
[Jpn name: Minamiakaeso]
KAUM–I. 134713, 170.9 mm SL, KAUM–I. 134714, 152.1 mm SL, KUN-P 55655, 94.0 mm SL, KUN-P
55678, 160.2 mm SL, Nakasato; KAUM–I. 134719, 117.9 mm SL, Akaren.
Synodus variegatus (Lacepède, 1803)[Jpn name: Hitosujieso]
Kamohara (1957): 1 unregistered specimen, 232 mm TL, Wan. This study: KAUM–I. 146792, 104.4 mm SL,
Akaren.
ORDER OPHIDIIFORMESFAMILY CARAPIDAE
Encheliophis vermicularis Müller, 1842
[Jpn name: Amamikakureuo]
Kamohara & Yamakawa (1965): 3 unregistered specimens, 83–113 mm TL, Wan.
FAMILY DINEMATICHTHYIDAEAlionematichthys sp. (unidentified)[Jpn name: N/A]
KAUM–I. 146625, 49.6 mm SL, Kamikatetsu.
Diancistrus fuscus (Fowler, 1946)[Jpn name: Sangoitachiuo]
Kamohara & Yamakawa (1965, as Brotulina fusca): 7 unregistered specimens, 27–108 mm TL, Somachi.
ORDER LOPHIIFORMESFAMILY ANTENNARIIDAE
Antennarius randalli Allen, 1970
[Jpn name: Himehiratakaeruanko]
Yoshida & Motomura (2016): KAUM–I. 72320, 12.3 mm SL, Shiomichi. Senou (2018): KAUM–I. 72320,
12.3 mm SL, Shiomichi. Senou (2019): KAUM–I. 72320, 12.3 mm SL, Shiomichi. This study: KAUM–I.
72320, 12.3 mm SL, Shiomichi.
Antennarius striatus (Shaw, 1794)[Jpn name: Kaeruanko]
Kamohara (1957, as Antennarius tridens): 4 unregistered specimens, 22–45 mm TL, Wan. Senou (2013a):
Kikai Island.
Histrio histrio (Linnaeus, 1758)[Jpn name: Hanaokoze]
Kamohara & Yamakawa (1968b, as Histrio histrio histrio): 1 unregistered specimen, 17 mm TL, Kikai Island.

Marine and freshwater fishes of Kikai Island

9

ORDER BERYCIFORMESFAMILY HOLOCENTRIDAE
Myripristis berndti Jordan & Evermann, 1903
[Jpn name: Akamatsukasa]
KAUM–I. 72322, 97.2 mm SL, KAUM–I. 135163, 114.7 mm SL, KAUM–I. 136732, 80.4 mm SL, Akaren;
KAUM–I. 146821, 92.2 mm SL, KAUM–I. 146845, 121.9 mm SL, Somachi.
Myripristis greenfieldi Randall & Yamakawa, 1996
[Jpn name: Tsumarimatsukasa]
KAUM–I. 134558, 85.5 mm SL, KAUM–I. 134559, 67.9 mm SL, KAUM–I. 134560, 49.9 mm SL, KAUM–
I. 134561, 49.8 mm SL, KAUM–I. 146779, 95.4 mm SL, KAUM–I. 146780, 110.9 mm SL, KAUM–I.
146781, 82.0 mm SL, KAUM–I. 146847, 81.1 mm SL, KAUM–I. 146848, 77.3 mm SL, KUN-P 55660, 76.8mm SL, KUN-P 55670, 57.5 mm SL, Akaren; KUN-P 55716, 37.7 mm SL, KUN-P 55775, 45.9 mm SL, Somachi.
Myripristis kuntee Valenciennes, 1831
[Jpn name: Kurobimatsukasa]
KAUM–I. 134555, 76.0 mm SL, KAUM–I. 146778, 85.5 mm SL, Akaren; KAUM–I. 146846, 85.5 mm SL,
Somachi.
Myripristis murdjan (Forsskål, 1775)[Jpn name: Yogorematsukasa]
KAUM–I. 72515, 102.4 mm SL, KAUM–I. 134556, 95.8 mm SL, KAUM–I. 134557, 69.1 mm SL, Akaren;
KAUM–I. 136724, 43.6 mm SL, Shiomichi.
Neoniphon sammara (Forsskål, 1775)[Jpn name: Ukeguchiittodai]
KAUM–I. 15175, 102.4 mm SL, KAUM–I. 15204, 100.0 mm SL, KAUM–I. 15231, 86.3 mm SL, Kikai Island; KAUM–I. 134683, 35.9 mm SL, Shiomichi; KAUM–I. 146559, 54.7 mm SL, KAUM–I. 146748, 54.0mm SL, Keraji; KAUM–I. 146761, 57.9 mm SL, Tekuzuku.
Ostichthys kaianus (Günther, 1880)[Jpn name: Kaiebisu]
Nakae (2018): KAUM–I. 68646, 242.7 mm SL, Kikai Island. Nakae (2019): KAUM–I. 68646, 242.7 mm
SL, Kikai Island. This study: KAUM–I. 68646, 242.7 mm SL, Kikai Island.
Sargocentron diadema (Lacepède, 1802)[Jpn name: Nijiebisu]
Kamohara (1957, as Holocentrus diadema): 1 unregistered specimen, 80 mm TL, Somachi.
Sargocentron ittodai (Jordan & Fowler, 1902)[Jpn name: Teriebisu]
KAUM–I. 8108, 81.4 mm SL, Urahara; KAUM–I. 15213, 92.9 mm SL, KAUM–I. 15234, 74.1 mm SL,
KAUM–I. 15237, 92.0 mm SL, KAUM–I. 15256, 50.3 mm SL, Kikai Island; KAUM–I. 72310, 106.8 mm
SL, KAUM–I. 136702, 92.8 mm SL, Shiomichi; KAUM–I. 72516, 112.5 mm SL, KAUM–I. 72517, 112.3mm SL, KAUM–I. 72518, 98.4 mm SL, KAUM–I. 72519, 91.4 mm SL, KAUM–I. 134554, 64.8 mm SL,
KAUM–I. 136731, 114.9 mm SL, KAUM–I. 146803, 98.5 mm SL, Akaren; KAUM–I. 134798, 56.6 mm SL,
KAUM–I. 147642, 91.1 mm SL, Keraji; KAUM–I. 146509, 60.6 mm SL, Onotsu; KAUM–I. 146822, 90.2
mm SL, Somachi.

10

K. Fujiwara and H. Motomura

Sargocentron praslin (Lacepède, 1802)[Jpn name: Kurobiebisu]
KAUM–I. 134682, 33.7 mm SL, KAUM–I. 136703, 82.7 mm SL, Shiomichi; KAUM–I. 146266, 30.7 mm
SL, Ikeji.
Sargocentron punctatissimum (Cuvier, 1829)[Jpn name: Hoshiebisu]
KAUM–I. 8113, 83.3 mm SL, Urahara; KAUM–I. 15214, 74.2 mm SL, KAUM–I. 15263, 46.6 mm SL, Kikai Island; KAUM–I. 135171, 115.3 mm SL, KAUM–I. 136721, 60.2 mm SL, KAUM–I. 136722, 56.7 mm
SL, KAUM–I. 136723, 41.7 mm SL, KAUM–I. 136996, 123.2 mm SL, Shiomichi; KAUM–I. 146263, 94.9mm SL, Ikeji; KUN-P 55714, 45.5 mm SL, KUN-P 55715, 64.0 mm SL, KUN-P 55725, 44.3 mm SL, Somachi.
ORDER GASTEROSTEIFORMESFAMILY CENTRISCIDAE
Aeoliscus strigatus (Günther, 1861)[Jpn name: Hekoayu]
KAUM–I. 134840, 125.5 mm SL, KAUM–I. 134841, 130.8 mm SL, KAUM–I. 134842, 129.9 mm SL,
KAUM–I. 134843, 129.4 mm SL, KAUM–I. 134844, 132.0 mm SL, KAUM–I. 134845, 135.4 mm SL,
KAUM–I. 134846, 130.1 mm SL, KUN-P 55727, 134.5 mm SL, KUN-P 55727, 134.5 mm SL, KUN-P
55727, 134.5 mm SL, KUN-P 55727, 134.5 mm SL, KUN-P 55728, 105.5 mm SL, KUN-P 55729, 131.3mm SL, KUN-P 55730, 129.8 mm SL, KUN-P 55731, 63.4 mm SL, KUN-P 55732, 139.7 mm SL, KUN-P
55767, 131.3 mm SL, KUN-P 55768, 138.5 mm SL, KUN-P 55769, 128.5 mm SL, KUN-P 55770, 125.6mm SL, KUN-P 55771, 147.0 mm SL, KUN-P 56303, 126.5 mm SL, KUN-P 56304, 106.1 mm SL, KUN-P
56305, 117.5 mm SL, KUN-P 56306, 136.5 mm SL, KUN-P 56307, 139.8 mm SL, KUN-P 56308, 121.8mm SL, KUN-P 56309, 135.1 mm SL, KUN-P 56311, 125.3 mm SL, KUN-P 56312, 94.2 mm SL, KUN-P
56313, 95.8 mm SL, KUN-P 56314, 129.6 mm SL, KUN-P 56315, 112.5 mm SL, KUN-P 56316, 112.7mm SL, KUN-P 56317, 113.2 mm SL, KUN-P 56318, 119.4 mm SL, KUN-P 56319, 126.1 mm SL, KUN-P
56320, 133.6 mm SL, KUN-P 56321, 130.1 mm SL, KUN-P 56322, 94.9 mm SL, KUN-P 56323, 110.7 mm
SL, KUN-P 56467, 115.9 mm SL, KUN-P 56468, 113.0 mm SL, KUN-P 56469, 94.8 mm SL, Somachi.
FAMILY SYNGNATHIDAE
Corythoichthys haematopterus (Bleeker, 1851)[Jpn name: Ishiyoji]
KAUM–I. 134761, 174.1 mm SL, Keraji.
Corythoichthys schultzi Herald, 1953
[Jpn name: Kuchinagaishiyoji]
KAUM–I. 146533, 77.8 mm SL, Shiomichi.
Choeroichthys sculptus (Günther, 1870)[Jpn name: Chigoyoji]
Kamohara (1957): 1 unregistered specimen, 66 mm TL, Wan. Kamohara & Yamakawa (1965): 1 unregistered specimen, 65 mm TL, Araki.
Doryrhamphus (Doryrhamphus) excisus excisus Kaup, 1856
[Jpn name: Hibashiyoji]
Kamohara (1957, as Doryrhamphus melanopleura): 1 unregistered specimen, 65 mm TL, Somachi; 4 unreg-

Marine and freshwater fishes of Kikai Island

11

istered specimens, 48–65 mm TL, Wan. Kamohara & Yamakawa (1965, as Doryrhamphus melanopleura): 2
unregistered specimens, 48–53 mm TL, Tekuzuku; 3 unregistered specimens, 52–62 mm TL, Araki; 3 unregistered specimens, 51–58 mm SL, Somachi. Matsunuma (2018b): KAUM–I. 72317, 49.3 mm SL, Shiomichi.
Matsunuma (2019b): KAUM–I. 72317, 49.3 mm SL, Shiomichi. This study: KAUM–I. 72317, 49.3 mm
SL, Shiomichi; KAUM–I. 134613, 51.2 mm SL, Onotsu; KAUM–I. 146684, 33.0 mm SL, Araki; KAUM–I.
146757, 33.1 mm SL, KAUM–I. 146758, 24.2 mm SL, Keraji.
Doryrhamphus (Dunckerocampus) dactyliophorus (Bleeker, 1853)[Jpn name: Kasumioiran-yoji]
KAUM–I. 146478, 105.5 mm SL, Tekuzuku.
Halicampus brocki (Herald, 1953)[Jpn name: Nokogiriumiyakko]
KAUM–I. 134762, 88.8 mm SL, Keraji.
Micrognathus andersonii (Bleeker, 1858)[Jpn name: Kammuriyoji]
KAUM–I. 146760, 51.0 mm SL, Keraji.
Microphis (Oostethus) brachyurus brachyurus (Bleeker, 1854)[Jpn name: Tenguyoji]
KAUM–I. 7630, 127.0 mm SL, Urahara; KAUM–I. 135024, 101.3 mm SL, Shiramizu; KAUM–I. 135154,
84.5 mm SL, Shitooke.
Phoxocampus belcheri (Kaup, 1856)[Jpn name: Boyoji]
KAUM–I. 134830, 55.7 mm SL, Nakasato; KAUM–I. 146727, 59.8 mm SL, Tekuzuku.
ORDER MUGILIFORMESFAMILY MUGILIDAE
Chelon lauvergnii (Eydoux & Souleyet, 1850)[Jpn name: Sesujibora]
Kamohara (1957, as Liza carinata): many unregistered specimens, 68–180 mm TL, Kikai Island (including
Amami-oshima Island).
Chelon macrolepis (Smith, 1846)[Jpn name: Kobora]
KAUM–I. 134508, 52.8 mm SL, KAUM–I. 134509, 58.1 mm SL, KAUM–I. 134510, 51.3 mm SL, KAUM–
I. 134511, 60.7 mm SL, Akaren; KAUM–I. 134570, 100.9 mm SL, Shitooke; KUN-P 55709, 110.0 mm SL,
KUN-P 55713, 72.8 mm SL, Somachi; KUN-P 55790, 130.6 mm SL, Tekuzuku.
Crenimugil crenilabis (Forsskål, 1775)[Jpn name: Furaibora]
Kamohara & Yamakawa (1965): many unregistered specimens, 83–155 mm TL, Somachi. This study:
KUN-P 55721, 38.4 mm SL, KUN-P 55722, 24.2 mm SL, KUN-P 55726, 28.3 mm SL, Somachi.
Crenimugil seheli (Forsskål, 1775)[Jpn name: Taiwammenada]12

K. Fujiwara and H. Motomura

KAUM–I. 134500, 32.3 mm SL, KAUM–I. 134501, 32.2 mm SL, KAUM–I. 134502, 30.6 mm SL, KAUM–
I. 134503, 34.0 mm SL, KAUM–I. 134516, 30.8 mm SL, KAUM–I. 134517, 30.4 mm SL, KAUM–I.
134518, 28.8 mm SL, KAUM–I. 134519, 30.4 mm SL, KAUM–I. 134520, 28.5 mm SL, Akaren; KAUM–I.
135160, 30.8 mm SL, KAUM–I. 135161, 30.3 mm SL, Shitooke; KAUM–I. 136727, 32.1 mm SL, Shiomichi.
Ellochelon vaigiensis (Quoy & Gaimard, 1825)[Jpn name: Onibora]
KAUM–I. 134505, 27.9 mm SL, KAUM–I. 134506, 26.6 mm SL, KAUM–I. 134507, 29.2 mm SL, KAUM–
I. 134521, 22.4 mm SL, KAUM–I. 134522, 24.8 mm SL, KAUM–I. 134523, 22.7 mm SL, KAUM–I.
134524, 26.7 mm SL, Akaren; KAUM–I. 134674, 97.4 mm SL, Shiomichi; KUN-P 55723, 23.4 mm SL,
KUN-P 55724, 39.6 mm SL, Somachi.
Mugil cephalus cephalus Linnaeus, 1758
[Jpn name: Bora]
Kamohara (1957, as Mugil cephalus): many unregistered specimens, 90–155 mm TL, Kikai Island (including Amami-oshima Island). Kamohara & Yamakawa (1968a, as Mugil cephalus): 5 unregistered specimens,
45–50 mm TL, Kikai Island. This study: KAUM–I. 135156, 22.4 mm SL, KAUM–I. 135157, 24.4 mm SL,
KAUM–I. 135158, 21.0 mm SL, Shitooke.
Plicomugil labiosus (Valenciennes, 1836)[Jpn name: Waniguchibora]
KAUM–I. 134504, 22.5 mm SL, KAUM–I. 134512, 26.4 mm SL, KAUM–I. 134513, 24.2 mm SL, KAUM–
I. 134514, 25.0 mm SL, KAUM–I. 134515, 27.3 mm SL, Akaren; KAUM–I. 146574, 60.5 mm SL, KAUM–
I. 146575, 26.4 mm SL, Somachi.
ORDER ATHERINIFORMESFAMILY ATHERINIDAE
Hypoatherina lunata Sasaki & Kimura, 2012
[Jpn name: Okinawatogoro]
KAUM–I. 134689, 68.8 mm SL, KAUM–I. 136728, 25.8 mm SL, Shiomichi.
FAMILY NOTOCHEIRIDAE
Iso flosmaris Jordan & Starks, 1901
[Jpn name: Naminohana]
Kamohara & Yamakawa (1968b): 11 unregistered specimens, 25–36 mm TL, Kikai Island. Senou (2013b):
Kikai Island.
ORDER CYPRINODONTIFORMESFAMILY POECILIIDAE
Gambusia affinis affinis (Baird & Girard, 1853)[Jpn name: Kadayashi]
Nakajima et al. (2020, as Gambusia affinis): Urahara.
Poecilia reticulata Peters, 1859
[Jpn name: Guppi]
KAUM–I. 4213, 2 specimens, 7.5–17.9 mm SL, KAUM–I. 4215, 40.5 mm SL, KAUM–I. 4216, 27.3 mm
SL, KAUM–I. 4217, 32.3 mm SL, KAUM–I. 4218, 25.6 mm SL, KAUM–I. 4219, 25.7 mm SL, KAUM–
I. 4220, 14.6 mm SL, KAUM–I. 4221, 15.5 mm SL, KAUM–I. 4222, 16.4 mm SL, KAUM–I. 8146, 31.4

Marine and freshwater fishes of Kikai Island

13mm SL, KAUM–I. 8147, 21.5 mm SL, KAUM–I. 137400–137499, 18.2–22.2 mm SL, KAUM–I. 137500–
137599, 19.2–30.2 mm SL, KAUM–I. 137600–137699, 15.0–23.7 mm SL, KAUM–I. 137700–137799,
14.6–24.0 mm SL, KAUM–I. 137800–137899, 13.3–33.5 mm SL, KAUM–I. 137900–137999, 11.5–30.7mm SL, KAUM–I. 137800–138099, 10.8–27.3 mm SL, Urahara; KAUM–I. 135000, 20.7 mm SL, KAUM–
I. 135000, 20.7 mm SL, KAUM–I. 135000, 20.7 mm SL, KAUM–I. 135000, 20.7 mm SL, KAUM–I.
135000, 20.7 mm SL, KAUM–I. 135000, 20.7 mm SL, KAUM–I. 135000, 20.7 mm SL, KAUM–I. 135000,
20.7 mm SL, KAUM–I. 135000, 20.7 mm SL, KAUM–I. 135000, 20.7 mm SL, KAUM–I. 135000, 20.7mm SL, KAUM–I. 135001, 20.1 mm SL, KAUM–I. 135002, 31.2 mm SL, KAUM–I. 135003, 19.2 mm SL,
KAUM–I. 135004, 18.6 mm SL, KAUM–I. 135005, 20.7 mm SL, KAUM–I. 135006, 18.4 mm SL, KAUM–
I. 135007, 18.6 mm SL, KAUM–I. 135008, 18.1 mm SL, KAUM–I. 135009, 18.4 mm SL, KAUM–I.
135010, 17.0 mm SL, KAUM–I. 135011, 20.0 mm SL, KAUM–I. 135012, 15.9 mm SL, KAUM–I. 135013,
17.9 mm SL, KAUM–I. 135014, 17.1 mm SL, KAUM–I. 135015, 16.0 mm SL, KAUM–I. 135016, 20.3mm SL, KAUM–I. 135017, 28.4 mm SL, KAUM–I. 135018, 23.9 mm SL, KAUM–I. 135019, 21.7 mm SL,
KAUM–I. 135020, 21.6 mm SL, KAUM–I. 135021, 25.9 mm SL, KAUM–I. 135022, 24.5 mm SL, KAUM–
I. 135023, 19.3 mm SL, KAUM–I. 135155, 22.1 mm SL, KAUM–I. 136500–136599, 17.2–21.7 mm SL,
KAUM–I. 136600–136699, 17.4–20.7 mm SL, KAUM–I. 136809, 12.5 mm SL, KAUM–I. 136810, 11.4mm SL, KAUM–I. 136811, 13.5 mm SL, KAUM–I. 136812, 11.0 mm SL, KAUM–I. 136813, 12.0 mm SL,
KAUM–I. 136814, 12.7 mm SL, KAUM–I. 136815, 11.9 mm SL, KAUM–I. 136816, 13.2 mm SL, KAUM–
I. 136817, 11.5 mm SL, KAUM–I. 136818, 11.9 mm SL, KAUM–I. 136819, 15.0 mm SL, KAUM–I.
136820, 12.5 mm SL, KAUM–I. 136821, 14.9 mm SL, KAUM–I. 136822, 12.5 mm SL, KAUM–I. 136823,
11.9 mm SL, KAUM–I. 136824, 14.1 mm SL, KAUM–I. 136825, 16.2 mm SL, KAUM–I. 136826, 14.1mm SL, KAUM–I. 136827, 13.1 mm SL, KAUM–I. 136828, 12.8 mm SL, KAUM–I. 136829, 12.6 mm SL,
KAUM–I. 136830, 11.4 mm SL, KAUM–I. 136831, 15.5 mm SL, KAUM–I. 136832, 13.2 mm SL, KAUM–
I. 136833, 13.4 mm SL, KAUM–I. 136834, 15.6 mm SL, KAUM–I. 136835, 13.5 mm SL, KAUM–I.
136836, 13.0 mm SL, KAUM–I. 136837, 15.0 mm SL, KAUM–I. 136838, 15.8 mm SL, KAUM–I. 136839,
13.0 mm SL, KAUM–I. 136840, 12.9 mm SL, KAUM–I. 136841, 13.3 mm SL, KAUM–I. 136842, 10.8mm SL, KAUM–I. 136843, 10.7 mm SL, KAUM–I. 136844, 10.9 mm SL, KAUM–I. 136845, 12.5 mm SL,
KAUM–I. 136846, 13.5 mm SL, KAUM–I. 136847, 12.1 mm SL, KAUM–I. 136848, 12.3 mm SL, KAUM–
I. 136849, 11.8 mm SL, KAUM–I. 136850, 13.5 mm SL, KAUM–I. 136851, 15.3 mm SL, KAUM–I.
136852, 14.0 mm SL, KAUM–I. 136853, 15.6 mm SL, KAUM–I. 136854, 14.7 mm SL, KAUM–I. 136855,
13.1 mm SL, KAUM–I. 136856, 12.0 mm SL, KAUM–I. 136857, 14.2 mm SL, KAUM–I. 136858, 14.6mm SL, KAUM–I. 136859, 12.5 mm SL, KAUM–I. 136860, 14.0 mm SL, KAUM–I. 136861, 12.5 mm SL,
KAUM–I. 136862, 14.0 mm SL, KAUM–I. 136863, 13.0 mm SL, KAUM–I. 136864, 14.6 mm SL, KAUM–
I. 136865, 14.4 mm SL, KAUM–I. 136866, 12.6 mm SL, KAUM–I. 136867, 13.8 mm SL, KAUM–I.
136868, 12.7 mm SL, KAUM–I. 136869, 12.6 mm SL, KAUM–I. 136870, 12.4 mm SL, KAUM–I. 136871,
15.5 mm SL, KAUM–I. 136872, 13.8 mm SL, KAUM–I. 136873, 14.8 mm SL, KAUM–I. 136874, 13.4mm SL, KAUM–I. 136875, 13.7 mm SL, KAUM–I. 136876, 15.1 mm SL, KAUM–I. 136877, 12.1 mm SL,
KAUM–I. 136878, 12.8 mm SL, KAUM–I. 136879, 13.8 mm SL, KAUM–I. 136880, 13.8 mm SL, KAUM–
I. 136881, 12.1 mm SL, KAUM–I. 136882, 12.2 mm SL, KAUM–I. 136883, 12.8 mm SL, KAUM–I.
136884, 13.4 mm SL, KAUM–I. 136885, 13.6 mm SL, KAUM–I. 136886, 13.5 mm SL, KAUM–I. 136887,
13.2 mm SL, KAUM–I. 136888, 13.1 mm SL, KAUM–I. 136889, 12.8 mm SL, KAUM–I. 136890, 12.5mm SL, KAUM–I. 136891, 12.4 mm SL, KAUM–I. 136892, 11.9 mm SL, KAUM–I. 136893, 12.4 mm SL,
KAUM–I. 136894, 8.2 mm SL, KAUM–I. 136895, 6.9 mm SL, KAUM–I. 136896, 15.4 mm SL, KAUM–I.
136897, 11.0 mm SL, KAUM–I. 136898, 11.9 mm SL, KAUM–I. 136899, 11.5 mm SL, KAUM–I. 137000–
137099, 13.5–26.2 mm SL, KAUM–I. 137100–137199, 10.8–30.3 mm SL, KAUM–I. 137200–137299,
10.4–17.6 mm SL, KAUM–I. 137300–137399, 9.4–14.3 mm SL, KUN-P 55779, 19.0 mm SL, KUN-P
55780, 22.7 mm SL, KUN-P 55781, 19.1 mm SL, KUN-P 55782, 17.2 mm SL, KUN-P 55783, 17.6 mm SL,
KUN-P 55784, 18.2 mm SL, KUN-P 55785, 19.4 mm SL, KUN-P 55786, 31.0 mm SL, KUN-P 55787, 32.6mm SL, KUN-P 55789, 19.5 mm SL, KUN-P 55791, 18.7 mm SL, KUN-P 56441, 16.0 mm SL, KUN-P
56442, 19.7 mm SL, KUN-P 56443, 20.0 mm SL, KUN-P 56444, 18.2 mm SL, KUN-P 56445, 16.6 mm SL,
KUN-P 56446, 15.9 mm SL, KUN-P 56447, 15.2 mm SL, KUN-P 56448, 13.3 mm SL, KUN-P 56449, 14.7

14

K. Fujiwara and H. Motomura

mm SL, KUN-P 56450, 13.6 mm SL, KUN-P 56451, 14.9 mm SL, KUN-P 56452, 15.8 mm SL, KUN-P
56453, 12.6 mm SL, KUN-P 56454, 16.5 mm SL, KUN-P 56455, 12.6 mm SL, KUN-P 56456, 8.0 mm SL,
KUN-P 56457, 13.9 mm SL, KUN-P 56458, 15.7 mm SL, KUN-P 56459, 15.8 mm SL, KUN-P 56460, 14.9mm SL, KUN-P 56461, 13.1 mm SL, KUN-P 56462, 12.3 mm SL, KUN-P 56463, 11.6 mm SL, KUN-P
56464, 11.2 mm SL, KUN-P 56465, 15.8 mm SL, KUN-P 56466, 13.2 mm SL, Isaneku.
ORDER BELONIFORMESFAMILY ADRIANICHTHYIDAE
Oryzias latipes (Temminck & Schlegel, 1846)[Jpn name: Minamimedaka]
Hayashi (1995): YCM-P 31004, 22 specimens, 22.8–32.9 mm SL, Araki. Senou (2013c): Kikai Island. This
study: KAUM–I. 135140, 25.6 mm SL, KAUM–I. 135141, 26.4 mm SL, KAUM–I. 135142, 24.0 mm SL,
KAUM–I. 135143, 24.6 mm SL, KAUM–I. 135144, 25.4 mm SL, KAUM–I. 135145, 25.6 mm SL, KAUM–
I. 135146, 26.8 mm SL, KAUM–I. 135147, 26.3 mm SL, KAUM–I. 135148, 29.8 mm SL, KAUM–I.
135149, 28.1 mm SL, Shiomichi.
FAMILY HEMIRAMPHIDAE
Hyporhamphus dussumieri (Valenciennes, 1847)[Jpn name: Marusayori]
KAUM–I. 134678, 116.7 mm SL, KAUM–I. 136726, 63.4 mm SL, KAUM–I. 136995, 169.0 mm SL,
KUN-P 55755, 78.1 mm SL, Shiomichi.
FAMILY EXOCOETIDAE
Cheilopogon agoo agoo (Temminck & Schlegel, 1846)[Jpn name: Tobiuo]
KAUM–I. 27153, 224.3 mm SL, Kikai Island.
FAMILY BELONIDAE
Tylosurusacus melanotus (Bleeker, 1850)[Jpn name: Tenjikudatsu]
Kamohara (1957): 1 unregistered specimen, 475 mm TL, Onotsu.
ORDER SCORPAENIFORMESFAMILY SCORPAENIDAE
Caracanthus maculatus (Gray, 1831)[Jpn name: Dangokoze]
KAUM–I. 146685, 31.4 mm SL, Araki.
Caracanthus unipinna (Gray, 1831)[Jpn name: Watagedangokoze]
KAUM–I. 146468, 23.5 mm SL, KAUM–I. 146484, 25.8 mm SL, Tekuzuku.
Dendrochirus zebra (Cuvier, 1829)[Jpn name: Kirimmino]
KAUM–I. 146600, 47.3 mm SL, Tekuzuku; KAUM–I. 146666, 25.5 mm SL, Araki.
Parascorpaena mossambica (Peters, 1855)[Jpn name: Nettaifusakasago]
KAUM–I. 134608, 74.3 mm SL, Onotsu.

Marine and freshwater fishes of Kikai Island

15

Pterois radiata Cuvier, 1829
[Jpn name: Kimiokoze]
Kamohara (1957): 3 unregistered specimens, 43–80 mm TL, Wan. Kamohara & Yamakawa (1965): 1 unregistered specimen, 40 mm TL, Araki; 4 unregistered specimens, 48–84 mm SL, Somachi. This study: KAUM–
I. 147641, 134.9 mm SL, Keraji.
Pterois volitans (Linnaeus, 1758)[Jpn name: Hanaminokasago]
Kamohara (1957): 1 unregistered specimen, 67 mm TL, Wan. This study: KAUM–I. 146265, 73.0 mm SL,
Ikeji; KAUM–I. 146665, 57.5 mm SL, Araki; KAUM–I. 147622, 68.1 mm SL, Kamikatetsu.
Scorpaenodes guamensis (Quoy & Gaimard, 1824)[Jpn name: Guamukasago]
Motomura (2018): KAUM–I. 72333, 33.1 mm SL, Onotsu. Motomura (2019a): KAUM–I. 72333, 33.1 mm
SL, Onotsu. This study: KAUM–I. 36034, 31.0 mm SL, KAUM–I. 36035, 19.2 mm SL, Akaren; KAUM–I.
72333, 33.1 mm SL, KAUM–I. 134609, 47.8 mm SL, Onotsu; KAUM–I. 134647, 45.4 mm SL, KAUM–I.
134825, 20.2 mm SL, Nakasato; KAUM–I. 134695, 49.8 mm SL, KAUM–I. 136700, 73.2 mm SL, KAUM–
I. 136701, 64.7 mm SL, KAUM–I. 136713, 56.2 mm SL, Shiomichi; KAUM–I. 135031, 51.5 mm SL,
KAUM–I. 135088, 27.6 mm SL, Araki; KAUM–I. 146259, 59.0 mm SL, KAUM–I. 146260, 28.4 mm SL,
Ikeji; KAUM–I. 146632, 62.0 mm SL, Kamikatetsu; KUN-P 55717, 82.6 mm SL, Somachi.
Scorpaenodes varipinnis Smith, 1957
[Jpn name: Seboshiisokasago]
Motomura (2018): KAUM–I. 72371, 39.1 mm SL, Keraji. Motomura (2019a): KAUM–I. 72371, 39.1 mm
SL, Keraji. This study: KAUM–I. 72371, 39.1 mm SL, Keraji.
Scorpaenopsis diabolus (Cuvier, 1829)[Jpn name: Niraikasago]
KAUM–I. 134822, 169.8 mm SL, Nakasato.
Scorpaenopsis neglecta Heckel, 1839
[Jpn name: Satsumakasago]
Kamohara (1957, as Scorpaenopsis gibbosa): 2 unregistered specimens, 60–62 mm TL, Wan. This study:
KAUM–I. 146584, 47.6 mm SL, Kamikatetsu; KAUM–I. 146745, 54.9 mm SL, Keraji; KUN-P 55676, 57.6mm SL, KUN-P 55677, 77.9 mm SL, Somachi.
Scorpaenopsis macrochir Ogilby, 1910
[Jpn name: Marusubekasago]
Motomura (2018): KAUM–I. 72370, 67.0 mm SL, Keraji. Motomura (2019a): KAUM–I. 72370, 67.0 mm
SL, Keraji. This study: KAUM–I. 72370, 67.0 mm SL, Keraji.
Scorpaenopsis possi Randall & Eschmeyer, 2002
[Jpn name: Mimitogeonikasago]
KPM-NI 26344, 105.9 mm SL, Kikai Island.
Scorpaenopsis ramaraoi Randall & Eschmeyer, 2002
[Jpn name: Inukasago]
KAUM–I. 134919, 166.2 mm SL, KAUM–I. 134920, 121.4 mm SL, Shiomichi.

16

K. Fujiwara and H. Motomura

Sebastapistes cyanostigma (Bleeker, 1856)[Jpn name: Kasurifusakasago]
Kamohara & Yamakawa (1968b, as Scorpaena albibrunnea): 2 unregistered specimens, 73–76 mm TL, Kikai Island. This study: KAUM–I. 146657, 28.3 mm SL, Araki.
Sebastapistes fowleri (Pietschmann, 1934)[Jpn name: Puchifusakasago]
KAUM–I. 146653, 26.2 mm SL, KAUM–I. 146659, 24.3 mm SL, Araki.
Sebastapistes strongia (Cuvier, 1829)[Jpn name: Madarafusakasago]
KAUM–I. 36033, 35.5 mm SL, Akaren; KAUM–I. 135128, 48.9 mm SL, Araki.
Sebastapistes tinkhami (Fowler, 1946)[Jpn name: Niramikasago]
KAUM–I. 146658, 26.6 mm SL, Araki.
Taenianotus triacanthus Lacepède, 1802
[Jpn name: Hadakahaokoze]
KAUM–I. 146466, 74.8 mm SL, Tekuzuku; KAUM–I. 146510, 38.2 mm SL, Onotsu.
FAMILY SYNANCEIIDAE
Synanceia verrucosa Bloch & Schneider, 1801
[Jpn name: Onidarumaokoze]
Kamohara (1957): 1 unregistered specimen, 60 mm TL, Wan. This study: KAUM–I. 134918, 192.5 mm SL,
Shiomichi.
FAMILY TETRAROGIDAE
Ablabys taenianotus (Cuvier, 1829)[Jpn name: Tsumajirokoze]
Kamohara (1957, as Amblyapistus taenianotus): 1 unregistered specimen, 75 mm TL, Wan; 1 unregisteredspecimen, 60 mm SL, Somachi. Kamohara & Yamakawa (1965, as Amblyapistus taenianotus): 2 unregistered specimens, 48–50 mm TL, Somachi. This study: KAUM–I. 134921, 45.7 mm SL, Shiomichi.
FAMILY PLATYCEPHALIDAE
Cociella punctata (Cuvier, 1829)[Jpn name: Minamiinegochi]
KAUM–I. 134673, 143.7 mm SL, Shiomichi; KAUM–I. 134726, 277.7 mm SL, Somachi.
Thysanophrys chiltonae Schultz, 1966
[Jpn name: Kuroshimagochi]
KAUM–I. 146525, 23.6 mm SL, Shiomichi.
ORDER PERCIFORMESFAMILY ACROPOMATIDAE
Verilus pacificus (Mochizuki, 1979)[Jpn name: Bakemutsu]
Inaba et al. (2016): KAUM–I. 54407, 345.9 mm SL, Kikai Island. Inaba (2018a): KAUM–I. 54407, 345.9mm SL, Kikai Island. Inaba (2019a): KAUM–I. 54407, 345.9 mm SL, Kikai Island. This study: KAUM–I.
54407, 345.9 mm SL, Kikai Island.

Marine and freshwater fishes of Kikai Island

17

FAMILY SERRANIDAE
Aporops bilinearis Schultz, 1943
[Jpn name: Yamatotogemegisu]
Kamohara (1957, as Aporops japonicus): BSKU 5425, 65 mm TL, Wan. Kamohara & Yamakawa (1968b): 3
unregistered specimens, 71–89 mm TL, Kikai Island. Senou (2013d): Kikai Island.
Cephalopholis igarashiensis Katayama, 1957
[Jpn name: Shimahata]
KAUM–I. 72512, 272.1 mm SL, Kikai Island.
Cephalopholis miniata (Forsskål, 1775)[Jpn name: Yukatahata]
Kamohara (1957, as Cephalopholis miniatus): 1 unregistered specimen, 250 mm TL, Somachi.
Cephalopholis sonnerati (Valenciennes, 1828)[Jpn name: Azahata]
Sakurai (2018a): KAUM–I. 72478, 295.2 mm SL, Shiramizu. Sakurai (2019a): KAUM–I. 72478, 295.2 mm
SL, Shiramizu. This study: KAUM–I. 72478, 295.2 mm SL, Shiramizu.
Cephalopholis urodeta (Forster, 1801)[Jpn name: Nijihata]
KAUM–I. 146652, 61.5 mm SL, Araki; KAUM–I. 146762, 43.5 mm SL, Tekuzuku.
Epinephelus areolatus (Forsskål, 1775)[Jpn name: Omonhata]
Kamohara (1957, as Epinephelus areolatus areolatus): 1 unregistered specimen, 71 mm TL, Somachi.
Epinephelus coeruleopunctatus (Bloch, 1790)[Jpn name: Hakutenhata]
Kamohara (1957, as Epinephelus summana caeruleopunctatus): 3 unregistered specimens, 40–212 mm TL,
Wan; 1 unregistered specimen, 235 mm TL, Somachi. Kamohara & Yamakawa (1968a): 2 unregistered specimens, 90–164 mm TL, Kikai Island.
Epinephelus cyanopodus (Richardson, 1846)[Jpn name: Tsuchihozeri]
KAUM–I. 134902, 154.5 mm SL, Somachi.
Epinephelus fasciatus (Forsskål, 1775)[Jpn name: Akahata]
KAUM–I. 146606, 55.5 mm SL, KAUM–I. 146607, 46.7 mm SL, Tekuzuku; KAUM–I. 146648, 50.6 mm
SL, Araki.
Epinephelus fuscoguttatus (Forsskål, 1775)[Jpn name: Akamadarahata]
Kamohara (1957): 1 unregistered specimen, 390 mm TL, Wan. Kamohara & Yamakawa (1968a): 4 unregistered specimens, 142–247 mm TL, Kikai Island.
Epinephelus hexagonatus (Forster, 1801)[Jpn name: Ishigakihata]

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K. Fujiwara and H. Motomura

Kamohara & Yamakawa (1968a): 2 unregistered specimens, 137–165 mm TL, Kikai Island. This study:
KAUM–I. 15526, 119.2 mm SL, Keraji; KAUM–I. 146650, 55.7 mm SL, Araki.
Epinephelus howlandi (Günther, 1873)[Jpn name: Hiregurohata]
KAUM–I. 134912, 78.1 mm SL, KAUM–I. 134913, 58.5 mm SL, KAUM–I. 136997, 98.8 mm SL, KAUM–
I. 136998, 102.3 mm SL, Shiomichi; KAUM–I. 146628, 54.4 mm SL, Kamikatetsu.
Epinephelus macrospilos (Bleeker, 1855)[Jpn name: Kibirehata]
KAUM–I. 135135, 122.5 mm SL, Shiomichi.
Epinephelus melanostigma Schultz, 1953
[Jpn name: Sumitsukihata]
Senou (2013d): Kikai Island.
Epinephelus merra Bloch, 1793
[Jpn name: Kammonhata]
KAUM–I. 5113, 104.7 mm SL, Kikai Island; KAUM–I. 43785, 84.3 mm SL, Keraji; KAUM–I. 134562,
144.7 mm SL, KAUM–I. 134563, 127.4 mm SL, KAUM–I. 134564, 115.0 mm SL, KAUM–I. 134565,
127.2 mm SL, KAUM–I. 146789, 94.8 mm SL, KAUM–I. 146790, 109.1 mm SL, KAUM–I. 146798, 100.1mm SL, KUN-P 55668, 115.5 mm SL, KUN-P 55669, 88.1 mm SL, Akaren; KAUM–I. 134668, 159.3 mm
SL, KAUM–I. 134669, 108.7 mm SL, KAUM–I. 134897, 101.1 mm SL, KAUM–I. 134898, 142.3 mm
SL, KAUM–I. 134899, 138.5 mm SL, KAUM–I. 134900, 120.3 mm SL, KAUM–I. 134901, 114.2 mm SL,
KAUM–I. 146810, 106.3 mm SL, Somachi; KAUM–I. 134730, 155.3 mm SL, KAUM–I. 146508, 41.4mm SL, Onotsu; KAUM–I. 134911, 111.0 mm SL, Shiomichi; KAUM–I. 146581, 51.1 mm SL, KAUM–I.
146627, 52.1 mm SL, Kamikatetsu.
Epinephelus tauvina (Forsskål, 1775)[Jpn name: Hitomihata]
Kamohara & Yamakawa (1968a): 12 unregistered specimens, 36–162 mm TL, Kikai Island.
Grammistes sexlineatus (Thunberg, 1792)[Jpn name: Nunosarashi]
Kamohara & Yamakawa (1965, as Grammistes sexlineatus proerythraeus): 1 unregistered specimen, 28 mm
TL, Somachi. This study: KAUM–I. 134998, 90.1 mm SL, Nakaguma; KAUM–I. 146698, 66.9 mm SL, Kamikatetsu; KUN-P 55733, 94.3 mm SL, Aden.
Pseudanthias squamipinnis (Peters, 1855)[Jpn name: Kingyohanadai]
Kamohara (1957): 2 unregistered specimens, 100–105 mm TL (as Franzia squamipinnis), Wan; 2 unregistered specimens, 112–120 mm TL (as Franzia cheirospilos), Wan.
Variola louti (Forsskål, 1775)[Jpn name: Barahata]
KAUM–I. 134896, 128.0 mm SL, KAUM–I. 134948, 132.4 mm SL, Somachi.

Marine and freshwater fishes of Kikai Island

19

FAMILY PSEUDOCHROMIDAE
Amsichthys knighti (Allen, 1987)[Jpn name: Ometanabatamegisu]
Yoshida (2018a): KAUM–I. 72312, 29.2 mm SL, Shiomichi. Yoshida (2019a): KAUM–I. 72312, 29.2 mm
SL, Shiomichi. This study: KAUM–I. 72312, 29.2 mm SL, Shiomichi.
Labracinus cyclophthalmus (Müller & Troschel, 1849)[Jpn name: Megisu]
Kamohara (1957, as Dampieria melanotaenia): 1 unregistered specimen, 71 mm TL, Somachi. Kamohara& Yamakawa (1968a, as Dampieria melanostigma): 5 unregistered specimens, 40–60 mm TL, Kikai Island.
This study: KAUM–I. 146796, 103.0 mm SL, Akaren; KAUM–I. 147603, 113.4 mm SL, KAUM–I. 147604,
105.2 mm SL, KAUM–I. 147605, 114.3 mm SL, Araki.
Pictichromis porphyreus (Lubbock & Goldman, 1974)[Jpn name: Kurenainisesuzume]
KAUM–I. 134731, 42.8 mm SL, Keraji; KAUM–I. 146593, 27.1 mm SL, Kamikatetsu; KAUM–I. 146613,
30.9 mm SL, Tekuzuku.
Pseudochromis cyanotaenia Bleeker, 1857
[Jpn name: Ryukyunisesuzume]
Aoyagi (1941, as Pseudochromis kikaii): YCM-P 6886, 32.8 mm SL, Kikai Island. Kamohara (1957): several unregistered specimens, 34–49 mm TL (as Pseudochromis kikaii) and 42–53 mm TL (as Pseudochromis
cyanotaenia), Kikai Island. Kamohara & Yamakawa (1965): 4 unregistered specimens, 42–48 mm TL (asPseudochromis cyanotaenia), Somachi; many unregistered specimens, 33–55 mm TL (as Pseudochromis
kikaii), Somachi. Hayashi (1995): YCM-P 6886, 32.8 mm SL, Kikai Island. Hayashi et al. (2013): Kikai Island. Yoshida (2018a): KAUM–I. 72357, 36.4 mm SL, Keraji. Yoshida (2019a): KAUM–I. 72357, 36.4 mm
SL, Keraji. This study: KAUM–I. 72314, 43.3 mm SL, Shiomichi; KAUM–I. 72357, 36.4 mm SL, Keraji;
KAUM–I. 146701, 21.1 mm SL, Kamikatetsu.
Pseudochromis luteus Aoyagi, 1943
[Jpn name: Koganenisesuzume]
Aoyagi (1943): 1 unregistered specimen, 44.2 mm SL, Kikai Island. Kamohara (1957): 2 unregistered specimens, 52–53 mm TL, Wan; 1 unregistered specimen, 52 mm TL, Somachi. Kamohara & Yamakawa (1965):
2 unregistered specimens, 51–52 mm TL, Tekuzuku; 7 unregistered specimens, 48–54 mm TL, Somachi.
Hayashi et al. (2013): Kikai Island.
Pseudochromis marshallensis Schultz, 1953
[Jpn name: Hoshinisesuzume]
Yoshida (2018a): KAUM–I. 72326, 42.7 mm SL, Onotsu. Yoshida (2019a): KAUM–I. 72326, 42.7 mm SL,
Onotsu. This study: KAUM–I. 72358, 32.9 mm SL, KAUM–I. 134741, 43.8 mm SL, KAUM–I. 134763,
24.1 mm SL, KAUM–I. 134799, 45.6 mm SL, Keraji; KAUM–I. 72326, 42.7 mm SL, KAUM–I. 134605,
38.4 mm SL, KAUM–I. 146505, 34.4 mm SL, Onotsu; KAUM–I. 146615, 30.4 mm SL, Tekuzuku; KAUM–
I. 146678, 38.5 mm SL, Araki.
Pseudoplesiops rosae Schultz, 1943
[Jpn name: Tanabatamegisu]
KAUM–I. 146519, 22.9 mm SL, Shiomichi; KAUM–I. 146622, 23.9 mm SL, Tekuzuku.

20

K. Fujiwara and H. Motomura

Pseudochromis tapeinosoma Bleeker, 1853
[Jpn name: Katsuironisesuzume]
Kamohara (1957): 1 unregistered specimen, 44 mm TL, Somachi.
FAMILY PLESIOPIDAE
Assessor randalli Allen & Kuiter, 1976
[Jpn name: Tsubametanabatauo]
KAUM–I. 72364, 23.5 mm SL, Keraji; KAUM–I. 146518, 24.3 mm SL, KAUM–I. 146526, 18.9 mm SL,
Shiomichi.
Belonepterygion fasciolatum (Ogilby, 1889)[Jpn name: Togetanabatauo]
Kamohara & Yamakawa (1968b, as Ernogrammoides fasciatus): 1 unregistered specimen, 31 mm TL, Kikai
Island.
Calloplesiops altivelis (Steindachner, 1903)[Jpn name: Shimofuritanabatauo]
KAUM–I. 134835, 91.3 mm SL, Shitooke.
Plesiops coeruleolineatus Rüppell, 1835
[Jpn name: Tanabatauo]
Yoshida (2018b): KAUM–I. 72187, 64.8 mm SL, Urahara. Yoshida (2019b): KAUM–I. 72187, 64.8 mm SL,
Urahara. This study: KAUM–I. 8093, 43.3 mm SL, KAUM–I. 8104, 24.0 mm SL, KAUM–I. 8105, 41.3 mm
SL, KAUM–I. 8106, 40.8 mm SL, KAUM–I. 8107, 57.3 mm SL, KAUM–I. 8119, 25.0 mm SL, Shitooke;
KAUM–I. 72187, 64.8 mm SL, Urahara; KAUM–I. 72365, 48.0 mm SL, KAUM–I. 72366, 23.8 mm SL,
KAUM–I. 134735, 48.0 mm SL, KAUM–I. 134736, 17.7 mm SL, Keraji; KAUM–I. 134602, 50.7 mm SL,
KAUM–I. 134603, 52.4 mm SL, KAUM–I. 134604, 39.7 mm SL, KAUM–I. 146504, 26.9 mm SL, Onotsu; KAUM–I. 134991, 21.0 mm SL, KAUM–I. 134992, 36.7 mm SL, Aden; KAUM–I. 135028, 38.8 mm
SL, Araki; KAUM–I. 136725, 34.9 mm SL, Shiomichi; KAUM–I. 146268, 46.3 mm SL, Ikeji; KAUM–I.
146577, 26.2 mm SL, KAUM–I. 146633, 37.6 mm SL, KAUM–I. 146634, 21.0 mm SL, KAUM–I. 146635,
52.8 mm SL, KAUM–I. 146636, 47.3 mm SL, KAUM–I. 146709, 50.1 mm SL, KAUM–I. 146710, 44.2 mm
SL, Kamikatetsu.
FAMILY PRIACANTHIDAE
Heteropriacanthus carolinus (Cuvier, 1829)[Jpn name: Gomahirekintoki]
KPM-NI 26353, 125.8 mm SL, Kikai Island.
Priacanthus hamrur (Forsskål, 1775)[Jpn name: Hosekikintoki]
Kawaji (2018): KAUM–I. 72756, 186.3 mm SL, Keraji. Kawaji (2019): KAUM–I. 72756, 186.3 mm SL,
Keraji. This study: KAUM–I. 15528, 133.4 mm SL, KAUM–I. 72755, 164.5 mm SL, KAUM–I. 72756,
186.3 mm SL, Keraji.
FAMILY APOGONIDAE
Apogon crassiceps Garman, 1903
[Jpn name: Akafujitenjikudai]
Yoshida (2018c): KAUM–I. 72196, 38.3 mm SL, Urahara. Yoshida (2019c): KAUM–I. 72196, 38.3 mm
SL, Urahara. This study: KAUM–I. 72195, 37.2 mm SL, KAUM–I. 72196, 38.3 mm SL, Urahara; KUN-P
55737, 23.6 mm SL, Aden.

Marine and freshwater fishes of Kikai Island

21

Apogon kominatoensis Ebina, 1935
[Jpn name: Kominatotenjikudai]
Yoshida (2018c): KAUM–I. 72353, 35.8 mm SL, Keraji. Yoshida (2019c): KAUM–I. 72353, 35.8 mm
SL, Keraji. This study: KAUM–I. 72353, 35.8 mm SL, Keraji; KAUM–I. 134636, 40.7 mm SL, Onotsu;
KAUM–I. 146642, 26.9 mm SL. Kamikatetsu.
Apogon sp.
[Jpn name: Ryukyuishimochi]
Hayashi (1995, as Apogon erythrinus): YCM-P 4366, 7 specimens, 27.5–44.3 mm SL, Kikai Island. Yoshida(2018c): KAUM–I. 72328, 34.2 mm SL, Onotsu. Yoshida (2019c): KAUM–I. 72328, 34.2 mm SL, Onotsu.
This study: KAUM–I. 72328, 34.2 mm SL, KAUM–I. 146499, 33.9 mm SL, KAUM–I. 146500, 33.4 mm
SL, Onotsu.
Apogon spp. (unidentified)[Jpn name: N/A]
KAUM–I. 134732, 39.9 mm SL, KAUM–I. 134767, 34.7 mm SL, Keraji; KAUM–I. 134936, 42.5 mm SL,
Nakaguma; KAUM–I. 146642, 26.9 mm SL, KAUM–I. 146711, 41.5 mm SL, Kamikatetsu.
Cercamia eremia (Allen, 1987)[Jpn name: Sakuratenjikudai]
Yoshida (2018c): KAUM–I. 72349, 40.4 mm SL, Keraji. Yoshida (2019c): KAUM–I. 72349, 40.4 mm SL,
Keraji. This study: KAUM–I. 72349, 40.4 mm SL, KAUM–I. 72350, 41.5 mm SL, KAUM–I. 134856, 23.0mm SL, KAUM–I. 134857, 31.8 mm SL, Keraji; KAUM–I. 134894, 27.1 mm SL, Sadeku.
Cheilodipterus intermedius Gon, 1993
[Jpn name: Sudareyaraiishimochi] (Fig. 3A)
Underwater photograph, off Araki.
Cheilodipterus macrodon (Lacepède, 1802)[Jpn name: Ryukyuyaraiishimochi]
Yoshida (2018c): KAUM–I. 72319, 49.0 mm SL, Shiomichi. Yoshida (2019c): KAUM–I. 72319, 49.0 mm
SL, Shiomichi. This study: KAUM–I. 72319, 49.0 mm SL, Shiomichi; KAUM–I. 72342, 154.7 mm SL, Keraji.
Cheilodipterus quinquelineatus Cuvier, 1828
[Jpn name: Yaraiishimochi]
KAUM–I. 136725, 34.9 mm SL, Shiomichi; KAUM–I. 146498, 98.3 mm SL, Onotsu; KAUM–I. 146696,
50.9 mm SL, KAUM–I. 146705, 70.2 mm SL, Kamikatetsu.
Foa fo Jordan & Seale, 1905
[Jpn name: Taiwammatoishimochi]
KAUM–I. 134685, 22.9 mm SL, KAUM–I. 134686, 18.9 mm SL, KAUM–I. 134697, 19.9 mm SL, Shiomichi.
Fowleria isostigma (Jordan & Seale, 1906)[Jpn name: Nahamatoishimochi]
Kamohara & Yamakawa (1968a, as Apogon isostigma): 1 unregistered specimen, 23 mm TL, Kikai Island.
This study: KAUM–I. 134882, 20.1 mm SL, Keraji; KAUM–I. 134889, 18.4 mm SL, Aden; KAUM–I.
146586, 14.7 mm SL, KAUM–I. 146629, 64.5 mm SL, Kamikatetsu.

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K. Fujiwara and H. Motomura

Nectamia fusca (Quoy & Gaimard, 1825)[Jpn name: Hososujinamidatenjikudai]
Yoshida (2018c): KAUM–I. 72347, 68.2 mm SL, KAUM–I. 72348, 50.3 mm SL, Keraji. Yoshida (2019c):
KAUM–I. 72347, 68.2 mm SL, KAUM–I. 72348, 50.3 mm SL, Keraji. This study: KAUM–I. 72347, 68.2mm SL, KAUM–I. 72348, 50.3 mm SL, KAUM–I. 134759, 22.8 mm SL, KAUM–I. 146565, 31.5 mm SL,
Keraji; KAUM–I. 134595, 64.8 mm SL, Onotsu; KAUM–I. 134694, 50.3 mm SL, KAUM–I. 136710, 66.3mm SL, KAUM–I. 136711, 52.0 mm SL, KAUM–I. 136712, 43.9 mm SL, Shiomichi; KAUM–I. 135033,
55.3 mm SL, KAUM–I. 135034, 46.6 mm SL, KAUM–I. 135038, 50.7 mm SL, Araki; KAUM–I. 146267,
36.3 mm SL, Ikeji; KAUM–I. 146707, 67.5 mm SL, Kamikatetsu; KUN-P 55675, 51.3 mm SL, Somachi;
KUN-P 55735, 76.0 mm SL, Nakaguma.
Nectamia savayensis (Günther, 1872)[Jpn name: Namidatenjikudai]
KAUM–I. 134933, 48.6 mm SL, KAUM–I. 134934, 38.0 mm SL, KAUM–I. 134935, 32.8 mm SL, Nakaguma.
Ostorhinchus cookii (Macleay, 1881)[Jpn name: Sujiishimochi]
Hayashi (1995, as Apogon robustus): YCM-P 4399, 2 specimens, 45.1–52.9 mm SL, Kikai Island. Yoshida(2018c): KAUM–I. 72186, 61.6 mm SL, Urahara. Yoshida (2019c): KAUM–I. 72186, 61.6 mm SL, Urahara.
This study: KAUM–I. 72186, 61.6 mm SL, Urahara; KAUM–I. 135035, 45.1 mm SL, KAUM–I. 135036,
42.5 mm SL, KAUM–I. 135037, 49.1 mm SL, Araki; KAUM–I. 136716, 69.1 mm SL, KAUM–I. 136717,
58.6 mm SL, Shiomichi; KUN-P 55774, 44.0 mm SL, KUN-P 55674, 62.7 mm SL, Somachi.
Ostorhinchus doederleini (Jordan & Snyder, 1901)[Jpn name: Osujiishimochi]
KAUM–I. 72335, 11.6 mm SL, Onotsu; KAUM–I. 134532, 92.2 mm SL, Akaren.
Ostorhinchus nigrofasciatus (Lachner, 1953)[Jpn name: Minamifutosujiishimochi]
KAUM–I. 72327, 53.6 mm SL, KAUM–I. 134632, 64.4 mm SL, KAUM–I. 134633, 54.7 mm SL, KAUM–
I. 134634, 48.3 mm SL, KAUM–I. 134635, 35.4 mm SL, Onotsu; KUN-P 55738, 27.8 mm SL, Aden.
Ostorhinchus novemfasciatus (Cuvier, 1828)[Jpn name: Tasujiishimochi]
Hayashi (1995, as Apogon novemfasciatus): YCM-P 4367, 3 specimens, 49.2–69.1 mm SL, Kikai Island.
This study: KAUM–I. 134931, 67.3 mm SL, KAUM–I. 134932, 61.3 mm SL, Nakaguma; KAUM–I.
135123, 26.9 mm SL, Araki; KAUM–I. 136729, 31.6 mm SL, Shiomichi.
Ostorhinchus properuptus (Whitley, 1964)[Jpn name: Kinsenishimochi]
Kamohara & Yamakawa (1968a, as Apogon cyanosoma): 4 unregistered specimens, 45–49 mm TL, Kikai
Island. This study: KAUM–I. 134803, 15.8 mm SL, Keraji; KAUM–I. 146467, 37.9 mm SL, KAUM–I.
146480, 42.5 mm SL, KAUM–I. 146483, 51.9 mm SL, KAUM–I. 146487, 19.2 mm SL, KAUM–I. 146488,
15.7 mm SL, KAUM–I. 146489, 14.8 mm SL, Tekuzuku; KAUM–I. 146501, 48.6 mm SL, Onotsu.
Ostorhinchus taeniophorus (Regan, 1908)[Jpn name: Misujitenjikudai]
Yoshida (2018c): KAUM–I. 72316, 52.7 mm SL, Shiomichi. Yoshida (2019c): KAUM–I. 72316, 52.7 mm

Marine and freshwater fishes of Kikai Island

23

SL, Shiomichi. This study: KAUM–I. 72316, 52.7 mm SL, Shiomichi; KAUM–I. 134531, 74.3 mm SL,
Akaren; KAUM–I. 134995, 32.7 mm SL, Aden; KAUM–I. 146502, 39.8 mm SL, Onotsu; KUN-P 55697,
21.4 mm SL, Nakasato.
Pristiapogon exostigma (Jordan & Starks, 1906)[Jpn name: Yukataishimochi]
Kamohara & Yamakawa (1968a, as Apogon exostigma): 1 unregistered specimen, 52 mm TL, Kikai Island.
This study: KAUM–I. 134768, 23.5 mm SL, KAUM–I. 134872, 81.3 mm SL, KAUM–I. 134883, 26.7 mm
SL, KAUM–I. 134884, 27.0 mm SL, Keraji; KAUM–I. 146708, 62.2 mm SL, Kamikatetsu; KAUM–I.
146708, 62.2 mm SL, Tekuzuku; KAUM–I. 146777, 84.6 mm SL, Akaren.
Pristiapogon fraenatus (Valenciennes, 1832)[Jpn name: Hitosujiishimochi]
KAUM–I. 146494, 24.4 mm SL, Tekuzuku; KAUM–I. 146591, 20.3 mm SL, Kamikatetsu.
Pristiapogon kallopterus (Bleeker, 1856)[Jpn name: Kasuriishimochi]
KAUM–I. 134625, 80.6 mm SL, KAUM–I. 134626, 90.6 mm SL, Onotsu; KAUM–I. 134743, 34.9 mm SL,
Keraji; KAUM–I. 134785, 69.4 mm SL, KAUM–I. 146811, 85.2 mm SL, KAUM–I. 146818, 96.7 mm SL,
KAUM–I. 146819, 93.1 mm SL, KUN-P 55718, 39.6 mm SL, KUN-P 55719, 43.4 mm SL, KUN-P 55720,
42.1 mm SL, Somachi.
Pseudamia gelatinosa Smith, 1956
[Jpn name: Numeritenjikudai]
KAUM–I. 146697, 15.9 mm SL, Kamikatetsu.
Pseudamiops gracilicauda (Lachner, 1953)[Jpn name: Kudaribozugisumodoki]
KAUM–I. 134744, 33.1 mm SL, Keraji.
FAMILY CARANGIDAE
Caranx melampygus Cuvier, 1833
[Jpn name: Kasumiaji]
KAUM–I. 134675, 95.8 mm SL, KAUM–I. 136706, 67.7 mm SL, Shiomichi; KAUM–I. 134838, 180.8 mm
SL, KAUM–I. 134950, 621.6 mm SL, KUN-P 55682, 72.0 mm SL, Somachi.
Caranx sexfasciatus Quoy & Gaimard, 1825
[Jpn name: Gingameaji]
Kamohara & Yamakawa (1968a): 1 unregistered specimen, 70 mm TL, Kikai Island. This study: KAUM–I.
134681, 50.5 mm SL, Shiomichi; KAUM–I. 146799, 258.2 mm SL, Somachi.
Trachinotus baillonii (Lacepède, 1801)[Jpn name: Kobanaji]
KAUM–I. 134693, 51.3 mm SL, Shiomichi; KUN-P 55673, Somachi.
Trachinotus blochii (Lacepède, 1801)[Jpn name: Marukoban]
KAUM–I. 134672, 126.7 mm SL, KAUM–I. 134692, 56.6 mm SL, KAUM–I. 136704, 49.4 mm SL,
KAUM–I. 136705, 51.8 mm SL, KUN-P 55686, 107.9 mm SL, Shiomichi.

24

K. Fujiwara and H. Motomura

FAMILY BRAMIDAE
Eumegistus illustris Jordan & Jordan, 1922
[Jpn name: Chikamechiopia]
Hata et al. (2015b): KAUM–I. 68443, 549.0 mm SL, Kikai Island. Hata (2018a): KAUM–I. 68443, 549.0mm SL, Kikai Island. Hata (2019a): KAUM–I. 68443, 549.0 mm SL, Kikai Island. This study: KAUM–I.
68443, 549.0 mm SL, Kikai Island.
FAMILY LUTJANIDAE
Aphareus furca (Lacepède, 1801)[Jpn name: Ishifuedai]
Kamohara (1957): 1 unregistered specimen, 285 mm TL, Wan.
Aphareus rutilans Cuvier, 1830
[Jpn name: Oguchiishichibiki]
Kamohara (1957): 1 unregistered specimen, 400 mm TL, Somachi.
Aprion virescens Valenciennes, 1830
[Jpn name: Aochibiki]
KAUM–I. 134715, 125.5 mm SL, Akaren.
Etelis coruscans Valenciennes, 1862
[Jpn name: Hamadai]
Kamohara (1957, as Etelis carbunculus): 2 unregistered specimens, 240–400 mm TL, Somachi. Hata
(2018b): KAUM–I. 72788, 330.1 mm SL, Kikai Island. Hata (2019b): KAUM–I. 72788, 330.1 mm SL, Kikai Island. This study: KAUM–I. 72788, 330.1 mm SL, Kikai Island.
Lutjanus argentimaculatus (Forsskål, 1775)[Jpn name: Gomafuedai]
Kamohara (1957): 1 unregistered specimen, 124 mm TL, Somachi.
Lutjanus bohar (Forsskål, 1775)[Jpn name: Barafuedai]
KAUM–I. 146788, 97.4 mm SL, Akaren.
Lutjanus fulviflamma (Forsskål, 1775)[Jpn name: Nisekurohoshifuedai]
Kamohara (1957): 3 unregistered specimens, 92–162 mm TL, Somachi.
Lutjanus fulvus (Forster, 1801)[Jpn name: Okifuedai]
Kamohara (1957, as Lutjanus vaigiensis): 1 unregistered specimen, 103 mm TL, Somachi. Kamohara &Yamakawa (1965, as Lutjanus vaigiensis): 1 unregistered specimen, 315 mm TL, Somachi. This study:
KAUM–I. 38208, 85.0 mm SL, KAUM–I. 38209, 92.0 mm SL, KAUM–I. 38210, 114.6 mm SL, Kikai Island; KAUM–I. 134676, 52.0 mm SL, KUN-P 55777, 45.5 mm SL, Shiomichi; KAUM–I. 134729, 116.6mm SL, Onotsu; KAUM–I. 135080, 67.5 mm SL, KAUM–I. 135085, 45.5 mm SL, Araki.
Lutjanus gibbus (Forsskål, 1775)[Jpn name: Himefuedai]

Marine and freshwater fishes of Kikai Island

25

KAUM–I. 134533, 89.6 mm SL, KAUM–I. 134534, 97.6 mm SL, KAUM–I. 134535, 84.4 mm SL, KUN-P
55661, 89.6 mm SL, KUN-P 55662, 93.4 mm SL, Akaren; KAUM–I. 134923, 93.0 mm SL, KAUM–I.
147628, 103.3 mm SL, Somachi; KAUM–I. 135067, 89.4 mm SL, KAUM–I. 135078, 86.4 mm SL, KAUM–
I. 135079, 63.8 mm SL, Araki; KUN-P 55679, 50.4 mm SL, Nakasato.
Lutjanus kasmira (Forsskål, 1775)[Jpn name: Yosujifuedai]
KAUM–I. 134536, 76.3 mm SL, KAUM–I. 134537, 83.4 mm SL, KAUM–I. 134540, 88.2 mm SL, KAUM–
I. 134541, 118.4 mm SL, KAUM–I. 134542, 102.3 mm SL, KUN-P 55664, 101.1 mm SL, KUN-P 55667,
101.3 mm SL, Akaren.
Lutjanus monostigma (Cuvier, 1828)[Jpn name: Ittenfuedai]
Kamohara & Yamakawa (1965): many unregistered specimens, 100–265 mm TL, Somachi.
Lutjanus rufolineatus (Valenciennes, 1830)[Jpn name: Kyusenfuedai]
Kamohara (1957): 1 unregistered specimen, 233 mm TL, Somachi.
Macolor macularis Fowler, 1931
[Jpn name: Hohosujitarumi] (Fig. 3B)
Underwater photograph, off Tekuzuku.
Paracaesio caerulea (Katayama, 1934)[Jpn name: Aodai]
Kamohara (1957): 1 unregistered specimen, 315 mm TL, Somachi. Hata (2018b): KAUM–I. 72510, 220.7mm SL, Kikai Island. Hata (2019b): KAUM–I. 72510, 220.7 mm SL, Kikai Island. This study: KAUM–I.
72510, 220.7 mm SL, Kikai Island.
Pristipomoides argyrogrammicus (Valenciennes, 1832)[Jpn name: Hanafuedai]
Kamohara (1957, as Pristipomoides amoenus): 1 unregistered specimen, 280 mm TL, Somachi. Hata
(2018b): KAUM–I. 72302, 180.0 mm SL, Kikai Island. Hata (2019b): KAUM–I. 72302, 180.0 mm SL, Kikai Island. This study: KAUM–I. 72302, 180.0 mm SL, Kikai Island.
Pristipomoides filamentosus (Valenciennes, 1830)[Jpn name: Ohime]
Kamohara (1957): 1 unregistered specimen, 210 mm TL, Somachi.
Pristipomoides sieboldii (Bleeker, 1855)[Jpn name: Himedai]
Hata (2018b): KAUM–I. 72511, 252.7 mm SL, Kikai Island. Hata (2019b): KAUM–I. 72511, 252.7 mm SL,
Kikai Island. This study: KAUM–I. 72511, 252.7 mm SL, Kikai Island.
FAMILY CAESIONIDAE
Pterocaesio tile (Cuvier, 1830)[Jpn name: Kumasasahanamuro]
KAUM–I. 146646, 85.2 mm SL, KAUM–I. 146647, 88.8 mm SL, KAUM–I. 147625, 96.2 mm SL, Kamikatetsu.

26

K. Fujiwara and H. Motomura

Pterocaesio trilineata Carpenter, 1987
[Jpn name: Issentakasago]
Kamohara (1957): 2 unregistered specimens, 150–163 mm TL, Wan.
FAMILY GERREIDAE
Gerres longirostris (Lacepède, 1801)[Jpn name: Tsupparisagi]
KAUM–I. 135066, 213.3 mm SL, Araki.
Gerres oblongus Cuvier, 1830
[Jpn name: Nagasagi]
KAUM–I. 134706, 40.8 mm SL, KAUM–I. 134671, 89.1 mm SL, Shiomichi.
Gerres oyena (Forsskål, 1775)[Jpn name: Minamikurosagi]
KAUM–I. 134690, 79.0 mm SL, Shiomichi.
FAMILY HAEMULIDAE
Plectorhinchus vittatus (Linnaeus, 1758)[Jpn name: Musujikoshodai]
Hata (2018c): KAUM–I. 72455, 157.6 mm SL, Shiramizu. Hata (2019c): KAUM–I. 72455, 157.6 mm SL,
Shiramizu. This study: KAUM–I. 72455, 157.6 mm SL, Shiramizu.
FAMILY NEMIPTERIDAE
Scolopsis bilineata (Bloch, 1793)[Jpn name: Futasujitamagashira]
KAUM–I. 146740, 24.9 mm SL, Keraji.
Scolopsis lineata Quoy & Gaimard, 1824
[Jpn name: Yokoshimatamagashira]
KAUM–I. 134904, 60.9 mm SL, KAUM–I. 136707, 54.7 mm SL, Shiomichi.
FAMILY SPARIDAE
Amamiichthys matsubarai (Akazaki, 1962)[Jpn name: Hoshirenko]
Nakamura (2018a): KAUM–I. 53888, 364.6 mm SL, KAUM–I. 72509, 267.3 mm SL, Kikai Island. Nakamura (2019a): KAUM–I. 53888, 364.6 mm SL, KAUM–I. 72509, 267.3 mm SL, Kikai Island. This study:
KAUM–I. 53888, 364.6 mm SL, KAUM–I. 72509, 267.3 mm SL, Kikai Island.
Dentex abei Iwatsuki, Akazaki & Taniguchi, 2007
[Jpn name: Kibireakarenko]
Nakamura (2018a): KAUM–I. 72513, 213.4 mm SL, Kikai Island. Nakamura (2019a): KAUM–I. 72513,
213.4 mm SL, Kikai Island. This study: KAUM–I. 72513, 213.4 mm SL, Kikai Island.
Pagrus major (Temminck & Schlegel, 1843)[Jpn name: Madai]
Hayashi & Hagiwara (2013a): Kikai Island.

Marine and freshwater fishes of Kikai Island

27

FAMILY LETHRINIDAE
Gnathodentex aureolineatus (Lacepède, 1802)[Jpn name: Nokogiridai]
KAUM–I. 15533, 112.4 mm SL, Keraji; KAUM–I. 135164, 104.7 mm SL, Akaren; KAUM–I. 146808, 120.2mm SL, KAUM–I. 146809, 85.0 mm SL, KAUM–I. 146820, 87.0 mm SL, Somachi.
Gymnocranius superciliosus Brosa, Béarez, Paijo & Chen, 2013
[Jpn name: Hikimayumeichi]
Shibuya et al. (2020): KAUM–I. 134725, 206.7 mm SL, Akaren. This study: KAUM–I. 134725, 206.7 mm
SL, Akaren.
Monotaxis grandoculis (Forsskål, 1775)[Jpn name: Yokoshimakurodai]
KAUM–I. 135065, 93.9 mm SL, Araki; KAUM–I. 146807, 110.3 mm SL, KAUM–I. 147627, 128.4 mm SL,
Somachi.
FAMILY MULLIDAE
Mulloidichthys flavolineatus flavolineatus (Lacepède, 1801)[Jpn name: Montsukiakahimeji]
Yamakawa (1969, as Mulloidichthys samoensis): 2 unregistered specimens, 72–88 mm TL, Kikai Island.
This study: KAUM–I. 72413, 249.8 mm SL, Keraji; KAUM–I. 72499, 222.7 mm SL, Onotsu; KAUM–I.
135026, 72.7 mm SL, KAUM–I. 136715, 78.3 mm SL, Shiomichi; KAUM–I. 134646, 113.8 mm SL, Nakasato; KAUM–I. 146776, 103.8 mm SL, Akaren; KUN-P 55710, 117.2 mm SL, KUN-P 55711, 86.1 mm SL,
KUN-P 55712, 77.2 mm SL, Somachi.
Mulloidichthys vanicolensis (Valenciennes, 1831)[Jpn name: Akahimeji]
KAUM–I. 146812, 123.8 mm SL, Somachi.
Parupeneus barberinus (Lacepède, 1801)[Jpn name: Osujihimeji]
Yamakawa (1969, as Pseudupeneus barberinus): 1 unregistered specimen, 38 mm TL, Kikai Island. This
study: KAUM–I. 134717, 196.6 mm SL, Akaren.
Parupeneus ciliatus (Lacepède, 1802)[Jpn name: Horaihimeji]
KAUM–I. 87310, 78.8 mm SL, Kikai Island; KAUM–I. 134914, 164.3 mm SL, Shiomichi; KAUM–I.
146261, 97.5 mm SL, Ikeji; KAUM–I. 147624, 131.0 mm SL, Kamikatetsu.
Parupeneus cyclostomus (Lacepède, 1801)[Jpn name: Marukuchihimeji]
Kamohara (1957, as Pseudupeneus cyclostomus): several unregistered specimens, 123–227 mm TL, Kikai
Island (including Amami-oshima Island).
Parupeneus multifasciatus (Quoy & Gaimard, 1825)[Jpn name: Ojisan]
Bandai (2018): KAUM–I. 72457, 175.6 mm SL, Shiramizu. Bandai (2019): KAUM–I. 72457, 175.6 mm
SL, Shiramizu. This study: KAUM–I. 72457, 175.6 mm SL, Shiramizu; KAUM–I. 72485, 167.1 mm SL,

28

K. Fujiwara and H. Motomura

Aden; KAUM–I. 134543, 107.5 mm SL, KAUM–I. 134544, 93.3 mm SL, KAUM–I. 134545, 127.5 mm
SL, KAUM–I. 134546, 126.6 mm SL, KAUM–I. 134547, 105.9 mm SL, KAUM–I. 134548, 102.1 mm
SL, KAUM–I. 134549, 88.6 mm SL, KAUM–I. 134550, 149.1 mm SL, KAUM–I. 134551, 121.5 mm
SL, KAUM–I. 134552, 113.2 mm SL, KAUM–I. 134553, 96.7 mm SL, KAUM–I. 134722, 144.5 mm SL,
KAUM–I. 146784, 97.1 mm SL, KAUM–I. 146785, 105.4 mm SL, KAUM–I. 146786, 83.4 mm SL, KUN-P
55658, 127.6 mm SL, KUN-P 55659, 108.9 mm SL, Akaren; KAUM–I. 134665, 191.5 mm SL, KAUM–I.
134666, 118.5 mm SL, KAUM–I. 134667, 99.6 mm SL, KAUM–I. 146816, 91.4 mm SL, KAUM–I. 147632,
91.8 mm SL, KAUM–I. 147633, 77.5 mm SL, KAUM–I. 147634, 94.3 mm SL, KAUM–I. 147635, 120.3mm SL, Somachi; KAUM–I. 146824, 136.3 mm SL, KAUM–I. 146825, 89.6 mm SL, Araki.
Parupeneus pleurostigma (Bennett, 1831)[Jpn name: Ryukyuhimeji]
Kamohara (1957): 1 unregistered specimen, 165 mm TL, Wan. This study: KAUM–I. 15527, 132.4 mm SL,
Keraji.
Parupeneus spilurus (Bleeker, 1854)[Jpn name: Okinahimeji]
Kamohara & Yamakawa (1965): 1 unregistered specimen, 200 mm TL, Somachi.
Upeneus tragula Richardson, 1846
[Jpn name: Yomehimeji]
KAUM–I. 15291, 180.8 mm SL, Sakamine.
FAMILY PEMPHERIDAE
Pempheris adusta Bleeker, 1877
[Jpn name: Ryukyuhatampo]
Kamohara (1957, as Pempheris oualensis): 2 unregistered specimens, 67–75 mm TL, Somachi. Kamohara& Yamakawa (1965, as Pempheris oualensis): 5 unregistered specimens, 75–155 mm TL, Somachi. Koeda(2018): KAUM–I. 72334, 8.7 mm SL, Onotsu. Koeda (2019): KAUM–I. 72334, 8.7 mm SL, Onotsu. This
study: KAUM–I. 72334, 8.7 mm SL, Onotsu.
Pempheris schwenkii Bleeker, 1855
[Jpn name: Minamihatampo]
Koeda (2018): KAUM–I. 72331, 8.0 mm SL, Onotsu. Koeda (2019): KAUM–I. 72331, 8.0 mm SL, Onotsu.
This study: KAUM–I. 72331, 8.0 mm SL, Onotsu; KAUM–I. 134873, 111.3 mm SL, Keraji.
FAMILY MONODACTYLIDAE
Monodactylus argenteus (Linnaeus, 1758)[Jpn name: Himetsubameuo]
Kamohara (1957): 5 unregistered specimens, 95–120 mm TL, Kikai Island (including Amami-oshima Island). This study: KAUM–I. 134679, 38.8 mm SL, KUN-P 55688, 38.5 mm SL, Shiomichi.
FAMILY CHAETODONTIDAE
Chaetodon argentatus Smith & Radcliffe, 1911
[Jpn name: Kagamichochouo]
Kamohara & Yamakawa (1968a): 1 unregistered specimen, 45 mm TL, Kikai Island. Fujiwara (2018):
KAUM–I. 72341, 76.7 mm SL, Keraji. Fujiwara (2019): KAUM–I. 72341, 76.7 mm SL, Keraji. This study:
KAUM–I. 72341, 76.7 mm SL, Keraji; KAUM–I. 146764, 62.9 mm SL, Tekuzuku.

Marine and freshwater fishes of Kikai Island

29

Chaetodon auriga Forsskål, 1775
[Jpn name: Togechochouo]
Kamohara (1957): 2 unregistered specimens, 54–55 mm TL, Wan. This study: KAUM–I. 72474, 100.9 mm
SL, Shiramizu; KAUM–I. 135132, 59.9 mm SL, Shiomichi.
Chaetodon auripes Jordan & Snyder, 1901
[Jpn name: Chochouo]
Kamohara (1957, as Chaetodon collaris): several unregistered specimens, 130–170 mm TL, Somachi.
This study: KAUM–I. 72754, 145.7 mm SL, Keraji; KAUM–I. 146775, 133.8 mm SL, Akaren; KAUM–I.
146805, 134.9 mm SL, Somachi.
Chaetodon citrinellus Cuvier, 1831
[Jpn name: Gomachochouo]
Kamohara (1957): many unregistered specimens, 60–83 mm TL, Kikai Island. Kamohara & Yamakawa(1968a): 3 unregistered specimens, 33–40 mm TL, Kikai Island. This study: KAUM–I. 15295, 85.7 mm SL,
KAUM–I. 15403, 91.2 mm SL, Sakamine; KAUM–I. 134627, 78.1 mm SL, Onotsu; KAUM–I. 134797, 38.0mm SL, Keraji.
Chaetodon daedalma Jordan & Fowler, 1902
[Jpn name: Yuzen] (Fig. 3C)
Underwater photograph, off Ikeji.
Remarks: At Kikai Island, only a single ca. 3 cm SL juvenile (taken at Ikeji on 15 August 2018) has been
confirmed to date (J. Yoda pers. comm.).
Chaetodon lunula (Lacepède, 1802)[Jpn name: Chohan]
Kamohara & Yamakawa (1965): 3 unregistered specimens, 43–90 mm TL, Araki; 3 unregistered specimens,
59–87 mm TL, Somachi. Fujiwara (2018): KAUM–I. 72459, 95.3 mm SL, Shiramizu. Fujiwara (2019):
KAUM–I. 72459, 95.3 mm SL, Shiramizu. This study: KAUM–I. 72458, 92.2 mm SL, KAUM–I. 72459,
95.3 mm SL, Shiramizu; KAUM–I. 135169, 26.6 mm SL, Shiomichi.
Chaetodon lunulatus Quoy & Gaimard, 1825
[Jpn name: Misujichochouo]
KAUM–I. 15368, 94.7 mm SL, Sakamine; KAUM–I. 15538, 101.6 mm SL, KAUM–I. 87266, 91.0 mm SL,
KAUM–I. 134737, 26.1 mm SL, KAUM–I. 134738, 16.4 mm SL, KAUM–I. 134764, 17.3 mm SL, KAUM–
I. 146545, 18.8 mm SL, KAUM–I. 147615, 88.2 mm SL, Keraji.
Chaetodon melannotus Bloch & Schneider, 1801
[Jpn name: Akebonochochouo]
Fujiwara (2018): KAUM–I. 72454, 126.4 mm SL, Shiramizu. Fujiwara (2019): KAUM–I. 72454, 126.4 mm
SL, Shiramizu. This study: KAUM–I. 72454, 126.4 mm SL, Shiramizu; KAUM–I. 72492, 125.4 mm SL,
Onotsu.
Chaetodon speculum Cuvier, 1831
[Jpn name: Tonosamadai]
Fujiwara (2018): KAUM–I. 72479, 137.1 mm SL, Aden. Fujiwara (2019): KAUM–I. 72479, 137.1 mm SL,
Aden. This study: KAUM–I. 72479, 137.1 mm SL, Aden.

30

K. Fujiwara and H. Motomura

Chaetodon ulietensis Cuvier, 1831
[Jpn name: Sudarechochouo]
KAUM–I. 147623, 129.6 mm SL, Kamikatetsu.
Chaetodon unimaculatus Bloch, 1787
[Jpn name: Ittenchochouo]
Kamohara (1957): many unregistered specimens, 65–144 mm TL, Kikai Island (including Amami-oshimaIsland).
Chaetodon vagabundus Linnaeus, 1758
[Jpn name: Furaichochouo]
Kamohara (1957): 6 unregistered specimens, 46–140 mm TL, Kikai Island. Kamohara & Yamakawa (1965):
1 unregistered specimen, 86 mm TL, Somachi; 1 unregistered specimen, 52 mm SL, Araki. This study:
KAUM–I. 72496, 101.2 mm SL, Onotsu; KAUM–I. 134680, 50.6 mm SL, KAUM–I. 135170, 38.2 mm SL,
Shiomichi; KAUM–I. 146817, 127.4 mm SL, Somachi; KAUM–I. 147614, 81.1 mm SL, Keraji.
Forcipiger flavissimus Jordan & McGregor, 1898
[Jpn name: Fueyakkodai]
Kamohara (1957, as Forcipiger longirostris): 5 unregistered specimens, 110–115 mm TL, Wan (including
Amami-oshima Island). This study: KAUM–I. 15356, 104.2 mm SL, Sakamine; KAUM–I. 15531, 108.6 mm
SL, Keraji.
Hemitaurichthys polylepis (Bleeker, 1857)[Jpn name: Kasumichochouo]
Kamohara (1957, as Hemitaurichthys zoster): many unregistered specimens, 112–152 mm TL, Kikai Island(including Amami-oshima Island).
Heniochus chrysostomus Cuvier, 1831
[Jpn name: Minamihatatatedai]
KAUM–I. 146645, 53.8 mm SL, Kamikatetsu.
Heniochus monoceros Cuvier, 1831
[Jpn name: Onihatatatedai]
Kamohara & Yamakawa (1968b): 1 unregistered specimen, 45 mm TL, Kikai Island.
FAMILY POMACANTHIDAE
Centropyge heraldi Woods & Schultz, 1953
[Jpn name: Herarudokoganeyakko]
KAUM–I. 146614, 45.4 mm SL, Tekuzuku.
Centropyge vrolikii (Bleeker, 1853)[Jpn name: Namerayakko]
KAUM–I. 146651, 39.9 mm SL, KAUM–I. 146656, 37.5 mm SL, Araki.
Pomacanthus imperator (Bloch, 1787)[Jpn name: Tatejimakinchakudai]Kamohara & Yamakawa (1968b, as Holacanthus imperator): 1 unregistered specimen, 39 mm TL, Kikai Island.

Marine and freshwater fishes of Kikai Island

31

Pomacanthus semicirculatus (Cuvier, 1831)[Jpn name: Sazanamiyakko]
Kamohara (1957, as Holacanthus semicirculatus): 5 unregistered specimens, 36–125 mm TL, Onotsu. Wibowo (2018): KAUM–I. 72318, 49.6 mm SL, Shiomichi. Wibowo (2019): KAUM–I. 72318, 49.6 mm SL,
Shiomichi. This study: KAUM–I. 72318, 49.6 mm SL, Shiomichi.
FAMILY CIRRHITIDAE
Amblycirrhitus unimaculus (Kamohara, 1957)[Jpn name: Kurohoshigombe]
Kamohara (1957, as Cirrhitoidea unimacula): BSKU 6387, 85 mm TL, Somachi. Hayashi & Hagiwara(2013b, as Amblycirrhitus unimacula): Kikai Island.
Cirrhitichthys aprinus (Cuvier, 1829)[Jpn name: Minamigombe]
KAUM–I. 146677, 45.1 mm SL, Araki.
Cirrhitichthys falco Randall, 1963
[Jpn name: Sarasagombe]
KAUM–I. 146692, 53.2 mm SL, Kamikatetsu.
Cirrhitus pinnulatus pinnulatus (Forster, 1801)[Jpn name: Isogombe]
Kamohara & Yamakawa (1965, as Cirrhitus pinnulatus): 6 unregistered specimens, 56–158 mm TL, Somachi. Hayashi & Hagiwara (2013b, as Cirrhitus pinnulatus): Kikai Island.
Paracirrhites arcatus (Cuvier, 1829)[Jpn name: Meganegombe]
KAUM–I. 146655, 53.7 mm SL, KAUM–I. 146672, 26.6 mm SL, KAUM–I. 146679, 40.7 mm SL, Araki;
KAUM–I. 146763, 27.9 mm SL, Tekuzuku.
Paracirrhites forsteri (Schneider, 1801)[Jpn name: Hoshigombe]
KAUM–I. 134795, 35.0 mm SL, KAUM–I. 134796, 29.4 mm SL, Keraji; KAUM–I. 146683, 76.2 mm SL,
Araki.
FAMILY LATRIDAE
Goniistius zonatus (Cuvier, 1830)[Jpn name: Takanohadai]
Kamohara & Yamakawa (1965): 1 unregistered specimen, 75 mm TL, Wan.
FAMILY CICHLIDAE
Oreochromis niloticus (Linnaeus, 1758)[Jpn name: Nairuteirapia]
Nakajima et al. (2020): Urahara. This study: KAUM–I. 135134, 117.6 mm SL, Shiomichi.
FAMILY POMACENTRIDAE
Abudefduf caudobimaculatus Okada & Ikeda, 1939
[Jpn name: Shiritensuzumedai]
Wibowo et al. (2017): KPM-NI 26335, 69.3 mm SL, Kikai Island; Iwatsubo (2018): KPM-NI 26335, 69.3mm SL, Kikai Island. Iwatsubo (2019): KPM-NI 26335, 69.3 mm SL, Kikai Island.

32

K. Fujiwara and H. Motomura

Abudefduf notatus (Day, 1870)[Jpn name: Isosuzumedai]
Kamohara (1957): 1 unregistered specimen, 62 mm TL, Onotsu; 1 unregistered specimen, 85 mm TL, Somachi. Kamohara & Yamakawa (1965): 1 unregistered specimen, 80 mm TL, Tekuzuku; 1 unregistered specimen, 72 mm TL, Somachi. Iwatsubo (2018): KAUM–I. 72505, 102.4 mm SL, Onotsu. Iwatsubo (2019):
KAUM–I. 72505, 102.4 mm SL, Onotsu. This study: KAUM–I. 72505, 102.4 mm SL, Onotsu.
Abudefduf septemfasciatus (Cuvier, 1830)[Jpn name: Shichisensuzumedai]
Kamohara (1957): 4 unregistered specimens, 67–83 mm TL, Onotsu. Kamohara & Yamakawa (1965): 7
unregistered specimens, 76–92 mm TL, Somachi. This study: KAUM–I. 72507, 116.2 mm SL, Onotsu;
KAUM–I. 135089, 19.3 mm SL, KAUM–I. 135090, 15.9 mm SL, KAUM–I. 135091, 16.3 mm SL, KAUM–
I. 135092, 18.4 mm SL, KAUM–I. 135093, 14.2 mm SL, Araki.
Abudefduf sexfasciatus (Lacepède, 1801)[Jpn name: Rokusensuzumedai]
Kamohara (1957): 2 unregistered specimens, 70–83 mm TL, Onotsu. Kamohara & Yamakawa (1968a): 3
unregistered specimens, 45–54 mm TL, Kikai Island. This study: KAUM–I. 134538, 73.8 mm SL, KAUM–
I. 134539, 75.3 mm SL, KUN-P 55665, 75.8 mm SL, Akaren; KAUM–I. 134938, 37.1 mm SL, Nakaguma;
KAUM–I. 135074, 54.6 mm SL, KAUM–I. 135075, 61.5 mm SL, Araki; KAUM–I. 146713, 50.7 mm SL,
Kamikatetsu; KAUM–I. 147640, 93.4 mm SL, Somachi.
Abudefduf sordidus (Forsskål, 1775)[Jpn name: Shimasuzumedai]
KAUM–I. 8111, 53.3 mm SL, Urahara.
Abudefduf vaigiensis (Quoy & Gaimard, 1825)[Jpn name: Oyabitcha]
KAUM–I. 8112, 39.1 mm SL, KAUM–I. 72192, 42.6 mm SL, Urahara; KAUM–I. 72300, 84.6 mm SL, Onotsu; KAUM–I. 135073, 68.9 mm SL, Araki; KAUM–I. 147639, 93.6 mm SL, Somachi; KUN-P 55654, 20.8mm SL, Nakasato.
Amphiprion clarkii (Bennett, 1830)[Jpn name: Kumanomi]
Iwatsubo (2018): KAUM–I. 72367, 25.0 mm SL, Keraji. Iwatsubo (2019): KAUM–I. 72367, 25.0 mm SL,
Keraji. This study: KAUM–I. 72367, 25.0 mm SL, Keraji; KAUM–I. 146611, 33.3 mm SL, Tekuzuku;
KAUM–I. 146688, 22.7 mm SL, Araki.
Amphiprion frenatus Brevoort, 1856
[Jpn name: Hamakumanomi]
KAUM–I. 134874, 17.1 mm SL, KAUM–I. 134875, 78.9 mm SL, KAUM–I. 134876, 60.8 mm SL, Keraji;
KAUM–I. 146496, 62.1 mm SL, Tekuzuku.
Amphiprion perideraion Bleeker, 1855
[Jpn name: Hanabirakumanomi]
KAUM–I. 134890, 67.6 mm SL, KAUM–I. 134891, 52.1 mm SL, KAUM–I. 134892, 18.5 mm SL, Sadeku;
KAUM–I. 146520, 23.1 mm SL, Shiomichi; KAUM–I. 146716, 31.5 mm SL, Tekuzuku.

Marine and freshwater fishes of Kikai Island

33

Chromis atripectoralis Welander & Schultz, 1951
[Jpn name: Aobasuzumedai]
KAUM–I. 146571, 25.4 mm SL, Keraji.
Chromis chrysura (Bliss, 1883)[Jpn name: Amamisuzumedai]
Kamohara (1957, as Chromis isharae): 1 unregistered specimen, 132 mm TL, Wan. This study: KPM-NI
48951, 119.1 mm SL, Kikai Island; KAUM–I. 147631, 90.2 mm SL, Somachi.
Chromis margaritifera Fowler, 1946
[Jpn name: Shikokusuzumedai]
KAUM–I. 134637, 49.4 mm SL, Onotsu; KAUM–I. 146459, 19.0 mm SL, KAUM–I. 146460, 15.3 mm SL,
KAUM–I. 146461, 17.1 mm SL, KAUM–I. 146481, 51.0 mm SL, Tekuzuku; KAUM–I. 146661, 30.2 mm
SL, KAUM–I. 146662, 22.0 mm SL, Araki.
Chromis ovatiformes Fowler, 1946
[Jpn name: Marusuzumedai]
KAUM–I. 146462, 51.9 mm SL, KAUM–I. 146463, 24.6 mm SL, KAUM–I. 146464, 17.1 mm SL, KAUM–
I. 146604, 25.0 mm SL, KAUM–I. 146605, 21.5 mm SL, Tekuzuku; KAUM–I. 146669, 59.2 mm SL, Araki.
Chromis vanderbilti (Fowler, 1941)[Jpn name: Himesuzumedai]
KAUM–I. 146654, 24.1 mm SL, KAUM–I. 146689, 47.7 mm SL, Araki.
Chromis viridis (Cuvier, 1830)[Jpn name: Debasuzumedai]
KAUM–I. 134747, 23.3 mm SL, KAUM–I. 134748, 13.5 mm SL, KAUM–I. 134749, 11.9 mm SL, Keraji.
Chromis xouthos Allen & Erdmann, 2005
[Jpn name: Himawarisuzumedai]
Koeda et al. (2016): KAUM–I. 82088, 111.9 mm SL, Kikai Island. Iwatsubo (2018): KAUM–I. 82088, 111.9mm SL, Kikai Island. Iwatsubo (2019): KAUM–I. 82088, 111.9 mm SL, Kikai Island. This study: KAUM–I.
82088, 111.9 mm SL, Kikai Island.
Chromis yamakawai Iwatsubo & Motomura, 2013
[Jpn name: Kihoshisuzumedai]
KAUM–I. 134661, 76.2 mm SL, Nakasato; KAUM–I. 146804, 92.9 mm SL, Somachi.
Chrysiptera biocellata (Quoy & Gaimard, 1825)[Jpn name: Sujibuchisuzumedai]
Kamohara (1957, as Abudefduf biocellatus): 1 unregistered specimen, 48 mm TL, Wan. Kamohara & Yamakawa (1965, as Abudefduf biocellatus): 2 unregistered specimens, 40–45 mm TL, Araki. This study:
KAUM–I. 134905, 25.6 mm SL, KAUM–I. 134906, 57.1 mm SL, KAUM–I. 136718, 52.2 mm SL, Shiomichi; KAUM–I. 134993, 20.8 mm SL, KAUM–I. 134994, 16.4 mm SL, Aden; KAUM–I. 135095, 64.8mm SL, KAUM–I. 135096, 26.2 mm SL, KAUM–I. 135097, 17.0 mm SL, KAUM–I. 135125, 18.4 mm SL,
Araki; KAUM–I. 135111, 42.4 mm SL, KAUM–I. 135112, 25.2 mm SL, KAUM–I. 135113, 16.3 mm SL,
KAUM–I. 147613, 72.3 mm SL, Keraji.

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Chrysiptera caeruleolineata (Allen, 1973)[Jpn name: Aosujisuzumedai]
Hayashi (1995): YCM-P 31346, 42.6 mm SL, Kikai Island.
Chrysiptera cyanea (Quoy & Gaimard, 1825)[Jpn name: Rurisuzumedai]
Kamohara (1957, as Abudefduf cyaneus): 1 unregistered specimen, 62 mm TL, Somachi. Kamohara & Yamakawa (1968a, as Abudefduf assimilis): 2 unregistered specimens, 36–38 mm TL, Kikai Island. This study:
KAUM–I. 146746, 36.0 mm SL, Keraji.
Chrysiptera glauca (Cuvier, 1830)[Jpn name: Nezusuzumedai]
Kamohara & Yamakawa (1968a, as Abudefduf glaucus): 4 unregistered specimens, 22–42 mm TL, Kikai
Island. Iwatsubo (2018): KAUM–I. 72193, 68.5 mm SL, Urahara. Iwatsubo (2019): KAUM–I. 72193, 68.5mm SL, Urahara. This study: KAUM–I. 72193, 68.5 mm SL, Urahara; KAUM–I. 134940, 15.2 mm SL,
KAUM–I. 134941, 21.4 mm SL, KAUM–I. 134942, 12.2 mm SL, Nakaguma; KAUM–I. 135061, 51.2 mm
SL, KAUM–I. 135062, 21.6 mm SL, KAUM–I. 135063, 18.5 mm SL, KAUM–I. 135094, 19.7 mm SL, Araki; KUN-P 55650, 21.2 mm SL, KUN-P 55650, 21.2 mm SL, KUN-P 55650, 21.2 mm SL, KUN-P 55651,
22.1 mm SL, KUN-P 55652, 26.2 mm SL, KUN-P 55653, 28.6 mm SL, KUN-P 55656, 67.6 mm SL, KUN-P
55657, 54.9 mm SL, Nakasato.
Chrysiptera leucopoma (Cuvier, 1830)[Jpn name: Miyakokisensuzumedai]
Kamohara (1957, as Abudefduf albofasciatus): 3 unregistered specimens, 34–66 mm TL, Kikai Island.
Chrysiptera rex (Snyder, 1909)[Jpn name: Remonsuzumedai]
Kamohara (1957, as Abudefduf rex): 5 unregistered specimens, 58–71 mm TL, Somachi. This study:
KAUM–I. 134880, 44.7 mm SL, KAUM–I. 146546, 31.8 mm SL, KAUM–I. 146728, 33.6 mm SL, Keraji;
KAUM–I. 146718, 33.3 mm SL, Tekuzuku.
Chrysiptera starcki (Allen, 1973)[Jpn name: Senakirurisuzumedai]
KAUM–I. 146612, 44.2 mm SL, Tekuzuku.
Chrysiptera unimaculata (Cuvier, 1830)[Jpn name: Ichimonsuzumedai]
KAUM–I. 134922, 17.8 mm SL, Shiomichi; KAUM–I. 146547, 25.1 mm SL, Keraji.
Dascyllus aruanus (Linnaeus, 1758)[Jpn name: Misujiryukyusuzumedai]
KAUM–I. 134750, 44.7 mm SL, KAUM–I. 134751, 30.7 mm SL, KAUM–I. 134752, 21.7 mm SL, KAUM–
I. 134753, 22.1 mm SL, KAUM–I. 134754, 18.4 mm SL, KAUM–I. 134755, 19.0 mm SL, KAUM–I.
134756, 15.2 mm SL, KAUM–I. 134786, 38.5 mm SL, KAUM–I. 134787, 32.1 mm SL, KAUM–I. 135114,
36.3 mm SL, KAUM–I. 135115, 29.2 mm SL, KAUM–I. 135116, 15.7 mm SL, Keraji; KAUM–I. 146506,
32.6 mm SL, Onotsu.

Marine and freshwater fishes of Kikai Island

35

Dascyllus trimaculatus (Rüppell, 1829)[Jpn name: Mitsuboshikurosuzumedai]
KAUM–I. 134831, 17.7 mm SL, Shitooke; KAUM–I. 146687, 13.5 mm SL, Araki; KAUM–I. 147629, 88.0mm SL, KAUM–I. 147630, 88.0 mm SL, Somachi.
Neoglyphidodon melas (Valenciennes, 1830)[Jpn name: Kurosuzumedai]
KAUM–I. 134878, 32.6 mm SL, Keraji.
Neoglyphidodon nigroris (Cuvier, 1830)[Jpn name: Hirenagasuzumedai]
KAUM–I. 134721, 85.2 mm SL, Akaren; KAUM–I. 134740, 29.4 mm SL, KAUM–I. 134792, 51.2 mm SL,
KAUM–I. 146540, 25.0 mm SL, Keraji.
Plectroglyphidodon imparipennis (Vaillant & Sauvage, 1875)[Jpn name: Iwasakisuzumedai]
Kamohara (1957, as Abudefduf iwasakii): 3 unregistered specimens, 53–57 mm TL, Somachi. This study:
KAUM–I. 146603, 46.0 mm SL, KAUM–I. 146766, 35.9 mm SL, Tekuzuku; KAUM–I. 146691, 41.7 mm
SL, Kamikatetsu.
Plectroglyphidodon johnstonianus Fowler & Ball, 1924
[Jpn name: Rurimeishigakisuzumedai]
KAUM–I. 146541, 64.8 mm SL, Keraji.
Plectroglyphidodon lacrymatus (Quoy & Gaimard, 1825)[Jpn name: Rurihoshisuzumedai]
Iwatsubo (2018): KAUM–I. 72332, 39.0 mm SL, Onotsu. Iwatsubo (2019): KAUM–I. 72332, 39.0 mm SL,
Onotsu. This study: KAUM–I. 72332, 39.0 mm SL, KAUM–I. 134587, 16.4 mm SL, KAUM–I. 134588,
50.7 mm SL, KAUM–I. 134589, 14.3 mm SL, KAUM–I. 134631, 78.7 mm SL, Onotsu; KAUM–I. 72351,
46.3 mm SL, KAUM–I. 134758, 51.0 mm SL, KAUM–I. 134788, 51.6 mm SL, KAUM–I. 146548, 15.9 mm
SL, KAUM–I. 146564, 47.0 mm SL, Keraji; KAUM–I. 146465, 16.8 mm SL, Tekuzuku; KAUM–I. 146637,
22.2 mm SL, Kamikatetsu; KAUM–I. 146660, 36.5 mm SL, KUN-P 55765, 22.7 mm SL, Araki.
Plectroglyphidodon leucozonus (Bleeker, 1859)[Jpn name: Hakusensuzumedai]
KUN-P 55734, 61.4 mm SL, Aden.
Plectroglyphidodon phoenixensis (Schultz, 1943)[Jpn name: Fuenikkususuzumedai]
Aonuma (2013): Kikai Island.
Pomacentrus amboinensis Bleeker, 1868
[Jpn name: Nisenettaisuzumedai]
Kamohara (1957, as Chromis dimidiatus): 1 unregistered specimen, 78 mm TL, Wan. This study: KAUM–I.
146542, 52.3 mm SL, Keraji; KAUM–I. 146693, 26.9 mm SL, Kamikatetsu.

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Pomacentrus bankanensis Bleeker, 1854
[Jpn name: Meganesuzumedai]
Kamohara & Yamakawa (1965, as Pomacentrus dorsalis): 1 unregistered specimen, 60 mm TL, Somachi.
Iwatsubo (2018): KAUM–I. 72313, 43.8 mm SL, Shiomichi. Iwatsubo (2019): KAUM–I. 72313, 43.8 mm
SL, Shiomichi. This study: KAUM–I. 72313, 43.8 mm SL, Shiomichi; KAUM–I. 134596, 63.7 mm SL, Onotsu; KAUM–I. 134746, 22.6 mm SL, Keraji; KAUM–I. 146706, 63.3 mm SL, Kamikatetsu.
Pomacentrus chrysurus Cuvier, 1830
[Jpn name: Ojirosuzumedai]
Yamakawa (1969, as Pomacentrus flavicauda): 4 unregistered specimens, 48–61 mm TL, Kikai Island. This
study: KAUM–I. 134612, 52.5 mm SL, Onotsu; KAUM–I. 134655, 12.4 mm SL, Nakasato; KAUM–I.
135083, 65.3 mm SL, Araki.
Pomacentrus lepidogenys Fowler & Bean, 1928
[Jpn name: Asadosuzumedai]
KAUM–I. 8098, 25.4 mm SL, Urahara.
Pomacentrus nagasakiensis Tanaka, 1917
[Jpn name: Nagasakisuzumedai]
KAUM–I. 135122, 45.5 mm SL, Araki; KAUM–I. 146507, 64.5 mm SL, Onotsu.
Pomacentrus philippinus Evermann & Seale, 1907
[Jpn name: Fuiripinsuzumedai]
KAUM–I. 134789, 31.8 mm SL, Keraji.
Pomacentrus taeniometopon Bleeker, 1852
[Jpn name: Sumizomesuzumedai]
KAUM–I. 146827, 70.6 mm SL, KAUM–I. 146828, 64.5 mm SL, Araki.
Pomachromis richardsoni (Snyder, 1909)[Jpn name: Okinawasuzumedai]
Kamohara & Yamakawa (1968b, as Abudefduf richardsoni): 1 unregistered specimen, 32 mm TL, Kikai Island.
Stegastes albifasciatus (Schlegel & Müller, 1840)[Jpn name: Kiobisuzumedai]
KAUM–I. 134860, 67.2 mm SL, KAUM–I. 134877, 77.1 mm SL, KAUM–I. 146543, 56.9 mm SL, Keraji;
KAUM–I. 135127, 16.3 mm SL, Araki; KAUM–I. 146482, 31.5 mm SL, Tekuzuku.
Stegastes fasciolatus (Ogilby, 1889)[Jpn name: Fuchidorisuzumedai]
Iwatsubo (2018): KAUM–I. 72329, 29.4 mm SL, Onotsu. Iwatsubo (2019): KAUM–I. 72329, 29.4 mm
SL, Onotsu. This study: KAUM–I. 72329, 29.4 mm SL, Onotsu; KAUM–I. 134757, 24.1 mm SL, KAUM–
I. 134865, 18.1 mm SL, Keraji; KAUM–I. 135082, 62.3 mm SL, KAUM–I. 146649, 60.3 mm SL, Araki;
KAUM–I. 146601, 52.4 mm SL, KAUM–I. 146610, 29.8 mm SL, Tekuzuku.
Stegastes insularis Allen & Emery, 1985
[Jpn name: Yoronsuzumedai]
KAUM–I. 134590, 45.3 mm SL, Onotsu.

Marine and freshwater fishes of Kikai Island

37

Stegastes nigricans (Lacepède, 1802)[Jpn name: Kurosorasuzumedai]
KAUM–I. 134861, 27.1 mm SL, KAUM–I. 134862, 34.7 mm SL, KAUM–I. 134879, 34.4 mm SL, Keraji;
KAUM–I. 135131, 15.0 mm SL, Araki.
Stegastes obreptus (Whitley, 1948)[Jpn name: Aisuzumedai]
KAUM–I. 135084, 54.2 mm SL, Araki.
Stegastes punctatus (Quoy & Gaimard, 1825)[Jpn name: Hananagasuzumedai]
KAUM–I. 134790, 61.9 mm SL, KAUM–I. 134791, 50.7 mm SL, Keraji.
FAMILY TERAPONTIDAE
Terapon jarbua (Forsskål, 1775)[Jpn name: Kotohiki]
KAUM–I. 134525, 38.7 mm SL, Akaren.
FAMILY KUHLIIDAE
Kuhlia mugil (Forster, 1801)[Jpn name: Gin-yugoi]
Nakamura (2018b): KAUM–I. 72198, 168.8 mm SL, Onotsu. Nakamura (2019b): KAUM–I. 72198, 168.8mm SL, Onotsu. This study: KAUM–I. 72198, 168.8 mm SL, KAUM–I. 72199, 175.6 mm SL, Onotsu;
KAUM–I. 134707, 19.1 mm SL, Shitooke.
Kuhlia rupestris (Lacepède, 1802)[Jpn name: Okuchiyugoi]
KAUM–I. 132978, 265.7 mm SL, KAUM–I. 132979, 212.0 mm SL, Isaneku; KAUM–I. 134709, 108.8 mm
SL, KAUM–I. 134710, 54.8 mm SL, KUN-P 55683, 44.4 mm SL, KUN-P 55684, 56.1 mm SL, KUN-P
55689, 39.4 mm SL, Urahara; KAUM–I. 134958, 50.8 mm SL, KAUM–I. 134959, 41.9 mm SL, Shiramizu;
KAUM–I. 135159, 34.8 mm SL, Shitooke.
FAMILY KYPHOSIDAE
Kyphosus cinerascens (Forsskål, 1775)[Jpn name: Tenjikuisaki]
KAUM–I. 135166, 116.0 mm SL, KAUM–I. 136999, 117.7 mm SL, Shiomichi.
Kyphosus vaigiensis (Quoy & Gaimard, 1825)[Jpn name: Isuzumi]
Kamohara (1957, as Kyphosus lembus): 1 unregistered specimen, 101 mm TL, Somachi. Nakamura (2018c):
KAUM–I. 72301, 201.4 mm SL, Kikai Island. Nakamura (2019c): KAUM–I. 72301, 201.4 mm SL, Kikai
Island. This study: KAUM–I. 72301, 201.4 mm SL, Kikai Island.
FAMILY MICROCANTHIDAE
Microcanthus strigatus (Cuvier, 1831)[Jpn name: Kagokakidai]
KPM-NI 26382, 110.9 mm SL, Kikai Island.

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K. Fujiwara and H. Motomura

FAMILY GIRELLIDAE
Girella leonina (Richardson, 1846)[Jpn name: Kuromejina]
Kamohara (1957, as Girella melanichthys): 4 unregistered specimens, 118–160 mm TL, Somachi. This
study: KAUM–I. 8585, 66.5 mm SL, Kikai Island.
Girella mezina Jordan & Starks, 1907
[Jpn name: Okinamejina]
Kamohara (1957): many unregistered specimens, 85–170 mm TL, Kikai-jjima Island (including Amami-oshima Island). Kamohara & Yamakawa (1965): Kikai Island. This study: KAUM–I. 87280, 129.5 mm SL,
Kikai Island.
FAMILY POLYNEMIDAE
Polydactylus sexfilis (Valenciennes, 1831)[Jpn name: Nan-yoagonashi]
KAUM–I. 134837, 72.7 mm SL, Somachi.
FAMILY LABRIDAE
Anampses caeruleopunctatus Rüppell, 1829
[Jpn name: Buchisusukibera]
Kamohara (1957): 2 unregistered specimens, 124–206 mm TL, Wan.
Anampses meleagrides Valenciennes, 1840
[Jpn name: Hokutobera]
Kamohara (1957): 1 unregistered specimen, 127 mm TL, Wan. This study: KAUM–I. 15382, 126.6 mm SL,
Sakamine.
Anampses twistii Bleeker, 1856
[Jpn name: Hoshisusukibera]
Kamohara (1957): 1 unregistered specimen, 115 mm TL, Onotsu.
Bodianus bilunulatus (Lacepède, 1801)[Jpn name: Kitsunebera]
Kamohara (1957, as Lepidaplois hirsutus): 1 unregistered specimen, 186 mm TL, Wan.
Bodianus perditio (Quoy & Gaimard, 1834)[Jpn name: Takibera]
KAUM–I. 146797, 100.5 mm SL, Akaren.
Cheilinus chlorourus (Bloch, 1791)[Jpn name: Akatemmochinouo]
Kamohara (1957, as Cheilinus chlorourus): 1 unregistered specimen, 94 mm TL, Somachi.
Cheilinus fasciatus (Bloch, 1791)[Jpn name: Yashabera] (Fig. 3D)
Underwater photograph, off Kamikatetsu.

Marine and freshwater fishes of Kikai Island

39

Cheilinus trilobatus Lacepède, 1801
[Jpn name: Mitsubamochinouo]
Fukui (2018): KAUM–I. 72498, 201.1 mm SL, Onotsu. Motomura (2019b): KAUM–I. 72498, 201.1 mm
SL, Onotsu. This study: KAUM–I. 72498, 201.1 mm SL, KAUM–I. 134591, 41.4 mm SL, Onotsu.
Choerodon fasciatus (Günther, 1867)[Jpn name: Shichisembera] (Fig. 3E)
Underwater photograph, off Araki.
Coris aygula Lacepède, 1801
[Jpn name: Kammuribera]
Hayashi (1995): YCM-P 30418, 86.5 mm SL, Kikai Island. Fukui (2018): KAUM–I. 72352, 86.3 mm SL,
Keraji. Motomura (2019b): KAUM–I. 72352, 86.3 mm SL, Keraji. This study: KAUM–I. 15521, 153.4mm SL, KAUM–I. 15523, 145.0 mm SL, KAUM–I. 15524, 143.5 mm SL, KAUM–I. 72352, 86.3 mm SL,
KAUM–I. 134881, 70.7 mm SL, Keraji; KAUM–I. 134716, 142.8 mm SL, Akaren.
Coris dorsomacula Fowler, 1908
[Jpn name: Sujibera]
KAUM–I. 146724, 41.9 mm SL, Tekuzuku.
Coris gaimard (Quoy & Gaimard, 1824)[Jpn name: Tsuyubera]
KAUM–I. 146609, 46.6 mm SL, Tekuzuku; KAUM–I. 146749, 45.3 mm SL, Keraji.
Gomphosus varius Lacepède, 1801
[Jpn name: Kugibera]
Kamohara (1957): 1 unregistered specimen, 48 mm TL, Somachi. Kamohara & Yamakawa (1965): 1 unregistered specimen, 128 mm TL, Somachi.
Halichoeres biocellatus Schultz, 1960
[Jpn name: Nishikikyusen]
KAUM–I. 15192, 90.3 mm SL, Keraji.
Halichoeres chrysus Randall, 1981
[Jpn name: Koganekyusen]
KAUM–I. 146479, 29.8 mm SL, Tekuzuku.
Halichoeres hortulanus (Lacepède, 1801)[Jpn name: Tokarabera]
Kamohara (1957, as Halichoeres centriquadrus): 1 unregistered specimen, 122 mm TL, Somachi. Kamohara& Yamakawa (1965, as Halichoeres centriquadrus): 6 unregistered specimens, 39–119 mm TL, Somachi.
Fukui (2018): KAUM–I. 72330, 44.3 mm SL, Onotsu. Motomura (2019b): KAUM–I. 72330, 44.3 mm SL,
Onotsu. This study: KAUM–I. 72330, 44.3 mm SL, Onotsu; KAUM–I. 146549, 21.0 mm SL, KAUM–I.
146741, 20.7 mm SL, Keraji.
Halichoeres margaritaceus (Valenciennes, 1839)[Jpn name: Akanijibera]
Kamohara (1957): 2 unregistered specimens, 48–55 mm TL, Wan; 3 unregistered specimens, 55–73 mm TL,
Somachi. This study: KAUM–I. 15530, 106.5 mm SL, Keraji.

40

K. Fujiwara and H. Motomura

Halichoeres marginatus Rüppell, 1835
[Jpn name: Kanokobera]
Kamohara (1957): 1 unregistered specimen, 162 mm TL, Somachi. Kamohara & Yamakawa (1965): 1 unregistered specimen, 165 mm TL, Somachi. This study: KAUM–I. 15363, 138.5 mm SL, KAUM–I. 15380,
106.9 mm SL, Sakamine; KAUM–I. 72356, 45.8 mm SL, Keraji; KAUM–I. 134649, 28.2 mm SL, KUN-P
55645, 34.6 mm SL, Nakasato; KAUM–I. 134944, 24.3 mm SL, Nakaguma.
Halichoeres melanochir Fowler & Bean, 1928
[Jpn name: Munatembera]
KAUM–I. 15402, 121.2 mm SL, Sakamine; KAUM–I. 72343, 141.8 mm SL, KAUM–I. 134804, 55.3 mm
SL, Keraji; KAUM–I. 134833, 38.0 mm SL, Shitooke.
Halichoeres melanurus (Bleeker, 1851)[Jpn name: Kazarikyusen]
KAUM–I. 15298, 79.1 mm SL, Sakamine.
Halichoeres prosopeion (Bleeker, 1853)[Jpn name: Munatemberadamashi]
Kamohara (1957): 1 unregistered specimen, 68 mm TL, Somachi.
Halichoeres scapularis (Bennett, 1832)[Jpn name: Seitembera]
Kamohara (1957): 1 unregistered specimen, 61 mm TL, Wan. Kamohara & Yamakawa (1965): 2 unregistered specimens, 55–62 mm TL, Araki; 5 unregistered specimens, 48–70 mm TL, Somachi.
Halichoeres tenuispinis (Günther, 1862)[Jpn name: Hombera]
Kamohara (1957): 1 unregistered specimen, 81 mm TL, Somachi. Shimada (2013): Kikai Island.
Halichoeres trimaculatus (Quoy & Gaimard, 1834)[Jpn name: Mitsuboshikyusen]
Kamohara & Yamakawa (1965): 1 unregistered specimen, 60 mm TL, Tekuzuku; 1 unregistered specimen,
73 mm TL, Araki. This study: KAUM–I. 135101, 42.5 mm SL, Araki; KAUM–I. 135027, 79.8 mm SL,
Shiomichi; KAUM–I. 146742, 26.3 mm SL, KAUM–I. 146756, 42.7 mm SL, Keraji.
Hemigymnus fasciatus (Bloch, 1792)[Jpn name: Shimatarekuchibera]
KPM-NI 26368, 169.7 mm SL, Kikai Island.
Hemigymnus melapterus (Bloch, 1791)[Jpn name: Tarekuchibera]
KUN-P 55760, 17.6 mm SL, Araki.
Hologymnosus annulatus (Lacepède, 1801)[Jpn name: Namerabera]
KAUM–I. 134928, 159.4 mm SL, Somachi.
Hologymnosus doliatus (Lacepède, 1801)[Jpn name: Shirotasukibera] (Fig. 3F)
Underwater photograph, off Tekuzuku.

Marine and freshwater fishes of Kikai Island

41

Labrichthys unilineatus (Guichenot, 1847)[Jpn name: Kurobera]
Fukui (2018): KAUM–I. 72369, 36.1 mm SL, Keraji. Motomura (2019b): KAUM–I. 72369, 36.1 mm SL,
Keraji. This study: KAUM–I. 72369, 36.1 mm SL, KAUM–I. 134800, 23.8 mm SL, KAUM–I. 134801, 20.4mm SL, KAUM–I. 134802, 27.8 mm SL, Keraji.
Labroides bicolor Fowler & Bean, 1928
[Jpn name: Somewakebera]
Kamohara (1957): 1 unregistered specimen, 112 mm TL, Wan.
Labroides dimidiatus (Valenciennes, 1839)[Jpn name: Honsomewakebera]
Kamohara (1957): 1 unregistered specimen, 72 mm TL, Somachi; 1 unregistered specimen, 65 mm TL, Wan.
Kamohara & Yamakawa (1965): 2 unregistered specimens, 46–53 mm TL, Tekuzuku; 1 unregistered specimen, 65 mm SL, Somachi. This study: KAUM–I. 146624, 14.5 mm SL, Tekuzuku.
Macropharyngodon meleagris (Valenciennes, 1839)[Jpn name: Nodogurobera]
Kamohara (1957, as Macropharyngodon pardalis): 1 unregistered specimen, 58 mm TL, Somachi. This
study: KAUM–I. 134614, 16.1 mm SL, Onotsu; KAUM–I. 146717, 39.0 mm SL, Tekuzuku.
Pteragogus sp.
[Jpn name: N/A]
Kamohara (1957, as Duymaeria flagellifera): 1 unregistered specimen, 85 mm TL, Somachi.
Remarks. The taxonomy of Pteragogus spp. distributed in the Ryukyu Islands is currently problematic, precluding a definitive species identification.
Pseudocheilinus hexataenia (Bleeker, 1857)[Jpn name: Nisemochinouo]
KAUM–I. 146544, 52.3 mm SL, Keraji.
Pseudocheilinus octotaenia Jenkins, 1901
[Jpn name: Yasujinisemochinouo]
KAUM–I. 146585, 15.6 mm SL, Kamikatetsu.
Pseudocoris yamashiroi (Schmidt, 1931)[Jpn name: Yamashirobera]
Hayashi (1995): YCM-P 30417, 107.7 mm SL, Kikai Island.
Stethojulis bandanensis (Bleeker, 1851)[Jpn name: Akaobibera]
Hayashi (1995): YCM-P 30420, 78.2 mm SL, Kikai Island. This study: KAUM–I. 135126, 46.3 mm SL,
Araki.
Stethojulis interrupta terina Jordan & Snyder, 1902
[Jpn name: Kaminaribera]
Kamohara (1957, as Stethojulis kalosoma): 1 unregistered specimen, 126 mm TL, Wan. Kamohara & Yamakawa (1965, as Stethojulis kalosoma): 1 unregistered specimen, 102 mm TL, Araki; 1 unregistered specimen, 102 mm TL, Somachi.

42

K. Fujiwara and H. Motomura

Stethojulis maculata Schmidt, 1931
[Jpn name: Sumitsukikaminaribera]
Hayashi (1995): YCM-P 30416, 114.3 mm SL, Kikai Island. This study: KAUM–I. 15529, 120.0 mm SL,
Keraji.
Stethojulis strigiventer (Bennett, 1833)[Jpn name: Harasujibera]
Kamohara & Yamakawa (1968a): 1 unregistered specimen, 85 mm TL, Kikai Island. This study: KAUM–I.
72197, 55.4 mm SL, Urahara.
Stethojulis trilineata (Bloch & Schneider, 1801)[Jpn name: Onibera]
Kamohara (1957, as Stethojulis phekadopleura): 5 unregistered specimens, 58–120 mm TL, Wan. Kamohara& Yamakawa (1968a): 1 unregistered specimen, 130 mm TL, Kikai Island.
Thalassoma cupido (Temminck & Schlegel, 1845)[Jpn name: Nishikibera]
Hayashi (1995): YCM-P 30419, 90.9 mm SL, Kikai Island. Fukui (2018, as Thalassoma trilobatum):
KAUM–I. 72339, 23.6 mm SL, Onotsu. Motomura (2019b): KAUM–I. 72339, 23.6 mm SL, Onotsu. This
study: KAUM–I. 19866, 76.7 mm SL, Keraji; KAUM–I. 72339, 23.6 mm SL, Onotsu.
Thalassoma hardwicke (Bennett, 1830)[Jpn name: Senasujibera]
Kamohara (1957): 1 unregistered specimen, 112 mm TL, Somachi. Kamohara & Yamakawa (1965): 2 unregistered specimens, 51–60 mm TL, Araki; 1 unregistered specimen, 108 mm TL, Somachi. Fukui (2018):
KAUM–I. 72375, 28.2 mm SL, KAUM–I. 72385, 49.2 mm SL, Keraji. Motomura (2019b): KAUM–I.
72375, 28.2 mm SL, KAUM–I. 72385, 49.2 mm SL, Keraji. This study: KAUM–I. 72375, 28.2 mm SL,
KAUM–I. 72381, 33.5 mm SL, KAUM–I. 72385, 49.2 mm SL, KAUM–I. 72386, 43.0 mm SL, Keraji.
Thalassoma lutescens (Lay & Bennett, 1839)[Jpn name: Yamabukibera]
Kamohara & Yamakawa (1965): 1 unregistered specimen, 88 mm TL, Somachi. Hayashi (1995): YCM-P
30415, 39.0 mm SL, Kikai Island. This study: KAUM–I. 15536, 123.6 mm SL, KAUM–I. 19865, 117.7mm SL, KAUM–I. 134871, 23.5 mm SL, Keraji; KAUM–I. 134662, 123.6 mm SL, KAUM–I. 134663, 95.6mm SL, KAUM–I. 134664, 82.8 mm SL, KAUM–I. 134929, 93.0 mm SL, KAUM–I. 146782, 154.6 mm
SL, KAUM–I. 146814, 149.4 mm SL, KAUM–I. 146815, 94.9 mm SL, KAUM–I. 147636, 147.3 mm SL,
KAUM–I. 147637, 104.6 mm SL, KAUM–I. 147638, 91.6 mm SL, KUN-P 55663, 94.3 mm SL, KUN-P
55666, 88.5 mm SL, Somachi; KAUM–I. 135068, 92.0 mm SL, Araki; KAUM–I. 146783, 101.3 mm SL,
KAUM–I. 146791, 83.1 mm SL, Akaren.
Thalassoma purpureum (Forsskål, 1775)[Jpn name: Kinubera]
KAUM–I. 72321, 22.4 mm SL, Somachi.
FAMILY SCARIDAE
Calotomus carolinus (Valenciennes, 1840)[Jpn name: Taiwambudai]
Sakurai (2018b): KAUM–I. 72467, 241.3 mm SL, Somachi. Sakurai (2019b): KAUM–I. 72467, 241.3 mm
SL, Somachi. This study: KAUM–I. 72467, 241.3 mm SL, Somachi; KAUM–I. 72489, 202.7 mm SL, Aden.

Marine and freshwater fishes of Kikai Island

43

Cetoscarus ocellatus (Valenciennes, 1840)[Jpn name: Irobudai]
Sakurai (2018b): KAUM–I. 72500, 343.7 mm SL, Onotsu. Sakurai (2019b): KAUM–I. 72500, 343.7 mm
SL, Onotsu. This study: KAUM–I. 72500, 343.7 mm SL, Onotsu; KAUM–I. 130225, 416.0 mm SL, Kikai
Island.
Chlorurus bowersi (Snyder, 1909)[Jpn name: Omonhagebudai] (Fig. 3G)
Underwater photograph, off Keraji.
Chlorurus frontalis (Valenciennes, 1840)[Jpn name: Onihagebudai]
Sakurai (2018b): KAUM–I. 72468, 177.0 mm SL, Somachi. Sakurai (2019b): KAUM–I. 72468, 177.0 mm
SL, Somachi. This study: KAUM–I. 72468, 177.0 mm SL, Somachi.
Chlorurus microrhinos (Bleeker, 1854)[Jpn name: Nan-yobudai]Kamohara (1957, as Chlorurus microrhinus): 1 unregistered specimen, 190 mm TL, Wan. This study:
KAUM–I. 72506, 151.7 mm SL, Onotsu.
Chlorurus spilurus (Valenciennes, 1840)[Jpn name: Hagebudai]Kamohara (1957, as Xanothon sordidus): 1 unregistered specimen, 305 mm TL, Wan. This study: KAUM–I.
134642, 274.1 mm SL, Onotsu; KAUM–I. 147621, 226.0 mm SL, Kamikatetsu.
Leptoscarus vaigiensis (Quoy & Gaimard, 1824)[Jpn name: Mizorebudai]Kamohara (1957, as Leptoscarus caeruleopunctatus): 1 unregistered specimen, 123 mm TL, Wan.
Scarus forsteni (Bleeker, 1861)[Jpn name: Ichimonjibudai]Kamohara (1957, as Callyodon forsteni): 1 unregistered specimen, 195 mm TL, Wan. Sakurai (2018b):
KAUM–I. 72456, 233.8 mm SL, Somachi. Sakurai (2019b): KAUM–I. 72456, 233.8 mm SL, Somachi. This
study: KAUM–I. 72456, 233.8 mm SL, Somachi; KAUM–I. 72488, 254.7 mm SL, Aden.
Scarus hypselopterus Bleeker, 1853
[Jpn name: Kibirebudai]
Sakurai (2018b): KAUM–I. 72508, 168.2 mm SL, Onotsu. Sakurai (2019b): KAUM–I. 72508, 168.2 mm
SL, Onotsu. This study: KAUM–I. 72508, 168.2 mm SL, Onotsu.
Scarus prasiognathos Valenciennes, 1840
[Jpn name: Nishikibudai]
KAUM–I. 72487, 162.6 mm SL, Aden.
Scarus psittacus Forsskål, 1775
[Jpn name: Omubudai]Kamohara (1957, as Xanothon oktodon): 1 unregistered specimen, 91 mm TL, Wan; 1 unregistered specimen,
119 mm TL, Somachi. Sakurai (2018b): KAUM–I. 72411, 157.9 mm SL, Keraji. Sakurai (2019b): KAUM–I.
72411, 157.9 mm SL, Keraji. This study: KAUM–I. 72411, 157.9 mm SL, KAUM–I. 146569, 44.8 mm SL,

44

K. Fujiwara and H. Motomura

Keraji; KAUM–I. 72471, 167.6 mm SL, KAUM–I. 72475, 219.1 mm SL, Somachi; KAUM–I. 72486, 185.1mm SL, Aden; KAUM–I. 135081, 210.0 mm SL, Araki; KAUM–I. 146630, 61.6 mm SL, Kamikatetsu.
Scarus rivulatus Valenciennes, 1840
[Jpn name: Sujibudai]
KAUM–I. 134917, 318.2 mm SL, Shiomichi.
Scarus rubroviolaceus Bleeker, 1847
[Jpn name: Nagabudai]Kamohara & Yamakawa (1965): 2 unregistered specimens, 343–345 mm TL, Somachi. This study: KAUM–
I. 72466, 326.9 mm SL, Shiramizu; KAUM–I. 146638, 26.4 mm SL, Kamikatetsu.
Scarus xanthopleura Bleeker, 1853
[Jpn name: Akabudai]
KAUM–I. 131313, 454.0 mm SL, Kikai Island.
FAMILY PINGUIPEDIDAE
Parapercis cylindrica (Bloch, 1792)[Jpn name: Dandaratoragisu]
Kamohara (1957, as Cilias synaphodesmus): 1 unregistered specimen, 113 mm TL, Somachi. This study:
KAUM–I. 134903, 73.3 mm SL, Shiomichi; KAUM–I. 135071, 96.1 mm SL, KAUM–I. 135072, 98.6 mm
SL, Araki; KAUM–I. 146557, 71.5 mm SL, KAUM–I. 146558, 72.9 mm SL, KAUM–I. 146566, 65.4 mm
SL, Keraji.
Parapercis millepunctata (Günther, 1860)[Jpn name: Wanuketoragisu]
KAUM–I. 134930, 103.5 mm SL, Somachi; KAUM–I. 146795, 124.0 mm SL, Akaren.
Parapercis pacifica Imamura & Yoshino, 2007
[Jpn name: Ogurotoragisu]
KAUM–I. 134530, 122.5 mm SL, KAUM–I. 134720, 97.5 mm SL, Akaren; KAUM–I. 146582, 68.7 mm SL,
Kamikatetsu; KAUM–I. 146823, 89.6 mm SL, Araki.
Parapercis xanthozona (Bleeker, 1849)[Jpn name: Ojirotoragisu]
Kamohara & Yamakawa (1968a): 1 unregistered specimen, 67 mm TL, Kikai Island.
FAMILY CREEDIIDAE
Limnichthys fasciatus Waite, 1904
[Jpn name: Tobigimpo]
Kamohara & Yamakawa (1968b): 3 unregistered specimens, 25–46 mm TL, Kikai Island. This study:
KAUM–I. 146587, 25.0 mm SL, Kamikatetsu.
FAMILY TRIPTERYGIIDAE
Enneapterygius etheostoma (Jordan & Snyder, 1902)[Jpn name: Hebigimpo]
Tashiro (2018): KAUM–I. 72325, 18.3 mm SL, Onotsu. Tashiro (2019): KAUM–I. 72325, 18.3 mm SL, Onotsu. This study: KAUM–I. 8092, 39.5 mm SL, Urahara; KAUM–I. 36037, 21.6 mm SL, Akaren; KAUM–I.
72325, 18.3 mm SL, Onotsu; KAUM–I. 135046, 35.2 mm SL, Araki.

Marine and freshwater fishes of Kikai Island

45

Enneapterygius flavoccipitis Shen, 1994
[Jpn name: Erimakihebigimpo]
Tashiro (2018): KAUM–I. 72345, 20.2 mm SL, Keraji. Tashiro (2019): KAUM–I. 72345, 20.2 mm SL, Keraji. This study: KAUM–I. 72345, 20.2 mm SL, Keraji; KAUM–I. 146527, 18.6 mm SL, Shiomichi; KAUM–
I. 146589, 18.5 mm SL, Kamikatetsu; KAUM–I. 146617, 16.3 mm SL, KAUM–I. 146726, 16.0 mm SL,
Tekuzuku.
Enneapterygius phoenicosoma Motomura, Ota & Meguro, 2015
[Jpn name: Akahebigimpo]
KAUM–I. 146752, 24.8 mm SL, Keraji.
Enneapterygius sp. 1[Jpn name: N/A]
Tashiro (2018): KAUM–I. 72307, 18.3 mm SL, Somachi. Tashiro (2019): KAUM–I. 72307, 18.3 mm SL,
Somachi. This study: KAUM–I. 72307, 18.3 mm SL, Somachi.
Enneapterygius sp. 2[Jpn name: N/A]
Tashiro (2018): KAUM–I. 72344, 23.5 mm SL, Keraji. Tashiro (2019): KAUM–I. 72344, 23.5 mm SL, Keraji. This study: KAUM–I. 72344, 23.5 mm SL, Keraji; KAUM–I. 146671, 18.5 mm SL, Araki; KAUM–I.
146694, 22.6 mm SL, Kamikatetsu.
Enneapterygius tutuilae Jordan & Seale, 1906
[Jpn name: Segurohebigimpo]
Tashiro (2018): KAUM–I. 72305, 19.4 mm SL, Somachi. Tashiro (2019): KAUM–I. 72305, 19.4 mm SL,
Somachi. This study: KAUM–I. 72305, 19.4 mm SL, Somachi; KAUM–I. 72346, 19.8 mm SL, KAUM–I.
72383, 19.4 mm SL, KAUM–I. 72384, 19.6 mm SL, KAUM–I. 134781, 17.8 mm SL, KAUM–I. 134782,
18.5 mm SL, KAUM–I. 134805, 18.4 mm SL, Keraji; KAUM–I. 134615, 18.1 mm SL, KAUM–I. 134620,
19.6 mm SL, Onotsu.
Helcogramma fuscipectoris (Fowler, 1946)[Jpn name: Kuromasuku]
Tashiro (2018): KAUM–I. 72323, 14.6 mm SL, KAUM–I. 72324, 18.2 mm SL, Onotsu. Tashiro (2019):
KAUM–I. 72323, 14.6 mm SL, KAUM–I. 72324, 18.2 mm SL, Onotsu. This study: KAUM–I. 72323, 14.6mm SL, KAUM–I. 72324, 18.2 mm SL, Onotsu.
Helcogramma inclinata (Fowler, 1946)[Jpn name: Ayahebigimpo]
KAUM–I. 72306, 42.7 mm SL, Somachi.
Norfolkia brachylepis (Schultz, 1960)[Jpn name: Kokutennisehebigimpo]
Tashiro (2018): KAUM–I. 72303, 38.1 mm SL, KAUM–I. 72304, 37.4 mm SL, Somachi; KAUM–I. 72354,
10.5 mm SL, Keraji. Tashiro (2019): KAUM–I. 72303, 38.1 mm SL, KAUM–I. 72304, 37.4 mm SL, Somachi; KAUM–I. 72354, 10.5 mm SL, Keraji. This study: KAUM–I. 72303, 38.1 mm SL, KAUM–I. 72304,
37.4 mm SL, Somachi; KAUM–I. 72354, 10.5 mm SL, Keraji.
Norfolkia thomasi Whitley, 1964
[Jpn name: Nisehebigimpo]
KAUM–I. 72308, 23.5 mm SL, Somachi.

46

K. Fujiwara and H. Motomura

FAMILY BLENNIIDAE
Aspidontus taeniatus Quoy & Gaimard, 1834
[Jpn name: Nisekurosujigimpo]
KAUM–I. 146695, 91.7 mm SL, Kamikatetsu.
Atrosalarias holomelas (Günther, 1872)[Jpn name: Indokaeruo]
Kawama (2018): KAUM–I. 72372, 42.2 mm SL, Keraji. Kawama (2019): KAUM–I. 72372, 42.2 mm SL,
Keraji. This study: KAUM–I. 72372, 42.2 mm SL, Keraji.
Blenniella bilitonensis (Bleeker, 1858)[Jpn name: Hohogurogimpo]
Kamohara (1957, as Istiblennius bilitonensis): 1 unregistered specimen, 122 mm TL, Somachi.
Blenniella periophthalmus (Valenciennes, 1836)[Jpn name: Hanakaeruo]
Kamohara & Yamakawa (1965, as Istiblennius periophthalmus): 1 unregistered specimen, 115 mm TL, Somachi.
Cirripectes castaneus (Valenciennes, 1836)[Jpn name: Tategamikaeruo]
KAUM–I. 146476, 52.0 mm SL, KAUM–I. 146477, 27.8 mm SL, KAUM–I. 146619, 45.6 mm SL, KAUM–
I. 146765, 43.5 mm SL, KAUM–I. 146767, 35.6 mm SL, Tekuzuku; KAUM–I. 146682, 43.1 mm SL, Araki.
Cirripectes imitator Williams, 1985
[Jpn name: Amimeminokaeruo]
KAUM–I. 146681, 60.8 mm SL, Araki; KAUM–I. 146704, 79.1 mm SL, Keraji.
Crossosalarias macrospilus Smith-Vaniz & Springer, 1971
[Jpn name: Erigurogimpo]
KAUM–I. 72188, 39.0 mm SL, Urahara; KAUM–I. 146517, 39.7 mm SL, Shiomichi.
Ecsenius bicolor (Day, 1888)[Jpn name: Futairokaeruo]
KAUM–I. 134765, 29.7 mm SL, KAUM–I. 134766, 29.5 mm SL, KAUM–I. 146755, 40.0 mm SL, Keraji;
KAUM–I. 146486, 36.7 mm SL, Tekuzuku.
Ecsenius oculus Springer, 1971
[Jpn name: Goishigimpo]
KAUM–I. 146512, 23.2 mm SL, Tekuzuku; KAUM–I. 146555, 34.5 mm SL, Keraji.
Ecsenius yaeyamaensis (Aoyagi, 1954)[Jpn name: Ishigakikaeruo]
Kamohara & Yamakawa (1965, as Istiblennius yaeyamaensis): 1 unregistered specimen, 44 mm TL, Somachi. Yamakawa (1969, as Istiblennius yaeyamaensis): 1 unregistered specimen, 23 mm TL, Kikai Island.
Kawama (2018): KAUM–I. 72363, 38.0 mm SL, Keraji. Kawama (2019): KAUM–I. 72363, 38.0 mm SL,
Keraji. This study: KAUM–I. 72363, 38.0 mm SL, KAUM–I. 134783, 32.4 mm SL, KAUM–I. 134784, 21.6mm SL, KAUM–I. 134818, 48.3 mm SL, KAUM–I. 134819, 32.5 mm SL, KAUM–I. 134885, 36.8 mm SL,
KAUM–I. 146753, 30.0 mm SL, Keraji; KAUM–I. 146497, 43.7 mm SL, Tekuzuku; KAUM–I. 146576,
32.5 mm SL, Kamikatetsu.

Marine and freshwater fishes of Kikai Island

47

Enchelyurus kraussii (Klunzinger, 1871)[Jpn name: Kurogimpo]
KUN-P 55649, 27.7 mm SL, Nakasato.
Entomacrodus decussatus (Bleeker, 1858)[Jpn name: Kikaikaeruo]
Aoyagi (1954, as Salarias kikaiensis): YCM-P 30815, 112.6 mm SL, YCM-P 308161, 2 specimens, 94.4–
101.7 mm SL, Kikai Island. Kamohara & Yamakawa (1968a, as Istiblennius kikaiensis): 1 unregistered specimen, 100 mm TL, Kikai Island. Hayashi (1995, as Istiblennius chrysospilos): YCM-P 30815, 112.6 mm SL,
YCM-P 308161, 2 specimens, 94.4–101.7 mm SL, Kikai Island.
Entomacrodus niuafoouensis (Fowler, 1932)[Jpn name: Keshogimpo]
KPM-NI 41248, 41.9 mm SL, Kikai Island.
Exallias brevis (Kner, 1868)[Jpn name: Sedakagimpo]
KAUM–I. 15535, 88.0 mm SL, Keraji; KAUM–I. 146475, 41.7 mm SL, Tekuzuku.
Istiblennius edentulus (Forster & Schneider, 1801)[Jpn name: Nisekaeruo]
KAUM–I. 135043, 87.7 mm SL, KAUM–I. 135044, 72.7 mm SL, KAUM–I. 135045, 29.0 mm SL, KAUM–
I. 135100, 77.6 mm SL, Araki; KUN-P 55647, 38.7 mm SL, KUN-P 55648, 24.5 mm SL, Nakasato; KUN-P
55741, 21.4 mm SL, Aden.
Istiblennius lineatus (Valenciennes, 1836)[Jpn name: Senkaeruo]
Kamohara (1957): 2 unregistered specimens, 90–97 mm TL, Wan. This study: KAUM–I. 134981, 77.9 mm
SL, Aden; KAUM–I. 135041, 80.3 mm SL, KAUM–I. 135042, 62.3 mm SL, Araki.
Meiacanthus atrodorsalis (Günther, 1877)[Jpn name: Ogonnijigimpo]
Kawama (2018): KAUM–I. 72311, 72.5 mm SL, Somachi. Kawama (2019): KAUM–I. 72311, 72.5 mm SL,
Somachi. This study: KAUM–I. 72311, 72.5 mm SL, Somachi.
Meiacanthus kamoharai Tomiyama, 1956
[Jpn name: Kamoharagimpo]
Kamohara (1957): 1 unregistered specimen, 89 mm TL, Wan. Kamohara & Yamakawa (1965): 1 unregistered specimen, 100 mm TL, Somachi. This study: KAUM–I. 15532, 79.8 mm SL, Keraji.
Nannosalarias nativitatis (Regan, 1909)[Jpn name: Hinagimpo]
KPM-NI 41250, 25.7 mm SL, Kikai Island.
Omobranchus elongatus (Peters, 1855)[Jpn name: Gomakumogimpo]
Aoyagi (1955, as Petroscirtes elegans): YCM-P 31147, 36.8 mm SL, Kikai Island. Hayashi (1995): YCM-P
31147, 36.8 mm SL, Kikai Island.

48

K. Fujiwara and H. Motomura

Omobranchus loxozonus (Jordan & Starks, 1906)[Jpn name: Kumogimpo]
Kamohara (1957): 1 unregistered specimen, 45 mm TL, Wan. This study: KAUM–I. 134980, 48.7 mm SL,
KAUM–I. 134996, 43.9 mm SL, KAUM–I. 134997, 33.9 mm SL, Aden.
Parablennius yatabei (Jordan & Snyder, 1900)[Jpn name: Isogimpo]
Aoyagi (1955, as Blennius tasmanianus): YCM-P 31147, 36.8 mm SL, Kikai Island. Hayashi (1995): YCM-P
31446, 62.4 mm SL, Kikai Island.
Parenchelyurus hepburni (Snyder, 1908)[Jpn name: Karasugimpo]
Yamakawa (1971, as Omobrachus hepburni): 1 unregistered specimen, 26 mm TL, Kikai Island. This study:
KAUM–I. 135049, 26.0 mm SL, KAUM–I. 135050, 36.2 mm SL, Araki.
Petroscirtes breviceps (Valenciennes, 1836)[Jpn name: Nijigimpo]
Kamohara (1957, as Dasson trossulus): 1 unregistered specimen, 91 mm TL, Wan.
Petroscirtes mitratus Rüppell, 1830
[Jpn name: Hatatategimpo]
KAUM–I. 134863, 13.6 mm SL, Keraji.
Plagiotremus tapeinosoma (Bleeker, 1857)[Jpn name: Tenkurosujigimpo]
Kamohara & Yamakawa (1965, as Aspidontus tapeinosoma): 1 unregistered specimen, 69 mm TL, Somachi.
This study: KAUM–I. 146550, 84.4 mm SL, KAUM–I. 146551, 75.5 mm SL, Keraji.
Praealticus bilineatus (Peters, 1868)[Jpn name: Tamagimpo]
Kamohara (1957, as Istiblennius margaritarius): 2 unregistered specimens, 44–81 mm TL, Wan; 1 unregistered specimen, 75 mm TL, Onotsu. Kawama (2018): KAUM–I. 72189, 37.9 mm SL, Urahara. Kawama(2019): KAUM–I. 72189, 37.9 mm SL, Urahara. This study: KAUM–I. 72189, 37.9 mm SL, Urahara;
KAUM–I. 134972, 56.3 mm SL, KAUM–I. 134973, 59.1 mm SL, KAUM–I. 134974, 55.1 mm SL, KAUM–
I. 134975, 51.2 mm SL, Aden; KAUM–I. 135047, 52.4 mm SL, KAUM–I. 135048, 49.5 mm SL, Araki.
Praealticus tanegasimae (Jordan & Starks, 1906)[Jpn name: Tanegimpo]
Hayashi (1995): YCM-P 31290, 2 specimens, 64.3–66.2 mm SL, Kikai Island. KAUM–I. 134970, 49.8 mm
SL, KAUM–I. 134976, 55.1 mm SL, KAUM–I. 134977, 60.9 mm SL, KAUM–I. 134978, 49.8 mm SL,
KAUM–I. 134979, 51.9 mm SL, Aden.
Rhabdoblennius nitidus (Günther, 1861)[Jpn name: Rosokugimpo]
KPM-NI 41249, 33.5 mm SL, Kikai Island.
Salarias fasciatus (Bloch, 1786)[Jpn name: Yaeyamagimpo]
Kamohara & Yamakawa (1965): 4 unregistered specimens, 73–144 mm TL, Tekuzuku; 1 unregistered specimen, 145 mm TL, Araki; 3 unregistered specimens, 107–110 mm TL, Somachi.

Marine and freshwater fishes of Kikai Island

49

Salarias luctuosus Whitley, 1929
[Jpn name: Shimagimpo]
KAUM–I. 8109, 52.2 mm SL, KAUM–I. 8110, 32.2 mm SL, Urahara; KAUM–I. 134688, 25.7 mm SL,
Shiomichi; KAUM–I. 134820, 25.8 mm SL, KAUM–I. 146570, 31.3 mm SL, KAUM–I. 146754, 38.6mm SL, Keraji; KAUM–I. 134937, 52.6 mm SL, KAUM–I. 134943, 49.1 mm SL, Nakaguma; KAUM–I.
135039, 43.1 mm SL, KAUM–I. 135040, 34.4 mm SL, Araki; KUN-P 55694, 35.9 mm SL, KUN-P 55695,
32.1 mm SL, Nakasato; KUN-P 55747, 39.9 mm SL, KUN-P 55748, 27.0 mm SL, KUN-P 55749, 22.7 mm
SL, Aden.
Salarias sinuosus Snyder, 1908
[Jpn name: Hirenagakaeruo]
Kamohara & Yamakawa (1965, as Negoscartes sinuosus): 1 unregistered specimen, 62 mm TL, Somachi.
Xiphasia matsubarai Okada & Suzuki, 1952
[Jpn name: Himeunagigimpo]
Shibuya & Motomura (2020): KAUM–I. 134670, 249.7 mm SL, Onotsu. This study: KAUM–I. 134670,
249.7 mm SL, Onotsu.
FAMILY GOBIESOCIDAE
Lepadichthys misakius (Tanaka, 1908)[Jpn name: Misakiubauo]
Kamohara & Yamakawa (1965, as Lepadichthys frenatus): 1 unregistered specimen, 44 mm TL, Somachi.
FAMILY ELEOTRIDAE
Eleotris acanthopoma Bleeker, 1853
[Jpn name: Chichibumodoki]Akihito (1967): 1 unregistered specimen, 41 mm SL, Akaren. This study: KAUM–I. 134578, 54.9 mm SL,
KAUM–I. 134579, 35.4 mm SL, KAUM–I. 134580, 35.9 mm SL, KAUM–I. 134581, 22.2 mm SL, KAUM–
I. 134582, 20.9 mm SL, KAUM–I. 134583, 20.2 mm SL, Shitooke.
Eleotris fusca (Bloch & Schneider, 1801)[Jpn name: Tenjikukawaanago]
KAUM–I. 134584, 90.2 mm SL, Shitooke; KAUM–I. 134961, 57.2 mm SL, KAUM–I. 134962, 51.5 mm
SL, KAUM–I. 134963, 99.8 mm SL, Shiramizu.
Eleotris melanosoma Bleeker, 1853
[Jpn name: Okamehaze]
KAUM–I. 134907, 12.2 mm SL, Shiomichi.
FAMILY XENISTHMIDAE
Xenisthmus clarus (Jordan & Seale, 1906)[Jpn name: Yanagihaze]
Kamohara & Yamakawa (1968b): 1 unregistered specimen, 45 mm TL, Kikai Island.
Xenisthmus sp. 1[Jpn name: Kuroyanagihaze]
KAUM–I. 134908, 20.8 mm SL, KAUM–I. 134909, 21.4 mm SL, Shiomichi.

50

K. Fujiwara and H. Motomura

FAMILY GOBIIDAE
Amblygobius phalaena (Valenciennes, 1837)[Jpn name: Sarasahaze]
KAUM–I. 134826, 90.0 mm SL, Nakasato; KAUM–I. 136730, 74.3 mm SL, KAUM–I. 146534, 77.4 mm
SL, Shiomichi; KAUM–I. 146562, 41.2 mm SL, KAUM–I. 146750, 73.1 mm SL, Keraji; KAUM–I. 146595,
61.5 mm SL, Tekuzuku.
Asterropteryx semipunctata Rüppell, 1830
[Jpn name: Hoshihaze]
KAUM–I. 8097, 32.4 mm SL, KAUM–I. 8118, 26.8 mm SL, Urahara; KAUM–I. 134815, 24.8 mm SL,
KAUM–I. 146561, 26.4 mm SL, Keraji; KAUM–I. 134988, 24.3 mm SL, KAUM–I. 134989, 27.3 mm SL,
KAUM–I. 134990, 22.2 mm SL, KUN-P 55742, 30.1 mm SL, Aden; KAUM–I. 135129, 24.6 mm SL, Araki.
Bathygobius coalitus (Bennett, 1832)[Jpn name: Kuroyahazuhaze]
KAUM–I. 134829, 68.2 mm SL, Nakasato; KAUM–I. 134946, 19.5 mm SL, Nakaguma; KAUM–I. 134982,
77.8 mm SL, KAUM–I. 134983, 51.8 mm SL, KAUM–I. 134984, 52.4 mm SL, KAUM–I. 134985, 59.4mm SL, KAUM–I. 134986, 48.4 mm SL, KAUM–I. 134987, 26.9 mm SL, Aden; KAUM–I. 135057, 62.6mm SL, KAUM–I. 135102, 66.4 mm SL, KAUM–I. 135103, 26.4 mm SL, KAUM–I. 135104, 26.0 mm SL,
Araki; KUN-P 55643, 34.5 mm SL, KUN-P 55644, 40.8 mm SL, Nakasato; KUN-P 55778, 54.1 mm SL,
Shiomichi.
Bathygobius cocosensis (Bleeker, 1854)[Jpn name: Sujikumohaze]
Hagiwara (2018): KAUM–I. 72194, 35.0 mm SL, Urahara. Hagiwara (2019): KAUM–I. 72194, 35.0 mm
SL, Urahara. This study: KAUM–I. 8095, 38.7 mm SL, KAUM–I. 72194, 35.0 mm SL, Urahara; KAUM–
I. 134866, 32.9 mm SL, Isago; KAUM–I. 135058, 33.8 mm SL, KAUM–I. 135059, 31.5 mm SL, Araki;
KUN-P 55642, 15.7 mm SL, Nakasato; KAUM–I. 146560, 29.8 mm SL, KAUM–I. 146563, 26.6 mm SL,
Keraji.
Bathygobius cyclopterus (Valenciennes, 1837)[Jpn name: Yahazuhaze]
KAUM–I. 134659, 24.9 mm SL, KAUM–I. 134660, 22.4 mm SL, Nakasato; KAUM–I. 134700, 56.3 mm
SL, KAUM–I. 134701, 44.5 mm SL, Shiomichi; KAUM–I. 135060, 39.7 mm SL, Araki.
Bathygobius fuscus (Rüppell, 1830)[Jpn name: Kumohaze]
KAUM–I. 134576, 32.4 mm SL, Shitooke; KAUM–I. 134656, 31.5 mm SL, KAUM–I. 134657, 23.7 mm
SL, Nakasato; KAUM–I. 135105, 22.7 mm SL, KAUM–I. 135106, 21.1 mm SL, KAUM–I. 135107, 27.0mm SL, KAUM–I. 135108, 24.8 mm SL, KAUM–I. 135109, 31.4 mm SL, Araki; KAUM–I. 135110, 31.0mm SL, Keraji.
Cabillus tongarevae (Fowler, 1927)[Jpn name: Yorimehaze]
KAUM–I. 134576, 23.7 mm SL, Keraji.
Callogobius hasseltii (Bleeker, 1851)[Jpn name: Okinawahaze]Marine and freshwater fishes of Kikai Island

51

Kamohara (1957): 1 unregistered specimen, 61 mm TL, Wan. This study: KAUM–I. 146503, 20.2 mm SL,
Onotsu; KAUM–I. 146567, 29.2 mm SL, KAUM–I. 146568, 22.6 mm SL, Keraji; KUN-P 55758, 24.0 mm
SL, Araki; KUN-P 55788, 34.8 mm SL, Tekuzuku.
Callogobius maculipinnis (Fowler, 1918)[Jpn name: Katahohaze]
KAUM–I. 134616, 40.6 mm SL, KAUM–I. 134641, 28.8 mm SL, Onotsu; KAUM–I. 134814, 23.3 mm SL,
KAUM–I. 134886, 39.7 mm SL, Keraji.
Callogobius okinawae (Snyder, 1908)[Jpn name: Namerahaze]
KAUM–I. 134703, 42.5 mm SL, KAUM–I. 134910, 26.3 mm SL, Shiomichi; KAUM–I. 135051, 33.3 mm
SL, Araki.
Callogobius tanegasimae (Snyder, 1908)[Jpn name: Tanehaze]
KAUM–I. 134571, 50.5 mm SL, Shitooke.
Cryptocentrus albidorsus (Yanagisawa, 1978)[Jpn name: Shirobihaze]
Hagiwara (2018): KAUM–I. 72362, 62.5 mm SL, Keraji. Hagiwara (2019): KAUM–I. 72362, 62.5 mm SL,
Keraji. This study: KAUM–I. 72362, 62.5 mm SL, Keraji.
Cryptocentrus nigrocellatus (Yanagisawa, 1978)[Jpn name: Kurohoshihaze]
Yanagisawa (1978, as Mars nigrocellatus): SMBL 293, 65.7 mm SL, SMBL 294, 65.7 mm SL, Shiramizu.
Eviota abax (Jordan & Snyder, 1901)[Jpn name: Isohaze]
Kamohara (1957, as Eviota abax abax): 3 unregistered specimens, 24–28 mm TL, Wan. This study: KUN-P
55639, 13.1 mm SL, KUN-P 55641, 15.6 mm SL, KUN-P 55698, 22.0 mm SL, Nakasato.
Eviota distigma Jordan & Seale, 1906
[Jpn name: Kobitoisohaze]
KAUM–I. 134617, 11.1 mm SL, KAUM–I. 134618, 13.5 mm SL, Onotsu.
Eviota fasciola Karnella & Lachner, 1981
[Jpn name: Toranokoisohaze]
KAUM–I. 146680, 12.6 mm SL, Araki.
Eviota japonica Jewett & Lachner, 1983
[Jpn name: Minamiisohaze]
KAUM–I. 8102, 17.5 mm SL, KAUM–I. 8103, 15.0 mm SL, KAUM–I. 8409, 18.4 mm SL, KAUM–I. 8410,
19.2 mm SL, KAUM–I. 72191, 18.4 mm SL, Urahara; KAUM–I. 134684, 18.4 mm SL, Shiomichi; KAUM–
I. 134775, 14.0 mm SL, KAUM–I. 134776, 12.6 mm SL, KAUM–I. 146736, 13.1 mm SL, Keraji; KAUM–
I. 134827, 19.2 mm SL, KAUM–I. 134828, 13.3 mm SL, Nakasato; KAUM–I. 134947, 17.7 mm SL, Nakaguma; KAUM–I. 135053, 18.2 mm SL, KAUM–I. 135054, 14.7 mm SL, KAUM–I. 135055, 13.5 mm SL,
KAUM–I. 135056, 9.8 mm SL, KAUM–I. 135098, 17.1 mm SL, KAUM–I. 135099, 13.7 mm SL, Araki;
KAUM–I. 146643, 9.4 mm SL, Kamikatetsu; KAUM–I. 146719, 13.1 mm SL, Tekuzuku.

52

K. Fujiwara and H. Motomura

Eviota latifasciata Jewett & Lachner, 1983
[Jpn name: Obiisohaze]
KAUM–I. 134777, 11.5 mm SL, Keraji; KAUM–I. 146675, 11.1 mm SL, Araki.
Eviota melasma Lachner & Karnella, 1980
[Jpn name: Akahoshiisohaze]
KAUM–I. 146554, 18.6 mm SL, Keraji.
Eviota pellucida Larson, 1976
[Jpn name: Kojikaisohaze]
Hagiwara (2018): KAUM–I. 72376, 21.3 mm SL, Keraji. Hagiwara (2019): KAUM–I. 72376, 21.3 mm SL,
Keraji. This study: KAUM–I. 72376, 21.3 mm SL, KAUM–I. 72377, 26.1 mm SL, KAUM–I. 134773, 20.9mm SL, KAUM–I. 134774, 19.2 mm SL, KAUM–I. 134809, 17.3 mm SL, KAUM–I. 146732, 13.1 mm SL,
KAUM–I. 146733, 17.9 mm SL, KAUM–I. 146734, 18.6 mm SL, KAUM–I. 146735, 16.7 mm SL, Keraji;
KAUM–I. 146532, 22.0 mm SL, KAUM–I. 146536, 12.4 mm SL, KAUM–I. 146537, 14.0 mm SL, KAUM–
I. 146538, 16.6 mm SL, KAUM–I. 146539, 15.7 mm SL, Shiomichi; KAUM–I. 146590, 14.0 mm SL,
KAUM–I. 146592, 17.3 mm SL, Kamikatetsu; KAUM–I. 146721, 19.7 mm SL, Tekuzuku.
Eviota perspicilla Fujiwara, Suzuki & Motomura, 2019
[Jpn name: Hoderiisohaze]
KAUM–I. 146495, 13.6 mm SL, Tekuzuku; KAUM–I. 146737, 11.3 mm SL, Keraji.
Eviota prasina (Klunzinger, 1871)[Jpn name: Nan-yomidorihaze]
Hagiwara (2018): KAUM–I. 72190, 20.6 mm SL, Urahara. Hagiwara (2019): KAUM–I. 72190, 20.6 mm
SL, Urahara. Fujiwara et al. (2019): KAUM–I. 72190, 20.6 mm SL, Urahara. This study: KAUM–I. 72190,
20.6 mm SL, Urahara; KAUM–I. 146700, 10.9 mm SL, Kamikatetsu; KUN-P 55691, 16.5 mm SL, KUN-P
55692, 18.0 mm SL, KUN-P 55693, 16.5 mm SL, KUN-P 55696, 17.4 mm SL, KUN-P 55702, 16.6 mm
SL, KUN-P 55703, 14.7 mm SL, KUN-P 55704, 13.0 mm SL, KUN-P 55705, 12.4 mm SL, KUN-P 55706,
12.3 mm SL, Nakasato; KUN-P 55750, 14.1 mm SL, KUN-P 55751, 14.5 mm SL, KUN-P 55752, 11.5 mm
SL, KUN-P 55753, 12.2 mm SL, KUN-P 55754, 11.1 mm SL, Aden; KUN-P 55756, 14.1 mm SL, KUN-P
55757, 15.1 mm SL, KUN-P 55761, 11.6 mm SL, KUN-P 55762, 10.1 mm SL, KUN-P 55763, 13.1 mm SL,
KUN-P 55764, 11.8 mm SL, Araki.
Eviota toshiyuki Greenfield & Randall, 2010
[Jpn name: Midorihaze]
KAUM–I. 134772, 13.8 mm SL, Keraji; KAUM–I. 146673, 13.0 mm SL, Araki.
Fusigobius duospilus Hoese & Reader, 1985
[Jpn name: Sehoshisankakuhaze]
KAUM–I. 134769, 32.0 mm SL, KAUM–I. 134770, 33.5 mm SL, KAUM–I. 134887, 45.1 mm SL, Keraji;
KAUM–I. 146492, 29.9 mm SL, KAUM–I. 146493, 26.5 mm SL, Tekuzuku.
Fusigobius gracilis (Randall, 2001)[Jpn name: Sesujisankakuhaze]
KAUM–I. 134888, 31.3 mm SL, Keraji; KAUM–I. 146491, 33.8 mm SL, Tekuzuku; KAUM–I. 146524,
31.5 mm SL, KAUM–I. 146529, 22.5 mm SL, Shiomichi; KAUM–I. 146580, 28.6 mm SL, Kamikatetsu.

Marine and freshwater fishes of Kikai Island

53

Fusigobius humeralis (Randall, 2001)[Jpn name: Kataboshisankakuhaze]
KAUM–I. 72379, 24.8 mm SL, Keraji; KAUM–I. 146530, 24.0 mm SL, Shiomichi; KAUM–I. 146644, 17.5mm SL, Kamikatetsu.
Fusigobius neophytus (Günther, 1877)[Jpn name: Sankakuhaze]
Kamohara (1957, as Gobius neophytus): 2 unregistered specimens, 40–56 mm TL, Wan; 2 unregistered specimens, 47–68 mm TL, Somachi. This study: KAUM–I. 8408, 23.8 mm SL, KAUM–I. 8094, 45.7 mm SL,
KAUM–I. 8099, 48.1 mm SL, KAUM–I. 8100, 30.0 mm SL, Urahara; KAUM–I. 134638, 50.4 mm SL, Onotsu; KAUM–I. 134658, 23.7 mm SL, KUN-P 55700, 38.9 mm SL, KUN-P 55701, 28.0 mm SL, Nakasato;
KAUM–I. 134821, 30.7 mm SL, KAUM–I. 146751, 52.2 mm SL, Keraji; KAUM–I. 134945, 19.5 mm SL,
Nakaguma; KAUM–I. 135124, 24.3 mm SL, KUN-P 55759, 30.2 mm SL, Araki; KAUM–I. 136708, 50.0mm SL, KAUM–I. 136709, 41.4 mm SL, Shiomichi; KUN-P 55743, 44.4 mm SL, KUN-P 55744, 31.7 mm
SL, KUN-P 55745, 33.5 mm SL, KUN-P 55746, 33.4 mm SL, Aden.
Gnatholepis anjerensis (Bleeker, 1851)[Jpn name: Omonhaze]
Kamohara (1957, as Gobius knighti): 1 unregistered specimen, 53 mm TL, Somachi. Kamohara & Yamakawa (1965, as Gobius knighti): 1 unregistered specimen, 45 mm TL, Araki; 1 unregistered specimen, 35 mm
SL, Somachi. This study: KAUM–I. 134864, 21.9 mm SL, Keraji.
Gnatholepis cauerensis (Bleeker, 1853)[Jpn name: Kataboshiomonhaze]
Hagiwara (2018): KAUM–I. 72378, 26.7 mm SL, Keraji. Hagiwara (2019): KAUM–I. 72378, 26.7 mm SL,
Keraji. This study: KAUM–I. 72378, 26.7 mm SL, KAUM–I. 134771, 27.9 mm SL, KAUM–I. 134807,
27.4 mm SL, KAUM–I. 134808, 30.2 mm SL, Keraji; KAUM–I. 134834, 31.1 mm SL, Shitooke; KAUM–I.
146521, 35.4 mm SL, KAUM–I. 146522, 28.6 mm SL, KAUM–I. 146528, 23.1 mm SL, Shiomichi; KUN-P
55699, 36.4 mm SL, Nakasato.
Gobiodon heterospilos Bleeker, 1856
[Jpn name: Ichimonjikobanhaze]
KAUM–I. 146471, 17.1 mm SL, KAUM–I. 146725, 14.3 mm SL, Tekuzuku.
Gobiodon quinquestrigatus (Valenciennes, 1837)[Jpn name: Futairosangohaze]
KAUM–I. 134812, 20.7 mm SL, KAUM–I. 134813, 19.9 mm SL, Keraji; KAUM–I. 146472, 16.0 mm SL,
KAUM–I. 146485, 22.8 mm SL, Tekuzuku.
Gobiodon rivulatus (Rüppell, 1830)[Jpn name: Tasujikobanhaze]
Kamohara & Yamakawa (1968b): 1 unregistered specimen, 31 mm TL, Kikai Island. This study: KAUM–I.
146469, 28.1 mm SL, KAUM–I. 146470, 21.0 mm SL, Tekuzuku; KAUM–I. 146676, 20.6 mm SL, Araki.
Gobiopsis arenaria (Snyder, 1908)[Jpn name: Isagohaze]
Kamohara & Yamakawa (1968b, as Pipidonia arenarius): 1 unregistered specimen, 36 mm TL, Kikai Island.

54

K. Fujiwara and H. Motomura

Istigobius decoratus (Herre, 1927)[Jpn name: Hoshikazarihaze]
KAUM–I. 134640, 48.2 mm SL, Onotsu; KAUM–I. 146513, 13.1 mm SL, Araki; KAUM–I. 146552, 80.4mm SL, KAUM–I. 146553, 65.0 mm SL, Keraji.
Istigobius ornatus (Rüppell, 1830)[Jpn name: Kazarihaze]
Kamohara & Yamakawa (1968a, as Gobius ornatus): 20 unregistered specimens, 12–18 mm TL, Kikai Island. This study: KAUM–I. 134699, 71.0 mm SL, Shiomichi.
Kelloggella quindecimfasciata (Fowler, 1946)[Jpn name: Hadakahaze]
KAUM–I. 134598, 19.6 mm SL, KAUM–I. 134599, 19.5 mm SL, KUN-P 55638, 17.7 mm SL, KUN-P
55640, 10.6 mm SL, Nakasato.
Luciogobius sp.
[Jpn name: N/A]
KAUM–I. 135162, 38.4 mm SL, KAUM–I. 138101, 34.2 mm SL, KAUM–I. 138102, 24.5 mm SL, KAUM–
I. 138103, 19.8 mm SL, KAUM–I. 138104, 28.5 mm SL, Shiomichi.
Remarks: This species is possibly an undescribed species.
Mugilogobius chulae (Smith, 1932)[Jpn name: Namihaze]
KAUM–I. 134847, 12.8 mm SL, KAUM–I. 134848, 13.0 mm SL, Shiramizu.
Mugilogobius sp. 1[Jpn name: Izumihaze]
KAUM–I. 134572, 33.2 mm SL, KAUM–I. 134573, 29.0 mm SL, KAUM–I. 134574, 22.4 mm SL, Shitooke; KAUM–I. 134956, 34.2 mm SL, KAUM–I. 134957, 20.9 mm SL, Shiramizu.
Oligolepis acutipennis (Valenciennes, 1837)[Jpn name: Noborihaze]
KAUM–I. 134849, 38.6 mm SL, Urahara; KAUM–I. 134964, 25.4 mm SL, Shiomichi.
Pandaka sp.
[Jpn name: Gomahaze]
KAUM–I. 134566, 13.0 mm SL, KAUM–I. 134567, 13.0 mm SL, KAUM–I. 134568, 12.5 mm SL, KAUM–
I. 134569, 13.1 mm SL, Shitooke; KAUM–I. 135120, 9.4 mm SL, KAUM–I. 135121, 6.7 mm SL, Keraji.
Pandaka trimaculata Akihito & Meguro, 1975
[Jpn name: Mitsuboshigomahaze]
Akihito et al. (2013): Kikai Island.
Paragobiodon echinocephalus (Rüppell, 1830)[Jpn name: Darumahaze]
KAUM–I. 134639, 26.2 mm SL, Onotsu; KAUM–I. 134816, 25.4 mm SL, Keraji.
Paragobiodon melanosoma (Bleeker, 1853)[Jpn name: Kurodarumahaze]
KAUM–I. 146739, 23.6 mm SL, Keraji.

Marine and freshwater fishes of Kikai Island

55

Pleurosicya micheli Fourmanoir, 1971
[Jpn name: Akasujiumitakehaze]
KAUM–I. 146686, 10.0 mm SL, Araki.
Pleurosicya muscarum (Jordan & Seale, 1906)[Jpn name: Umitakehaze]
KAUM–I. 146686, 13.1 mm SL, Keraji.
Priolepis cincta (Regan, 1908)[Jpn name: Benkeihaze]
Kamohara (1957, as Gobius eugenius): 2 unregistered specimens, 50–52 mm TL, Wan; 2 unregistered specimens, 43–46 mm TL, Somachi. Hagiwara (2018): KAUM–I. 72368, 18.4 mm SL, Keraji. Hagiwara (2019):
KAUM–I. 72368, 18.4 mm SL, Keraji. This study: KAUM–I. 72368, 18.4 mm SL, Keraji; KAUM–I.
134619, 27.2 mm SL, Onotsu; KAUM–I. 146621, 28.8 mm SL, KAUM–I. 146623, 14.7 mm SL, Tekuzuku;
KAUM–I. 146640, 20.6 mm SL, Kamikatetsu.
Priolepis latifascima Winterbottom & Burridge, 1993
[Jpn name: Futosujiirezumihaze]
KAUM–I. 146641, 15.4 mm SL, KAUM–I. 146699, 14.2 mm SL, Kamikatetsu.
Priolepis semidoliata (Valenciennes, 1837)[Jpn name: Irezumihaze]
KAUM–I. 8101, 30.2 mm SL, Urahara; KAUM–I. 134654, 19.8 mm SL, KAUM–I. 134824, 15.5 mm SL,
Nakasato; KAUM–I. 135052, 16.9 mm SL, Araki; KUN-P 55740, 10.7 mm SL, Aden.
Redigobius bikolanus (Herre, 1927)[Jpn name: Hinahaze]
KAUM–I. 134575, 25.4 mm SL, Shitooke.
Rhinogobius brunneus (Temminck & Schlegel, 1845)[Jpn name: Kuroyoshinobori]
KAUM–I. 4214, 2 specimens, 15.2–15.6 mm SL, KAUM–I. 7631, 42.6 mm SL, KAUM–I. 7632, 36.7 mm
SL, KAUM–I. 135150, 34.3 mm SL, KAUM–I. 135151, 44.1 mm SL, KAUM–I. 135152, 49.3 mm SL,
KAUM–I. 135153, 44.6 mm SL, Urahara.
Rhinogobius similis Gill, 1859
[Jpn name: Gokurakuhaze]
KAUM–I. 7629, 39.6 mm SL, KAUM–I. 8145, 33.9 mm SL, Shiomichi; KAUM–I. 134850, 32.0 mm SL,
Urahara; KAUM–I. 134955, 63.9 mm SL, Shiramizu.
Stenogobius sp.
[Jpn name: Tanekawahaze]
KAUM–I. 134965, 28.5 mm SL, KAUM–I. 134966, 25.3 mm SL, Shiomichi.
Trimma caesiura Jordan & Seale, 1906
[Jpn name: Benihaze]
Kamohara (1957, as Eviota caesiura): 1 unregistered specimen, 23 mm TL, Wan. Kamohara & Yamakawa(1965, as Eviota caesiura): 3 unregistered specimens, 22–34 mm TL, Somachi. Hagiwara (2018): KAUM–I.
72359, 26.5 mm SL, Keraji. Hagiwara (2019): KAUM–I. 72359, 26.5 mm SL, Keraji. This study: KAUM–
I. 72359, 26.5 mm SL, KAUM–I. 72360, 25.6 mm SL, KAUM–I. 72361, 20.4 mm SL, KAUM–I. 134778,

56

K. Fujiwara and H. Motomura

25.7 mm SL, KAUM–I. 134779, 17.5 mm SL, KAUM–I. 134780, 14.6 mm SL, KAUM–I. 134810, 18.4 mm
SL, KAUM–I. 134811, 21.1 mm SL, KAUM–I. 146738, 17.4 mm SL, Keraji; KAUM–I. 146523, 29.0 mm
SL, Shiomichi; KAUM–I. 146579, 17.7 mm SL, KAUM–I. 146588, 15.4 mm SL, Kamikatetsu.
Trimma caudomaculatum Yoshino & Araga, 1975
[Jpn name: Aogihaze]
KAUM–I. 134733, 25.5 mm SL, KAUM–I. 134734, 23.3 mm SL, KAUM–I. 134742, 21.3 mm SL, KAUM–
I. 134858, 18.5 mm SL, KAUM–I. 134859, 20.9 mm SL, KAUM–I. 134867, 26.5 mm SL, KAUM–I.
134868, 21.8 mm SL, KAUM–I. 134869, 22.1 mm SL, KAUM–I. 134870, 22.5 mm SL, KAUM–I. 146703,
25.3 mm SL, KAUM–I. 146729, 16.5 mm SL, KAUM–I. 146730, 17.1 mm SL, KAUM–I. 146731, 17.5 mm
SL, Keraji.
Trimma maiandros Hoese, Winterbottom & Reader, 2011
[Jpn name: Aobenihaze]
KAUM–I. 146535, 13.1 mm SL, Shiomichi; KAUM–I. 146618, 14.6 mm SL, Tekuzuku.
Trimma naudei Smith, 1957
[Jpn name: Chigobenihaze]
KAUM–I. 146578, 28.4 mm SL, Kamikatetsu.
Trimma okinawae (Aoyagi, 1949)[Jpn name: Okinawabenihaze]
KAUM–I. 134895, 17.0 mm SL, Sadeku; KAUM–I. 146473, 20.8 mm SL, KAUM–I. 146474, 16.7 mm SL,
KAUM–I. 146597, 8.2 mm SL, KAUM–I. 146608, 18.3 mm SL, KAUM–I. 146616, 19.2 mm SL, KAUM–
I. 146720, 16.4 mm SL, Tekuzuku; KAUM–I. 146531, 18.5 mm SL, Shiomichi; KAUM–I. 146667, 22.1 mm
SL, KAUM–I. 146668, 23.2 mm SL, Araki.
Trimmatom sp.
[Jpn name: Shimaisohaze]
Akihito et al. (2013): Kikai Island. Hagiwara (2018): KAUM–I. 72315, 23.3 mm SL, Shiomichi. Hagiwara(2019): KAUM–I. 72315, 23.3 mm SL, Shiomichi. This study: KAUM–I. 72315, 23.3 mm SL, Shiomichi.
Trypauchenopsis intermedia Volz, 1903
[Jpn name: Higewarasubo]
KAUM–I. 134851, 87.1 mm SL, KAUM–I. 134852, 83.7 mm SL, KAUM–I. 134853, 76.1 mm SL, KAUM–
I. 134854, 41.6 mm SL, KAUM–I. 134855, 23.2 mm SL, Urahara; KAUM–I. 134967, 71.3 mm SL, KAUM–
I. 134969, 49.8 mm SL, Aden; KAUM–I. 134968, 65.8 mm SL, Shiomichi.
Valenciennea sexguttata (Valenciennes, 1837)[Jpn name: Mizutamahaze] (Fig. 4A)
KAUM–I. 146744, 80.9 mm SL, Keraji.
Remarks: The specimen represents the first record of V. sexguttata from the Amami Islands.
Valenciennea strigata (Broussonet, 1782)[Jpn name: Akahachihaze]
Kamohara (1957, as Calleleostris strigatus): 1 unregistered specimen, 86 mm TL, Wan. This study: KAUM–
I. 146596, 97.0 mm SL, Tekuzuku.
Yongeichthys nebulosus (Forsskål, 1775)[Jpn name: Tsumugihaze]Marine and freshwater fishes of Kikai Island

57

KAUM–I. 134698, 67.3 mm SL, KAUM–I. 136720, 84.8 mm SL, KUN-P 55687, 73.6 mm SL, KUN-P
55772, 87.1 mm SL, KUN-P 55773, 76.4 mm SL, KUN-P 55776, 84.6 mm SL, Shiomichi.
FAMILY PTERELEOTRIDAE
Nemateleotris magnifica Fowler, 1938
[Jpn name: Hatatatehaze]
KAUM–I. 146723, 30.9 mm SL, Tekuzuku.
Parioglossus formosus (Smith, 1931)[Jpn name: Yosujihaze]
KUN-P 55635, 14.2 mm SL, KUN-P 55636, 27.4 mm SL, KUN-P 55637, 14.2 mm SL, Nakasato.
Parioglossus raoi (Herre, 1939)[Jpn name: Miyarabihaze]
KAUM–I. 135117, 24.0 mm SL, KAUM–I. 135118, 10.0 mm SL, KAUM–I. 135119, 10.2 mm SL, Keraji.
Ptereleotris evides (Jordan & Hubbs, 1925)[Jpn name: Kuroyurihaze]
Yamakawa (1971): 1 unregistered specimen, 27.5 mm TL, Kikai Island.
Ptereleotris sp. (unidentified)[Jpn name: N/A]
KAUM–I. 134704, 20.0 mm SL, Shiomichi.
FAMILY EPHIPPIDAE
Platax orbicularis (Forsskål, 1775)[Jpn name: Nan-yotsubameuo]
KPM-NI 26399, 46.1 mm SL, KPM-NI 26400, 42.4 mm SL, Kikai Island.
Platax pinnatus (Linnaeus, 1758)[Jpn name: Akakukuri]
KAUM–I. 15534, 68.6 mm SL, Keraji.
FAMILY SIGANIDAE
Siganus argenteus (Quoy & Gaimard, 1825)[Jpn name: Hanaaigo]
KAUM–I. 15373, 143.2 mm SL, Sakamine; KAUM–I. 135064, 257.2 mm SL, Araki.
Siganus spinus (Linnaeus, 1758)[Jpn name: Amiaigo]
Kamohara (1957): many unregistered specimens, 100–160 mm TL, Kikai Island (including Amami-oshimaIsland). This study: KAUM–I. 134915, 103.8 mm SL, Shiomichi; KAUM–I. 146264, 92.4 mm SL, Ikeji;
KAUM–I. 147616, 162.5 mm SL, Keraji.
FAMILY ZANCLIDAE
Zanclus cornutus (Linnaeus, 1758)[Jpn name: Tsunodashi]
Kuriiwa (2018): KAUM–I. 72472, 145.8 mm SL, Shiramizu. Kuriiwa (2019): KAUM–I. 72472, 145.8 mm

58

K. Fujiwara and H. Motomura

SL, Shiramizu. This study: KAUM–I. 72469, 146.0 mm SL, KAUM–I. 72472, 145.8 mm SL, KAUM–
I. 72473, 146.1 mm SL, KAUM–I. 72476, 128.7 mm SL, KAUM–I. 72477, 128.0 mm SL, Shiramizu;
KAUM–I. 72503, 132.7 mm SL, KAUM–I. 134622, 06.6 mm SL, KAUM–I. 134623, 60.6 mm SL, Onotsu.
FAMILY ACANTHURIDAE
Acanthurus dussumieri Valenciennes, 1835
[Jpn name: Nisekanranhagi]
Sakurai (2018c): KAUM–I. 72470, 185.5 mm SL, Shiramizu. Sakurai (2019c): KAUM–I. 72470, 185.5 mm
SL, Shiramizu. This study: KAUM–I. 72412, 147.5 mm SL, Keraji; KAUM–I. 72470, 185.5 mm SL, Shiramizu; KAUM–I. 134723, 114.8 mm SL, Akaren.
Acanthurus japonicus (Schmidt, 1931)[Jpn name: Namidakurohagi] (Fig. 3H)
Underwater photograph, off Tekuzuku.
Acanthurus mata (Cuvier, 1829)[Jpn name: Hiraniza]
KAUM–I. 72482, 117.4 mm SL, Aden; KAUM–I. 72497, 175.9 mm SL, KAUM–I. 72502, 142.5 mm SL,
Onotsu.
Acanthurus nigricauda Duncker & Mohr, 1929
[Jpn name: Kuromontsuki]
Sakurai (2018c): KAUM–I. 72493, 201.9 mm SL, Onotsu. Sakurai (2019c): KAUM–I. 72493, 201.9 mm SL,
Onotsu. This study: KAUM–I. 72493, 201.9 mm SL, Onotsu.
Acanthurus nigrofuscus (Forsskål, 1775)[Jpn name: Naganiza]
Kamohara (1957, as Acanthurus elongatus): 1 unregistered specimen, 175 mm TL, Somachi. This study:
KAUM–I. 15386, 96.4 mm SL, Sakamine; KAUM–I. 15525, 120.9 mm SL, KAUM–I. 42787, 85.2 mm SL,
KAUM–I. 134793, 42.3 mm SL, Keraji; KAUM–I. 72465, 136.1 mm SL, Shiramizu; KAUM–I. 134593,
62.0 mm SL, KAUM–I. 134594, 46.0 mm SL, KAUM–I. 134610, 37.6 mm SL, KAUM–I. 134611, 39.0 mm
SL, KAUM–I. 134629, 78.2 mm SL, Onotsu; KAUM–I. 146690, 58.4 mm SL, Araki; KAUM–I. 146793,
128.1 mm SL, KAUM–I. 146794, 105.1 mm SL, Akaren; KAUM–I. 146813, 118.4 mm SL, Somachi.
Acanthurus olivaceus Bloch & Schneider, 1801
[Jpn name: Montsukihagi]
KAUM–I. 146773, 135.3 mm SL, KAUM–I. 146787, 96.9 mm SL, Akaren; KAUM–I. 146802, 146.4 mm
SL, Somachi.
Acanthurus thompsoni (Fowler, 1923)[Jpn name: Ohagurohagi]
Kamohara (1957, as Acanthurus philippinus): 2 unregistered specimens, 112–128 mm TL, Wan; 1 unregistered specimen, 130 mm TL, Somachi. This study: KAUM–I. 15522, 118.9 mm SL, Keraji.
Acanthurus triostegus (Linnaeus, 1758)[Jpn name: Shimahagi]
KAUM–I. 72480, 123.8 mm SL, Aden; KAUM–I. 134621, 21.3 mm SL, Onotsu; KAUM–I. 134653, 20.8mm SL, KUN-P 55646, 21.0 mm SL, Nakasato; KAUM–I. 134708, 21.2 mm SL, Shitooke; KAUM–I.
135032, 32.2 mm SL, KAUM–I. 135086, 37.8 mm SL, KAUM–I. 135087, 28.4 mm SL, Araki; KAUM–I.
146262, 58.5 mm SL, Ikeji.

Marine and freshwater fishes of Kikai Island

59

Acanthurus xanthopterus Valenciennes, 1835
[Jpn name: Kurohagi]
KAUM–I. 134696, 23.7 mm SL, KAUM–I. 134916, 99.7 mm SL, KAUM–I. 136719, 93.0 mm SL, Shiomichi.
Ctenochaetus binotatus Randall, 1955
[Jpn name: Kokutensazanamihagi]
KAUM–I. 134606, 32.8 mm SL, Onotsu; KAUM–I. 146602, 31.5 mm SL, KAUM–I. 146620, 27.9 mm SL,
Tekuzuku.
Ctenochaetus striatus (Quoy & Gaimard, 1825)[Jpn name: Sazanamihagi]
Kamohara & Yamakawa (1965): 2 unregistered specimens, 165–192 mm TL, Somachi. Sakurai (2018c):
KAUM–I. 72481, 108.4 mm SL, Onotsu. Sakurai (2019c): KAUM–I. 72481, 108.4 mm SL, Onotsu. This
study: KAUM–I. 72481, 108.4 mm SL, KAUM–I. 134592, 38.2 mm SL, KAUM–I. 134607, 37.4 mm SL,
KAUM–I. 134628, 67.5 mm SL, Onotsu; KAUM–I. 135029, 61.0 mm SL, KAUM–I. 135030, 43.5 mm SL,
Araki.
Naso lituratus (Forster, 1801)[Jpn name: Miyakotenguhagi]
Kamohara (1957, as Callicanthus lituratus): 1 unregistered specimen, 115 mm TL, Wan. This study:
KAUM–I. 72461, 139.2 mm SL, KAUM–I. 72462, 159.6 mm SL, KAUM–I. 72463, 152.3 mm SL, Shiramizu; KAUM–I. 72483, 142.5 mm SL, KAUM–I. 72484, 143.9 mm SL, Aden; KAUM–I. 72501, 228.2 mm
SL, Onotsu.
Naso unicornis (Forsskål, 1775)[Jpn name: Tenguhagi]
Kamohara & Yamakawa (1965): 1 unregistered specimen, 340 mm TL, Somachi. Sakurai (2018c): KAUM–I.
72340, 89.5 mm SL, Keraji. Sakurai (2019c): KAUM–I. 72340, 89.5 mm SL, Keraji. This study: KAUM–I.
72340, 89.5 mm SL, KAUM–I. 147612, 118.0 mm SL, Keraji; KAUM–I. 135133, 66.6 mm SL, Shiomichi.
Naso vlamingii (Valenciennes, 1835)[Jpn name: Sazanamitosakahagi]
KAUM–I. 146806, 122.8 mm SL, Somachi.
Zebrasoma scopas (Cuvier, 1829)[Jpn name: Gomahagi]
Sakurai (2018c): KAUM–I. 72355, 36.4 mm SL, Keraji. Sakurai (2019c): KAUM–I. 72355, 36.4 mm SL,
Keraji. This study: KAUM–I. 15537, 113.3 mm SL, KAUM–I. 72355, 36.4 mm SL, KAUM–I. 134739, 26.2mm SL, KAUM–I. 146702, 25.8 mm SL, KAUM–I. 146747, 24.5 mm SL, Keraji; KAUM–I. 146515, 24.9mm SL, KAUM–I. 146516, 23.9 mm SL, Shiomichi.
Zebrasoma velifer (Bloch, 1795)[Jpn name: Hirenagahagi]
Sakurai (2018c): KAUM–I. 72494, 176.8 mm SL, KAUM–I. 72495, 180.2 mm SL, Onotsu. Sakurai (2019c):
KAUM–I. 72494, 176.8 mm SL, KAUM–I. 72495, 180.2 mm SL, Onotsu. This study: KAUM–I. 72494,
176.8 mm SL, KAUM–I. 72495, 180.2 mm SL, Onotsu; KAUM–I. 135165, 52.2 mm SL, KAUM–I. 146514,
21.0 mm SL, Shiomichi; KAUM–I. 146556, 34.3 mm SL, Keraji.

60

K. Fujiwara and H. Motomura

FAMILY ISTIOPHORIDAE
Tetrapturus angustirostris Tanaka, 1915
[Jpn name: Furaikajiki]
Hata & Motomura (2015): KAUM–I. 68262, 1268.0 mm SL, Kikai Island. Nakamura (2018d): KAUM–
I. 68262, 1268.0 mm SL, Kikai Island. Nakamura (2019d): KAUM–I. 68262, 1268.0 mm SL, Kikai Island.
This study: KAUM–I. 68262, 1268.0 mm SL, Kikai Island.
FAMILY SPHYRAENIDAE
Sphyraena barracuda (Edwards, 1771)[Jpn name: Onikamasu]
KPM-NI 26379, 249.4 mm SL, Kikai Island.
Sphyraena forsteri Cuvier, 1829
[Jpn name: Omekamasu]
Senou (2013e): Kikai-jiam Island. This study: KAUM–I. 134526, 455.8 mm SL, KAUM–I. 134527, 366.4mm SL, KAUM–I. 134727, 479.8 mm SL, KAUM–I. 134728, 384.9 mm SL, Akaren.
Sphyraena putnamae Jordan & Seale, 1905
[Jpn name: Okamasu]
Kamohara (1957, as Sphyraena jelllo): 1 unregistered specimen, 375 mm TL, Wan.
FAMILY SCOMBRIDAE
Grammatorcynus bilineatus (Rüppell, 1836)[Jpn name: Nijosaba]
KAUM–I. 134949, 410.5 mm SL, Somachi.
Katsuwonus pelamis (Linnaeus, 1758)[Jpn name: Katsuo]
Inaba (2018b): KAUM–I. 72408, 418.0 mm SL, Kikai Island. Inaba (2019b): KAUM–I. 72408, 418.0 mm
SL, Kikai Island. This study: KAUM–I. 72408, 418.0 mm SL, Kikai Island.
Thunnus alalunga (Bonnaterre, 1788)[Jpn name: Binnaga]
Inaba (2018b): KAUM–I. 73258, 533.0 mm SL, Kikai Island. Inaba (2019b): KAUM–I. 73258, 533.0 mm
SL, Kikai Island. This study: KAUM–I. 73258, 533.0 mm SL, Kikai Island.
Thunnus obesus (Lowe, 1839)[Jpn name: Mebachi]
Inaba (2018b): KAUM–I. 76058, 484.1 mm SL, Kikai Island. Inaba (2019b): KAUM–I. 76058, 484.1 mm
SL, Kikai Island. This study: KAUM–I. 76058, 484.1 mm SL, Kikai Island.
ORDER PLEURONECTIFORMESFAMILY BOTHIDAE
Bothus pantherinus (Rüppell, 1830)[Jpn name: Togedarumagarei]
KAUM–I. 134691, 57.3 mm SL, Shiomichi; KUN-P 55671, 29.6 mm SL, KUN-P 55690, 50.6 mm SL, Somachi.

Marine and freshwater fishes of Kikai Island

61

FAMILY SOLEIDAE
Aseraggodes cheni Randall & Senou, 2007
[Jpn name: Mushikuisasaushinoshita]
KAUM–I. 146490, 51.3 mm SL, Tekuzuku.
Pardachirus pavoninus (Lacepède, 1802)[Jpn name: Minamiushinoshita]
KAUM–I. 134702, 66.4 mm SL, Shiomichi; KAUM–I. 134839, 124.4 mm SL, KUN-P 55672, 104.0 mm
SL, KUN-P 55707, 136.1 mm SL, KUN-P 55708, 123.6 mm SL, Somachi.
Soleichthys heterorhinos (Bleeker, 1856)[Jpn name: Sazanamiushinoshita]
Kamohara (1957, as Solea heterorhina): 2 unregistered specimens, 104–114 mm TL, Wan; 1 unregisteredspecimen, 127 mm SL, Somachi. Kamohara & Yamakawa (1965, as Solea heterorhina): 1 unregistered specimen, 150 mm TL, Tekuzuku. This study: KAUM–I. 134760, 87.0 mm SL, Keraji; KAUM–I. 146715, 46.2mm SL, Kamikatetsu.
ORDER TETRAODONTIFORMESFAMILY BALISTIDAE
Balistapus undulatus (Park, 1797)[Jpn name: Kumadori]
KAUM–I. 147602, 160.0 mm SL, Araki.
Balistoides conspicillum (Bloch & Schneider, 1801)[Jpn name: Mongarakawahagi]
KAUM–I. 19842, 117.6 mm SL, Keraji; KAUM–I. 72491, 139.4 mm SL, Onotsu; KUN-P 55680, 73.0 mm
SL, Akaren.
Balistoides viridescens (Bloch & Schneider, 1801)[Jpn name: Gomamongara]
Matsuura (2018a): KAUM–I. 72415, 433.0 mm SL, Keraji. Matsuura (2019a): KAUM–I. 72415, 433.0 mm
SL, Keraji. This study: KAUM–I. 72415, 433.0 mm SL, Keraji.
Melichthys vidua (Richardson, 1845)[Jpn name: Kuromongara]
KAUM–I. 15194, 176.9 mm SL, Kikai Island.
Odonus niger (Rüppell, 1836)[Jpn name: Akamongara]
Kamohara (1957, as Odonus erythrodon): 1 unregistered specimen, 260 mm TL, Wan.
Pseudobalistes flavimarginatus (Rüppell, 1829)[Jpn name: Kiherimongara]
Matsuura (2018a): KAUM–I. 72409, 212.2 mm SL, Keraji. Matsuura (2019a): KAUM–I. 72409, 212.2 mm
SL, Keraji. This study: KAUM–I. 72409, 212.2 mm SL, Keraji; KAUM–I. 134644, 125.5 mm SL, Onotsu.
Rhinecanthus aculeatus (Linnaeus, 1758)[Jpn name: Murasamemongara]62

K. Fujiwara and H. Motomura

KAUM–I. 15007, 70.2 mm SL, KAUM–I. 15171, 70.4 mm SL, KAUM–I. 15193, 87.1 mm SL, KAUM–
I. 15203, 67.0 mm SL, KAUM–I. 15208, 81.0 mm SL, KAUM–I. 15217, 97.5 mm SL, KAUM–I. 15219,
86.2 mm SL, KAUM–I. 15222, 80.4 mm SL, KAUM–I. 15224, 77.1 mm SL, KAUM–I. 15225, 74.9 mm
SL, KAUM–I. 15226, 75.9 mm SL, KAUM–I. 15252, 71.0 mm SL, KAUM–I. 15253, 64.2 mm SL, Kikai
Island; KAUM–I. 134600, 47.7 mm SL, Onotsu; KAUM–I. 134711, 160.8 mm SL, KAUM–I. 134712, 134.4mm SL, Nakasato; KAUM–I. 135069, 135.1 mm SL, KAUM–I. 135076, 140.0 mm SL, KAUM–I. 135077,
157.0 mm SL, KAUM–I. 146826, 92.0 mm SL, Araki; KAUM–I. 147600, 105.0 mm SL, KAUM–I. 147601,
123.0 mm SL, Sakamine.
Rhinecanthus rectangulus (Bloch & Schneider, 1801)[Jpn name: Tasukimongara]
Kamohara & Yamakawa (1965): 1 unregistered specimen, 70 mm TL, Somachi. This study: KAUM–
I. 15169, 92.6 mm SL, KAUM–I. 15196, 122.0 mm SL, KAUM–I. 15216, 116.2 mm SL, Kikai Island;
KAUM–I. 72452, 129.1 mm SL, KAUM–I. 72453, 122.4 mm SL, KAUM–I. 72460, 115.9 mm SL, Shiramizu; KAUM–I. 134601, 41.1 mm SL, Onotsu; KAUM–I. 146598, 54.4 mm SL, Tekuzuku; KAUM–I.
146831, 93.7 mm SL, KAUM–I. 146832, 114.4 mm SL, Somachi.
Sufflamen chrysopterum (Bloch & Schneider, 1801)[Jpn name: Tsumajiromongara]
KAUM–I. 134597, 69.1 mm SL, Onotsu; KAUM–I. 134925, 117.9 mm SL, KAUM–I. 134926, 123.3 mm
SL, KAUM–I. 146829, 114.3 mm SL, KAUM–I. 146830, 134.3 mm SL, Somachi; KAUM–I. 146583, 43.4mm SL, Kamikatetsu; KAUM–I. 146743, 58.5 mm SL, Keraji.
Sufflamen fraenatum (Latreille, 1804)[Jpn name: Meganehagi]
Kamohara (1957, as Balistes capistratus): 1 unregistered specimen, 175 mm TL, Wan. This study: KAUM–I.
134724, 152.6 mm SL, Akaren; KAUM–I. 134927, 174.7 mm SL, Somachi.
FAMILY MONACANTHIDAE
Aluterus scriptus (Osbeck, 1765)[Jpn name: Soshihagi]
KAUM–I. 72490, 316.0 mm SL, Onotsu.
Cantherhines dumerilii (Hollard, 1854)[Jpn name: Hakuseihagi]
Matsuura (2018b): KAUM–I. 72410, 197.9 mm SL, Keraji. Matsuura (2019b): KAUM–I. 72410, 197.9 mm
SL, Keraji. This study: KAUM–I. 72504, 242.5 mm SL, KAUM–I. 134643, 167.7 mm SL, Onotsu; KAUM–
I. 72410, 197.9 mm SL, Keraji; KAUM–I. 134924, 151.6 mm SL, Somachi.
Cantherhines pardalis (Rüppell, 1837)[Jpn name: Amimeumazurahagi]
KAUM–I. 72464, 120.0 mm SL, Shiramizu; KAUM–I. 146674, 68.1 mm SL, Araki.
Paramonacanthus oblongus (Temminck & Schlegel, 1850)[Jpn name: Yosogi] (Fig. 4B)
KAUM–I. 15205, 91.9 mm SL, KAUM–I. 15209, 88.5 mm SL, KAUM–I. 15211, 98.0 mm SL, KAUM–I.
15218, 98.3 mm SL, Kikai Island.
Remarks: The specimens represent the first records of P. oblongus from the Amami Islands. All specimens
were transferred from the Faculty of Fisheries, Kagoshima University. Detail collection data unknown.

Marine and freshwater fishes of Kikai Island

63

FAMILY OSTRACIIDAE
Lactoria cornuta (Linnaeus, 1758)[Jpn name: Kongofugu]
KAUM–I. 135168, 43.3 mm SL, Shiomichi.
Ostracion cubicum Linnaeus, 1758
[Jpn name: Minamihakofugu]
KAUM–I. 15201, 95.5 mm SL, KAUM–I. 15220, 57.3 mm SL, KAUM–I. 15221, 54.2 mm SL, Kikai Island;
KAUM–I. 134624, 14.3 mm SL, Onotsu; KAUM–I. 146594, 16.7 mm SL, Kamikatetsu.
Ostracion meleagris Shaw, 1796
[Jpn name: Kurohakofugu]
KAUM–I. 146599, 49.0 mm SL, Tekuzuku.
FAMILY TETRAODONTIDAE
Arothron hispidus (Linnaeus, 1758)[Jpn name: Sazanamifugu]
Kamohara (1957, as Tetraodon hispidus): 2 unregistered specimens, 85–98 mm TL, Wan. This study:
KAUM–I. 15009, 48.1 mm SL, KAUM–I. 15215, 15.9 mm SL, Kikai Island; KAUM–I. 134648, 60.2 mm
SL, Nakasato; KAUM–I. 134817, 128.3 mm SL, Keraji; KAUM–I. 134939, 20.4 mm SL, Nakaguma.
Arothron nigropunctatus (Bloch & Schneider, 1801)[Jpn name: Kokutenfugu]
KAUM–I. 72338, 87.9 mm SL, Onotsu; KAUM–I. 147620, 180.8 mm SL, Kamikatetsu.
Canthigaster amboinensis (Bleeker, 1864)[Jpn name: Gomafukinchakufugu]
Kamohara (1957): 1 unregistered specimen, 82 mm TL, Onotsu; 4 unregistered specimens, 32–58 mm TL,
Somachi. Kamohara & Yamakawa (1968a): 1 unregistered specimen, 78 mm TL, Kikai Island. This study:
KAUM–I. 15310, 92.3 mm SL, Sakamine.
Canthigaster janthinoptera (Bleeker, 1855)[Jpn name: Shiborikinchakufugu]
KAUM–I. 72309, 50.9 mm SL, Shiomichi; KAUM–I. 146511, 55.9 mm SL, Onotsu; KAUM–I. 146664,
22.7 mm SL, Araki.
Canthigaster rivulata (Temminck & Schlegel, 1850)[Jpn name: Kitamakura]
KAUM–I. 15195, 27.7 mm SL, Kikai Island.
Canthigaster valentini (Bleeker, 1853)[Jpn name: Shimakinchakufugu]
Kamohara & Yamakawa (1968a): 1 unregistered specimen, 62 mm TL, Kikai Island. This study: KAUM–
I. 134630, 54.4 mm SL, Onotsu; KAUM–I. 134745, 22.1 mm SL, KAUM–I. 134806, 28.5 mm SL, Keraji;
KAUM–I. 134832, 22.7 mm SL, Shitooke; KAUM–I. 134893, 55.2 mm SL, Sadeku; KAUM–I. 135070,
64.1 mm SL, Araki; KAUM–I. 146458, 23.8 mm SL, Tekuzuku; KAUM–I. 146631, 65.3 mm SL, Kamikatetsu; KAUM–I. 146800, 73.5 mm SL, KAUM–I. 146801, 70.0 mm SL, Somachi.

64

K. Fujiwara and H. Motomura

Chelonodontops patoca (Hamilton, 1822)[Jpn name: Okinawafugu]
KAUM–I. 15200, 104.4 mm SL, KAUM–I. 15202, 108.7 mm SL, Kikai Island.
Lagocephalus cheesemanii (Clarke, 1897)[Jpn name: Kurosabafugu]
KAUM–I. 15207, 105.4 mm SL, Kikai Island.
FAMILY DIODONTIDAE
Diodon holocanthus Linnaeus, 1758
[Jpn name: Harisembon]
KAUM–I. 134953, 153.5 mm SL, KAUM–I. 134954, 168.6 mm SL, Somachi.
Diodon hystrix Linnaeus, 1758
[Jpn name: Nezumifugu]
KAUM–I. 134951, 242.5 mm SL, Shiomichi.
Diodon liturosus Shaw, 1804
[Jpn name: Hitozuraharisembon]
KAUM–I. 134823, 199.0 mm SL, Nakasato; KAUM–I. 147617, 194.2 mm SL, Kamikatetsu; KUN-P 55766,
183.7 mm SL, Araki.
Acknowledgments
We are especially grateful to H. Yonemori and J. Yoda (Yonemori Diving Service, Kikaijima) and M. Matsuoka (Kagoshima University) for providing underwater photographs and assistance in collecting fishes at
Kikai Island, T. Komagoe, H. Sasaki, and Y. Yasuda (KIKAI institute for Coral Reef Sciences) for kind support during the ichthyofaunal surveys on the island, H. Senou (KPM) for the opportunity to examine KPM
specimens, M. Matsunuma (Kindai University), S. Harazaki (Diving Service “Mori to Umi”, Yakushima),
staff of Kagoshima City Aquarium (Kagoshima), and students and volunteers of the Kagoshima University
Museum for providing species identifications and assisting in collecting fishes. G. Hardy (New Zealand) readthe manuscript and provided help with English. This study was supported in part by a Grant-in-Aid fromthe Japan Society for the Promotion of Science for JSPS Fellows (DC1: 19J21103); JSPS KAKENHI Grant
Numbers JP23580259, JP26450265, and 20H03311; the JSPS Core-to-Core Program: B Asia-Africa Science
Platforms; the “Biological Properties of Biodiversity Hotspots in Japan” project of the National Museum of
Nature and Science, Tsukuba, Japan; and “Establishment of Glocal Research and Education Network in the
Amami Islands” project of Kagoshima University adopted by the Ministry of Education, Culture, Sports,
Science and Technology, Japan.
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Matsuura, K. 2019a. Balistidae. Pp. 403–404 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun
Kaihatsu Center, Kagoshima.
Matsuura, K. 2019b. Monacanthidae. Pp. 412–413 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.).
Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon
Shimbun Kaihatsu Center, Kagoshima.
Mochida, I. and H. Motomura. 2018. An annotated checklist of marine and freshwater fishesof Tokunoshima island in
the Amami Islands, Kagoshima, southern Japan, with 202 new records. Bulletin of the Kagoshima University
Museum, 10: 1–80.
Motomura, H. 2016. The ichthyofauna of Yoron-jima Island in the southern extremity of the Amami Islands, Japan,
including comparisons with similar nearbyregions. Pp. 71–78. in Kawai, K., R. Terada and S. Kuwamura(eds.). TheAmami Islands: Culture, Society, Industry and Nature. Hokuto Shobou, Kyoto.
Motomura, H. 2018. Scorpaenidae. Pp. 68–75 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka
City Museum, Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National Museumof Nature and Science, Tsukuba.
Motomura, H. 2019a. Scorpaenidae. Pp. 70–78 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun
Kaihatsu Center, Kagoshima.
Motomura, H. 2019b. Labridae. Pp. 256–282 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun
Kaihatsu Center, Kagoshima.
Motomura, H. 2020. List of Japan’s all fish species. Current standard Japanese and scientific names of all fish species
recorded from Japanese waters. Online ver. 5. Available at https://www.museum.kagoshima-u.ac.jp/staff/
motomura/jaf.html (accessed 17 Nov. 2020).

Marine and freshwater fishes of Kikai Island

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Motomura, H. and S. Ishikawa (eds.). 2013. Fish collection building and procedures manual, English edition. The Kagoshima University Museum, Kagoshima and the Research Institute for Humanity and Nature, Kyoto. 70 pp.
Motomura, H. and K. Matsuura (eds.). 2010. Fishes of Yaku-shima island – A World Heritage island in the Osumi
Group, Kagoshima Prefecture, southern Japan. National Museum of Nature and Science, Tokyo. viii + 264
pp., 704 figs.
Motomura, H. and K. Matsuura (eds.). 2014. Field guide to fishes of Yoron Island in the middle of the Ryukyu Islands,
Japan. The Kagoshima University Museum, Kagoshima and the National Museum of Nature and Science,
Tsukuba. 648 pp.
Motomura, H. and K. Uehara. 2020. An annotated checklist of marine and freshwater fishes of Okinoerabu Island in
the Amami Islands, Kagoshima, southern Japan, with 361 new records. Bulletin of the Kagoshima University
Museum, 12: 1–125.
Nakae, M. 2018. Holocentridae. Pp. 43–50 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka City
Museum, Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National Museum of
Nature and Science, Tsukuba.
Nakae, M. 2019. Holocentridae. Pp. 45–52 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun
Kaihatsu Center, Kagoshima.
Nakae, M., H. Motomura, K. Hagiwara, H. Senou, K. Koeda, T. Yoshida, S. Tashiro, B. Jeong, H. Hata, Y. Fukui, K.
Fujiwara, T. Yamakawa, M. Aizawa, G. Shinohara and K. Matsuura. 2018. An annotated checklist of fishes ofAmami-oshima Island, the Ryukyu Islands, Japan. Memoirs of the National Museum of Nature and Science,
Tokyo 52: 205–361.
Nakajima, J., K. Ooi, M. Tomi, and T. Ijichi. 2020. Record of Carassius langsdorfii and C. auratus (Cypiriniformes,
Cyprinidae) in Kikai Island, Amami Islands, Kagoshima Prefecture, Japan. Izunuma-Uchinuma Wetland Researches 14: 103–111.
Nakamura, J. 2018a. Sparidae. Pp. 178–179 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka City
Museum, Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National Museum of
Nature and Science, Tsukuba.
Nakamura, J. 2018b. Kuhliidae. Pp. 247–248 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka
City Museum, Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National Museumof Nature and Science, Tsukuba.
Nakamura, J. 2018c. Kyphosidae. Pp. 249–250 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka
City Museum, Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National Museumof Nature and Science, Tsukuba.
Nakamura, J. 2018d. Istiophoridae. P. 387 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka City
Museum, Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National Museum of
Nature and Science, Tsukuba.
Nakamura, J. 2019a. Sparidae. Pp. 181–182 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun
Kaihatsu Center, Kagoshima.
Nakamura, J. 2019b. Kuhliidae. Pp. 250–251 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun
Kaihatsu Center, Kagoshima.
Nakamura, J. 2019c. Kyphosidae. Pp. 252–253 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun
Kaihatsu Center, Kagoshima.

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Nakamura, J. 2019d. Istiophoridae. P. 393 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun
Kaihatsu Center, Kagoshima.
Nelson, J. S. 2006. Fishes of the world. Fourth edition. John Wiley & Sons, Inc., New Jersey. xv + 601 pp.
Sakurai, Y. 2018a. Serranidae. Pp. 81–100 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka City
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Nature and Science, Tsukuba.
Sakurai, Y. 2018b. Scaridae. Pp. 279–290 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka City
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Nature and Science, Tsukuba.
Sakurai, Y. 2018c. Acanthuridae. Pp. 377–386 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka
City Museum, Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National Museumof Nature and Science, Tsukuba.
Sakurai, Y. 2019a. Serranidae. Pp. 84–103 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun
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Sakurai, Y. 2019b. Scaridae. Pp. 283–294 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun
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Sakurai, Y. 2019c. Acanthuridae. Pp. 382–392 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun
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Senou, H. 2013a. Antennariidae. Pp. 537–542, 1883–1885 in Nakabo, T. (ed.). Fishes of Japan with pictorial keys tothe species. Third edition. Tokai University Press, Hadano.
Senou, H. 2013b. Notocheiridae. Pp. 645, 1921–1922 in Nakabo, T. (ed.). Fishes of Japan with pictorial keys to thespecies. Third edition. Tokai University Press, Hadano.
Senou, H. 2013c. Adrianichthyidae. Pp. 649–650, 1923–1927 in Nakabo, T. (ed.). Fishes of Japan with pictorial keys
to the species. Third edition. Tokai University Press, Hadano.
Senou, H. 2013d. Serranidae. Pp. 757–802, 1960–1971 in Nakabo, T. (ed.). Fishes of Japan with pictorial keys to thespecies. Third edition. Tokai University Press, Hadano.
Senou, H. 2013e. Sphyraenidae. Pp. 1636–1639, 2219–2221 in Nakabo, T. (ed.). Fishes of Japan with pictorial keys tothe species. Third edition. Tokai University Press, Hadano.
Senou, H. 2018. Antennariidae. Pp. 40–41 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka City
Museum, Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National Museum of
Nature and Science, Tsukuba.
Senou, H. 2019. Antennariidae. Pp. 42–43 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun
Kaihatsu Center, Kagoshima.
Shibuya, S., Y. Ogata, R. Miki, H. Wada, and H. Motomura. 2020. Distributional records of Gymnocranius superciliosus (Perciformes: Lethrinidae) from southern Kyushu and the Ryukyu Islands, Japan. Fauna Ryukyuana 55:
9–16.
Shibuya, S. and H. Motomura. 2020. First Ryukyu record of Xiphasia matsubarai (Perciformes: Blenniidae) from Kikai-jima island in the Amami Islands, Japan, with the discovery of a healthy live individual from a tidal pool.
Ichthy, Natural History of Fishes of Japan 1: 29–31.
Shimada, K. 2013. Labridae. Pp. 1088–1136, 2045–2056 in Nakabo, T. (ed.). Fishes of Japan with pictorial keys tothe species. Third edition. Tokai University Press, Hadano.

Marine and freshwater fishes of Kikai Island

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Tashiro, S. 2018. Tripterygiidae. Pp. 295–307 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka
City Museum, Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National Museumof Nature and Science, Tsukuba.
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Yamakawa, T. 1971. Additional records of Marine fishes from Amami (IV). Reports of the Usa Marine Biological Station, Kochi University 18 (2): 1–21.
Yanagisawa, Y. 1978. Studies on the interspecific relationship between gobiid fishes and snapping shrimp. I. Gobiid
fishes associated with snapping shrimps in Japan. Publications of the Seto Marine Biological Laboratory 24:
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Yoshida, T. 2018a. Pseudochromidae. Pp. 102–107 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.).
Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka City Museum, Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National
Museum of Nature and Science, Tsukuba.
Yoshida, T. 2018b. Plesiopidae. Pp. 108–110 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka
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Yoshida, T. 2018c. Apogonidae. Pp. 113–142 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima, Yokosuka
City Museum, Yokosuka, Kanagawa Prefectural Museum of Natural History, Odawara, and National Museumof Nature and Science, Tsukuba.
Yoshida, T. 2019a. Pseudochromidae. Pp. 105–110 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.).
Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon
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Yoshida, T. 2019c. Apogonidae. Pp. 116–145 in Motomura, H., K. Hagiwara, H. Senou and M. Nakae (eds.). Identification guide to fishes of the Amami Islands in the Ryukyu Archipelago, Japan. The Minaminippon Shimbun
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Yoshida, T. and H. Motomura. 2016. First record of Antennarius randalli (Lophiiformes: Antennariidae) from Kikai-jima island, Amai Islands, Kagoshima Prefecture, southern Japan. Nature of Kagoshima 42: 45–48.

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Fig. 1. Maps of the Ryukyu
Archipelago (upper) and theAmami Islands (lower). The
island marked in red is Kikai
Island.

Fig. 2. Map of Kikai Island(entire island = Kikai Town).
Names used in text are shown
here.

K. Fujiwara and H. Motomura

Marine and freshwater fishes of Kikai Island

A

E

B

F

C

G

D

H

73

Fig. 3. Fishes recorded from Kikai Island on the basis of underwater photographs. A, Cheilodipterusintermedius (Apogonidae), Araki; B, Macolor macularis (Lutjanidae), Tekuzuku; C, Chaetodon daedalma(Chaetodontidae), Ikeji; D, Cheilinus fasciatus (Labridae), Kamikatetsu; E, Choerodon fasciatus (Labridae),
Araki; F, Hologymnosus doliatus (Labridae), Tekuzuku; G, Chlorurus bowersi (Scaridae), Keraji; H,
Acanthurus japonicus (Acanthuridae), Tekuzuku. Photo C by J. Yoda; all other photos by M. Matsuoka.

A

B

Fig. 4. Fishes recorded from Kikai Island, representing the first records from Amami Islands, Ryukyu Islands,
Japan. A, Valenciennea sexguttata (Gobiidae), KAUM–I. 146744, 80.9 mm SL, Keraji; B, Paramonacanthus
oblongus (Monacanthidae), KAUM–I. 15205, 91.9 mm SL, Kikai Island.

Ichthyological Publications by the Kagoshima University Museum2009 (March)
Motomura, H. (ed.). Fish collection building
and procedures manual. Japanese edition.
Kagoshima University Museum, Kagoshima. 70 pp.

2010 (March)
Motomura, H. and K. Matsuura (eds.). Fishes of Yaku-shima Island - A World Heritage
island in the Osumi Group, Kagoshima Prefecture, southern Japan. National Museumof Nature and Science, Tokyo. viii + 264
pp., 704 figs.

2011 (Nobember)
Matsunuma, M., H. Motomura, K. Matsuura, N. A. M. Shazili and M. A. Ambak (eds.).
Fishes of Terengganu - east coast of Malay
Peninsula, Malaysia. National Museumof Nature and Science, Tokyo, Universiti
Malaysia Terengganu, Terengganu, and Kagoshima University Museum, Kagoshima.
viii + 251 pp., 678 figs.

2013 (February)
Motomura, H. and S. Ishikawa (eds.). Fishcollection building and procedures manual.
English edition. Kagoshima University
Museum, Kagoshima and Research Institutefor Humanity and Nature, Kyoto. 70 pp.

2013 (March)
Motomura, H., S. Dewa, K. Furuta and K.
Matsuura (eds.). Fishes of Iou-jima andTake-shima islands, Mishima, Kagoshima,
Japan. Kagoshima University Museum, Kagoshima and National Museum of Natureand Science, Tsukuba. 390 pp., 883 figs.

2013 (September)
Yoshida, T., H. Motomura, P. Musikasin-thorn and K . Matsuura (eds.). Fishesof northern Gulf of Thailand. National
Museum of Nature and Science, Tsukuba,
Research Institute for Humanity and Nature,
Kyoto, and Kagoshima University Museum,Kagoshima. viii + 239 pp., 621 figs.

2014 (March)
Motomura, H. and K. Matsuura (eds.). Field
guide to fishes of Yoron Island in the middleof the Ryukyu Islands, Japan. Kagoshima
University Museum, Kagoshima andNational Museum of Nature and Science,
Tsukuba. 648 pp., 1808 figs.

2016 (July)
Iwatsubo, H., S. Kato and H. Motomura(eds.). Field guide to fishes of Ei in southern Kyushu, Japan. Kagoshima Museumof Aquatic Biodiversity, Makura-zaki, Kagoshima University Museum, Kagoshima,and Seahorse Ways, Minami-kyushu. 80
pp., 192 figs.

Most publications listed above can be downloaded fromhttp://www.museum.kagoshima-u.ac.jp/staff/motomura/dl.html

Ichthyological Publications by the Kagoshima University Museum2016 (August)Matsunuma, M., Y. Fukui and H. Motomura(eds.). Freshwater fishes of Kagoshima
City, southern Kyushu, Japan. Kagoshima
University Museum, Kagoshima. 86 pp.,
221 figs.

2016 (September)
Koeda, K., Y. Hibino, T. Yoshida, Y. Kimura, R. Miki, T. Kunishima, D. Sasaki, T. Furukawa, M. Sakurai, K. Eguchi, H. Suzuki,
T. Inaba, T. Uejo, S. Tanaka, M. Fujisawa, H.
Wada and T. Uchiyama. Annotated checklist of fishes of Yonaguni-jima island, thewesternmost island in Japan. Kagoshima
University Museum, Kagoshima. v + 120
pp., 523 figs.

2017 (January)
Motomura, H., U. B. Alama, N. Muto, R.
Babaran, and S. Ishikawa (eds.). Commer-cial and bycatch market fishes of Panay
Island, Republic of the Philippines. The
Kagoshima University Museum, Kagoshima, University of the Philippines Visayas,
Iloilo, and Research Institute for Humanity
and Nature, Kyoto. 246 pp., 911 figs.

2017 (February)
Motomura, H. and S. Harazaki. Annotatedchecklist of marine and freshwater fishesof Yaku-shima island in the Osumi Islands,
Kagoshima, southern Japan, with 129 newrecords. Bulletin of the Kagoshima University Museum, 9: 1–183.

2017 (October)
Iwatsubo, H. and H. Motomura (eds.). Field
guide to fishes of Kagoshima Bay in southern Kyushu, Japan. Kagoshima Museum of
Aquatic Biodiversity, Kago-shima and theKagoshima University Museum, Kagoshima. 302 pp., 618 figs.

2017 (December)
Kimura, Y., Y. Hibino, R. Miki, T. Minetoma and K. Koeda (eds.) Field guide tofishes of Kuchinoerabu-jima island in theOsumi Group, Kagoshima, southern Japan.
Kagoshima University Museum, Kagoshima. 200 pp.

2018 (March)
Koeda, K., H. Hata, M. Yamada and H.
Motomura (eds.). Field guide to fishes landed at Uchinoura Fishing Port, Kagoshima,
Japan. Kagoshima University Museum,Kagoshima. 520 pp., 1059 figs.

2018 (April)Motomura, H., K. Hagiwara, H. Senou
and M. Nakae (eds.). Identification guideto fishes of the Amami Islands, Japan. Kagoshima University Museum, Kagoshima,
Yokosuka City Museum, Yokosuka,Kanagawa Prefectural Museum of Natural History,
Odawara, and National Museumof Natureand Science, Tsukuba. 414 pp., 3290 figs.

Most publications listed above can be downloaded fromhttp://www.museum.kagoshima-u.ac.jp/staff/motomura/dl_en.html

Ichthyological Publications by the Kagoshima University Museum2018 (September)
Mochida, I. and H. Motomura. An annotated checklist of marine and freshwater
fishes of Tokunoshima island in the Amami
Islands, Kagoshima, southern Japan, with202 new records. Bulletin of the Kagoshima
University Museum, 10: 1–80.

2019 (February)
Motomura, H., K. Hagiwara, H. Senou and
M. Nakae (eds.). Identification guide tofishes of the Amami Islands in the Ryukyu
Archipelago, Japan. The Minaminippon
Shimbun Kaihatsu Center, Kagoshima. 436
pp., 3370 figs.

2020 (February)
Sonoyama, T., K. Ogimoto, S. Hori, Y.
Uchida and M. Kawano. An annotatedchecklist of marine fishes of the Sea of Japan off Yamaguchi Prefecture, Japan, with74 new records. Bulletin of the Kagoshima
University Museum, 11: 1–152.

2020 (March)
Motomura, H. and K. Uehara. An annotatedchecklist of marine and freshwater fishes of
Okinoerabu Island in the Amami Islands,
Kagoshima, southern Japan, with 361 newrecords. Bulletin of the Kagoshima University Museum, 12: 1–125.

2020 (May)
Motomura, H. List of Japan’s all fish species. Current standard Japanese and scientific names of all fish species recorded fromJapanese waters. The Kagoshima University
Museum, Kagoshima. 560 pp.

2020 (September)
Motomura, H., T. Yamamoto and S. Tagane(eds.). Field guide to wildlife of the Takaono River mouth and its vicinity areas, Izumi
City, northwestern Kagoshima Prefecture,
Japan. The Kagoshima University Museum,Kagoshima. 316 pp.

2020 (November)
Koeda, K., H. Hata, M. Yamada and H.
Motomura (eds.). Fishes from markets inOsumi Peninsula, Kagoshima, Japan. The
Kagoshima University Museum, Kagoshima. 634 pp.

2020 (December)
Fujiwara, K. and H. Motomura. An annotated checklist of marine and freshwater
fishes of Kikai Island in the Amami Islands,
Kagoshima, southern Japan, with 259 newrecords. Bulletin of the Kagoshima University Museum, 14: 1–73.

Most publications listed above can be downloaded fromhttp://www.museum.kagoshima-u.ac.jp/staff/motomura/dl_en.html

A total of 530 species, including 259 species that represent
the first reliable records from Kikai Island on the basis ofcollected specimens and/or underwater photographs,
are listed with citation of literature, registration numbers,
sizes, localities on the island, and some remarks.