October 2024
April 2025
October 2025

For reference

Graduate Institute for Advanced Studies, SOKENDAI[Genetics]
Application Guidelines
Contents
I.

Outline: Graduate Institute for Advanced Studies
p. 2

II.

Five-year Doctoral Program: Application Procedures and Important Notes
p. 3

III. Three-year Doctoral Program: Application Procedures and Important Notes
p. 17
IV. Majors and Research Subjects of the Facultyp. 31
The following Application Guidelines in English are provided for the purpose of non-JapaneseApplicants' convenience only. In the event of any inconsistency between the Japanese Application
Guidelines and the English version, the Japanese version shall prevail.
Applicants have to require to obtain a booklet of Application Forms. Please contact the address
below to ask for a booklet or any further information.

[Contact Information]
Student Affairs Section, Academic and Student Affairs Division
The Graduate University for Advanced Studies, SOKENDAI
Shonan Village, Hayama, Miura, Kanagawa 240-0193 JAPANE-mail: gakusei@ml.soken.ac.jp
Telephone number +81-46-858-1525/1526

1

Ⅰ. Outline: Graduate Institute for Advanced Studies

Admission Policy<What SOKENDAI expect of our grad students>
The Graduate University for Advanced Studies, SOKENDAI, seeks students who have a strong interest in
research, who constantly hone their abundant intellect and sensitivity with the aim of conducting researchthat will open up a new era, while taking a “bird's-eye view” of the entire field of study, and who have the will
and enthusiasm to be active on the international stage.
<Basic policy for the selection of our grad students>
In selecting students for admission, SOKENDAI places importance on basic academic ability and logicalthinking ability to actively promote research in the cutting-edge 20 research institutes affiliated to SOKENDAI.
In order to properly judge such abilities, various selections will be made according to the respective fields ofspecialization.
About SOKENDAI

https://www.soken.ac.jp/en/features/
Faculty Directory
https://www.soken.ac.jp/en/faculty-directory/index.html
Coursehttps://www.soken.ac.jp/en/education/curriculum/course/

Requirements for Graduation
https://www.soken.ac.jp/en/education/curriculum/requirement/
Degree: Apply to Doctor (Katei-Hakase)https://www.soken.ac.jp/en/education/degree/doctor/index.html

2

Ⅱ. Five-year Doctoral Program: Application Procedures and Important Notes

1

Number of Students Accepted
Number of Students Accepted
ProgramApril Admission

October Admission

Informatics

Around 8

A few

Statistical Science

Around 2

A few

Particle and Nuclear Science

Around 6

A few

Accelerator Science

Around 2

A few

Astronomical Science

Around 5

Fusion Science

Around 3

A few

Space and Astronautical Science

Around 4

A few

Molecular Science

Around 7

Materials structure Science

Around 2

Polar Science

Around 2

Basic Biology

Around 5

A few

Physiological Sciences

Around 3

A few

Genetics

Around 6

A few

Integrative Evolutionary Science

Around 3

A few

Total

2

A few

58

Qualifications for Application

Applicants must fulfill one of the following conditions
Applicants who are not required to have qualification screening;(1) Applicants who have graduated or are expected to graduate from a Japanese university specified in Article
83 of the School Education Law by the preceding month of enrollment.
(2) Applicants on whom a Bachelor’s degree has been conferred or is expected to be conferred by thepreceding month of enrollment in accordance with Article 104, Paragraph 4 of the School Educational Law.
*1(3) Applicants who have completed or are expected to complete 16-year course of school education in aforeign country by the preceding month of enrollment.
(4) Applicants who have completed or are expected to complete 16-year course of school education in aforeign country by taking the correspondence courses provided by a school in said foreign country while
residing in Japan by the preceding month of enrollment.
3(5) Applicants who have completed or are expected to complete a program in an educational institute in Japan
designated separately by the Minister of Education, Culture, Sports, Science and Technology, that provides
courses of a foreign university within the 16-year school education of said foreign country, by the preceding
month of the enrollment. This applies solely to those who have completed 16-year course of school
education in said foreign country.
(6) Applicants who have been conferred or are expected to be conferred a degree equivalent to a Bachelor’s
degree by the preceding month of enrollment by completing an educational course of three or more years
in a foreign university or in a foreign educational institute; which shall be evaluated by an organization
approved by a relevant official institution in the country for their education and research activities, orrecognized as so by the Minister of Education, Culture, Sports, Science and Technology. The course shall
include a correspondence course which enables students to study in Japan by a university or an
educational institute which is approved by the educational system of the country and satisfy the above
requirements.
(7) Applicants who have completed or are expected to complete the specialized course of a vocational school
designated separately by the Minister of Education, Culture, Sports, Science and Technology in accordancewith Enforcement Regulation of the School Education Law, Article 155, paragraph 1, item 5 (limited tocourses for which the term of study is four years or more, and which satisfies the standards determined by
the Minister of Education, Culture, Sports, Science and Technology) on or after the date determined by the
Minister of Education, Culture, Sports, Science and Technology, or by the preceding month of enrollment.
(8) Applicants who are designated by the Minister of Education, Culture, Sports, Science and Technology inaccordance with Enforcement Regulation of the School Education Law, Article 155, paragraph 1, item 6(i.e., Ministry of Education Notification number 5, 1953) *2
Applicants who are required to have qualification screening before the general application can besubmitted;(9) Applicants who fulfill any of the following provisions (a) to (c) below, and have been recognized by
SOKENDAI as having acquired the specified credits with excellent results;(a) Applicants who have completed 15-year course of school education in a foreign country by the end of
preceding month of enrollment,(b) Applicants who have completed 15-year course of school education in a foreign country by taking thecorrespondence courses provided by a school in said foreign country while residing in Japan,(c) Applicants who have completed or are expected to complete a program in an educational institute inJapan designated separately by the Minister of Education, Culture, Sports, Science and Technology,
that provides courses of a foreign university within the 15-year school education of said foreign country,
by the preceding month of the enrollment. This applies solely to those who have completed 15-year
course of school education in said foreign country.
(10) Applicants who have been or will have been enrolled in a Japanese university specified in Article 83 of the
School Education Law for at least three years by the end of preceding month of enrollment, and have beenrecognized by SOKENDAI as having acquired the specified credits of said university with excellent results.
4(11) Applicants who have entered a graduate school other than SOKENDAI in accordance with the Article 102,
paragraph 2 of the School Education Law, and have been recognized by SOKENDAI as having theappropriate academic ability to follow the content of graduate coursework.
(12) Applicants who have been recognized as having academic ability equivalent to a university graduate or
higher by the individual screening of Admission Qualifications of SOKENDAI, and attain the age of 22 by
the end of preceding month of enrollment. *3*1

Applicants to whom the provision (2) above apply are those on whom a Bachelor’s degree has been

conferred or are expected to be conferred by National Institute for Academic Degrees and Quality Enhancementof Higher Education (formerly, National Institution for Academic Degrees and University Evaluation).
*2

Applicants to whom the provision (8) above apply are those who have graduated or are expected tograduate from a university under Old University Ordinances, or Daigakko under orders for organization and actsof establishment of government ministries or agencies.
*3

Applicants to whom the provision (12) above apply are those who have graduated or are expected tograduate from junior college, technical college, vocational school, other schools, Japan campus of foreign
university, foreigners’ school in Japan and other educational institutes, and who have been recognized by
SOKENDAI as having academic ability equivalent to a university graduate or higher by individual screening.
If you have any questions regarding the qualification for admission, please contact the Student Affairs Section(E-mail: gakusei@ml.soken.ac.jp or telephone no.: +81-46-858-1525/1526) in advance.

3

Qualification Screening for Application

Applicants who intend to apply under the provisions (9) to (12) in “2 Qualifications for Application” are required
to submit the following documents to the Student Affairs Section by the designated deadline. Application
documents shall be withheld until the qualification screening is completed.
(1) Documents Required for Qualification Screening for Application(a) Application for Certifying Applicant’s Qualification (Form 7-1)(b) Application documents (as described in “5 Application Documents” below)
Examination fee, however, should be paid after application qualification is approved.
(c) Other documents required by the program. For details, please contact to the program office.
(a), (b) and (c) above must be submitted all together.
(2) Application Period for Qualification Screening(Only for the applicants who intend to apply under the provisions (9) to (12) in “2 Qualification forApplication”)
First round
Second round

Application Period
October 2024 Admission
June 3 (Monday) to
April 2025 Admission
June 6 (Thursday), 2024
April 2025 Admission
November 5 (Tuesday) to
October 2025 Admission
November 7 (Thursday), 2024

The application must arrive no later than the last day of the application period without fail.
5

It must be received during 09:00～12:00, 13:00～17:00 on weekdays of the application period.
(3) Procedure for Qualification Screening
The complete set of application documents should be sent by postal mail in a commercially available
envelope (33cm x 24cm) with the prescribed label in the booklet pasted. Please be sure to send it byregistered express mail. Applicants should write “Application Documents and Application for Certifying
Applicant’s Qualification Enclosed” in red ink on the face of the envelope. The application must reach
SOKENDAI no later than the last day of the application period.
(4) Result of Qualification Screening for Application
The result of Qualification Screening for Application shall be notified prior to the application period.
Applicants who have been approved by the screening should send in the payment for the examination fee
in accordance with Form 8. After remittance is completed, Form 8 on which the remittance receipt is
attached should be submitted.

4

Application Procedure

Applicants who intend to apply under the provisions (1) to (8) in “2 Qualifications for Application” are required tosubmit the following documents to the Student Affairs Section by the designated deadline.
(1) Application Period
First round
Second round

Application Period
October 2024 Admission
June 27 (Thursday) to
April 2025 Admission
July 3 (Wednesday), 2024
April 2025 Admission
December 5 (Thursday) to
October 2025 Admission
December 11 (Wednesday), 2024

The application must arrive no later than the last day of the application period without fail.
It must be received during 09:00～12:00, 13:00～17:00 on weekdays of the application period.
(2) Application Method
In principle, the complete set of application documents should be sent by postal mail in acommercially available envelope (33cm x 24cm) with the prescribed label in the booklet pasted.
Please be sure to send it by registered express mail. The application must reach SOKENDAI no later
than the last day of the application period. Please note that application documents will NOT be accepted byany parent institute or program office.
(Note)
Before submitting, please be sure to fill in and enclose all the necessary documents.
Please allow enough time to send the application documents, considering mail delivery conditions and thecase of insufficient documents.
Mailing address:
Student Affairs Section, Academic and Student Affairs Division
The Graduate University for Advanced Studies, SOKENDAI
Shonan Village, Hayama, Miura, Kanagawa 240-0193 JAPAN
Telephone number +81-46-858-1525/1526
6

5

Application Documents
Prior contact with the prospective supervisor of your choice is required in order to apply.
Please read through the “Important Notes for Applicants” beforehand. Also, application forms must becompleted with a black or blue pen or ballpoint pen (erasable pens are not acceptable).
(1) Application form and admission ticket for the examination (Form 1-A)
Please attach two identical photographs (4.5cm by 3.5cm taken within the past three months, upper
body, full-faced with no hat) on Form 1-A. Please read through the application guidelines and sign yourname in agreement with the contents.
(2) Academic transcripts (original)(a) Applicants who intend to apply under the provision (1) in “2 Qualifications for Application” must
submit academic transcripts from the undergraduate school. In case any of the credits wasapproved after transferred to the undergraduate school, academic transcripts from the technicalcollege or other college are also required.
(b) Applicants who intend to apply under the provision (2) in “2 Qualifications for Application” must
submit all the academic transcripts concerning the Bachelor’s degree.
(c) Applicants who intend to apply under the provisions (3) to (9) in “2 Qualifications for Application”
must submit the academic transcripts from the foreign undergraduate school.
(d) Applicants who intend to apply under the provisions (10) to (12) in “2 Qualifications for Application”
must submit the academic transcript from their final academic background.
(3) Certificate of (expected) graduation (original)(a) Applicants who intend to apply under the provision (1) in “2 Qualifications for Application” must
submit the certificate of graduation from the university which a Bachelor’s degree has been
conferred from.
(b) Applicants who intend to apply under the provision (2) in “2 Qualifications for Application” must
submit the certificate from National Institute for Academic Degrees and Quality Enhancement ofHigher Education (formerly, National Institute for Academic Degrees and University Evaluation).
(c) Applicants who intend to apply under the provisions (3) to (9) in “2 Qualifications for Application”
must submit the certificate of (expected) graduation from the foreign undergraduate school.
(d) Applicants who intend to apply under the provision (10) in “2 Qualifications for Application” must
submit the school-certificate, and applicants who intend to apply under the provisions (11) and (12)in “2 Qualifications for Application” must submit the certificate of graduation of their final academic
background.
(4) Statement of Purpose (Form 2)
Applicants who apply to a second-choice program must copy and prepare the form for the secondchoice program separately.
(5) Examination fee of 30,000 yen
For payment details, please refer to Form 8. MEXT scholarship students do not have to bear theexamination fee, however, they need to submit a certificate of MEXT scholarship student status.
7(6) Recipient’s address label (Form 9)
Applicants who live outside Japan must contact the Student Affairs Section in advance. [E-mail:gakusei@ml.soken.ac.jp](7) Confirmation Letter regarding the Applicability of the Specific Categories for Compliance with Article
25(1) and (2) of the Foreign Exchange and Foreign Trade Act (Form10)
Applicants residing in Japan must submit Form 10. Please refer to the simple check flow chart (*) tocomplete the form. For details, please contact the Research Coordination Section. [E-mail:kenkyo@ml.soken.ac.jp]* https://www.soken.ac.jp/en/admission/file/tokuteiruikei-E.pdf(8) TOEFL/TOEIC/IELTS score
Please select how to submit your English score on Form 13-1. Refer to “Important Notes for Applicants”.
(9) Books, papers, or lectures that were particularly interesting to you (Form 14-1)
Please refer to “Important Notes for Applicants”.
(10)LETTER PACK PLUS
We will send Admission ticket for the examination by "LETTER PACK PLUS." Please purchase"LETTER PACK PLUS" at the post office in Japan. Applicants who live outside Japan must contact theStudent Affairs Section in advance. [E-mail: gakusei@ml.soken.ac.jp](11)Curriculum Vitae (Form 1-2, only for international applicants and Japanese applicants who havereceived their education outside Japan)(12)Documents certifying the applicant’s research abilities, such as thesis, research report and othermaterials in which research capabilities are indicated.
Applicants who apply to a second-choice program must submit another set of copy of the documents
for the second-choice program separately.
(13)Permission for Studying While in Employment (Form 6)
Applicants who are currently employed full-time are required to submit Permission for Studying Whilein Employment (Form 6). In case the permission cannot be obtained or the applicant intends to resign
before enrollment, s/he may instead submit a statement of reason with her/his signature.
(14)A copy of Residence Card (international applicants residing in Japan) or
A copy of passport (international applicants residing outside Japan at the time of application)
Notes:i.

Incomplete documents shall not be accepted. No documents shall be returned.

ii.

In case the applicant’s name has changed after marriage, etc., a copy of family register should be
attached.

iii.

Form 2 and Form14-1 are also downloadable at our website.
https://www.soken.ac.jp/en/admission/application_info/genetics/index.html

iv.

Application documents should be written in Japanese or English. If you submit the certificate neither
in Japanese nor English, please also attach the certificate in Japanese or English.

v.

Applicants who are (were) in the doctoral or master's program must submit a transcript of the doctoral8

or master's program. Applicants who have graduated from a doctoral or master's program must
submit a certificate of graduation. Applicants who have withdrawn from the doctoral or master's
program must submit a withdrawal certificate.

6

Screening Procedures
Screening will be conducted based on submitted application documents and results of academic tests. For
the details of the screening methods for this program, please refer to “Important Notes for Applicants”.
If you apply to more than two programs, please note that screening is conducted individually, and the date orvenue may differ depending on the screening.

First round
Second round

October 2024 Admission
April 2025 Admission
April 2025 Admission
October 2025 Admission

Examination Date
July 28 (Sun) and July 29 (Mon), 2024
Spare date: July 30 (Tue), 2024
January 20 (Mon) and January 21 (Tue), 2025
Spare date: January 22 (Wed), 2025

(Note) Detailed information about the time and venue of the examination will be provided with theadmission ticket for the exam. In case the ticket shall not be delivered one week prior to the date of
examination, please contact the Student Affairs Section. Please see the following website for the delivery
schedule of admission ticket.
https://www.soken.ac.jp/en/admission/application_info/genetics/index.html
Venue of the examination

7

Program

Location and Directions

Genetics

・The entrance examination will be conducted through the Internet remotely.
National Institute of Genetics (NIG)
1111 Yata, Mishima, Shizuoka 411-8540
From the bus rotary on the South Exit of Mishima Station (JR Tokaido line), takea bus bound for “Yanagigochil” at No.5 bus stop and get off at “Idenken mae” (infront of NIG), or take a taxi (10 minutes) from Mishima Station.
The NIG free shuttle bus runs between NIG and the North Exit of Mishima Station
on weekdays.

Announcement of Results
First round
Second round

Notification
October 2024 Admission
April 2025 Admission
April 2025 Admission
October 2025 Admission

Early August 2024
Early February 2025

Further details will be sent to applicants.
Results will be mailed to the successful applicants. Announcement of results will be made on the SOKENDAI
website (https://www.soken.ac.jp/en/admission/general_admission/result/index.html), however, results shallbe confirmed by mailed notification. Inquiries regarding the results by telephone or other means will not be
responded to.

8

Admission Procedures(1) Admission period is scheduled as below:
9Late September 2024 for the enrollment of October 2024
Early to mid-March 2025 for the enrollment of April 2025
Late September 2025 for the enrollment of October 2025
Successful applicants must complete the admission procedures during the prescribed period. Further
details will be notified to successful applicants separately.
(2) Fees required for admission are as follows.
Entrance Fee: JPY282, 000
Tuition Fee for six months: JPY267, 900
Student Insurance Fee for five years: JPY5,750(Personal Accident Insurance for Students Pursuing Education and Research)
Note:(a) In case the entrance or tuition fees are revised at the time of or during enrollment, the revised
fees shall be applied from the date of revision.
(b) Entrance fees shall not be refunded under any circumstances once the payment is made.
Premium for the student insurance, however, may be refunded only if applicants decline theadmission by the cut-off dates as blow:
September 30, 2024 for the enrollment of October 2024
March 31, 2025 for the enrollment of April 2025
September 30, 2025 for the enrollment of October 2025
(3) Applicants who are currently employed full-time must submit the “Letter of Approval” issued by the employer
that acknowledges the enrollment while employed. Resignation certificate must be submitted if you resign
before you enroll at SOKENDAI.
(4) Applicants who are enrolled at a school other than SOKENDAI at the time of application (not including
applicants who will have graduated/completed the school before you enroll at SOKENDAI) must submit thecertificate of withdrawal from said school.
(5) Foreign nationals are strongly advised to obtain a student visa unless a particular reason would prohibit
them from doing so. Detailed information on how to obtain this type of visa is available on the SOKENDAI’s
website: http://www.soken.ac.jp/en/campuslife/international/immigration/

9Long-term course system
Long-term course system is a planned course of study for a certain period of time beyond the standard
period of study upon request from the student, who has an occupation or other circumstances. For further
information, please contact the Educational Affairs Section.
E-mail: kyomu@ml.soken.ac.jp, or Fax: +81-46-858-1632

10

General Notes(1) Before applying and taking the entrance examination, applicants should read through “Important Notes forApplicants”.
10

(2) Submitted documents shall not be returned. No changes or alternations to the submitted documents shallbe accepted after filing.
(3) Admission might be revoked in case of any false entry or act of dishonesty on application documents andother documents.
(4) Applicants who wish to transfer to SOKENDAI from other graduate school must contact Student Affairs
Section before the last day of the application period.
(5) Applicants with disabilities who require special consideration at the examination and after enrollment areadvised to inform SOKENDAI three months prior to the application period.
(6) Applicants should inform the Student Affairs Section if they wish to withdraw their application.
(7) In case the applicant has changed the mailing address after submitting the application documents, please
inform the Student Affairs Section of the change.
E-mail: gakusei@ml.soken.ac.jp, or Fax: +81-46-858-1632(8) Students are not allowed to simultaneously register at other universities while studying at SOKENDAI.
(9) For applicants with foreign nationality, if your name contains characters from your native language, thesecharacters may be replaced with the Roman alphabet for administrative purposes.
(10) Please be sure to check our website for any changes in the method of admission selection due to the impact
of new coronavirus infection or other factors.

11

Security Export Control
Based on the Foreign Exchange and Foreign Trade Act (*), SOKENDAI has established regulations andconducts examinations for the provision of technology and the acceptance of researchers and students. For
this reason, if it is deemed that there is a risk of falling under regulations, you may be required to change
your desired education or research content. Security Export Control regulations has nothing to do with
passing or failing the entrance examination. Please consult the program office for further details.
* https://www.meti.go.jp/policy/anpo/englishpage.html

12

Privacy Policy(1) Any personal information including applicant’s name and address submitted to SOKENDAI as part of theapplication documents shall be used during the application process such as applicant/examination
procedures, notification of results and admission procedures. After enrollment, personal information shallalso be used for student affairs (school register and course registration), student services (health care,tuition exemption and scholarship application, and career support) and administrative purposes ofprocessing payments for entrance and tuition fees.
(2) Personal information obtained in the screening process such as examination results, shall be used for
aggregate analyses of examination results and research for use in the screening process.

11

Important Notes for Applicants to Genetics Program(Five-year Doctoral Program)(1) When you apply to the Genetics Program, please contact the PI of the lab you wish to belong to in order todiscuss the research with them. As for each lab’s research, please read “Majors and Research Subjects of
the Faculty” on pages 27-31 or see the faculty introduction on the Genetics Program website.
(https://www.nig.ac.jp/nig/phd-program/faculty)
Faculty members who are scheduled to retire within the standard period of study (5 years from the time ofadmission) cannot be appointed as supervisors.
(2) Fill your prospective supervisor’s name in “1st Choice of Supervisor” of Form 1-A (Application Form). You
may appoint another supervisor as “2nd Choice” in the Genetics Program.
(3) You may apply for up to two programs at the same time, and the second choice can be chosen from the
Basic Biology Program or the Physiological Science Program or the Integrative Evolutionary Science
Program. However, since the selection process is conducted for each program, please be aware of theexamination schedules and venues when applying for two programs.
(4) In Form 2 “Statement of Purpose”, state “the reason why you would like to research in the Genetics Program”
based on your past experience in approximately 1,000 words in English.
(5) In Form 14-1 (Books, papers, or lectures that were particularly interesting to you), list three titles and explain
what you found interesting in several lines.
(6) All lectures in the Genetics Program are given in English. A score of TOEFL, TOEIC or IELTS test is used
to evaluate your English ability. Please submit your score record/official certificate in accordance with “Howto submit TOEFL/TOEIC/IELTS score” on the later page.
(7) The entrance examination for the Genetics Program is held over 2 days and is composed of a written exam
(held on the first day) and an oral exam (held on the second day). Both the written and oral portions of theentrance examination shall be conducted remotely via the Internet. Please refer to the notification that will
be posted prior to the examination for details on the schedule and how the examination will be conducted.
Written Exam (on the first day)Applicants take exam at home or equivalent location.
Applicants receive a question sheet and an answer sheet as a password-locked electronic file via email
prior to the day of the examination. On the exam day, applicants will be notified of the password at thestarting time of the exam and must complete the exam within 3 hours. Answers may be entered directly
into the electronic file (Microsoft Word or pdf), handwritten on a printed copy of the answer sheet, or a
combination of the two.
After the 3 hours have concluded (the end of the written exam), applicants must submit their completed
answer sheets by email within 20 minutes. (Handwritten answer sheets should first be scanned orotherwise converted to an electronic file).

12Oral Exam (on the second day)Applicants participate from home or equivalent location. The oral exam will be conducted online via either
Zoom or WebEx and is attended by all faculty.
For the first 5 minutes, the applicant makes a presentation summarizing either their past research or afuture research proposal, followed by a Q&A session on the applicant’s presentation, written exam, andproposed research. A total of 30 minutes is allotted for each applicant interview.
Applicants will prepare and submit in advance a 1 page (A4) document of resource text and/or graphics forfaculty to refer to during the initial 5-minute presentation. (Refer to the notification that will be posted prior
to the examination for submission deadline and other details.)
Please prepare an environment where you can use the Internet; receive and view the written exam question
sheet (electronic file); complete and submit the written exam answer sheet (electronic file); and attend a
Zoom or Webex web meeting (the meeting room URL will be informed in advance).
When taking the examination at a location other than your home, you must secure an environment where
the contents of the examination cannot be known to anyone but yourself.
Please inform us ahead of time of the location where you will take the examination.
Connection tests will be conducted in advance. Please inform us of an e-mail address and telephone
number where we will be able to contact you at any time on the previous day and the day of the examinationin case of any connection issues.
(8) The entrance examination can be taken either in Japanese or English. You may take the written exam ortake the interview in English, if you wish to do so.
(9) In “Ⅱ-6 Screening Procedures”, three dates are set as examination dates. The third day is spare and theexamination is usually carried out within the first two days.
(10)Criteria for evaluation and judgment<Criteria for evaluation>
The result of the written examination, interview and submitted documents shall be evaluatedcomprehensively.
<Criteria for acceptance>
Applicants who are considered to be competent to write a doctoral dissertation in five years and to obtainenough research abilities as PhD.
(11)If circumstances require a planned course of study beyond the standard term of study, students may beallowed to use the long-term enrollment system. For details, please refer to the Genetics Program website.
(https://www.nig.ac.jp/nig/phd-program/courses-top/stretched-graduate-program)If you wish to apply for this system at the time of application, please be sure to consult with your prospective
supervisor in advance.
(12)Students can earn a salary of 710,000 JPY per year through the Research Assistant (RA) system.
(13)Please contact the Academic Services Division, National Institute of Genetics for any enquiries about the
Genetics Program.
13

Tel: +81-55-981-6720 Fax: +81-55-981-6715 E-mail: info-soken@nig.ac.jp
For further information about the Genetics Program and NIG, please refer to the website below.
https://www.nig.ac.jp/nig/phd-program/main-page-top/main-page

14

How to submit TOEFL/TOEIC/IELTS score
All lectures in the Genetics Program are given in English. A score of TOEFL, TOEIC or IELTS test is used toevaluate your English ability.
On Form 13-1 (How to evaluate English proficiency), please select how you would like to submit yourEnglish test score. When you submit your score, please note the following points and prepare accordingly.
(1) Eligible Tests- TOEFL iBT Test
- TOEFL iBT Home Edition- TOEIC Listening & Reading Test
- TOEIC Listening & Reading IP Test
- IELTS (Academic Module)
Either of the above tests must have been taken within two years prior to the written exam date.
(2) How to Submit Score
Please submit one of the following by the method selected in Form 13-1 (How to evaluate English
proficiency).
- Official Score Report / Test Taker Score Report of the TOEFL-iBT Test
- Official Score Report / Test Taker Score Report of the TOEFL-iBT Home Edition Test
- Official Score Certificate of the TOEIC Listening & Reading Test
- Official Score Certificate of the TOEIC Listening & Reading IP Test
- Test Report Form for IELTS (Academic Module)
Submit the original at the time of application.
If the original cannot be submitted at the time of application, the original must arrive at the Genetics Program(Academic Services Division, National Institute of Genetics) by post strictly by the submission due date in
the table below. (In this case, please indicate in Attached Form 13-1 of the Application Forms.)
TOEFL-iBT Test and TOEFL iBT Home Edition : You can arrange with ETS to send your Official Score
Report directly to SOKENDAI. In this case, you do not need to submit the original at the time of
application.
Institution Code: 7564
Institution Name: SOKENDAI (The Graduate University for Advanced Studies)
IELTS (Academic Module): You can arrange with the IELTS administrator to send your Test Report Form15

directly to SOKENDAI. In this case, you do not need to submit the original at the time of application.
Organization name: SOKENDAI (The Graduate University for Advanced Studies).

first roundsecond round

October 2024 Admission
April 2025 Admission
April 2025 Admission
October 2025 Admission

Submission Due Date
July 24 (Wed), 2024
January 15 (Wed), 2025

Note:
Dates and venues for these English tests are limited. If you intend to apply to the Genetics Program, werecommend that you take one of the English tests ahead of time. Check the official websites for the details of
TOEFL-iBT Test, TOEIC Listening & Reading Test, or IELTS (Academic Module).
(TOEFL-iBT)

https://www.toefl-ibt.jp/index.html(TOEIC)

https://www.iibc-global.org/english.html(IELTS)https://www.eiken.or.jp/ielts/en

16

Ⅲ. Three-year Doctoral Program: Application Procedures and Important Notes

1

Number of Students Accepted
Number of Students Accepted
ProgramApril Admission
Anthropological Studies

Around 4

Japanese Studies

Around 3

Japanese History

Around 3

Japanese Literature

Around 2

Japanese Language Science

Around 3

Informatics

Around 12

A few

Statistical Science

Around 6

A few

Particle and Nuclear Science

Around 1

A few

Accelerator Science

Around 1

A few

Astronomical Science

Around 1

A few

Fusion Science

Around 2

A few

Space and Astronautical Science

Around 3

A few

Molecular Science

Around 5

A few

Materials structure Science

Around 1

A few

Global Environmental Studies

Around 2

Polar Science

Around 1

A few

Basic Biology

Around 3

A few

Physiological Sciences

Around 6

A few

Genetics

Around 2

A few

Integrative Evolutionary Science

Around 1

A few

Total

2

October Admission

62

Qualifications for Application
Applicants must fulfill one of the following conditions
Applicants who are not required to have qualification screening;(1) Applicants who hold or are expected to take a Master’s degree or a Professional degree by the preceding
month of enrollment.
(2) Applicants on whom a Master’s degree or a degree equivalent to a Professional degree has been17

conferred or is expected to be conferred in a foreign country by the preceding month of enrollment.
(3) Applicants who have completed the correspondence courses provided by a school in a foreign country
while residing in Japan, and have been conferred or are expected to be conferred a Master’s degree ora degree equivalent to a Professional degree by the preceding month of enrollment.
(4) Applicants who have completed or are expected to complete a program in an educational institute inJapan that provides courses from a foreign graduate school within the school education system of said
foreign country, and that is designated separately by the Minister of Education, Culture, Sports, Scienceand Technology, and have been conferred or are expected to be conferred a Master’s degree or a degree
equivalent to a Professional degree by the preceding month of enrollment.
(5) Applicants who have completed a course at the United Nations University and have been conferred orare expected to be conferred a degree equivalent to a Master’s degree by the preceding month ofenrollment, in accordance with the Enforcement Regulation of the School Education Law, Article 156,
Item 4.
(6) Applicants who have completed a course of study at a school in a foreign country or an educational
institute that has graduate school in a foreign country or the United Nations University, and have passed
or will have passed the examinations and screening equivalent to those specified in Article 16-2 of theStandards for Establishment of Graduate Schools (1974 Ministry of Education, Science and Culture
Ordinance Number 28), and who have been or will have been recognized by SOKENDAI as having
academic abilities equivalent to those who have a Master’s degree.
Applicants who are required to have qualification screening before the general application can besubmitted;(7) Applicants who have graduated from a university and have been or will have been engaged in researchat a university or a research institute for at least two years by the preceding month of the enrollment, andhave been recognized by SOKENDAI as having academic ability equivalent to or superior to those whohave a Master’s degree or a Professional degree based on said research achievement.
(8) Applicants who have completed 16-year course of school education in a foreign country or 16-year courseof school education by taking the correspondence courses provided by a school in a foreign country while
residing in Japan, and thereafter, have been or will have been engaged in research at a university or a
research institute for at least two years by the preceding month of the enrollment, and have beenrecognized by SOKENDAI as having academic ability equivalent to or superior to those who have a
Master’s degree or a Professional degree based on said research achievement.
(9) Applicants who have been recognized by SOKENDAI as having academic ability equivalent to or superior
to those who have a Master’s degree or a Professional degree by the individual screening of Admission
Qualifications of SOKENDAI, and attain the age of 24 by the end of preceding month of enrollment. ** Applicants who have completed a six-year course of medical, dental, pharmaceutical or veterinary schools are
required to be recognized by SOKENDAI as having academic ability equivalent to or superior to those who havea Master’s degree or a Professional degree by the individual screening of Admission Qualifications of18

SOKENDAI. For details, please refer to ‘3. Qualification Screening for Application’.
If you have any question regarding the qualification for admission, please contact the Student Affairs Section(E-mail: gakusei@ml.soken.ac.jp or telephone no.: +81-46-858-1525/1526) in advance.

3

Qualification Screening for Application(1) Applicants who intend to apply under the provisions (7) to (9) in “2 Qualifications for Application” are required
to submit the following documents to Student Affairs Section by the designated deadline. Application
documents shall be withheld until the qualification screening is completed.
(2) Applicants who wish to apply under the provisions (7) to (9) in “2 Qualifications for Application” are required
to submit the following documents for screening to be recognized as “having academic ability equivalent tothose who have a Master’s degree or a Professional degree”. An interview may be conducted if necessary.
(a)Documents Required for Qualification Screening for Applicationi.

Application for Certifying Applicant’s Qualification (Form 7-2)

ii.

Application Documents as specified in “5 Application Documents”

Examination fee, however, should be paid after the application is approved.
(i.) and (ii.) above should be submitted all together. Please note that applicants might be asked to submit
other documents by the program.
(b)

Application Period regarding Qualification Screening(Only for the applicants who intend to apply under the provisions (7) to (9) in “2 Qualification forApplication”)
First round
Second round

Application Period
October 2024 Admission
June 3 (Monday) to
April 2025 Admission
June 6 (Thursday), 2024
April 2025 Admission
November 5 (Tuesday) to
October 2025 Admission
November 7 (Thursday), 2024

Application must arrive no later than the last day of the application period without fail.
It must be received during 09:00～12:00, 13:00～17:00 on weekdays of the application period.
(c) Procedure for Qualification Screening
The complete set of application documents should be sent by postal mail in a commercially available
envelope (33cm x 24cm) with the prescribed label in the booklet pasted. Please be sure to send it byregistered express mail. Applicants should write “Application Documents and Application for Certifying
Applicant’s Qualification Enclosed” in red ink on the face of the envelope. If the application documents
do not fit in an envelope, they may be sent in a box. However, even in this case, please send the
documents by traceable mail. The application must reach SOKENDAI no later than the last day of theapplication period.
(d) Result of Qualification Screening for Application
The result of Qualification Screening for Application shall be notified prior to the application period.
Applicants who have been approved by the screening should complete payment for the examination fee
accordance with Form 8. After the remittance is completed, Form 8 on which the remittance receipt is
19attached should be submitted to Student Affairs Section.

4

Application Procedure

Applicants who intend to apply under the provisions (1) to (6) in “2 Qualifications for Application” are required tosubmit the following documents to the Student Affairs Section by the designated deadline.
(1) Application Period
First round
Second round

Application Period
October 2024 Admission
June 27 (Thursday) to
April 2025 Admission
July 3 (Wednesday), 2024
April 2025 Admission
December 5 (Thursday) to
October 2025 Admission
December 11 (Wednesday), 2024

The application must arrive no later than the last day of the application period without fail.
It must be received during 09:00～12:00, 13:00～17:00 on weekdays of the application period.
(2) Application Method
In principle, the complete set of application documents should be sent by postal mail in acommercially available envelope (33cm x 24cm) with the prescribed label in the booklet pasted.
Please be sure to send it by registered express mail. If the application documents do not fit in an
envelope, they may be sent in a box. However, even in this case, please send the documents by traceable
mail. The application must reach SOKENDAI no later than the last day of the application period. Please
note that application documents will not be accepted by any parent institute or program office.
(Note)
Before submitting, please be sure to fill in and enclose all the necessary documents.
Please allow enough time to send the application documents, considering mail delivery conditions and thecase of insufficient documents.
Mailing address:
Student Affairs Section, Academic and Student Affairs Division
The Graduate University for Advanced Studies, SOKENDAI
Shonan Village, Hayama, Miura, Kanagawa 240-0193 JAPAN
Telephone number +81-46-858-1525/1526

5

Application Documents

Prior contact with the prospective supervisor of your choice is required in order to apply.
Please read through the “Important Notes for Applicants” beforehand. Also, application forms must be completedwith a black or blue pen or ballpoint pen (erasable pens are not acceptable).
(1) Application form and admission ticket for the examination (Form 1-B)
Please attach two identical photographs (4.5cm by 3.5cm taken within the past three months, upper-body,full-faced with no hat) on Form 1-B. Please read through the application guidelines and sign your name inagreement with the contents.
(2) Academic transcripts (original)(a) Applicants who intend to apply under the provision (1) in “2 Qualifications for Application” must submit
20

the academic transcript from the graduate school or the Professional graduate school, and that from theundergraduate school. In case any of the credits was approved after transferred to the undergraduate
school, academic transcripts from the technical college or other college are also required.
(b) Applicants who intend to apply under the provisions (2) to (6) in “2 Qualifications for Application” must
submit the academic transcript from the graduate school in said foreign country (a Master’s degree or a
Professional degree) and that from the undergraduate school concerning the Bachelor’s degree.
(c) Applicants who intend to apply under the provisions (7) to (9) in “2 Qualifications for Application” must
submit the academic transcript from their final academic background.
(3) Certificate of (expected) graduation (original)
The certificate should state conferment of the Master’s degree or the Professional degree.
(a) Applicants who intend to apply under the provision (1) in “2 Qualifications for Application” must submit acertificate of (expected) graduation from the graduate school which a Master’s degree or a Professional
degree has been conferred from.
(b) Applicants who intend to apply under the provision (2) to (5) in “2 Qualifications for Application” must
submit the certificate of (expected) graduation from the foreign graduate school.
(c) Applicants who intend to apply under the provisions (6) to (9) in “2 Qualifications for Application” must
submit a certificate of graduation of their final academic background.
(4) Summary of Previous Research Experience (Form 3)(5) List of Research Presentations and Publications (Form 4)(6) Research Proposal (Form 5)
Applicants who apply to a second-choice program must copy and prepare the form for the second-choiceprogram separately.
(7) Examination fee of 30,000yen
For payment details, please refer to Form 8. MEXT scholarship students do not have to bear theexamination fee, however, they need to submit a certificate of MEXT scholarship student status.
(8) Recipient’s address label (Form 9)
Applicants who live outside Japan must contact the Student Affairs Section in advance. [E-mail:gakusei@ml.soken.ac.jp](9) Confirmation Letter regarding the Applicability of the Specific Categories for Compliance with Article 25(1)and (2) of the Foreign Exchange and Foreign Trade Act (Form10)
Applicants residing in Japan must submit Form 10. Please refer to the simple check flow chart (*) tocomplete the form. For details, please contact the Research Coordination Section. [E-mail:kenkyo@ml.soken.ac.jp]* https://www.soken.ac.jp/en/admission/file/tokuteiruikei-E.pdf(10)TOEFL/TOEIC/IELTS score
Please select how to submit your English score on Form 13-2. Refer to “Important Notes for Applicants”.
(11)Books, papers or lectures that were particularly interesting to you (Form 14-2)
Please refer to the “Important Notes for Applicants”.
21

(12)LETTER PACK PLUS
We will send Admission ticket for the examination by "LETTER PACK PLUS." Please purchase "LETTERPACK PLUS" at the post office in Japan. Applicants who live outside Japan must contact the Student Affairs
Section in advance. [E-mail: gakusei@ml.soken.ac.jp](13)Curriculum Vitae (Form 1-2, only for international applicants and Japanese applicants who have received
their education outside Japan)(14)Applicants who have a Master’s degree or who have submitted the Master’s thesis should submit two copiesof Master’s thesis. Please specify “Master’s thesis” on the upper-right corner of the front page in red ink.
Applicants who apply to a second-choice program must submit another set of copy of the Master’s thesis
for the second-choice program separately.
(15)Applicants who have any publication of scientific paper or treatise should submit two copies of the
publication.
Applicants who apply to a second-choice program must submit another set of copy of the publication for thesecond-choice program separately.
(16)Permission for Studying While in Employment (Form 6)
Applicants who are currently employed full-time are required to submit Permission for Taking the Entrance
Examination (Form 6). In case the permission cannot be obtained or the applicant intends to resign before
enrollment, s/he may instead submit a statement of reason sealed by her/himself. In this case, “Letter of
Permission to be enrolled while employed” or “Certificate of Resign” must be submitted at the time ofadmission.
(17)A copy of Residence Card (International applicants residing in Japan) ora copy of passport (international applicants residing outside Japan at the time of application)
Notes
i.

Incomplete documents shall not be accepted. No documents shall be returned.

ii.

In case the applicant’s name has changed after marriage, etc., a copy of family register should be
attached.

iii.

Form 3, 4, 5 and 14-2 are also downloadable at our website.
https://www.soken.ac.jp/en/admission/application_info/genetics/index.html

iv.

Application documents should be written in Japanese or English. If you submit the certificate neither inJapanese nor English, please also attach the certificate in Japanese or English.

v.

Applicants who are (were) in the doctoral program must submit a transcript of the doctoral program.
Applicants who have graduated from a doctoral program must submit a certificate of graduation.
Applicants who have withdrawn from the doctoral program must submit a withdrawal certificate.

6

Screening Procedures
Screening will be conducted based on submitted application documents and results of academic tests. For
details of the screening methods for this program, please refer to “Important Notes for Applicants”.
If you apply to more than two programs, please note that screening is conducted individually, and the date or22

venue may differ depending on the screening.
October 2024 Admission
April 2025 Admission
April 2025 Admission
October 2025 Admission

First round
Second roundExamination Date

July 28 (Sun) and July 29 (Mon), 2024
Spare date: July 30 (Tue), 2024
January 20 (Mon) and January 21 (Tue), 2025
Spare date: January 22 (Wed), 2025

(Note) Detailed information about the time and place of the examination will be provided with the admission
ticket for the exam. In case the ticket shall not be delivered one week prior to the date of examination,please contact the Student Affairs Section. Please see the following website for the delivery schedule ofadmission ticket.
https://www.soken.ac.jp/en/admission/application_info/genetics/index.html
Venue of the examination

7

Program

Location and Directions

Genetic

・The entrance examination will be conducted through the Internet remotely.
National Institute of Genetics (NIG)
1111 Yata, Mishima, Shizuoka 411-8540
From the bus rotary on the South Exit of Mishima Station (JR Tokaido line), take a bus
bound for “Yanagigochi” at No.5 bus stop and get off at “Idenken mae” (in front of NIG), or
take a taxi (10 minutes) from Mishima Station.
The NIG free shuttle bus runs between NIG and the North Exit of Mishima Staion on
weekdays.

Announcement of Results
First round
Second round

Notification
October 2024 Admission
April 2025 Admission
April 2025 Admission
October 2025 Admission

Early August 2024
Early February 2025

Further details will be sent to applicants.
Results will be mailed to the successful applicants. Announcement of results will be made on the SOKENDAI
website (https://www.soken.ac.jp/en/admission/general_admission/result/index.html), however, results shallbe confirmed by mailed notification. Inquiries regarding the results by telephone or other means will not be
responded.

8

Admission Procedures(1) Admission period is scheduled as below:
Late September 2024 for the enrollment of October 2024
Early to mid-March 2025 for the enrollment of April 2025
Late September 2025 for the enrollment of October 2025
Successful applicants must complete the admission procedures during the prescribed period. Further
details will be notified to successful applicants separately.
(2) Fees required for admission are as follows.
Entrance Fee: JPY 282,000
Tuition Fee for six months: JPY 267,900
Student Insurance Fee for three years: JPY 3,620
23

(Personal Accident Insurance for Students Pursuing Education and Research)
Note:(a) In case the entrance or tuition fees are revised at the time of or during enrollment, the revised fees
shall be applied from the date of revision.
(b) Entrance fees shall not be refunded under any circumstances once the payment is made. Premiumfor the student insurance, however, may be refunded only if applicants decline the admission by the
cut-off dates as blow:
September 30, 2024 for the enrollment of October 2024
March 31, 2025 for the enrollment of April 2025
September 30, 2025 for the enrollment of October 2025
(3) Applicants who are currently employed full-time should submit the “Letter of Approval” issued by theemployer that acknowledges the enrollment while employed. Resignation certificate must be submitted if
you resign before you enroll at SOKENDAI.
(4) Applicants who are enrolled at a school other than SOKENDAI at the time of application (not including those
who will have graduated from/completed the school before you enroll at SOKENDAI) must submit thecertificate of withdrawal from said school.
(5) Foreign nationals are strongly advised to obtain a student visa unless a particular reason would prohibit
them from doing so. Detailed information on how to obtain this type of visa is available on the SOKENDAI’s
website: http://www.soken.ac.jp/en/campuslife/international/immigration/

9Long-term course system
Long-term course system is a planned course of study for a certain period of time beyond the standard
period of study upon request from the student, who has an occupation or other circumstances. For further
information, please contact the Educational Affairs Section.
E-mail: kyomu@ml.soken.ac.jp, or Fax: +81-46-858-1632

10(1)

General Notes
Before applying and taking the entrance examination, applicants should read through “Important Notes
for Applicants”.

(2)

Submitted documents shall not be returned. No changes or alternations to the submitted documents will
be accepted after filing.

(3) Admission might be revoked in case of any false entry or act of dishonesty on application documents andother documents.
(4) Applicants who wish to transfer to SOKENDAI from other graduate school must contact Student Affairs
Section before the last day of the application period.
(5) Applicants with disabilities who require special consideration at the examination and after enrollment areadvised to inform SOKENDAI three months prior to the application period.
(6) Applicants should inform the Student Affairs Section if they wish to withdraw their application.
24

(7) In case the applicant has changed the mailing address after submitting the application documents, please
inform the Student Affairs Section of the change.
E-mail: gakusei@ml.soken.ac.jp, or Fax: +81-46-858-1632(8) Students are not allowed to simultaneously register at other universities while studying at SOKENDAI.
(9) For applicants with foreign nationality, if your name contains characters from your native language, thesecharacters may be replaced with the Roman alphabet for administrative purposes.
(10)Please be sure to check our website for any changes in the method of admission selection due to the impact
of new coronavirus infection or other factors.

11

Security Export Controls
Based on the Foreign Exchange and Foreign Trade Act (*), SOKENDAI has established regulations andconducts examinations for the provision of technology and the acceptance of researchers and students. For
this reason, if it is deemed that there is a risk of falling under regulations, you may be required to change
your desired education or research content. Security Export Control regulations has nothing to do with
passing or failing the entrance examination. Please consult the program office for further details.
* https://www.meti.go.jp/policy/anpo/englishpage.html

12

Privacy Policy(1) Any personal information including applicant’s name and address submitted to SOKENDAI as part of theapplication documents shall be used during the application process such as applicant/examination
procedures, notification of results and admission procedures. After enrollment, personal information shallalso be used for student affairs (school register and course registration), student services (health care,tuition exemption and scholarship application, and career support) and administrative purposes ofprocessing payments for entrance and tuition fees.
(2) Personal information obtained in the screening process such as examination results, shall be used for
aggregate analyses of examination results and research for use in the screening process.

25

Important Notes for Applicants to Genetics Program(Three-year Doctoral Program)(1) When you apply to the Genetics Program, please contact the PI of the lab you wish to belong to in order
to discuss the research with them. As for each lab’s research, please read “Majors and Research subjectsof the Faculty” on pages 27-31 or see the faculty introduction on the Genetics Program website.
(https://www.nig.ac.jp/nig/phd-program/faculty)
Faculty members who are scheduled to retire within the standard period of study (3 years from the time ofadmission) cannot be appointed as supervisors.
(2) Fill your prospective supervisor’s name in “1st Choice of Supervisor” of Form 1-B (Application Form). You
may appoint another supervisor as “2nd Choice” in the Genetics Program.
(3) You may apply for up to two programs at the same time, and the second choice can be chosen from the
Basic Biology Program or the Physiological Sciences Program or the Integrative Evolutionary Science
Program. However, since the selection process is conducted for each program, please be aware of theexamination schedules and venues when applying for two programs.
(4) Enter the name of the laboratory (supervisor) where you have actually conducted your research activities(if you had more than one supervisor, please enter all of them) on the "Personal History" page which is thereverse side of Form 1-B (Application form).
(5) You can add charts/figures/tables to explain your research activities in Form 3 (Summary of Previous
Research Experience) and Form 5 (Summary of Prospective Research). All charts and figures must fit
within the form. A separate sheet of paper cannot be added.
(6) Select five scientific papers you have read recently that you found particularly interesting, and enter theauthor's name, title, journal name, volume, page, and year of publication, following the example in Form
14-2 (Papers you read recently that were particularly interesting to you), and briefly explain what you
found interesting about them.
(7) All lectures in the Genetics Program are given in English. A score of TOEFL, TOEIC, or IELTS is used toevaluate your English ability. Please submit your score record/official certificate in accordance with “Howto submit TOEFL/TOEIC/IELTS score” on the later page.
(8) The entrance examination for the Genetics Program is held over 2 days and is composed of a written exam
(held on the first day) and an oral exam (held on the second day). Both the written and oral portions of theentrance examination shall be conducted remotely via the Internet. Please refer to the notification that will
be posted prior to the examination for details on the schedule and how the examination will be conducted.
Written Exam (on the first day)Applicants take exam at home or equivalent location.
Applicants receive a question sheet and an answer sheet as a password-locked electronic file via email
prior to the day of the examination. On the exam day, applicants will be notified of the password at thestarting time of the exam and must complete the exam within 3 hours. Answers may be entered directly
26into the electronic file (Microsoft Word or pdf), handwritten on a printed copy of the answer sheet, or a
combination of the two.
After the 3 hours have concluded (the end of the written exam), applicants must submit their completed
answer sheets by email within 20 minutes. (Handwritten answer sheets should first be scanned orotherwise converted to an electronic file).
Oral Exam (on the second day)Applicants participate from home or equivalent location. The oral exam will be conducted online via either
Zoom or WebEx and is attended by all faculty.
For the first 5 minutes, the applicant makes a presentation summarizing either their past research or afuture research proposal, followed by a Q&A session on the applicant’s presentation, written exam, andproposed research. A total of 30 minutes is allotted for each applicant interview.
Applicants will prepare and submit in advance a 1 page (A4) document of resource text and/or graphics forfaculty to refer to during the initial 5-minute presentation. (Refer to the notification that will be posted prior
to the examination for submission deadline and other details.)
Please prepare an environment where you can use the Internet; receive and view the written exam question
sheet (electronic file); complete and submit the written exam answer sheet (electronic file); and attend a
Zoom or Webex web meeting (the meeting room URL will be informed in advance).
When taking the examination at a location other than your home, you must secure an environment where
the contents of the examination cannot be known to anyone but yourself.
Please inform us ahead of time of the location where you will take the examination.
Connection tests will be conducted in advance. Please inform us of an e-mail address and telephone
number where we will be able to contact you at any time on the previous day and the day of theexamination in case of any connection issues.
(9) The entrance examination can be taken in Japanese or English. You may take the written exam or takethe interview in English, if you wish to do so.
(10)In “Ⅲ-6 Screening Procedures”, three days are set as examination dates. The third day is spare and theexamination is usually carried out within the first two days.
(11)Criteria for evaluation and judgment
Criteria for evaluation: The suitability of the applicant as a scientist will be evaluated based on the resultof the written examination, interview and submitted documents.
Criteria for acceptance: Applicants who are considered to be competent to write a dissertation in three
years and to obtain enough research abilities as PhD.
(12)If circumstances require a planned course of study beyond the standard term of study, students may beallowed to use the long-term enrollment system. For details, please refer to the Genetics Programwebsite.
(https://www.nig.ac.jp/nig/phd-program/courses-top/stretched-graduate-program)27

If you wish to apply for this system at the time of application, please be sure to consult with yourprospective supervisor in advance.
(13)Students can earn a salary of 780,000 JPY per year through the Research Assistant (RA) system.
Students of the three-year doctoral program may earn 1,060,000 JPY in the first year by adding up
282,000 JPY for the entrance fee.
(14)Please contact the Academic Services Division, National Institute of Genetics for any enquiries about the
Genetics Program.
Tel: +81-55-981-6720 Fax: +81-55-981-6715 E-mail: info-soken@nig.ac.jp
For further information about the Genetics Program and NIG, please refer to the website below.
https://www.nig.ac.jp/nig/phd-program/main-page-top/main-page28

How to submit TOEFL/TOEIC/IELTS score
All lectures in the Genetics Program are given in English. A score of TOEFL, TOEIC or IELTS test is used toevaluate your English ability.
On Form 13-2 (How to evaluate English proficiency) please select how you would like to submit your English
test score. When you submit your score, please note the following points and prepare accordingly.
(1) Eligible Tests- TOEFL iBT Test
- TOEFL iBT Home Edition- TOEIC Listening & Reading Test
- TOEIC Listening & Reading IP Test
- IELTS (Academic Module)
Either of the above tests must have been taken within two years prior to the written exam date(2) How to Submit Score
Please submit one of the following by the method selected in Form 13-2 (How to evaluate English
proficiency).
- Official Score Report/ Test Taker Score Report of the TOEFL-iBT Test
- Official Score Report/ Test Taker Score Report of the TOEFL-iBT Home Edition Test
- Official Score Certificate of the TOEIC Listening & Reading Test
- Official Score Certificate of the TOEIC Listening & Reading IP Test
- Test Report Form for IELTS (Academic Module)
Submit the original at the time of application.
If the original cannot be submitted at the time of application, the original must arrive at the Genetics
Program (Academic Services Division, National Institute of Genetics) by post strictly by the submission
due date in the table below. (In this case, please indicate in Attached Form 13-2 of the Application Forms.)
TOEFL-iBT Test and TOEFL iBT Home Edition: You can arrange with ETS to send your Official Score
Report directly to SOKENDAI. In this case, you do not need to submit the original at the time of
application.
Institution Code: 7564
Institution Name: SOKENDAI (The Graduate University for Advanced Studies)
IELTS (Academic Module): You can arrange with the IELTS administrator to send your Test Report Form29

directly to SOKENDAI. In this case, you do not need to submit the original at the time of application.
Organization name: SOKENDAI (The Graduate University for Advanced Studies)

Submission Due Date

first roundsecond round

October 2024 Admission
April 2025 Admission
April 2025 Admission
October 2025 Admission

July 24 (Wed), 2024
January 15 (Wed), 2025

Note:
Dates and venues for these English tests are limited. If you intend to apply to the Genetics Program, werecommend that you take one of English tests ahead of time. Check the official websites for the details of
TOEFL-iBT Test, TOEIC Listening & Reading Test, or IELTS (Academic Module).
(TOEFL-iBT)

https://www.toefl-ibt.jp/index.html(TOEIC)

https://www.iibc-global.org/english.html(IELTS)https://www.eiken.or.jp/ielts/en/

30

Ⅳ. Majors and Research Subjects of the Faculty[Genetics Program]This is a list of research groups in the Genetics Program.
For more detailed information about research, please visit the website of the National Institute of Genetics(https://www.nig.ac.jp/nig/).
*Faculty members who are scheduled to retire within the standard period of study (5 years from the time of admission for
the 5-year PhD program, 3 years from the time of admission for the 3-year PhD program) cannot be appointed assupervisors.
AKASHI, Hiroshi Laboratory
Population genetics and genome evolution
Our research focuses on identifying mechanisms of genome evolution in microbes as well as multicellular eukaryotes. Our
work often combines theoretical/computational population genetics and large scale data analyses (bioinformatics) to test
evolutionary ideas. Current topics of interest include: weak selection in genome evolution (e. g. related to chromatin structure,codon usage), global constraints that act on proteomes, and methods to infer ancestral states and to estimate adaptive
evolution.
(+81-55-981-6793, hiakashi@nig.ac.jp, http://www.evogennig.org/)ASAKAWA, Kazuhide Laboratory
Understanding the molecular and cellular basis of ALS
The primary objective of our laboratory is to understand the molecular and cellular underpinnings of the motor system andthe disease mechanism of amyotrophic lateral sclerosis (ALS), a currently incurable motor neuron disease. To achieve thisgoal, we analyze motor circuits at the behavioral, circuit, cellular, and molecular levels using zebrafish as a primary model
system, which offers advantages for neuroimaging and optogenetic manipulation of the nervous system. By focusing on key
aspects such as synapse formation, phase separation, RNA metabolism, energy homeostasis, DNA damage, and ploidy,
we aim to elucidate the unknown fundamental properties and functional interactions of motor neurons and their surrounding
cells. Through multi-angled and in-depth basic research, we seek to understand the vulnerability of motor neurons,
contributing to the identification of the root causes of ALS and the development of therapeutic strategies.
（+81-55-981-6887, kasakawa(at)nig.ac.jp, https://www.asakawalab.com/）
ARITA, Masanori Laboratory
Evolutionary network study based on genome and metabolome
Our research theme is metabolomics, genomics and network biology. We do not perform biological experiments:
computational analysis of genome and metabolome is our expertise. The target species range from bacteria, fungi, tohigher animals, to investigate how genome evolves in general. We look forward to accepting students with a broader visionand higher motivation.
(+81-55-981-9449, arita@nig.ac.jp)
IKEO, Kazuho Laboratory
Study for molecular basis of organismal evolution based on genomic sequence and gene expression profile
We study the molecular basis and evolutionary history for the acquisition of novel characters based on comparative
genomics and gene expression profiles. In particular, we are currently focusing more on elucidation of (1) Biodiversity ofmarine ecosystem using metagenome data, (2) Evolution of central nervous system and sensory organs from the view pointof gene expression change, (3) Molecular evolutionary process of dosage compensation, and (4) Evolutionary basis ofsymbiosis between Cnidarians and algae. We also work on (5) Method development for large scale sequence data, (6)Comparative study of large scale sequence data sets for the study of bioinformatics field.
(+81-55-981-6851, kikeo@nig.ac.jp, http://molevo.sakura.ne.jp/DnaData_lab/)*Retiring at the end of March, 2028
IWASATO, Takuji Laboratory
Neuronal Circuit Development in the Mouse Brain
To understand development of complex yet sophisticated neuronal circuits underlying higher brain function of mammals,
integrative studies which cover from molecules to whole animals are indispensable. By taking advantage of mouse genetic
technologies and resources which have been tremendously improved in the past decades, we are studying mechanisms ofdevelopment and function of mammalian neuronal circuits. In the somatosensory system of the mouse, formation andrefinement of neuronal circuits which connect the peripheral sensory organ and cortex can be detected morphologically as"barrel" patterning. We are studying molecular and cellular mechanisms of barrel patterning as a model of activity-dependent31

circuit maturation, by developing and using our original mouse genetic and imaging systems.
(+81-55-981-6773, tiwasato@nig.ac.jp, http://www.iwasato-lab.sakuraweb.com)*Retiring at the end of March, 2029
KANEMAKI, Masato Laboratory
DNA replication and new genetic technologies
Our research goal is to understand the mechanisms contributing stability of the human genome that prevent cancer
predisposition and genetic disorders. We are particularly interested in DNA replication which is essential for cell proliferationand often becomes a source of genome instability. For this purpose, we use new genetic technologies such as the
CRISPR/CAS9-based genome editing and the auxin-inducible degron (AID) system, the latter of which was developed in
our lab. These technologies allow us to generate conditional-knockout cell lines of human cells. We also aim to develop andimprove genetic technologies for the studies of human cells and in mice. Our current projects are as follows.
・Understanding the mechanisms of DNA replication in human cells
・Improvement of the AID system
・Applying the AID system to stem cells and animals(+81-55-981-5830, mkanemak@nig.ac.jp)
KAWAMOTO, Shoko Laboratory
Research on utilization of biological resource and database
Our laboratory has been working in research and development of databases and information system for the national bioresource project (NBRP) and genetic resource center in NIG. We are continuing to improve the quality of databases forresource user. We are also developing genome databases of wide variety of model organisms. Recently we are building a
research registry for making use of model animals for medical research.
(+81-55-981-6885, skawamot@nig.ac.jp, http://www.nbrp.jp)
KITANO, Jun Laboratory
Genetic mechanisms of adaptation and speciation
How are new species formed? How do animals adapt to novel environments? We investigate the genetic mechanisms
underlying adaptation and speciation using stickleback fishes as a model. Stickleback fishes have achieved tremendous
diversification during the last few million years, resulting in the evolution of divergent morphs. We use integrative approaches
to investigate the following topics.
●Genetic mechanisms of speciation
We are investigating the molecular mechanisms of reproductive isolation between sympatric morphs of sticklebacks.
●Genetic mechanisms of adaptation and phenotypic plasticity
Both genetic changes and phenotypic plasticity contribute to phenotypic changes that occur after colonization of novel
environments. We are investigating the molecular mechanisms underlying adaptive evolution and plastic changes.
●Mechanisms of anthropogenic evolution
Our third project is aimed at applying the knowledge of evolutionary genetics to animal conservation and ecologicalmanagement. We are investigating the genetic and ecological mechanisms by which invasive stickleback populations adapt
to novel environments.
(+81-55-981-9415, jkitano@nig.ac.jp, https://www.nig.ac.jp/labs/EcoGene/index-e.html)
KIMURA, Akatsuki Laboratory
Understanding Cell Architecture through quantification and modeling
Cells are the minimal unit of life, and are beautiful architecture in nature. One of the biggest mysteries in Cell Biology is‘how a huge number of tiny macromolecules assemble into a cell with organized and dynamic structure that performs aharmonized function.’ To tackle this question, we are constructing quantitative 4-dimensional models of cells that explain
and predict the structure and function of Cell Architecture. These models serve as compilation of our current understandingof the cell and, more importantly, clarify future questions to be addressed. We are using the nematode Caenorhabditis
elegans embryo as a model system. The ongoing research projects are followings:• Modeling the forces determining the intracellular positioning of the centrosomes.
• Modeling the dynamics of intracellular organelles during oocyte-to-embryo transition• Modeling the cell positioning during embryogenesis• Prediction and measurement of the forces inside the cells(+81-55-981-5854, akkimura@nig.ac.jp, http://cellarchlab.galaxy.bindcloud.jp)
KURAKU, Shigehiro Laboratory
Understanding animal evolution with the mechanism of genomic readout
Our understanding of organismal characteristics is not fully achieved by sequencing the whole genome and is limited by
the scarcity of our knowledge about the interplay between different genes and their products as well as other genomic
components. Our laboratory aims at elucidating the evolutionary history of complex life by scrutinizing the genome evolution
mechanisms based on accumulating knowledge of genomic readout in cellular and developmental processes. Typicalapproaches in our research include molecular phylogenetics involving diverse vertebrate species containing non-traditionalmodel species and genome-wide molecular profiling. Skills of genome informatics and biodiversity literacy obtained throughtraining with us will broaden professional career opportunities in the changing society oriented towards evidence and32sustainability.
(+81-55-981-6801, skuraku@nig.ac.jp)
KUROKAWA, Ken Laboratory
Unveiling microbial community dynamics
We are interested in understanding about microbial genome evolution and microbial community dynamics, and we are
currently reaching out in the following two major research directions;
I. Facilitate the development of an integrated database “MicrobeDB.jp”
II. Microbial community dynamics.
Our research interests blend a background in microbial genomics and metagenomics with bioinformatics and integrated
database developments that are just now allowing the prospect of illuminating microbial community dynamics. We are trying
to gain a better understanding of how microbial diversity maintain as well as how it emerged.
(+81-55-981-9437, kkurokawa@nig.ac.jp, http://microbedb.jp/)
SAITO, Kuniaki Laboratory
RNA biology and epigenetics in Drosophila
Germ cells are the only type of cell that can transmit all genetic information to the next generation. Recent studies revealed
that epigenetic codes, including histone modifications and small RNA-mediated regulatory events, are critical for theinheritance of genomic information from parents to progeny. However, it still remains obscure what molecules are involvedin and how epigenetic codes are established and inherited. In order to uncover these issues, we are currently engaged instudying RNA-mediated regulatory pathways in Drosophila by genetic, biochemical and bioinformatic techniques.
(+81-55-981-6823, saitok@nig.ac.jp, http://ksaitolab.org/)
SASAKI, Mariko Laboratory
Revealing mechanisms and functions of extrachromosomal circular DNAs
Cancer cells often accumulate extrachromosomal circular DNAs that are derived from chromosomes. These circular DNAs
contain proto-oncogenes or drug-resistant genes and can contribute to the onset and progression of tumorigenesis andconfer resistance to chemotherapy. Other types of extrachromosomal circular DNAs have been observed in normal humancells as well as in other organisms but their functions remain largely unclear. We aim to understand mechanisms underlying
formation of different types of extrachromosomal circular DNAs and their functions.
(+81-55-981-6817, m_sasaki@nig.ac.jp, https://sasaki-lab-en.labby.jp/)
SATO, Yutaka Laboratory
Molecular genetics of plant embryogenesis
The goal of our research is to elucidate the mechanism of plant embryogenesis. We are focusing on processes of thepatterning of apical-basal or dorsal-ventral axis formation, and the organogenesis during early stages of rice embryogenesis.
We are taking a molecular genetic approach using a series of rice embryogenesis defective mutants as well as comparativeembryology and genomics approaches in grass species. We are also responsible for managing, preservation, propagation,and distribution of rice genetic resources of wild rice species collected in the NIG under the NBRP.
(+81-55-981-6808, yusato@nig.ac.jp, https://www.nig.ac.jp/nig/research/organization-top/organization/sato)
SAWA, Hitoshi Laboratory
Mechanisms of asymmetric division in C. elegans
For normal development and homeostasis of animals, behavior (division, differentiation, migration and death) of cells mustbe tightly controlled. We are studying cell behaviors using the nematode C. elegans in which individual cells can be easily
observed in living animals. We are particularly focusing on cell polarization, asymmetric division and dynamics of epigenetic
status.
(+81-55-981-6845, hisawa@nig.ac.jp, http://square.umin.ac.jp/Nemalab/index-e.html)*Retiring at the end of March, 2028

SHIMAMOTO, Yuta Laboratory
Biophysics of cell division, nuclear dynamics and development
The creation of multicellular organisms involves cell division, growth and differentiation, which are precisely coordinatedin time and space. Many of these processes are epigenetically regulated by the cytoskeleton, such as actin, microtubules
and lamins. We study their spatiotemporal dynamics using live cell imaging and subcellular micromanipulation. Our recent
studies encompass three main areas: (i) high-resolution imaging and biophysical manipulation of embryonic nuclei during
the preimplantation development of mice, (ii) quantitative analysis of meiotic spindle assembly using machine learning andcell-free Xenopus egg extracts, and (iii) in vitro reconstitution of spindles using laser-based nanomanipulation and singlemolecule imaging. By integrating biophysics, cell biology, biochemistry and materials science, our goal is to achieve aquantitative understanding of the complex inner workings of cells during development and to help develop new therapeuticstrategies.
33

(+81-55-981-6784, yuta.shimamoto@nig.ac.jp, https://shimamotolab.weebly.com/)
NAKAMURA, Yasukazu Laboratory
Genome sequencing and database construction as an infrastructure for life science
Ultra high-throughput sequencing technologies allow biologists to obtain larger amounts of nucleotide sequence data.
Reliable database operation and high-quality annotation supply are essential. However, the explosive growth in the volume
of information is making it increasingly difficult to provide experimental biologists with information from databases. In addition,
the lack of sequence annotations in databases and the amplification of errors in the description of feature information are
becoming problems. To overcome these problems, we are developing DFAST: an automatic annotation system ofprokaryotes for DDBJ, as well as determining the genomes of various species that have important roles in evolutionary
research, industry and medicine, such as liverworts, citrus fruits, domesticated cats, lactic acid bacteria, and algae, andconstructing high-quality genome databases for the species.
(+81-55-981-3502, yn@nig.ac.jp, http://ynlab.info)
NONOMURA, Ken-ichi Laboratory
Molecular cytogenetics of plant germ-cell development
Primordial germ cell is differentiated from hypodermis of stamen (male) and pistil (female) primordia in angiosperm species.
Primordial germ cells are divided mitotically several times, and produce into meiocytes and nursery cells. Meiosis is one of
the essential events of genetics, because it generates a new gene combination different from that of parents.
It has remained to be largely unknown how flowering plants generate and maintain germ cells, and how they undergo
meiosis. To answer these questions, we have analyzed molecular functions of genes and proteins relating to early steps ofplant germ-cell initiation, development and/or meiosis, mainly by using mutant lines of rice, a monocotyledonous model plant.
We have also conducted the germplasm center of wild relatives and local varieties of rice in collaboration with the Plant
Genetics Lab., under the funding support of the National Bioresource Project, Japan (NBRP).
(+81-55-981-6872, knonomur@nig.ac.jp, http://nonomuralab-nig.sakura.ne.jp/NonoLab_HP/top_e.html)
HIRATA, Tatsumi Laboratory
Vertebrate neural network formation
Precise neuronal connections are the basis for the complex brain function. The fully functional brain is constructed througha series of carefully controlled developmental processes including neuronal differentiation, migration, axon outgrowth, andtarget recognition. We are exploring genetic mechanisms governing the developmental processes in vertebrate nervous
systems.
• Central Olfactory Projection Olfactory information is transferred and processed in the olfactory bulb of the brain.
Development of afferent projections from this first-order center has been studied, using knockout mice for axon guidance
molecules.
• Neuronal Migration During development, the guidepost neurons, “lot cells”, for olfactory bulb axons show a dynamic ventral
migration over the telencephalon. We are investigating mechanisms of this unique neuronal migration.
• Axon Outgrowth and Pausing Axon tip-enriched protein M6a is implicated in axon outgrowth and pausing. We are analyzing
physiological functions of this protein.
• Evolution of the neocortical layer structure The layer structure in the neocortex is unique to mammals. The evolutionary
scenarios are explored through comparisons of developmental processes in the brain structures of different vertebrate
species.
(+81-55-981-6721, tathirat@nig.ac.jp, https://www.nig.ac.jp/labs/Brain/hirata_lab/TOP.html)*Retiring at the end of March, 2029
FUKUSHIMA, Kenji Laboratory
Unraveling evolutionary marvels in plants and beyond
Evolution has led to remarkable innovations in the structures and functions of organisms, sometimes reaching levels thatseem almost unbelievable. Our research aims to demystify these evolutionary marvels. We integrate computational andexperimental approaches to study a wide range of organisms, particularly focusing on plants. Our primary interest lies in
insect-trapping carnivorous plants, which derive nutrients from their captured prey. By studying these unique plants, ourresearch enhances understanding of key concepts in evolutionary biology, including convergent evolution, co-option, drastic
morphological changes, preadaptation, phenotypic plasticity, biological interactions, and whole-genome duplication.
(+81-55-981-6751, kenji.fukushima@nig.ac.jp, https://kenji-fukushima-lab.github.io/)
MAESHIMA, Kazuhiro Laboratory
3D-organization and behavior of human genome chromatin
Our research interest is understanding how a long string of genomic DNA is three-dimensionally organized and behavesin the cell, and how the organized genome functions during cellular proliferation, differentiation, and development. We areusing a novel combination of molecular cell biology and biophysics to elucidate 3D-organization and behavior of genome
chromatin. Current on-going projects are as follows:• Single molecule/nucleosome imaging in live cells• Quantitative analysis of imaging data
• Super-resolution imaging of higher-order chromatin structures in cell differentiation
Computational modeling of chromatin in the cell34

(+81-55-981-6864, kmaeshim@nig.ac.jp, http://maeshima-lab.sakuraweb.com)
MIYAGISHIMA, Shin-ya Laboratory
Coordinating mechanisms of eukaryotic cell and organelle/endosymbiont proliferation
Mitochondria and chloroplasts are energy-converting organelles in eukaryotic cells. Both originated more than one billion
years ago when bacterial cells were engulfed by primitive eukaryotic cells. Besides these organelles, there are many
examples of endosymbioses which have integrated new functions into host cells. In order to maintain a permanentendosymbiotic relationship, endosymbionts/organelles must be replicated and inherited into each daughter cell during hostcell division. We have shown that chloroplasts and mitochondria use similar division systems, both of which are derived fromthe ancestral bacterial endosymbionts and the eukaryotic host.
The major goal of our study is to understand how two different cells are integrated into a new cell by coordinated proliferationof a host and an endosymbiotic cell. To this end, we are investigating (1) how eukaryotic host cells regulate proliferation oforganelles/endosymbionts, (2) how activities of organelles/endosymbionts affect proliferation of the host cells, and (3) how
these systems have evolved and contributed to eukaryotic evolution.
(+81-55-981-9411, smiyagis@nig.ac.jp, http://miyagishima.sakura.ne.jp)
MURAYAMA, Yasuto Laboratory
Molecular mechanism of chromosome organization and segregation
Spatiotemporal chromosome organization is fundamental for gene expression, DNA replication, repair and chromosome
segregation. SMC proteins are one of central chromosomal organizers. We study the molecular mechanism of SMC proteinsby reconstituting their DNA loading reaction in test tube. We are now combining our reconstitution system with single
molecule approaches.
(+81-55-981-6810, ystmurayama@nig.ac.jp)
MORI, Hiroshi Laboratory
Genome biology to understand the organismal diversity in environments
Organisms inhabit various environments and exhibit remarkable genome diversity. Our main research goal is to understandthe relationships between genome diversity and habitat diversity. Currently, we are focusing on microbes. We use various
bioinformatics and statistical methodologies related to comparative genomics and metagenomics for our research. We arealso developing some bioinformatics methods and tools for genomics and metagenomics, and applying these methods forvarious collaborative researches in the Advanced Genomics Center.
(+81-55-981-6852, hmori@nig.ac.jp, https://www.genome.id)
YAMAMICHI, Masato Laboratory
Eco-evolutionary dynamics: interactions of rapid evolution and ecological processes
Organisms modify their phenotypes flexibly in response to environmental changes. Such rapid adaptation to environments
can prevent population decline and extinction. Our laboratory combines mathematical models, microcosm experiments, andmeta-analyses to investigate the effects of rapid evolution and phenotypic plasticity on population and community dynamics
as well as complex eco-evolutionary feedbacks. Our aim is revealing dynamics of biodiversity through exploring interactions
and similarities between intraspecific genetic diversity and species diversity in biological communities.
(+81-55-981-6741, myamamichi@nig.ac.jp, https://sites.google.com/view/yamamichi-lab/)
YONEHARA, Keisuke Laboratory
Multiscale understanding of the structural principles of mammalian sensory system
Extraction of information about the environment by the sensory system is important for animal’s survival. We aim tounderstand the fundamental principles and diversity of sensory function emergence and its underlying structure by studying
the visual system of mice and monkeys at multi-scales, including genes, molecules, cell types, circuits, neural computation,and behaviors. For this, we combine various techniques such as genetics, two-photon imaging, electrophysiology,transsynaptic labeling, single-cell transcriptomics, and machine learning. Our studies would pave the way to identifying thecell types responsible for sensory diseases and their repair.
(+81-55-981-6792, keisuke.yonehara@nig.ac.jp, https://yoneharalab.org/)35