^d a ef)2F67U=*u L) )½E :¹adº»¼ Black SprucePicea mariana )2F2O ´*+ῌPµ Q*,ῌR S¶*-ῌ·¸TRU*.
,- ,***+ῌ ῏῍ῐ῎01)2(349:;<-=3(5(>AB C)DEFGH; )I=3(5(./5678 ?@ ;J67 )GKL> /MF. .L NOC+*῍ PQ)R(/STUV)WXYYMFLQ ZD 

5[GH/MF  8\3 ]^_`ab67)!>5c^d a ef P!

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v=32+ rior Highlands, Yukon Flats

WUqrXL/3Tn&WP;KQ!EFTt5 . ewUM )!>5cGH *u's+ 2*F  -x/; y cz Interior Forested Low-{|+)*F}RG~  ; p W.blands and Uplands )/v= 0 ,-9TJ p#$%)!X3 -,:)1EF Black Spruce Picea mari-P2*E/Mc  LQ y=- Gana )2F2ῌ P3EF Black-HP4EFy!;/PG+ G!

Spruce ;2*{5X3TP\F[/8)$/MFF/M++ ^d a C 9 ..῍ PQ(6/7Ey=./ 2 )*+ ^d a C

3(F+. y=- : /;!m *u;ef )2(367 Stand replacing) ¡<*=U¢-=ﬁre ;$5$)!X3£?)<-= ¤¥ ,@¦bA *u)B§X3 n;6h *u)- +**¨[ 1©/GKF67)*ª32F's+CUDh *uEF*>6(«GῌG¬U­+Hv=Ly!UI,ª3(F y *c5 Black Spruce )2(3 + 67I®

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Muneto Hirobe, Satoru Hobara, Naoko Tokuchi and Yojiro Matsuura : The E#ects of a Stand Replacing Fire onSoil Nitrogen Availability in the Boreal Forest in Interior Alaska : A Preliminary Survey in the Black Spruce(Picea mariana) Forest in the Poker Flat Research Range*+ 
 *,   *-   
.
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E4 5 N > O P IARC ῌ Q  R S 4 5 T  * +UVK EJAXA

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JKLM/

0F -* cm EH

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'VY zo{(fYY *\Ymz+   P9P;7 0/ *1῏N, +.1 ,2῏W EK/n ,E:Y -6789WY +31+ UM ,***¡¢YZK£  ./ 0¤1E E-. 12¥¦§+-,-.+ mm, 2 ¥¨cE 2 ª5 1 ª­ + 2¥¨c2¥¨cgn©!3 ./ 4K./+/.-2«wU0~/¬./E VGnE Y The ,,2«wU0Alaska Climate Research Center, http : //climate.

~/YZK£ ®\ Z/E 452gi.alaska.edu/Climate/Normals/interior.html 4zo{E nK6¯¢ C,**. °;<6=>?8±²E¤³ /* ´En7µ¶o Y nYªG:·8   Boundary fire  Kzmz ¸w 00¤1 ¸Vzo ¹9© 0** km º»{N8 K¼¢:YZK;M/,,..

,xKAlaska Fire Service, http : //ﬁre.ak.blm.

Y,gov - <fY Black Spruce  ῌ y½  +KJ

0def]^ ,**/  2  +      No. 1+ ῍,**2῎15

-  CD FH  */ cm

 CD

/IJIKL'M KL'M

XYZ[\]^_`a4 2 + d 'M g FH;<E FG+ HF.!A+NOPQRSTUVDIM WM bc' !8G + <4EgRh.PgiDIj  "#$%&'()*+,-.! /0* 1=+>?@! A<B.1=>?<4 2 

ῌ+'5 e'MNG$f!C$'I.

 . k HkRh-.M /Ml ῌg

 Hk-.M

+*.* cm 2+345-.4 67 +8'1

9:1;<;;<  !WM FH

 !

m !  Black Spruce    g no Pleurozium schreberi p Hylocomium splendens .q rs!tuYvwxy + "z{|}~;<1# Cladina  Cladonia$' $z2tuYvwxy #IM Cetratia .q 3I;4 %y&d'M /0* tuYvwx#IM.g Sphagnum  3I&g  kg!+   Black Spruce kg+.1/ *.0+ mg-N g-resin+ kg *.*3@!$ '''e'Mkf4    n@!A+kh! ;/  Hk A

!z(.)G' A<B.'M+3CM AIDCD g2' ;!%G9:'+

*.*, mg-N g-resin+N.NO+-.!A+#IMM  kj'M Black Spruce  g Black Spruce ;< *#' S¡¢ £¤¥¦§Y¨©¦ª+E«@! +8GMl " 'M ¬­ , ¦§Y¨©¦ -* cm +1,k+®¯#I!$ -°§±²t³2k µ@!. ¶NOP+·$¸E¹kf!+,kg xº+'»/0¼+11½@!<B.!$FH´ "# ;< FH

CDKL'M FHcm ¡  + 

 mg-N kg soil  g-C kg soil+ g-N kg soil+ῌ
+FH

+*0+4--*242.,14,,4*+/40*4//,14/+4*-

0/4*0403,0,43+,4+-42*4/0+34**4+-  n16rs!1=d'M WM AID1, No. 1+ ῍,**2῎;<

*  /-¡C%Gd'MFH-+42+/41.,-4*043*++4+*4+/-24+*4+3,14/+-42+334-/,41342*+42.
+342+4/* ,      !"# $% + $&'()*+  FH %,-23-4 56 &'()*+789C%,-D  E/9#QR

97#

:;<=7 9>8 /?!@ABFGHIJ KL9  E?MND OP + FH %STUOP%NVW

XYTZ[\]^_`a5 Q OPbZcd!

!&eCfJD/?!@AE? B7# ,_`aghijk84C Elmnopqrs4D ,R$%>BRtuvwxyz!'!&RQ {|OPXYTD}~Z[-4!

6w;a~ jk84

 OP@HR-4QRC>++ J ῌL >I9CD! FH %!&RQ/?!@AE? 7# )Z[ DFGHiC~C1, +-s48:;D >OP)   >ghi_`!RQ

Black Spruce D  FG-Z~-TMN! Q++XYT%'9;asa QR

QjkC¡¢>CC ++£7e¤¥¦d 9§OP¨©ªU!D  E/ «¬9­R>£84./01 No. 1+ ῍,**2῎17   FH  , !"#$%&'() FH %*+ !,-./ῌ῏῎῍+Chapin, F.S. et al. (,**0) Floristic

diversity and vegetation distribution in the Alaskansoils. Glob. Change Biol., 2, 21,ῌ22..

2Saugier, B.

et al. (,**+) Estimations of global terrestrial productiv-

boreal forest. In “Alaska’s changing boreal forest”

ity : Converging toward a single number? In “Terres-(Chapin, F.S. et al. eds.), 2+ῌ33, Oxford University Press,trial Global Productivity” (Roy, J. et al. eds.), /.-ῌ//1,New York.

,Flanagan, P.W. and Van Cleve, K.

(+32-) Nutrient cycling in relation to decompositionand organic matter quality in taiga ecosystems. Can. J.
-For. Res., +-, 13/ῌ2+1.

Gallant, A.L. et al. (+330)

Ecoregions of Alaska. U.S. Geological Survey Profes.sional Paper +/01.

Kasischke, E.S. et al. (,**0)3Academic Press, San Diego.

Serreze, M.C. et al.

(,***) Observational evidence of recent change in thenorthern high-latitude environment. Climatic Change,+*.0, +/3ῌ,*1.

Smithwick, E.A.H. et al. (,**/) Pos-tﬁre soil N cycling in northern conifer forests a#ectedby severe, stand-replacing wildﬁres. Ecosystems, 2,++Fire trends in the Alaskan boreal forest. In “Alaska’s+0-ῌ+2+.

changing boreal forest” (Chapin, F.S. et al. eds.), ,2/ῌtems in interior Alaska. BioScience, --, -3ῌ...

Van Cleve, K. (+32-) Taiga ecosys+,-*+, Oxford University Press, New York.

/ 0

Van Cleve, K. et al. (+320) Bioassay of forest ﬂoor

1 +33. 2345678"9: ;<=>56

nitrogen supply for plant growth. Can. J. For. Res., +0,?@ .. 2+ῌ3+.

0Matsuura, Y. (,**/) Forest ﬁree#ects on soils at Poker Flat Research Range, interior+-,*ῌ+-,0.

+-

Van Cleve, K. et al. (+33*) Responseof black spruce (Picea mariana) ecosystems to soil

Alaska-a preliminary survey for monitoring study-.

temperature modiﬁcations in interior Alaska. Can. J.

Proc. GCCA0, ,+/ῌ,+0.

For. Res., ,*, +/-*ῌ+/-/.

1Ne#, J.C. and Hooper, D.

+.Yarie, J. and Billings

U. (,**,) Vegetation and climate controls on potential

S. (,**,) Carbon balance of taiga forest within Alas-CO,, DOC and DON production in northern latitude

ka : Present and future. Can. J. For. Res., ,-, +**+ῌ+*+..

18ABCD No. 1+ ῌ,**2῍