South Paciflc Study Vo1 21,No l,2000

A New Sじ 相 ap S from Taiwan and Japan
̀漁(Diptera,Bombyliidac)

Akira NAGATOM11),Nagahisa TAMAKI幼,and Neal L.EvENHUIS〕

Abstract

One species ofttsん

r9ρ4'diStributcd in Taiwan and Japan is hcrc namcd′

was dcscribcd as SIs,opr/s sp C bascd on Only malc spccimcns in NACATOMÌ′3 females of this spccics havc bccn found in Japan and its idcntiサ

Kcy words:Diptcra,Bombyliidac,S19′′27α
̀as ncw to scicnce This spccicsα′ (1991)Rcccntly 6 males andhaS bCCn confll■ llcd

r9ρ″S,OnC ncw spccics,Taiwan and Japan

Introduction
ln NAGATOMÌ′α′.(1991),シthought to bc possibly identical withめS″S Sp.C was described and illustratcd but wasぇs″"夕9ρ雰 力 ′″2οSα″郷 (ENDERLEIN,1926).Thc fCmaleof sp く〕was unknown at that tirne Bascd on thrcc fcmalc spccilncns reccntly coHcctcd inJapan,sp.C is named′ブγαιas new to science.

葛Figs l‑3sysr′ 9ρ 認Figs 4‑5

portion of abdomen,latcral vicw;3,apical poltin of abdomcn,vcntral vicwα′,1991),4,abdominal temlinalia,latcralン S″て
ψクSノ″ὰsp nov(=Sp C in NAGATOMÌ′vicw;5,apical portion of stemum 8,ventral vlew′ブ αι sp nov(=Sp C in NAGATOMI

1)4307,Murasakibarll,Kagoshima 890‑0082,Japangull,Saitama
2)332‑122,Maekllbo,Moroyama,Inlma―
3)J Linslcy Grcssitt Ccnter for Rcscarch in Entomology

ι′α′,1991), 1,lateral vicw;2,apical

350‑0442,Japan, B i s h o p M u s c u m , H o n o l u l u , H va a、i i 9 6 8 1 7 ‑ 0 9 1 6 , U S ASouth Paciic Sttldy Vo1 21,No l,2000

Srrap S Ji αι sp.nov。 (Figs. 1‑5)
め考

お S″qりZだ Sp.C,NAGATOMI,LIu,TAMAKI and EvENHUIS,1991,South Paciflc Study,12:105.Distibution:Taiwan and Japan(Honshu).Japancse name:Taiwan haraboso tsuriabu.
sノ′9ρクS Sp. C in

Thc description and inustrations of this specics wcre given asシNAGATOMI。( 1 9 9 1 ) a n d a r e n o t r e p c a t e d h e r e . T h e f e m a l e w a s u n k n o w n i n t h e p a s t b u t
α α′therc arc 3 fernale specilncns now on hand.Except for gcnitalia,no signiflcant differcnce
is scen between male and female.
αοた′ブ
タὰare very similar to those of∫''NAGATOMI,LIu,
MATSUMURA,1916
butaredistinguishedfromzッわブ
TAMAKI and EvENHUIS,1991 and S s夕The lnale gcnitalia of 61

the latter two by ha宙。(1991)[key 10(COupletι′α′ng the charactcrs shown in NAGATOMIll)]・

αι are silnilar to thosc
Arnong the species irom Taiwan,the extemal characters of̀l′JγofSαθたガ but are distinguishcd fronl the lattcr by having thc following characters: thel試eral border of mesonotulm(eXCCpt humeral and postalar calli and a spot just behhumcral callus)dark brown to black,that is,middle latcral yellowish brourn spot absent;hind tibia(eXCCpt apical part)darkCned and the larger parts ofrnid and hind trochantcrs darkbrown to black.
γαθ is characterizcd as follows:
ι′α′.(1991),S′ ブ

Among the 12 species in NAGATOMI

a middle latcral yellowish brown spot(befOre wing basc)is abSent on mcsonotum.
Figures l‑5 show thc wholc shapc, apical portion of the abdomen and that of thestemum 8 in the fernale.The dorsum of the male thorax and malc genitalia are shown inθ′α′.(1991)

flgs.171‑183 in NAGATOMIThe specimens newly found(6material(6♂ ,3♂ )Of Sp・C in NAGATOMÌ′♀ )it thc descriptions of thc Japancse

α′ (1991).

Distribution.Taiwan and Japan(HonShu).

島uted′雰 Species dis仕s′9′Japancse name:Taiwan haraboso tsuriabu.This meansaルarc notin Taiwan in addition to Japan.It has been judged by♂usfrom
thatTaiwan
2fromJapanlf
diffcrents e p a r a t e d s p e c i i c a l l y a n d s u b s p e c i i, c3 a♀l l y f r o m 1 2 ♂
uncxpectcdly,the Japanese name above should bc amended.
Holotype:♀

,Oshikiri, Kounan―

Paratypcs(6♂

,2♀

):4♂

Ogawa― machi,Saitama Prcf,31.宙,1♀

machi, Saitama Pref,22.vii.1994,M.UcHIDA.
,samc data as holotype;1♂

i.1997,T.UsuI;1♂

,1♀ ,Nishimrusato,,OasaO,Kumagaya City,SaitamaPrel,27.v五 .1999,N.TAMAKI.
from Japan and Taiwan)mentioned as sp.C in NAGATOMI

Thc specimens(8♂

αι.
pes of Sル″(1991)are alSO designated as paraりThc holotype(♀

)and One paraりTokyo Paratypes are preseⅣ
Kagoshima and thc pr市pe(♂

)arc deposited in National Science Museum,ed in Bishop Muscum,Honolulu,Kagoshima Univcrsity,

atc collcction of one of us(TAMAKI).

α α′

NACATOMI,TANAKA and EvENHUIS:A New SIs,optrs from Taiwan and Japan(Diptcra,Bombyliidae) 17

This species is dedicatcd to Dr.

Ningwu LIu for her contribution to the study ofBombyliidac.

Discusslon
This species,whosc fcrnalc was unknown in the past,was dcscribed and illustrated inNAGATOMIθ
Sル′α′.(1991)aSンSrr"ン S Sp・C・ Which was thought to be possibly identical withつοSα″郷 (E■lDERLEIN,1926).
One of thc most important distinguishing characters inめ

portion in thc fcmale stemum 8 oTAGATOMIHENNIG (1941)treated s′郷.
力 ″ οSαHc wrote,

ざ″リ ツS iS the shape of apical

α α′.,1991;LIu and NAGATOMI,1992).

αεγ″′″α′"s(ENDERLEIN, 1926)as a synOnym of∫】Dic Gestalt der weiblichcn Subgenitalplattc ist sO vaHabel, dass ich

αεγ″ブ″α″s,dic auf Grund dieses Merknals vonヵ溜り
οSα′夕
S abgettent wllrde,nicht als
eigene Art anerkenncs kann." However, thc fernale stemum 8 of rl
different in shapc from Sしαcッ″′″α′γs is sosα″γS that it is very difflcult to believe that they areわ′″,ο

conspeciflc.
Compare the following statcments:ENDERLEIN(1926)on Sdcs

αεク 滋α′夕s:

Subgcnitalplattc♀ stark zugcspitzt(in spitze ausgezogen)" ENDERLEN(1926)on Sヵ溜ワ
οSα′郷 :

Subgenitalplatte ♀
deshinten gerade abgcsmtzt und an dcn Scitenecken mit je einemzahnartigen,nach hinten uberstehcndcn Fortsatz."
The apical portion of fernale stemum 8 in

∫ ルγαθ is blackcncd, not pointed but

truncate and without any lateral processes(Fig.5).ThuS,S′」a p a n ) , m a y e a s i l y b e s e p a r a t e d f r o mα∫εzブクαθ(diStributed in Taiwan and′αr 7 r s a n d ∫ヵ ′″οS ακνS , b O t h o f w h i c h a r cfrom Talwan.
EvENHUIS and GREATHEAD(1999)On,ノら′″2ο
sα777S: Distribution:0五 ental:Taiwan.

Palacarctic:Japan."The rccord from Japan is based On sp.C in NAGATOMI

θ′α′ (1991).

So the rccord fronl Japan should be deleted
According to flgs. 3‑4 in HENNIG (1941), no Signiflcant diffcrcnce may bc sccn
be小″een Sヵ′″,ο
Sα″γS(SO deterlnined by HEllNIG, 1941)and S′′γαθ in the shape ofgonostylus and cercus(including blackened tubercle).It iS nOt deterrnined whether flgures
3‑4 by HENNIG(1941)represent∫
There are many specics ofめ″αrys or∫ ルタαθ.
ル rttοSα′郷 ,∫ α ″ブ
̀ッざ″q,夕 S from China and Taiwan.The number of species

listed in EvENHUIS and GREATHEAD(1999)amOuntS t0 44,of、

vhich 14 wcrc describcd byC YANG(1995),C.YANG and Du(1991),D.YANG(1998),and D.YANG and C YANG
(1991, 1995, 1997, 1998). For sollrces of thc above literature, see EvENHUIS andGREATHEAD(1999).A thorough revision ofthe Chincsc species is not yet done.However,、ve venture to name Sルγαι as new to sciencc.

South Paciflc Study Vo1 21, No l, 2000

Acknowledgments:Wc lnust exprcss our sincere thanks to NIIcssrs.M.UcHIDA and T.UsuIfor supplying thc mate五al examined and to Mr.S.HIsAMATSU fOr his encouragenlment andhelp.

Referellces
Subfamilie Systropodinac(Dipt.).

ENDERLEIN,G.1926.Zuc Kemtnis der Bombyliiden―Wien.Entomol.Ztg.,43,69‐ 92.

EvENWIS,N.L.and GREATHEAD,D.J.1999.World catalog of bce flies(Diptcra:Bombyliidac).756pp,Backhuys Publishers,Lciden.
HENNIG,W.1941.Verzcichnis dcr Diptcrcn von Fon■osa.Entomol.Bcih.Berlin¨Dahlem,8, 1‑239.
species(Diptera,LIu,N.and NAGATOMI,A.1992.The female genitalia of eight Sys″
Bombyliidac).Jpn.J.Entomol.,60,731‑748.
NAGATOMI,A.,LIu,N。

,TAMAKI,N.and EvENIIUIS,No L.1991.The genus Sys″

"需″ 郷 ■om

Japan,Korea,Taiwan and Thailand(Diptera,Bombyliidac).SOuth Paciic Study,12,
23‐112.