῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎῎ ῌῌ - +    !

"#$%&' &( )*+)! ,-./ῌ0)1*+)! 234 "56784 9:;<=>? @:)AB! 567.CDB4 EC-F@G

HIJKLMNKL OPQRKL&ST: ) S

U# ! .IJVW)8+4 X4 "YZ'.MN
VB- ! X4 [\H"-.]^_`a-@:DU1G- #4
bJ-.]^_`a-@:cd B_`a-8+U#e!
Wf 4 g-hiV 4 MNKL.jklLmHSnoj$LpqrstDB4 @:uojMjv#wxyOPQRKLV #pqrzHW)X U# {4 @:u|b}zH~- 8 U#d! U-4 Ehleringer Y& +33- .MNKL+{4 ojMjv ( MjNP )AB

intrinsic water-use e$ciency ! @:Cja) OPQRKLV )* U#!

UU-.4 U )4 UX-)qpIJKLpMj
NP )AB4 hiD!
,   UU-4 + a ¡¢IJVW)#£{¤¥4 ¦a)&§)Y! 9 ¨y4 U©ª-« Shorea leprosula
pIJKL. 3 mmolCO, m¬, s¬+. mmolH, Om¬, s¬+4 ,**,a ­ . -4 pMNKL.

4 ,**,b ­ , - ! US.4 9® + m,

{- 3¯+*¬0 a ¡¢£{¤¥p- .¯+*¬- a° U«D!  ¡¢ y *.**,,/ : + ?+ : ... -4 + ± ¡¢Yoosuke Matsumoto : Leaf Physiology of Tropical Forest Trees (-) Intrinsic
Water-use E$ciency²8³ ´ µ!¶J·¸}¹ºµ!·¸"»  ! ¼ No. // ῌ,**,῍67ῌ ῌ ῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏



 ... 

  + 

 ! "C0 H+, O0# $ %
 ,,00.  H, O 
  + &'()*+,-. + /0+,-. + 12+,- 345678

9: 5 +,;<,=>?@A  . 9BC8$DEV +YZ]^_`abcdeb*YZ[\wexyz{;W$¯°. |±;
W$²°+h S O³,};W+ M  s c a : S c a p h i u m m a c r opodum, lep : Shorea leprosula, aza :Azadirachta excelsa, ass : Shorea
assamica, odo : Hopea odorata, tec :Tectona grandis, aur : Acacia auriculiformis, man : Acacia mangium

I=>?@AFGH

JKL +M

N$OPQRSTUS - V + ;WXYZ[\YZ]^_`abcdeb*fg$h% ViDjklm. n;Wop
FG  YZ]^_`abcdeb*qYZ[\ZUDEopFGr5. st,H uv+U 
V + i . wexyz{;W|}~;W +fg$ MZU,5. W+M;W NZU  )iD ;

Acasia mangium, A. auriculformis, Tectona grandis, Hopea odorata ")% wexyz{;W+M ),  #. Azadirachta excelsa ,[\+* / ;W+M . ]^_`abcdeb * + ,. Aza. excelsa   -+* mmolH, Om, s+   A. mangium   +,,/*mmolH, Om, s+ +ZUUM V , ;WXopFG ", molCO,ῌmolH, O+  . +}% # $h% Vi . Scaphium macropodum " 12 +*0# +Y8opFG¡5. Shorea assamica " +0 +*0# +Y8¢S +opFG,
]^_`abcdeb*+M SR. [\¢Sca. macropo-dum "V + £¤# +opFGY8ZU5S¥¦,. ]^_`abcdeb*8§¨© SR+M  Sca. macropodum ,ª «¬M ^­+. ®¡ +,,** m68´ µ ¶ · No. // ῍,**,῎῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏ ῌῑῐῒ῍῎῏ῌῌl ,YZDmnop1 9IWUE:RSTUVqr;k IJKLMN's "Q
'tu RSTUVWX'v wx*9l , Fyz{|}:+. Scaphium macropodum+.. Ptychopyxis caput-medusae,. Hevea brasiliensis+/. Shorea platyclados-. Shorea parvifolia+0. Dipterocarpus sublamellatus.. Dryobalanops aromatica+1. Neobalanocarps heimii/. Dipterocarpus kerrii+2. Terminalia catappa
0. Gonystylus a$ne+3. Shorea leprosula
1. Azadirachta excelsa,*. Dipterocarpus oblongifolius
2. Shorea ovalis,+. Acacia mangium
3. Dipterocarpus cornutus,,. Shorea macroptera+*. Shorea curtisii,-. Hopea nervosa++. Acacia auriculiformis,.. Tectona grandis+,. Xanthophyllum amoenum,/. Shorea assamica+-. Hopea odorata

  ./ m  2* cm  ! "#$%&'( )* +,-./01' +*ῌ 2&(-3456'78 9PROSEA +33.: ;<= >!?* S. leprosula 9,,.*@+*A0: 'BCDE +3 FG !H* IJKLMNOP' -,.*@+*A0 "Q ' ./.0@+*A0 RSTUVWX' -+.3@+*A0 YZ[\OP' -..1@+*A0! Hevea brasiliensis 9]^_`aM: bcdefgGh WXij* . ;k T. catappa A. mangium DY Z f g No. // ῍,**,῎69ῌ ῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏ῌῑῐῒ῍῎῏ῌ T. grandis  -   +2 ,+ ,. 


H. brasiliensis  , +**!"#$ %&'()*+(,-./$01.2 . 3 Dryobalanops aromatica "#$0456789' 2*CD!:;<=% >?@A (5(BEFG S. assamica  HI(  JBB K + G(D"#$NOPQRS(GFEF

TUVWX.YZO.[\ ] ^\_  `ab T. grandis ],. _ L(GF M
 A. mangium ],+ _TUVWX.YZO.[c B\ ]K +_ Hdefg\hf\I iUjkl& G((T. grandis mn o! "LJ :;Fpq2

B r.s . tuv#wxy$ 2*% ]{& +330_3 +1z.- m D '3h(( (#|}F%H (" |}~e( H F) mnUj T. grandis k*+ `

TUVWX.YZO.[\h F [#[,\-.( k(iUj/*wB

0 ^\(def1\( T. grandis  `( w" GjM)h( |}(" iU"#$04233g45 v!G T. grandis  bG~ GwB<=%B 6H7&f 88F 9M (HH A. auriculformis   oh ] -2+*0 ++ _  A. mangium  bB :  H [ ¡¢£();¤(k¥¦<J \"Ff`H =§ ]Ang andMaruyama +33._ %B ¨©> A. mangium ?¨G( ª«¬@­B E A. mangium ¨© A. auriculformis ª«A®¯ °r±²³sB%B"#$043g45&C´µ¶5 Hopea odorata <='ῌD'`H )h( E'7ῌfFD +- · oJ/F¸ defgNOPQRS/G¹wB3 ]K + º»_70   ¼ No. // ῍,**,῎῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏῏ ῌῌ.  -  ,/ .+ ῌ  +333 ,***  ! +/"2*#+*$0 %& '( -.) 1#+*$0 ,*".*#+*$0 *+,-

./



! +*"3*#+*$0 %& '( ...0#+*$0 .*"/*#+*$0 *+,-%& 0 
 -1234567 8 89: ;<=>?>@ABC/-*DE/F7G07HI0
JK/F7G HLMG //0 
 , /-´µ/-  ,/ Q ,* RG '(N O8P7 O8

P7*+70

SM H
TUVWXYZ[G0'(NRG ...0#+*$0OP7\  +/ Q +, ]G ^- _`-ab[cde/. 7f>8 '(ghRG /-*,F iaGj kl O8 imn

imo pqrsZJ*tuRvH BABw

xyz{|} imn~8GI06* imA pq0im}GI0*u0L5>G + Ang L.H. Maruyama (+33.) Growth and photosynthesis ofAcacia auriculiformis and Acacia mangium seedlings planted on sand tailings. Proc.
of the International Workshop of Bio-Refor Kangar 01ῌ1+ , Ehleringer R. J.
Anthony H.E. & Farquhar D.G. (Ed.)(+33-) Stable isotopes and plant carbon-water
relations. ///pp. Academic Press San Diego -  +330 ) ar + : ,,,ῌ,,2 .  ++  +333  .+ DG1W ) ¡a@A .+ , : ++-ῌ+,+ /ῌLai Hoe Ang ,*** G1W
ῌ¢£¤¥¦§¨©ª ) TROPICS3 - : +3/ῌ,*3 0 ,**,a D«r + ¬l­) ab /- : 1-ῌ2* 1  ,**,b D«r + ®¯0 ®°±¯²³U) ab /. : 1+ῌ102 PROSEA (+33.) Plant resources of South-EastAsia / (+) Timber trees : Major
commercial timbers. -13ῌ-2.
  a b No. // ῍,**,῎71