Bull. Natn. Sci. Mus., Tokyo, Ser. B, 27(2, 3), pp. 65- 78 , September 22, 200 1

Mosses of Mts. Abukuma and its adjacent area, northern Honshu,
Japan. 2. Pleurocarpous mosses
Masanobu Higuchi
Deaprtment of Botany, National Science Museum, 4- 1- 1 Amakubo, Ts ukuba,lbaraki 305-0005 , Japan . E-mail : higuchi@kahaku.go.jp

Abstract This study deals with the moss flora of Mts. Abukuma and its adj ace nt
area, northern Honshu, Japan. The pleurocarpous mosses recognized by 1100 specimens comprise 19 fam ilies, 6 1 genera and 11 8 species. Totaly 22 families, 78 generaand 180 species of pleurocarpous mosses are recorded in thi s area. The pleurocarpous moss flora of this area is characteri zed by the following features. (I) It is principally of East Asian and circumboreal species. (2) Fourteen species having their
northernmost locality in Japan are recogni zed.
Key words:
areabryophytes, pleurocarpous mosses, Japan, Mts. Abukuma, limestone

OOJL. f4 '¥:t~ ~PJig ~::. J: -0 I B2fs:91J i'JO) El ~ 5l::f4 '¥: EtJ ttiS"~Jf~ J 1J~:i.fs: 1+1 JlUI: :g:~O) ~iiJ fEU.ll llJ :lil1.
:a:-$ -L, c -t -0 :lil1. ~ ~::. .t3 "' -r IJLP.X: 6, 7 :¥-&u::.~1ifli ~ h , ~H.iH§ ~::.-::> " ' -r 0) Ji.X:-* 0) - :g:B:a:- 3€
3\H.J: (timD , 1995 ) . :i.fs:~~Jt-c· (j: Jltr$1H::. ff.% ~ , tMP1 1:f iJ~'fflJE;J L!:tJWI*%" :a:- £ l::. iNIJ ~ -t -0
1 Hl O)J.il:tffi\1::. Jf~ P.X:-9 -0 UlUHJi (pleurocarpous mosses) 0) ::I '7 -t / < / ;;f, / :7·-tt-f.fi.l' G ~ 3 :7::I''7 f4 i ""f·O)*j!j* 1::. -::> v' -r$R-ff-t -0 .

~1t, H:J.TO) 11 tm?.JT-c·{f·:d.: .

A.

tliii'J!!f. IEH W7d?! II!TtiLAi~ ( ::OIA"E ±tfl. ~) , f!\!!~ 580m,

1994 if. 8J=l 22 B .

B.

tliii'J!!f.IE H W*~ II!T nf! ~ i\: ( ::OJA"E ±tfl. ~) , t!\!!~ 600 m ,1994 if. 8 J=l 22 B .

(::OIA"E :lil1.t9.X) , t!\!!~ 650m, 1994 if. 8 J=l 22B.
D. tliii'J!!f. IEHW?ttl ii!Tfili -5'¥ (::OIA"E:lil1.t9.X) , f!\!!~ 850 m , 1994 if. 8J=l23B.

c.

tliii'J!!f. IEH W*~ IIIT Jt!.nf!~i~ti-

E.

tliii'J!!f. IEH Wi~tl ii!T.%!. 1\: (::OJA"E ±tfl. ~) , t!l!!~ 750m , 1994 :¥- 8 J=l 23 B .

F.

:tliii'J!Jf.v 'b ~ rtr~: # JII i~'d~ .

t!\!!~ 200m,

1994 :¥- 8 J=J 24 B .

timD.iEf§

66
G.

ti'il~~v'V ~ m1fFfillft®ii~~. f~$j 160- 300m,

H.

ti'il~~v' b ~ mlm~II!T:flll, f~$j 320---400 m,

1994 if 8 .Fl 25 8I.

ti'il~~W. '$Uflti!li* IIIT* P' JII i~i,~, t~$i 150m ,1994if8.Fl268o

1994 if 8 .Fl 24 8J.

ti'il~~:tEI.~W/l¥:~111TJL:E (E!A:'Btthi:§X ) , t~$i i00-200m ,K.

:tiil~~nli:II!Tm t ~m:;~~. t~$i 150- 220m , 1994if8 .FJ 28 8~~ G ~1.t.:*"J 1100 ogO)jj':f-!1- ~ 'b t ~:::~lHJT ~ 1-T -::d.:n -r v' ~001994 if8.Fl 278o0~-r O)f~J.fs: (J: OO JL: f.t $ t!J1:~iiJ~iO)f~J.fs:

0

Jt ~*$!.1$i5'~1:!lmf.t$f.40)f~J.fs:J! (:::~31.66 ~ h -r v' ~, 1984
if'::: ~~t.Jrf'il ttht9X:"C·~31.· l t.:t~J.fs: 'b .g.t>-tt-r WH!T l t.:

J!jH:::~31.66 G

0

it.:,

0

4"@lO)~]t"C· ,~Ji'imO) 6li ~ 1!;;f, / 7"+Tf.41.J' G t: 3 7 :I·' 7f.ti "C'O) 19 l4(:::i:>v''lti'il ~ ~lf.t.

:I '7-\' / <' /u 61 ~ 11 8fi0)1:1f ~~If ~?.!. l t.: o ::. 0)¥;!1-*!~:(f i "(· (:::J.js::l1Qj:§j( (:::~t~?.!. ~ ht.:

!J!Hf-!tJHi 22f.478 ~ 180 ti t t;,: ~.~-'J:*O):im I'J "(·~ ~01-:J. T(::: lliiJJiU.J!l: LlJ±th lHFJalillttht9X:O) El f.¥/<~~-t7'Jr,

0

fll'!!F:M¥R t f'il:f* ,:::, 4-@lWH!T,:::ffl v, t.:~r-l"C·IHt ~r. ~ nt;,: 1.1, "') t.:l.lr:tiil ~

*

*"* n

~ tniiJ ~t-? 1t!J, 0) Elf~~::: J: I'J:l1h t9X: J: I'J~-r v, ~ ti 'b El f.W< ~::: ?i' 66 t.: H 0) ~c J"IHJ:
lwatsuki (1991 ) (::: J: 0 f.: o ~ ~ C li~ O)~cJ'Ij(j: 7 !v 7 7 « ·:; r )lifl"(· ~ ~ o ifl~~O)fZ ~c(J:Brummitt & Powell ( 1992) ~::: J: 0 t.:

0$ ~, to~ ~:::~:7[ ~ 4-@lftll ~ ~ ht.:1'.1f ~Mii.J C: ~1ttth.9:,t~J.fs:il):Ji} ( 7000ilJ:iJ'(il984if(:::~31.•~n,O)fltr ':::

* ~n ~ 1i l t.:ti,J::tiil~~~!i' im "C· ~ ~lt~:k$(:::1* ~ ~h'lv'~ )Climaciaceae~~lt.:. $ ~

0

t /~

Musci

::::J rJ t' ...1 ? /:I '7-\' / '7 /Climacium japonicum Lindb.

0:?- / "/-if l'l-;f, / 7"-IT~ tllii:±.l.. oC (23389, 23584, 24 104,24234 ), I (2445 1).

1:: :/'~ :::i'-TH

Hedwigiaceae

Hedwigia ciliata (Hedw.) P.Beauv.

t:-'.) ~ :1··7

ti'il ~l,lfHMiiJ ~t ( 1987).

Cryphaeaceae "J Jv :::i' 7-H

Forsstroemia cryphaeoides Cardot

t: 7- A ;;t..' :1"7

~tff:,~Z::E _t o A (23356), B (23418+

Anomodon minor subsp. integerrimus, 24133), H (24402), I (23848).
Forsstroemiajaponica(Besch.)Paris
Forsstroemia neckeroides Broth. 7

t:-t1 r :I"7 tH~'f:_t o £(23485,24 179,24192).
r A ;.t..' :1··7 ~t~'f:, :'5 _t o 0 (7188), E (23555 , 23578),

J (7158) .

Forsstroemia trichomitria (Hedw.) Lindb.

A ;f..' :1·· 7~t ~'f:,*-i. E , :'5 .1.. . A (23346), 0(7187, 7 189, 23453), E (23483, 23526, 23572), G (23679), H (24403), I (24467), J (23923, 23970,
24499).

Leucodontaceae

Dozyajaponica Sande Lac.
Leucodon lute us Besch.

1)

A :1"7

1' 5I 7- :j' 7-H~j~'f: _to') A :1·· 7 "f: r·· ~G (23667).

ti'il ~ ~tllil~iiJ~t ( 1987, as Leucodon exaltatus).

67

Trachypodaceae 1>. ~ T :::i" 7-H
Duthiella speciosissima Broth. ex Cardot ? "/.b. 7 ·::J7 tlli ~~ l@:~ [t (1987).
Trachypus bicolor Reinw. & Hornsch. .b. ".) T ::f7 tlli ~~lMio[t ( 1987).

Pterobryaceae
Eumyurium sinicum (Mitt.) Nog. T '7 :::l7
Pterobryum arbuscula Mitt. ~ .b. 0 :t 7

1::: 1>. D :J. 7-H
tlli~~t@:~[t (1987).

;jjj f1': _t . G (23664, 23702), I (23860, 23865 ,

24434).

Meteoriaceae
Barbellafiagellifera (Cardot) Nog.

I\ 1 1::: 'E :J.7-l;j.

.:t- 3 A ~ 1 1- :::l7 t5i: , 7 ~ /A , :1 / 7 1) - 1- ~_t .

I (23854, 23857, 24432 , 24436).
Meteorium buchananii (Broth .) Broth. subsp. helminthocladulum (Cardot) Nog.
7

;jjj ~~':'fi L:1 J \ 1 ~ -'f: :1··

I (23863 , 24458), J (7159, 7161, 7162).

Neckeraceae ~ 7 :1.. 7;]'4-\'? 1- ~ 7 ::J7 ;jjj~~' tit ' i ::O , :'fi L
E (23498, 23533), G (23631 , 23669), H (23719), J (239 18).
Homaliadelphus targionianus (Mitt.) Dixon & P.de la Varde var. targionianus -7 7- ~ 7 :1.. 7~U~, 'i::O, :5 L
H (23758 , 23806), J (7111, 23960, 23979, 24500).
Homaliadelphus targionianus var. rotunda/us Nog. ~ ;J. -7 7- ~ 7 :1··7 MD (1991). :;$:~
tiLl: v ·;; F-r··--7 7··-:; 7 W'UliTEI~ f:!iH!f)i5jff1:1: ~io~!f 2000 ) f:.t:>v'"l , *@.il&JTI:t!Hi<*@.il&JTI:t!'l- I 'J;JO f: t§'J:E ~-r ", J.J .
Homaliodendronflabellatum (Sw.) M.Fleisch. .::t--7.. 7- ~ 7 :1.. 7 ,r;;o_t. I (23853).
Neckera .fiexiramea Cardot :1 ;J. 1) /A -:::l7 tlli~~tMio[t (1987) .
Neckera goughiana Mitt.
'J ~ 7 ::f7 'i::O, :'fi_t . D (7190, 23441 , 23467), E (23496,
23504, 24174), J (7168, 7 176, 23959, 24492).
Neckera humilis Mitt. 7- -1'~ 7 :1·· 7;jjj ~~, ffij::0 _t. E (24188), H (2440 I), I(23869, 23904, 24430, 24446), J (71 12, 23911 ).
Neckera konoi Broth. ex Cardot -7 tJ i' ;J. 1) /A :1.. 7 tlli~~li~io[t ( 1987).
Neckerapennata Nog. J \ i' ~ 7 :1··7 tlli ~~li~P.J [t (1987).
*Neckerapusilla Mitt. ~ ;J. ~ 7 :1.. 7 ;jjtfl': _L A (23348), I (23864).
Neckera yezoana Besch. ~ '/ ~ 7 :1.. 7 tM~~ L A (23340), C (24247), E (23573, 24202),
G (23686), I (23850), J (23947).
Neckeropsis nitidula (Mitt.) M.Fleisch. 1) ;J-<" / :1.. 7 ;jjtfl': , 'i::O, :5 _t . G (23655, 23666,
24290), I (23822, 23843 , 24460).
Thamnobryum alopecurum (Hedw.) Nieuwl. .:f- ') i' / ;;f :1·· 7::0 _t . B (23423), C(24235 , 24239), J (7119, 23922).
*Thamnobryum coreanum (Cardot) Nog. & Z.lwats. :1? / 1- 7 / ;;f :1·· 7::0 _t. C(24087).
Homalia trichomanoides (Hedw.) Schimp. var. japonica (Besch.) S.He

n

*
*

*, 'i

'i

'i

68
Thamnobryum planifrons (Broth. & Yasuda) Nog. & Z.lwats.

~7

r 7/ :t -:J.. T

tlil~IJ1l.t@:~bl &t (1987).

Thamnobryum plicatulum (Sande Lac.) Z.Iwats.

=\'- ·'f'f- ~ 7·· -:J.. T(23362), D (24152), E (23539, 4180), G (23652).
Thamnobryum subseriatum (Mitt. ex Sande Lac.) B. C. Tan!lii £ L . A (24059), C:t :t r 7 / :t -:1··7:fM~~.

!lii£,:'fi L. C (24238), D (23446), H (23760), I (23887).
Lembophyllaceae !-- 7 / :t :::i.7-H

r 7 / :t -:1·· T

tlil ~ iJ1(. :f@: ~ &t

T .I. ;f 7 Y -:1·· T

tlil ~ IJ1l. t@: ~b; &t

Dolichomitra cymbifolia (Lindb.) Broth. var. cymbifolia(1987).
Dolichomitra cymbifolia var. subintegerrima Okam.

I\( 1987).
Dolichomitriopsis diversiformis (Mitt.) Nog. -:J 7 -lT -:1·· 7~~, ;f~, !lii £, :'fi __1:. E(24181), F (23617), G (23641, 23677, 23703), I (23819, 23846, 23907, 24429, 24455 , 24457+
Hypnum plumaeforme var. minus).
Isoth ecium subdiversiforme Broth. ~ ;' -:1 7 -lT -:1··7 tli\ ~IJ1l.:f@:~ [t ( 1987).

wt

Hookeriaceae 1' 7·7 :::i' 7-H

Hookeria acutifolia Hook. & Grev.

7 7·· 7 -:J.. T

tlil~IJ1l.tMbJ &t (1987).

Hypopterygiaceae 7 '>-\"' 7 :::i.7-H

Hypopterygium fauriei Besch.
23872), J (23983) .

7 ;;· -1' 7 -:1.. T

Theliaceae

Fauriella tenuis (Mitt.) Cardot

1¥~*, E , :'fi__1:. H (23766), I (23845,

t: 7 :::i' 7-H.I./f·r; o -:J -:1··~r-t F=f

I (23900, 23906, 24456).
Myurella sibirica (Mii1l.Hal.) Reimers!liiwt*, -j¥1]*, !lii £__1::. G (24333),

v 1 Y -:1··7 :'fi L. D (7175, 7 177, 7179), E (23502,

23527, 24169).

;;tq_tt;;J:-: td: -z"tlil~IJ1l. "("'j:#f:J!WI& ilJ ;>j> G ¥1H5 ~ n -c V' t.:t.~ r ( tlil~IJ1l.tMbJ&t 1987) ' ~m: r:p :g:~(7) E }1( :'fi±t!rt9X ': :73'::1'!1 L -c v, J.J -: t.~r~t~r. ~ t.:. ;1Js:f.t ,;;~:
v ·; r-··-r-7 7···; 7 (l~~IT§?'!!:f:!i1:31JaJlffj:_j:_~~Jf; 2000 ) l:i>v'""C, ~i~ITI:'i:J'l-f.t ( *~i~c.

@1 (7)W\llit ': J:. I'J , ~or m:~l'l: i'i1i
ITI:t!'!- I ~ )nl:fg7£~tL-cv'l.J .

Fabroniaceae :J :::i' ;;!, :::i' 7-H

wt*,

Schwetschkeopsis fabronia (Schwiigr.) Broth. 1 Y, 7 -:J.. T!lii£, :'fi__l:. A (23338,
23358), D (23456, 24157), E (23 510), I (23849, 24437), J (23946, d23948), K (24016, 24024,
24025).
Schwetschkeopsis robustula (Broth.) Ando :t :t!::: ;' !::: T -:1··7 !lii £, :'fi__l:. B (24125), G(23694), I (24470) .

69Leskeaceae

r'J .A 7' [:] :::i' -7-l'i-*Okamuraea brachydictyon (Cardot) Nog.
'J :t fJ .L... 7 :.i 7
tRt ~, !5 J::.. E (24186), F(23604), H (23770) .
Okamuraea brevipes Broth. ex Okam . ::I~/ :t fJ .L... 7 ::1··7 :f:;,!~;!.~;j:~!tP.J ~t (1987).
Okamuraea hakoniensis (Mitt.) Broth. :t fJ .L... 7 ::1··7 :t:;,!l%J,!f.l.;J:~!tP.J~t (1987).
Pseudoleskeopsis zippelii (Dozy & Molk.) Broth. 7 +T 1 r ::1" 7 ~iD , !5 J::.. G (24305 ,
24319), J (244 71 ), K (24026).
Pterigy nandrumfi.liforme Hedw. ;f, j v 1 r ::J"7 !fiJ::.. I (24438) .
Thuidiaceae :/ / 7' :::i' 7HAnomodon abbreviatus Mitt.

~ -\'7.:\='';t'? ~ ::J"7.:C F ::\'-:t:;,!l%J,!f.l.;J:~!tP.J ~t (1987).

Anomodon giraldii MUll. Ha!. :t :t .:¥' ;t' '7 ~ ::1" 7 .:C F ::\'- tRt ~, ~iD , D :11[, !5 J::.. C(23360, 23397, 24246), D (7 184, 23438, 23462, 24149), E (23515, 23517, 23525 , 23552, 23553,
24161 , 24164, 24177), H (23726, 24349, 24354 ), I (23905), J (23987).
tRt ~~, ~iD, !5 J::. . C (24241 ), D
Anomodon longifolius (Brid.) Hartm. ::\'- :J. 1 r ::1·· 7(7 193,23444, 23458), E(23531 , 23547) .
Anomodon minor (Hedw.) Lindb. subsp. integerrimus (Mitt.) Z.Iwats . .:¥' ;t' '7 ~ ::I" 7 .:C F ::t'tRt~~. ~iD ,D:f1[ , !51:.. B (23418+Forsstroemia cryphaeoides, 23427, 24113), C (24232), D(24 151), E (23556, 24206, 24208), H (23732, 23733 , 23773 , 23803, 23805, 23809, 24375), 1(7117, 7160, 23958, 23962), K (24023).
;j:~, ~iD, !5 J::.. C (24096), D
Anomodon rugelii (Muii.Hal.) Keissl. .I.'/ 1 r ::1·· 7(24137), E (23497), 1 (7114).
Anomodon thraustus MUll. Ha!. ::I '7 / ::\'- :J. 1 r ::J"7 :t:; .il%J.!f.l. 1Mia ~t (1987).
Anomodon viticulosus (Hedw.) Hook. & Tay!. ::\'-A j ::\'- :J. 1 r ::1·· 7 :t:;,!l%J,!f.l.t~!tm~t (1987) .
~f. D, !5 J::. . C (23405), F (23607,
Boulaya mittenii (Broth .) Cardot T '\' ;t' A 7..' ::1" 724275 , 24278).
B1yonoguchia molkenboeri (Sande Lac.) Z.lwats. & lnoue~
7"::1"7 ffi]*, Jlilit~±.
~iD J::.. C (23585), E (23521 , 24196), 1 (24475).
Claopodium aciculum (Broth.) Broth. I \ 1) ::I"7 :t:; .!l%J.!f.l.t~ !tm ~t ( 1987).
Claopodium assurgens (Sull. & Lesq.) Cardot '7::\'-1\l\ 1) ::1·· 7 tRt~~. :'51:.. D (23459), E(24176, 24213 , 24214, 24216).

*/ /

*

'J I\ 1) ::1··7 :t:; ,!l%J,!f.l. ;j:~~ia ~t (1987).
Claopodium gracillimum (Cardot & Ther.) Nog,
Claopodium pellucinerve (Mitt.) Best 7 r I \ 1) ::I"7 tRt~, ~iD , !51:. . A (23354), D(24150), E (23495 , 23561, 24218), H (23720, 23723).
~f. D, !5 J::.. D
Claopodium prionophyllum (MUII.Hal.) Broth. T fJ" A ''./I \ 1) ::1·· 7(23452), E (24189, 24205), H (24353 , 24356), 1 (7125 , 24488) .
Haploc/adium angustifolium (Hampe & MUII.Hal.) Broth. / ~ I\.=. '7 ::1·· 7 tRU~, 1l~~iD, !51:. . A (23337, 23350), B (23436), D (24154), E (23562), F (24260), G (23713 , 24324),
H (23738), 1 (23914).
Haplocladium microphyllum (Hedw.) Broth. ::I ;I. I '\'::\'- ;J. ::1·· 7~~, ffi] *, !fi J::. . C

*,

wJ

70
(24089), D (23469), E (23560, 23571), G (23649, 23688), H (23744), I (23894).
Haplocladium perparaphyllum R. Watanabe -:c r'; 3 r'; :I / ·{ / ;f ;;. :I.. 7"( 1987).
Haplo caldium strictu/urn (Cardot) Reimers

A:,/ Y / 7 .. :1.. 7"

Haplohymenium longinerve (Broth.) Broth.

:t; ~!!¥'.- M: ~m at:t; ~J!¥1.-tW!~at (1987).

T ti A:/ 1 1- :1··7

l!ii:tl J:. . F (23606, 23610,

24276).
Haplohymenium pseudo-triste (Miill.Hal.) Broth. :I J{ / 1 1- -:::lJr -E J:. . 1 (23986).
Haplohym enium sieboldii (Dozy & Molk.) Dozy & Molk. 1 '7 1 1- :1.. 7" .:C F .:f l!ii:tl J:. . E(24222).
Haplohym enium triste (Ces.) Kindb. 1 '7 1 1- :1·· 7 ~H1, iflj*J:. . A (23347, 23357), F(23603 ), H (23 730), I (24431 ).
Herpetineuron toccoae (Sull. & Lesq.) Cardot 7 -t / :1.. 7" ~H~ , iflj *J:. . G (23665), I(24466), K (240 18).
Heterocladium angustifolium (Dixon) R.Watanabe
J { 1 1- '/ Jv :1·· 7"-E Ji fill (1981 ).

* '/Heterocladium capillaceum Broth. ex Iisiba

1-t / 1 1- ·:; Jv:I'Jr:t;.~J!¥1.-tMm at (1987).

Heterocladium tenellum Deguchi & Suzuki :t;~J!¥1.- tW! ~mat (1987).
Miyabea fruticella (Mitt.) Broth. ~ -'t X.. :1··7 ~t~~ . -f¥1]* , l!ii:tl J:. . F (23602, 24282), I(24462), J (24493).
Miyabea rotundifolia Cardot

7 Jv J { ~ -'t X.. :1·· 7"

tiffi D ( 1991 ).

Thuidium contortulum (Mitt.) A.Jaeger i, :/ v Y / 7' :1'7" -E Ji fill ( 1981 ).
Thuidium cymbifolium (Dozy & Molk.) Dozy & Molk. 1::: J.. Y / 7' :1·· 7" tlt l!ii :tl, -E J:. .
A (23325 , 24071 ), B (24131 ), C (24097), D (24155), E (24185), G (23700, 24337), 1 (23952).
*Thuidium glaucinoides Broth. 1) :1. r'J ;f :1. r'J Y / 7" :1·· 7" l!ii 1J , -E J:. . A (23333), B(24112), E (23503 , 24203), H (24399).
Thuidium kan edae Sakurai
1- -'t 7 Y / 7" :1·· 7"~t ~ ,-j¥1] * ,± , l!ii D , -E J:. . A(24060), B (24135), C (24240), E (23537), G (23656), I (23841), H (23745 , 23761 , 23799, 24355,
24367, 24371).
Thuidium pristocalyx (Miill.Hal.) A.1aeger 7 :t Y / 7" :1·· 7" l!ii D , -E J:. . D (23439), E(23550, 24209), F (23620), G (24313), J (7115 , 7154, 23981 , 24502).
Thuidium pygmaeum Bruch & Schimp. ~ ;) / :I Y / 7·' :I.. 7" l!ii 1J ,(23993).
Thuidium recognitum (Hedw.) Lindb. var. delicatulum (Hedw.) Warnst.

7"

C (23409), G (24344).
Thuidium sparsifolium (Mitt.) A.Jaeger-E _t . H (24352), J:I J ~· / ..I. 'f Y /7'' :1·'+'V *Y / 7'':I··~r l!ii :tl , -EJ:. . A (23327 + Eu-

rhynchium savatieri, 23328), B (24129), E (23492, 23518, 23563, 24190, 24211 ), G (23642, 23645 ,
23650, 24346), H (23721 , 23731, 23743, 23748, 24374, 24390, 24392 + Cratoneuron filicinum), J(7144, 23917), K (240 14, 24019, 24043) .
Thuidium subglaucinum Cardot :t :t 7 :t Y / 7" :1. 7" l!ii:tl, -EJ:. . C (23370), G (24320),

J (23928).
Thuidium submicropteris Cardot:I

7 / 1 ~ Y / 7" :1.. 7" l!ii:tl J:. . E (23487).

71
Amblystegiaceae-t' T :¥. :J.-7-H

Amblystegium serpens (Hedw.) Schimp.

~ ;J. "tf';f':::f'7

ti'ii~~;f@:!Jo/.J~t (1987).

Amblystegium varium (Hedw.) Lindb.

T ~ A ;;· f T :¥' ::J 7

~m 0

*

( 1992).

Calliergonella cuspidata (Hedw.) Loeske f ') /::J.. 7 ti ~~;f@:!Jo/.J~t (1987) .
Campyliadelphus chrysophyllus (Brid.) Kanda ::J :tf;f,J\1 :::1··7 tliUf:, ifrl*,±, -'5J::.

D (7 173), E (2348 1, 23505 , 24160), H (23762, 23765, 23771 , 23811), J (7142), K (24012) .
Campy lium hispidulum (Brid.) Mitt. "tf'::f·::J-·7-=E V:f- 13'~*1:_· A (23342+Brachythecium p lumosum) .
Campylium squarrosulum (Besch. & Cardot) Kanda( 1987).
Cra toneuron filicinum (Hedw.) Spruce'J ') I \ f~ ;;:.: / Jf :::1.. 7

T .:¥. :::1·· 7

ti ~ ~ 1@: !Jo/.J ~t± , ~i.E, -'5 J::. A (23318,

tlit~f:,24055 , 24073), D (7171), H (23736, 23737, 23756, 23796, 24361, 24370, 24372, 24373 , 24378 ,
24379, 24392+Thuidium sparsifolium) , J (7 151 , 7156, 23945), K (24006).
Hygrohypnum ochraceum (Wi1son) Loeske '7 A :f- / ;J. 1) :::1.. 7
ti ~~ ;f@:!jo/.Jtl_; (1987).
Leptodictyum humile (P Beauv.) H.A.Crum /\f7ff'.:¥·::J.. 7
ti ~~ ;f@:!Jo/.J ~t (1987, asLeprodictyum kochii).

Brachytheciaceae
Brachythecium auriculatum A.Jaeger
um auriculatum).
Brachy thecium brotheri Paris7' ;;t :¥. x :J. -7-H
ti~~1MP.I~t (1987, as Camptorheci-·lf7 / :f- :::1.. 7

7 71\ ~ / :J :::1··7~;f@:±J::.

C (23406).

T :ti ~ / ;;· :::1··7 1.lt ifil*, ± , ~i.E , ::J /7 ')- Ht, -'5J::. A (23343), B (24120), D (23451, 23475 , 24145, 24153), E (2349 1, 23542,
2420 I), G (23660, 24303 , 24327), I (23816), J (23949, 23991 ), K (24002, 24005).
Brachy rhecium coreanum Cardot ::J < / ~ / ;;· :::1.. 7
ti ~~ 1@:!Jo/.J ~t (1987).
Brachythecium buchananii (Hook.) A.Jaeger

Brachythecium garovaglioides MUII.Hal.

7 ~ "J '/::J.. 7

ti~~j:@:~P.I~t ( 1987, as Brachythe-cium wichurae) .
Brachythecium glareosum (Spr.) Bruch & Schimp.

/ 7··7- ::J.. 7

Brachythecium helminthocladum Broth. & Paris~ .:C ~ / ;;· ::J.. 7

ti~~j:@:~P.Itl.; (1987).
~i.E,::J / 7 ') -r ~.

-'5J::. A (24074), D (23468), E (23516), J (23978 , 24479, 24503), K (24003).
Brachy thecium kuroishicum Besch.

7 0 1 / ~ / :J :::1·· 7

Ut~,

if~* J::.

(24227).
*Brachythecium moriense Besch. ~ / :J :::1.. 7 jrj*l:_· E (24204).
Brachy rhecium plumosum (Hedw.) Schimp. var. plumosum I \ ;f, ~ / :J :::1.. 7

C (24 100), Ejr)*, ~1@:±,~i.E, -'5J::. A (23342 +Campylium hispidulum) , C (23369+Homomallium connexum , 24105), E(24 199), F (24264, 24273 ), G (23639, 23643 , 24294, 24298, 24300, 24301 , 24345), H (23788,
24382, 24387), I (23899).
Brachythecium plumosum var. scariosifolium Cardot :f- 1::: / ~ "J :J :::1·· 7
ti ~ ~ 1@: ~m ~t( 1987).
Brachythecium populeum (Hedw.) Schimp. var. populeum

7 :t .:¥' .)( :::1·· 7;f.lt ~f.E, -'5 J:: .

72
B (24114, 24117, 24124), C (24230), D (23466, 24138, 24144), E (2419 1, 24207), H (23729,
23753, 24358, 24364).
tlli !flh ~T 7f .I./ 7 :t .:¥' Y. -::1·· 7'

Brachy th ecium populeum var. longisetum Bruch & Schimp .
t@:~at (1987).

Brachy th ecium pulchellum Broth. & Paris '/ 1' ~ ':J ·;) -::1· 7' tlli !flh ~;tMiaat (1987).
Brachy th ecium reflexum (Starke) Schimp. 7 7 .I./ ~ ':J "/' -::1"7' ir~*..l.. D (23478).
Brachythecium rivulare Schimp. 7:::... -::1"7' ilii::fi ...t. A (24057 + Bryhnia novae-angliae).
Brachythecium rutabulum (Hedw.) ~ D / \ / 7 -IT -::1··7 Jm' ;j:@:±..l. . B (23426).
Brachythecium uyematui Broth. ex Cardot-::1 ~ ':J "/' -::1·· 7'

jjt ~f:, il!i E ...t . C (23368), E

1'/;f,-::J.. 'i'jji*, ;t.R, jl]*, ±, ilii::fi, :'fi(24175 , 24224).

Bryhnianovae-angliae(Suii.&Lesq .)Grout...t .

A (24052, 24057 +Brachythecium rivulare), C (23392 , 24079, 24082, 24099), D (7194,

24141 , 24156), E (23568 , 24197), G (23712, 24293 , 24317), H (24404), K (24046).
tlli!fih~;j:@:~at ( 1987) ~.::. J:: tUi',;;f::lt!!.~i.P G ;f, "/' v f/ ;f, -::1··7' (Bryhnia sublaevifolia Broth.

& Paris) 7Jrfa1!r~h 'lv'.QiJr, ;;f:fi:liNoguchi (1991) (.::.J:: I'J 1' / ;f,-::J"JrO)~ / :::...L. t ~ht.:. .
Bryhnia tenerrima Broth. & Yasuda~} 1' / ;f, -::1··7':'fi...t. G (24297), H (23764).

Bryhnia tokubuchii (Broth.) Paris .I. '/1' / ;f, -::1"7' tllilfih~f@:~ at ( 1987).
Bryhnia trichomitria Dixon & Ther. .:\'- / .:C '7 1' / ;f, -::1·· 7' ;!'R, il!i E, :'fi ...t. D (7183 ,23437), E (23513 , 23541 , 23558, 24210).

T tf;~· ~If;\ -::1"7'Cirriphyllum piliferum (Hedw.) Grout

Jl!l; ;j:@:±...t . C (23388).

**Eurhynchium angustirete (Broth.) T.J.Kop.
'J ;\') Jv!\ ~ -::1··7' ;!'R...t. E (23 575).
Eurhynchium eustegium (Besch.) Dixon :t :::.. ~ ':J ;;· -::1·· 7' jjUf: , Jm' ;j:@: ± , il!i E ..l. . C(23363 , 24090, 24106), H (23759), I (23875).
'/ 7 '/ T .:¥' -::1''7' .:C F.:\'-

Eurhynchium hians (Hedw.) Sande Lac.

ilii::fi, :'fi ..l..

A (23352),

B (23434, 241 09), C (24229), G (24314, 24334), H (23741 ), J (23938), K (24036).

Eurhynchium savatieri Schimp. ex Besch.

jlj * ...t . A (233 19, 23327 +':J 7 '/ T .:¥' -::1" 7'

Thuidium sparsifolium, 24058 , 24066, 24067 , 24068) .
.:t-7·· 1) 7 .:¥' -::1·'7' ~ (Eurhn chium ) ~= li, tlli !fih~;j:@:~at ( 1987) 1.::. J:: hli';;$::lt!!.~7'.1' G 7 tJ -IT-::J"':J Jv/\ ~ -::1"7' (Eurhy nchium striatum (Hedw.) Schimp.) 'b ii:T .Q.:::. t 1.::.1;t-::> "( v' .Q 7Jr,::<js:fi:li 8 ::<js: ~= iiT .Q .:::. t 7Jr§1 r"~1Jl. ~ h "( .H I') (cf. Noguchi 199 1), ::;$: § ~~-c· li lW<)'~ L t.:. .

Homalothecium laevisetum Sande Lac.

7 ':J 7''-IT -::1·· 7'

wt~f:,

il!i ::fi, :'fi ...t . A (2334 1), B(24118), D (23440, 24143), E (23490), J (7 148, 23980).

Myuroclada maximoviczii (Borcz.) Steere & WB.Schofield;f, ;t.' ~ / :t -::1··7'

jjUf:, ±, :'fi...t. C (23367 , 24088), D (23448, 24139), H (23749), K (24030).

Palamocladium leskeoides (Hook.) E .G.Britton

7 ':J 7·· -IT -::1" 7' .:C ]--·· .:\'-~i E,:'fi ...t.

H(23754), J (7 169, 23963 , 23977).

Platyhypnidium riparioides (Hedw.) Dixon

7 :t /\1 -::1··7'~iE, :'fi..l. . A (23 329, 24054),

C (23393), F (24266), G (23648, 24304, 24335), H (23793 , 24360, 24362, 24389, 24391), J (7 153,
23932, 24472), K (24029).

Rhynchostegium contractum Cardot-IT 1 ~ .:r. '7 T / 7· -::1· 7'

jlj*_t · A (2332 1).

73
71-\' ~·7

Rhynchostegium inc/inatum (Mitt.) A.Jaeger

t~~~f~~[t, (1987).

t~ ~ ~f~ ~ [t, (1987) ~.:: J: tU;f, 21': itJ2 t~ J: V) I \ ;t, 71 -\' ~· 7 (Rhynchostegium plumosum
Ther.) O)~cf~iJr~ .Q ;~;r, /.f':fiL:t Noguchi (1991) ~.:: J: V) 71-\' ~··7 0) "./ / ::: L. t ~ ht..::.

Rhynchostegium pallidifolium (Mitt.) A.Jaeger~ 71-\' ~·7

ifil*. ± ..1:. . C (23380, 23412),

I (23902).
Rhynchostegium rotundifolium (Brid.) Bruch & Schimp. '7 Jv I { 71-\' ~·7 MD (1996).
71-\' ~·7 ~ (Rhynchostegium) ~.:: (;;!:, t~ ~~f~~P;[t, (1987) ~.:: J: hii'/.f':±i!!~/.P G / ~ .:¥' 1) 71-\' ~-· 7 (Rhynchostegium serrulatum (Hedw.) A. Jaeger) ~ i'ff T .Q .::. t ~.:: 7J:- -:: > -c lt' .Q 1Jf, /.f':fi (;;!:
B/.f':~.::i'ff-t .Q .::. t ;~;rlitif:r.,~t.ll. ~ h -c i3 V) (cf. Robinson 1987), /.f\:13 f~-c·u:~ff<jH.J..::.
Entodontaceae ") -\1 :::]" -Tl'l1::: P /\ 'J-\' ~··7

Entodon challengeri (Paris) Cardot:fflt*J:.. A (23336, 23349, 23359), D'i(23471).
*Entodon concinnus (De Not.) Paris subsp. caliginosus (Mitt.) Mizush.

'7 Jv I~- 'J-\' ~· 7{] 1:. E (23557).
B21': i'ff 0) ·;; -\' ~· 'T ~ (Entodon) (;;!: Mizushima ( 1960) ~.:: J: V) 1iJf~ ~ h, -t- 0) 9'1-c· < Jv 1 { ·;;-t ~·7 O)t~:r:~. *"*~~. -ttlff~. in~~. ~~~. s ilitf~O)~/.f':iJf~ 1ffl ~ n -c v, .Q . -tO)f&. LlJ~~ (:7](~ 1971 ) , i%S9iD~ (J:.H1ili 1979) , ~-~~ (7klff1ili 1987 ) , :%:~£~ (MD 2000) iJ' G¥1H5·1Jr~ V), .::. h 1-c· ~.::9;o G h -c v' .Q 001*J0)7J';fj] ~~ I ~.::;7F-t . /.f':f!L:U~ <7-\' iJ' G 9'1 00, -5i't3, B21': ~.:: iJ'iT -c 7J';fj) L, 71-lfliJC$' -T I \ 1 ~·7 ~.:: 1J;J, -c i3 V) , 'J -\' ~·7 ~0)11QO)fi t ii*O)Jl~~iJr!J:JTJii-c· 2-4 *Hl!l?l.l\\iH.:: 7J:- .Q .::. t -c·IR~IJ ~ h.Q. 001*1 -c· (i 3:. t L -c n
Jj(:fiitJJ.~~.::~ G h, ~ lmfi ~.~- L t..::i~;J:N.Pin.::i> v'-c ~ jl{;/1: t Dfii'h.Q 1J}j(:fi0)1.i~Ji1Jr~ G h.Q t~ rJT-c-~IJ. OOI*J~.::i>it.Q~t~lt7J:".Q. 7J:-i>, +*~O)iwiiLlJJ: 1J ~¥1H!:r~tL-cv't..::1Jr(i~lff 1975 ) ' +*~ .31: 9'1 :'R:t~~~g ~.:: 1!f:tg ~ h -c lt \ .Q -t- O)¥Jii!:f0) ~ t ~.:: 7j:- "? t..::t~/.f': ~ Wnl"'t..:: t .::. 0 , -t- n '± < Jv 1 ~- -:; -\' ~·· 7 -c· i±7J:- < ;t 'J ~ ·;; -\' ~-· 7 -c· ~ .Q .::. t H1H?.?. L t..:: •Entodon curvatirameus Cardot'7 71" 1) 'J-\' ~··7t~~~f~~[t, (1987).

*, 'i {], :E ..1:..

Entodon jlavescens (Hook.) A.Jaeger .I. .Y ·;; -\' ~·· 7
ffij
24253), G (23677, 24336), H (24393), I (23839, 23842).
Entodon giraldii Mii1l.Hal. ~ ~-·}. 'J-\' ~· 7 t~ ~~ l~~ [t, (1987).
Entodon luridus (Griff.) A.Jaeger
Entodon scabridens Lindb.

7

717 7

F (23618,

r -IT T 'i ~· 7 t~ ~~;j:@:~[t, (1987).
r ·;;-\' ~·7 t~~~;j:@:~[t, (1987).

Entodon sullivantii (Miill. Ha!.) Lindb. var. versicolor (Besch.) Mizush.

*, 'i {],*'J ~ ·;;-\' ~·7:fflt

E ..1:. . B (24115, 24116), F (23598, 24263), G (23644, 23707, 24302), H (23727,
24351), 1 (23912, 23940, 24473), K (24004, 24032).
Plagiotheciaceae-tt T ~ ::f -Tl'l-'7 Jv 7 -IT ~··7

Jli/;l@:±J:.. C (23365).
Plagiothecium euryphyllum (Cardot & Ther.) Z.Iwats. :t:t-!T-J-5'"~"7-'f: F.:f :fflt*. ifU*,
Plagiothecium cavifolium (Brid.) Z.Iwats.

!5..1:. . C (23379, 24077), G (24342), I (23824).
Plagiothecium laetum Schimp.
curvifolium ).

T / 7'"-!T T 5'" ~· 7t~~~M~[t, (1987, as Plagiothecium

74

-r

~I. B :<jq: H ~t J., '"<' Jv ;\ 'J-\' ::t 7 (e ) 0)5)-;;fp (Mizushima (1960) 1ili f: 'b !::<).
Fig. 1. Distribution of Entodon concinnus subsp. ca/iginosus (e ) in Japan (after Mizushima( 1960), et al.).

75
Plagiothecium nemorale (Mitt.) A.Jaeger~ -t? 1T T 5'.. ::tl;-

i!t~f:.

IIRJt~±,

~in,

EL .

IIRlt~± ,

~in,

EL.

B (23433), C (23386, 23391 ), F (23597, 24265), G (24295).
j-jj' I\ '/ :::i. 7 H

Sematophyllaceae
Brotherella henonii (Duby) M.Fieisch.

i!Uf:, f;'l]*,tJ tJ.. ~ ::fl;-

E (23580, 24225), G (24283), I (23830, 23832 , 23874,23880,23889,23901,24454, 24461).
Pylaisiadelpha tenuirostris (Bruch & Schimp. ex Sull .) W.R.Buck ::I -'ET- 1 1- ::I.. I;- i!t*,f;'lj * L. B (23431), C (23375, 23377, 23378, 23394, 24102, 24242), D (7195, 7196, 23470,
234 76, 234 77, 24159), E (23577, 24223 , 24226), F (23594), I (23890, 24428), K (24020).
Sematophyllum subhumile (Miill.Hal.) M.Fieisch. subsp. japonicum (Broth.) Seki T ff;\ / :::1··

7

tlU%W-tMiiJ~t (1987) .

Hypnaceae 1\1 :::i'7H7-lt :::1··~;-

Callicaladium haldanianum (Grev.) H.A.Crum

i!t~f:.

1¥U*L. C (23374, 23381 ,

24085, 24236, 24248), E (23538), 1 (711 0, 23925).
Ctenidium capillifolium (Mitt.) Broth. 7 / J J\ :::1··~;- ±, ~in, EL. G (23661, 23699,
2431 0), H (23798, 23800), I (23825, 23833, 23885, 23886, 24418), 1 (7137, 7150, 23954, 23957,
23971, 23992), K (23998, 24008, 24045) .
Ctenidium hastile (Mitt.) Lindb. ::I 7 / J J\ :::1··~;- E JH@ (1981).
Ctenidium percrassum Sakurai :t:::.. 7 / J J\ :I.. I;- tii!D (1995).
EL . G (23673 ,Ectropothecium obtusulum (Cardot) Z.Iwats. :::.. 7·· J\ 5' 7 T if :I.. 724322).
Ectropothecium zollingeri (Miill.Hal.) A.Jaeger
24047).
Glossadelphus ogatae Broth. & Yasudafi ( ~i~JE'/:J'!.I~J! )EL.

K (24042,

EL. G (23653).
1T 'j J \7 '/ ::I :::1·· 7 E J'l1@ ( 1981, as Symphy-') 7 / 1::: 7 / ;t' ::I.. I;-

Gollania japonica (Cardot) Ando & Higuchi
odonjaponicum).
::ljq_it.;!: v-:; F-r·- 5' 7···; 7:t :t 1::: 7 "J ¥ :::1·· 7

WV$Hf§~1*=81.fflJ!ff~I=A:.~iiJ~JI!; 2000)

t:t§''i:E~tL--cv'6.

t:Hv'""C,~i~!Etl'!­*, m

Gollania ruginosa (Mitt.) Broth. / '7 7 ·; ::I :::1·· 7'f¥Ut~, ~in L. A (23332,
24072), C (24233), E (23523, 23524, 23536), G (23685, 23687), H (23775, 23802), 1 (23950), K(24013).
Gollania taxiphylloides An do & Higuchi:f- -\' 7 J \ 7 ·; ::I :::1·· 723551 ), H (23724, 23735, 23755), 1 (23984) .
Herzogiella perrobusta (Broth. & Cardot) Z.Iwats.

~

~in,

+ J 7 1 + 1 :I.. I;-

EL.

E (23 549,

i!t*, ±L. B(23430), F (24258), 1 (7135).
Herzogiella turfacea (Lindb.) Z.lwats. '/ 7 -'E J \ 1 :::1··~;- i!Uf: L . C (241 01 ).
Homomallium connexum (Cardot) Broth. .I. '/:f- '.J. 5' :::1··~;- i!t*, ~in, EL. B (23429,
23435), C (23369+Brachythecium plumosum), D (23473, 24136), H (23740, 23746, 24357,
24369), 1 (7113, 23961,23975 , 24480, 24497), K (24021) .
Homomalliumjaponico-adnatum (Broth.) Broth. 't? 1- :f- '.J. 5' :::1··~;- tlii~W-tMiiJ~t (1987).

76
Honda e/la caperata (Mitt.) Ando, B.C. Tan & Z.lwats.

*/~tfl':,-'1"·::!7:'fi..t . E (23528),1(7132, 7170,23969,23973,23982).

Hypnum calcicolum Ando
Hypnum densirameum Andot~ ~W-t!Hm[t ( 1987).

7-7- 7"; \ 1 '::f7

Hypnum cupressiforme Hedw.

/\1l::: ; ·\' :f7t~~W-t@:!l'm[t ( 1987).

7 7-!f-J\ 1 ::f7 MD (1996).
l::: 7J\1 ::f7 ll!iu, :'fi..t. C (23385), G (23658), I

Hypnum erectiusculum Sull. & Lesq.

(23884), 1 (23934).
HypnumfaurieiCardot

J\1 ::J"7 t~~l!f-t!Hm[t(1987).
Hypnum /indbergii Mitt. .I. 'fJ\ 1 ::J"7 :'fi..t. G (23691).
Hypnum oldhamii (Mitt.) A.1aeger & Sauerb. l::: /- J\ 1 ::J"7 t~~W-t!Hm[t (1987).
Hypnum plumaeforme Wi1son var. plumaeforme J \ 1 :::1·· 7 ff'ij *, ll!i u, :'fi ..t. F (23591 ,::J? /

23624, 24249, 24261, 24268, 24440), G (23629, 23632, 23674, 23678 , 23692, 24292, 24307,
24332, 23794), I (23826, 23873, 23895, 2444~ 24450), 1 (7131 , 713~ 23916, 23927, 23933~ K(24011).
Hypnum plumaeforme var. minus Ando ::J J\ 1 :::1··7 ~tfl':..t. I (24457 + Dolichomitriopsis
diversiformis) .
Hypnum sakuraii (Sakurai) An do:t :t "":::... J \ 1 :::1·· 7

ll!i u, :'fi ..t . G (23690, 24323 ,

24338).
Hypnum subimponens Lesq . subsp. ulophyllum (Mii1l.Hal.) An do 1'? J \ 1 ::J"7 :'fi ..t. G(24311).
Hypnum tristo-viride (Broth.) Paris 1
J\ 1 :::1·· 7
ff'ij *, !lii ..t. E (23534, 23535,
24198), G (23682), H (23797), I (23831), 1 (7120, 23913 , 23997).
Isopterygiopsis mue/leriana (Schimp.) Z.Iwats. .:r. 'f / l::: 7 / ;t;' ::J"7 nmD (1996).
Platygy rium repens (Brid.) Bruch & Schimp. 1 7. -IT-t -'!" :::1··7 ff'~*..t. E (23570).
Pseudotaxiphy llum densum (Cardot) Z.lwats. l::: -'J"; \ 1 7- 1 :::1·· 7 :'fi Ji 1lh ( 1981 , as Isoptery-rugium densum).
Pseudotaxiphy/lum pohliaecarpum (Sull. & Lesq .) Z.Iwats.

(23627), I (23817, 23893), 1(23942).
Py laisia brotheri Besch. 3f- 7. ::J"7 t~~!Jf-t@:!tm[t (1987).
Rhytidiadelphus japonicus (Reimers) T.J.Kop.
::J 7 -IT ::J" 7(24 194), G (23704), J (23926).
Rhytidiadelphus triquetrus (Hedw.) Warnst.
Taxiphy/lum aomoriense (Besch.) Z.lwats.

7 iJ 1 7-1 ::J"7±, :'fi..t . G

C (23361, 23408, 24093), E:t :t 7 -IT ::J"7 t~ ~W- t@:!l'm[t ( 1987).
7 :t .:C 1) -IT T -'!" ::J"7 ~t fl': , :'fi ..t. A (23355),

C (23384, 24098), D (7174, 23457, 24148), E (24187), G (23695), H (23742), J (7167, 23985).
Taxiphy llum cuspidifolium (Cardot) Z.lwats.
r iJ" 1) ; \ 1 7- 1 :::1" 7 !lii u, :'fi ..t. H(23 728), 1 (7143 , 23988).
Taxiphyllum pilosum (Broth. & Yasuda) Z.lwats. 7 -IT T -'!" :::1··7 t~ ~W-t@:!l'm[t ( 1987).
Taxiphy llum taxirameum (Mitt.) M.F1eisch. 3f- 1' 7 J \ :::1·· 7~t f1':, ± , ll!i :0, :'fi ..t. B(24132), D (7178), E (24221 ), G (23651 ), H (23725, 23813), I (2384 7).
Vesiculariafiaccida (Sull. & Lesq.) Z.lwats . l::: T-IT T -'!" :::1··7 ±..t. C (24086).

77

Hylocomiaceae l:: :3 7 :::j. 7-H

Hy locomiastrum pyrenaicum (Spruce) Broth.

~ -'V? 1) :2 7 r:· ::f-7

MiD ( 1996).

Hylocomium splendens (Hedw.) Schimp. 1 r; Jf v ::I -7 tlil ~ ~trnm at ( 1987).
Loeskeobryum cavifolium (Sande Lac .) M.Fleisch . ex Broth. 7 r ') :2 7 r:· ::l-7
1:.. G (24341 ), I (23840, 23844, 23871 , 24422).

:f.lt fi.u*

.z- 0) Ji'iJ :ill ±t!!.:t9X ~.:::, ~[ill 0) k:a t -:. tL i -c· 0) ~2 f.iH·-@- b -t±- 180 fi 0) ~lHH.ff-'V/"'/ ;t, / :7"-ltf47J' G ~ 3 :7 ::I"-7 N i -c- ~ ?;-tr) 0)5J';fjJ7Jr~t~2 ~ tLt-=. -:. tL G ~.:::
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1992. Authors of Plant Names. 732 pp. Royal Botanic Gardens,

1987. tll\ ll1H'l\-til!~m~.t . 481 pp. tffii~M\Hi!i!W~Hi~!v~~~ . t~i1l;.

tilllD.ii1§ , 1995. JliiJJi:\':l>~tltlift.t3 J: u·.z-O)Ji'iJiU:Iili:Ji&O)~H.fl. @:I JL: N "F:t~!WiisWt% 28 73- 82.
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lwatsuki, Z., 1991. Catalog of the Mosses of Japan. 182 pp. Hattori Botanical Laboratory, Nichinan .
..t;fHf · 'ti.'!Jij._;}( · l±lDfili:~lj · tffi D.!E f~ · fti,~ Jil = E1~ · 'fi.FJ'!ff;t!lJJ · *D t~'i: · ~tJilf<\1'1: · illHtl[-!H · ftc!Jiyilf* · llliiflf= ·~HIM, 1979. B ::<fs:~$s'J:Jict~m 7@l *~t.::.t3o '·ot:~~tLt.:Jm007<JliJ
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J~ :IJ"~ IT El ?~ f~ ~l!Uil] !ff 1£ g::_ ~i!J ~!1l Ui ) , 2ooo. cJ ilT · B :::$: 0) 1.-@. ii!X 0) .t3 f" tL 0) if:> J.> ~T-1£ l£ ~- v ·:; r-··-r· - ?' 7· ·:; 7- 9 fil!tiiJ 11 ( ff.ffi ~'~Hi1!~ml-:L!7l- l. 429 pp. El ~J;i~H / 7-, Jf[JK .
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7dH'£ · * DtL~'i: · +illi ft~ , 1987. Ill !Mis£Jl · l~ 'J:Ji I. l!F.~l.lfl-ilt!Mi£J!EI~!f<. t'f.~~t@!fm ~t.cJ aT!:!Hi#;~~~ (*i l , i!f.~~t@~iiJ~t.cJilT~. pp.395-415. t'f.~~Noguchi, A. , 1991. Illustrated Moss Flora of Japan. 4: 743- 1012. Hattori Botanical Laboratory, Nichinan.
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