Bull. Natn. Sci. Mus., Tokyo, Ser. B, 32(2), pp. 67–83, June 22, 2006

Mosses of Mts. Yatsugatake, Central Japan
Masanobu Higuchi1 and Tomotsugu Arikawa2
1

Department of Botany, National Science Museum,
4–1–1 Amakubo, Tsukuba, Ibaraki 305–0005 Japan
E-mail: higuchi@kahaku.go.jp
2
Department of Biology, Keio University,
4–1–1 Hiyoshi, Kohoku-ku, Yokohama, Kanagawa 223–8521 Japan
E-mail: arikawa@hc.cc.keio.ac.jp

Abstract The moss flora of Mts. Yatsugatake, central Japan, was investigated in 2003 and 2004.
The moss specimens collected by Higuchi in 1985 were also examined. The mosses recognizedcomprise 34 families, 97 genera and 174 species. For each species, substrate, locality, specimen
number and distribution were given. In total the moss flora of Mts. Yatsugatake consists of 291
species in 135 genera and 39 families, which is characterized by the richness of rare and/or endangered species as well as species number.
Key words : bryophytes, mosses, flora, Mts. Yatsugatake, Japan.

This study deals with the moss flora of Mts.
Yatsugatake, central Japan, based on the collections made under a research program, “Taxonomic research of the flora in areas with remarkablebiodiversity in Japan,” of the National Science
Museum, Tokyo. Although the outline of thebryophyte flora in Japan is becoming clear (cf.
Iwatsuki 2004, Yamada & Iwatsuki 2006), there
has never been enough knowledge on the local
flora. The lack of the information on bryophytes
in local areas in Japan has become a bottlenecktowards the development of an effective program
to protect and conserve the local flora and vegetation. In 2002 and 2003 we made a field research and collected bryophytes mainly from thesummit area of Mts. Yatsugatake. During the research in 2003, we found a rare and interesting
species, Takakia lepidozioides S. Hatt. & Inoue,
in the mountains for the first time (Higuchi &Arikawa 2005).
The Yatsugatake Mountains are located in central Honshu, Japan, and stretch ca. 20 km northand south, and consist of ten mountains higer
than 2,500 m above sea level (Fig. 1). The highest
peak is Mt. Akadake (35°58N, 138°22E) at
2,889 m above sea level. The mountains are of
volcanic origin and located in the so-called Fossa

magna region. Three vertical vegetation zones
are distinguished on the mountains: (1) the montane deciduous forest zone ranging below 1800 m
alt. where develops secondary forest and Fagus
crenata forest, (2) the subalpine coniferous forest
zone ranging 1800–2500 m alt. where develops
Tsuga deiversifolia forest lower and Abies
veitchii and A. mariesii forest upper and (3) the
alpine zone ranging above 2500 m alt. where develops Pinus pumila scrub (cf. Furuki & Inoue1984).
For the bryophyte flora of Mts. Yatsugatake,the hepatic flora was investigated and reported by
Furuki & Inoue (1984). There is no extensive
study of the moss flora of the mountains exceptthat of Kiguchi et al. (2006) who reported preliminary lists of the moss flora. As far as weknow, the first record of mosses from Mts. Yatsugatake was made by Okamura (1917) who described Calliergon perdecurrens Broth. ex S.
Okam. (Hygrohypnum ochraceum (Wilson)
Loeske). The taxa newly described from themountains are as follows: Dicranella ditrichoides
Broth. (1921), Dicranum recurvicaule Sakurai(1952a) (Dicranum nipponense Besch.),Entodon rubrissimus Sakurai (1953) (Entodon
flavescens (Hook.) A.Jaeger), Kiaeria blyttii

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Masanobu Higuchi and Tomotsugu Arikawa

Fig. 1. Map showing the area investigated. I: Northern area (north of Mugikusa Pass). II: Central area (between
Mugikusa Pass and Natsuzawa Pass). III: Southern area (south of Natsuzawa Pass). Solid line indicates
2,000 m above sea level.

Mosses of Mts. Yatsugatake(Schimp.) Broth. var. secundifolia Sakurai(1952b) (Dicranum muehlenbeckii Bruch &Schimp.), Kiaeria falcata (Hedw.) I.Hagen var.
serratifolia Sakurai (1952b), Orthotrichum
takakii Sakurai (1952c) (Arctoa fulvella(Dicks.) Bruch & Schimp. var. longisetacea(Cardot) Iisiba), Plagiobryum japonicum Nog.
(1952) and Pohlia yanoi H.Ochi (1955) (Pohlia
nutans (Hedw.) Lindb.).
The purpose of this study is (1) to investigate
the moss flora of Mt. Yatsugatake, (2) to compile
it based on the specimens collected and the literature reported and (3) to make clear the characteristics of the moss flora of the mountains.
Materials and Methods
Field studies were carried out in 2003 and2004 mainly above the subalpine coniferous forest zone, and a total of ca. 800 bryophyte specimens were collected. We also examined about
380 spcimens collected by the first author from
Mts. Yatsugatake in 1985 and two specimens collected by Dr. Furuki in 1981, which are preserved in the herbarium of Hiroshima University(HIRO). Among them about 850 specimens weremosses.
The main sites investigated are divided into
three areas as follows (Fig. 1).
I: Northern area (north of Mugikusa Pass).
August 21, 1985: Mugikusa Pass – Mt. Shimagare – Nanatsuike – Amaike – Mugikusa Pass,
2,150–2,400 m alt., Higuchi 12392–12527.
September 30, 2004: Tsuboniwa – Mt. Yokodake (Mt. Kitayokodake), 2,260–2,360 m alt.,
Higuchi 42818–42830, Arikawa 4461–4474.
October 1, 2004: Nanatsuike Pond – Mt. Yokodake – Mt. Odake – Futagoike Pond,
2,160–
2,410 m alt., Higuchi 42831–42915, Arikawa
4475–4557.
October 2, 2004: Futagoike Pond – Mt. Futagoyama – Kikkoike Pond – Futagoike Pond, 1,970–
2,250 m alt., Higuchi 42916–43021, Arikawa
4558–4662.
October 3, 2004: Kikkoike, 2,030 m alt.,
Higuchi 43022–43031, Arikawa 4663–4670.

69

II: Central area (between Mugikusa Pass and
Natsuzawa Pass).
August 20, 1985: Natsuzawa Pass – Mt.
Neishidake – Mt. Tengudake – Nakayama Pass –Shirakomaike, 2,150–2,600 m alt., Higuchi
12275–12391.
October 18, 2004: Shibunoyu – Nakayama
Pass, 1,860–2,240 m alt., Higuchi 43032–43043.
October 19, 2004: Nakayama Pass – Mt. Tengudake – Mt. Neishidake – Natsuzawa Pass,
2,450–2,600 m alt., Higuchi 43044–43084.
October 20, 2004: Nakayama Pass, 2,380–
2,400 m alt., Higuchi 43095–43149.
October 21, 2004: Nakayama Pass – Shibunoyu, 2,000–2,370 m alt., Higuchi 43150–43161.
III: Southern area (south of Natsuzawa Pass).
August 18, 1985: Minoto – Mt. Akadake,
1,700–2,890 m alt., Higuchi 12148–12186.
August 19, 1985: Mt. Akadake – Mt. Ioudake,
2,600–2,800 m alt., Higuchi 12187–12274.
October 14, 2003: Minoto – Akadake-kosen,
1,500–1,700 m alt., Higuchi 41449–41450.
October 15, 2003: Akadake-kosen – Daidoshin, 2,260–2,600 m alt., Higuchi 41451–
41524.
October 16, 2003: Akadake-kosen – Mt. Yokodake, 2,300–2,750 m alt., Higuchi 41525–41577.
October 17, 2003: Akadake-kosen – Gyojagoya, 2,250–2,350 m alt., Higuchi 41578–
41617.
October,18, 2003: Akadake-kosen – Minoto,
1500–1850 m alt., Higuchi 41618–41620.
Results and Discussion
The mosses recognized in this study comprise
34 families, 97 genera and 174 species. Recently
Kiguchi et al. (2006) reported 254 species in 125
genera and 37 families of mosses from Mts. Yatsugatake. Consequently the moss flora of Mts.
Yatsugatae comprises 291 species in 135 generaand 39 families.

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Masanobu Higuchi and Tomotsugu Arikawa

1. Comparison of the moss floras between
Mts. Yatsugatake and other mountainous regions in JapanTo make clear the characteristics of the moss
flora of Mts. Yatsugatake, it was compared withthose of other mountaineous regions in Japan,such as (1) Mts. Hakkoda (Deguchi 1981), (2)Mt. Hakusan (Akiyama 1984), (3) Mt. Ontake(Ando & Sasaki 1958, Noguchi 1958) and (4)Mt. Ishizuchi (Higuchi 2000). Although theSouthern Japan Alps and Mt. Tsurugizan haverich moss floras (cf. Takaki et al. 1970, Deguchi1982), those were not considered here because of
their geology including limestone areas. In Mts.
Hakkoda the highest peak is Mt. Odake(40°39N, 140°52E) at 1,584 m above sea level.
In Mt. Hakusan the highest peak is Mt. Gozengamine (36°9N, 136°46E) at 2,702 m above sea
level. In Mt. Ontake the highest peak is Mt.
Kengamine (35°53N, 137°28E) at 3,067 m
above sea level. In Mt. Ishizuchi the highest peakis Mt. Tengudake (33°46N, 133°6E) at 1,982 m
above sea level.
Comparison of the moss floras between Mts.
Yatsugatake and above regions indicates that themountains have comparatively rich and uniquemoss flora. Although the location and the landarea are different among them, the moss flora ofMts. Yatsugatake was the richest. Of four regions
Mt. Ontake was the most similar for the species
composition. There were about 150 species common to both regions. This similarity may be reasonable because of the location and the altitudeof the two regions.
There are 78 species present in Mts. Yatsugatake but not in other four reigions, which was
the highest number (e.g. the next is 45 species in
Mts. Hakkoda). This 78 species include several
elements of distribution. They are classified
roughly into the following floristic groups: circumboreal species (33 spp., 42%), East Asian
species including Sino-Japanese floristic element(20 spp., 26%), species endemic to Japan (10spp., 13%), worldwide species (7 spp., 9%), or
others (8 spp., 10%). The circumboreal species,
widely occurring in the Northern Hemisphere,

are represented by Brachydontium trichodes,
Myurella julacea, Plagiopus oederianus, etc. The
East Asian species include Didymodon constrictus, Mielichhoferia sasaokae, etc. The species
endemictoJapan
includeDicranella
ditrichoides, Diphyscium satoi, Plagiobryum
japonicum, etc. The worldwide species, occurring widely without relation to any continent orvegetation types, are represented by Bartramia
halleriana, Bryoerythrophyllum recurvirostre,etc.
2. Noteworthy species
Suzuki & Iwatsuki (2003) described Seligeria
pseudodonniana Tad.Suzuki & Z.Iwats. based on
the specimens from “Mts. Yatsu, Mt. Iwô-dake.”
This species is similar to Seligeria donniana(Sw.) Müll.Hal. in the absence of peristome
teeth, but is distinguished from the latter by having differentiated annulus and large, thin-walled
exothecial cells. Furuki & Higuchi (1983) reported Seligeria donniana (Sw.) Müll.Hal. based on
the specimen (Furuki 2448) from Nakayama Passof Mts. Yatsugatake. We examined the specimen
and confirmed that it was not S. donniana, but S.
pseudodonniana. This species is endemic to Mts.
Yatsugatake.
Among the species recorded in Mts. Yatsugatake, seventeen are species listed as endangered
bryophytes in Japan (cf. Environmental Agencyof Japan 2000), which are Andreaea nivalis(CREN), Aongstroemia julacea (VU), Brachydontium trichodes (CREN), Campylopus fragilis (VU), Helodium sachalinense (CREN),
Mielichhoferia sasaokae (CREN), Myurella julacea (CREN), Myurella sibirica (CREN),
Myurella tenerrima (CREN), Oedipodium griffithianum (CREN), Oreas martiana (VU), Plagiomnium tezukae (CREN), Plagiothecium
neckeroideum (CREN), Rhodobryum ontariense (VU), Schistostega pennata (CREN),
Takakia lepidozioides (CREN) and Tortulasinensis (CREN).
3. Enumeration of species
In the following enumeration species recorded

Mosses of Mts. Yatsugatakefrom Mts. Yatsugatake are listed. The families,
genera and species are arranged alphabetically.
Each species is referred by collecting sites(I–III), substrates and collector’s numbers of
Higuchi (abbreviated as mh) and Arikawa (as ta).
The complete set of the specimens are preservedin the herbarium of the Department of Botany,
National Science Museum (TNS). An asterisk (*)
preceding a species indicates a species to be
added to the list of the mosses of Mts. Yatsugatake by Kiguchi et al. (2006).

7142856, 42913; on boulder, mh-42927. II: on
rock, mh-12364; on boulder, mh-43111,
43112; on fallen log, mh-43157. III: on rock,mh-12187; on rock-cliff, mh-41531; on boulder, mh-41470.
Aulacomniaceae
Aulacomnium heterostichum (Hedw.) Bruch &Schimp. Kiguchi et al. (2006).
Bartramiaceae

Amblystegiaceae
Campyliadelphus chrysophyllus (Brid.) R.S.
Chopra Kiguchi et al. (2006).
Campylium hispidulum (Brid.) Mitt. I: on treetrunk, mh-43026.
Campylium squarrosulum (Broth. & Cardot)
Kanda Kiguchi et al. (2006).
*Cratoneuron filicinum (Hedw.) Spruce III: on
soil, mh-12247.
Drepanocladus fluitans (Hedw.) Warnst. I: on
soil, mh-12461, 12462, 12477, 12521, 42837(Polytrichum commune); on humus, mh12522, 42854. III: on humus, mh-12164.
Hygrohypnum ochraceum (Wilson) Loeske
Okamura (1917), Kiguchi et al. (2006).
Notes. Okamura (1917) described Calliergon
perdecurrens Broth. ex S.Okam. from “Prov. Shinano: Mt. Yatsugadake.” Later Kanda (1976) reduced it to a synonym of Hygrohypnum
ochraceum.
Sanionia uncinata (Hedw.) Loeske I: on treetrunk, mh-12438; on decaying log, mh-42957(Buxbaumia aphylla), ta-4466; on humus,
mh-42944, ta-4574, 4654. II: on decaying
log, mh-12379. III: on rock, mh-12176; onhumus, mh-12154, 12205, 12273, 12280.
Andreaeaceae
Andreaea nivalis Hook. II: on boulder, mh12371.
Andreaea rupestris Hedw. var. fauriei (Besch.)
Takaki I: on rock, ta-4481; on rock-cliff, mh-

Bartramia halleriana Hedw. Kiguchi et al.
(2006).
Bartramia ithyphylla Brid. III: on rock-cliff,
mh-41499 (Anomobryum filiforme, Encalypta ciliata); on soil, mh-12236 (Encalypta ciliata), 12237 (Thuidium cymbifolium).
Bartramia pomiformis Hedw. I: on boulder, ta4660; on soil, mh-42907; on humus, mh42820, ta-4524; on decorticated log, ta-4465.
II: on rock-cliff, mh-43034, 43057. III: on
rock-cliff, mh-12268.
Philonotis fontana (Hedw.) Brid. III: on rockcliff, mh-12174; on humus, mh-12232.
Plagiopus oederianus (Sw.) H.A.Crum &L.E. Anderson III: on rock-cliff, mh-41511(Myurella julacea, Thuidium cymbifolium).
Brachytheciaceae
Brachythecium brotheri Par. I: on boulder,
mh-42937; on humus, mh-42878.
*Brachythecium buchananii (Hook.) A.Jaeger
II: on humus, mh-43059 (Brachythecium reflexum), 43127.
Brachythecium coreanum Cardot Kiguchi etal. (2006).
Brachythecium helminthocladum Broth. &Paris Kiguchi et al. (2006).
Brachythecium plumosum (Hedw.) Schimp. I:
on boulder, mh-42928, 42984. II: on rockcliff, mh-43102; on boulder, mh-43063,
43094; on stone-works, mh-43132.
Brachythecium populeum (Hedw.) Bruch &72

Masanobu Higuchi and Tomotsugu Arikawa

Schimp. Kiguchi et al. (2006).
*Brachythecium reflexum (Starke) Schimp. I: on
boulder, mh-42950, ta-4579; on humus, mh42922 (Hypnum subimponens subsp. ulophyllum), 42951, 42982, ta-4572. II: on boulder, mh43061, 43122; on humus, mh-43059 (Brachythecium buchananii), 43124. III: on boulder,mh-41451.
Brachythecium salebrosum (F.Weber & Mohr.)
Bruch & Schimp. Kiguchi et al. (2006).
*Brachythecium uyematsui Broth. ex Cardot
III: on rock-cliff, mh-12231, 41497, 41518,
41562.
Brachythecium uncinifolium Broth. & Paris
III: on rock-cliff, mh-41615 (Mnium heterophyllum); on boulder, mh-41608.
Bryhnia hultenii Bartr. ex Grout Kiguchi et al.
(2006).
Bryhnia novae-angliae (Sull. & Lesq. ex Sull.)Grout Kiguchi et al. (2006).
Cirriphyllum cirrosum (Schwägr.) Grout III:on rock-cliff, mh-41550, 41553; on soil, mh12221 (Bryum paradoxum), 12222.
*Rhynchostegium pallidifolium (Mitt.) A.Jaeger
II: on humus, mh-43126.
Rhynchostegium riparioides (Hedw.) Cardot
Kiguchi et al. (2006).
Bryaceae
Anomobryum filiforme (Griff.) A.Jaeger III:on rock-cliff, mh-12258, 41499 (Bartramia
ithyphylla, Encalypta ciliata).
Brachymenium nepalense Hook. Kiguchi etal. (2006).
*Bryum caespiticium L. ex Hedw. III: on rockcliff, mh-41502.
Bryum capillare Hedw. Kiguchi et al. (2006).
Bryum pallens (Brid.) Sw. ex Roehl. Kiguchi
et al. (2006).
*Bryum paradoxum Schwägr. III: on soil, mh12202, 12221 (Cirriphyllum cirrosum),
12254, 12262.
Mielichhoferia sasaokae Broth. Kiguchi et al.
(2006).
Plagiobryum japonicum Nog. II: on rock-cliff,

mh-43091. III: on rock, mh-12286; on rockcliff, mh-41532.
Notes. Noguchi (1952) described Plagiobryum
japonicum Nog. from “Honsyu: Mt. Yatsugatake (Iwôdake ca. 2700 m).”Plagiobryum zierii (Hedw.) Lindb. Kiguchi etal. (2006).
Pohlia cruda (Hedw.) Lindb. I: on boulder, ta4644.
Pohlia elongata Hedw. I: on soil, ta-4610.
Pohlia longicollis (Hedw.) Lindb. I: on rockcliff, mh-42827, 42883, 42898; on soil, mh12395, 42979 (Dicranella heteromalla); on
decorticated log, ta-4462. II: on soil, mh12370; on root, mh-43095. III: on rock, mh12192; on rock-crevice, mh-12283, 41492,
41561; on soil, mh-12319, 12322; on decaying
log, mh-12155; on humus, mh-12326.
Pohlia nutans (Hedw.) Lindb. I: on rock-cliff,
mh-42911 (Bartramiopsis lescurii, Takakia
lepidozioides); on soil, mh-42923. II: on
rock-cliff, mh-43037; on rock-crevice, mh43056; on soil, mh-43128; on humus, mh43123.
Notes. Ochi (1955) described Pohlia yanoi
H.Ochi from “Prov. Shinano: Mt. YatsugatakeYokodake, alt. ca. 2,700 m,” and later he (1959)
reduced it to a synonym of Pohlia schimperi(Müll.Hal.) A.L.Andrews which is now treated asa synonym of Pohlia nutans (cf. Iwatsuki 2004).
*Pohlia proligera (Kindb.) Lindb. ex Arn. I: on
soil, mh-42921, 42954.
*Pohlia revolvens (Cardot) Nog. II: on rockcliff, mh-12332, 43069; on boulder, mh-43049(Pogonatum sphaerothecium). III: on rockcliff, mh-41563.
Pohlia wahlenbergii (F.Weber & Mohr) A.L.Andrews III: on rock-cliff, mh-41547.
Rhodobryum ontariense (Kindb.) Kindb.
Kiguchi et al. (2006).
Rhodobryum roseum (Hedw.) Limpr. II: on
humus, mh-43079.
Bryoxiphiaceae
Bryoxiphium norvegicum (Brid.) Mitt. subsp.

Mosses of Mts. Yatsugatakejaponicum (Berggr.) A.Löve & D.Löve I: on
rock-cliff, mh-42899, ta-4544; on boulder, mh42992, ta-4618. II: on rock-cliff, mh-43050,
43081. III: on rock-cliff, mh-12149, 41503.
Buxbaumiaceae
Buxbaumia aphylla Hedw. I: on soil, mh42861, 42941, 42970, ta-4477, 4510, 4602; ondecaying log, mh-42957 (Sanionia uncinata); on rotten log, mh-42826, ta-4586. II: on
soil, mh-43131. III: on soil, mh-12301.
Buxbaumia minakatae Okam. I: on humus,
mh-42980; on rotten log, ta-4630.
Diphyscium foliosum (Hedw.) Mohr I: on soil,
mh-42865, ta-4622; on boulder, ta-4623. III:on rock-cliff, mh-41605.
Diphyscium satoi Tuzibe Kiguchi et al. (2006).
Climaciaceae
Climacium dendroides (Hedw.) F.Weber & Mohr
III: on soil, mh-41519, 41555; on humus, mh12229, 12230.
Climacium japonicum Lindb. Kiguchi et al.
(2006).
Dicranaceae
Aongstroemia julacea (Hook.) Mitt. III: on
rock-cliff, mh-12200.
Arctoa fulvella (Dicks.) Bruch & Schimp.
var. fulvella II: on boulder, mh-12366(Racomitrium nitidulum), 43134. III: on
rock, mh-12285; on rock-cliff, mh-12175,
12194 (Grimmia apiculata, Pogonatum
sphaerothecium), 12195 (Grimmia longirostris).
*Arctoa fulvella (Dicks.) Bruch & Schimp. var.
longisetacea (Cardot) Iisiba Sakurai (1952c).
Notes. When Sakurai (1952c) described Orthotrichum takakii Sakurai, he cited a specimen
from “Prov. Shinano, mt. Yatsugatke, Yokodake,
2800 m) as a syntype. Later Noguchi (1968) reduced it to a synonym of Arctoa fulvella (Dicks.)
Bruch & Schimp. var. longisetacea (Cardot) Iisi-

73

ba.
Campylopus gracilis (Mitt.) A.Jaeger Kiguchi
et al. (2006).
Campylopus japonicus Broth. Kiguchi et al.
(2006).
Cynodontium fallax Limpr. Kiguchi et al.
(2006).
Cynodontium gracilescens (F.Weber & Mohr)
Schimp. II: on boulder, mh-43148; on
humus, mh-43064.
Cynodontium polycarpum (Hedw.) Schimp.
III: on boulder, mh-12288.
Cynodontium strumiferum (Hedw.) Lindb.
Kiguchi et al. (2006).
Cynodontium tenellum (Schimp.) Limpr. Narita (1950), Sakurai (1953).
Dicranella
ditrichoides
Broth. Brotherus(1921), Kiguchi et al. (2006).
Dicranella heteromalla (Hedw.) Schimp. I: on
rock-cliff, mh-42892; on boulder, mh-42825,
42906 (Takakia lepidozioides); on soil, mh12437 (Bartramiopsis lescurii), 42979(Pohlia longicollis); on decorticated log, ta4463; on rotten stump, ta-4650. II: on soil,
mh-43036, 43150; on root, mh-43153(Tetraphis pellucida). III: on rock-cliff,
mh-41458(Distichium
capillaceum,
Ditrichum divaricatum), 41459 (Distichium
capillaceum); on soil, mh-12310, 41461; onroot, mh-43153 (Tetraphis pellucida).
Dicranella subulata (Hedw.) Schimp. II: on
boulder, mh-43055.
*Dicranodontium asperulum (Mitt.) Broth. I:
on rock-cliff, mh-42903; on humus, mh42877. II: on soil, mh-43145.
Notes. This species is distinguished fromJapanese species of Dicranodontium by having
flexuouse laeaves, lax areolation at sheathingbase of leaves and serrulate margins of sheathingbase of leaves.
Dicranodontium denudatum (Brid.) E.G.Britton
ex Williams Kiguchi et al. (2006).
Dicranoloma cylindrothecium (Mitt.) Sakurai
Kiguchi et al. (2006).
Dicranoweisia crispula (Hedw.) Lindb. ex Milde
Kiguchi et al. (2006).

74

Masanobu Higuchi and Tomotsugu Arikawa

Dicranum flagellare Hedw. Kiguchi et al.
(2006).
Dicranum fulvum Hook. Kiguchi et al. (2006).
Dicranum fuscescens Turner III: on humus,
mh-12302, 12325.
Dicranum groenlandicum Brid. Kiguchi et al.
(2006).
Dicranum hamulosum Mitt. I: on tree-trunk,mh-12417, 42840, 42931, ta-4568, 4593. II:on boulder, mh-43070; on rotten log, mh43115.
Dicranum japonicum Mitt. I: on rock-cliff,
mh-42889; on boulder, mh-42952; on humus,
mh-12515, 42948, 42995. II: on humus, mh43035.
Dicranum leiodontum Cardot II: on humus,
mh-43086. III: on rock-crevice, mh-41574;
on humus, mh-12216, 12278.
Dicranum majus Turner I: on humus, mh12409, 12480, 42846, 42871, 42908 (Hylocomium splendens), ta-4513. II: on humus,
mh-12290, 43066 (Rhytidiadelphus triquetrus), 43121. III: on humus, mh-12220,
41473, 41598.
Dicranum mayrii Broth. III: on tree-trunk, mh12321.
*Dicranum muehlenbeckii Bruch & Schimp.
I: on tree-trunk, mh-12413, 42917, 43002.
II: on decaying log, mh-12345.
Notes. Sakurai (1952) described Kiaeria
blyttii (Schimp.) Broth. var. secundifolia Sakurai
from “Prov. Kai, mt. Yatsugatake, 2800 m,”
which is treated as a synonym of Dicranum
muehlenbeckii Bruch & Schimp. (cf. Iwatsuki
2004)*Dicranum nipponense Besch. Sakurai (1952).
Notes. Sakurai (1952) described Dicranum
recurvicaule Sakurai from “Prov. Shinano, mt.
Yatsugatake,” which was reduced to a synonymof Dicranum nipponense Besch. by Takaki(1964).
Dicranum polysetum Sw. Narita (1950, as Dicranum bergeri Blandow), Kiguchi et al.
(2006).
Dicranum setifolium Cardot Kiguchi et al.
(2006).

Dicranum viride (Sull. & Lesq.) Lindb. var.
hakkodense (Cardot) Takaki I: on tree-trunk,mh-12432, 12527, 42841, 42958, ta-4578; on
decaying log, mh-12448. II: on tree-trunk,mh-12296, 12324, 12346, 43140.
Kiaeria falcata (Hedw.) I.Hagen var. serratifolia
Sakurai Sakurai (1952).
Notes. Sakurai (1952) described Kiaeria
blyttii (Schimp.) Broth. var. serratifolia Sakurai
from “Prov. Kai, mt. Yatsugatake, 2500 m.”
Oncophorus wahlenbergii Brid. var. wahlenbergii I: on boulder, mh-43000, ta-4629.
II: on rock-cliff, mh-43101, 43104; on soil,
mh-12374. III: on tree-trunk, mh-41475; on
decaying log, mh-12156.
Oncophorus wahlenbergii var. perbrevipes
Deguchi & H.Suzuki Kiguchi et al. (2006).
Oreas martiana (Hoppe & Hornsch. ex Hornsch.) Brid. Kiguchi et al. (2006).
Oreoweisia laxifolia (Hook.) Kindb. Kiguchi etal. (2006).
Paraleucobryum longifolium (Hedw.) Loeske
Kiguchi et al. (2006).
Rhabdoweisia crispata (With.) Lindb. I: on
rock-cliff, mh-42828, 42901; on rock-crevice,
mh-12452; on boulder, mh-42993. II: on
rock-cliff, mh-43041; on soil, mh-12361,
43144; on root, mh-43082. III: on rock-cliff,
mh-41534; on soil, mh-41478.
Ditrichaceae
Ceratodon purpureus (Hedw.) Brid. II: on soil,
mh-12317.
Distichium capillaceum (Hedw.) Bruch &Schimp. III: on rock-cliff, mh-12217, 12256,
41458 (Dicranella heteromalla, Ditrichum
divaricatum), 41459 (Dicranella heteromalla), 41491, 41513, 41514, 41560, 41569.
*Ditrichum crispatissimum (Müll.Hal.) Paris
III: on rock-crevice, mh-12250.
Ditrichum divaricatum Mitt. I: on soil, mh42942, ta-4509; on boulder, ta-4546. II: on
soil, mh-12375. III: on rock-cliff, mh-41458(Dicranella heteromalla, Distichium capillaceum).

Mosses of Mts. Yatsugatake

Ditrichum heteromallum (Hedw.) E.G.Britton
I: on boulder, mh-12441, 42819, 42823; onsoil, mh-42939. II: on soil, mh-12337,
12353, 12357, 12385. III: on rock-crevice,
mh-12190.
Ditrichum macrorhynchum Broth. ex Cardot
Kiguchi et al. (2006).
*Ditrichum pallidum (Hedw.) Hampe I: on
soil, mh-42864. II: on soil, mh-43054,
43139. III: on rock-crevice, mh-12152,
12173, 41471; on soil, mh-12279, 41453.
Ditrichum rhynchostegium Kindb. Kiguchi etal. (2006).
*Saelania glaucescens (Hedw.) Broth. III: on
soil, mh-41449 (Fissidens teysmannianus).
Encalyptaceae
Encalypta ciliata Hedw. III: on rock-cliff, mh41400 (Anomobryum filiforme, Bartramia
ithyphylla), 41485, 41505 (Bryoerythrophyllum brachystegium, Desmatodon laureri),
41507 (Bryoerythrophyllum brachystegium);on soil, mh-12236 (Bartramia ithyphylla),
12238 (Desmatodon laureri), 12243, 12245,
41508 (Tortula sinensis).
Entodontaceae
Entodon concinnus (De Not.) Paris subsp. caliginosus (Mitt.) Mizush. Kiguchi et al. (2006).
Entodon flavescens (Hook.) A.Jaeger Sakurai(1953).
Notes. Sakurai (1953) described Entodon
rubrissimus Sakurai from “Prov. Shinano,
Nobeyama (1500 m).” Later Buck (1980) reduced it to a synonym of Entodon flavescens.
Erpodiaceae
Glyphomitrium humillimum
Kiguchi et al. (2006).

(Mitt.)

CardotFissidentaceae
Fissidens bryoides Hedw.

III: on rock-cliff,75

mh-12266, 41557, 41558
Fissidens dubius P.Beauv. Kiguchi et al. (2006).
Fissidens gymnogynus Besch. Kiguchi et al.
(2006).
*Fissidens taxifolium Hedw. I: on boulder,
mh-42998.
*Fissidens teysmannianus Dozy & Molk. I: on
rock, ta-4628. III: on soil, mh-41449 (Saelania glaucescens).
Grimmiaceae
Campylostelium saxicola (F.Weber & Mohr)
Bruch & Schimp. Kiguchi et al. (2006).
*Coscinodon humilis Nog. III: on rock-cliff,
mh-12171, 12207 (Grimmia longirostris).
Grimmia apiculata Hornsch. II: on rock-cliff,
mh-43085. III: on rock-cliff, mh-12198,
12203; on rock-crevice, mh-12194 (Arctoa
fulvella, Pogonatum sphaerothecium).
Grimmia donniana Sm. III: on rock-cliff, mh41536, 41551.
Grimmia elongata Kaulf. II: on rock-crevice,
mh-43044 (Arctoa fulvella).
Grimmia incurva Schwägr. Kiguchi et al.
(2006).
Grimmia longirostris Hook. I: on boulder, mh12495. III: on rock, mh-12188, 12189,
12191, 12263; on rock-cliff, mh-12195 (Arctoa fulvella), 12207 (Coscinodon humilis),12209, 12241, 12335, 41482, 41495, 41611(Schistidium strictum); on boulder, mh41573.
Grimmia pilifera P.Beauv. Kiguchi et al. (2006).
Racomitrium anomodontoides Cardot III: on
rock, mh-12153.
Racomitrium atroviride Cardot Kiguchi et al.
(2006).
Racomitrium barbuloides Cardot I: on boulder, mh-12435, 42855, 42973, 43028; on soil,
ta-4536; on humus, mh-42983 (Plagiomnium acutum). II: on humus, mh-43136. III:on rock, mh-12265.
Racomitrium fasciculare (Hedw.) Brid. I: on
rock, ta-4484, 4485; on boulder, mh-42938,
43029, ta-4538; on tree-trunk, mh-42868; on

76

Masanobu Higuchi and Tomotsugu Arikawa

decaying log, mh-42824. II: on boulder, mh43053. III: on rock, mh-12151, 12157,
12295; on boulder, mh-41469.
Racomitrium heterostichum (Hedw.) Brid. I:
on rock, ta-4482.
Racomitrium laetum Besch. & Cardot I: on
boulder, mh-42961, ta-4545. III: on humus,
mh-12276.
Racomitrium lanuginosum (Hedw.) Brid. I: on
rock, mh-12492; on humus, mh-42869. II:on boulder, mh-43135. III: on boulder, mh41572; on humus, mh-12208.
Racomitrium muticum (Kindb.) Frisvoll
Kiguchi et al. (2006).
Racomitrium nitidulum Cardot I: on boulder,mh-12494, 42987. II: on rock-cliff, mh43087; on boulder, mh-12366 (Arctoa fulvella). III: on rock, mh-12248, 12271.
Schistidium confertum (Funck) Bruch &Schimp. Kiguchi et al. (2006).
Schistidium strictum (Turn.) Loeske ex Martensson II: on boulder, mh-43113. III: on rockcliff, mh-12170, 12251, 41490, 41506 (Amphidium lapponicum), 41556, 41610, 41611(Grimmia longirostris); on rock-crevice, mh12249.

43119, 43161. III: on humus, mh-12186,
41521, 41597.
Pleurozium schreberi (Brid.) Mitt. I: on soil,
ta-4573; on humus, mh-12411, 42860, 42943,
43005, ta-4469, 4501, 4652; on tree-trunk,mh-12428; on fallen log, mh-12518; on decaying log, mh-43021. II: on decaying log, mh12355 (Hypnum pallescens); on humus, mh12359, 43116. III: on rock-cliff, mh-41607;
on tree-trunk, mh-12320; on humus, mh12185, 12233, 41474.
Rhytidiadelphus japonicus (Reimers) T.J.Kop.
I: on humus, mh-42862, 42925. II: on
humus, mh-43117, 43118. III: on humus, mh12291.
*Rhytidiadelphus squarrosus (Hedw.) Warnst.
I: on boulder, ta-4563.
Rhytidiadelphus triquetrus (Hedw.) Warnst. I:
on boulder, mh-42960; on humus, mh-42859,
ta-4502, 4508, 4590, 4612. II: on humus,
mh-43099. III: on humus, mh-12148, 12227,
41450.
Rhytidium rugosum (Hedw.) Kindb. II: on
rock-cliff, mh-43103; on humus, mh-43142.
III: on humus, mh-12204, 41504.
Hypnaceae

Hylocomiaceae
Hylocomiastrum
himalayanum(Mitt.)
M.Fleisch. ex Broth. I: on humus, mh42836, 43016. II: on humus, mh-43138.
III: on humus, mh-41590; on decaying log,
mh-12391. III: on humus, mh-12165, 41479,
41596.
Hylocomiastrum
pyrenaicum(Spruce)
M.Fleisch. ex Broth. I: on humus, mh-42985(Helodium paludosum), 42986 (Plagiomnium acutum). II: on boulder, mh-43098. III:on humus, mh-41566.
Hylocomiastrum umbratum (Hedw.) M.Fleisch.
ex Broth. Kiguchi et al. (2006).
Hylocomium splendens (Hedw.) Schimp. I: on
boulder, mh-42991, ta-4646; on humus, mh42858, 42908 (+Dicranum majus), 42945, ta4504, 4591, 4613. II: on humus, mh-12356,

Callicladium haldanianum (Grev.) H.A.Crum
I: on tree-trunk, mh-42969 (Iwatsukiella leucotricha, Pylaisiadelpha tenuirostris); on decaying log, mh-12504, 12525; on tree-trunk,mh-42916. II: on boulder, mh-43155.
Gollania varians (Mitt.) Broth. Kiguchi et al.
(2006).
*Herzogiella turfacea (Lindb.) Z.Iwats. I: on
humus, mh-42981.
Hypnum callichroum Brid. subsp. japonicum
Ando II: on rock-cliff, mh-43042; on soil,
mh-12373, 12376. III: on boulder, mh12159.
Hypnum cupressiforme Hedw. Kiguchi et al.
(2006).
Hypnum dieckii Renauld & Cardot Kiguchi etal. (2006).
Hypnum fujiyamae (Broth.) Paris III: on root,Mosses of Mts. Yatsugatake

mh-41465.
Hypnum lindbergii Mitt. I: on boulder, mh43030, ta-4665. III: on humus, mh-12180.
Hypnum pallescens (Hedw.) P.Beauv. I: on
rock-cliff, mh-42886 (Pogonatum sphaerothecium); on tree-trunk, mh-12505, 12510,
12526, 42844, 42946, 42953, 42966, 42989,
43031, ta-4493, 4500, 4569, 4597, 4598, 4594
(Plagiothecium neckeroideum); on branch,
mh-12487, 42830; on fallen tree, mh-12425,
12426. II: on tree-trunk, mh-12380, 43040;
on branch, mh-12377, 43080; on decaying log,
mh-12355 (Pleurozium schreberi); on decaying stump, mh-12316. III: on tree-trunk, mh41480, 41585 (Ulota reptans).
Hypnum plicatulum (Lindb.) A.Jaeger & Sauerb.
I: on tree-trunk, mh-12520, 42851; on root,
mh-12496; on decaying log, mh-12445. II:on tree-trunk, mh-12342, 12351, 12354,
12381, 43097; on fallen log, mh-12347,
43107; on decaying log, mh-12343. III: on
rock-cliff, mh-12199; on boulder, mh-42936;
on tree-trunk, mh-12184, 12297; on decaying
log, mh-12160, 12300, 12313; on humus, mh43047.
Hypnum sakuraii (Sakurai) Ando Kiguchi etal. (2006).
Hypnum subimponens Lesq. subsp. ulophyllum(Müll.Hal.) Ando I: on humus, mh-42922(Brachythecium reflexum). III: on boulder,mh-12298; on humus, mh-12327.
Hypnum tristo-viride (Broth.) Paris I: on rockcliff, mh-12450; on tree-trunk, mh-12511,
42924. II: on rock-cliff, mh-43159; on boulder, mh-12360. III: on decaying log, mh41464.
*Platygyrium repens (Brid.) Bruch & Schimp.
I: on decaying log, mh-12516.
Podperaea krylovii (Podp.) Z.Iwats. & Glime
Kiguchi & Iwatsuki (2005).
Ptilium crista-castrensis (Hedw.) De Not. I: on
boulder, mh-42959; on humus, mh-42866, ta4503. II: on humus, mh-43110. III: on
humus, mh-12287, 12309, 41606.
Pylaisia intricata (Hedw.) Schimp.
Notes. Kiguchi et al. (2006) reported this77

species as Pylaisiella intricata (Hedw.) Grout.
Pylaisia polyantha (Hedw.) Schimp.
Notes. Kiguchi et al. (2006) reported thisspecies as Pylaisiella polyantha (Hedw.) Grout.
*Pylaisia stereodontoides Broth. & Yasuda ex
Iisiba I: on tree-trunk, mh-12517, 43022, ta4605, 4663. III: on tree-trunk, mh-12167(Ulota japonica), 12168, 41613, 41619(Orthotrichum sordidum).
Notes. Pylaisia stereodontoides is characterized by delicate appearance, smaller and concave
branch leaves, long-acuminate exostome teeth
and fragmentary endostome (Arikawa 2004).
This species is distributed in Japan (Honshu andHokkaido) and Korea, and its southern limit inJapan is Mt. Jizogatake in Yamanashi Prefecture.
Lembophyllaceae
Dolichomitriopsis diversiformis (Mitt.) Nog. I:
on tree-trunk, mh-42990.
Isothecium subdiversiforme Broth. Kiguchi etal. (2006).
Leskeaceae
Iwatsukiella leucotricha (Mitt.) W.R.Buck &H.A.Crum I: on tree-trunk, mh-12431,
42842, 42935, 42969 (Callicaldium haldanianum, Pylaisiadelphus tenuirostris); on decaying log, mh-42956. II: on tree-trunk, mh12314, 12389 (Ulota crispa), 43073(Rigodiadelphus robustus); on fallen log,
mh-12339, 43065. III: on tree-trunk, mh12183, 41476.
*Lescuraea incurvata (Hedw.) E.Lawton III:on rock-cliff, mh-41494.
Lescuraea mutabilis (Brid.) Lindb. ex I.Hagen
I: on tree-trunk, mh-42843.
Lescuraea saxicola (Schimp.) Milde III: on
rock-cliff, mh-12172, 41496 (Pterigynandrum filiforme).
Lindbergia brachyptera (Mitt.) Kindb. Kiguchi
et al. (2006).
Orthoamblystegium spurio-subtile (Broth. &Par.) Kanda & Nog. Kiguchi et al. (2006).

78

Masanobu Higuchi and Tomotsugu Arikawa

Pterigynandrum filiforme Hedw. III: on rockcliff, mh-41496 (Lescuraea saxicola); on
rock-crevice, mh-12264.
Rigodiadelphus robustus (Lindb.) Nog. I: on
tree-trunk, mh-42845, ta-4492, 4535; on root,
ta-4507; on branch, mh-43018; on fallen log,
mh-43014. II: on tree-trunk, mh-43073(Iwatsukiella leucotricha), 43074, 43075.
III: on tree-trunk, mh-12274, 41564, 41571;
on branch, mh-12306.
Leucodontaceae*Leucodon sapporensis Besch.
trunk, mh-43025.

I: on tree-

Mniaceae*Mnium heterophyllum (Hook.) Schwägr. III:
rock-cliff, mh-41615 (Brachythecium uncinifolium).
Mnium hornum Hedw. Kiguchi et al. (2006).
Mnium lycopodioides Schwägr. I: on treetrunk, mh-43010. III: on rock-cliff, mh41510; on soil, mh-12260.
Mnium spinosum (Voit) Schwägr. Kiguchi etal. (2006).
Plagiomnium acutum (Lindb.) T.J.Kop. I: on
boulder, ta-4614; on humus, mh-42983(Racomitrium barbuloides), 42986 (Hylocomium pyrenaicum). III: on boulder, mh41612.
Plagiomnium confertidens (Lindb. & Arnott)
T.J.Kop. Kiguchi et al. (2006).
*Plagiomnium cuspidatum (Hedw.) T.J.Kop.
II: on humus, mh-43062.
Plagiomnium medium (Bruch & Schimp.)
T.J.Kop. Kiguchi et al. (2006).
*Plagiomnium tezukae (Sakurai) T.J.Kop. II:on boulder, mh-43108.
Plagiomnium vesicatum (Besch.) T.J.Kop.
Kiguchi et al. (2006).
Pseudobryum speciosum (Mitt.) T.J.Kop. I: on
humus, mh-42821, 42893, 43003, ta-4467,
4637. II: on fallen log, mh-43106; on humus,
mh-12367, 43038, 43151. III: on humus,

mh-12307, 41523.
Rhizomnium hattorii T.J.Kop. Kiguchi et al.
(2006).
Rhizomnium magnifolium (Horik.) T.J.Kop. I:
on soil, ta-4514; on humus, mh-12410, 12473,
42839, 42863, 42971, ta-4497, 4640. II: on
humus, mh-43052, 43077. III: on rock-cliff,
mh-41600; on humus, mh-12299, 41455.
Rhizomnium nudum (E.G.Britton & R.S.
Williams) T.J.Kop. Kiguchi et al. (2006).
Rhizomnium striatulum (Mitt.) T.J.Kop. I: on
boulder, mh-12497, 43012; on decaying log,
mh-12503; on decorticated log, ta-4626; onrotten log, mh-42999. II: on decaying log,
mh-12368, 12369; on rotten log, mh-43033,
43158.
Rhizomnium tuomikoskii T.J.Kop. Kiguchi etal. (2006).
Trachycystis flagellaris (Sull. & Lesq.) Lindb.
I: on rock, ta-4651; on boulder, ta-4566, 4620,
4621; on soil, mh-42920; on decaying log, mh12499, 12500. II: on fallen log, mh-43105.
Trachycystis ussuriensis (Maack & Regel)
T.J.Kop. Kiguchi et al. (2006).
Neckeraceae
Homalia trichomanoides (Hedw.) Bruch &Schimp. var. japonica (Besch.) S.He
Kiguchi et al. (2006).
Neckera borealis Nog. Kiguchi et al. (2006).
Neckera pennata Hedw. Kiguchi et al. (2006).
Orthotrichaceae
Amphidium lapponicum (Hedw.) Schimp. II:
on rock-cliff, mh-43093. III: on rock-crevice,
mh-41554; on rock-cliff, mh-12242, 12252,
12255, 12261, 41472, 41484, 41506 (Schistidium strictum), 41546.
Amphidium mougeotii (Bruch & Schimp.)
Schimp. Kiguchi et al. (2006).
*Orthotrichum consobrinum Cardot I: on treetrunk, mh-43024.
Orthotrichum erubescens Müll.Hal. Kiguchi
et al. (2006).

Mosses of Mts. Yatsugatake

Orthotrichum sordidum Sull. & Lesq. I: on
tree-trunk, ta-4582, 4598. III: on tree-trunk,mh-41619 (Pylaisia stereodontoides).
Ulota crispa (Hedw.) Brid. I: on tree-trunk,mh-12433, 12506, 43023 (Ulota drummondii), 43027 (Ulota drummondii), ta4664. II: on tree-trunk, mh-12389 (Iwatsukiella leucotricha), 12390, 43143. III: on
tree-trunk, mh-12224, 41583 (Ulota reptans).
Ulota drummondii (Hook. & Grev. ex Grev.)
Brid. I: on tree-trunk, mh-43023 (Ulota
crispa), 43027 (Ulota crispa). III: on decaying log, mh-12293.
Ulota japonica (Sull. & Lesq.) Mitt. III: on
tree-trunk, mh-12167 (Pylaisia stereodontoides).
Ulota reptans Mitt. I: on tree-trunk, mh-42934,
42967. III: on tree-trunk, mh-12182, 12514,
41462, 41583 (Ulota crispa), 41585 (Hypnum pallescens).
Plagiotheciaceae
Isopterygiopsis muelleriana (Schimp.) Z.Iwats.
I: on rock-cliff, mh-42872; on soil, mh-42964.
II: on soil, mh-12358. III: on rock-cliff, mh12253.
*Isopterygiopsis pulchella (Hedw.) Z.Iwats.
III: on rock-crevice, mh-41549.
Plagiothecium cavifolium (Brid.) Z.Iwats. I:
on rock-cliff, mh-42882; on boulder, mh43004. II: on rock-cliff, mh-43083; onhumus, mh-43078. III: on humus, mh-12294.
Plagiothecium denticulatum (Hedw.) Schimp.
I: on rock-cliff, mh-43020. III: on old plantsof Pogonatum japonicum, mh-12334.
Plagiothecium euryphyllum (Cardot & Thér.)
Z.Iwats. I: on humus, mh-42838, 42955; ontree-trunk, ta-4512.
Plagiothecium lateum Schimp. I: on root, mh42918. III: on decaying log, mh-12318(Hypnum pallescens).
Plagiothecium neckeroideum Bruch & Schimp.
I: on humus, ta-4515, 4520, 4641; on treetrunk, ta-4594 (Hypnum pallescens). III:79

rock-cliff, mh-41530; on soil, mh-12303.
Plagiothecium nemorale (Mitt.) A.Jaeger I: on
rock, ta-4648; on boulder, mh-43008, ta-4619,
4636.
Pleuroziopsidaceae
Pleuroziopsis ruthenica (Weinm.) Kindb. ex
E.G.Britton I: on humus, mh-42867, 42894,
43017, ta-4511, 4643. II: on humus, mh12378, 43076, 43141. III: on humus, mh12161, 41589.
Polytrichaceae
Atrichum rhystophyllum (Müll.Hal.) Paris
Kiguchi et al. (2006).
Atrichum undulatum (Hedw.) P.Beauv. var. undulatum I: on soil, mh-42919.
Atrichum undulatum var. gracilisetum Besch.
Kiguchi et al. (2006).
Bartramiopsis lescurii (James) Kindb. I: on
rock, ta-4533; on boulder, ta-4537; on rockcliff, mh-42911 (Pohlia nutans, Takakia lepidozioides); on soil, mh-12430, 12437 (Dicranella heteromalla), 42977, ta-4470, 4539; on
humus, mh-42822. III: on rock-cliff, mh41594 (Pogonatum urnigerum); on soil, mh12158, 41530 (Plagiothecium neckeroideum).
Oligotrichum aligerum Mitt. I: on soil, mh12436, 12349, 42835, 42940, 42965, ta-4559.
II: on soil, mh-43129.
Oligotrichum parallelum (Mitt.) Kindb. II: on
rock-cliff, mh-43039; on soil, mh-43130. III:on soil, mh-41603.
Pogonatum alpinum (Hedw.) Röhl. II: on rockcliff, mh-43051, 43100. III: on rock-cliff,
mh-12267, 41559; on boulder, mh-41609; on
soil, mh-12239, 12289; on humus, mh-12219,
12275.
Pogonatum contortum (Brid.) Lesq. I: on soil,
mh-42831, 42897, 42933, 42976, ta-4474,
4478, 4560; on humus, 4649. II: on boulder,
mh-43156. III: on soil, mh-41456.
Pogonatum dentatum (Brid.) Brid. II: on soil,
mh-43084. III: on rock-crevice, mh-12193.

80

Masanobu Higuchi and Tomotsugu Arikawa

Pogonatum japonicum Sull. & Lesq. I: on soil,
mh-42926, 43067, ta-4554; on humus, mh42880, ta-4642. II: on soil, mh-12344,
43149. III: on soil, mh-12292.
Pogonatum neesii (Müll.Hal.) Dozy Kiguchi etal. (2006).
Pogonatum sphaerothecium Besch. I: on rockcliff, mh-42886 (Hypnum pallescens). II:
on rock-cliff, mh-43060; on boulder, mh43049 (Pohlia revolvens). III: on rockcliff, mh-12333, 41575; on rock-crevice, mh12194 (Arctoa fulvella, Grimmia apiculata);
on boulder, mh-41528.
Pogonatum urnigerum (Hedw.) P.Beauv. I: on
boulder, mh-42829. III: on rock-cliff, mh41594 (Bartramiopsis lescurii); on humus,
mh-12277.
Polytrichum commune Hedw. I: on soil, mh12484, 42837 (Drepanocladus fluitans),
42932, ta-4473, 4499. II: on soil, mh-43072.
III: on soil, mh-12181, 41527; on decaying
stump, mh-41599.
Polytrichum formosum Hedw. I: on humus,
mh-42852.
Polytrichum juniperinum Hedw. Kiguchi et al.
(2006).
Polytrichum norvegicum Hedw. Kiguchi et al.
(2006).
Polytrichum ohioense Renauld & Cordot
Kiguchi et al. (2006).
Polytrichum piliferum Hedw. Kiguchi et al.
(2006).
Pottiaceae
Anoectangium stracheyanum Mitt. Kiguchi etal. (2006).
Anoectangium thomsonii Mitt. Kiguchi et al.
(2006).
Barbula unguiculata Hedw. Kiguchi et al.
(2006).
Bryoerythrophyllum brachystegium (Besch.)
K.Saito III: on rock-cliff, mh-41498, 41501,
41505 (Desmatodon laureri, Encalypta ciliata), 41507 (Encalypta ciliata), 41517.
Bryoerythrophyllum gymnostomum (Broth.)

Chen III: on rock-cliff, mh-41512 (Didymodon constrictus, Myurella julacea).
Bryoerythrophyllum
recurvirostre(Hedw.)
P.C.Chen Kiguchi et al. (2006).
Bryoerythrophyllum rubrum (Jur.) P.C.Chen var.
minus K.Saito Kiguchi et al. (2006).
Desmatodon laureri (Schultz) Bruch & Schimp.
III: on rock-crevice, mh-12234, 12238 (Encalypta ciliata), 12259.
Didymodon constrictus (Mitt.) K.Saito III: on
rock-cliff, mh-41512 (Bryoerythrophyllum
gymnostomum, Myurella julacea).
Didymodon rigidicaulis (Müll.Hal.) K.Saito
Kiguchi et al. (2006).
Didymodon rigidulus Hedw. var. gracilis (Schreich ex Hook. & Grev.) R.H.Zander Kiguchi
et al. (2006).
Gymnostomum aurantiacum (Mitt.) Paris
Kiguchi et al. (2006).
Oxystegus tenuirostris (Hook. & Taylor)
A.J.E.Smith I: on soil, mh-42975, 42988.
II: on rock-cliff, mh-43109; on rock-crevice,
mh-41493.
Tortella tortuosa (Hedw.) Limpr. Kiguchi et al.
(2006).
Tortula norvegica (F.Weber) Wahlenb. ex Lindb.
Kiguchi et al. (2006).
Tortula sinensis (Müll.Hal.) Broth. III: on
rock-cliff, mh-12240; on soil, mh-41508(Encalypta ciliata).
Trichostomum brachydontium Bruch Kiguchi
et al. (2006).
Weisiopsis anomala Broth. & Paris Kiguchi etal. (2006).
Weissia controversa Hedw. Kiguchi et al.
(2006).
Pterobryaceae
Pterobryum arbuscula Mitt.
mh-43001.

I: on tree-trunk,

Schistostegaceae
Schistostega pennata (Hedw.) F.Weber & Mohr
I: on soil, mh-42832, 42876, 42885, ta-4517.

Mosses of Mts. Yatsugatake

II: on soil, mh-12382, 12352.
mh-41466, 41525.

III: on soil,

Seligeriaceae
Blindia japonica Broth. Kiguchi et al. (2006).
Brachydontium noguchii Z.Iwats., Tad.Suzuki &H.Kiguchi Iwatsuki et al. (2004), Kiguchi etal. (2006).
Notes. When Iwatsuki et al. (2004) described Brachydontium noguchii Z.Iwats.,
Tad.Suzuki & H.Kiguchi, they cited specimens
collected from Mts. Yatsugatake as one ofparatypes.
Brachydontium trichodes (F.Weber) Milde I:
on rock, ta-4519; on boulder, mh-42881. II:
on rock, T. Furuki-2431.
Seligeria pseudodonniana Tad.Suzuki &Z.Iwats. II: on rock-cliff, mh-12329, 12330,
12362, 43058. III: on boulder, mh-41576,
41577.
Sematophyllaceae*Brotherella henonii (Duby) M.Fleisch. I: on
humus, ta-4528. II: on rotten log, mh-43032.
*Brotherella recurvans (Mich.) M.Fleisch. I:
on soil, mh-12451; on decaying log, mh12449. II: on decaying log, mh-43154. III:on rock-cliff, mh-41533.
Heterophyllium affine (Hook.) M.Fleisch. I:
on rock-cliff, mh-42910; on tree-trunk, mh42929, 42930, ta-4564; on decaying log, mh12501; on rotten log, mh-43015; on root, ta4645. II: on tree-trunk, mh-12372; on decaying log, mh-12365; on rotten log, mh-43114.
Pylaisiadelpha tenuirostris (Bruch & Schimp.)
W.R.Buck I: on tree-trunk, mh-42968, 42969(Callicladium haldanianum, Iwatsukiella
leucotricha) , ta-4576, 4601.
Sphagnaceae
Sphagnum capillifolium (Ehrh.) Hedw. I: on
humus, mh-12457, 42818 (Sphagnum fimbriatum), 42833, 42847, 42914.

81

Notes. Kiguchi et al. (2006) reported thisspecies as Sphagnum nemoreum Scop. from central region of Mts. Yatsugatake.
*Sphagnum fimbriatum Wilson ex Wilson &Hook.f. I: on humus, mh-12523, 12524,
42818 (Sphagnum capillifolium).
Sphagnum girgensohnii Russow I: on humus,
mh-12446, 12453, 12474, 12481, 42870,
42972, 42974, 42994, ta-4461, 4475, 4556,
4653. II: on humus, mh-12383, 43090,
43120, 43137, 43152. III: on rock, mh12179; on humus, mh-12162, 12281, 41454,
41529, 41540, 41593.
*Sphagnum quinquefarium (Lindb. ex Braithw.)
Warnst. I: on humus, mh-42853, 42915.
Sphagnum squarrosum Crome I: on humus,
mh-43019. III: on humus, mh-12163.
Sphagnum tenellum Ehrh. ex Hoffm. Kiguchi
et al. (2006).
Splachnaceae
Oedipodium griffithianum (Dicks.) Schwägr.
Kiguchi et al. (2006).
*Splachnum ampullaceum Hedw. Nishimura& Inoue (1981).
Tetraplodon angustatus (Hedw.) Bruch &Schimp. II: on stump, mh-12349; on humus,
mh-43125, 43147. III: on boulder, mh41539.
Tetraplodon mnioides (Hedw.) Bruch & Schimp.
II: on stump, mh-12348.
Takakiaceae*Takakia lepidozioides S.Hatt. & Inoue I: on
rock-cliff, mh-42873, 42874, 42890, 42891,
42896, 42900, 42902, 42906 (Dicranella
heteromalla), 42911 (Bartramiopsis lescurii,
Pohlia nutans), ta-4521, 4527, 4534, 4542,
4555.
Notes. Higuchi & Arikawa (2005) reported
this species for the first time from Mt. Yokodake(sometimes called as “Mt. Kita-yokodake” when
it is distinguished from Mt. Yokodake situated insouthern area) of Mts. Yatsugatake, where is the82

Masanobu Higuchi and Tomotsugu Arikawa

southernmost locality in Japan.
Tetraphidaceae
Tetraphis geniculata Girg. ex Milde I: on boulder, ta-4661; on decaying log, mh-12502; ondecorticated log, ta-4631; on rotten log, mh12416. II: on root, mh-43043, 43160. III:on rotten log, mh-41526.
Tetraphis pellucida Hedw. III: on root, mh43153 (Dicranella heteromalla).
Tetrodontium brownianum (Dicks.) Schägr. var.
repandum (Funck) Limpr. I: on rock, ta4516, 4518, 4658; on rock-cliff, mh-42875,
42879; on boulder, mh-42997, 43011, ta-4617.
II: on rock-cliff, mh-43046, 43068, 43071,
43092; on boulder, mh-43096. III: on rockcliff, mh-12328, 12331, 41457, 41537, 41565.
Theliaceae
Myurella julacea (Schwägr.) Bruch, Schimp. &Gümbel III: on rock-cliff, mh-41486, 41489,
41511 (Plagiopus oederianus, Thuidium
cymbifolium), 41512 (Bryoerythrophyllum
gymnostomum,
Didymodon
constrictus),
41515.
Myurella sibirica (Müll.Hal.) Reimers Kiguchi
et al. (2006).
Myurella tenerrima (Brid.) Lindb. Kiguchi etal. (2006).
Thuidiaceae
Abietinella abietina (Hedw.) M.Fleisch. III: on
humus, mh-12218; on boulder, mh-41481; onrock-cliff, mh-41516.
Anomodon rugelii (Müll.Hal.) Keissl. I: on decaying log, mh-43009.
Boulaya mittenii (Broth.) Cardot I: on treetrunk, mh-43002 (Dicranum muehlenbeckii),
ta-4667. II: on tree-trunk, mh-12384.
Bryonoguchia molkenboeri (Sande Lac.)
Z.Iwats. & Inoue III: on humus, mh-41620.
Claopodium pellucinerve (Mitt.) Best III: on
rock-crevice, mh-12244.

Claopodium prionophyllum (Müll.Hal.) Broth.
Kiguchi et al. (2006).
Haplocladium microphyllum (Hedw.) Broth.
Kiguchi et al. (2006).
Haplocladium
perparaphyllum
R.Watan.
Kiguchi et al. (2006).
*Haplohymenium pseudo-triste (Müll.Hal.)
Broth. III: on rock-cliff, mh-41483.
Haplohymenium sieboldii (Dozy & Molk.) Dozy
& Molk. Kiguchi et al. (2006).
*Helodium paludosum (Austin) Broth. I: on
humus, mh-42985 (Hylocomiastrum pyrenaicum).
Helodiumsachalinense(Lindb.)Broth.
Kiguchi et al. (2006).
Pelekium versicolor (Müll.Hal.) Touw Kiguchi
et al. (2006).
Rauiella fujisana (Paris) Reimers Kiguchi etal. (2006).
Thuidium cymbifolium (Dozy & Molk.) Dozy &Molk. III: on rock-cliff, mh-12235; on soil,
mh-12237 (Bartramia ithyphylla).
Thuidium lepidoziaceum Sakurai Kiguchi etal. (2006).
Thuidium philibertii Limpr. Kiguchi et al.
(2006).
Thuidium pristocalyx (Müll.Hal.) A.Jaeger
Kiguchi et al. (2006).
Thuidium recognitum (Hedw.) Lindb. var. delicatulum (Hedw.) Warnst. Kiguchi et al.
(2006).
Thuidium tamariscinum (Hedw.) Bruch &Schimp. Kiguchi et al. (2006).
Trachypodaceae
Duthiella speciosissima Broth. ex Cardot
Kiguchi et al. (2006).
Trachypus bicolor Reinw. & Hornsch. Kiguchi
et al. (2006).
Acknowledgements
We wish to thank Prof. H. Deguchi and Dr. T.
Furuki for the specimens and Mr. R. Suda for his
help in the field research in 1985. This study wasMosses of Mts. Yatsugatake

supported in part by Grant-in Aid (no. 13640707
to Higuchi) from the Ministry of Education, Culture, Sports, Science and Technology, Japan
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