Bull. Natl. Mus. Nat. Sci., Ser. A, 47(1), pp. 7–17, February 22, 2021
DOI: 10.50826/bnmnszool.47.1_7

A List of Pselaphine Species (Insecta, Coleoptera, Staphylinidae)
Collected from Mt. Phia Oac, North Vietnam
in 2017 with a Corrected Checklistof the Pselaphine Species Known from Vietnam
Shûhei Nomura1* and Thai Hong Pham2, 3
1

Department of Zoology, National Museum of Nature and Science,
4–1–1 Amakubo, Tsukuba, Ibaraki 305–0005, Japan
E-mail: nomura@kahaku.go.jp*: Corresponding author
2
Mientrung Institute for Scientiﬁc Research, Vietnam Academyof Science and Technology, Hue, Vietnam
3
Vietnam National Museum of Nature & Graduate School of Science and Technology,
Vietnam Academy of Science and Technology, Hanoi, Vietnam
E-mail: phamthai@vnmn.vast.vn(Received 14 September 2020; accepted 23 December 2020)

Abstract A list of pselaphine species collected from Mt Phia Oac, Cao Bang Province, North
Vietnam in 2017 is present. Collecting data of eighteen pselaphine species are listed up. Three pselaphine species, Pseudoplectus sp. 1, Triomicrus sp. 1 and Nomuraius piaoacus are observed by
SEM. A corrected checklist of the Vietnamese species of the subfamily Pselaphinae including 73
genera, 116 species is supplemented.
Key words : Pselaphinae, Staphylinidae, fauna, Vietnam, SEM.

Introduction
This paper ﬁrstly presents an annotated list
including collecting data of 18 pselaphine species taken from Mt Phia Oac, Cao Bang Province, North Vietnam in 2017, together with thehabitus-photos of all collected species, anddetailed SEM-photos of 3 species (Pseudoplectus
sp. 1, Triomicrus sp. 1 and Nomuraius piaoacus)
for identiﬁcation in species level. After the publication of a list of Vietnamese pselaphine species
by Nomura (2013), some newly recorded or
newly described Vietnamese species were added,
partly by conducting another faunal study of ours
(Nomura and Pham, 2019), and partly by
unknown papers to us (Kurbatov, Cuccodoro andIvan Löbl, 2007; Kurbatov and Sabella, 2008).
© 2021 National Museum of Nature and Science

Thus, second, in this paper, a corrected checklistof the Vietnamese species of the subfamily Pselaphinae including 73 genera, 116 species arealso supplemented.
Materials and Methods
The collecting survey was conducted from19th to 23rd Aug. 2017 at several points of Mt.
Phia Oac (＝Pia Oac), Cao Bang Province, North
Vietnam (Figs. 2A–D), whose geographical position is shown in Fig. 1. The pselaphine specimens were collected by the following methods.
1) Some portable light traps each with a ﬂuorescent tube 4W in the system of Dr. Yuta Nakase(＝NLT) were used (Fig. 2E). 2) Other some pselaphines were collected from leaf litter laid on
the ﬂoor of forests using simpliﬁed Winkler
apparatus (SWA) (Fig. 2F).

8

S. Nomura and T. H. Pham

Fig. 1. The location of Mt. Phia Oac on the map of N Vietnam.

For the SEM observation (Figs. 4–5), all specimens were air dried, uncoated, and illustrated
with an SEM ﬁt with a digital microscope system(KEYENCE VHX-2000＋VHX-D510) under AV
0.9–2.0 kv. Collected specimens are tentatively
preserved in collection of the department of
Zoology, National Museum of Nature and Science (NMNS), Tsukuba, Japan.

Results
1) The List of Pselaphine Species Collectedfrom Mt. Phia Oac, North Vietnam in 2017
In Aug. 2017, the ﬁrst author Nomura visited
Mt. Phia Oac (＝ Pia Oac), Cao Bang Prov., N
Vietnam, and collected eighteen species of pselaphines. The second author Pham supported
Nomura to collect Pselaphines in Mt. Phia Oac.
The annotated list including collecting data ofPselaphinae Species Collected from Mt. Phia Oac

9Fig. 2. Habitats and conditions of the traps in the survey at N Vietnam in 2017. A. A view of Mt. Phia Oac; B. A
view of the summit of Mt. Phia Oac; C. A forest view around the summit; D. A view of W (tungusten) mine on
the halfway up to Mt. Phia Oac; E. The portable light trap set in Mt. Phia Oac.; F. Simpliﬁed Winkler apparatuses used in the survey.

these species are herewith given.
Supertribe Euplectitae
1. Pyxidicerina, gen. and sp. undet 1 (Fig. 3A)Specimens examined. 1 male, W (: wolfranium＝ tungsten) mine, St. A, alt. 1,672 m, by SWA,
20. viii.; 1 female, SE slope, St. A, alt. 1,287 m,
by SWA, 21. viii.
Remarks. The subtribe Pyxidicerina is separated from the other subtribe Bythinnoplectina by
having a pair of deep cavities where the maxil-

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S. Nomura and T. H. Pham

Fig. 3. Photos of pselaphine species collected in the survey. A. Pyxidicerina, gen. and sp. undet 1; B. Pseudoplectus? sp. 1, male; C. ditto, female; D. Trimium? sp. 1; E. Intestinarius sp. 1; F. I. sp. 2; G. Trisinus sp. 1; H.
T. sp. 2; I. T. sp. 3; J. Sathytes sp. 1; K. Reichenbachia sp. 1; L. Trissemus sp. 1; M. Batraxis sp. 1; N. Triomicrus sp. 1, male; O. ditto, female; P. Plagiophorus sp. 1; Q. P. sp. 2; R. Pseudophanias sp. 1; S. Pselaphodes
sp. 1; T. Nomuraius piaoacus Yin et Lee, 2013, male; U. ditto, female.

lary palpi are contracted not on the both lateralsides (Bythinoplectina), but on the frontal side. It
is common in Southeast Asia as shown in
Nomura et al. (2010) (15 spp. from Thailand),and Nomura (2019) (3 spp. from Laos).

than those of the females. In both sexes, the pronotum is rounded on both sides, with a pair ofbasilateral and a basimedian fovea, and a pair ofshort sulci connecting basilateral and basimedianfoveae.

2. Pseudoplectus? sp. 1 (Figs. 3B–C, 4)
Specimens examined. 1 male, Nr. Summit, St.
A, alt. ca 1,900 m, by SWA, 22. viii.; 2 females,
W mine, St. B, alt. 1,672 m, by SWA, 20. viii.
Remarks. This genus is characterized by thevery small and elongate body. After the SEM
obserbation, the male and the female characters
were examined as shown in Fig. 4. The male iseasily distinguished from the female by the sexual patch laid on the frons and the strongly thickened mid femur. The male eyes are clearly larger

3. Trimium? sp. 1 (Fig. 3D)Specimens examined. 1 female, W mine, St. A,
alt. 1,672 m, by SWA, 20. viii.
Remarks. The genus Trimium is the type genusof the subtribe Trimiina, which is distinct in thetribe Trichonychini by having the antennal club
formed only by the terminal segment (antennomere XI).

Pselaphinae Species Collected from Mt. Phia Oac

11

Fig. 4. SEM photos of Pseudoplectus? sp. 1. A–C. male; D–F. female. A, D. habitus in dorsal view; B, E. headand pronotum; C, F. head enlarged.

Supertribe Batrisitae
4. Intestinarius sp. 1 (Fig. 3E)
Specimen examined. 1 female, W mine, St. B,
alt. 1,672 m, by SWA, 20. viii.
Remarks. This genus was deﬁned by Kurbatov(2007) with the type species, Batrisus quinquesulcatus Raﬀray, 1894 known from Singapore
and seven new species were described. Later, Yin
et al. (2011) added three species from China.
Then, eleven species have been known in thisgenus. Although no named species of this genus
has been known from Vietnam according to
Nomura (2013), some unnamed species are
already recognized in Vietnam (Nomura, unpub-

lished).
5. Intestinarius sp. 2 (Fig. 3F)
Specimen examined. 1 male, Nr. Summit, St.
A, alt. ca 1,900 m, by SWA, 22. viii.
Remarks. This genus is known from China,
Laos, Malaysia and Singapore. Two unnamed
species collected in Mt. Phia Oac were both collected from high altitude area. This second species is very characteristic in the large head andanteriorly narrowed elytra.
6. Trisinus sp. 1 (Fig. 3G)Specimens examined. 1 male, Mt. Phia Oac,alt. ca 1,600 m, by NLT-LP (1 m above the12

S. Nomura and T. H. Pham

Fig. 5. SEM photos of Triomicrus spp. A, E. T. vietus Löbl et al. from Mt. Tam Dao, male; B, F. ditto, female; C,
G. T. sp. 1 from Mt. Phia Oac, male; D, H. ditto, female. A–D. head and pronotum in dorsal view; E–H. apicalsegments of antenna.

ground), 19–20. viii.; 2 females, W mine, St. A,
alt. 1,672 m, by SWA, 20. viii.
Remarks. The genus Trisinus was establishedwith the type species, T. cratocerus Raﬀray, 1894
collected from Singapore. The east Asian genus
Batrisoplisus was synonymized with Trisinus by
Yin et al. (2012). In this time, 26 species were
included in this genus.

21–22. viii.
Remarks. This genus is very characteristic by
the thick body densely covered with coarse punctures and strongly modiﬁed antennal club in the
male. It is broadly distributed in tropical to East
Asia and distinctly diversiﬁed.

7. Trisinus sp. 2 (Fig. 3H)Specimen examined. 1 female, SE slope, St. A,
alt. 1,287 m, by SWA, 21. viii.
8. Trisinus sp. 3 (Fig. 3I)
Specimen examined. 1 male, 2 females, Nr.
Summit, St. A, alt. ca 1,900 m, by SWA, 22. viii;
1 female, same data as above, but St. B.

10. Reichenbachia sp. 1 (Fig. 3K)
Specimens examined. 1 male, Mt. Phia Oac,alt. ca 1,600 m, by NLT-HP (4 m above theground), 19–20. viii.
Remarks. This genus is highly diversiﬁed in
the Palearctic and Oriental Regions. According
to Nomura (2013), four species of this genus has
been known from Vietnam.

9. Sathytes sp. 1 (Fig. 3J)Specimen examined. 1 male, SE slope, alt.
1,287 m, by NLT-LP (1 m above the ground),

11. Trissemus sp. 1 (Fig. 3 L)Specimen examined. 1 male, SE slope, alt.
1,287 m, by NLT-LP (1 m above the ground),Supertribe Goniaceritae

Pselaphinae Species Collected from Mt. Phia Oac

13

Fig. 6. SEM photos of Nomuraius piaoacus Yin et Li. A–C. male; D–F. female. A, D. Head and pronotum in
dorsal view; B, E. Ditto, enlarged; C, F. antennal club.

21–22. viii.
Remarks. This genus is very closely allied to
the former genus, but is separated by having theelytra each with 3 basal foveae. It includes ﬁve
Vietnamese species according to Nomura (2013).
12. Batraxis sp. 1 (Fig. 3M)Specimen examined. 1 male, SE slope, alt.
1,287 m, by NLT-LP (1 m above the ground),
21–22. viii.
Remarks. The genus Batraxis is easily separated from the other Brachyglutine genera byhaving the thick and robust body, the slender andsometimes swelling legs and the asymmetricalterminal segment of antenna.

13. Triomicrus sp. 1 (Figs. 3N–O, 5C–D, G–H)
Specimens examined. 1 male, SE slope, St. A,
alt. 1,287 m, by SWA, 20. viii.; 1 female, W
mine, St. B, alt. 1,622 m, by SWA, 20. viii.
Remarks. This genus is distributed in East to
Southeast Asia. As to the Vietnamese fauna one
species of this genus, T. vietus Löbl et al., 1998
were described from Mt. Tam Dao, Vinh Phu
Province, N Vietnam. This species collected fromMt Phia Oac was easily separated from T. vietus
by the more sparsely and more weakly punctuated head and pronotum as shown in Fig. 4.
14. Plagiophorus sp. 1 (Fig. 3P)Specimen examined. 1 female, W mine, St. A,

14

S. Nomura and T. H. Phamalt. 1,622 m, by SWA, 20. viii.
Remarks. This genus is very characteristic in
the small, robust and densely punctuated body,the seven- or ten-segmented antenna, and theconjoined abdominal segments III to VII. Two
species of this genus were collected from Mt.
Phia Oac in this study, both of which have
10-segmented antennae.
15. Plagiophorus sp. 2 (Fig. 3Q)
Specimens examined. 1 female, W mine, St. A,
alt. 1,622 m, by SWA, 20. viii.; 1 female, Nr.
Summit, St. A, alt. ca 1,900 m, by SWA, 22. viii.
Supertribe Pselaphitae
16. Pseudophanias sp. 1 (Fig. 3R)Specimen examined. 1 female, SE slope, St. A,
alt. 1,287 m, by SWA, 21. viii.
Remarks. The genus Pseudophanias wasestablished for the type species, P. malaianus
Raﬀray, 1890 described from Penang Is., Malaysia. It is characterized by the middle- to largesized and stout body, very small and symmetricalmaxillary palpus with fusiform parpomere 4.
17. Pselaphodes sp. 1 (Fig. 3S)Specimen examined. 1 male, SE slope, alt.
1,287 m, by NLT-LP (1 m above the ground),
21–22. viii.
Remarks. This genus is characterized by thevery large and broad body and the externallyexpanded or projected palpomeres 3 and 4, andthe metasternum lacking median fovea. It has
been known from East, Southeast and South
Asia.
18. Nomuraius piaoacus Yin et Li, 2013 (Figs.
3T–U)
Specimens examined. 1 female, SE slope, St.
A, alt. 1,287 m, by SWA, 21. viii.; 1 male, 1female, same as above, but st. B; 1 male, 1female, W mine, St. B, alt. 1,622 m, by SWA, 20.
viii.
Remarks. The genus Nomuraius was deﬁned
by Hlaváč (2003) with the type species, N. viet-

namicus Hlaváč, 2003 described from Mt. Tam
Dao, Vinh Phu Province, N Vietnam. It is distinct
in the very large body with very long antennae
and legs, and the posteriorly prolonged tarsomere
2 of each leg. This species was described by Yin
and Li (2013) from Mt. Phia Oac.
2) The Corrected Checklist of the Pselpahine
Species known from Vietnam until 2019.
Nomura (2013) listed up 97 spp. in 66 generaof the pselaphine species known from Vietnam.
However, it did not cover all known species from
Vietnam. Nomura and Pham (2019) corrected thechecklist with an addition of fourteen species,
and the number of the Vietnamese species
became 111. In these inventory works, the important papers, Kurbatov et al. (2007) and Kurbatov
and Sabella (2008) were still overlooked.
Kurbatov et al. (2007) revised the taxonomyof the genus Morana and the allied genera in the
supertribe Goniaceritae, the tribe Iniocyphini, thesubtribe Natypleurina. Four species in 2 genera
were added to the Vietnamese pselaphine fauna.
Kurbatov and Sabella (2008) taxonomicallyrevised the genus Atychodea of the tribe Tychiniin the supertribe Goniaceritae. By this paper,
they added two new species of Atychodea from
Vietnam. As the result, 116 species in 73 generaof pselaphines have been known from Vietnam
up to the present.
The asterisk used in this species list means asfollows:*: The newly added species by Nomura andPham (2019).
**: The presently supplemented species.
***: This species was recorded as Ctenistidius
coomani Jeannel, 1957 in Nomura (2013). However, the genus Ctenistidius Jeannel, 1957 wassynonymized with Ctenisoschema Jeannel, 1956
in Besuchet (2008). This nomenclatural act was
overlooked by Nomura (2013).
Tribe Dimerini
Octomicrites bifrons (Jeannel, 1952)
Octomicrus longulus longulus Schaufuss,
1877

Pselaphinae Species Collected from Mt. Phia Oac

Tuberoplectus cucphuongensis Nomura,
2010
Supertribe Euplectitae
Tribe Bythinoplectini
Subtribe Bythinoplectina
Bolbozethus boosteri Coulon, 1982
Zethopsus opacus (Schaufuss, 1877)
Subtribe Pyxidicerina
Pyxidicerinus barbieri Jeannel, 1952
Tribe Trichonychini
Subtribe Bibloporina
Aphilia angusticollis Jeannel, 1952
Subtribe Panaphantina
Bibloplectinus solskyi (Schaufuss, 1877)
Euplectodina hipposideros (Schaufuss,
1877)
Hypoplectus (Hypoplectinus) barbieri(Jeannel, 1952)
H. (H.) laticollis (Jeannel, 1952)
H. (H.) depressifrons Jeannel, 1957
Methorius truncaticollis Jeannel, 1952
Philiopsis obscura Raﬀray, 1898
Supertribe Batrisitae
Tribe Batrisini
Subtribe Batrisina
Ancistromus longulus Jeannel, 1957
Arthromelodes cariei Jeannel, 1952
A. reichenbachi (Motschulsky, 1851)*
Batricrator trabisoides Jeannel, 1957
Batriscenodes siamensis (Raﬀray, 1904)
B. excisus (Schaufuss, 1877)
Batriscenites morulus (Reitter, 1882)
Batrisiella barbieri (Jeannel, 1952)
B. micromelus (Jeannel, 1952)
B. coomani (Jeannel, 1957)
Batrisocenus annamita Raﬀray, 1904*
B. cribricollis Jeannel, 1957
Coryphomodes coomani Jeannel, 1957
Cratna abdominalis Löbl, 1975*
C. cicatricosa Raﬀray, 1918*
Nenemeca barbieri Jeannel, 1952
Physomerinus septemfoveolatus (Schaufuss,
1877)

15

Sathytes clavatus (Jeannel, 1957)
Sinotrisus vietnamensis Yin et Nomura,
2012
Supertribe Goniaceritae
Tribe Arnyllini
Harmophorus barbieri (Jeannel, 1952)
Harmophorus ciliatus (Raﬀray, 1894)
Tribe Brachyglutini
Subtribe Brachyglutina
Atenisodus longicornis (Raﬀray, 1895)
Barbiera frontalis Jeannel, 1952
Batraxis curvispina Raﬀray, 1893
B. raﬀrayana (Blattný, 1925)*
B. variabilis Raﬀray, 1890.
Berlaraxis coomani Jeannel, 1957
B. punctata Jeannel, 1957
Comatopselaphus parcepunctatus Raffray, 1904
Eupinidius barbieri Jeannel, 1952
Eupines sphaerica (Motschulsky, 1851)
Prosthecarthron sauteri Raﬀray, 1914
Reichenbachella budha (Raﬀray, 1891)
R. coomani Jeannel, 1957
R. rufa (Schmidt-Göbel, 1838)
Reichenbachia baumeisteri Schaufuss,
1877
Rybaxis dentipes (Jeannel, 1952)
R. villosa Raﬀray, 1904*
Triomicrus vietus Löbl, Kurbatov et
Nomura, 1998
Trissemus mamilla (Schaufuss, 1877)
T. oblongulus Jeannel, 1957
T. eurycnemis Jeannel, 1957
T. coomani Jeannel, 1957
T. implicitus (Raﬀray, 1912)
Tribe Iniocyphini
Subtribe Natypleurina
Morana belajevae Kurbatov et Cuccodoro, 2007**: 620 Ma Da, 55 km NE
Ho Chi Minh
M. hoplomacha Kurbatov et Cuccodoro,
2007**: 637 Cat Tien, 120 km NNE Ho
Chi Minh

16

S. Nomura and T. H. PhamM. lucipeta Kurbatov et Cuccodoro,
2007**: 639 Cat Tien, 120 km NNE Ho
Chi Minh
Nippiliphus napolovi Kurbatov et Cuccodoro, 2007**: 669 Env. Tam Dao,
800 m, 55 km NNW Hanoi
Tribe Proterini
Bythinoderes marginata Raﬀray, 1896
Pareuplectops coomani Jeannel, 1957
P. fartor Kurbatov et Cuccodoro, 2009*
P. tenasserimi (Blattný, 1925)*
Tribe Tychini
Atychodea fedorenkoi Kurbatov et al.,2008**: 34 Cat Tien, 120 km NNE Ho
Chi Minh
A. vicina Kurbatov et al., 2008**: 35 Cat
Tien, 120 km NNE Ho Chi Minh
Supertribe Pselaphitae
Tribe Pselaphini
Curculionellus setosus Jeannel, 1952
Mentraphus pselaphoides Sharp, 1883
Pselaphaulax bifoveolatus (Shaufuss,
1877)
P. articularis (Schaufuss, 1877)
Pselaphidius multangulus (Schaufuss,
1877)
P. parvipalpis Reitter, 1883
P. pilicollis Reitter, 1882
Pselaphogenius vietnamicus Nomura,
2003
Pselaphus pubescens Raﬀray, 1904*
Tyraphus testaceus Schaufuss, 1877
T. semiopacus Schaufuss, 1877
T. pilosus Raﬀray, 1891
T. baeri Raﬀray, 1891
T. coomani Jeannel, 1957
T. brevicornis Jeannel, 1957
Tribe Hybocepharini
Apharina conicicollis (Schaufuss, 1877)
Mestogaster barbieri Jeannel, 1952
Stipesa cariniventris (Schaufuss, 1877)
S. ampliventris (Schaufuss, 1877)

Tribe Tmesiphorini
Raphitreodes dentimanus (Raﬀray, 1890)
Tribe Tyrini
Subtribe Centrophthalmina
Centrophthalmina gracilicornis Jeannel,
1952
C. laevis (Raﬀray, 1896)
Centrophthalmus clementis Schaufuss,
1877
C. herferi Blattný, 1925
C. punctipennis Schaufuss, 1877
C. sinensis Raﬀray, 1904
Enantius brevipalpis Jeannel, 1952
Subtribe Tyrina
Lasinus mandarinus Raﬀray, 1890
Megatyrus laqueus Hlaváč et Nomura,
2003
M. coni Hlaváč et Nomura, 2003
Nomuraius vietnamicus Hlaváč, 2002
N. piaoacus Yin et Li, 2013*
Paralasinus antennatus Hlaváč et
Nomura, 2001
P. humilis Hlaváč et Nomura, 2001
P. namcatienicus Hlaváč et Nomura, 2001
P. thayerae Hlaváč et Nomura, 2001
Sublipalpis myrmecophilus (Raﬀray,1904)*
Tyrodes setosus Jeannel, 1957
Tribe Ctenistini
Ctenistes mitis Schaufuss, 1882
Ctenisoschema
coomani(Jeannel,***
1957)
Enoptostomus crassicornis (Raﬀray,
1904)
Poroderus maindroni (Raﬀray, 1909)*.
Tribe Odontalgini
Odontalgus gracilis Raﬀray, 1904*
Odontalgus vestitus Schaufuss, 1886
Supertribe Clavigeritae
Tribe Clavigerini
Subtribe Clavigerodina
Microdiartiger thinhi Nomura, 2007

Pselaphinae Species Collected from Mt. Phia Oac

Subtribe Mastigerini
Cerylambus reticulatus (Raﬀray, 1895)
Mastiger brevicornis Raﬀray, 1890*
Tribe Colilodionini
Colilodion thienmu Nomura et Sugaya,
2007
Acknowledgements
We wish to express our hearty thanks to Mr.
Nguyen Van Dat (VNMN) for his kind support toour ﬁeldwork in Mt. Phia Oac.
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