Bull. Natl. Mus. Nat. Sci., Ser. B, 34(3), pp. 103–111, September 22, 2008

Type Collection in the Herbarium of
National Museum of Nature and Science (TNS)
Ericales, Primulales and Lamiales
Atsushi Ebihara and Kyoko Miyashita
Department of Botany, National Museum of Nature and Science,
Amakubo 4–1–1, Tsukuba-shi, 305–0005 Japan

Abstract Type specimens of three angiosperm orders (Ericales, Primulales and Lamiales) deposited in the herbarium of National Museum of Nature and Science (TNS) were explored and reexamined. In total, 109 sheets are confirmed as type collections for 68 taxa including 43 holotypes.
The 109 sheets consist of 3 Clethraceae, 24 Ericaceae, 1 Epacridaceae, 4 Myrsinaceae, 20 Primulaceae, 9 Boraginaceae, 5 Verbenaceae and 43 Labiatae.
Key words : Ericales, Lamiales, Primulales, TNS-herbarium, type collection.

CLETHRACEAE F. (Family No.) 228
Clethra L.
Clethra barbinervis Siebold et Zucc. forma
grandis Hayashi
in Bull. Gov. For. Exp. Stat. 77: 30, fig. 20(1955).
Jap. Name. Ohba-ryobu.
Japan, Hondo, Prov. Iwaki [Fukushima
Pref.], Iwaki-gun, Kawabe-mura, Mt. Megane,
1953-7-10, Y. Hayashi s.n. [TNS-VS-255849
Holotype].
Note. Holotype was originally deposited in the
Herbarium of Government Forest Experimental Station according to the original description, but was later transferred to TNS.
Clethra stolonifela Nakaiin Bull. Natl. Sci. Mus., Tokyo 33: 19 (1953).
Jap. Name. Chabo-ryobu.
Japan, Hondo, Prov. Iwashiro [Fukushima
Pref.], Oze, monte Hiutji [Hiuchi], alt.
1850 m, 1950-8-27, N. Maruyama s.n. [TNSVS-84014 Syntype]. Japan, Prov. Tôtômi
[Shizuoka Pref.], Shuchi-gun, urbis Misakubo, Mt. Shirakurayama, alt. 1800 m, 194910-25, Y. Hayashi s.n. [TNS-VS-81077 Syntype].

Note. TNS-VS-84014 was designated as“florifer.-typus” and TNS-VS-81077 as “typus
fructus”. Both specimens should be syntypes.
ERICACEAE F. 230
Hugeria Small
Hugeria incisa F.Maek.
in Bot. Mag. (Tokyo) 47: 614, fig. 15 (1933).
Jap. Name. Hagoromo-akushiba.
Japan, Hondo-borealis, Prov. Uzen [Yamagata Pref.], Urbe Higashine, Rokuta, 1932-8-20,
S. Okuyama s.n. [TNS-VS-41677 Isotype].
Note. Holotype in TI (according to the originaldescription).

Menziesia Sm.
Menziesia ciliicalyx (Miq.) Maxim. var. lasiostipes Ohwi
in Fl. Jap. (Ohwi) [Japanese ed.] 883 (1953).
Jap. Name. Kenaga-usugi-youraku.
Japan, Prov. Mikawa [Aichi Pref.], Mt.
Goban-ishi, 1934-8-21, Y. Kato s.n. [TNS-VS45752Holotype].
Menziesia goyozanensis M.Kikuchiin J. Jap. Bot. 37: 355, figs. 1–2 (1962).

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Atsushi Ebihara and Kyoko Miyashita

Jap. Name. Goyozan-youraku.
Japan, Prov. Rikuzen [Iwate Pref.], monte
Goyozan, 1962-6-27, M. Kikuchi s.n. [TNSVS-216513 Isosyntype].
Menziesia katsumatae M.Tash. et H.Hatta
in Bot. Mag. (Tokyo) 99: 29, figs. 1–2 (1986).
Jap. Name. Hozaki-tsurigane-tsutsuji.
Japan, Honshū, Gifu Pref., Gujo-gun, Hachimanchō, Mt. Takagasan, 1974-6-14, H. Katsumata and M. Tashiro 5641 [TNS-VS458266Holotype]. H. Katsumata & M.
Tashiro 5642 [TNS-VS-467801 Isotype].
Menziesia yakushimensis M.Tash. et H.Hatta
in Bot. Mag. (Tokyo) 99: 33, figs. 3–5 (1986).
Jap. Name. Yakushima-youraku.
Japan, Kyushu, Kagoshima Pref., Yaku-shimaIsland, Mt. Nagatadake, 1973-6-15, H. Hatta
s.n. [TNS-VS-458268 Holotype; TNS-VS458267Isotype].
Phyllodoce Salisb.
Phyllodoce aleutica (Spreng.) A.Heller var.
marmorata Toyok.
in Symb. Asahikawa 3: 128 (1968).
Jap. Name. Nishiki-tsugazakura.
Japan, Prov. Ishikari, in monte Asahi montium
Taisetsusan, 1966-7-21, H. Toyokuni and Y.
Toyokuni s.n. [TNS-VS-285851 Holotype].
Phyllodoce alpina Koidz. forma viridiflora
Nakai
in J. Jap. Bot. 14: 742 (1938).
Jap. Name. Seika-ko-tsugazakura.
Japan, Honshu, Prov. Etchu [Pref. Toyama],
Mt. Tateyama, 1938-8-17, Y. Katô 13 [TNSVS-59174 Isotype].
Note. Holotype probably in TI.
Rhododendron L.
Rhododendron albrechtii Maxim. forma albiflorum T.Yamaz.
in Rev. Rhodod. Jap., Taiwan, Kor. Sakhal. 121(1996).
Jap. Name. Shirobana-murasaki-yashio-tsutsuji.
Japan, Pref. Akita, Senhokugun, Kakunodate-machi, Shiraiwadake, alt. 1230 m, 1974-6-10,
S. Saito s.n. [TNS-VS-322660 Isotype].
Note. Holotype in TI (according to the originaldescription).
Rhododendron amanoi Ohwi
in Bull. Natl. Sci. Mus., Tokyo n.s. 1: 4
(1954).
Jap. Name. Sakishima-tsutsuji.
Japan, Ryukyu, Ins. Iriomote, Inaba, secundum fluvii Urauchi, 1953-6-27, T. Amano
7185 [TNS-VS-102296 Holotype].
Rhododendron eriocarpum (Hayata) Nakai var.
tawadae Ohwi
in Bull. Natl. Sci. Mus., Tokyo n.s. 1: 4
(1954).
Jap. Name. Senkaku-tsutsuji.
Japan, Ryukyu [Okinawa Pref.], ins. Uwotsuri, Mt. Uwotsuridake, alt. 300–320 m, 19524-17, S. Tawada 29 [TNS-VS-102295 Holotype; TNS-VS-102293 and TNS-VS-102294
Isotype].
Rhododendron keiskei Miq. var. hypoglaucum
Suto et T.Suzuki
in Trans. Nat. Hist. Soc. Taiwan 22: 23 (1932).
Jap. Name. Urajiro-hikage-tsustuji.
Japan, Prov. Shimotuke [Tochigi Pref.],monte Ozaku-san, 1931-5-?, E. Kitamura
ST3902 [TNS-VS-55292 Isotype].
Note. Holotype in TAI (according to the original description).
Rhododendron reticulatum D.Don ex G.Don
var. ciliatum Nakai
in Bot. Mag. (Tokyo) 40: 486 (1926).
Jap. Name. Arage-mitsuba-tsutsuji.
Japan, Prov. Suwo [Yamaguchi Pref.], Yoshiki-gun, oppido Ōuchi, Hikami, 1892?-?-?, J.
Nikai 423 [TNS-VS-46989 Isotype].
Note. Holotype in TI (according to the originaldescription). The collection date is not accurately shown on the label — flowering in
1892-4-20, unfolding young leaves in 1892-422, fruiting in 1892-6-17. This specimen has
flowers, leaves and fruit.
Rhododendron schlippenbachii Maxim. var.
ciliatum Nakaiin Bull. Natl. Sci. Mus., Tokyo 33: 19 (1953).

TNS type collections

South Korea, Zennan, monte Gesshutusan,
1936-8-5, M. Bok (S. Hozawa) s.n. [TNS-VS84733Holotype].
Rhododendron serpyllifolium Miq. forma amagianum Hayashi
in Bull. Gov. For. Exp. Stat. 170: 81, t. 1, fig.
51 (1964).
Jap. Name. Amagi-unzen-tsutsuji.
Japan, Hondo, Prov. Idzu [Shizuoka
Pref.], Tagata-gun, Amagi-yugashima-cho, Mt.
Amagi, 1962-5-4, Y. Hayashi s.n. [TNS-VS255847Holotype].
Rhododendron tschonoskii Maxim. forma
roseotinctum Kurash., N.Kobayashi et
Yukawain Ann. Tsukuba Bot. Gard. 19: 6 (2000).
Jap. Name. Tsumabeni-choji-kometsutsuji.
Japan, Nagano Pref., Komoro-shi, Mt. Kurofu,
alt. 2300 m, 1997-7-16, Y. Kurashige & T.
Yukawa 351 [TNS-VS-9508515 Holotype].
Rhododendron wadanum Makino forma
kaiense Hiyama
in J. Jap. Bot. 28: 217 (1953).
Jap. Name. Kai-mitsuba-tsutsuji.
Japan, Hondo, Prov. Kai [Yamanashi Pref.],
Mt. Mitsutôge, 1935-5-12, K. Hiyama s.n.
[TNS-VS-105557 Holotype].
Tripetaleia Siebold et Zucc.
Tripetaleia paniculata Siebold et Zucc. var. fruticulata Nakaiin Bull. Natl. Sci. Mus., Tokyo 33: 20 (1953).
Jap. Name. Akizaki-hotsutsuji.
Japan, Hondo, Prov. Etjigo [Niigata Pref.],
Higashi-kambara-gun, urbis Tugawa, Mt. Kirinzan, 1950-10-7, T. Nakai and N. Maruyamas.n. [TNS-VS-83989 and TNS-VS-83990 Syntypes; TNS-VS-83988 Isosyntype].
Note. TNS-VS-83989 and -83990 are designated as “Holotype of flowers” and TNS-VS83990 as “Holotype of fruits”, respectively.
Since they are “not clearly labeled as being
part of a single specimen” (ICBN 2006, Art.
8.3.), TNS-VS-83989 and -83990 should be
syntypes. The collection date is different from

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that in the original description (1950-10-10).
Tritomodon Turcz.
Tritomodon campanulatus (Miq.) F.Maek. ex
Okuyama forma ellipticum Okuyama
in J. Jap. Bot. 30: 40 (1955).
Jap. Name. Koban-sarasa-dodan.
Japan, Hondo, Prov. Musashi [Tokyo Pref.],
Komazawa, Toritsu-engei [Tokyo Metropolitan Horticulture High School], 1952-5-13, T.
Oba 2523 [TNS-VS-90280 Holotype].
Note. A cultivated plant.
EPACRIDACEAE F. 232
Cyathodes Labill.
Cyathodes mariannensis Kaneh.
in Bot. Mag. (Tokyo) 48: 734, fig. 5 (1934).
Mariana Islands (USA), Aramagan Island,
1933-7-6, R. Kanehira 2182 [TNS-VS-11160
Isotype].
Note. Holotype probably in FU.
MYRSINACEAE F. 234Anamtia Koidz.
Anamtia stolonifera Koidz.
in Bot. Mag. (Tokyo) 37: 40 (1923).
Jap. Name. Tsuru-manryo.
Japan, Honshû, Prov. Yamato [Nara Pref.],
Yoshino-gun, Imoyama near Kamiichi, 192210-9, G. Koizumi s.n. [TNS-VS-364347 Isotype].
Note. Holotype probably in KYO.
Ardisia Sw.
Ardisia sciophila T.Suzuki
in J. Jap. Bot. 13: 507 (1937).
Taiwan, Taihoku-shû, Bunzan-gun, Agyoku,
Tôgo-kei, Habun-Baron, 1936-6-7, T. Suzuki
ST15040 [TNS-VS-60019 Isotype].
Note. Holotype in TAI (according to the original description).

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Atsushi Ebihara and Kyoko Miyashita

Bladhia Thunb.
Bladhia violacea T.Suzuki
in Trans. Nat. Hist. Soc. Taiwan 25: 41 (1935).
Taiwan, Pref. Taihoku-Shû, inter Tyakon et Rimogan, 1934-9-4, T. Suzuki 11697 [TNS-VS64491Isotype].
Note. Holotype in TAI (according to the original description).
Rapanea Aubl.
Rapanea okabeana Tuyama
in Bot. Mag. (Tokyo) 52: 465, figs. 2b, 3(1938).
Jap. Name. Maruba-taimin-tachibana.
Japan, Bonin Islands, Isl. Titizima (Chichijima), Takeda-Bokuzyo, 1938-3-1, T. Tuyamas.n. [TNS-VS-59478 Isotype].
Note. Holotype in TI (according to the originaldescription).
PRIMULACEAE F. 235Lysimachia L.
Lysimachia clethroides Duby var. crassifoliosa
Konta
in Bull. Natl. Sci. Mus., Tokyo, Ser. B 31: 138,
figs. 4, 8 (2005).
Jap. Name. Isokaze-okatoranoo.
Japan, Honshu, Pref. Shizuoka, Shimoda City,
Suzaki Peninsula, Suzaki, Region II, Suzakigoyotei, alt. 50 m, 2005-7-2, F. Konta 24430-a[TNS-VS-749838 Holotype],F.
Konta
24430-b[TNS-VS-749840 Isotype],
F.
Konta 24430-c [TNS-VS-749837 Isotype], F.
Konta 24430-d [TNS-VS-749839 Isotype].
Same locality, 2005-6-19, F. Konta 24395-a[TNS-VS-749602 Paratype],F.
Konta
24395-b [TNS-VS-749601 Paratype], F.
Konta 24395-c [TNS-VS-749561 Paratype],
F. Konta 24395-d [TNS-VS-749562 Paratype].
Primula L.
Primula candicans W.W.Sm.
in Trans. & Proc. Bot. Soc. Edinburgh 35: 190(1950).
China, South East Tibet, Sobhe La, near
Tongyub Dzong, 1947-5-21, F. Ludlow, G.
Sherriff and H. H. Elliot 13749 [TNS-VS107000 and TNS-VS-107017Isotypes].
Note. Holotype in BM (according to the original description).
Primula cuneifolia Ledeb. var. hakusanensis(Franch.) Makino forma albiflora Makino
in J. Jap. Bot. 1: 20 (1917).
Jap. Name. Shirobana-nankin-kozakura.
Japan, Prov. Iwashiro [Fukushima Pref.], Mt.
Iide, 1908-8-16, collector unknown [TNS-VS14794Holotype].
Primula cuneifolia Ledeb. var. heterodonta(Franch.) Makino forma albiflora Makino
in J. Jap. Bot. 1: 20 (1917).
Jap. Name. Shirobana-michinoku-kozakura.
Japan, Prov. Mutsu [Aomori Pref.], Mt.
Iwaki, 1885-8-?, H. Sakurai s.n. [TNS-VS9889Holotype].
Primula hisauchii Miyabe
in Trans. Sapporo Nat. Hist. Soc. 16: 187(1940).
Jap. Name. Kiyosato-kozakura.
Japan, Honshu, Prov. Kai [Yamanashi Pref.],
Kiyosato-mura, 1936-6-14, K. Hisauchi 1415
[TNS-VS-52354Holotype].
Primula kisoana Miq. var. shikokiana Makino
in J. Jap. Bot. 1: 19 (1917).
Jap. Name. Shikoku-kakkoso.
Japan, Shikoku, Prov. Iyo [Ehime Pref.],around Kuma, 1885-5-15, K. Fudzino s.n.
[TNS-VS-9899Syntype].
Note. Another syntype collected by K. Okudaira is not found in TNS.
Primula miyabeana T.Ito et Kawak.
in Miyabe Festschrift 1, pl. 29 (1911).
Taiwan, montem Morrison (Niitaka-yama), alt.
12000 ft., 1908-11-?, T. Kawakami and U.
Mori s.n. [TNS-VS-61543 Holotype].
Primula modesta Bisset et S.Moore var.
shikokumontana Miyabe et Tatew.
in Trans. Sapporo Nat. Hist. Soc. 16: 188(1940).
Jap. Name. Ishizuchi-kozakura.

TNS type collections

Japan, Shikoku, Prov. Iyo [Ehime Pref.], Mt.
Ishizuchi, 1885-8-?, T. Makino s.n. [TNS-VS9891Holotype].
Note. The spelling of the variety name is originally “shikoku-montana”.
Primula reinii Franch et Sav. var. ovatifolia
Ohwi
in Fl. Jap. (Ohwi) [Japanese ed.] 926 (1953).
Jap. Name. Nagaba-koiwazakura.
Japan, Prov. Totomi [Shizuoka Pref.],
Shuchi-gun, Totyuyama, 1949-10-29, Y.
Hayashi s.n. [TNS-VS-81076 Holotype].
Primula rhodotricha Nakai et F. Maek.
in Bull. Natl. Sci. Mus., Tokyo 29: 85, t. 3(1950).
Jap. Name. Chichibu-iwazakura.
Japan, Hondo, Prov. Musashi [Saitama
Pref.], Chichibu, in montis rupitus calcinis
montis Bukô, 1949-5-11, T. Nakai s.n. [TNSVS-112138 Holotype; TNS-VS-112139 
Isotype].
Primula rhodotricha Nakai et F. Maek. forma
leucantha Nakaiin Bull. Natl. Sci. Mus., Tokyo 29: 86 (1950).
Jap. Name. Shirobana-chichibu-iwazakura.
Japan, Hondo, Prov. Musashi [Saitama
Pref.], Chichibu, in montis rupitus calcinis
montis Bukô, 1949-5-11, T. Nakai s.n. [TNSVS-112137 Holotype].
BORAGINACEAE F. 256
Cynoglossum L.
Cynoglossum alpestre Ohwi
in Acta Phytotax. Geobot. 2: 150 (1933).
Taiwan, Taichushu, Mt. Nanko-taisan,
Bunakkei, 1933-7-?, J. Ohwi 4076 [TNS-VS43024Isotype].
Note. Holotype probably in KYO.
Macrotomia DC.
Macrotomia nepalensis Kitam.
in Peoples Nepal Himalaya 3: 423 (1957).
Nepal, Thaple Himal, 1953-7-30, S. Nakaos.n. [TNS-VS-263950 Isotype].
Note. Holotype in KYO (according to the orig-

107

inal description).
Mertensia RothMertensia pterocarpa (Turcz.) Tatew. et Ohwi
var. yezoensis Tatew. et Ohwi
in Acta Phytotax. Geobot. 2: 106 (1933).
Jap. Name. Ezo-ruriso.
Japan, Yezo [Hokkaido Pref.], Prov. Ishikari,
Sorachigun, Mt. Ashibetsu, 1915-7-30, H.
Koidzumi 74009 [TNS-VS-712373 Isotype].
Note. Holotype in KYO (Yonekura and Kajita,
2003–).
Omphalodes Mill.
Omphalodes laevisperma Nakai
in J. Jap. Bot. 23: 17 (1949).
Jap. Name. Echigo-ruriso.
Japan, Hondo, Prov. Etjigo [Niigata Pref.],
tractus Higashi-Kambara, Tunaki, 1948-5-30,
Y. Ikegami 10319 [TNS-VS-79399 Holotype]. Japan, Hondo, Prov. Etjigo [Niigata
Pref.], tractus Higashi-Kambara, Akatani,
1948-5-30, Y. Ikegami 10321 [TNS-VS-79400
Paratype]. Japan, Prov. Etjigo [Niigata
Pref.], tractus Higashi-Kambara, Tugawa,
1948-5-16, Y. Ikegami 10269 [TNS-VS79401Paratype].
Omphalodes prolifera Ohwi
in Bull. Natl. Sci. Mus., Tokyo n.s. 3: 98, fig. 1(1956).
Jap. Name. Hai-ruriso
Japan, Honshu, Owari Prov. [Aichi Pref.],
Shinanomachi, Mt. Jokojisan, 1955-6?-10 to17, K. Inami s.n. [TNS-VS-133753 Syntype]. Same locality, 1956-5-13, K. Inami s.n.
[TNS-VS-133754 Syntype].
Note. TNS-VS-133753 and TNS-VS-133754
are designated as types of fruit and flowers, respectively. There is some confusion about thecollection date of TNS-VS-133753 — “55. 1017 May” is written by hand on the originallabel, and the same date is shown on the secondary (herbarium) label. But, the latter wascorrected to “At. 17, 1955”. The collection
date of the type of fruit is “June 10–17, 1955”

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Atsushi Ebihara and Kyoko Miyashita

according to the original description.
Trigonotis Steven
Trigonotis coronata Ohwi
in Bot. Mag. (Tokyo) 44: 571 (1930).
Jap. Name. Koshiji-tabirako.
Japan, Hondo, Prov. Echigo [Niigata Pref.],
Mt. Naebayama, 1927-7-15, J. Ohwi s.n.
[TNS-VS-219386 Isotype].
Note. Holotype in KYO (according to the original description).
VERBENACEAE F. 258Callicarpa L.
Callicarpa dichotoma (Lour.) K.Koch forma
albifructus Okuyama
in Col. Ill. Wild Pl. Jap. 4: 137, pl. 351-2,
index 16 (1959).
Jap. Name. Shiratama-koshikibu.
Japan, Prov. Settsu [Hyogo Pref.], Nagaomura, Yamamoto, 1954-11-21, M. Togashi s.n.
[TNS-VS-110624 Holotype]. Japan, Chiba
Pref. Tsudanuma, 1957-9-27, S. Okuyama
20540 [TNS-VS-284546 Paratype].
Note. Cultivated plants. TNS-VS-284546 is
not directly cited in the original description,
but a specimen of the plant was used for theplate (photograph).
Callicarpa japonica Thunb. var. luxurians Rehder forma albifrons Konta
in Bull. Natl. Sci. Mus., Tokyo, Ser. B 31: 147,
figs. 15-6, 7 (2005).
Jap. Name. Shirobana-o-murasakishikibu.
Japan, Honshu, Pref. Shizuoka, Shimoda City,
Suzaki Peninsula, Tsumekizaki, Region III,
alt. ca. 30 m, 2005-7-2, F. Konta 24428-a[TNS-VS-749810 Holotype], 24428-b [TNSVS-749709 Isotype], 24428-c [TNS-VS749832 Isotype].
Note. “F. Konta 24428-a” is also cited as an
isotype, but is probably a misprint.

LABIATAE F. 260
Ajuga L.
Ajuga elatior Ohwi
in Bull. Natl. Sci. Mus., Tokyo n.s. 1: 4, fig, 3(1954).
Jap. Name. Oni-kiranso.
Japan, Ryukyu, ins. Tokunoshima, Kametoku,
1951-5-17, T. Amano 6434 [TNS-VS-90154
Holotype; TNS-VS-90153 Isotype].
Note. While the original label is not found in
the holotype, the photograph of the type in theoriginal description matches TNS-VS-90154.
Ajuga nipponensis Nakino forma nivea Hiyama
in J. Jap. Bot. 28: 94 (1953).
Jap. Name. Shirobana-junihitoe.
Japan, Hondo, Prov. Musashi [Tokyo Pref.],
1952-5-?, Y. Fukuhara 6434 [TNS-VS133460Holotype].
Note. A cultivated plant, originally collected at
Shirokane, Shiba, Tokyo.
Amethystanthus Nakai
Amethystanthus excisinflexus Nakaiin Bull. Natl. Sci. Mus., Tokyo 33: 21 (1953).
Jap. Name. Tairin-yamahakka.
Japan, Hondo, Prov. Etjigo [Niigata Pref.],
Higashi-kambara-gun, Ryokanose-mura,
Kanose, 1950-10-8, T. Nakai and N. Maruyama s.n. [TNS-VS-83730 Holotype; TNSVS-83729 & TNS-VS-83731 Isotype].
Note. A putative hybrid between Amethystanthus excisus Nakai (Rabdosia excisa(Maxim.) H.Hara) and A. inflexus Nakai (R.
inflexa (Thunb.) H.Hara).
Chelonopsis longipes Makino
in Bot. Mag. (Tokyo) 12: 57 (1898)
based on Chelonopsis moschata Miq. var.
longipes Makino, nom. nud.
in Bot. Mag. (Tokyo) 6: 54 (1892).
Jap. Name. Tani-jakoso.
Japan, Shikoku, Prov. Tosa [Kochi Pref.],
Tosa-yama[-go], 1892-10-?, T. Makino s.n.
[TNS-VS-10512Possible isosyntype].
Note. The collection date is 1892-10-6 in theoriginal description.

TNS type collections

Elsholtzia Willd.
Elsholtzia nipponica Ohwi
in Acta Phytotax. Geobot. 5: 180 (1935).
Jap. Name. Futobo-naginatakoju.
Japan, Hondo, Yamashiro [Kyoto Pref.], Kibune, 1935-10-14, G. Ikeo (J. Ohwi 9567)[TNS-VS-224649a and TNS-VS-224649b
Isotypes].
Note. Holotype in KYO (according to the original description). Ohwi’s collection number is
not indicated in these isotypes.
Isodon (Schrad. ex Benth.) Spach
Isodon kameba Okuyama ex Ohwi
in Bull. Natl. Sci. Mus., Tokyo 33: 85 (1953).
Jap. Name. Kameba-hikiokoshi.
Japan, Hondo, Prov. Shinano [Nagano
Pref.], Manzatôge, 1942-8-29, S. Okuyama
7175 [TNS-VS-78143 Holotype].
Isodon kameba Okuyama ex Ohwi var. latiofolius (Okuyama) Okuyama forma albescens
Okuyamain Bull. Natl. Sci. Mus., Tokyo n.s. 3: 16(1956).
Jap. Name. Shirobana-koshin-yamhakka.
Japan, Honshu, Prov. Shinano [Nagano
Pref.], Shimoina-gun, Ohshikamura, Jigokudani, 1930-9-21, H. Koidzumi 26960 [TNSVS-109907 Holotype], H. Koidzumi 26959[TNS-VS-109906 Isotype].
Isodon kameba Okuyama ex Ohwi var. latiofolius (Okuyama) Okuyama forma komaensis
Okuyamain Bull. Natl. Sci. Mus., Tokyo n.s. 3: 16(1956).
Jap. Name. Koma-yamahakka.
Japan, Honshu, Prov. Shinano [Nagano
Pref.], monte Koma, 1911-8-17, J. Nikai 2209
[TNS-VS-46647 Holotype].
Isodon kameba Okuyama ex Ohwi var. latiofolius (Okuyama) Okuyama forma ternatus
Okuyamain Bull. Natl. Sci. Mus., Tokyo n.s. 3: 16(1956).
Jap. Name. Kurumaba-koshin-yamahakka.

109

Japan, Honshu, Prov. Shinano [Nagano
Pref.], monte Shiomi, 1951-7-28, collector unknown (“ex Y. Hayashi”) [TNS-VS-92834 
Holotype].
Isodon trichocarpus (Maxim.) Kudo forma crythranthus Ikegami
in J. Jap. Bot. 32: 128 (1957).
Jap. Name. Akabana-hikiokoshi.
Japan, Niigata [Pref.], Minami-uwonuma,
Tsuchidaru, Yomogi-toge, alt. 1250 m, 1955-88, Y. Ikegami 29917 [TNS-VS-123854Holotype].
Isodon umbrosus (Maxim.) H.Hara var. latiofolius Okuyama
in J. Jap. Bot. 26: 319 (1951).
Jap. Name. Koshin-yamahakka.
Japan, Honshu, Prov. Kai [Yamanashi Pref.],
Sagashio-kosen, 1943-9-11, Y. Satake & S.
Okuyama 4969 [TNS-VS-78124 Holotype].
Japan, Honshu, Prov. Kai [Yamanashi Pref.],
Mt. Mitsu-toge, 1948-9-1, K. Hiyama s.n.
[TNS-VS-80036Paratype]. Japan, Honshu,
Prov. Shinano [Nagano Pref.], Kitazawatôge, 1938-8-5, S. Okuyama 4311 [TNS-VS72449a, TNS-VS-72449b and TNS-VS72449cParatypes].
Isodon ohwii Okuyama
in Bull. Natl. Sci. Mus., Tokyo n.s. 3: 16(1956).
Jap. Name. Kurobana-yamahakka.
Japan, Honshu, Prov. Echigo [Niigata Pref.],Oyu, 1949-9-?, J. Ohwi s.n. [TNS-VS-85008
Holotype; TNS-VS-85007 Isotype].
Note. A putative hybrid between I. inflexus(Thunb.) Kudô and I. trichocarpus (Maxim.)
Kudô.
Isodon suzukii Okuyama
in Bull. Natl. Sci. Mus., Tokyo n.s. 3: 17(1956).
Jap. Name. Kameba-yamahakka.
Japan, Honshu, Prov. Iwaki [Fukushima
Pref.], monte Kochiyama, 1950-9-20, S. Suzuki s.n. [TNS-VS-83948 Holotype].
Note. A putative hybrid between I. kameba
Okuyama ex Ohwi and I. inflexus (Thunb.)
Kudô.

110

Atsushi Ebihara and Kyoko Miyashita

Isodon togashii Okuyama
in Bull. Natl. Sci. Mus., Tokyo n.s. 3: 16(1956).
Jap. Name. Koshiji-hikiokoshi.
Japan, Honshu, Prov. Kodzuke [Gunma
Pref.], Shimidzu-toge, 1951-9-16, M. Togashi
s.n. [TNS-VS-109911 Holotype; TNS-VS109910, TNS-VS-109912, TNS-VS-109913,
TNS-VS-109914, TNS-VS-109915 and TNSVS-109916 Isotypes].
Note. A putative hybrid between I. excisinflexus Nakai (Honda) and I. kameba.
Mentha L.
Mentha afghanica Murata
in Acta Phytotax. Geobot. 17: 73 (1958).
Afghanistan, Eastern Prov., near Chaghasarai,
1955-7-26, S. Kitamura s.n. [TNS-VS249737Isotype].
Note. Holotype in KYO (according to the original description).
Micromeria Benth.
Micromeria nepalensis Kitam. et Murata
in Acta Phytotax. Geobot. 16: 3 (1955).
Nepal, Bangu Khola, alt. 3400 m, 1953-7-23,
S. Nakao s.n. [TNS-VS-263933 Possible isotype].
The collection date is different from that in theoriginal description (1953-7-11).
Holotype in KYO (according to the originaldescription).
Nepeta L.
Nepeta subsessilis Maxim. forma latifolia Nakai
in J. Jap. Bot. 23: 15 (1949).
Jap. Name. Hiroha-misogawaso.
Japan, Hondo, Prov. Rikutyû [Iwate Pref.],
Isawa, monte Yakeisidake, 1942-8-30, H.
Iwabuchi s.n. [TNS-VS-72730 Holotype].
Phlomis L.
Phlomis kawaguchii Murata
in Acta Phytotax. Geobot. 16: 15 (1955).

China, Tibet, circa Lhasa, Pulunka temple,
1914-8-20, E. Kawaguchi 79 [TNS-VS103284Holotype].
Prunellopsis Kudô
Prunellopsis prunelliformis (Maxim.) Kudô
forma lilacina Nakaiin Bull. Natl. Sci. Mus., Tokyo 29: 97 (1950).
Jap. Name. Usuiro-hakusan-utsubo.
Japan, Hondo, Prov. Shinano [Nagano
Pref.], Kita-azumi-gun, monte Happôzan,1948-8-4, Y. Hayashi s.n. [TNS-VS-81068 
Holotype].
Salvia L.
Salvia omerocalyx Hayata var. prostrata Satakein J. Jap. Bot. 17: 724 (1941).
Jap. Name. Hai-tamuraso.
Japan, Honshu, Prov. Etizen [Fukui Pref.],
Kurokawa, 1936-6-9, Z. Tashiro s.n. [TNSVS-56576 Holotype].
Salvia sakuensis Naruh. et Hihara
in J. Phytogeogr. Taxon. 49: 168 (2002).
Jap.Name. Saku-kibana-akigiri.
Japan, Nagano Pref., Saku-shi, Uchiyama, alt.
800 m, 2001-9-24, N. Naruhashi 1092401[TNS-VS-775650Isotype].
Scutellaria L.
Scutellaria indica L. var. japonica Franch. et
Sav. forma humilis Makino
in Bot. Mag. (Tokyo) 18: 46 (1904).
Jap. Name. Shisoba-tatsunami.
Japan, Prov. Tosa [Kochi Pref.], Akinokawamura, 1892-6-3, T. Makino s.n. [TNS-VS10753Isotype].
Note. Holotype in MAK (Yonekura and Kajita.
2003–). This is the basyonym of Scutellaria
laeteviolacea Koidz.
Scutellaria shikokiana Makino forma pubericaulis Ohwi
in Fl. Jap. (Ohwi) [Japanese ed.] 1000 (1953).
Jap. Name. Ke-miyama-namiki.
Japan, Kiushu, Osumi [Kagoshima Pref.],TNS type collections

Mt. Takakuma, 1913-7-24, Z. Tashiro s.n.
[TNS-VS-31057 Holotype].
Scutellaria transitra Makino
in Bot. Mag. (Tokyo) 18: 70 (1904).
Jap. Name. Yama-tatsunamiso.
Japan, Prov. Tosa [Kochi Pref.], Sakawa[mura], 1889-5-18, T. Makino s.n. [TNS-VS10758Isosyntype].
Stachys L.
Stachys takanoi Satakein J. Jap. Bot. 17: 721 (1941).
China, Manchukuo, Prov. Lung-chiang, Chêntung-hsien, Chiêu-t‘ai, 1940-7-15, T. Takano
68 [TNS-VS-13271 Holotype]. Prov. Lungchiang, Chên-tung-hsien, Ér-ching, 1940-7-26,
T. Takano 29 [TNS-VS-13272 Paratype],same locality, T. Takano 45 [TNS-VS13273Paratype].
Teucrium L.
Teucriumveronicoides
Maxim.
var.
brachytrichum Ohwi
in Fl. Jap. (Ohwi) [Japanese ed.] 998 (1953).
Jap. Name. Inu-nigakusa.
Japan, Hondo, Uzen [Yamagata Pref.],
Tachitanisawa, 1948-8-3, M. Kato s.n. [TNSVS-85735 Holotype].
Following specimens have not yet found in theherbarium, though they are cited as holotypes
and an isotype in the original description.

111

Enkianthus subsessilis (Miq.) Makino var. angustifolius H.Hatta et K.Yoshida
in Ann. Tsukuba Bot. Gard. 8: 11 (1989).
Jap. Name. Hosoba-abura-tsutsuji.
Japan, Gunma Pref., Onishi-machi, Nango,
Sanbagawa (cultivated at private garden of T.
Azami, Saitama Pref.), 1986-5-16, K. Yoshida& N. Kobayashi s.n. [TNS-VS-?Holotype].
Rhododendron sataense Nakaiin Bull. Natl. Sci. Mus., Tokyo 27: 33 (1949).
Jap. Name. Sata-tsutsuji.
Japan, Osumi [Kagoshima Pref.], Sata,
1948-4-16, T. Nakai & N. Maruyama s.n.
[TNS-VS-? Holotype].
Primula tosaensis Yatabe forma brachycarpa
H.Harain Iconogr. Pl. Asiae Orient. 1(4): 76 (1936).
Jap. Name. Shinano-kozakura.
Japan, Shinano [Nagano Pref.], Kosedozawa, 1928-7-15, H. Koidzumi 16639 [TNSVS-?Holotype].
Salvia japonica Thunb. forma albiflora Hiyama
in J. Jap. Bot. 17: 319 (1941).
Jap. Name. Shirobana-aki-no-tamuraso.
Japan, Sagami [Kanagawa Pref.], Arasiyama, 1938-10-9, K. Hiyama s.n. [TNS-VS-?
Holotype].
References
Yonekura, K. and T. Kajita. 2003–. BG Plants Name
Index (YList). http://bean.bio.chiba-u.jp/bgplants/
ylist_main.html.