Bull. Natl. Mus. Nat. Sci., Ser. B, 35(2), pp. 71–89, June 22, 2009

Notes on the Distribution of the Vandenboschia radicans Complex(Hymenophyllaceae) in Japan
Atsushi Ebihara
Department of Botany, National Museum of Nature and Science, Amakubo 4–1–1, Tsukuba, 305–0005 Japan
E-mail: ebihara@kahaku.go.jp

Abstract The distribution of eight species and four hybrids included in the Vandenboschia radicans complex within Japan is reviewed. Distribution maps for each taxon are provided based on
herbarium specimens, which are listed in order of municipality code.
Key words: Distribution, Hymenophyllaceae, Japan, Vandenboschia.

Introduction
Vandenboschia subgen. Vandenboschia (Hymenophyllaceae) distributed in world temperate/
subtropical regions encompasses a number ofmorphologically ill-defined species, and they aretentatively lumped under a species complex
named “Vandenboschia radicans complex”. As
for Japanese members of the complex, Ebihara
et al. (2005a) clarified their biological entities by
DNA/ploidy analyses, and Ebihara et al. (2009)
reexamined their taxonomic properties. Since thenumber of samples analyzed in these studies isstill limited, it is necessary to examine as many
herbarium specimens as possible to cover thewhole range of the Japanese archipelago and map
their localities for grasping the distribution ofeach newly defined taxon.
Materials and Methods
A total of 3589 sheets of herbarium specimens
tentatively identified as the Vandenboschia radicans complex were examined in the herbaria
KAG, KYO, MBK, RYU, TAI, TI and Yamaguchi
Museum (hereafter abbreviated as YMG, but unregistered in the Index Herbariorum). Identification was made with the knowledge on correlation
between morphology and genomic formula obtained from our previous studies (Ebihara et al.,2005a; 2009). In the present identification, twoamphidiploids, V. hokurikuensis and V. orientalis,

were embedded in V. stenosiphon and V.
quelpaertensis, respectively because it is difficult to discriminate amphidiploids from sterile
hybrids sharing the same genome combinations(but different genome dosage). As shown in thediscriminant analysis (Ebihara et al., 2009), someof the redefined taxa morphologically overlap
with each other. Therefore, descriptions and keys
do not always provide sufficient information for
their identification. Digital images of examined
specimens (only those deposited in TNS), tocompensate the difficulty, are accessible via online database (http://www.vspecimens.net/). All
the TNS specimens collected in Japan, and specimens of other herbaria with localities not overlapping with those of TNS specimens are citedwith their registration numbers and localities(municipalities). Their order generally follows
municipality code instituted by the Ministry ofInternal Affairs and Communications, Japan withexception of some islands.
Results and Discussion
Vandenboschia kalamocarpa (Hayata) Ebihara[Figs. 1, 13]
Distributed mostly on the Pacific side, around
35.5° N latitude line south. Known localities of
diploids are limited including Kii Peninsula,
Yaku Isl. and the Bonin islands, while triploids
show wider distribution. Frond length of diploids

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Atsushi Ebihara

usually under 10 cm, not more than 15 cm. All
the examined individuals of Okinawa Island andTaiwan are triploids. Rachis wings tend to bebroad and waved, but this character is not always
a reliable key for segregating this species from V.
stenosiphon. Distributed in continental China(Jiangxi Prov.) as well as Taiwan.
Vandenboschia nipponica (Nakai) Ebihara[Figs. 2, 14]An element of broad-leaved deciduous foreston the Sea of Japan side, ranging south to the
Kyushu mountain range. Diploids are recognizable by three-dimensionally arranged segments
and broadly triangular involucres. The lowest
pinnae are usually more or less shorter thanupper ones. Triploids which usually have subdeltoid fronds and less three-dimensionally arranged
segments are often difficult to segregate fromV. stenosiphon by morphological characters.
Ranging north to Shiribeshi Prov., Hokkaido
Pref., and south to Aira-cho, Kagoshima Pref.
There is some possibility of misidentification of
triploids as V. stenosiphon. Endemic to Japan,
and probably speciated from V. kalamocarpa
adapting to heavy-snow environment.
Vandenboschia birmanica (Bedd.) Ching[Figs. 3, 15]
A subtropical species, distributed only inwarm places on the Pacific side of the JapaneseArchipelago. The distribution largely overlaps
with that of V. kalamocarpa. The northern andeastern distribution limit is Shirakawa, Nishiizucho, Shizuoka Pref. There are a number of localities from southern Kyushu to Okinawa Isl., but
none known from Yaeyama Islands. Frond length
usually more than 30 cm, diameter of rhizome
approximately 1.5 mm — plants smaller thanthese sizes are probably the hybrid, V. quelpaertensis. There are at least two cytotypes,
diploid and triploid, but morphological difference
between them is unclear. Distributed in Taiwan,
China, India, Nepal, Bhutan, Myanmar (type locality), Thailand, Laos and Vietnam.

Vandenboschia subclathrata K. Iwats.
[Figs. 4, 16]
A local endemic species in Yaeyama Islands(Iriomote Isl. and Ishigaki Isl.). Many localities
are known in Iriomote Isl., while only one collection from the Ishigaki Isl.
Vandenboschia hokurikuensis Ebihara [Fig. 5]
A tetraploid species originated from hybridizations between V. kalamocarpa and V. nipponica.
So far known only from Japan.
Vandenboschia orientalis (C. Chr.) Ching[Fig. 6]
A tetraploid species originated from hybridizations between V. birmanica and V. kalamocarpa.
The distribution roughly corresponds to the area
where distributions of the two parental species
overlap. In most cases, segregation from V. 
quelpaertensis by morphological characters isdifficult, but Amami-oshima plants exceptionallyshow characteristic frond shape which is reminescent of V. subclathrata with widely spaced
segments. Endemic to Japan.
Vandenboschia miuraensis Ebihara [Fig. 7]
A tetraploid species originated from hybridizations between V. birmanica and V. nipponica. No
overlapping distribution of the two parents is
known so far.
Vandenboschia oshimensis (H. Christ) Ebihara[Fig. 8]A tetraploid species originated from hybridizations between V. birmanica and V. liukiuensis.
Distributed in Amami-oshima Isl. (type collection only) and Okinawa Island.
Vandenboschia stenosiphon (H. Christ)
Copel.
[Fig. 9]A sterile hybrid consisting of the genome of V.
kalamocarpa and V. nipponica. Widely and frequently distributed throughout Japan except in
the Ryukyu and Bonin Islands. Also distributedin Cheju-do Isl., South Korea.

Distribution of Vandenboschia in Japan

Vandenboschia quelpaertensis(Nakai)Ebihara[Fig. 10]A sterile hybrid consisting of the genome of V.
birmanica and V. kalamocarpa. Distributed mostly in coastal areas from 35.5°N latitude line south
to southern Ryukyus and to Bonin Islands. The
two plants*1 with 2n  108 chromosomes observed by Mitui (1968; 1973) are probably thishybrid. [*1 Yakushima Isl., Kagoshima Pref. (TNS-VS-321833) and Chichjima Isl., Bonin Islands, Tokyo Pref. ( TNS-VS-290706)] Also
distributed in Cheju-do Isl., South Korea, butrare.
Vandenboschia birmanica (Bedd.) Ching 
V. nipponica (Nakai) Ebihara[Fig. 11]A sterile hybrid consisting of the genome of V.
birmanica and V. nipponica. Known only from
Chiba and Kanagawa prefectures. As alreadynoted in Ebihara et al. (2005b), Miura Peninsula(Kanagawa Pref.) and Boso Peninsula (Chiba
Pref.) are the areas richest in genome combination of hybrids.
Vandenboschia birmanica (Bedd.) Ching 
V. kalamocarpa (Hayata) Ebihara  V. nipponica(Nakai) Ebihara[Fig. 12]A sterile hybrid consisting of genomes derived
from three parental species. Further localities
may have been overlooked since species recogni-

73tion based solely on morphological characters isoften unsuccessful.
Acknowledgement
The author thanks the curators of KAG, KYO,
MBK, RYU, TAI, TI and Yamaguchi Museum fortheir help in examining specimens.
References
Ebihara A., Ishikawa, H., Matsumoto, S., Lin, S.-J., Iwatsuki, K., Takamiya, M., Watano, M. and Ito, M. 2005a.
Nuclear DNA, chloroplast DNA, and ploidy analysis
clarified biological complexity of the Vandenboschia
radicans complex (Hymenophyllaceae) in Japan andadjacent areas. American Journal of Botany 92:
1535–1547.
Ebihara, A., Oka, T. and Matsumoto, S. 2005b. The diversity and reticulate evolution of the Vandenboschia radicans complex (Hymenophyllaceae) in Izu, Miura andBoso Peninsulas. Annals of the Tsukuba Botanical Garden (24): 17–25.
Ebihara, A., Matsumoto, S. and Ito, M. 2009. Taxonomyof the reticulate Vandenboschia radicans complex(Hymenophyllaceae) in Japan. Acta Phytotaxonomica
et Geobotanica 60: 27–41.
Mitui. K. 1968. Chromosomes and speciation in ferns.
Science Report of Tokyo Kyoiku Daigaku B 203:
285–333.
Mitui, K. 1973. A cytological survey on the pteridophytesof the Bonin Islands. Journal of Japanese Botany 48:
247–253.

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Atsushi Ebihara

Figs. 1–6. Distribution maps of the Vandenboschia radicans complex in Japan. Fig. 1. V. kalamocarpa. Fig. 2. V.
nipponica. Fig. 3. V. birmanica. Fig. 4. V. subclathrata. Fig. 5. V. orientalis. Fig. 6. V. hokurikuensis. Large
dot: localities of the specimens analyzed by molecular/ploidy analyses; small dot: localities of herbarium
specimens.

Distribution of Vandenboschia in Japan

Figs. 7–12. Distribution maps of the Vandenboschia radicans complex in Japan. Fig. 7. V. miuraensis. Fig. 8. V.
oshimensis. Fig. 9. V. stenosiphon. Fig. 10. V. quelpaertensis. Fig. 11. V. birmanica  V. nipponica. Fig.
12. V. birmanica  V. kalamocarpa  V. nipponica. Large dot: localities of the specimens analyzed by molecular/ploidy analyses; small dot: localities of herbarium specimens.

75

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Atsushi Ebihara

Figs. 13–16. Authentic specimens of four diploid species. Fig. 13. Vandenboschia kalamocarpa (Wakayama
Pref., TNS-VS-779034). Fig. 14. V. nipponica (Oita Pref., TNS-VS-779209). Fig. 15. V. birmanica (Shizuoka
Pref., TNS-VS-778646). Fig. 16. V. subclathrata (Iriomote Isl., Okinawa Pref., TNS-VS-778359). Scale 
5 cm.

Distribution of Vandenboschia in Japan
Appendix: Specimens examined
Vandenboschia kalamocarpa (Hayata) Ebihara (Figs. 1, 13)
Tokyo Pref. Kozu Isl./Kozushima-mura: TNS-VS802848; Mikura Isl./Mikurajima-mura: TNS-VS802956, TNS-VS-802957, TNS-VS-803020; Chichijima Isl./Ogasawara-mura: TNS-VS-52395, TNS-VS778664, TNS-VS-778665, TNS-VS-778666.
Shizuoka Pref. Fujieda-shi: TNS-VS-536194; Kamogun, Minamiizu-cho: KYO 00010399; Nishiizu-cho:
TNS-VS-449833,
TNS-VS-778642;
Tagata-gun,Nakaizu-cho: TNS-VS-457356, TNS-VS-536196;
Ogasa-gun, Daito-cho: TNS-VS-44338; Iwata-gun, Misakubo-cho: TNS-VS-1031905; Inasa-gun, Hosoe-cho:
TNS-VS-449829.
Mie Pref. Owase-shi: TNS-VS-130364, TNS-VS537847, TNS-VS-1031947, KYO 00010266, KYO
00010630, OSA 00111555; Hisai-shi: TNS-VS537913; Iinan-gun, Iitaka-cho: OSA 00093696; Takigun, Miyagawa-mura: TNS-VS-537846; Watarai-gun,Ochiyama-mura: TNS-VS-537833; Kitamuro-gun,Miyama-cho: KYO 00010263; Minamimuro-gun,
Kiho-cho: TNS-VS-779033, OSA s.n. (K. Seto 4471),
OSA s.n. (K. Seto 4473).
Kyoto Pref. Kasa-gun, Oe-cho: TNS-VS-194842.
Shiga Pref. Kanzaki-gun, Notogawa-cho: TI 00000574,
TI 00000575.
Osaka Pref. Izumi-shi: TNS-VS-778606, TNS-VS778607.
Nara Pref. Uda-gun, Muro-mura: KYO 00010367; Yoshino-gun, Yoshino-cho: TNS-VS-583792; Nishiyoshinomura: OSA 00051636; Kawakami-mura: TNS-VS779037.
Wakayama Pref. Kainan-shi: OSA 00131639; Hashimoto-shi: TNS-VS-937967; Tanabe-shi: TNS-VS-937936;
Shingu-shi: TNS-VS-779034, OSA 00131688; Nagagun, Kokawa-cho: OSA 00131641; Ito-gun, Katsuragicho: TNS-VS-937969, TNS-VS-937970; Arita-gun,Shimizu-cho: TNS-VS-937955, OSA 00131650;
Hidaka-gun, Ryujin-mura: TNS-VS-937963, OSA
00005004; Nishimuro-gun, Shirahama-cho: TNS-VS779184, TNS-VS-802937, TNS-VS-802938, TNS-VS802939, TNS-VS-802940, TNS-VS-802941, TNS-VS802943, OSA 00131655; Nakahechi-cho: OSA
00131661; Oto-mura: OSA 00131669; Hikigawacho: TNS-VS-937945; Susami-cho: OSA 00131686;
Kushimoto-cho: OSA 00131684; Higashimurogun, Nachikatsuura-cho: OSA 00047784, OSA
00131690; Kozagawa-cho: TNS-VS-1032029, OSA
00027053; Kumanogawa-cho: TNS-VS-937940, OSA
00131699, OSA 00131700; Hongu-cho: TNS-VS937938, TNS-VS-937960, OSA 00027873.
Tottori Pref. Iwami-gun, Kokufu-cho: TNS-VS-452644.
Shimane Pref. Hamada-shi: YMG 0049629; Hirata-shi:

77

TI 00000549; Yatsuka-gun, Yakumo-mura: TNS-VS778656, TNS-VS-9516603; Nita-gun, Yokota-cho:
OSA 00004998; Mino-gun, Mito-cho: YMG 0035709;
Kanoashi-gun, Nichihara-cho: YMG 0049215.
Hiroshima Pref. Saeki-gun, Itsukaichi-cho: TNS-VS9516546; Yamagata-gun, Kake-cho: TNS-VS-793393.
Yamaguchi Pref. Yamaguchi-shi: TNS-VS-50514;
Iwakuni-shi: YMG 0079581; Mine-shi: YMG 0040949;
Tsuno-gun, Kano-cho: YMG 0064779; Toyora-gun,Toyota-cho: TNS-VS-275298; Abu-gun, Ato-cho:
YMG 0023789.
Tokushima Pref. Myodo-gun, Sanagochi-son: KYO
00010477; Naka-gun, Aioi-cho: TNS-VS-779161;
Kito-son: TNS-VS-9516621; Kaifu-gun, Kainan-cho:
TNS-VS-779165, TNS-VS-779168, TNS-VS-779170,
TNS-VS-779171, TNS-VS-779172, TNS-VS-9516543,
TNS-VS-9516614.
Kagawa Pref. Okawa-gun, Shirotori-cho: TNS-VS9516643; Nagao-cho: TNS-VS-9516644; Shozu-gun,Uchinomi-cho: TNS-VS-9516629.
Ehime Pref. Matsuyama-shi: TNS-VS-525384; Niihamashi: KYO 00010486; Ozu-shi: TNS-VS-779167, TNSVS-793472; Shuso-gun, Tanbara-cho: TNS-VS-525387;
Kamiukena-gun, Mikawa-mura: KYO 00010478; Kitauwa-gun, Mima-cho: TNS-VS-793420; Matsunocho: TNS-VS-525392, TI 00000506; Hiyoshi-mura:
TNS-VS-525386, TNS-VS-525391; Minamiuwa-gun,Ipponmatsu-cho: KYO 00010481, KYO 00010485.
Kochi Pref. Nankoku-shi: MBK 0019994, OSA
00216532; Susaki-shi: KYO 00010464; Tosashimizushi: TNS-VS-779175, KYO 00010462; Aki-gun, Toyocho: TNS-VS-553418; Kami-gun, Monobe-son: TNSVS-553486; Tosa–gun, Tosa-cho: MBK 0052740;
Agawa-gun, Ikegawa-cho: TNS-VS-779177, KYO
00010463; Niyodogawa-cho: TNS-VS-63416, TNSVS-252872; Takaoka-gun, Higashitsuno-mura: TNSVS-553427; Sakawa-cho: TI 00000546; Ochi-cho:
TNS-VS-553487, TNS-VS-782817, TNS-VS-1031953,
TNS-VS-1031955.
Fukuoka Pref. Fukuoka-shi: TNS-VS590377, TNS-VS590378; Chikushino-shi: TNS-VS-590381; Chikushigun, Nakagawa-machi: TNS-VS-778629, TNS 9519705,
TNS 9519715; Kurate-gun, Miyata-machi: KYO
00010137; Tagawa-gun, Soeda-machi: KYO 00010215;
Chikujo-gun, Taihei-mura: TNS-VS-589856.
Saga Pref. Imari-shi: TNS-VS-536402; Takeo-shi: TNSVS-536399; Saga-gun, Fuji-cho: TNS-VS-536409;
Kanzaki-gun, Sefuri-mura: TNS-VS-536412; Fujitsugun, Tara-cho: TNS-VS-536406.
Nagasaki Pref. Nagasaki-shi: TNS-VS-519901, OSA
00046635; Kitatakaki-gun, Iimori-cho: TNS-VS480530; Iki Isl./Iki-gun, Gonora-cho: TNS-VS-793402.
Kumamoto Pref. Hitoyoshi-shi: TNS-VS-35816, KYO
00010140, KYO 00010168; Minamata-shi: YMG

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Atsushi Ebihara

0050013, KYO 00010110, KYO 00010112; Kamotogun, Kahoku-machi: TNS-VS-778338; Aso-gun, Ogunimachi: OSA s.n. (Z. Tashiro 1921-8-25); Kamimashiki-gun, Mifune-machi: TNS-VS-455305; Yabe-machi:
TNS-VS-455293; Yatsushiro-gun, Izumi-mura: TNSVS-802883; Kuma-gun, Nishiki-machi: KYO 00010180;
Mizukami-mura: KYO 00010138; Sagara-mura: KYO
00010166; Itsuki-mura: TNS-VS-762695, TNS-VS778520; Yamae-mura: TNS-VS-762671, TNS-VS778335, TNS-VS-802850.
Oita Pref. Saeki-shi: TNS-VS-456746; Kusu-gun, Kokonoe-machi: KYO 00010181; Shimoge-gun, Honyabakeimachi: TNS-VS-456749.
Miyazaki Pref. Miyazaki-shi: TNS-VS-459237; Miyakonojo-shi: TNS-VS-521720; Nichinan-shi: TNS-VS459234, TNS-VS-521776, TNS-VS-784673, TNS-VS9518558, TNS-VS-9518559; Saito-shi: TNS-VS521729; Ebino-shi: TNS-VS-77863, KYO 00010220;
Kitamorokata-gun,
Yamanokuchi-cho:
TNS-VS521726, TNS-VS-521732; Takazaki-cho: TNS-VS521731; Nishimorokata-gun, Takaharu-cho: TNS-VS521784; Suki-son: TNS-VS-521734; Higashimorokata-gun, Takaoka-cho: TI 00000543; Koyu-gun, Tsunocho: TNS-VS-459235; Higashiusuki-gun, Saigo-son:
TNS-VS-521738; Kitakata-cho: TNS-VS-521739; Shiiba-son: TNS-VS-521719; Nishiusuki-gun, Hinokagecho: KYO 00010126; Gokase-cho: TI 00042265.
Kagoshima Pref. Kanoya-shi: TNS-VS-453785, TNS-VS453786; Izumi-shi: KYO 00010116, KYO 00010117;
Okuchi-shi: TNS-VS-802926, KYO 00010188; Kasedashi: TNS-VS-453599; Ibusuki-gun, Kaimon-cho:
KYO 00010122; Hioki-gun, Higashiichiki-cho: TNSVS-453588; Satsuma-gun, Miyanojo-cho: TNS-VS453793; Shimokoshiki Isl./Shimokoshiki-son: TNSVS-802890, YMG 0050084; Isa-gun, Hishikaricho: TNS-VS-453585; Aira-gun, Kajiki-cho: KYO
00010199; Kamo-cho: TNS-VS-453790, TNS-VS1032006; Mizobe-cho: KYO 00010192; Yokogawacho: TNS-VS-453781, TNS-VS-453783; Kurino-cho:
TNS-VS-453612, TNS-VS-453614, TNS-VS-1032009,
KYO 00010196; Makizono-cho: TNS-VS-1031959,
KYO 00010211; Kirishima-cho: TNS-VS-453613;
So-gun, Takarabe-cho: TNS-VS-453624, TNS-VS453791, KYO 00010218; Shibushi-cho: TNS-VS453607; TNS-VS-453788; Kimotsuki-gun, Uchinouracho: TNS-VS-453590, TNS-VS-1032039, TNS-VS1032042; Koyama-cho: KYO 00010212; Onejime-cho:
TNS-VS-802899; Tashiro-cho: TNS-VS-276572, TNSVS-276573, TNS-VS-453604, TNS-VS-1031949,
TNS-VS-1031950, TNS-VS-1031982; Sata-cho: TNSVS-81922, TNS-VS-81923, TNS-VS-453603, TNSVS-453621; Yaku Isl./Kumage-gun, Kamiyaku-cho:
TNS-VS-368324, TNS-VS-453626, TNS-VS-453627,
NS-VS-775466, TNS-VS-778953, TNS-VS-778954,

TNS-VS-778955, TNS-VS-778956, TNS-VS-778961,
TNS-VS-778962, TNS-VS-778965, TNS-VS-802855,
TNS-VS-925912, TNS-VS-929501, TNS-VS-1031924,
TNS-VS-1032045, TNS-VS-1032046; Yaku Isl./Kumage-gun, Yaku-cho: TNS-VS-318048, TNS-VS453594, TNS-VS-453596, TNS-VS-453597, TNS-VS453897, TNS-VS-710322, TNS-VS-778948, TNS-VS778949, TNS-VS-778951, TNS-VS-778957, TNS-VS778960, TNS-VS-778964, TNS-VS-802862a/b, TNSVS-802863, TNS-VS-802864, TNS-VS-802876, TNSVS-802895, TNS-VS-802896, TNS-VS-1031918, TNSVS-1031919, TNS-VS-1031920, TNS-VS-1031937;
Tanegashima Isl./Nishinoomote-shi: TNS-VS-514287;
Kuroshima Isl./Kagoshima-gun Mishima-mura: TNSVS-453784.
Okinawa Pref. Naha-shi: TNS-VS-778360; Kunigamigun, Kunigami-son: TNS-VS-793441, KAG s.n. (S.
Hatusima 12851A), TI 00000589, TI 00000592; Higashi-son: TNS-VS-778362; Onna-son: TNS-VS518028.
Vandenboschia nipponica (Nakai) Ebihara (Figs. 2, 14)
Hokkaido Pref. Isoya-gun, Rankoshi-cho: TNS-VS9516590.
Aomori Pref. Aomori-shi: TNS-VS-524459, TNS-VS778927, TNS-VS-778935, TNS-VS-778937, TNS-VS778938, TNS-VS-778939, TI 00000234; Kamikitagun, Towadako-machi: TNS-VS-463347, TNS-VS524468, TNS-VS-710920, TNS-VS-778928, TNS-VS778936; Shimokita-gun, Ohata-machi: YMG 0031214.
Akita Pref. Oga-shi: TNS-VS-452382, TNS-VS-778931,
TNS-VS-778932, TNS-VS-778933, TNS-VS-778934:
Kitaakita-gun, Hinai-machi: TNS-VS-461404; Senboku-gun, Nishisenboku-machi: TNS-VS-548015.
Miyagi Pref. Toda-gun, Tajiri-cho: TNS-VS-9516641.
Yamagata Pref. Yonezawa-shi: TNS-VS-460993, KYO
00010230; Tsuruoka-shi: TNS-VS-460992, TNS-VS779189, TNS-VS-793419; Nishiokitama-gun, Iidemachi: TNS-VS-460985, KYO 00010231; Oguni-machi:
TNS-VS-303644, TNS-VS-459703, KYO 00010602; Higashitagawa-gun, Asahi-mura: TNS-VS-460706; Mogami-gun, Okura-mura: TNS-VS-384256; Akumi-gun,Yuza-machi: TNS-VS-448779; Hirata-machi: TNSVS-460998.
Fukushima Pref. Shirakawa-shi: TNS-VS-388510.
Chiba Pref. Kimitsu-shi: TNS-VS-779075, TNS-VS779083.
Niigata Pref. Nagaoka-shi: TNS-VS-778905; Murakamishi: TNS-VS-779205; Nishikanbara-gun, Yahiko-mura:
TNS-VS-779071; Maki-machi: TNS-VS-779062; Minamiuonuma-gun, Yuzawa-machi: TNS-VS-779066,
TNS-VS-927254; Nakakubiki-gun, Myokokogenmachi: TNS-VS-747790, TNS-VS-1031935; Iwafunegun, Asahi-mura: TNS-VS-793475.
Toyama Pref. Nakaniikawa-gun, Kamiichi-machi: TNS-Distribution of Vandenboschia in Japan
VS-778350; Tateyama-machi: KYO 00010600; Higashitonami-gun, Kamitaira-mura: TNS-VS-793464.
Ishikawa Pref. Kanazawa-shi: TI 00000240; Ishikawagun, Shiramine-mura: TNS-VS-540621, KYO
00010631.
Hyogo Pref. Sayo-gun, Nanko-cho: KYO 00010384.
Tottori Pref. Saihaku-gun, Daisen-cho: TNS-VS-388324,
TNS-VS-512953, TNS-VS-512956, KYO 00010620.
Kumamoto Pref. Kikuchi-shi: KYO 00010615, OSA
00008365; Aso-gun, Soyo-machi: TNS-VS-793453.
Oita Pref. Kusu-gun, Kusu-machi: TNS-VS-779209,
TNS-VS-779210, KYO 00010618; Hita-gun, Amagase-machi: TNS-VS-456735.
Kagoshima Pref. Aira-gun, Yokogawa-cho: TNS-VS453783.
Vandenboschia birmanica (Bedd.) Ching (Figs. 3, 15)
Shizuoka Pref. Kamo-gun, Nishiizu-cho: TNS-VS147168, TNS-VS-168022, TNS-VS-223704, TNS-VS331780, TNS-VS-778646, TNS-VS-1031971.
Wakayama Pref. Nishimuro-gun, Shirahama-cho: TNSVS-162628, KYO 00010565, TI 00042245.
Tokushima Pref. Kaifu-gun, Kainan-cho: TNS-VS779164.
Kumamoto Pref. Hitoyoshi-shi: TNS-VS-141834; Minamata-shi: TNS-VS-144661, KYO 00010524; Yatsushiro-gun, Sakamoto-mura: TNS-VS-293570.
Miyazaki Pref. Miyazaki-shi: TNS-VS-803076, KYO
00010523; Nichinan-shi: TNS-VS-680978, TNS-VS1031997.
Kagoshima Pref. Kagoshima-shi: TNS-VS-458392,
TNS-VS-778952, TNS-VS-793433; Ibusuki-shi: TNSVS-405007; Ibusuki-gun, Kiire-cho: TNS-VS-793442;
Kawanabe-gun, Chiran-cho: TNS-VS-9516571; Satsuma-gun, Kedoin-cho: TNS-VS-458393; Satsumacho: TNS-VS-777026; Isa-gun, Hishikari-cho: TNSVS-147404, TNS-VS-148279, TNS-VS-148280; Airagun, Aira-cho: KYO 00010525; So-gun, Shibushi-cho:
TNS-VS-35821, TNS-VS-102774, TNS-VS-187220;
Yaku Isl./Kumage-gun, Kamiyaku-cho: TNS-VS342344; Yaku Isl./Yaku-cho: KYO 00010026; Kuroshima Isl./Oshima-gun, Mishima-mura: KAG s.n. (S.
Sako 621), KAG s.n. (K. Maruno, 1980-11-3);
Nakanoshima Isl./Toshima-mura: TNS-VS-803077a/b/c; Kuchinoshima Isl./Toshima-mura: KAG s.n. (S.
Hatusima 16131); Takarajima Isl./Toshima-mura: KAG
s.n. (S. Sako 4489); Gaja Isl./Toshima-mura: KAG s.n.
(S. Sako & K. Kawanabe 2347); Amami-oshima
Isl./Naze-shi: TNS-VS-311716, TNS-VS-311723,
TNS-VS-518086, TNS-VS-518096, TNS-VS-518097,
TNS-VS-779191, TNS-VS-779192, TNS-VS-779193,
TNS-VS-779194, TNS-VS-779195, TNS-VS-779197,
TNS-VS-779198, TNS-VS-779200, TNS-VS-779201,
TNS-VS-312906; Amami-oshima Isl./Oshima-gun,

79

Uken-son: TNS-VS-28299, TNS-VS-140651; Sumiyoson: TNS-VS-311021, TNS-VS-311714, TNS-VS311715, TNS-VS-311717, TNS-VS-311721, TNS-VS311722, TNS-VS-453794, TNS-VS-465169, TNS-VS518085, TNS-VS-779196, TNS-VS-779199, TNS-VS1031897; Tokunoshima Isl./Amagi-cho.: RYU 34713.
Okinawa Pref. Nago-shi: TNS-VS-778358, RYU 29586,
RYU 28605, TI 00000193; Kunigami-gun, Kunigamison: TNS-VS-296632, TNS-VS-315138, TNS-VS316756, TNS-VS-775468, TNS-VS-778355, TNS-VS779185, TNS-VS-779186, TNS-VS-779187, TNS-VS779188, TNS-VS-1031978, KYO 00010001, RYU
22602, RYU 22603, RYU 42395, RYU 42396, RYU
17203, RYU 29598, RYU 40646, RYU 29581, RYU
6669, RYU 29606, RYU 29761, RYU 29578, RYU
42393, RYU 21913, RYU 17157, RYU 42398, RYU
42391; Onna-son: TNS-VS-216129; Ginoza-son: RYU
21958.
Vandenboschia subclathrata K.Iwats. (Figs. 4, 16)
Okinawa Pref. Ishigaki-shi: TNS-VS-803079; Yaeyamagun, Taketomi-cho: TNS-VS-103250, TNS-VS109026, TNS-VS-296621, TNS-VS-316737, TNS-VS316738, TNS-VS-316739, TNS-VS-316740, TNS-VS316741, TNS-VS-316742, TNS-VS-316743, TNS-VS368652, TNS-VS-368932, TNS-VS-369149, TNS-VS369697, TNS-VS-370494, TNS-VS-370511, TNS-VS370514, TNS-VS-370525, TNS-VS-466957, TNS-VS503478, TNS-VS-503479, TNS-VS-518027, TNS-VS551868, TNS-VS-551940, TNS-VS-561860, TNS-VS732047, TNS-VS-778352, TNS-VS-778353, TNS-VS778354, TNS-VS-778359, TNS-VS-936776, TNS-VS936777, TNS-VS-936830, TNS-VS-943079, TNS-VS1032051, TNS-VS-1032052, TNS-VS-1032053, TNSVS-1032054, TNS-VS-1032055, TNS-VS-9505717,
TNS-VS-9505809,
TNS-VS-9505810,
TNS-VS9506675, TNS-VS-9519977.
Vandenboschia hokurikuensis Ebihara (Fig. 5)
Saitama Pref. Chichibu-gun, Ogano-machi: TNS-VS779090.
Kanagawa Pref. Yokosuka-shi: TNS-VS-779232.
Niigata Pref. Niitsu-shi: TNS-VS-779060; Kamo-shi:
TNS-VS-779061.
Fukui Pref. Fukui-shi: TNS-VS-778339, TNS-VS778348; Yoshida-gun, Eiheiji-cho: TNS-VS-775470,
TNS-VS-778344, TNS-VS-778345, TNS-VS-778346,
TNS-VS-778347, TNS-VS-779049.
Shizuoka Pref. Shimoda-shi: TNS-VS-778640.
Aichi Pref. Nukata-gun, Nukata-cho: TNS-VS-778944.
Mie Pref. Iinan-gun, Iitaka-cho: TNS-VS-779053.
Kyoto Pref. Kyoto-shi: TNS-VS-779055.
Hyogo Pref. Hikami-gun, Aogaki-cho: TNS-VS-779059.
Nara Pref. Tenri-shi: TNS-VS-779035; Yoshino-gun,

80

Atsushi Ebihara

Nishiyoshino-mura: TNS-VS-779054.
Shimane Pref. Nima-gun, Yunotsu-machi: TNS-VS778653.
Okayama Pref. Katsuta-gun, Shoboku-cho: TNS-VS778365, TNS-VS-779057.
Hiroshima Pref. Hiba-gun, Takano-cho: TNS-VS779056.
Yamaguchi Pref. Yamaguchi-shi: TNS-VS-779058.
Vandenboschia orientalis (C.Chr.) Ching (Fig. 6)
Tokyo Pref. Hachijo Isl./Hachijo-machi: TNS-VS778667, TNS-VS-778668, TNS-VS-778669; Aogashima Isl./Aogashima-mura: TNS-VS-778911, TNSVS-778912.
Kanagawa Pref. Yokosuka-shi: TNS-VS-779234, TNSVS-779236.
Shizuoka Pref. Ito-shi: TNS-VS-778907, TNS-VS778908; Shimoda-shi: TNS-VS-778645, TNS-VS778909, TNS-VS-778910; Tagata-gun, Nakaizu-cho:
TNS-VS-778650.
Osaka Pref. Izumisano-shi: TNS-VS-778624; Minamikawachi-gun, Chihayaakasaka-mura: TNS-VS778613.
Tottori Pref. Tohaku-gun, Yurihama-cho: TNS-VS778654.
Shimane Pref. Yatsuka-gun, Shimane-cho: TNS-VS778662.
Oita Pref. Hita-gun, Amagase-machi: TNS-VS-779211.
Miyazaki Pref. Miyazaki-shi: TNS-VS-778633, TNSVS-778634.
Kagoshima Pref. Amami-oshima Isl./Naze-shi: TNS-VS311708, TNS-VS-453618, TNS-VS-779202, TNS-VS779204; Okuchi-shi: TNS-VS-778958; Amami-oshima
Isl./Oshima-gun, Sumiyo-son: TNS-VS-311713, TNSVS-1031898.
Okinawa Pref. Kunigami-gun, Kunigami-son: TNS-VS778356.
Vandenboschia miuraensis Ebihara (Fig. 7)
Kanagawa Pref. Yokohama-shi/Kamakura-shi: TNS-VS779179, TNS-VS-779181, TNS-VS-779182, TNS-VS779183; Yokosuka-shi: TNS-VS-777652; Fujisawa-shi:
TNS-VS-779180.
Vandenboschia oshimensis (H.Christ) Ebihara (Fig. 8)
Kagoshima Pref. Amami-oshima Isl.: KYO 00042390.
Okinawa Pref. Naha-shi: TNS-VS-778920, TNS-VS778921, RYU 28603; Kunigami-gun, Kunigami-son:
TNS-VS-778916, TNS-VS-778917, TNS-VS-778918,
TNS-VS-778919, KYO 00010038, KYO 00010042,
KYO 00010043, RYU 42392, RYU 17139, OSA
00111822, OSA 00155035.
Vandenboschia stenosiphon (H.Christ) Copel. (Fig. 9)

Hokkaido Pref. Hiyama-gun, Assabu-cho: TNS-VS9516588; Shimamaki-gun, Shimamaki-mura: TNS-VS802970, TNS-VS-802971, TNS-VS-802972; Iwanaigun, Kyowa-mura: TNS-VS-9516589; Furuu-gun,Kamoenai-mura: TNS-VS-9516599, KYO 00010224;
Yoichi-gun, Akaigawa-mura: TNS-VS-9516598; Esashigun, Nakatonbetsu-cho: TNS-VS-105904.
Aomori Pref. Aomori-shi: TNS-VS-524467, TI 00000218;
Hirosaki-shi: TNS-VS-524464; Higashitsugaru-gun,
Kanita-machi: TNS-VS-524465; Tairadate-mura: TNSVS-524457; Minmaya-mura: TNS-VS-366201, TNSVS-524461, TNS-VS-524462; Nishitsugaru-gun,
Fukaura-machi: TNS-VS-524458; Nakatsugaru-gun,Soma-mura: TNS-VS-524463; Kamikita-gun, Yokohama-machi: TNS-VS-524466; Shimokita-gun, Ohatamachi: TNS-VS-463346, TNS-VS-710917; Omamachi: TNS-VS-524460; Wakinosawa-mura: YMG
0031216.
Iwate Pref. Iwate-gun, Shizukuishi-cho: TNS-VS538453.
Miyagi Pref. Kesennuma-shi: TNS-VS-538452; Izumishi: TNS-VS-9516638; Igu-gun, Marumori-machi:
TNS-VS-9516637; Miyagi-gun, Matsushima-machi:
TNS-VS-50519, TNS-VS-9516639; Miyagi-machi:
TNS-VS-9516647; Kurihara-gun, Kurikoma-machi:
TNS-VS-286649; Hanayama-mura: TNS-VS-9516645;
Mono-gun, Naruse-cho: TNS-VS-9516646; Oshikagun, Onagawa-cho: TNS-VS-9516640.
Akita Pref. Akita-shi: TNS-VS-58801, TNS-VS-461406,
TNS-VS-548014; Honjo-shi: TNS-VS-778930; Kitaakita-gun, Takanosu-machi: TNS-VS-461401, TNSVS-461402, TNS-VS-461403; Hinai-machi: TNS-VS778929; Kamikoani-mura: TNS-VS-461400; Yamamoto-gun, Futatsui-machi: TNS-VS-461398, TNS-VS548016, TNS-VS-778926; Hachimori-machi: TNSVS-452378, TI 00042252; Yuri-gun, Kisakata-machi:
TNS-VS-452381, TNS-VS-724141; Chokai-machi:
TNS-VS-461405; Higashiyuri-machi: TNS-VS-452376;
Ochi-machi: TNS-VS-452379; Senboku-gun, Kyowamachi: TNS-VS-452380; Senboku-machi: TNS-VS461399; Ogachi-gun, Ogachi-machi: TNS-VS-452377.
Yamagata Pref. Yamagata-shi: TNS-VS-460989; Yonezawa-shi: TNS-VS-793440; Tsuruoka-shi: TNS-VS448794, TNS-VS-460988, TNS-VS-461002; Shinjoshi: TNS-VS-463793, TNS-VS-461000; Nishimurayama-gun, Nishikawa-machi: KYO 00010597; Mogamigun, Mamurogawa-machi: TNS-VS-460990, TNS-VS460991; Tozawa-mura: TNS-VS-461003; Nishiokitama-gun, Oguni-machi: TNS-VS-460705; Higashitagawa-gun, Haguro-machi: TNS-VS-460986; Asahimura: TNS-VS-461001, TNS-VS-793414, TI 00000226;
Nishitagawa-gun, Atsumi-machi: TNS-VS-460987,
TNS-VS-460994, TNS-VS-460997, TNS-VS-793415,
TNS-VS-793416, TNS-VS-793417, TNS-VS-793418;

Distribution of Vandenboschia in Japan
Akumi-gun, Yuza-machi: TNS-VS-460999; Yawatamachi: TNS-VS-460996, TNS-VS-460995.
Fukushima Pref. Fukushima-shi: TNS-VS-23864;
Aizuwakamatsu-shi: TNS-VS-457142; Iwaki-shi: TNSVS-253963, TNS-VS-341756, TNS-VS-457145, TNSVS-457146, TNS-VS-457147, TNS-VS-600859, TNSVS-793382, TNS-VS-793383, TNS-VS-793384, TNSVS-793386, TNS-VS-793413, TNS-VS-793469, TNSVS-793470; Minamiaizu-gun, Shimogo-machi: TNSVS-457151; Tadami-machi: TNS-VS-457143; Yamagun, Atsushiokano-mura: TNS-VS-457149, TNS-VS457150; Kitashiobara-mura: TNS-VS-457144; Yamato-machi: TNS-VS-793385; Onuma-gun, Aizutakadamachi: TNS-VS-457148, TNS-VS-457152.
Ibaraki Pref. Takahagi-shi: TNS-VS-454430; Kitaibarakishi: TNS-VS-454431; Kasama-shi: TNS-VS-454339;
Tsukuba-shi: KYO 00010430; Higashiibaraki-gun,
Katsura-mura: TNS-VS-454432; Kuji-gun, Satomimura: TNS-VS-454429; Niihari-gun, Yasato-machi:
TNS-VS-454338; Chiyoda-mura: TNS-VS-454428,
TNS-VS-454433.
Tochigi Pref. Utsunomiya-shi: TNS-VS-454199, TNSVS-212786; Haga-gun, Mashiko-machi: TI 00042271;
Motegi-machi: TNS-VS-454198; Nasu-gun, Batomachi: TNS-VS-449293.
Gunma Pref. Numata-shi: TNS-VS-124141; Tone-gun,
Minakami-machi: TNS-VS-793439.
Saitama Pref. Hanno-shi: TNS-VS-577976; Iruma-gun,Ogose-machi: TNS-VS-577281; Naguri-mura: TNSVS-577974,
TNS-VS-802973,
TNS-VS-802974,
TNS-VS-802975; Hiki-gun, Ogawa-machi: TNS-VS293112; Chichibu-gun, Ogano-machi: TNS-VS-779081;
Arakawa-mura: TNS-VS-577278, TNS-VS-577280; Higashichichibu-mura: TNS-VS-577279; Kodama-gun,
Kodama-machi: TNS-VS-577973; Kamikawa-mura:
TNS-VS-577975.
Chiba Pref. Kamogawa-shi: TNS-VS-936764; Futtsu-shi:
TNS-VS-540086, TNS-VS-540113, TNS-VS-793387,
TNS-VS-803043; Awa-gun, Tomiyama-machi: TNSVS-540081; Chikura-machi: TNS-VS-401365; Amatsukominato-machi: TNS-VS-389572, TNS-VS-803034,
TNS-VS-803038, TNS-VS-803042.
Tokyo Pref. Hachioji-shi: TNS-VS-142389, TNS-VS147879, TNS-VS-247872, TNS-VS-523522, TNS-VS441601, TNS-VS-459313, TNS-VS-779085, TNS-VS803017; Ome-shi: TNS-VS-793134, TNS-VS-459312;
Nishitama-gun, Itsukaichi-machi: TNS-VS-793467;
Okutama-machi: TNS-VS-802954; Hachijo Isl./Hachijo-machi: TNS-VS-81227, TNS-VS-266477.
Kanagawa Pref. Yokosuka-shi: TNS-VS-779231, TNSVS-779233; Kamakura-shi: TNS-VS-779223, TNSVS-779224, TNS-VS-779225, TNS-VS-779226, TNSVS-803004; Zushi-shi: TNS-VS-561708, TNS-VS647240, TNS-VS-779079, TNS-VS-779089, TNS-VS-

81

793127, TNS-VS-802979, TNS-VS-802982, TNS-VS802984, TNS-VS-802985, TNS-VS-802986, TNS-VS802987; Ashigarashimo-gun, Hakone-machi: TNS-VS1405, TNS-VS-43236, TNS-VS-115293, TNS-VS561716, TNS-VS-779219, TNS-VS-779220, TNSVS-779221, TNS-VS-779222, TNS-VS-9516538; Yugawara-machi: TNS-VS-314968, TNS-VS-779214,
TNS-VS-779215, TNS-VS-779216, TNS-VS-779217,
TNS-VS-779218,
TNS-VS-1031908,
TNS-VS1031952, TNS-VS-9516539, TNS-VS-9516540.
Niigata Pref. Nagaoka-shi: TNS-VS-778906; Kashiwazaki-shi: TNS-VS-9516533, TNS-VS-9516560, TNSVS-9516557; Shibata-shi: TNS-VS-925925, KYO
00010613; Niitsu-shi: TNS-VS-464870, TNS-VS779063; Itoigawa-shi: TNS-VS-779067, TNS-VS9516534, TNS-VS-9516552; Nakakanbara-gun, Muramatsu-machi: TNS-VS-321758, TNS-VS-779072,
TNS-VS-9516551; Nishikanbara-gun, Iwamuro-mura:
TNS-VS-927252; Minamikanbara-gun, Tagami-mura:
TNS-VS-254120; Shitada-mura: TNS-VS-463791, TI
00042267; Higashikanbara-gun, Tsugawa-machi:
TNS-VS-793474, TNS-VS-9516558; Kanose-machi:
TNS-VS-779070; Santo-gun, Teradomari-machi: TNSVS-9516597; Kitauonuma-gun, Yunotani-mura: TNSVS-80484; Irihirose-mura: TNS-VS-1031941, TNSVS-9516553; Minamiuonuma-gun, Yuzawa-machi:
TNS-VS-384590, TNS-VS-779064, TNS-VS-779065;
Shiozawa-machi: TNS-VS-9516554; Muika-machi:
TNS-VS-779068, TNS-VS-779069; Kariwa-gun,Takayanagi-machi:
TNS-VS-793437,
TNS-VS1031930; Nakakubiki-gun, Kakizaki-machi: TNSVS-102776, TNS-VS-793436, TNS-VS-1031940;
Nishikubiki-gun, No-machi: TNS-VS-9516559, KYO
00010601; Omi-machi: TNS-VS-136496, TNS-VS1031931, TNS-VS-9516555; Iwafune-gun, Arakawamachi: TNS-VS-9516569; Asahi-mura: TNS-VS9516556; Sanpoku-machi: TNS-VS-9516535; Sado
Isl./Ryotsu-shi: TNS-VS-9516568; Joetsu-shi: TNSVS-793435, TNS-VS-9516532; Sado Isl./Sado-gun,
Aikawa-machi: TNS-VS-9516567; Sado Isl./Manomachi: TNS-VS-9516596; Sado Isl./Ogi-machi: TNSVS-9516595, KYO 00010447; Sado Isl./Akadomarimura: TNS-VS-1031922.
Toyama Pref. Himi-shi: TNS-VS-465003; Namerikawashi: TNS-VS-533271, TNS-VS-9506772, TNS-VS9506863; Kurobe-shi: KYO 00010604; Oyabe-shi:
TNS-VS-533269; Kaminiikawa-gun, Oyama-machi:
TNS-VS-533272, RYU 2095; Nakaniikawa-gun,Kamiichi-machi: TNS-VS-152485, TNS-VS-153927,
TNS-VS-533274; Shimoniikawa-gun, Unazuki-machi:
TNS-VS-793428, TNS-VS-793429, TNS-VS-793430;
Asahi-machi: TNS-VS-533273, TNS-VS-533278,
KYO 00010398; Nei-gun, Yatsuo-machi: TNS-VS533270; Higashitonami-gun, Toga-mura: TNS-VS-

82

Atsushi Ebihara

778349.
Ishikawa Pref. Kanazawa-shi: TNS-VS-136493, TNSVS-540616, TNS-VS-540626, TNS-VS-925903, KYO
00010606; Nanao-shi: TNS-VS-540630, TNS-VS540632; Komatsu-shi: TNS-VS-540623, TNS-VS540624, TNS-VS-568246, TNS-VS-936838; Wajimashi: TNS-VS-540453, TNS-VS-540483; Suzu-shi:
TNS-VS-540457, TNS-VS-540484; Kaga-shi: TNSVS-309802, TNS-VS-540615; Hakui-shi: TNS-VS540631; Enuma-gun, Yamanaka-machi: TNS-VS540618, TNS-VS-540619, TNS-VS-540620, TNS-VS1031913, KYO 00010424; Nomi-gun, Tatsunokuchimachi: TNS-VS-540625; Ishikawa-gun, Tsurugimachi: TNS-VS-568245; Kawachi-mura: TNS-VS540617; Yoshinodani-mura: RYU 40598; Oguchimura: TNS-VS-540622; Kahoku-gun, Tsubata-machi:
TNS-VS-540627; Hakui-gun, Togi-machi: TNS-VS540452, TNS-VS-540628; Shika-machi: TNS-VS540451, TNS-VS-568242, TNS-VS-568243; Oshimizumachi: TNS-VS-540482; Kashima-gun, Kashimamachi: TNS-VS-540629; Fugeshi-gun, Monzen-machi:
TNS-VS-540458, TNS-VS-568856, TNS-VS-568857;
Yanagida-mura: TNS-VS-540455; Hosu-gun, Anamizumachi: TNS-VS-540456; Noto-cho: TNS-VS-540454.
Fukui Pref. Fukui-shi: TNS-VS-453198, TNS-VS453207; Tsuruga-shi: TNS-VS-453191; Takefu-shi:
TNS-VS-453184, TNS-VS-453194; Obama-shi: TNSVS-453187; Ono-shi: TNS-VS-453183, TNS-VS453196, TNS-VS-453200, TNS-VS-453201, TNS-VS592431, TNS-VS-745198, TNS-VS-1031925, TNSVS-1031932, TNS-VS-1031936, TNS-VS-1032013;
Katsuyama-shi: TNS-VS-453192, TNS-VS-453193,
TNS-VS-453197, TNS-VS-592434, TNS-VS-592436,
TNS-VS-592468, TNS-VS-778340, TNS-VS-778341,
TNS-VS-778342, TNS-VS-778343, TNS-VS-1031963,
TNS-VS-9516585, TNS-VS-9516586, KYO 00010393,
KYO 00010625; Asuwa-gun, Miyama-cho: TNS-VS453195, TNS-VS-453199; Yoshida-gun, Matsuokacho: TNS-VS-453203; Ono-gun, Izumi-mura: TNSVS-453205, TNS-VS-453206; Sakai-gun, Kanazu-cho:
TNS-VS-453185, TNS-VS-592467; Maruoka-cho:
TNS-VS-453204; Nanjo-gun, Imajo-cho: TNS-VS453209; Kono-mura: TNS-VS-453202, TNS-VS592432; Nyuu-gun, Asahi-cho: TNS-VS-453210; Mikata-gun, Mikata-cho: TNS-VS-453182, TNS-VS453208; Onyuu-gun, Kaminaka-cho: TNS-VS-453186;
Natasho-mura: TNS-VS-453181, TNS-VS-453190;
Oi-gun, Takahama-cho: TNS-VS-453188Oi-cho: TNSVS-453189.
Nagano Pref. Shimoina-gun, Achi-mura: TNS-VS421342; Shimominochi-gun, Sakae-mura: TNS-VS779073.
Gifu Pref. Tajimi-shi: TNS-VS-464109; Mino-shi: TNSVS-9516594; Gujo-shi: TNS-VS-778942; Fuwa-gun,

Tarui-cho: TNS-VS-802916; Sekigahara-cho: TNSVS-793431; Ibi-gun, Kuze-mura: TNS-VS-745436,
TNS-VS-9516580; Motosu-gun, Neo-mura: TNS-VS802845, TNS-VS-9516583; Yamagata-gun, Miyamacho: TNS-VS-9516581; Mugi-gun, Horado-mura:
TNS-VS-462930, TNS-VS-775473; Mugegawa-cho:
TNS-VS-462931, TNS-VS-778941; Gujo-gun, Hachiman-cho: TNS-VS-9516578, TNS-VS-9516579, TNSVS-9516584; Takasu-mura: TNS-VS-9516593; Minami-mura: TNS-VS-259148, TNS-VS-399051, TNSVS-9516591, KYO 00010622; Kamo-gun, Hichisocho: TNS-VS-802923, TNS-VS-9516582; Ena-gun,Kushihara-mura:
TNS-VS-462932;
Mashita-gun,
Kanayama-cho: TNS-VS-793444; Yoshiki-gun, Miyagawa-mura: TNS-VS-9516592.
Shizuoka Pref. Shizuoka-shi: TNS-VS-449821, TNSVS-536182; Hamamatsu-shi: TNS-VS-449826, TNSVS-462632; Numazu-shi: TNS-VS-545460; Shimizushi: TNS-VS-536189; Atami-shi: TNS-VS-779213;
Mishima-shi: TNS-VS-311219, TNS-VS-536233; Itoshi: TNS-VS-460365, TNS-VS-536238, TNS-VS802992; Fuji-shi: TNS-VS-449820, TNS-VS-449840,
TNS-VS-536202, TNS-VS-925895; Kakegawa-shi:
TNS-VS-925911; Tenryu-shi: TNS-VS-83862, TNSVS-449811; Shimoda-shi: TNS-VS-536225, TNS-VS778639, TNS-VS-778649; Susono-shi: TNS-VS536234, TNS-VS-925897; Kamo-gun, Kawazu-cho:
TNS-VS-332537, TNS-VS-449838, TNS-VS-449845,
TNS-VS-778648, TNS-VS-803009, TNS-VS-929513;
Minamiizu-cho: TNS-VS-311884; Matsuzaki-cho: TNSVS-449814, TNS-VS-925910; Nishiizu-cho: TNS-VS449837, TNS-VS-779131; Tagata-gun, Shuzenji-cho:
TNS-VS-536206; Amagiyugashima-cho: TNS-VS324793, TNS-VS-449816, TNS-VS-449846, TNS-VS802875, TNS-VS-802877, TNS-VS-925908, KYO
00010442; Tagata-gun, Nakaizu-cho: TNS-VS-449835,
TNS-VS-1032000; Ihara-gun, Fujikawa-cho: TNS-VS545459, TNS-VS-745195; Shida-gun, Okabe-cho:
TNS-VS-536190; Haibara-gun, Kawane-cho: TNSVS-545457; Nakakawane-cho: TNS-VS-463489, TNSVS-536205; Ogasa-gun, Hamaoka-cho: TNS-VS536191; Shuchi-gun, Mori-machi: TNS-VS-449822;
Iwata-gun, Tatsuyama-mura: TNS-VS-449813, TNSVS-449823; Sakuma-cho: TNS-VS-449809, TNS-VS449830, TNS-VS-535523, TNS-VS-536183, TNS-VS536201; Misakubo-cho: TNS-VS-1031962; Inasa-gun,
Inasa-cho: TNS-VS-449827, TNS-VS-536185; Mikkabicho: TNS-VS-536198, TNS-VS-9516450.
Aichi Pref. Toyohashi-shi: TNS-VS-1031961, TNS-VS9516549; Okazaki-shi: TNS-VS-463527, TNS-VS9516566; Toyota-shi: TNS-VS-463529, TNS-VS465147; Inuyama-shi: TNS-VS-465148; Shinshiro-shi:
TNS-VS-463611, TNS-VS-9516547; Chita-gun, Minamichita-cho: TNS-VS-9516550, TNS-VS-9516625;

Distribution of Vandenboschia in Japan
Hazu-gun, Kira-cho: TNS-VS-9516631; Nukata-gun,
Nukata-cho: TNS-VS-463526, TNS-VS-778943, TNSVS-778945, TNS-VS-778946, TNS-VS-9516561,
TNS-VS-9516565, TNS-VS-9516628; Higashikamogun, Asuke-cho: TNS-VS-463530, TNS-VS-9516562,
TNS-VS-9516624; Kitashitara-gun, Shitara-cho: TNSVS-9516634; Toyone-mura: KYO 00010426; Tomiyama-mura: TNS-VS-9516564, TNS-VS-9516632; Minamishitara-gun, Horai-cho: TNS-VS-45782, TNSVS-463528, TNS-VS-802912, TNS-VS-802913, TNSVS-1031943, TNS-VS-9516548, TNS-VS-9516563,
TNS-VS-9516577,
TNS-VS-9516633,
TNS-VS9516635, TNS-VS-9516636, KYO 00010610, KYO
00010428; Hoi-gun, Otowa-cho: TNS-VS-9516626.
Mie Pref. Ise-shi: TNS-VS-1031981, TNS-VS-1032017;
Nabari-shi: TNS-VS-247311, TNS-VS-537841; Owaseshi: TNS-VS-537832, TNS-VS-716318, TNS-VS803007, TNS-VS-1031917, TNS-VS-1031944, TNSVS-1031951, YMG 0126486; Toba-shi: TNS-VS537834; Kumano-shi: TNS-VS-247300, TNS-VS537843, KYO 00010272; Iga-shi: KYO 00010278;
Kuwana-gun, Tado-cho: TNS-VS-537839, TNS-VS803006; Age-gun, Geino-cho: TNS-VS-537919; Ichishigun, Misugi-mura: TNS-VS-63415; Iinan-gun, Iitakacho: TNS-VS-537844; Taki-gun, Miyagawa-mura:
TNS-VS-779044, KYO 00010261; Watarai-gun, Tamaki-cho: KYO 00010245; Nansei-cho: TNS-VS-537840;
Watarai-cho: TNS-VS-537917; Kitamuro-gun, Miyama-cho: TNS-VS-343889, TNS-VS-537916, KYO
00010250, KYO 00010264; Minamimuro-gun, Mihama-cho: YMG 0126491; Kiho-cho: TNS-VS537831, TNS-VS-568244.
Shiga Pref. Otsu-shi: TNS-VS-17706, KYO 00010375,
KYO 00010374; Hikone-shi: TNS-VS-585585; Sakata-gun, Ibuki-cho: TNS-VS-585586; Ika-gun, Nishiazai-cho: TNS-VS-585584.
Kyoto Pref. Kyoto-shi: TNS-VS-1407, TNS-VS-277535,
TNS-VS-1031939, KYO 00010363, KYO 00010376;
Kameoka-shi: TNS-VS-194843; Nagaokakyo-shi: TNSVS-277536; Yawata-shi: KYO 00010352; Otokunigun, Oyamazaki-mura: TNS-VS-277540; Kitakuwadagun, Miyama-cho: TNS-VS-247313, TNS-VS-802846;
Funai-gun, Wachi-cho: KYO 00010386; Kasa-gun,Oe-cho: TNS-VS-678726, TNS-VS-678727, KYO
00010355.
Osaka Pref. Takatsuki-shi: TNS-VS-247044; Izumisanoshi: TNS-VS-778609, TNS-VS-778610, TNS-VS778611; Kawachinagano-shi: TNS-VS-247333, TNSVS-775472, TNS-VS-778608, TNS-VS-778616, TNSVS-778617, TNS-VS-778618, TNS-VS-778625; Matsubara-shi: TNS-VS-379053; Izumi-shi: TNS-VS778604, TNS-VS-778605, TNS-VS-778612, TNS-VS778615; Mino-shi: TNS-VS-584448; Minamikawachigun, Kanan-cho: TNS-VS-778614.

83

Hyogo Pref. Kobe-shi: TNS-VS-554756, TNS-VS554757, TNS-VS-9516609, KYO 00010349; Nishinomiya-shi: KYO 00010377; Sasayama-shi: TNSVS-778619, KYO 00010350; Taka-gun, Kami-cho:
TNS-VS-584447, TNS-VS-584570; Kasai-gun, Hojocho: TNS-VS-554760; Shikama-gun, Yumesaki-cho:
TNS-VS-554759; Kanzaki-gun, Kanzaki-cho: KYO
00010388; Sayo-gun, Sayo-cho: TNS-VS-554755;
Kozuki-cho: TNS-VS-541004; Nanko-cho: TNS-VS141936, TNS-VS-584609, TNS-VS-802882; Shisogun, Yamasaki-cho: TNS-VS-584446; Yasutomi-cho:
TNS-VS-379054; Ichinomiya-cho: KYO 00010354;
Haga-cho: TNS-VS-584445, KYO 00010385; Mikatagun, Muraoka-cho: TNS-VS-584569, TNS-VS-778620,
TNS-VS-778621; Mikata-gun, Hamasaka-cho: TNSVS-778622, TNS-VS-778623; Onsen-cho: TNS-VS554758; Yabu-gun, Yabu-cho: TNS-VS-584444; Oyacho: TNS-VS-584571; Hikami-gun, Hikami-cho: KYO
00010379, KYO 00010360; Taki-gun, Tannan-cho:
TNS-VS-247312, KYO 00010371.
Nara Pref. Nara-shi: TNS-VS-583708, TNS-VS-583711,
TNS-VS-779041; Sakurai-shi: TNS-VS-583709; Udagun, Haibara-cho: TNS-VS-779047; Muro-mura:
TNS-VS-132713, TNS-VS-779038, TNS-VS-779042;
Yoshino-gun, Yoshino-cho: TNS-VS-779048, TNS-VS779203; Shimoichi-cho: TNS-VS-583707; Totsukawamura: TNS-VS-583712, TNS-VS-583713, TNS-VS779043,
TNS-VS-793443;
Shimokitayama-mura:
KYO 00010365; Kamikitayama-mura: TNS-VS779039; Kawakami-mura: TNS-VS-779036, TNS-VS779046; Higashiyoshino-mura: TNS-VS-779040, KYO
00010357.
Wakayama Pref. Kainan-shi: TNS-VS-937964; Nagagun, Momoyama-cho: TNS-VS-937968; Ito-guinKoya-cho: KYO 00010265; Arita-gun, Kibi-cho: TNSVS-937966; Shimizu-cho: TNS-VS-42978, TNS-VS937965, TI 00000210; Nishimuro-gun, Nakahechi-cho:
TNS-VS-937937; Kamitonda-cho: KYO 00010251;
Hikigawa-cho: TNS-VS-937947; Susami-cho: KYO
00010271; Higashimuro-gun, Nachikatsuura-cho:
TNS-VS-37586, TNS-VS-80483, TNS-VS-217096,
TNS-VS-937941, TNS-VS-937950, TNS-VS-1032021;
Kozagawa-cho: TNS-VS-1031926, TNS-VS-1031933,
TNS-VS-1031946, TNS-VS-1032022, YMG 0042719;
Kumanogawa-cho: TNS-VS-937957, TNS-VS-937958;
Honguu-cho: TNS-VS-802930, KYO 00010246, KYO
00010252; Kitayama-mura: TNS-VS-937959.
Tottori Pref. Tottori-shi: TNS-VS-452643, TNS-VS778658; Yonago-shi: TNS-VS-782819; Iwami-gun,
Kokufu-cho: TNS-VS-452642; Fukube-son: KYO
00010239; Yazu-gun, Hatto-cho: TNS-VS-452645;
Wakasa-cho: KYO 00010241; Mochigase-cho: TI
00042277; Chizu-cho: TNS-VS-452715; Tohaku-gun,Togo-cho: TNS-VS-452641; Misasa-cho: TNS-VS-

84

Atsushi Ebihara

144728, TNS-VS-452640, TNS-VS-452714, TNS-VS1031929; Kotora-cho: TNS-VS-778659; Saihaku-gun,
Saihaku-cho: TNS-VS-462672; Daisen-cho: TNS-VS265341, TNS-VS-452646; Hino-gun, Nichinan-cho:
TNS-VS-462671; Hino-cho: TNS-VS-452639; Mizokuchi-cho: TNS-VS-462670.
Shimane Pref. Matsue-shi: TNS-VS-9516602; Izumoshi: YMG 0050667Oda-shi: KYO 00010235, KYO
00010624; Yasugi-shi: YMG 0049298; Hirata-shi:
TNS-VS-330898, TNS-VS-384239; Yatsuka-gun, Shimane-cho: TNS-VS-9516604; Mihonoseki-cho: TNSVS-9516601; Shinji-cho: TNS-VS-9516541; Nogigun, Hirose-machi: TNS-VS-778655, TNS-VS778661; Nita-gun, Yokota-cho: TNS-VS-39380, TNSVS-9516605; Iishi-gun, Mitoya-cho: YMG 0049373;
Akagi-cho: TNS-VS-38891, KYO 00010626; Hikawagun, Taisha-machi: TNS-VS-9516610; Ochi-gun,
Daiwa-mura: TNS-VS-745593; Hasumi-mura: TNSVS-523951; Mino-gun, Mito-cho: YMG 0128490;
Kanoashi-gun, Tsuwano-cho: YMG 0126498, YMG
0126502; Nichihara-cho: YMG 0126496, YMG
0126503, YMG 0035314, YMG 0031243; Muikaichimachi: YMG 0045937; Oki-gun, Fuse-mura: TNS-VS778660, TI 00000237; Oki-gun, Goka-mura: YMG
0126492.
Okayama Pref. Okayama-shi: TNS-VS-451854; Takahashi-shi: TNS-VS-330890; Niimi-shi: TNS-VS451849, TNS-VS-537134, TNS-VS-595834, TNS-VS778372; Mitsu-gun, Mitsu-cho: TNS-VS-778370;
Kawakami-gun, Nariwa-cho: YMG 0126485; Atetsugun, Osa-cho: TNS-VS-451851, TNS-VS-537135;
Shingo-cho: TNS-VS-451846, TNS-VS-451852, KYO
00010237; Tetta-cho: TNS-VS-451848, TNS-VS451850; Tessei-cho: TNS-VS-451847; Maniwa-gun,
Katsuyama-cho: TNS-VS-463210, TNS-VS-595832;
Ochiai-cho: TNS-VS-451853; Yubara-cho: TNS-VS778369; Kuse-cho: TNS-VS-595833; Shinjo-son: TNSVS-537133; Chuuka-son: TNS-VS-745196, TNS-VS778371, TNS-VS-793461; Tomata-gun, Okutsu-cho:
TNS-VS-778364; Okutsu-cho: TNS-VS-778373; Kamisaibara-son: TNS-VS-778363; Kagamino-cho: TNSVS-778375; Katsuta-gun, Nagi-cho: YMG 0047450;
Shoboku-cho: TNS-VS-778366; Kume-gun, Kumecho: TNS-VS-778374.
Hiroshima Pref. Hiroshima-shi: TNS-VS-385310, TNSVS-387077, TNS-VS-793389, TNS-VS-793394, TNSVS-793395, TNS-VS-9516544; Otake-shi: TNS-VS385311, TNS-VS-793390; Aki-gun, Kamagari-cho:
TNS-VS-385312, TNS-VS-793388; Saeki-gun, Hatsukaichi-cho: TNS-VS-793398, TNS-VS-793434; Yukicho: TNS-VS-793397; Saeki-cho: TNS-VS-793396;
Asa-gun, Gion-cho: TNS-VS-9516545; Yamagata-gun,Kake-cho: YMG 0031245; Geihoku-cho: TNS-VS449319, TNS-VS-793392; Takata-gun, Takamiya-cho:

TNS-VS-793466; Jinseki-gun, Yuki-cho: TNS-VS793449; Hiba-gun, Takano-cho: TNS-VS-778367;
Hiwa-cho: TNS-VS-793468.
Yamaguchi Pref. Shimonoseki-shi: YMG 0079589; Yamaguchi-shi: TNS-VS-778368, TNS-VS-778376,
TNS-VS-778377, YMG 0001646; Hagi-shi: YMG
0034402; Tokuyama-shi: TNS-VS-745197, YMG
0072207, YMG 0126472, YMG 0079580, YMG
0085479, KYO 00010232; Iwakuni-shi: TNS-VS793445, TNS-VS-793446, TNS-VS-793447, TNS-VS793448, YMG 0071661; Nagato-shi: YMG 0079591;
Yanai-shi: TNS-VS-535242; Shinnanyo-shi: TNS-VS793455, YMG 0069294, YMG 0071205; Oshima-gun,
Oshima-cho: TNS-VS-793452; Kuga-gun, Hongo-son:
YMG 0073264; Nishiki-cho: TNS-VS-276002;
Mikawa-cho: YMG 0070591; Tsuno-gun, Kano-cho:
TNS-VS-276001; Saba-gun, Tokuji-cho: YMG
0079594, YMG 0017938, YMG 0079595; Toyora-gun,
Kikugawa-cho: YMG 0035910; Toyota-cho: YMG
0035270, YMG 0003578, YMG 0017940, YMG
0035154, YMG 0017961, YMG 0126477, YMG
0014012, YMG 0126461; Mine-gun, Mito-cho: YMG
0126499, YMG 0055280, YMG 0005265; Otsu-gun,Misumi-cho: YMG 0126473; Abu-gun, Kawakamison: YMG 0079596, YMG 0017945, YMG 0079593;
Abu-cho: YMG 0041079, TNS-VS-559738, TNS-VS559741, YMG 0034563, YMG 0059101, YMG
0025522; Asahi-son: YMG 0126464, YMG 0017933.
Tokushima Pref. Tokushima-shi: TNS-VS-9516622,
KYO 00010619; Naka-gun, Kaminaka-cho: TNS-VS174927, TNS-VS-779169, TNS-VS-9516612, TNSVS-9516613; Kisawa-son: TNS-VS-9516607, KYO
00010469; Kito-son: TNS-VS-321856, TNS-VS9516606; Kaifu-gun, Mugi-cho: TNS-VS-9516617;
Kainan-cho: TNS-VS-9516611; Oe-gun, Kawashimacho: TNS-VS-9516618; Yamakawa-cho: TNS-VS9516619; Mima-gun, Handa-cho: TNS-VS-9516615;
Sadamitsu-cho: TNS-VS-1031923; Anabuki-cho: TNSVS-585390; Miyoshi-gun, Yamashiro-cho: TNS-VS9516616.
Kagawa Pref. Shozu-gun, Uchinomi-cho: TNS-VS9516642; Mitoyo-gun, Mino-cho: TNS-VS-9516630.
Ehime Pref. Matsuyama-shi: TNS-VS-525389; Uwajimashi: TNS-VS-282160; Kawanoe-shi: KYO 00010475;
Hojo-shi: YMG 0074572; Shuso-gun, Tanbaracho: TNS-VS-779163; Kamiukena-gun, Omogo-mura:
TNS-VS-793463; Oda-cho: TNS-VS-525385; Higashiuwa-gun, Uwa-cho: TNS-VS-525388; Kitauwagun, Hiyoshi-mura: TNS-VS-525390; Minamiuwagun, Johen-cho: KYO 00010483.
Kochi Pref. Kochi-shi: TNS-VS-1031928, TNS-VS1031942, TNS-VS-1031945; Tosa-shi: TNS-VS54863; Nakamura-shi: TNS-VS-553426; Tosashimizushi: TNS-VS-553182, TNS-VS-779176; Aki-gun,

Distribution of Vandenboschia in Japan
Toyo-cho: TNS-VS-553408; Nahari-cho: TNS-VS553185; Kami-gun, Tosayamada-cho: TNS-VS-384351,
TNS-VS-553429; Monobe-son: TNS-VS-553183,
TNS-VS-1031956, TNS-VS-1031957, MBK 0027858;
Nagaoka-gun, Motoyama-cho: TNS-VS-553415, TNSVS-553631, TNS-VS-1031938, KYO 00010461; Tosagun, Tosa-cho: TNS-VS-553412, TNS-VS-1031910;
Okawa-mura: TNS-VS-553414; Agawa-gun, Ikegawacho: TNS-VS-553413, KYO 00010471; Gohokuson: TNS-VS-553416, TNS-VS-553417; Takaoka-gun,
Sakawa-cho: MBK 0019990, KYO 00010465; Ochi-cho:
TNS-VS-779178; Yusuhara-cho: TNS-VS-553184;
Onomi-son: TNS-VS-553411, TNS-VS-1032003,
TNS-VS-1032004, TNS-VS-1032028; Niyodo-mura:
TI 00000555; Hata-gun, Taisho-cho: TNS-VS-460837,
TNS-VS-1031927; Towa-son: TNS-VS-1031980.
Fukuoka Pref. Kitakyushu-shi: TNS-VS-589853; Kurume-shi: TNS-VS-1031999; Amagi-shi: TNS-VS590373, KYO 00010595; Maebaru-shi: TNS-VS778637; Chikushi-gun, Nakagawa-machi: TNS-VS778631; Kasuya-gun, Umi-machi: TNS-VS-590372;
Sasaguri-machi: TNS-VS-590371; Kaho-gun, Chikuhomachi: TNS-VS-512952; Itoshima-gun, Maebarumachi: TNS-VS-590379; Nijo-machi: TNS-VS590380; Yame-gun, Kurogi-machi: TNS-VS-590374;
Tachibana-machi: TNS-VS-590375; Yabe-mura: TNSVS-590370; Hoshino-mura: TNS-VS-590369; Tagawagun, Soeda-machi: TNS-VS-589854; Kawasaki-machi:
TNS-VS-589855; Miyako-gun, Saigawa-machi: TNSVS-590397.
Saga Pref. Imari-shi: TNS-VS-536403, TNS-VS-536404;
Saga-gun, Yamato-cho: TNS-VS-536411; Kanzakigun, Higashisefuri-son: TNS-VS-536397, TNS-VS778635; Higashimatsuura-gun, Kyuuragi-machi: TNSVS-536408; Kitahata-mura: TNS-VS-778628; Nishimatsuura-gun, Nishiarita-cho: TNS-VS-536401.
Nagasaki Pref. Nagasaki-shi: TNS-VS-461512, TNSVS-480531; Shimabara-shi: TNS-VS-793401, KYO
00010214; Omura-shi: TNS-VS-100193; Nishisonogigun, Seihi-cho: TNS-VS-793408; Sotome-cho: TNSVS-480562; Higashisonogi-gun, Higashisonogi-cho:
TNS-VS-480529, TNS-VS-480533; Kitatakaki-gun,Takaki-cho: TNS-VS-100195, TNS-VS-793400, TNSVS-1031934; Iki Isl./Iki-gun, Gonora-cho: TNS-VS1031909; Shimoagata-gun, Mitsushima-cho: TNS-VS793404; Kamiagata-gun, Kamitsushima-cho: TNS-VS793410.
Kumamoto Pref. Kumamoto-shi: TNS-VS-455285; Hitoyoshi-shi: TNS-VS-583585, TNS-VS-802929, KYO
00010139; Kikuchi-shi: TNS-VS-455287; Kamotogun, Kahoku-machi: TNS-VS-465093; Aso-gun, Asomachi: TNS-VS-455286; Minamioguni-machi: YMG
0081108; Namino-son: TNS-VS-463030; Kamimashiki-gun, Mifune-machi: TNS-VS-455297; Mashiki-85

machi: TNS-VS-455292, TNS-VS-455298; Yabemachi: TNS-VS-455300, TNS-VS-583583, TNS-VS583584, TNS-VS-778333; Yatsushiro-gun, Toyo-mura:
TNS-VS-455296; Ashikita-gun, Tsunagi-machi: TNSVS-455160; Kuma-gun, Nishiki-machi: TNS-VS376714; Taragi-machi: KYO 00010142; Mizukamimura: TNS-VS-455282, TNS-VS-583609; Itsuki-mura:
TNS-VS-332447, TNS-VS-793450; Yamae-mura:
TNS-VS-778334, TNS-VS-778336; Kuma-mura:
TNS-VS-455158.
Oita Pref. Oita-shi: TNS-VS-456729; Hita-shi: TNS-VS456753, YMG 0079607; Taketa-shi: TNS-VS-50515,
TNS-VS-456740, TNS-VS-456745; Usa-shi: TNS-VS456754; Nishikunisaki-gun, Kakaji-cho: TNS-VS456750; Higashikunisaki-gun, Aki-machi: TNS-VS456728; Oita-gun, Shonai-cho: TNS-VS-456748; Yufuin-cho: YMG 0079604; Minamiamabe-gun, Umemachi: TNS-VS-456732; Ono-gun, Mie-machi: TNSVS-456739, YMG 0042972; Kiyokawa-mura: TNSVS-456730, TNS-VS-456734; Ogata-machi: TNS-VS456731; Asaji-machi: TNS-VS-456755; Ono-machi:
YMG 0079608; Naoiri-gun, Kujuu-machi: TNS-VS456743; Kusu-gun, Kokonoe-machi: TNS-VS-456742;
Kusu-machi: TNS-VS-778632, TNS-VS-779206, TNSVS-779207, TNS-VS-779208, YMG 0079606, KYO
00010055; Hita-gun, Maetsue-mura: YMG 0126506;
Kamitsue-mura: TNS-VS-456738, YMG 0079605;
Amagase-machi: TNS-VS-456737, TNS-VS-456744,
TNS-VS-779212; Shimoge-gun, Yabakei-machi: TNSVS-456736; Yamakuni-machi: TNS-VS-456752.
Miyazaki Pref. Kobayashi-shi: TNS-VS-1031988, TNSVS-1032023; Ebino-shi: TNS-VS-521725, TNS-VS521777, TNS-VS-521778, TNS-VS-521785, TNSVS-762705, KYO 00010217; Minaminaka-gun, Kitago-cho: TNS-VS-521721; Kitamorokata-gun, Yamanokuchi-cho: KYO 00010171; Nishimorokata-gun,Suki-son: TNS-VS-521780; Koyu-gun, Kijo-son: TNSVS-521727; Higashiusuki-gun, Saigo-son: TNS-VS793458; Morotsuka-son: TNS-VS-521722; Shiiba-son:
TNS-VS-521781; Nishiusuki-gun, Takachiho-cho:
TNS-VS-459236, TNS-VS-521717, TNS-VS-793460,
KYO 00010145; Hinokage-cho: TNS-VS-521779,
TNS-VS-793459.
Kagoshima Pref. Kagoshima-shi: TNS-VS-778959;
Kanoya-shi: TNS-VS-453622; Izumi-shi: TNS-VS463092; Okuchi-shi: TNS-VS-762587, TNS-VS777635, TNS-VS-778950; Kokubu-shi: TNS-VS453625;
Tarumizu-shi:
TNS-VS-453611;
Satsumasendai-shi: TNS-VS-771413; Kawanabe-gun, Chiran-cho: TNS-VS-9516574, TNS-VS-9516587; Satsuma-gun, Satsuma-cho: TNS-VS-453782; Aira-gun,Mizobe-cho: TNS-VS-1032011; Yokogawa-cho: KYO
00010195; Makizono-cho: TNS-VS-1032008, KYO
00010216, KYO 00010190; So-gun, Kihoku-cho:86Atsushi Ebihara
TNS-VS-453610; Ariake-cho: TNS-VS-453787; Kimotsuki-gun, Uchinoura-cho: TNS-VS-453606, TNSVS-1031921; Koyama-cho: TNS-VS-453591; Onejime-cho: TNS-VS-453605.

Vandenboschia quelpaertensis (Nakai) Ebihara (Fig. 10)
Chiba Pref. Kamogawa-shi: TNS-VS-540084, TNS-VS724690, TNS-VS-793471; Kimitsu-shi: TNS-VS540082, TNS-VS-802952, TNS-VS-927455; Awa-gun,Maruyama-machi: TNS-VS-540083; Wada-machi:
TNS-VS-540112, TNS-VS-724691; Amatsukominatomachi: TNS-VS-1406, TNS-VS-71593, TNS-VS107105, TNS-VS-254504, TNS-VS-256654, TNS-VS540085, TNS-VS-604413, TNS-VS-690436, TNS-VS724694, TNS-VS-802851, TNS-VS-802953, TNS-VS803033, TNS-VS-803035, TNS-VS-803036, TNS-VS803037, TNS-VS-803039, TNS-VS-803040, TNS-VS803041, TNS-VS-927251, TNS-VS-927253, TNS-VS802852a/b.
Tokyo Pref. Ome-shi: TNS-VS-803016; Oshima Isl./
Oshima-machi: TNS-VS-647321, TNS-VS-803026;
Toshima Isl./Toshima-mura: TNS-VS-459314; Kozu
Isl./Kozushima-mura: TNS-VS-802847, TNS-VS802849; Miyake Isl./Miyake-mura: TNS-VS-43235,
TNS-VS-51481, TNS-VS-83866, TNS-VS-88952,
TNS-VS-88953, TNS-VS-459311, TNS-VS-803023,
TNS-VS-803024, TNS-VS-803025; Mikura Isl./Mikurajima-mura: TNS-VS-121028, TNS-VS-144379,
TNS-VS-647390, TNS-VS-802947, TNS-VS-802955,
TNS-VS-802958, TNS-VS-802959, TNS-VS-803018,
TNS-VS-803019, TNS-VS-803021, TNS-VS-803022,
TNS-VS-1031991; Hachijo Isl./Hachijo-machi: TNSVS-54857, TNS-VS-81228, TNS-VS-81229, TNS-VS121027, TNS-VS-181331, TNS-VS-369874, TNS-VS370361, TNS-VS-390724, TNS-VS-459315, TNS-VS747050, TNS-VS-775464, TNS-VS-802844, TNS-VS802948, TNS-VS-802949, TNS-VS-802950, TNS-VS802960, TNS-VS-802961, TNS-VS-802962, TNS-VS802963, TNS-VS-802965, TNS-VS-802966, TNS-VS802967, TNS-VS-802968, TNS-VS-803015, TNS-VS803044, TNS-VS-803045, TNS-VS-803046, TNS-VS803047, TNS-VS-803048, TNS-VS-803049, TNS-VS803050, TNS-VS-803051, TNS-VS-803052, TNS-VS803053, TNS-VS-803054, TNS-VS-803055, TNS-VS803056, TNS-VS-803057, TNS-VS-803058, TNS-VS803059, TNS-VS-803060; Aogashima Isl./Aogashimamura: TNS-VS-137186, TNS-VS-369693; Chichijima
Isl./Ogasawara-mura: TNS-VS-266465; TNS-VS266466; Hahajima Isl./Ogasawara-mura: TNS-VS290706, TNS-VS-523870, TNS-VS-778663, TNS-VS802885, TNS-VS-802886, TNS-VS-802887, TNS-VS803032.
Kanagawa Pref. Zushi-shi: TNS-VS-779080; Miura-gun,
Hayama-machi: TNS-VS-142426; Ashigarakami-gun,

Yamakita-machi: TNS-VS-561715, TNS-VS-9516537;
Ashigarashimo-gun, Yugawara-machi: TNS-VS-561707.
Fukui Pref. Oi-gun, Takahama-cho: TNS-VS-793465.
Gifu Pref. Yamagata-gun, Miyama-cho: TNS-VS802918.
Shizuoka Pref. Hamamatsu-shi: TNS-VS-925901, TNSVS-925902; Numazu-shi: TNS-VS-449843, TNS-VS536235; Atami-shi: TNS-VS-449831, TNS-VS536187; Ito-shi: TNS-VS-536236, TNS-VS-536237,
TNS-VS-557306, TNS-VS-802988, TNS-VS-802989,
TNS-VS-802990, TNS-VS-802991, TNS-VS-802993,
TNS-VS-802994, TNS-VS-802995, TNS-VS-802996,
TNS-VS-802997, TNS-VS-802998, TNS-VS-802999,
TNS-VS-803000, TNS-VS-803001, TNS-VS-803005,
TNS-VS-925917, TNS-VS-925918, TNS-VS-1032032;
Fuji-shi: TNS-VS-449825, TNS-VS-793456; Kakegawashi: TNS-VS-252868, TNS-VS-449810, TNS-VS536192, TNS-VS-536204, TNS-VS-925907, TNS-VS1031891, TNS-VS-1031993; Fujieda-shi: TNS-VS536195; Tenryu-shi: TNS-VS-449824, TNS-VS536197, TNS-VS-536199, TNS-VS-542744, TNS-VS925896, TNS-VS-925899, TNS-VS-925900; Shimodashi: TNS-VS-778644; Susono-shi: TNS-VS-946593;
Kamo-gun, Higashiizu-cho: TNS-VS-449836, TNSVS-536181, TNS-VS-802871, TNS-VS-802872, OSA
00207881; Kawazu-cho: TNS-VS-142061, TNS-VS288448, TNS-VS-312905, TNS-VS-323272, TNS-VS332534, TNS-VS-536188, TNS-VS-650035, TNS-VS802922, TNS-VS-803010, TNS-VS-803011, TNS-VS803012, TNS-VS-1031907, TNS-VS-9512582; Matsuzaki-cho: TNS-VS-288490, TNS-VS-536224, TNSVS-449817, TNS-VS-536227, TNS-VS-925909; Nishiizu-cho: TNS-VS-286887, TNS-VS-331680, TNS-VS332540, TNS-VS-449815, TNS-VS-778643, TNS-VS778647, TAI 106818; Kamo-mura: TNS-VS-155909,
TNS-VS-156526, TNS-VS-449819, TNS-VS-449842,
TNS-VS-531376, TNS-VS-536226, TNS-VS-802870;
Tagata-gun, Heda-mura: TNS-VS-210669, TNS-VS449834, TNS-VS-449841, TNS-VS-536186, TNS-VS939979; Nirayama-cho: TNS-VS-449844; Amagiyugashima-cho: TNS-VS-449818, TNS-VS-564258,
TNS-VS-802873, TNS-VS-802874, TNS-VS-802878,
TNS-VS-802879, TNS-VS-803002, TNS-VS-803003,
TNS-VS-925906; Nakaizu-cho: TNS-VS-566223, TNSVS-773768, TNS-VS-965065, TNS-VS-1031964,
TNS-VS-1031983, TNS-VS-1032001, KYO 00010451;
Fuji-gun, Shibakawa-cho: TNS-VS-793457; Ogasa-gun,Ogasa-cho: TNS-VS-545458; Daito-cho: TNS-VS44338, TNS-VS-536193; Shuchi-gun, Haruno-cho:
TNS-VS-536184; Iwata-gun, Toyoka-mura: TNS-VS449812, TNS-VS-536200, TNS-VS-925898; Sakumacho: TNS-VS-288445, TNS-VS-449828, TNS-VS536228, TNS-VS-802908, TNS-VS-802917, TNS-VS1031901, TNS-VS-1032044; Misakubo-cho: TNS-VS-

Distribution of Vandenboschia in Japan
802881; Inasa-gun, Mikkabi-cho: TNS-VS-925914.
Aichi Pref. Okazaki-shi: TNS-VS-463610; Toyohashi-shi:
TNS-VS-9516623; Kasugai-shi: TNS-VS-95166276.
Mie Pref. Ise-shi: TNS-VS-378982, TNS-VS-1031984;
Matsusaka-shi: TNS-VS-537920; Owase-shi: TNSVS-537836, TNS-VS-130364, TNS-VS-144738, TNSVS-793438, TNS-VS-802919, TNS-VS-802921, TNSVS-1031903, TNS-VS-1031912, TNS-VS-1031948,
TNS-VS-1032035, MBK 0030322, OSA s.n. (K. Seto
4229); Kumano-shi: TNS-VS-537842, TNS-VS802860a/b, TNS-VS-1031995, TNS-VS-1032018, TNSVS-1032019; Hisai-shi: TNS-VS-537914; Kuwana-gun,Tado-cho: TNS-VS-803008, TNS-VS-941848; Ichishigun, Ichishi-cho: TNS-VS-537912; Iinan-gun, Iitakacho: YMG 0039310; Taki-gun, Seiwa-mura: OSA
00005006; Miyagawa-mura: OSA 00100402; Wataraigun, Nansei-cho: KYO 00010249; Nanto-cho: TNSVS-537918; Omiya-cho: TNS-VS-537910; Kisei-cho:
TNS-VS-537845; Kitamuro-gun, Kiinagashima-cho:
TNS-VS-537835; Minamimuro-gun, Mihama-cho:
TNS-VS-537911; Kiho-cho: TNS-VS-464084, TNSVS-778925, TNS-VS-779032, TNS-VS-9508433.
Shiga Pref. Ika-gun, Kinomoto-cho: TNS-VS-802888,
TNS-VS-802889.
Kyoto Pref. Otokuni-gun, Kasa-gun, Oe-cho: TNS-VS194842, KYO 00010382, KYO 00010383.
Osaka Pref. Izumi-shi: TNS-VS-726370; Sennan-gun:
TNS-VS-1032002.
Hyogo Pref. Tsuna-gun, Tsuna-cho: TNS-VS-554754.
Nara Pref. Gose-shi: TNS-VS-937972; Uda-gun, Muromura: KYO 00010368; Yoshino-gun, Nishiyoshinomura: TNS-VS-583710.
Wakayama Pref. Hashimoto-shi: TNS-VS-937954, TNSVS-937971; Shingu-shi: TNS-VS-779045, TNS-VS9508434; Kaiso-gun, Shimotsu-cho: OSA 00131640;
Misato-cho: OSA 00005007, OSA 00026888; Aritagun, Kanaya-cho: TNS-VS-937956, OSA 00131648;
Hidaka-gun, Kawabe-cho: TNS-VS-937961; Nakatsumura: TNS-VS-937962; Nishimuro-gun, Shirahamacho: TNS-VS-37588, TNS-VS-775469, TNS-VS802931, TNS-VS-802932, TNS-VS-802933, TNS-VS802934, TNS-VS-802935, TNS-VS-802936, TNS-VS802942, TNS-VS-802944, TNS-VS-802945, TNS-VS802946, TNS-VS-937944; Nakahechi-cho: TNS-VS937939, OSA 00034222, OSA 00131659; Oto-mura:
OSA s.n. (K. Seto 13165), OSA 00131668, OSA
00131674; Hikigawa-cho: TNS-VS-937973; Susamicho: TNS-VS-937948, OSA 00052415, OSA 00131680,
OSA 00131681, OSA 00131682, OSA 00131685,
OSA 00152821; Kushimoto-cho: TNS-VS-937953;
Higashimuro-gun,Nachikatsuura-cho:
TNS-VS937949; Taiji-cho: KYO 00010254; Koza-cho: TNSVS-692575, TNS-VS-802920; Kozagawa-cho: TNSVS-937946, TNS-VS-937951, TNS-VS-937952, TNS-87

VS-1031998, KYO 00010358, OSA 00077294; Kumanogawa-cho: TNS-VS-937942, TNS-VS-937943.
Tottori Pref. Iwami-gun, Iwami-cho: TNS-VS-452716.
Shimane Pref. Hirata-shi: TNS-VS-9516600; Kanoashigun, Tsuwano-cho: YMG 0034347.
Hiroshima Pref. Hiroshima-shi: TNS-VS-1032048;
Saeki-gun, Yuki-cho: TNS-VS-793391.
Yamaguchi Pref. Yamaguchi-shi: TNS-VS-50514, TNSVS-275998, YMG 0079579, YMG 0086023; Tokuyama-shi: YMG 0126470; Iwakuni-shi: YMG 0077214,
YMG 0076017; Yanai-shi: TNS-VS-793476; Mine-shi:
YMG 0126482; Shinnanyo-shi: TNS-VS-793454; Kugagun, Nishiki-cho: YMG 0079585, YMG 0126466;Miwa-cho: YMG 0079577; Tsuno-gun, Kano-cho:
YMG 0079584, YMG 0079582; Saba-gun, Tokuji-cho:
YMG 0126468; Toyora-gun, Toyota-cho: YMG
0075601, YMG 0017098, YMG 0016162, YMG
0126476, YMG 0126478, YMG 0126480; Mine-gun,Shuho-cho: TNS-VS-275999, YMG 0045852, YMG
0126481; Otsu-gun, Misumi-cho: YMG 0126469;
Yuya-cho: YMG 0079590; Abu-gun, Kawakami-son:
TNS-VS-793462; Ato-cho: TNS-VS-50513, YMG
0019801.
Tokushima Pref. Tokushima-shi: TNS-VS-50516, KYO
00010476; Anan-shi: TNS-VS-9516608; Kaifu-gun,Kainan-cho: TNS-VS-779173, TNS-VS-779190, TNSVS-1032034, TNS-VS-1032049, OSA s.n. (K. Seto
12060); Shishikui-cho: TNS-VS-1031892, TNSVS-1032037, TNS-VS-9516542, TNS-VS-9516620;
Miyoshi-gun, Ikawa-cho: TNS-VS-1032038.
Ehime Pref. Niihama-shi: OSA 00005011; Ozu-shi:
TNS-VS-525393; Uma-gun, Shingu-mura: TNS-VS793451; Shuso-gun, Tanbara-cho: TNS-VS-793473;
Nishiuwa-gun, Honai-cho: YMG 0074574; Higashiuwagun, Nomura-cho: KYO 00010479, KYO 00010480;
Kitauwa-gun, Tsushima-cho: YMG 0074573, OSA
00005010; Minamiuwa-gun, Johen-cho: TNS-VS35819; Ipponmatsu-cho: KYO 00010522.
Kochi Pref. Muroto-shi: TNS-VS-460838, TNS-VS553409, TNS-VS-1032027, MBK 0046361; Aki-shi:
TNS-VS-553407; Sukumo-shi: TNS-VS-460839;
TNS-VS-553181, TNS-VS-779162; Tosashimizu-shi:
TNS-VS-553180, TNS-VS-553419, TNS-VS-647391,
TNS-VS-771529, TNS-VS-779174, TNS-VS-803013,
TNS-VS-803014, TNS-VS-1031906, TNS-VS-1032015,
TNS-VS-1032030,
TNS-VS-1032031,
TNS-VS1032033; Aki-gun, Toyo-cho: TNS-VS-128039, TNSVS-779166, TNS-VS-1031902, MBK 0049946; Yasuda-cho: TNS-VS-553410; Kitagawa-mura: OSA
00212426; Umaji-mura: TNS-VS-462524; Kami-gun,
Monobe-son: TNS-VS-1031911, MBK 0027859; Nagaoka-gun, Otoyo-cho: TNS-VS-1032026, KYO
00010468; Takaoka-gun, Ochi-cho: TNS-VS-318479,
TNS-VS-1031893, YMG 0081109; Higashitsuno-

88

Atsushi Ebihara

mura: TNS-VS-553427; Hayama-mura: TNS-VS553428.
Fukuoka Pref. Kitakyuushuu-shi: TNS-VS-463650, TNSVS-590387; Fukuoka-shi: TNS-VS-590378, TNS-VS590386; Kurume-shi: TNS-VS-590383, TNS-VS1031960; Yamada-shi: TNS-VS-463649; Chikushigun, Nakagawa-machi: TNS-VS-590376, TNS-VS590382; TNS-VS-778630, TNS-VS-778638, OSA
00003151; Kasuya-gun, Hisayama-machi: TNS-VS590385; Kaho-gun, Kaho-machi: TNS-VS-590384;
Chikuho-machi: YMG 0036457; Tagawa-gun, Soedamachi: TNS-VS-590398.
Saga Pref. Taku-shi: TNS-VS-272798; Imari-shi: TNSVS-536398, TNS-VS-941840; Takeo-shi: TNS-VS536400; Kashima-shi: TNS-VS-536407; Saga-gun,
Yamato-cho: TNS-VS-536410; Ogi-gun, Ogi-machi:
TNS-VS-536413; Higashimatsuura-gun, Kitahatamura: TNS-VS-536405, TNS-VS-778626, TNS-VS778627; Nishimatsuura-gun: TNS-VS-1031887.
Nagasaki Pref. Nagasaki-shi: TNS-VS-793405, TNSVS-793406, TNS-VS-793407; Shimabara-shi: TNSVS-50517; Isahaya-shi: TNS-VS-480532, TNS-VS793411; Omura-shi: TNS-VS-793409, TNS-VS-941849;
Minamitakaki-gun, Mizuho-cho: TNS-VS-793412; Minamimatsuura-gun, Arikawa-cho: TNS-VS-793403; Iki
Isl./Iki-gun, Gonora-cho: TNS-VS-1031896.
Kumamoto Pref. Hitoyoshi-shi: TNS-VS-35816, TNSVS-455159; Minamata-shi: TNS-VS-802909, TNS-VS802910, TNS-VS-802911; Hondo-shi: OSA 00005014;
Yamaga-shi: YMG 0081110; Amakusa-shi: TNS-VS762650; Shimomashiki-gun, Chuo-son: TNS-VS455302; Tomochi-machi: TNS-VS-455301; Kamotogun, Kikuka-machi: TNS-VS-465094; Kamimashikigun, Mifune-machi: TNS-VS-455304; Yatsushiro-gun,
Sakamoto-mura: TNS-VS-293595, TNS-VS-455291,
TNS-VS-455295, KYO 00010512; Miyahara-machi:
TNS-VS-455294; Toyo-mura: TNS-VS-112589, KYO
00010509; Izumi-mura: TNS-VS-455299; Ashikitagun, Ashikita-machi: TNS-VS-455290; Kuma-gun,
Nishiki-machi: TNS-VS-455283; Okaharu-mura: TNSVS-465095; Sue-mura: TNS-VS-455284; Kuma-mura:
TNS-VS-455289, KYO 00010125; Amakusa-gun, Ariake-machi: TNS-VS-455288; Kuratake-machi: KYO
00010154; Sumoto-machi: TNS-VS-455157; Amakusamachi: TNS-VS-778337, OSA 00005013.
Oita Pref. Oita-shi: TNS-VS-456741; Saiki-shi: TNSVS-456733, TNS-VS-456746; Bungotakada-shi: TNSVS-456751; Oita-gun, Shonai-cho: TNS-VS-456747;
Shimoge-gun, Yabakei-machi: KYO 00010133, KYO
00010134; Usa-gun, Ajimu-machi: TNS-VS-456756.
Miyazaki Pref. Miyazaki-shi: TNS-VS-521775, TNS-VS521783, TNS-VS-802928; Miyakonojo-shi: TNS-VS521736, TNS-VS-802843, KYO 00010521; Nobeokashi: TNS-VS-521728; Nichinan-shi: TNS-VS-83858,

TNS-VS-83861, TNS-VS-83864, TNS-VS-378984,
TNS-VS-378985, TNS-VS-521726, TNS-VS-521718,
TNS-VS-521735, TNS-VS-700553, TNS-VS-784659,
TNS-VS-784672, TNS-VS-1031985, TNS-VS-1032010,
TNS-VS-1032014, TNS-VS-9518557; Kobayashi-shi:
TNS-VS-521723; Kushima-shi: TNS-VS-521782;
Saito-shi: TNS-VS-521729, KYO 00010127; Ebinoshi: KYO 00010158; Miyazaki-gun, Tano-cho: TNSVS-521737; Minaminaka-gun, Kitago-cho: TNS-VS762768; Kitamorokata-gun, Yamanokuchi-cho: RYU
38174; Nishimorokata-gun, Takaharu-cho: TNS-VS521784; Nojiri-cho: TNS-VS-521724; Higashimorokata-gun, Takaoka-cho: TNS-VS-521733; Aya-cho:
TNS-VS-521730, KYO 00010056; Koyu-gun, Kijocho: TNS-VS-521740; Higashiusuki-gun, Togocho: TNS-VS-521741; Shiiba-son: TNS-VS-776801;
Nishiusuki-gun, Takachiho-cho: KYO 00010163.
Kagoshima Pref. Kagoshima-shi: TNS-VS-775465, TNSVS-925913, TNS-VS-1032005, TNS-VS-1032007;
Sendai-shi: TNS-VS-463089; Kanoya-shi: TNS-VS453786, TNS-VS-1031915; Okuchi-shi: TNS-VS453617, TNS-VS-762585, TNS-VS-762586, TNS-VS777321, TNS-VS-778963, TNS-VS-802925, KYO
00010504; Ibusuki-shi: TNS-VS-453619; Kokubu-shi:
KYO 00010489; Tarumizu-shi: TNS-VS-305445,
TNS-VS-802902, TNS-VS-802903, TNS-VS-802924,
TNS-VS-1032036, KYO 00010526; Kagoshima-gun,
Yoshida-cho: TNS-VS-453598; Mishima-mura: KYO
00010528; Ibusuki-gun, Kiire-cho: TNS-VS-453620;
Kaimon-cho: TNS-VS-710349, YMG 0031221;
Kawanabe-gun, Chiran-cho: TNS-VS-453600, TNSVS-453615, TNS-VS-9516570, TNS-VS-9516572,
TNS-VS-9516573, TNS-VS-9516576; Satsuma-gun,Iriki-cho: TNS-VS-463090, KYO 00010501; Miyanojo-cho: TNS-VS-539621, TNS-VS-802884, TNS-VS1031987, TNS-VS-1031996; Tsuruda-cho: TNS-VS777370, TNS-VS-777621, TNS-VS-802892, TNS-VS802893, TNS-VS-802894, TNS-VS-802900, TNSVS-802901; Kedoin-cho: TNS-VS-463088; KYO
00010505; Shimokoshiki Isl./Shimokoshiki-son: TNSVS-453589, TNS-VS-453792, TNS-VS-802891, TNSVS-803075; Satsuma-cho: TNS-VS-777032; Aira-gun,Kajiki-cho: TNS-VS-1031972, TNS-VS-1032012;
Aira-cho: TNS-VS-453602, TNS-VS-1031914, TNSVS-1031958, TNS-VS-1031989, KYO 00010194,
KYO 00010503; Kamo-cho: KYO 00010497; Mizobecho: TNS-VS-453789, KYO 00010197; Yokogawacho: TNS-VS-453781, KYO 00010498; Makizonocho: KYO 00010507; So-gun, Sueyoshi-cho: TNS-VS453608; Shibushi-cho: TNS-VS-453607; Ariake-cho:
TNS-VS-453609; Kimotsuki-gun, Uchinoura-cho:
TNS-VS-453590, TNS-VS-802880, TNS-VS-802898,
TNS-VS-1032040,
TNS-VS-1032041,
TNS-VS1032043, KYO 00010508; Koyama-cho: YMG 0047317,

Distribution of Vandenboschia in JapanKYO 00010161; Onejime-cho: OSA 00038194; Nejime-cho: TNS-VS-453623; Tashiro-cho: TNS-VS144654, TNS-VS-262554, TNS-VS-632744, TNS-VS925904, TNS-VS-1031889, TNS-VS-1031895, TNSVS-1031965, RYU 33444; Sata-cho: TNS-VS1031894, TNS-VS-1031899; Tanegashima Isl./Nishinoomote-shi: TNS-VS-155373, TNS-VS-453796,
TNS-VS-507272, TNS-VS-507273, TNS-VS-507279,
TNS-VS-507306, TNS-VS-507307, TNS-VS-507308,
TNS-VS-507310, TNS-VS-507311, TNS-VS-507313,
TNS-VS-507314, TNS-VS-507315, TNS-VS-507316,
TNS-VS-507317, TNS-VS-507318, TNS-VS-514282,
TNS-VS-514283, TNS-VS-514284, TNS-VS-514285,
TNS-VS-514286,
TNS-VS-745194,
TNS-VS9519978; KAG s.n. (Teruya 300); Tanegashima
Isl./Kumage-gun, Nakatane-cho: TNS-VS-507280,
TNS-VS-507309, TNS-VS-507312, TNS-VS-507319,
TNS-VS-508399, TNS-VS-784874; Tanegashima
Isl./Minamitane-cho: TNS-VS-453587, TNS-VS453795, TNS-VS-507274, TNS-VS-507275, TNS-VS507276, TNS-VS-507277, TNS-VS-507278, TNS-VS507305, TNS-VS-507401, TNS-VS-507535, TNS-VS507536; Yaku Isl./Kamiyaku-cho: TNS-VS-321833,
TNS-VS-453592, TNS-VS-453593, TNS-VS-453627,
TNS-VS-763169, TNS-VS-802853, TNS-VS-802854,
TNS-VS-802856, TNS-VS-802857, TNS-VS-802857,
TNS-VS-802858, TNS-VS-802869, TNS-VS-1031900,
TNS-VS-1031994; Yaku Isl./Yaku-cho: TNS-VS453595, TNS-VS-453601, TNS-VS-778947, TNS-VS802859, TNS-VS-802897, TNS-VS-1032047; Nazeshi: TNS-VS-1031990; Izumi-shi: TNS-VS-323310,
TNS-VS-453586, TNS-VS-455026, TNS-VS-463091,
TNS-VS-777401, TNS-VS-802914, TNS-VS-802915,
TNS-VS-802927, TNS-VS-927456, TNS-VS-927458;
Amami-oshima Isl./Amami-shi: TNS-VS-764148;
Amami-oshima Isl./Oshima-gun, Yamato-son: KYO
00010013; Amami-oshima Isl./Uken-son: TNS-VS311712,
TNS-VS-1031968,
TNS-VS-1031979;
Amami-oshima Isl./Setouchi-cho: YMG 0045350;
Amami-oshima Isl./Sumiyo-son: TNS-VS-311709,
TNS-VS-311710, TNS-VS-311711, KYO 00010321;
Amami-oshima Isl./China-cho: TNS-VS-1031970.
Okinawa Pref. Nago-shi: TNS-VS-778922, TNS-VS778923, TNS-VS-778924, TNS-VS-936778, TNS-VS1031976; Kunigami-gun, Kunigami-son: TNS-VS296993, TNS-VS-316755, TNS-VS-317901, TNS-VS-

89

369076, TNS-VS-369704, TNS-VS-369878, TNS-VS394800, TNS-VS-750475RYU 32779; Onna-son:
TNS-VS-518028; Ishigaki Isl./Ishigaki-shi: KYO
00010308, KYO 00010312; Uotsuri Isl./Ishigaki-shi:
TNS-VS-395413, RYU 33641; Yaeyama-gun, Taketomi-cho: TNS-VS-561859, TNS-VS-778361, TNS-VS936779, RYU 32895, KAG s.n. (S. Hatusima 23197),
KYO 00010337, OSA 00010647, OSA 00040253.
Vandenboschia birmanica (Bedd.) Ching  V. nipponica(Nakai) Ebihara (Fig. 11)
Chiba Pref. Kimitsu-shi: TNS-VS-724689, TNSVS724692, TNS-VS-779084, TNS-VS-802951, TNSVS-927520, TNS-VS-1031954, TNS-VS-1032025.
Kanagawa Pref. Yokohama-shi: TNS-VS-779087.
Vandenboschia birmanica (Bedd.) Ching  V. kalamocarpa (Hayata) Ebihara  V. nipponica (Nakai) Ebihara (Fig. 12)
Chiba Pref. Awa-gun, Wada-machi: TNS-VS-724693;
Amatsukominato-machi: TNS-VS-779074, TNS-VS779082.
Tokyo Pref. Aogashima Isl./Aogashima-mura: TNS-VS779088, TNS-VS-803027, TNS-VS-803028, TNS-VS803029, TNS-VS-803030, TNS-VS-803031.
Kanagawa Pref. Yokosuka-shi: TNS-VS-779235; Yokosuka-shi/Kamakura-shi: TNS-VS-779241, TNS-VS779242, TNS-VS-779243, TNS-VS-561706, TNS-VS779227, TNS-VS-779228, TNS-VS-779229, TNS-VS779230, TNS-VS-779237, TNS-VS-779238, TNS-VS779239; Kamakura-shi: TNS-VS-779240; Fujisawashi: TNS-VS-779086; Zushi-shi: TNS-VS-35962,
TNS-VS-83863, TNS-VS-83865, TNS-VS-184003,
TNS-VS-200153, TNS-VS-305012, TNS-VS-628017,
TNS-VS-779076, TNS-VS-779077, TNS-VS-779078,
TNS-VS-779091, TNS-VS-802865, TNS-VS-802866,
TNS-VS-802867, TNS-VS-802868, TNS-VS-802976,
TNS-VS-802977, TNS-VS-802978, TNS-VS-802980,
TNS-VS-802981, TNS-VS-802983, TNS-VS-803404,
TNS-VS-1031904,
TNS-VS-1031986,
TNS-VS1032016, TNS-VS-1032020, TNS-VS-1032024, TNSVS-9516536.
Shizuoka Pref. Shimoda-shi: TNS-VS-775471, TNS-VS778641; Tagata-gun, Nakaizu-cho: TNS-VS-778651.
Tottori Pref. Tohaku-gun, Yurihama-cho: TNS-VS778652, TNS-VS-778657.