# Checklist of the marine fishes from metropolitan France

Philippe Béarez, Patrice Pruvost, Eric Feunteun, Samuel P. Iglesias, Patrice Francour, Romain Causse, Jeanne de Mazieres, Sandrine Tercerie, Nicolas Bailly

► To cite this version:

Philippe Béarez, Patrice Pruvost, Eric Feunteun, Samuel P. Iglesias, Patrice Francour, et al.. Checklist of the marine fishes from metropolitan France. Cybium : Revue Internationale d'Ichtyologie, 2017, 41 (4), pp.351 - 371. 10.26028/cybium/2017-414-006. mnhn-02921938

HAL Id: mnhn-02921938

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Submitted on 26 Apr 2022

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<figure><img src="image_1.png" /></figure>

Distributed under a Creative Commons Attribution 4.0 International License
Checklist of the marine fishes from metropolitan France

by

Philippe BÉAREZ* (1, 8), Patrice PRUVOST (2), Éric FEUNTEUN (2, 3, 8), Samuel IGLÉSIAS (2, 4, 8), Patrice FRANCOUR (5), Romain CAUSSE (2, 8), Jeanne DE MAZIERES (6), Sandrine TERCERIE (6) & Nicolas BAILLY (7, 8)

<figure><img src="image_2.png" /></figure>

© SFI
Received: 4 Jul. 2017
Accepted: 21 Nov. 2017
Editor: G. Duhamel

**Abstract.** – A list of the marine fish species occurring in the French EEZ was assembled from more than 200 references. No updated list has been published since the 19th century, although incomplete versions were available in several biodiversity information systems. The list contains 729 species distributed in 185 families. It is a preliminary step for the Atlas of Marine Fishes of France that will be further elaborated within the INPN (the National Inventory of the Natural Heritage: https://inpn.mnhn.fr).

**Résumé.** – Liste des poissons marins de France métropolitaine.

Une liste des poissons marins se trouvant dans la Zone Économique Exclusive de France a été constituée à partir de plus de 200 références. Cette liste n'avait pas été mise à jour formellement depuis la fin du 19e siècle, bien que des versions incomplètes existent dans plusieurs systèmes d'information sur la biodiversité. La liste contient 729 espèces réparties dans 185 familles. C'est une étape préliminaire pour l'Atlas des Poissons marins de France qui sera élaboré dans le cadre de l'INPN (Inventaire National du Patrimoine Naturel : https://inpn.mnhn.fr).

**Key words**

Marine fishes
Northeast Atlantic
Mediterranean Sea
Metropolitan France
Taxonomic list

No recent faunistic work covers the fish species present only in the marine waters of the French Exclusive Economic Zone (EEZ) in Europe, contrary to the continental waters (Keith et al., 2011).

Since Gervais and Boulart (1876-1877) and Moreau (1881), French ichthyologists have considered with some reasons that a list of the marine fishes of France would present no interest from a zoogeographic point view, and that only an European list or an Atlantic list versus a Mediterranean list would be relevant (J.C. Hureau, pers. comm.). Hence, they were much involved in the publication of the CLOFNAM (Check-List of the Fishes of the North-eastern Atlantic and the Mediterranean) (Hureau and Monod, 1973-1979), and its subsequent fauna, the FNAM (Fishes of the North-eastern Atlantic and the Mediterranean) (Whitehead et al., 1984-1986). This remains the case for the most recent field guides

(e.g. Quéro et al., 2003; Louisy, 2015), in which the entire Europe is considered (Atlantic only for the former).

However, the Convention on Biological Diversity (United Nations, 1992) that France signed on 13 June 1992, ratified on 10 June 1994 (Loi n° 94-477 du 10 juin 1994¹) and set up in application since 29 September 1994 (Décret n° 95-140 du 6 février 1995²), requests that countries identify the components of their biological diversity, such as species and communities, important for its conservation and sustainable use. France meets this commitment by developing inventories of species, which is an objective transposed in the national Environment Code. The article L411-1A establishes a national inventory of natural heritage (Inventaire National du Patrimoine Naturel, INPN) including the

<footnote n="1">¹ https://www.legifrance.gouv.fr/affichTexte.do?cidTexte=JORFT EXT000000731610</footnote>

<footnote n="2">² https://www.legifrance.gouv.fr/affichTexte.do?cidTexte=JORFT EXT000000734825</footnote>

<footnote>(1) Archéozoologie, archéobotanique : sociétés, pratiques et environnements (AASPE UMR 7209), Sorbonne Universités, Muséum national d'Histoire naturelle, CNRS, 55 rue Buffon, CP 56, 75005 Paris, France. [bearez@mnhn.fr]</footnote>

<footnote>(2) Biologie des Organismes et Écosystèmes Aquatiques (BOREA UMR 7208), Sorbonne Universités, Muséum national d'Histoire naturelle, Université Pierre et Marie Curie, Université de Caen Normandie, Université des Antilles, CNRS, IRD, 43 rue Cuvier, CP 26, 75005 Paris, France. [pruvost@mnhn.fr] [causse@mnhn.fr]</footnote>

<footnote>(3) MNHN - Station Marine de Dinard, Centre de Recherche et d'Enseignement Sur les Systèmes Côtiers (CRESCO), 38 rue du Port Blanc, 35800 Dinard, France. [feunteun@mnhn.fr]</footnote>

<footnote>(4) MNHN - Station Marine de Concarneau, Place de la Croix, BP 225, 29182 Concarneau Cedex, France. [iglesias@mnhn.fr]</footnote>

<footnote>(5) Ecosystèmes Côtiers Marins et Réponses aux Stress (ECOMERS FRE 3729), Université Côte d'Azur, CNRS, 28 avenue Valrose, BP 71, 06108 Nice CEDEX 2, France. [patrice.francour@unice.fr]</footnote>

<footnote>(6) UMS 2006 Patrimoine Naturel, Muséum national d'Histoire naturelle, 36 rue Geoffroy-Saint-Hilaire, 75005 Paris, France. [jdemazieres@mnhn.fr] [tercerie@mnhn.fr]</footnote>

<footnote>(7) FishBase Information and Research Group, c/o IRRI, Khush Hall, Los Baños, 4031 Laguna, Philippines. [n.bailly@fin.ph]</footnote>

<footnote>(8) Société Française d'Ichthyologie, Muséum national d'Histoire naturelle, 43 rue Cuvier, 75005 Paris, France.</footnote>

<footnote>* Corresponding author [bearez@mnhn.fr]</footnote>
<figure><img src="image_3.png" /><figcaption>Figure 1. – Original pictures of some interesting records from the French Mediterranean (© P. Francour). A: *Didogobius splechtnai* (Scandola, 25 m), a rare species in France. B: *Gobius kolombatovici* (Scandola, 50 m), a recently described species. C: *Mycteroperca rubra* (Scandola, 15 m), an uncommon species mainly found in Corsica. D: *Fistularia commersonii*: a newcomer in the Mediterranean.</figcaption></figure>

biological and geological diversity of metropolitan France and its overseas territories. The INPN is carried out under the MNHN scientific responsibility with multiple nature stakeholders and is made available to the public through its Website (inpn.mnhn.fr).

This checklist of marine fish species was developed in the framework of the INPN as a preliminary step to establish and publish an Atlas of Marine Fishes of metropolitan France. Due to the importance of fishes for the fisheries including discards, fish farming, diving and game fishing, in the economy of the country, and for the ecological status assessment of marine waters for the Marine Strategy Framework Directive, a complete list of species, taking into account newly recorded and described species (Fig. 1), also seemed to be recommended.

The following checklist is intended to provide a national reference list that will be updated with new records in the future.

## MATERIAL AND METHODS

The checklist presented herein covers the marine waters of the French metropolitan Economic Exclusive Zone (EEZ) off the coast bordering the Northeast Atlantic Ocean, including the Channel and the North Sea, and the Mediterranean Sea, including Corsica (Fig. 2). This excludes the French overseas territories.

All French fishes belonging to the subphylum Vertebrata were considered, including Myxini, Petromyzonti, Elasmobranchii, Holocephali and Actinopterygii (Eschmeyer *et al.*, 2017).

The first objective was to establish a list of species objectively recorded in the French EEZ, that means specimens either cited in scientific publications or present in fish collections, based on all available historical data. For this, we first compared a primary list extracted from FishBase (Froese and Pauly, 2017), which included more than 560 species known to occur in the French EEZ, with the list issued from the national taxonomic register TAXREF v10.0 (Gargominy *et al.*, 2016). The list for European waters in FishBase was mainly updated from the list of fishes in the European Register of Marine Species (Van der Land *et al.*, 2001).
<figure><img src="image_4.png" /><figcaption>Figure 2. – Map of the study area with delimitation of the French economic exclusive zone (EEZ).</figcaption></figure>

We then looked for objective records for all these species. Some uncommon or rare species known from a few specimens in the Northeast Atlantic, or the Mediterranean Sea but with no records from French EEZ, were not considered. To complete this drafting list, we considered the records of specimens captured in the study area and deposited in the fish collections of the Muséum national d'Histoire naturelle, Paris and all museums in France registered in GICIM (Gestion Informatisée des Collections Ichthyologiques des Musées d'histoire naturelle de France).

We also considered other web resources such as the GBIF (Global Biodiversity Information Facility) using a Geographical Information System (QGIS 2.10) to select all the potential species observed in the EEZ area extended by 110 km (ca 470 km off the coast in total). The 188 uncovered species were checked in the literature resulting in the addition of 46 species to the previous list.

The report of the Ifremer surveys (Trenkel *et al.*, 2009) was also consulted, allowing us to add five more species to the checklist. We also considered regional and local checklists as well as many useful other regional publications. In

total, more than 200 books, articles and taxonomic revisions were consulted (the list of these references is available at http://sfi-cybium.fr/fr/checklist-marine-fishes-metropolitan-france).

We followed Van der Laan *et al.* (2017) for family-group names and the Catalog of Fishes (Eschmeyer *et al.*, 2017) for the scientific nomenclature and classification. Within each family, species were sorted alphabetically.

For each species, we provide English and French common names according to the FAO, the Aquatic Science and Fisheries Information System (ASFIS), or Quéro *et al.* (2003) when available. Several non-commercial species, without available common names, were given a new common name.

We do not consider subspecies anymore. Subspecies that were regarded as valid in most of the latest publications were elevated at species rank (e.g. *Trachyscorpia cristulata echinata* is listed as *Trachyscorpia echinata*). In case of doubtful validity, subspecies were synonymised with their nominal species, which was already done in the Catalog of Fishes and FishBase.
## RESULTS

We obtained a consolidated list of 729 marine fish species with objective reports of presence in the French EEZ, distributed in 185 families and 458 genera as indicated in table I.

<table><caption>Table I. – Number of orders, families, genera and species of fish from metropolitan France.</caption><thead><tr><th>Order</th><th>Families</th><th>Genera</th><th>Species</th></tr></thead><tbody><tr><td>Myxiniformes</td><td>1</td><td>1</td><td>1</td></tr><tr><td>Petromyzontiformes</td><td>1</td><td>2</td><td>2</td></tr><tr><td>Hexanchiformes</td><td>2</td><td>3</td><td>4</td></tr><tr><td>Lamniformes</td><td>5</td><td>8</td><td>9</td></tr><tr><td>Carcharhiniformes</td><td>6</td><td>10</td><td>23</td></tr><tr><td>Squaliformes</td><td>7</td><td>12</td><td>20</td></tr><tr><td>Squatiniformes</td><td>1</td><td>1</td><td>3</td></tr><tr><td>Torpediniformes</td><td>1</td><td>2</td><td>3</td></tr><tr><td>Rhinopristiformes</td><td>3</td><td>3</td><td>4</td></tr><tr><td>Rajiformes</td><td>2</td><td>8</td><td>24</td></tr><tr><td>Myliobatiformes</td><td>5</td><td>8</td><td>9</td></tr><tr><td>Chimaeriformes</td><td>2</td><td>4</td><td>7</td></tr><tr><td>Acipenseriformes</td><td>1</td><td>1</td><td>1</td></tr><tr><td>Elopiformes</td><td>1</td><td>1</td><td>1</td></tr><tr><td>Notacanthiformes</td><td>2</td><td>4</td><td>6</td></tr><tr><td>Anguilliformes</td><td>9</td><td>20</td><td>21</td></tr><tr><td>Saccopharyngiformes</td><td>1</td><td>1</td><td>1</td></tr><tr><td>Clupeiformes</td><td>2</td><td>6</td><td>8</td></tr><tr><td>Osmeriformes</td><td>7</td><td>28</td><td>35</td></tr><tr><td>Salmoniformes</td><td>1</td><td>1</td><td>2</td></tr><tr><td>Stomiiformes</td><td>4</td><td>21</td><td>32</td></tr><tr><td>Ateleopodiformes</td><td>1</td><td>1</td><td>1</td></tr><tr><td>Aulopiformes</td><td>11</td><td>16</td><td>20</td></tr><tr><td>Myctophiformes</td><td>2</td><td>18</td><td>34</td></tr><tr><td>Lampriformes</td><td>4</td><td>5</td><td>6</td></tr><tr><td>Gadiformes</td><td>7</td><td>32</td><td>54</td></tr><tr><td>Ophidiiformes</td><td>4</td><td>11</td><td>14</td></tr><tr><td>Batrachoidiformes</td><td>1</td><td>1</td><td>1</td></tr><tr><td>Lophiiformes</td><td>4</td><td>5</td><td>7</td></tr><tr><td>Gobiesociformes</td><td>1</td><td>5</td><td>8</td></tr><tr><td>Atheriniformes</td><td>1</td><td>1</td><td>3</td></tr><tr><td>Beloniformes</td><td>3</td><td>6</td><td>9</td></tr><tr><td>Stephanoberyciformes</td><td>1</td><td>2</td><td>4</td></tr><tr><td>Beryciformes</td><td>3</td><td>3</td><td>6</td></tr><tr><td>Cetomimiformes</td><td>1</td><td>1</td><td>1</td></tr><tr><td>Zeiformes</td><td>4</td><td>6</td><td>6</td></tr><tr><td>Gasterosteiformes</td><td>1</td><td>3</td><td>3</td></tr><tr><td>Syngnathiformes</td><td>3</td><td>6</td><td>14</td></tr><tr><td>Scorpaeniformes</td><td>10</td><td>20</td><td>30</td></tr><tr><td>Perciformes</td><td>49</td><td>142</td><td>249</td></tr><tr><td>Pleuronectiformes</td><td>6</td><td>23</td><td>37</td></tr><tr><td>Tetraodontiformes</td><td>4</td><td>6</td><td>6</td></tr></tbody></table>

Non-indigenous species, for which we have rare or occasional evidence in the French EEZ, are indicated with an asterisc.

Brackish and freshwater species that only enter sporadically the marine biota, that we defined as haline waters (salinity > 30 PSU), are listed in the Annex. For more details on these species, see Keith *et al.* (2011).

These results will be available and downloadable on the INPN website (https://inpn.mnhn.fr/programme/referentiel-taxonomique-taxref) in the next version of the national taxonomic register TAXREF v11.0 expected at the end of 2017.

### CLASS MYXINI

#### Order MYXINIFORMES

##### Family Myxinidae

*Myxine glutinosa* Linnaeus, 1758 – Hagfish – Myxine

### CLASS PETROMYZONTI

#### Order PETROMYZONTIFORMES

##### Family Petromyzontidae

*Lampetra fluviatilis* (Linnaeus, 1758) – River lamprey – Lamproie de rivière

*Petromyzon marinus* Linnaeus, 1758 – Sea lamprey – Lamproie marine

### CLASS ELASMOBRANCHII

#### Order HEXANCHIFORMES

##### Family Hexanchidae

*Heptranchias perlo* (Bonnaterre, 1788) – Sharpnose sevengill shark – Requin perlon

*Hexanchus griseus* (Bonnaterre, 1788) – Bluntnose sixgill shark – Requin griset

*Hexanchus nakamurai* Teng, 1962 – Bigeyed sixgill shark – Requin vache

##### Family Chlamydoselachidae

*Chlamydoselachus anguineus* Garman, 1884 – Frilled shark – Requin lézard

#### Order LAMNIFORMES

##### Family Odontaspididae

*Carcharias taurus* Rafinesque, 1810 – Sand tiger shark – Requin taureau

*Odontaspis ferox* (Risso, 1810) – Smalltooth sand tiger – Requin féroce

##### Family Mitsukurinidae

*Mitsukurina owstoni* Jordan, 1898 – Goblin shark – Requin lutin

##### Family Lamnidae

*Carcharodon carcharias* (Linnaeus, 1758) – Great white shark – Grand requin blanc

*Isurus oxyrinchus* Rafinesque, 1810 – Shortfin mako – Taupe bleu
* *Lamna nasus* (Bonnaterre, 1788) – Porbeagle – Requin-taupe commun

Family Cetorhinidae

*Cetorhinus maximus* (Gunnerus, 1765) – Basking shark – Pèlerin

Family Alopiidae

*Alopias superciliosus* Lowe, 1841 – Bigeye thresher – Renard à gros yeux

*Alopias vulpinus* (Bonnaterre, 1788) – Thresher – Renard

Order CARCHARHINIFORMES

Family Pentanchidae

*Apristurus aphyodes* Nakaya & Stehmann, 1998 – White ghost catshark – Holbiche spectre

*Apristurus laurussonii* (Saemundsson, 1922) – Iceland catshark – Holbiche d'Islande

*Apristurus melanoasper* Iglésias, Nakaya & Stehmann, 2004 – Black roughscale catshark – Holbiche noire

*Apristurus profundorum* (Goode & Bean, 1896) – Deep-water catshark – Holbiche papoila

*Galeus melastomus* Rafinesque, 1810 – Blackmouth catshark – Chien espagnol

*Galeus murinus* (Collett, 1904) – Mouse catshark – Chien islandais

Family Scyliorhinidae

*Scyliorhinus canicula* (Linnaeus, 1758) – Smallspotted catshark – Petite roussette

*Scyliorhinus stellaris* (Linnaeus, 1758) – Nursehound – Grande roussette

Family Pseudotriakidae

*Pseudotriakis microdon* de Brito Capello, 1868 – False catshark – Requin à longue dorsale

Family Triakidae

*Galeorhinus galeus* (Linnaeus, 1758) – Tope shark – Requin-hâ

*Mustelus asterias* Cloquet, 1819 – Starry smooth-hound – Émissole tachetée

*Mustelus mustelus* (Linnaeus, 1758) – Smooth-hound – Émissole lisse

*Mustelus punctulatus* Risso, 1827 – Blackspotted smooth-hound – Émissole pointillée

Family Carcharhinidae

*Carcharhinus brachyurus* (Günther, 1870) – Copper shark – Requin cuivre

*Carcharhinus brevipinna* (Valenciennes, 1839) – Spinner shark – Requin tisserand

*Carcharhinus limbatus* (Valenciennes, 1839) – Blacktip shark – Requin bordé

*Carcharhinus obscurus* (Lesueur, 1818) – Dusky shark – Requin de sable

*Carcharhinus plumbeus* (Nardo, 1827) – Sandbar shark – Requin gris

* *Galeocerdo cuvier* (Péron & Lesueur, 1822) – Tiger shark – Requin tigre

*Prionace glauca* (Linnaeus, 1758) – Blue shark – Peau bleue

Family Sphyrnidae

*Sphyrna lewini* (Griffith & Smith, 1834) – Scalloped hammerhead – Requin-marteau halicorne

*Sphyrna tudes* (Valenciennes, 1822)<fnref n="3" /> – Smalleye hammerhead – Requin-marteau à petits yeux

*Sphyrna zygaena* (Linnaeus, 1758) – Smooth hammerhead – Requin-marteau commun

Order SQUALIFORMES

Family Dalatiidae

*Dalatias licha* (Bonnaterre, 1788) – Kitefin shark – Squale liche

*Squaliolus laticaudus* Smith & Radcliffe, 1912 – Spined pygmy shark – Squale nain

Family Etmopteridae

*Etmopterus princeps* Collett, 1904 – Great lanternshark – Sagre rude

*Etmopterus pusillus* (Lowe, 1839) – Smooth lanternshark – Sagre nain

*Etmopterus spinax* (Linnaeus, 1758) – Velvet belly lantern shark – Sagre commun

Family Somniosidae

*Centroscymnus coelolepis* Barbosa du Bocage & de Brito Capello, 1864 – Portuguese dogfish – Pailona commun

*Centroselachus crepidater* (Barbosa du Bocage & de Brito Capello, 1864) – Longnose velvet dogfish – Pailona à long nez

*Scymnodon ringens* Barbosa du Bocage & de Brito Capello, 1864 – Knifetooth dogfish – Squale-grogneur commun

*Somniosus microcephalus* (Bloch & Schneider, 1801) – Greenland shark – Laimargue du Groenland

*Somniosus rostratus* (Risso, 1827) – Little sleeper shark – Laimargue de Méditerranée

Family Oxynotidae

*Oxynotus centrina* (Linnaeus, 1758) – Angular roughshark – Centrine commune

*Oxynotus paradoxus* Frade, 1929 – Sailfin roughshark – Humantin

Family Centrophoridae

*Centrophorus granulosus* (Bloch & Schneider, 1801) – Gulper shark – Squale-chagrin commun

*Centrophorus squamosus* (Bonnaterre, 1788) – Leafscale gulper shark – Squale-chagrin de l'Atlantique

*Deania calcea* (Lowe, 1839) – Birdbeak dogfish – Squale savate

*Deania profundorum* (Smith & Radcliffe, 1912) – Arrowhead dogfish – Squale-savate lutin

<footnote n="3">³ Most probably extinct in the French Mediterranean.</footnote>
## Family Squalidae

* *Squalus acanthias* Linnaeus, 1758 – Piked dogfish – Aiguillat commun

* *Squalus blainville* (Risso, 1827) – Longnose spurdog – Aiguillat coq

* *Squalus uyato* Rafinesque, 1810 – Little gulper shark – Petit squale-chagrin

## Family Echinorhinidae

* *Echinorhinus brucus* (Bonnaterre, 1788) – Bramble shark – Squale bouclé

## ORDER SQUATINIFORMES

### Family Squatinidae

* *Squatina aculeata* Cuvier, 1829⁴ – Sawback angelshark – Ange de mer épineux

* *Squatina oculata* Bonaparte, 1840 – Smoothback angelshark – Ange de mer ocellé

* *Squatina squatina* (Linnaeus, 1758) – Angelshark – Ange de mer commun

### Order TORPEDINIFORMES

#### Family Torpedinidae

* *Tetronarce nobiliana* (Bonaparte, 1835) – Atlantic torpedo – Torpille noire

* *Torpedo torpedo* (Linnaeus, 1758) – Common torpedo – Torpille ocellée

* *Torpedo marmorata* Risso, 1810 – Marbled torpedo – Torpille marbrée

### Order RHINOPRISTIFORMES

#### Family Rhinobatidae

* *Rhinobatos rhinobatos* (Linnaeus, 1758) – Common guitarfish – Raie-guitare commune

#### Family Glaucostegidae

* *Glaucostegus cemiculus* (Geoffroy Saint-Hilaire, 1817) – Blackchin guitarfish – Raie-guitare fouisseuse

#### Family Pristidae

* *Pristis pristis* (Linnaeus, 1758)<fnref n="5" /> – Common sawfish – Poisson-scie commun

* *Pristis pectinata* Latham, 1794⁶ – Smalltooth sawfish – Poisson-scie tident

### Order RAJIFORMES

#### Family Arhynchobatidae

* *Bathyraja pallida* (Forster, 1967) – Pale ray – Raie pâle

* *Bathyraja richardsoni* (Garrick, 1961) – Richardson's ray – Raie de Richardson

## Family Rajidae

* *Amblyraja radiata* (Donovan, 1808) – Starry ray – Raie radiée

* *Dipturus batis* (Linnaeus, 1758) – Blue skate – Pocheteau gris

* *Dipturus intermedius* (Parnell, 1837) – Intermediate skate – Pocheteau intermédiaire

* *Dipturus nidarosiensis* (Storm, 1881) – Norwegian skate – Pocheteau de Norvège

* *Dipturus oxyrinchus* (Linnaeus, 1758) – Longnose skate – Pocheteau noir

* *Leucoraja circularis* (Couch, 1838) – Sandy ray – Raie circulaire

* *Leucoraja fullonica* (Linnaeus, 1758) – Shagreen ray – Raie chardon

* *Leucoraja naevus* (Müller & Henle, 1841) – Cuckoo ray – Raie fleurie

* *Neoraja caerulea* (Stehmann, 1976) – Blue ray – Raie bleue

* *Raja asterias* Delaroche, 1809 – Starry ray – Raie étoilée

* *Raja brachyura* Lafont, 1873 – Blonde ray – Raie lisse

* *Raja clavata* Linnaeus, 1758 – Thornback ray – Raie bouclée

* *Raja microocellata* Montagu, 1818 – Small-eyed ray – Raie mêlée

* *Raja miraletus* Linnaeus, 1758 – Brown ray – Raie-miroir

* *Raja montagui* Fowler, 1910 – Spotted ray – Raie douce

* *Raja polystigma* Regan, 1923 – Speckled ray – Raie tachetée

* *Raja radula* Delaroche, 1809 – Rough ray – Raie râpe

* *Raja undulata* Lacepède, 1802 – Undulate ray – Raie brunette

* *Rajella bigelowi* (Stehmann, 1978) – Bigelow's ray – Raie à ventre noir

* *Rajella fyllae* (Lütken, 1887) – Round ray – Raie ronde

* *Rajella kukujevi* (Dolganov, 1985) – Mid-Atlantic skate – Raie pâlotte

* *Rostroraja alba* (Lacepède, 1803) – White skate – Raie blanche

### Order MYLIOBATIFORMES

#### Family Dasyatidae

* *Bathytoshia centroura* (Mitchill, 1815) – Roughtail stingray – Pastenague épineuse

* *Dasyatis pastinaca* (Linnaeus, 1758) – Common stingray – Pastenague commune

* *Dasyatis tortonesei* Capapé, 1975 – Tortonese's stingray – Pastenague de Tortonese

* *Pteroplatytrygon violacea* (Bonaparte, 1832) – Pelagic stingray – Pastenague violette

#### Family Gymnuridae

* *Gymnura altavela* (Linnaeus, 1758) – Spiny butterfly ray – Raie-papillon épineuse

<footnote n="4">⁴ Most probably extinct in the French Mediterranean.</footnote>

<footnote n="5">⁵ Most probably extinct in the French Mediterranean.</footnote>

<footnote n="6">⁶ Most probably extinct in the French Mediterranean.</footnote>
### Family Myliobatidae

* *Myliobatis aquila* (Linnaeus, 1758) – Common eagle ray – Aigle commun

* *Aetomylaeus bovinus* (Geoffroy Saint-Hilaire, 1817) – Bull ray – Aigle vachette

### Family Rhinopteridae

* *Rhinoptera marginata* (Geoffroy Saint-Hilaire, 1817) – Lusitanian cownose ray – Mourine lusitanienne

### Family Mobulidae

* *Mobula mobular* (Bonnaterre, 1788) – Devil ray – Mante

## CLASS HOLOCEPHALI

### Order CHIMAERIFORMES

#### Family Rhinochimaeridae

* *Harriotta raleighana* Goode & Bean, 1895 – Narrownose chimaera – Chimère de Raleigh

* *Rhinochimaera atlantica* Holt & Byrne, 1909 – Spearnose chimaera – Chimère à nez mou

#### Family Chimaeridae

* *Chimaera monstrosa* Linnaeus, 1758 – Rabbit fish – Chimère commune

* *Chimaera opalescens* Luchetti, Iglésias & Sellos, 2011 – Opale chimaera – Chimère opale

* *Hydrolagus affinis* (de Brito Capello, 1868) – Smalleyed rabbitfish – Chimère à petits yeux

* *Hydrolagus mirabilis* (Collett, 1904) – Large-eyed rabbitfish – Chimère à gros yeux

* *Hydrolagus pallidus* Hardy & Stehmann, 1990 – Pale rabbitfish – Chimère pâle

## CLASS ACTINOPTERYGII

### Order ACIPENSERIFORMES

#### Family Acipenseridae

* *Acipenser sturio* Linnaeus, 1758 – Sturgeon – Esturgeon

### Order ELOPIFORMES

#### Family Megalopidae

* *Megalops atlanticus* Valenciennes, 1847 – Tarpon – Tarpon atlantique

### Order NOTACANTHIFORMES

#### Family Halosauridae

* *Halosauropsis macrochir* (Günther, 1878) – Abyssal halosaur – Halosaure rayé

* *Halosaurus johnsonianus* Vaillant, 1888 – Halosaur – Halosaure

#### Family Notacanthidae

* *Notacanthus bonaparte* Risso, 1840 – Shortfin spiny eel – Poisson-tapir clair

* *Notacanthus chemnitzii* Bloch, 1788 – Snubnosed spiny eel – Poisson-tapir à grandes écailles

* *Polyacanthonotus challengeri* (Vaillant, 1888) – Longnose tapirfish – Poisson-tapir épineux

* *Polyacanthonotus rissoanus* (De Filippi & Verany, 1857) – Smallmouth spiny eel – Poisson-tapir à petites épines

### Order ANGUILLIFORMES

#### Family Synaphobranchidae

* *Simenchelys parasitica* Gill, 1879 – Snubnosed eel – Anguille à nez court

* *Synaphobranchus kaupii* Johnson, 1862 – Kaup's arrowtooth eel – Anguille égorgée

#### Family Muraenidae

* *Anarchias longicauda* (Peters, 1877) – Pygmy moray – Murène pygmée

* *Gymnothorax unicolor* (Delaroche, 1809) – Brown moray – Murène brune

* *Muraena helena* Linnaeus, 1758 – Mediterranean moray – Murène de Méditerranée

#### Family Derichthyidae

* *Derichthys serpentinus* Gill, 1884 – Narrownecked oceanic eel – Fouet petit-nez

* *Nessorhamphus ingolfianus* (Schmidt, 1912) – Duckbill oceanic eel – Fouet long-nez

#### Family Ophichthidae

* *Apterichtus anguiformis* (Peters, 1877) – Slender finless eel – Anguille-serpent à bouche noire

* *Apterichtus caecus* (Linnaeus, 1758) – European finless eel – Anguille-serpent aptère

* *Dalophis imberbis* (Delaroche, 1809) – Armless snake eel – Serpenton imberbe

* *Echelus myrus* (Linnaeus, 1758) – Painted eel – Serpenton miro

* *Ophichthus rufus* (Rafinesque, 1810) – Rufus snake eel – Serpenton jaune

* *Ophisurus serpens* (Linnaeus, 1758) – Serpent eel – Serpenton à long nez

* *Pisodonophis semicinctus* (Richardson, 1848) – Saddled snake eel – Serpenton à selles

#### Family Nettastomatidae

* *Nettastoma melanurum* Rafinesque, 1810 – Blackfin sorcerer – Sorcière à queue noire

#### Family Congridae

* *Ariosoma balearicum* (Delaroche, 1809) – Bandtooth conger – Congre des Baléares

* *Conger conger* (Linnaeus, 1758) – European conger – Congre d'Europe

* *Gnathophis mystax* (Delaroche, 1809) – Thinlip conger – Congre bec-fin

#### Family Nemichthyidae

* *Nemichthys scolopaceus* Richardson, 1848 – Slender snipe eel – Avocette ruban

#### Family Serrivomeridae

* *Serrivomer beanii* Gill & Ryder, 1883 – Stout sawpalate – Serrivomer trapu
### Family Anguillidae

*Anguilla anguilla* (Linnaeus, 1758) – European eel – Anguille d'Europe

### Order SACCOPHARYNGIFORMES

#### Family Eurypharyngidae

*Eurypharynx pelecanoides* Vaillant, 1882 – Pelican eel – Grand-gousier pélican

### Order CLUPEIFORMES

#### Family Clupeidae

*Alosa alosa* (Linnaeus, 1758) – Hallis shad – Grande alose

*Alosa fallax* (Lacepède, 1803) – Twaite shad – Alose feinte

*Clupea harengus* Linnaeus, 1758 – Atlantic herring – Hareng de l'Atlantique

*Sardina pilchardus* (Walbaum, 1792) – European pilchard – Sardine commune

*Sardinella aurita* Valenciennes, 1847 – Round sardinella – Allache

*Sprattus sprattus* (Linnaeus, 1758) – European sprat – Sprat

#### Family Engraulidae

*Engraulis albidus* Borsa, Collet & Durand, 2004 – White anchovy – Anchois blanc

*Engraulis encrasicolus* (Linnaeus, 1758) – European anchovy – Anchois

### Order OSMERIFORMES

#### Family Argentinidae

*Argentina silus* (Ascanius, 1775) – Greater argentine – Grande argentine

*Argentina sphyraena* Linnaeus, 1758 – Argentine – Petite argentine

*Glossanodon leioglossus* (Valenciennes, 1848) – Smalltoothed argentine – Argentine à petites dents

#### Family Microstomatidae

*Microstoma microstoma* (Risso, 1810) – Slender argentine – Serpe petite bouche

*Nansenia oblita* (Facciolà, 1887) – Forgotten argentine – Serpe oubliée

#### Family Bathylagidae

*Bathylagichthys greyae* (Cohen, 1958) – Grey's deepsea smelt – Garcette de Grey

*Bathylagus euryops* Goode & Bean, 1896 – Goiter blacksmelt – Garcette-goitre

*Dolicholagus longirostris* (Maul, 1948) – Longsnout blacksmelt – Garcette pâle

*Melanolagus bericoides* (Borodin, 1929) – Bigscale deepsea smelt – Garcette réticulée

#### Family Opisthoproctidae

*Dolichopteryx longipes* (Vaillant, 1888) – Brownsnout spookfish – Revenant télescope

*Opisthoproctus soleatus* Vaillant, 1888 – Barrel-eye – Revenant nocturne

### Family Alepocephalidae

*Alepocephalus agassizii* Goode & Bean, 1883 – Agassiz's slickhead – Alépocéphale obscur

*Alepocephalus bairdii* Goode & Bean, 1879 – Baird's slickhead – Alépocéphale de Baird

*Alepocephalus productus* Gill, 1883 – Smalley smooth-head – Alépocéphale émacié

*Alepocephalus rostratus* Risso, 1820 – Risso's smooth-head – Alépocéphale à bec

*Bathytroctes microlepis* Günther, 1878 – Smallscale smooth-head – Cassigné gros œil

*Conocara murrayi* (Koefoed, 1927) – Murray's smooth-head – Cassigné grosse tête

*Leptoderma macrops* Vaillant, 1886 – Grenadier smooth-head – Cassigné grenadier

*Narcetes stomias* (Gilbert, 1890) – Blackhead salmon – Cassigné à tête noire

*Photostylus pycnopterus* Beebe, 1933 – Starry smooth-head – Cassigné verruqueux

*Rouleina attrita* (Vaillant, 1888) – Softskin smooth-head – Cassigné noir d'encre

*Talismania antillarum* (Goode & Bean, 1896) – Antillean smooth-head – Cassigné ti-aile

*Xenodermichthys copei* (Gill, 1884) – Bluntsnout smooth-head – Gymnaste atlantique

#### Family Platyrotctidae

*Barbantus curvifrons* (Roule & Angel, 1931) – Palebelly searsid – Circé barbépine

*Holtbyrnia anomala* Krefft, 1980 – Bighead searsid – Circé armé

*Holtbyrnia macrops* Maul, 1957 – Bigeye searsid – Circé larme

*Maulisia argipalla* Matsui & Rosenblatt, 1979 – Palegold searsid – Circé troué

*Maulisia mauli* Parr, 1960 – Maul's searsid – Circé casqué

*Maulisia microlepis* Sazonov & Golovan, 1976 – Smallscale searsid – Circé éteint

*Mentodus rostratus* (Günther, 1878) – Beaked searsid – Circé barbe rude

*Normichthys operosus* Parr, 1951 – Multipore searsid – Circé troué

*Platyroctes apus* Günther, 1878 – Legless searsid – Circé élevé

*Sagamichthys schnakenbecki* (Krefft, 1953) – Schnakenbeck's searsid – Circé svelte

*Searsia koefoedi* Parr, 1937 – Koefoed's searsid – Circé commun

### Family Osmeridae

*Osmerus eperlanus* (Linnaeus, 1758) – European smelt – Éperlan européen
## Order SALMONIFORMES

### Family Salmonidae

*Salmo salar* Linnaeus, 1758 – Atlantic salmon – Saumon de l'Atlantique

*Salmo trutta* Linnaeus, 1758 – Sea trout – Truite de mer

## Order STOMIIFORMES

### Family Gonostomatidae

*Cyclothone acclinidens* Garman, 1899 – Benttooth bristlemouth – Cyclothone dentu

*Cyclothone braueri* Jespersen & Täning, 1926 – Garrick – Cyclothone de Brauer

*Cyclothone livida* Brauer, 1902 – Livid bristlemouth – Cyclothone livide

*Cyclothone microdon* (Günther, 1878) – Veiled anglemouth – Cyclothone brun

*Cyclothone pallida* Brauer, 1902 – Tan bristlemouth – Cyclothone à ventre clair

*Cyclothone pseudopallida* Mukhacheva, 1964 – Slender bristlemouth – Cyclothone à crâne ponctué

*Cyclothone pygmaea* Jespersen & Täning, 1926 – Pigmy bristlemouth – Cyclothone pygmée

*Gonostoma denudatum* Rafinesque, 1810 – Bigeye anglemouth – Gonostome gros œil

*Sigmops bathyphilus* (Vaillant, 1884) – Spark anglemouth – Gonostome noir

*Sigmops elongatus* (Günther, 1878) – Elongated bristlemouth – Gonostome à grandes dents

### Family Sternoptychidae

*Argyropelecus hemigymnus* Cocco, 1829 – Half-naked hatchetfish – Hache d'argent grande queue

*Argyropelecus olfersii* (Cuvier, 1829) – Silver hatchetfish – Hache d'argent commune

*Maurolicus muelleri* (Gmelin, 1789) – Silvery lightfish – Brossé améthyste

*Sternoptyx diaphana* Hermann, 1781 – Diaphanous hatchet fish – Hache d'argent diaphane

*Valenciennellus tripunctulatus* (Esmark, 1871) – Constellationfish – Brossé petits feux

### Family Phosichthyidae

*Polymetme corythaeola* (Alcock, 1898) – Plume lightfish – Luisant couronné

*Polymetme thaeocoryla* Parin & Borodulina, 1990 – Bright lightfish – Luisant grand feu

*Vinciguerria attenuata* (Cocco, 1838) – Slender lightfish – Luisant menton clair

*Vinciguerria poweriae* (Cocco, 1838) – Power's lightfish – Luisant nageoires claires

### Family Stomiidae

*Aristostomias tittmanni* Welsh, 1923 – Loosejaw – Dragagodet blanche tête

*Bathophilus nigerrimus* Giglioli, 1882 – Scaleless dragonfish – Dragon irisé

*Borostomias abyssorum* (Köhler, 1896) – Abyssal snaggletooth – Ledémon abyssal

*Borostomias antarcticus* (Lönnberg, 1905) – Snaggletooth – Ledémon double œil

*Chauliodus sloani* Bloch & Schneider, 1801 – Sloane's viperfish – Chauliode de Sloane

*Chirostomias pliopterus* Regan & Trewavas, 1930 – Branchedfin dragonfish – Dragon à nageoires ramifiées

*Flagellostomias boureei* (Zugmayer, 1913) – Longbarb dragonfish – Dragon à longue barbe

*Grammatostomias flagellibarba* Holt & Byrne, 1910 – Filamentousbarb dragonfish – Dragon à long filament

*Malacosteus niger* Ayres, 1848 – Spotlight loosejaw – Dragagodet imberbe

*Melanostomias bartonbeani* Parr, 1927 – Scaleless black dragonfish – Dragon à taches blanches

*Rhadinesthes decimus* (Zugmayer, 1911) – Slender snaggletooth – Dragon élancé

*Stomias boa* (Risso, 1810) – Boa dragonfish – Stomie commune

*Trigonolampa miriceps* Regan & Trewavas, 1930 – Threelight dragonfish – Dragon joue de feu

## Order ATELEOPODIFORMES

### Family Ateleopodidae

*Ijimaia loppei* Roule, 1922 – Loppe's tadpole fish – Atéléopode de Loppe

## Order AULOPIFORMES

### Family Aulopidae

*Aulopus filamentosus* (Bloch, 1792) – Royal flagfin – Limbert royal

### Family Chlorophthalmidae

*Chlorophthalmus agassizi* Bonaparte, 1840 – Shortnose greeneye – Éperlan du large

### Family Ipnopidae

*Bathypterois dubius* Vaillant, 1888 – Mediterranean spiderfish – Poisson-trépied grand noir

### Family Scopelarchidae

*Benthalbella infans* Zugmayer, 1911 – Zugmayer's pearleye – Œil-perlé à basse quille

### Family Notosudidae

*Scopelosaurus lepidus* (Krefft & Maul, 1955) – Blackfin waryfish – Guetteur à nageoire noire

### Family Synodontidae

*Synodus saurus* (Linnaeus, 1758) – Atlantic lizardfish – Poisson-lézard rayé

### Family Bathysauridae

*Bathysaurus ferox* Günther, 1878 – Deep-sea lizardfish – Bathysaure gris

### Family Paralepididae

*Arctozenus risso* (Bonaparte, 1840) – Spotted barracudina – Barracudine pintade
*Lestidiops affinis* (Ege, 1930) – Barracudina – Barracudine claire

*Lestidiops jayakari* (Boulenger, 1889) – Pacific barracudina – Barracudine à dos tacheté

*Lestidiops pseudosphyraenoides* (Ege, 1918) – Atlantic barracudina – Barracudine atlantique

*Lestidiops sphyrenoides* (Risso, 1820) – Shorthead barracudina – Barracudine à petite tête

*Macroparalepis affinis* Ege, 1933 – Naked barracudina – Barracudine nue

*Magnisudis atlantica* (Krøyer, 1868) – Duckbill barracudina – Barracudine à grande bouche

*Paralepis coregonoides* Risso, 1820 – Sharpchin barracudina – Barracudine petit bec

*Paralepis speciosa* Bellotti, 1878 – Mediterranean barracudina – Barracudine de Méditerranée

*Sudis hyalina* Rafinesque, 1810 – Barracudina – Barracudine à longues pectorales

### Family Anotopteridae

*Anotopterus pharao* Zugmayer, 1911 – Daggertooth – Pharaon

### Family Evermannellidae

*Evermannella balbo* (Risso, 1820) – Balbo sabretooth – Poisson à dents de sabre de Balbo

### Family Alepisauridae

*Alepisaurus ferox* Lowe, 1833 – Long snouted lancetfish – Lancier long-nez

## Order MYCTOPHIFORMES

### Family Neoscopelidae

*Neoscopelus macrolepidotus* Johnson, 1863 – Large-scaled lanternfish – Lanternon à grandes écailles

*Scopelengys tristis* Alcock, 1890 – Pacific blackchin – Lanternon à joue noire

### Family Myctophidae

*Benthosema glaciale* (Reinhardt, 1837) – Glacier lanternfish – Lanterne glaciaire

*Bolinichthys indicus* (Nafpaktitis & Nafpaktitis, 1969) – Smoothcheek lanternfish – Lanterne à joue lisse

*Bolinichthys supralateralis* (Parr, 1928) – Stubby lanternfish – Lanterne trapue

*Ceratoscopelus maderensis* (Lowe, 1839) – Madeira lanternfish – Lanterne de Madère

*Diaphus effulgens* (Goode & Bean, 1896) – Headlight lanternfish – Lanterne carrée

*Diaphus holti* Tåning, 1918 – Small lanternfish – Lanterne courte

*Diaphus metopoclampus* (Cocco, 1829) – Spothead lanternfish – Lanterne grosse tête

*Diaphus rafinesquii* (Cocco, 1838) – White-spotted lanternfish – Lanterne épaulette d'argent

*Electrona risso* (Cocco, 1829) – Electric lanternfish – Étincelle

*Hygophum benoiti* (Cocco, 1838) – Benoit's lanternfish – Lanterne de Benoit

*Hygophum hygomii* (Lütken, 1892) – Bermuda lanternfish – Lanterne des Bermudes

*Lampadena atlantica* Maul, 1969 – Atlantic sunbeam lanternfish – Lanterne soleil atlantique

*Lampadena chavesi* Collett, 1905 – Chaves' lanternfish – Lanterne de Chaves

*Lampadena luminosa* (Garman, 1899) – Luminous lanternfish – Lanterne lumineuse

*Lampadena speculigera* Goode & Bean, 1896 – Mirror lanternfish – Lanterne costumée

*Lampanyctus crocodilus* (Risso, 1810) – Jewel lanternfish – Lanterne crocodile

*Lampanyctus intricarius* Tåning, 1928 – Diamondcheek lanternfish – Lanterne à joue pailletée

*Lampanyctus festivus* Tåning, 1928 – Festive lanternfish – Lanterne festive

*Lampanyctus macdonaldi* (Goode & Bean, 1896) – MacDonald's lanternfish – Lanterne de MacDonald

*Lampanyctus pusillus* (Johnson, 1890) – Pygmy lanternfish – Lanterne pygmée

*Lobianchia dofleini* (Zugmayer, 1911) – Doflein's lanternfish – Lanterne de Doflein

*Lobianchia gemellarii* (Cocco, 1838) – Gemellar's lanternfish – Lanterne grande gueule

*Myctophum punctatum* Rafinesque, 1810 – Spotted lanternfish – Lanterne ponctuée

*Nannobrachium atrum* (Tåning, 1928) – Shortwing lanternfish – Lanterne à pectorale courte

*Notolychnus valdiviae* (Brauer, 1904) – Topside lanternfish – Lanterne de Valdivia

*Notoscopelus bolini* Nafpaktitis, 1975 – Bolin's lanternfish – Lanterne de Bolin

*Notoscopelus caudispinosus* (Johnson, 1863) – Spinetail lanternfish – Lanterne à queue épineuse

*Notoscopelus elongatus* (Costa, 1844) – Mediterranean lanternfish – Lanterne de Méditerranée

*Notoscopelus kroyeri* (Malm, 1861) – Krøyer's lanternfish – Lanterne de Krøyer

*Protomyctophum arcticum* (Lütken, 1892) – Arctic lanternfish – Lanterne arctique

*Symbolophorus veranyi* (Moreau, 1888) – Large-scale lanternfish – Lanterne à grandes écailles

*Taaningichthys bathyphilus* (Tåning, 1928) – Deepwater lanternfish – Lanterne à front lisse

## Order LAMPRIFORMES

### Family Lampridae

*Lampris guttatus* (Brünnich, 1788) – Opah – Opah

### Family Lophotidae

*Lophotus lacepede* Giorna, 1809 – Crested oarfish – Lophote
## Family Trachipteridae

*Trachipterus arcticus* (Brünnich, 1788) – Dealfish – Trachyptère atlantique

*Trachipterus trachypterus* (Gmelin, 1789) – Mediterranean dealfish – Trachyptère commun

*Zu cristatus* (Bonelli, 1819) – Scalloped ribbonfish – Trachyptère ventru

## Family Regalecidae

*Regalecus glesne* Ascanius, 1772 – King of herrings – Roi des harengs

## Order GADIFORMES

### Family Macrouridae

*Bathygadus melanobranchus* Vaillant, 1888 – Vaillant's grenadier – Grenadier de Vaillant

*Cetonurus globiceps* (Vaillant, 1884) – Globehead grenadier – Grenadier à tête globuleuse

*Coelorinchus caelorhincus* (Risso, 1810) – Hollowsnout grenadier – Grenadier raton

*Coelorinchus caudani* (Köhler, 1896) – Shortsnout grenadier – Grenadier petit nez

*Coelorinchus labiatus* (Köhler, 1896) – Spearsnouted grenadier – Grenadier long nez

*Coryphaenoides armatus* (Hector, 1875) – Abyssal grenadier – Grenadier abyssal

*Coryphaenoides brevibarbis* (Goode & Bean, 1896) – Shortbeard grenadier – Grenadier à barbe courte

*Coryphaenoides carapinus* Goode & Bean, 1883 – Carapine grenadier – Grenadier à barbillon court

*Coryphaenoides guentheri* (Vaillant, 1888) – Günther's grenadier – Grenadier de Günther

*Coryphaenoides leptolepis* Günther, 1877 – Ghostly grenadier – Grenadier petit front

*Coryphaenoides mediterraneus* (Giglioli, 1893) – Mediterranean grenadier – Grenadier camus

*Coryphaenoides rupestris* Gunnerus, 1765 – Roundnose grenadier – Grenadier de roche

*Coryphaenoides zaniophorus* (Vaillant, 1888) – Thickbeard grenadier – Grenadier barbichu

*Hymenocephalus italicus* Giglioli, 1884 – Glasshead grenadier – Grenadier plexiglas

*Kumba dentoni* Marshall, 1973 – Naked-head grenadier – Grenadier à tête nue

*Malacocephalus laevis* (Lowe, 1843) – Softhead grenadier – Grenadier barbu

*Nezumia aequalis* (Günther, 1878) – Common Atlantic grenadier – Grenadier lisse

*Nezumia sclerohynchus* (Valenciennes, 1838) – Roughtip grenadier – Grenadier imberbe

*Trachyrincus scabrus* (Rafinesque, 1810) – Roughsnout grenadier – Grenadier rude

## Family Moridae

*Antimora rostrata* (Günther, 1878) – Blue antimora – Moro bleu

*Eretmophorus kleinenbergi* Giglioli, 1889 – Oared mora – Moro rameux

*Gadella maraldi* (Risso, 1810) – Gadella – Moro imberbe

*Halargyreus johnsonii* Günther, 1862 – Slender codling – Moro svelte

*Lepidion guentheri* (Giglioli, 1880) – Günther's codling – Moro de Günther

*Lepidion lepidion* (Risso, 1810) – Mediterranean codling – Moro de Méditerranée

*Mora moro* (Risso, 1810) – Common mora – Moro commun

*Physiculus dalwigki* Kaup, 1858 – Black codling – Moro barbiche

## Family Melanonidae

*Melanonus zugmayeri* Norman, 1930 – Arrowtail – Mélanone

### Family Gadidae

*Gadiculus argenteus* Guichenot, 1850 – Silvery pout – Merlan argenté

*Gadus morhua* Linnaeus, 1758 – Atlantic cod – Morue de l'Atlantique

*Melanogrammus aeglefinus* (Linnaeus, 1758) – Haddock – Églefin

*Merlangius merlangus* (Linnaeus, 1758) – Whiting – Merlan

*Micromesistius poutassou* (Risso, 1827) – Blue whiting – Merlan bleu

*Pollachius pollachius* (Linnaeus, 1758) – Pollack – Lieu jaune

*Pollachius virens* (Linnaeus, 1758) – Saithe – Lieu noir

*Raniceps raninus* (Linnaeus, 1758) – Tadpole fish – Trident

*Trisopterus capelanus* (Lacepède, 1800) – Mediterranean poor cod – Capelan de Méditerranée

*Trisopterus esmarkii* (Nilsson, 1855) – Norway pout – Tacaud norvégien

*Trisopterus luscus* (Linnaeus, 1758) – Pouting – Tacaud commun

*Trisopterus minutus* (Linnaeus, 1758) – Poor cod – Capelan de l'Atlantique

## Family Lotidae

*Ciliata mustela* (Linnaeus, 1758) – Fivebeard rockling – Motelle à cinq barbillons

*Ciliata septentrionalis* (Collett, 1875) – Northern rockling – Motelle nordique

*Enchelyopus cimbrius* (Linnaeus, 1766) – Fourbeard rockling – Motelle à quatre barbillons

*Gaidropsarus biscayensis* (Collett, 1890) – Spotted rockling – Motelle mouchetée

*Gaidropsarus macrophthalmus* (Günther, 1867) – Bigeye rockling – Motelle à grands yeux
*Gaidropsarus mediterraneus* (Linnaeus, 1758) – Shore rockling – Motelle de Méditerranée

*Gaidropsarus vulgaris* (Cloquet, 1824) – Three-bearded rockling – Motelle commune

*Molva dypterygia* (Pennant, 1784) – Blue ling – Lingue bleue

*Molva macrophthalma* (Rafinesque, 1810) – Spanish ling – Lingue espagnole

*Molva molva* (Linnaeus, 1758) – Ling – Lingue franche

### Family Phycidae

*Phycis blennoides* (Brünnich, 1768) – Greater forkbeard – Mostelle de fond

*Phycis phycis* (Linnaeus, 1766) – Forkbeard – Mostelle de roche

### Family Merlucciidae

*Lyconus brachycolus* Holt & Byrne, 1906 – Grenadier hake – Merlu grenadier

*Merluccius merluccius* (Linnaeus, 1758) – European hake – Merlu européen

## Order OPHIDIIFORMES

### Family Ophidiidae

*Benthocometes robustus* (Goode & Bean, 1886) – Yellow cusk-eel – Fanfre jaune

*Lamprogrammus shcherbachevi* Cohen & Rohr, 1993 – Scaleline cusk-eel – Donzelle grand peigne

*Ophidion barbatum* Linnaeus, 1758 – Snake blenny – Donzelle à nageoires noires

*Ophidion rochei* Müller, 1845 – Roche's snake blenny – Donzelle à nageoires jaunes

*Parophidion vassali* (Risso, 1810) – Red snake blenny – Donzelle à nageoires rouges

*Spectrunculus grandis* (Günther, 1877) – Pudgy cusk-eel – Donzelle broche

### Family Parabrotulidae

*Parabrotula plagiophthalma* Zugmayer, 1911 – False cusk-eel – Gélatinette sombre

### Family Carapidae

*Carapus acus* (Brünnich, 1768) – Pearlfish – Aurin

*Echiodon dentatus* (Cuvier, 1829) – Toothed pearlfish – Fierasfer denté

*Echiodon drummondii* Thompson, 1837 – Atlantic pearlfish – Fierasfer atlantique

### Family Bythitidae

*Cataetyx alleni* (Byrne, 1906) – Allen's brotula – Donzelette de Allen

*Cataetyx laticeps* Koefoed, 1927 – Widehead brotula – Grande donzelette

*Grammonus ater* (Risso, 1810) – Black brotula – Fanfre noir

*Melodichthys hadrocephalus* Nielsen & Cohen, 1986 – Smalltail brotula – Donzelette petite queue

## Order BATRACHOIDIFORMES

### Family Batrachoididae

*Halobatrachus didactylus* (Bloch & Schneider, 1801) – Lusitanian toadfish – Poisson-crapaud lusitanien

## Order LOPHIIFORMES

### Family Lophiidae

*Lophius budegassa* Spinola, 1807 – Blackbellied angler – Baudroie rousse

*Lophius piscatorius* Linnaeus, 1758 – Angler – Baudroie commune

### Family Chaunacidae

*Chaunax pictus* Lowe, 1846 – Pink frogmouth – Baudroie rose

### Family Oneirodidae

*Lophodolos acanthognathus* Regan, 1925 – Whalehead dreamer – Rêveur à front haut

*Oneirodes carlsbergi* (Regan & Trewavas, 1932) – Longtail dreamer – Rêveur à grande queue

*Oneirodes eschrichtii* Lütken, 1871 – Cosmopolitan dreamer – Rêveur cosmopolite

### Family Ceratiidae

*Ceratias holboelli* Krøyer, 1845 – Kroyer's deep-sea angler fish – Noah

## Order GOBIESOCIFORMES

### Family Gobiesocidae

*Apletodon dentatus* (Facciolà, 1887) – Small-headed clingfish – Porte-écuelle à petite tête

*Apletodon incognitus* Hofrichter & Patzner, 1997 – Sea urchin clingfish – Porte-écuelle des oursins

*Diplecogaster bimaculata* (Bonnaterre, 1788) – Two spotted clingfish – Porte-écuelle à deux taches

*Gouania willdenowi* (Risso, 1810) – Blunt-snouted clingfish – Porte-écuelle jaune

*Lepadogaster candolii* Risso, 1810 – Connemarra clingfish – Lépadogastère de Candolle

*Lepadogaster lepadogaster* (Bonnaterre, 1788) – Shore clingfish – Lépadogastère commun

*Lepadogaster purpurea* (Bonnaterre, 1788) – Cornish clingfish – Lépadogastère pourpre

*Opeatogenys gracilis* (Canestrini, 1864) – Spindly clingfish – Porte-écuelle grêle

## Order ATHERINIFORMES

### Family Atherinidae

*Atherina boyeri* Risso, 1810 – Big-scale sand smelt – Joël

*Atherina hepsetus* Linnaeus, 1758 – Mediterranean sand smelt – Sauclet

*Atherina presbyter* Cuvier, 1829 – Sand smelt – Prêtre
## Order BELONIFORMES

### Family Scomberesocidae

* *Scomberesox saurus* (Walbaum, 1792) – Atlantic saury – Balaou atlantique

### Family Belonidae

* *Belone belone* (Linnaeus, 1760) – Garfish – Orphie

* *Belone svetovidovi* Collette & Parin, 1970 – Short-beaked garfish – Aiguille

* *Tylosurus imperialis* (Rafinesque, 1810) – Agujón needlefish – Aiguille voyeuse

### Family Exocoetidae

* *Cheilopogon cyanopterus* (Valenciennes, 1847) – Margined flyingfish – Exocet codène

* *Cheilopogon heterurus* (Rafinesque, 1810) – Blotchwing flyingfish – Exocet grande aile

* *Exocoetus volitans* Linnaeus, 1758 – Tropical two-wing flyingfish – Exocet commun

* *Hirundichthys rondeletii* (Valenciennes, 1847) – Black wing flyingfish – Exocet aile noire

* *Hirundichthys speculiger* (Valenciennes, 1847) – Mirrorwing flyingfish – Exocet miroir

## Order STEPHANOBERYCIFORMES

### Family Melamphaidae

* *Poromitra capito* Goode & Bean, 1883 – Ridgehead – Lubin rhino

* *Poromitra crassiceps* (Günther, 1878) – Crested bigscale – Lubin la crête

* *Poromitra megalops* (Lütken, 1878) – Largeye bigscale – Lubin gros œil

* *Scopelogadus beanii* (Günther, 1887) – Bean's bigscale – Lubin de Bean

## Order BERYCIFORMES

### Family Trachichthyidae

* *Hoplostethus atlanticus* Collett, 1889 – Orange roughy – Hoplostète orange

* *Hoplostethus cadenati* Quéro, 1974 – Black slimehead – Hoplostète noir

* *Hoplostethus mediterraneus* Cuvier, 1829 – Mediterranean slimehead – Hoplostète argenté

### Family Anoplogastridae

* *Anoplogaster cornuta* (Valenciennes, 1833) – Common fangtooth – Ogre

### Family Berycidae

* *Beryx decadactylus* Cuvier, 1829 – Alfonsino – Béryx commun

* *Beryx splendens* Lowe, 1834 – Splendid alfonsino – Béryx élancé

## Order CETOMIMIFORMES

### Family Rondeletiidae

* *Rondeletia loricata* Abe & Hotta, 1963 – Redmouth whalefish – Rondelet diable

## Order ZEIFORMES

### Family Oreosomatidae

* *Allocttus verrucosus* (Gilchrist, 1906) – Warty dory – Arrose verruqueuse

* *Neocyttus helgae* (Holt & Byrne, 1908) – False boarfish – Arrose sourcilleuse

### Family Parazenidae

* *Cyttopsis rosea* (Lowe, 1843) – Rosy dory – Saint-Pierre rouge

### Family Grammicolepididae

* *Grammicolepis brachiusculus* Poey, 1873 – Thorny tinselfish – Gramolette

### Family Zeidae

* *Zenopsis conchifer* (Lowe, 1852) – Silvery John dory – Saint-Pierre argenté

* *Zeus faber* Linnaeus, 1758 – John dory – Saint-Pierre

## Order GASTEROSTEIFORMES

### Family Gasterosteidae

* *Gasterosteus aculeatus* Linnaeus, 1758 – Three-spined stickleback – Épinoche à trois épines

* *Pungitius pungitius* (Linnaeus, 1758) – Ninespine stickleback – Épinochette

* *Spinachia spinachia* (Linnaeus, 1758) – Sea stickleback – Épinoche de mer

## Order SYNGNATHIFORMES

### Family Fistulariidae

* *Fistularia commersonii* Rüppell, 1838 – Bluespotted cornetfish – Poisson-flûte à taches bleues

### Family Centriscidae

* *Macroramphosus scolopax* (Linnaeus, 1758) – Longspine snipefish – Bécasse de mer

### Family Syngnathidae

* *Entelurus aequoreus* (Linnaeus, 1758) – Snake pipefish – Entélure

* *Hippocampus guttulatus* Cuvier, 1829 – Long-snouted seahorse – Hippocampe à long nez

* *Hippocampus hippocampus* (Linnaeus, 1758) – Short snouted seahorse – Hippocampe à nez court

* *Nerophis lumbriciformis* (Jenyns, 1835) – Worm pipefish – Nérophis lombricoïde

* *Nerophis maculatus* Rafinesque, 1810 – Spotted pipefish – Nérophis tacheté

* *Nerophis ophidion* (Linnaeus, 1758) – Straightnose pipefish – Nérophis à nez droit

* *Syngnathus abaster* Risso, 1827 – Black-striped pipefish – Syngnathe de lagune
*Syngnathus acus* Linnaeus, 1758 – Greater pipefish –
Sygnathe aiguille

*Syngnathus phlegon* Risso, 1827 – Spiny pipefish –
Sygnathe épineux

*Syngnathus rostellatus* Nilsson, 1855 – Lesser pipefish –
Sygnathe perçat

*Syngnathus tenuirostris* Rathke, 1837 – Narrow-snouted
pipefish – Sygnathe à nez grêle

*Syngnathus typhle* Linnaeus, 1758 – Broadnosed pipefish –
Siphonostome

## Order SCORPAENIFORMES

### Family Sebastidae

*Helicolenus dactylopterus* (Delaroche, 1809) – Blackbelly
rosefish – Sébaste chèvre

*Sebastes norvegicus* (Ascanius, 1772) – Golden redfish –
Sébaste doré

*Trachyscorpia echinata* (Köhler, 1896) – Spiny scorpionfish –
Rascasse épineuse

### Family Scorpaenidae

*Scorpaena elongata* Cadenat, 1943 – Slender rockfish –
Rascasse rose

*Scorpaena loppei* Cadenat, 1943 – Cadenat's rockfish –
Rascasse de Cadenat

*Scorpaena maderensis* Valenciennes, 1833 – Madeira
rockfish – Rascasse de Madère

*Scorpaena notata* Rafinesque, 1810 – Small red scorpionfish –
Rascasse pustuleuse

*Scorpaena porcus* Linnaeus, 1758 – Black scorpionfish –
Rascasse brune

*Scorpaena scrofa* Linnaeus, 1758 – Red scorpionfish –
Rascasse rouge

### Family Dactylopteridae

*Dactylopterus volitans* (Linnaeus, 1758) – Flying gurnard –
Grondin volant

### Family Triglidae

*Chelidonichthys cuculus* (Linnaeus, 1758) – Red gurnard –
Grondin rouge

*Chelidonichthys lucerna* (Linnaeus, 1758) – Tub gurnard –
Grondin perlon

*Chelidonichthys obscurus* (Walbaum, 1792) – Longfin
gurnard – Grondin sombre

*Eutrigla gurnardus* (Linnaeus, 1758) – Grey gurnard –
Grondin gris

*Lepidotrigla cavillone* (Lacepède, 1801) – Large-scaled
gurnard – Grondin cavillone

*Lepidotrigla dieuzeidei* Blanc & Hureau, 1973 – Spiny
gurnard – Grondin de Dieuzeide

*Trigla lyra* Linnaeus, 1758 – Piper gurnard – Grondin lyre

*Trigloporus lastoviza* (Bonnaterre, 1788) – Streaked gurnard – Grondin camard

### Family Peristediidae

*Peristedion cataphractum* (Linnaeus, 1758) – African
armoured searobin – Malarmat

### Family Cottidae

*Micrenophrys lilljeborgii* (Collett, 1875) – Norway bullhead –
Chabot nain

*Myoxocephalus scorpius* (Linnaeus, 1758) – Shorthorn
sculpin – Chaboisseau à épines courtes

*Taurulus bubalis* (Euphrasen, 1786) – Longspined bullhead –
Chabot de mer

### Family Psychrolutidae

*Cottunculus microps* Collett, 1875 – Polar sculpin – Cotte
polaire

*Cottunculus thomsonii* (Günther, 1882) – Pallid sculpin –
Cotte blême

### Family Agonidae

*Agonus cataphractus* (Linnaeus, 1758) – Hooknose – Souris
de mer

*Leptagonus decagonus* (Bloch & Schneider, 1801) –
Atlantic poacher – Agone atlantique

### Family Cyclopteridae

*Cyclopterus lumpus* Linnaeus, 1758 – Lumpfish – Lompe

### Family Liparidae

*Eutelichthys leptochirus* Tortonese, 1959 – Mediterranean
snailfish – Limace de Méditerranée

*Liparis liparis* (Linnaeus, 1766) – Striped seasnail – Limace
de mer

*Liparis montagui* (Donovan, 1804) – Montagu's seasnail –
Limace anicotte

## Order PERCIFORMES

### Family Moronidae

*Dicentrarchus labrax* (Linnaeus, 1758) – European seabass –
Bar

*Dicentrarchus punctatus* (Bloch, 1792) – Spotted seabass –
Bar tacheté

### Family Polyprionidae

*Polyprion americanus* (Bloch & Schneider, 1801) –
Wreckfish – Cernier commun

### Family Serranidae<fnref n="7" />

*Anthias anthias* (Linnaeus, 1758) – Swallowtail seaperch –
Barbier commun

*Epinephelus aeneus* (Geoffroy Saint-Hilaire, 1817) – White
grouper – Mérou blanc

*Epinephelus caninus* (Valenciennes, 1843) – Dogtooth
grouper – Mérou gris

<footnote n="7">⁷ Hyporthodus nigritus (Holbrook, 1855), cited by Quéro *et al.* (2000) is not included since only one large adult was recorded once from Brittany but never again. The occurrence of the species in Europe was also rejected by the IUCN Grouper Specialist Group.</footnote>
*Epinephelus costae* (Steindachner, 1878) – Goldblotch grouper – Mérou badèche

*Epinephelus marginatus* (Lowe, 1834) – Dusky grouper – Mérou brun

*Mycteroperca rubra* (Bloch, 1793) – Mottled grouper – Badèche rouge

*Serranus atricauda* Günther, 1874 – Blacktail comber – Serran à queue noire

*Serranus cabrilla* (Linnaeus, 1758) – Comber – Serran-chevrette

*Serranus hepatus* (Linnaeus, 1758) – Brown comber – Serran tambour

*Serranus scribe* (Linnaeus, 1758) – Painted comber – Serran écriture

### Family Callanthiidae

*Callanthias ruber* (Rafinesque, 1810) – Parrot seaperch – Barbier perroquet

### Family Apogonidae

*Apogon imberbis* (Linnaeus, 1758) – Cardinalfish – Apogon coq

### Family Epigonidae

*Epigonus constanciae* (Giglioli, 1880) – Spiny cardinalfish – Sonneur épineux

*Epigonus denticulatus* Dieuzeide, 1950 – Pencil cardinal – Sonneur nain

*Epigonus telescopus* (Risso, 1810) – Black cardinalfish – Sonneur commun

### Family Pomatomidae

*Pomatomus saltatrix* (Linnaeus, 1766) – Bluefish – Tassergal

### Family Echeneidae

*Echeneis naucrates* Linnaeus, 1758 – Live sharksucker – Grand rémora

*Phtheirichthys lineatus* (Menzies, 1791) – Slender suckerfish – Rémora grêle

*Remora brachyptera* (Lowe, 1839) – Spearfish remora – Rémora des espadons

*Remora remora* (Linnaeus, 1758) – Sharksucker – Rémora commun

### Family Carangidae

*Campogramma glaycos* (Lacepède, 1801) – Vadigo – Liche lirio

*Caranx crysos* (Mitchill, 1815) – Blue runner – Carangue coubali

*Caranx fischeri* Smith-Vaniz & Carpenter, 2007 – Longfin crevalle jack – Carangue de Fischer

*Decapterus macarellus* (Cuvier, 1833) – Mackerel scad – Comète maquereau

*Lichia amia* (Linnaeus, 1758) – Leerfish – Liche

*Naucrates ductor* (Linnaeus, 1758) – Pilotfish – Poisson pilote

*Pseudocaranx dentex* (Bloch & Schneider, 1801) – White trevally – Carangue dentue

**Seriola carpenteri* Mather, 1971 – Guinean amberjack – Sériole guinéenne

*Seriola dumerili* (Risso, 1810) – Greater amberjack – Sériole couronnée

**Seriola fasciata* (Bloch, 1793) – Lesser amberjack – Sériole babiane

*Seriola rivoliana* Valenciennes, 1833 – Longfin yellowtail – Sériole limon

*Trachinotus ovatus* (Linnaeus, 1758) – Pompano – Palomine

*Trachurus mediterraneus* (Steindachner, 1868) – Mediterranean horse mackerel – Chinchard à queue jaune

*Trachurus picturatus* (Bowdich, 1825) – Blue jack mackerel – Chinchard du large

*Trachurus trachurus* (Linnaeus, 1758) – Atlantic horse mackerel – Chinchard commun

### Family Coryphaenidae

*Coryphaena equiselis* Linnaeus, 1758 – Pompano dolphinfish – Coryphène dauphin

*Coryphaena hippurus* Linnaeus, 1758 – Common dolphinfish – Coryphène commune

### Family Bramidae

*Brama brama* (Bonnaterre, 1788) – Atlantic pomfret – Grande castagnole

*Pterycombus brama* Fries, 1837 – Atlantic fanfish – Castagnole voilier

*Taractes asper* Lowe, 1843 – Rough pomfret – Castagnole rugueuse

### Family Lobotidae

*Lobotes surinamensis* (Bloch, 1790) – Tripletail – Croupia roche

### Family Haemulidae

*Parapristipoma octolineatum* (Valenciennes, 1833) – African striped grunt – Grondeur rayé

*Pomadasys incisus* (Bowdich, 1825) – Bastard grunt – Grondeur métis

### Family Sparidae

*Boops boops* (Linnaeus, 1758) – Bogue – Bogue

*Dentex dentex* (Linnaeus, 1758) – Common dentex – Denti

*Dentex gibbosus* (Rafinesque, 1810) – Pink dentex – Denté bossu

*Dentex macrophthalmus* (Bloch, 1791) – Large-eye dentex – Denté à gros yeux

*Dentex maroccanus* Valenciennes, 1830 – Morocco dentex – Denté du Maroc

*Diplodus annularis* (Linnaeus, 1758) – Annular seabream – Sparaillon

*Diplodus cadenati* de la Paz, Bauchot & Daget, 1974 – Moroccan white seabream – Sar commun du Maroc

*Diplodus cervinus* (Lowe, 1838) – Zebra seabream – Sar tambour

*Diplodus puntazzo* (Walbaum, 1792) – Sharpsnout seabream – Sar à museau pointu
*Diplodus sargus* (Linnaeus, 1758) – White seabream – Sar commun

*Diplodus vulgaris* (Geoffroy Saint-Hilaire, 1817) – Common two-banded seabream – Sar à tête noire

*Lithognathus mormyrus* (Linnaeus, 1758) – Sand steenbras – Marbré

*Oblada melanura* (Linnaeus, 1758) – Saddled seabream – Oblade

*Pagellus acarne* (Risso, 1827) – Axillary seabream – Pageot acarné

*Pagellus bellottii* Steindachner, 1882 – Red pandora – Pageot à tache rouge

*Pagellus bogaraveo* (Brünnich, 1768) – Blackspot seabream – Pageot rose

*Pagellus erythrinus* (Linnaeus, 1758) – Common pandora – Pageot commun

*Pagrus caeruleostictus* (Valenciennes, 1830) – Bluespotted seabream – Pagre à points bleus

*Pagrus pagrus* (Linnaeus, 1758) – Red porgy – Pagre commun

*Sarpa salpa* (Linnaeus, 1758) – Salema – Saupe

*Sparus aurata* Linnaeus, 1758 – Gilthead seabream – Dorade royale

*Spondyliosoma cantharus* (Linnaeus, 1758) – Black seabream – Dorade grise

### Family Centracanthidae

*Centracanthus cirrus* Rafinesque, 1810 – Curled picarel – Picarel guetteur

*Spicara maena* (Linnaeus, 1758) – Blotched picarel – Mendole

*Spicara smaris* (Linnaeus, 1758) – Picarel – Picarel

### Family Sciaenidae

*Argyrosomus regius* (Asso, 1801) – Meagre – Maigre

*Sciaena umbra* Linnaeus, 1758 – Brown meagre – Corb

*Umbrina canariensis* Valenciennes, 1843 – Canary drum – Ombrine bronze

*Umbrina cirrosa* (Linnaeus, 1758) – Shi drum – Ombrine côtière

*Umbrina ronchus* Valenciennes, 1843 – Fusca drum – Ombrine fusca

### Family Mullidae

*Mullus barbatus* Linnaeus, 1758 – Red mullet – Rouget de vase

*Mullus surmuletus* Linnaeus, 1758 – Surmullet – Rouget de roche

### Family Kyphosidae

*Kyphosus sectatrix* (Linnaeus, 1758) – Bermuda sea chub – Calicagère blanche

### Family Cepolidae

*Cepola macrophthalma* (Linnaeus, 1758) – Red bandfish – Cépole

### Family Mugilidae

*Chelon labrosus* (Risso, 1827) – Thicklip grey mullet – Mulet lippu

*Liza aurata* (Risso, 1810) – Golden grey mullet – Mulet doré

*Liza ramada* (Risso, 1827) – Thinlip grey mullet – Mulet porc

*Liza saliens* (Risso, 1810) – Leaping mullet – Mulet sauteur

*Mugil cephalus* Linnaeus, 1758 – Flathead grey mullet – Mulet à grosse tête

*Oedalechilus labeo* (Cuvier, 1829) – Boxlip mullet – Mulet labéon

### Family Pomacentridae

*Chromis chromis* (Linnaeus, 1758) – Damselfish – Castagnole

### Family Labridae

*Acantholabrus palloni* (Risso, 1810) – Scale-rayed wrasse – Roucaou

*Centrolabrus exoletus* (Linnaeus, 1758) – Rock cook – Centrolabre

*Coris julis* (Linnaeus, 1758) – Rainbow wrasse – Girelle

*Ctenolabrus rupestris* (Linnaeus, 1758) – Goldsinny wrasse – Rouquié

*Labrus bergylta* Ascanius, 1767 – Ballan wrasse – Vieille commune

*Labrus merula* Linnaeus, 1758 – Brown wrasse – Merle

*Labrus mixtus* Linnaeus, 1758 – Cuckoo wrasse – Vieille coquette

*Labrus viridis* Linnaeus, 1758 – Green wrasse – Labre vert

*Lappanella fasciata* (Cocco, 1833) – Sharp-toothed wrasse – Labre iris

*Symphodus bailloni* (Valenciennes, 1839) – Baillon's wrasse – Crénilabre de Baillon

*Symphodus cinereus* (Bonnaterre, 1788) – Grey wrasse – Crénilabre cendré

*Symphodus doderleinii* Jordan, 1890 – Doderlein's wrasse – Crénilabre de Doderlein

*Symphodus mediterraneus* (Linnaeus, 1758) – Axillary wrasse – Crénilabre méditerranéen

*Symphodus melanocercus* (Risso, 1810) – Blacktailed wrasse – Crénilabre à queue noire

*Symphodus melops* (Linnaeus, 1758) – Corkwing wrasse – Crénilabre mélops

*Symphodus ocellatus* (Linnaeus, 1758) – Ocellated wrasse – Crénilabre ocellé

*Symphodus roissali* (Risso, 1810) – Five-spotted wrasse – Crénilabre à cinq taches

*Symphodus rostratus* (Bloch, 1791) – Pointed-snout wrasse – Sublet

*Symphodus tinca* (Linnaeus, 1758) – Peacock wrasse – Crénilabre paon

*Thalassoma pavo* (Linnaeus, 1758) – Ornate wrasse – Girelle paon
*Xyrichtys novacula* (Linnaeus, 1758) – Pearly razorfish –
Rason

**Family Scaridae**

*Sparisoma cretense* (Linnaeus, 1758) – Parrotfish –
Perroquet vieillard

**Family Zoarcidae**

*Lycenchelys alba* (Vaillant, 1888) – White eelpout – Lycode
blanche

*Melanostigma atlanticum* Koefoed, 1952 – Atlantic soft
pout – Mollase atlantique

*Pachycara bulbiceps* (Garman, 1899) – Snubnose eelpout –
Lycode obèse

*Pachycara crassiceps* (Roule, 1916) – Brown eelpout –
Lycode brun noir

*Zoarces viviparus* (Linnaeus, 1758) – European eelpout –
Loquette d'Europe

**Family Stichaeidae**

*Chirolophis ascanii* (Walbaum, 1792) – Yarrell's blenny –
Toupet marbré

**Family Pholidae**

*Pholis gunnellus* (Linnaeus, 1758) – Rock gunnel – Gonelle

**Family Anarhichadidae**

*Anarhichas denticulatus* Krøyer, 1845 – Northern wolffish –
Loup gélatineux

*Anarhichas lupus* Linnaeus, 1758 – Atlantic wolffish – Loup
atlantique

**Family Chiasmodontidae**

*Pseudoscopelus obtusifrons* (Fowler, 1934) – Swallowerfish
– Gargare

**Family Ammodytidae**

*Ammodytes marinus* Raitt, 1934 – Lesser sand-eel – Lançon
nordique

*Ammodytes tobianus* Linnaeus, 1758 – Small sand-eel –
Lançon équille

*Gymnammodytes cicerelus* (Rafinesque, 1810) –
Mediterranean sand-eel – Cicerelle de Méditerranée

*Gymnammodytes semisquamatus* (Jourdain, 1879) – Smooth
sand-eel – Cicerelle de l'Atlantique

*Hyperoplus immaculatus* (Corbin, 1950) – Greater sand-eel
– Lançon jolivet

*Hyperoplus lanceolatus* (Le Sauvage, 1824) – Great sand-
eel – Lançon commun

**Family Trachinidae**

*Echiichthys vipera* (Cuvier, 1829) – Lesser weever – Petite
vive

*Trachinus araneus* Cuvier, 1829 – Spotted weever – Vive-
araignée

*Trachinus draco* Linnaeus, 1758 – Greater weever – Grande
vive

*Trachinus radiatus* Cuvier, 1829 – Starry weever – Vive à
tête rayonnée

**Family Uranoscopidae**

*Uranoscopus scaber* Linnaeus, 1758 – Stargazer –
Uranoscope

**Family Tripterygiidae**

*Tripterygion delaisi* Cadenat & Blache, 1970 – Black-faced
blenny – Triptérygion jaune

*Tripterygion melanurum* Guichenot, 1850 – Black-spotted
triplefin – Triptérygion nain

*Tripterygion tripteronotum* (Risso, 1810) – Red-black
triplefin – Triptérygion à bec

**Family Clinidae**

*Clinitrachus argentatus* (Risso, 1810) – Cline – Cline
argenté

**Family Blenniidae**

*Aidablennius sphynx* (Valenciennes, 1836) – Sphinx blenny
– Blennie sphinx

*Blennius ocellaris* Linnaeus, 1758 – Butterfly blenny –
Blennie papillon

*Coryphoblennius galerita* (Linnaeus, 1758) – Montagu's
blenny – Blennie coiffée

*Lipophrys pholis* (Linnaeus, 1758) – Shanny – Blennie
mordocet

*Lipophrys trigloides* (Valenciennes, 1836) – Trigloid blenny
– Blennie trigloïde

*Microlipophrys canevae* (Vinciguerra, 1880) – Yellow cheek
blenny – Blennie de Caneva

*Microlipophrys dalmatinus* (Steindachner & Kolombatovic,
1883) – Dalmatian blenny – Blennie dalmate

*Microlipophrys nigriceps* (Vinciguerra, 1883) – Black-
headed blenny – Blennie à tête noire

*Parablennius gattorugine* (Linnaeus, 1758) – Tompot
blenny – Blennie gattorugine

*Parablennius incognitus* (Bath, 1968) – Mystery blenny –
Blennie diabolo

*Parablennius pilicornis* (Cuvier, 1829) – Ringneck blenny –
Blennie pilicorne

*Parablennius rouxi* (Cocco, 1833) – Long-stripped blenny –
Blennie de Roux

*Parablennius ruber* (Valenciennes, 1836) – Portuguese
blenny – Blennie rouge

*Parablennius sanguinolentus* (Pallas, 1814) – Rusty blenny
– Blennie palmicorne

*Parablennius tentacularis* (Brünnich, 1768) – Tentacled
blenny – Blennie tentaculée

*Parablennius zvonimiri* (Kolombatovic, 1892) – Zvonimir's
blenny – Blennie de Zvonimir

*Salaria basilisca* (Valenciennes, 1836) – Basilisk blenny –
Blennie basilic

*Salaria pavo* (Risso, 1810) – Peacock blenny – Blennie
paon

*Scartella cristata* (Linnaeus, 1758) – Molly miller – Blennie
chevelue
## Family Callionymidae

*Callionymus fasciatus* Valenciennes, 1837 – Banded dragonet – Dragonnet fascié

*Callionymus lyra* Linnaeus, 1758 – Dragonet – Dragonnet lyre

*Callionymus maculatus* Rafinesque, 1810 – Spotted dragonet – Dragonnet tacheté

*Callionymus pusillus* Delaroche, 1809 – Sailfin dragonet – Dragonnet élégant

*Callionymus reticulatus* Valenciennes, 1837 – Reticulated dragonet – Dragonnet réticulé

*Callionymus risso* Lesueur, 1814 – Risso's dragonet – Dragonnet de Risso

*Synchiropus phaeton* (Günther, 1861) – Phaeton dragonet – Dragonnet phaéton

## Family Gobiidae

*Aphia minuta* (Risso, 1810) – Transparent goby – Gobie nonnat

*Buenia affinis* Iljin, 1930 – De Buen's goby – Gobie réticulé

*Buenia jeffreysii* (Günther, 1867) – Jeffreys' goby – Gobie de Jeffreys

*Chromogobius quadrivittatus* (Steindachner, 1863) – Chestnut goby – Gobie à quatre bandes

*Chromogobius zebratus* (Kolombatovic, 1891) – Kolombatovic's goby – Gobie grêle zébré

*Corcyrogobius liechtensteini* (Kolombatovic, 1891) – Liechtenstein's goby – Gobie-corail nain

*Crystallogobius linearis* (Düben, 1845) – Crystal goby – Gobie cristal

*Deltentosteus collonianus* (Risso, 1820) – Toothed goby – Gobie denté

*Deltentosteus quadrimaculatus* (Valenciennes, 1837) – Four-spotted goby – Gobie à quatre taches

*Didogobius schlieweni* Miller, 1993 – Schliewen's goby – Gobie nain à taches blanches

*Didogobius splechtnai* Ahnelt & Patzner, 1995 – Splechtna's goby – Gobie nain à barres brunes

*Gammogobius steinitzi* Bath, 1971 – Steinitz's goby – Gobie de Steinitz

*Gobius ater* Bellotti, 1888 – Bellotti's goby – Gobie de Bellotti

*Gobius auratus* Risso, 1810 – Golden goby – Gobie doré

*Gobius bucchichi* Steindachner, 1870 – Bucchich's goby – Gobie moucheté

*Gobius cobitis* Pallas, 1814 – Giant goby – Gobie céphalote

*Gobius cruentatus* Gmelin, 1789 – Red-mouthed goby – Gobie ensanglanté

*Gobius fallax* Sarato, 1889 – Sarato's goby – Gobie de Sarato

*Gobius gasteveni* Miller, 1974 – Steven's goby – Gobie de Steven

*Gobius geniporus* Valenciennes, 1837 – Slender goby – Gobie à joues poreuses

*Gobius incognitus* Kovačić & Šanda, 2016 – Unknown goby – Gobie méconnu

*Gobius kolombatovici* Kovačić & Miller, 2000 – Kolombatovic's goby – Gobie à taches oranges

*Gobius niger* Linnaeus, 1758 – Black goby – Gobie noir

*Gobius paganellus* Linnaeus, 1758 – Rock goby – Gobie paganel

*Gobius roulei* de Buen, 1928 – Roule's goby – Gobie de Roule

*Gobius vittatus* Vinciguerra, 1883 – Striped goby – Gobie rayé

*Gobius xanthocephalus* Heymer & Zander, 1992 – Yellow-headed goby – Gobie à tête jaune

*Gobiusculus flavescens* (Fabricius, 1779) – Two-spotted goby – Gobie à deux taches

*Lebetus guilleti* (Le Danois, 1913) – Guillet's goby – Gobie nain de Guillet

*Lebetus scorpioides* (Collett, 1874) – Diminutive goby – Gobie nabot

*Lesueurigobius friesii* (Malm, 1874) – Fries' goby – Gobie de Fries

*Lesueurigobius suerii* (Risso, 1810) – Lesueur's goby – Gobie de Lesueur

*Millerigobius macrocephalus* (Kolombatović, 1891) – Large-headed goby – Gobie nain à grosse tête

*Odondebuenia balearica* (Pellegrin & Fage, 1907) – Coralline goby – Gobie des Baléares

*Pomatoschistus bathi* Miller, 1982 – Bath's goby – Gobie de Bath

*Pomatoschistus lozanoi* (de Buen, 1923) – Lozano's goby – Gobie de Lozano

*Pomatoschistus marmoratus* (Risso, 1810) – Marbled goby – Gobie marbré

*Pomatoschistus microps* (Krøyer, 1838) – Common goby – Gobie tacheté

*Pomatoschistus minutus* (Pallas, 1770) – Sand goby – Gobie des sables

*Pomatoschistus norvegicus* (Collett, 1902) – Norwegian goby – Gobie norvégien

*Pomatoschistus pictus* (Malm, 1865) – Painted goby – Gobie varié

*Pomatoschistus quagga* (Heckel, 1837) – Quagga goby – Gobie quagga

*Pseudaphya ferreri* (de Buen & Fage, 1908) – Ferrer's goby – Gobie-nageur de Ferrer

*Thorogobius ephippiatus* (Lowe, 1839) – Leopard-spotted goby – Gobie léopard

*Thorogobius macrolepis* (Kolombatovic, 1891) – Large-scaled goby – Gobie à grandes écailles

*Vanneaugobius dollfusi* Brownell, 1978 – Dollfus' goby – Gobie nain de Dollfus

*Zebrus zebrus* (Risso, 1827) – Zebra goby – Gobie zébré
*Zosterisessor ophiocephalus* (Pallas, 1814) – Grass goby –
Gobie lote

Family Siganidae

*Siganus luridus* (Rüppell, 1829) – Dusky spinefoot – Sigan
sombre

Family Luvaridae

*Luvarus imperialis* Rafinesque, 1810 – Luvar – Louvareau

Family Sphyraenidae

*Sphyraena sphyraena* (Linnaeus, 1758) – European barracuda – Bécune européenne

*Sphyraena viridensis* Cuvier, 1829 – Yellowmouth barracuda – Bécune à bouche jaune

Family Gempylidae

*Lepidocybium flavobrunneum* (Smith, 1843) – Escolar – Escolier noir

*Nesiarchus nasutus* Johnson, 1862 – Black gemfish – Escolier long nez

*Promethichthys prometheus* (Cuvier, 1832) – Roudi escolar – Escolier clair

*Ruvettus pretiosus* Cocco, 1833 – Oilfish – Rouvet

Family Trichiuridae

*Aphanopus carbo* Lowe, 1839 – Black scabbardfish – Poisson-sabre noir

*Benthodesmus simonyi* (Steindachner, 1891) – North Atlantic frostfish – Poisson-sabre ganse

*Lepidopus caudatus* (Euphrasen, 1788) – Silver scabbardfish – Poisson-sabre argenté

*Trichiurus lepturus* Linnaeus, 1758 – Largehead hairtail – Poisson-sabre commun

Family Scombridae

*Acanthocybium solandri* (Cuvier, 1832) – Wahoo – Thazard bâtard

*Auxis rochei* (Risso, 1810) – Bullet tuna – Bonitou

*Auxis thazard* (Lacepède, 1800) – Frigate tuna – Auxide

*Euthynnus alletteratus* (Rafinesque, 1810) – Little tunny – Thonine commune

*Katsuwonus pelamis* (Linnaeus, 1758) – Skipjack tuna – Listao

*Orcynopsis unicolor* (Geoffroy Saint-Hilaire, 1817) – Plain bonito – Palomette

*Sarda sarda* (Bloch, 1793) – Atlantic bonito – Bonite à dos rayé

*Scomber colias* Gmelin, 1789 – Atlantic chub mackerel – Maquereau espagnol

*Scomber scombrus* Linnaeus, 1758 – Atlantic mackerel – Maquereau commun

*Scomberomorus tritor* (Cuvier, 1832) – West African Spanish mackerel – Thazard blanc

*Thunnus alalunga* (Bonnaterre, 1788) – Albacore – Germon

*Thunnus obesus* (Lowe, 1839) – Bigeye tuna – Thon obèse

*Thunnus thynnus* (Linnaeus, 1758) – Atlantic bluefin tuna – Thon rouge de l'Atlantique

Family Xiphiidae

*Xiphias gladius* Linnaeus, 1758 – Swordfish – Espadon

Family Istiophoridae

*Istiophorus platypterus* (Shaw, 1792) – Sailfish – Voilier

*Kajikia albida* (Poey, 1860) – Atlantic white marlin – Makaire blanc de l'Atlantique

*Makaira nigricans* Lacepède, 1802 – Blue marlin – Makaire bleu

*Tetrapturus belone* Rafinesque, 1810 – Mediterranean spearfish – Marlin de Méditerranée

*Tetrapturus pfluegeri* Robins & de Sylva, 1963 – Longbill spearfish – Makaire bécune

Family Centrolophidae

*Centrolophus niger* (Gmelin, 1789) – Rudderfish – Centrolophe noir

*Hyperoglyphe perciformis* (Mitchill, 1818) – Barrelfish – Centrolophe épineux

*Schedophilus medusophagus* (Cocco, 1839) – Cornish blackfish – Rouffe des méduses

*Schedophilus ovalis* (Cuvier, 1833) – Imperial blackfish – Rouffe impérial

Family Nomeidae

*Cubiceps gracilis* (Lowe, 1843) – Driftfish – Dérivant grêle

*Psenes pellucidus* Lütken, 1880 – Bluefin driftfish – Dérivant gélatineux

Family Tetragonuridae

*Tetragonurus cuvieri* Risso, 1810 – Smalleye squaretail – Tétragonure de Cuvier

Family Stromateidae

*Stromateus fiatola* Linnaeus, 1758 – Blue butterfish – Fiahole

Family Caproidae

*Antigonia capros* Lowe, 1843 – Deepbody boarfish – Sanglier chevrette

*Capros aper* (Linnaeus, 1758) – Boarfish – Sanglier

Order PLEURONECTIFORMES

Family Citharidae

*Citharus linguatula* (Linnaeus, 1758) – Spotted flounder – Feuille

Family Scophthalmidae

*Lepidorhombus boscii* (Risso, 1810) – Four-spot megrim – Cardine à quatre taches

*Lepidorhombus whiffiagonis* (Walbaum, 1792) – Megrim – Cardine franche

*Phrynorhombus norvegicus* (Günther, 1862) – Norwegian topknot – Targie naine

*Scophthalmus maximus* (Linnaeus, 1758) – Turbot – Turbot

*Scophthalmus rhombus* (Linnaeus, 1758) – Brill – Barbue

*Zeugopterus punctatus* (Bloch, 1787) – Topknot – Targeur

*Zeugopterus regius* (Bonnaterre, 1788) – Eckström's topknot – Targie unimaculée
## Family Bothidae

* *Arnoglossus imperialis* (Rafinesque, 1810) – Imperial scaldfish – Arnoglosse impérial

* *Arnoglossus kessleri* Schmidt, 1915 – Scaldback – Arnoglosse de Kessler

* *Arnoglossus laterna* (Walbaum, 1792) – Scaldfish – Arnoglosse lanterne

* *Arnoglossus rueppelii* (Cocco, 1844) – Rüppell's scaldback – Arnoglosse de Rüppell

* *Arnoglossus thori* Kyle, 1913 – Thor's scaldfish – Arnoglosse de Thor

* *Bothus podas* (Delaroche, 1809) – Wide-eyed flounder – Rombou podas

## Family Pleuronectidae

* *Glyptocephalus cynoglossus* (Linnaeus, 1758) – Witch flounder – Plie cynoglosse

* *Hippoglossoides platessoides* (Fabricius, 1780) – American plaice – Balai

* *Hippoglossus hippoglossus* (Linnaeus, 1758) – Atlantic halibut – Flétan de l'Atlantique

* *Limanda limanda* (Linnaeus, 1758) – Common dab – Limande

* *Microstomus kit* (Walbaum, 1792) – Lemon sole – Limande-sole

* *Platichthys flesus* (Linnaeus, 1758) – European flounder – Flet d'Europe

* *Pleuronectes platessa* Linnaeus, 1758 – European plaice – Plie d'Europe

## Family Soleidae

* *Bathysolea profundicola* (Vaillant, 1888) – Deepwater sole – Sole de profondeur

* *Buglossidium luteum* (Risso, 1810) – Solenette – Petite sole jaune

* *Dicologlossa cuneata* (Moreau, 1881) – Wedge sole – Céteau

* *Microchirus azevia* (de Brito Capello, 1867) – Bastard sole – Sole-perdrix brune

* *Microchirus hexophthalmus* (Bennett, 1831) – Ocellated wedge sole – Sole-perdrix à six yeux

* *Microchirus ocellatus* (Linnaeus, 1758) – Foureyed sole – Sole-perdrix ocellée

* *Microchirus variegatus* (Donovan, 1808) – Thickback sole – Sole-perdix commune

* *Monochirus hispidus* Rafinesque, 1814 – Whiskered sole – Sole velue

* *Pegusa impar* (Bennett, 1831) – Adriatic sole – Sole-pole de Méditerranée

* *Pegusa lascaris* (Risso, 1810) – Sand sole – Sole-pole

* *Pegusa nasuta* (Pallas, 1814) – Blackhand sole – Sole-pole sombre

* *Solea senegalensis* Kaup, 1858 – Senegalese sole – Sole du Sénégal

* *Solea solea* (Linnaeus, 1758) – Common sole – Sole commune

* *Synapturichthys kleinii* (Risso, 1827) – Klein's sole – Sole tachetée

## Family Cynoglossidae

* *Symphurus ligulatus* (Cocco, 1844) – Elongate tonguesole – Plagusie longue

* *Symphurus nigrescens* Rafinesque, 1810 – Spotted tonguesole – Plagusie sombre

## Order TETRAODONTIFORMES

### Family Balistidae

* *Balistes capriscus* Gmelin, 1789 – Grey triggerfish – Baliste cabri

### Family Monacanthidae

* *Aluterus monoceros* (Linnaeus, 1758) – Unicorn leatherjacket – Bourse loulou

### Family Tetraodontidae

* *Lagocephalus lagocephalus* (Linnaeus, 1758) – Oceanic puffer – Compère océanique

* *Sphoeroides pachygaster* (Müller & Troschel, 1848) – Blunthead puffer – Compère émoussé

### Family Molidae

* *Mola mola* (Linnaeus, 1758) – Ocean sunfish – Poisson-lune

* *Ranzania laevis* (Pennant, 1776) – Slender sunfish – Poisson-lune tronqué

*Note:* The fish classification is being revised on the light of several recent works and re-assessments. Catalog of Fishes has adopted a new one in its last online version dated 1st November 2017 at a time when the paper was already accepted. We recommend to the users of this list to be cautious when referring to the classification we used in this paper. However new changes will be progressively integrated in the list available from the INPN website.

**Acknowledgements.** – We would like to thank the INPN for funding and the SFI for supervising the project. We are indebted to several persons who helped us during the course of the study, especially Salomé Fabri-Ruiz, Maria Mateo-Santos, Pierre Noël and Charles Muñoz-Nate.

## REFERENCES

ESCHMEYER W.N., FRICKE R. & VAN DER LAAN R. (eds), 2017. - Catalog of Fishes: Genera, species, references. (http://researcharchive.calacademy.org/research/ichthyology/catalog/fishcatmain.asp). Electronic version accessed 22 Mar. 2017.

FROESE R. & PAULY D. (eds), 2017. - FishBase. World Wide Web electronic publication. www.fishbase.org, version Feb. 2017.

GARGOMINY O., TERCERIE S., RÉGNIER C., RAMAGE T., SCHOELINCK C., DUPONT P., VANDEL E., DASZKIEWICZ P. & PONCET L., 2016. - TAXREF v10.0, référentiel taxonomique pour la France : méthodologie, mise en œuvre et
diffusion. Rapport SPN 2016-101. Paris: MNHN. https://inpn.mnhn.fr/programme/referentiel-taxonomique-taxref

GERVAIS H. & BOULART R., 1876-1877. - Les poissons : Synonymie, Description, Mœurs, Frai, Pêche, Iconographie, des Espèces composant plus particulièrement la Faune française. 2 parties en 3 volumes, 232 p., 300 p., 292 p., 260 pl. Paris: J. Rothschild.

HUREAU J.C. & MONOD T. (eds), 1973-1979. - Check-List of the Fishes of the North-Eastern Atlantic and of the Mediterranean (CLOFNAM I and II). 2 vol. Paris: UNESCO.

KEITH P., PERSAT H., FEUNTEUN É. & ALLARDI J. (Coord.), 2011. - Les Poissons d'Eau douce de France. Collection Inventaires et Biodiversité. 552 p. Mèze & Paris: Biotope - Muséum national d'Histoire naturelle.

LOUISY P., 2015. - Europe and Mediterranean Marine Fish Identification Guide. 512 p. Paris: Les Éditions Ulmer.

MOREAU E., 1881. - Histoire naturelle des Poissons de la France. 3 vol., 478 p., 572 p., 697 p. Paris: Masson.

QUÉRO J.C., DU BUIT H., LABORDE J.L. & VAYNE J.J., 2000. - Observations ichthyologiques effectuées en 1999. Ann. Soc. Sci. Nat. Charente-Mar., 8(9): 1039-1045.

QUÉRO J.C., PORCHÉ P. & VAYNE J.J., 2003. - Guide des Poissons de l'Atlantique européen. 465 p. Les Guides du naturaliste. Lonay & Paris: Delachaux et Niestlé.

TRENKEL V., BERTHELE O., LORANCE P. et al. [14 authors], 2009. - Atlas des grands Invertébrés et Poissons observés par les Campagnes scientifiques. Atlas de distribution. Bilan 2008. 100 p. Nantes: Ifremer.

UNITED NATIONS, 1992. - Convention on biological Diversity. Rio-de-Janeiro, 22 May 1992, 28 p. https://www.cbd.int/doc/legal/cbd-en.pdf

VAN DER LAAN R., ESCHMEYER W.N. & FRICKE R., 2017. - Family-group names. (http://www.calacademy.org/scientists/catalog-of-fishes-family-group-names/). Electronic version accessed 22 Mar. 2017.

VAN DER LAND J., COSTELLO M.J., ZAVODNIK D., SANTOS R.S., PORTEIRO F.M., BAILLY N., ESCHMEYER W.N. & FROESE R., 2001. - Pisces. In: European Register of Marine Species. A check-list of the marine species in Europe and a bibliography of guides to their identification (Costello M.J., Emblow C.S. & White R., eds), Patrimoines naturels, 50: 357-374. Paris: MNHN.

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Annex. - Complementary list of brackish and freshwater species that only enter sporadically the marine biota. Non-native species are indicated with an asterisk (*)

## CLASS ACTINOPTERYGII
### Order ACIPENSERIFORMES

#### Family Acipenseridae

*Acipenser baerii* Brandt, 1869 - Siberian sturgeon - Esturgeon de Sibérie

*Acipenser ruthenus* Linnaeus, 1758 - Sterlet sturgeon - Sterlet

### Order SALMONIFORMES
#### Family Salmonidae

*Oncorhynchus kisutch* (Walbaum, 1792)<fnref n="8" /> - Coho salmon - Saumon argenté

*Oncorhynchus mykiss* (Walbaum, 1792) - Rainbow trout - Truite arc-en-ciel

## Order CYPRINODONTIFORMES
### Family Cyprinodontidae

*Aphanius fasciatus* (Valenciennes, 1821) - Mediterranean banded pupfish - Aphanius de Corse

*Aphanius iberus* (Valenciennes, 1846)<fnref n="9" /> - Spanish pupfish - Aphanius d'Espagne

#### Family Poeciliidae

*Gambusia holbrooki* Girard, 1859 - Eastern mosquitofish - Gambusie de Holbrook

## Order GASTEROSTEIFORMES
### Family Gasterosteidae

*Pungitius laevis* (Cuvier, 1829) - Smoothtail ninespine stickleback - Épinochette à peau douce

Some primarily freshwater species are sometimes cited geographically from "marine" environment. However, it is often by accident (fishes washed out by a flood, and found at sea in the plume of freshwater at the surface), or in very restricted areas under the strong influence of a freshwater source. We did not include those in this list.

<footnote n="8">⁸ Was introduced for aquaculture, but the recorded escapees did not establish a population.</footnote>

<footnote n="9">⁹ Most probably extinct in France.</footnote>
<figure><img src="image_5.png" /></figure>