# An annotated list of the Blattodea (Insecta) described by Henri de Saussure. Part 4: the Blaberidae subfamilies Blaberinae, Geoscapheinae and Panesthiinae

John Hollier, Judith Marshall, Frédéric Legendre

► To cite this version:

John Hollier, Judith Marshall, Frédéric Legendre. An annotated list of the Blattodea (Insecta) described by Henri de Saussure. Part 4: the Blaberidae subfamilies Blaberinae, Geoscapheinae and Panesthiinae. Revue Suisse de Zoologie, 2025, 132 (1), pp.65-77. 10.35929/rsz.0140. mnhn-05156152

HAL Id: mnhn-05156152

https://mnhn.hal.science/mnhn-05156152v1

Submitted on 10 Jul 2025

HAL is a multi-disciplinary open access
archive for the deposit and dissemination of sci-
entific research documents, whether they are pub-
lished or not. The documents may come from
teaching and research institutions in France or
abroad, or from public or private research centers.

L'archive ouverte pluridisciplinaire HAL, est
destinée au dépôt et à la diffusion de documents
scientifiques de niveau recherche, publiés ou non,
émanant des établissements d'enseignement et de
recherche français ou étrangers, des laboratoires
publics ou privés.

<figure><img src="image_1.png" /></figure>

Distributed under a Creative Commons Attribution 4.0 International License
<figure><img src="image_2.png" /></figure>

# An annotated list of the Blattodea (Insecta) described by Henri de Saussure. Part 4: the Blaberidae subfamilies Blaberinae, Geoscapheinae and Panesthiinae

Authors: Hollier, John, Marshall, Judith, and Legendre, Frédéric

Source: Revue suisse de Zoologie, 132(1): 65-77

Published By: Muséum d'histoire naturelle, Genève

URL: https://doi.org/10.35929/RSZ.0140

The BioOne Digital Library (https://bioone.org/) provides worldwide distribution for more than 580 journals and eBooks from BioOne's community of over 150 nonprofit societies, research institutions, and university presses in the biological, ecological, and environmental sciences. The BioOne Digital Library encompasses the flagship aggregation BioOne Complete (https://bioone.org/subscribe), the BioOne Complete Archive (https://bioone.org/archive), and the BioOne eBooks program offerings ESA eBook Collection (https://bioone.org/esa-ebooks) and CSIRO Publishing BioSelect Collection (https://bioone.org/csiro-ebooks).

Your use of this PDF, the BioOne Digital Library, and all posted and associated content indicates your acceptance of BioOne's Terms of Use, available at www.bioone.org/terms-of-use.

Usage of BioOne Digital Library content is strictly limited to personal, educational, and non-commercial use. Commercial inquiries or rights and permissions requests should be directed to the individual publisher as copyright holder.

BioOne is an innovative nonprofit that sees sustainable scholarly publishing as an inherently collaborative enterprise connecting authors, nonprofit publishers, academic institutions, research libraries, and research funders in the common goal of maximizing access to critical research.
# An annotated list of the Blattodea (Insecta) described by Henri de Saussure.
## Part 4: the Blaberidae subfamilies Blaberinae, Geoscapheinae and Panesthiinae

John Hollier<fnref n="1" />*, Judith Marshall<fnref n="2" /> & Frédéric Legendre<fnref n="3" />

**Abstract:** Henri de Saussure described 57 species and one subspecies in the subfamilies Blaberinae, Geoscapheinae and Panesthiinae of the family Blaberidae, nine of them in collaboration with Leo Zehntner. These species are listed alphabetically with an account of the type specimens found in the Muséum d'histoire naturelle de Genève. The location of type specimens in other museums is given where known. The current binominal combination for each species is provided.

**Keywords:** Cockroaches, Geneva, London, Paris, type specimens.

## INTRODUCTION

Henri de Saussure (1829-1905) was a prolific Genevan naturalist who described over three thousand species and many higher taxa (Hollier & Hollier, 2013). His interest in cockroaches began with his expedition to the Antilles and Mexico in 1854-1856 (Hollier & Hollier, 2012), and his first publications on the group were descriptions of species he had captured during his trip. He returned to this insect group in the 1890s, partly working with the young Swiss entomologist Leo Zehntner (1864-1961) who acted as Saussure's assistant before going on to a distinguished career as a tropical applied entomologist (Hollier & Hollier, 2018). Saussure (alone and with Zehntner) is the author of 578 species level names in the Blattodea. The early descriptions are generally rudimentary and in most of his descriptions Saussure did not designate holotypes or even enumerate the number of specimens studied. He did not always state where the specimens were deposited.

Saussure described 57 species and one subspecies of cockroach currently placed in the Blaberidae subfamilies Blaberinae, Geoscapheinae and Panesthiinae. These are listed herein.

## ARRANGEMENT AND FORMAT

The subfamilies and species are listed alphabetically following the format used in Hollier *et al.* (2020).

The following abbreviations are used:

CSF Cockroach Species Files Online (Beccaloni, 2014)

MHNG Muséum d'histoire naturelle de Genève

MHNN Muséum d'histoire naturelle de Neuchâtel

MLUH Martin-Luther-Universität Halle

MNHN Muséum national d'Histoire naturelle, Paris

NHMUK Natural History Museum, London

## CATALOGUE

### BLABERINAE

**angulata** Saussure, 1864b: 257 [*Monastria*]

Le Brésil; Bahia. Unspecified number of ♀.

The MHNG collection contains one ♀ specimen (MHNG-ARTO-29132) with labels: “♀ Bahia, Brésil, Mr H de Saussure” [handwritten on ruled white card]; “Monastria, angulata Sauss, Brésil” [handwritten on green card]; “Monastria, angulata, Sss.” [handwritten on green paper];

<footnote n="1">¹ Muséum d'histoire naturelle de Genève, C.P. 6434, CH-1211 Geneva 6, Switzerland</footnote>

<footnote n="2">² Natural History Museum, Cromwell Road, London SW7 5BD, UK</footnote>

<footnote n="3">³ Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national d'Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, C.P. 50, 57 rue Cuvier, F-75005 Paris, France</footnote>

<footnote>* Corresponding author: john.hollier@geneve.ch</footnote>
"MHNG" [printed on white paper]; "Syntypus" [printed on red paper]. Specimen directly pinned. The ends of the antennae and the tarsi of the right hind leg are lost. The left front wing is angled forward. The type series is undefined, and the specimen must be regarded as a syntype. Box Blattodea 50.

*Monastria angulata* Saussure, 1864

**bilobata** Saussure, 1864a: 345 [Brachycolla]

Brasilia. Unspecified number of ♀.

Saussure (1870) stated that the type series was in the MNHN and gave the locality as "Province des Mines". The MHNG collection contains two specimens. One (MHNG-ARTO-22607) was collected by Fruhstorfer (1866-1922) and cannot be a syntype. The other, an immature specimen (MHNG-ARTO-22606) has a label indicating that it came from the collection of Brunner von Wattenwyl and has the same data label at the specimens of *D. brunneri* Saussure & Zehntner, 1895, and so it is probably not a syntype. Box Blattodea 49. The MNHN collection contains three ♀ specimens (MNHN-EP-EP1202, MNHN-EP-EP1203 and MNHN-EP-EP1204) referred to as syntypes in their online database.

*Parahormetica bilobata* (Saussure, 1864)

**brasiliana** Saussure, 1864a: 347 [Blabera]

Brasilia. Unspecified series.

Saussure (1864b) mentioned only the ♂. The MHNG collection contains one ♂ specimen (MHNG-ARTO-22689) with labels: "♂ Brésil, coll. Guérin" [handwritten on ruled white card]; "Blabera, Brasiliana, Brésil ♂ Sauss." [handwritten on green card]; "Blabera, brasiliana ♂ Sauss." [handwritten on green paper]; "Syntypus" [printed on red paper]. Specimen set with wings spread. The ends of the antennae, the tibia and tarsi of the left front leg, the tarsi of the left middle leg and the tarsi of the left hind leg are missing. Box Blattodea 52.

*Blaberus brasilianus* Saussure, 1864

**brunneri** Saussure, 1869a: 113 [Blabera]

Brasilia. Unspecified number of ♀.

Saussure (1870) stated that he had seen one ♀, without a head, in the MNHN.

No specimens found in the MHNG collection. The holotype (MNHN-EP-EP8454) is in the MNHN.

*Hemiblabera brunneri* (Saussure, 1869)

**brunneri** Saussure & Zehntner, 1895a: 57 [Dasyposoma]

Brasilia; Lagos [sic]. Unspecified number of ♀.

The MHNG collection contains two specimens. A ♀ specimen (MHNG-ARTO-22625) with labels: "609 56 Lages, Brésil, Mr Brunner d. W." [handwritten on ruled white card]; "Lages, Brésil" [handwritten on green paper]; "Dasyposoma Brunneri ♀ S. et Z." [handwritten on green paper]; "Syntypus" [printed on red paper].

Specimen directly pinned. The end of the left antenna and the complete right antenna are lost.

A ♀ specimen (MHNG-ARTO-22626) with labels: "609 56 Lages, Brésil, Mr Brunner d. W." [handwritten on ruled white card]; "Dasyposoma Brunneri ♂ [sic] Sauss et Z." [handwritten on green paper]; "Syntypus" [printed on red paper]. Specimen directly pinned. The ends of both antennae and the entire right hind leg are missing. Box Blattodea 49.

*Hormetica brunneri* (Saussure & Zehntner, 1895)

**burmeisteri** Saussure, 1864b: 254 [Monachoda]

Replacement name for *M. grossa* Burmeister, 1838 (nec Thunberg). The type series is presumably with Burmeister's collection in the MLUH.

*Monachoda burmeisteri* Saussure, 1864

**capucina** Saussure, 1862: 234 [Blabera]

Brasilia. Unspecified series.

Saussure (1864b) only mentioned the ♂ and gave the locality as Bahia.

The MHNG collection contains one ♂ specimen (MHNG-ARTO-22645) with labels: "Bahia" [printed on white card]; "♂ Brésil M H de Saussure" [handwritten on ruled white card]; "Hemiblabera capucina Sauss. ♂" [handwritten on green paper]; "Syntypus" [printed on red paper]. Specimen directly pinned, with left front wing spread. The head and underside of the prothorax are lost, and the body has been repaired with glue. The type series is unspecified and so the specimen is considered a syntype. Box Blattodea 49.

*Hemiblabera capucina* (Saussure, 1862)

**cicatricosa** Saussure, 1869a: 113 [Parahormetica]

Ager Argentinus. Unspecified number of ♂.

The MHNG collection contains one ♂ specimen (MHNG-ARTO-22608) with labels: "600 84 Rep. Argent., Mr Claraz" [handwritten on ruled white card]; "Hormetica cicatricosa, République Argentine" [handwritten on green paper"; "Parahormetica cicatricosa ♂ Sauss." [handwritten on green paper]; "Syntypus" [printed on red paper]. Specimen directly pinned. The tips of the antennae are missing. The type series is unspecified and so the specimen is regarded as a syntype.

Box Blattodea 49.

*Parahormetica cicatricosa* Saussure, 1869

**claraziana** Saussure, 1864a: 348 [Blabera]

Uruguay. Unspecified number of ♂ and ♀.

Saussure (1864b) gave measurements of both sexes and stated that he received three ♂ and four ♀ collected at "La Plata, province d'Entrerios" by Claraz.

Among the specimens placed under this name in the MHNG collection there are three ♂, three ♀ and two juvenile specimens with labels indicating that they had been sent by Claraz and are likely to represent the type
series. A ♂ specimen (MHNG-ARTO-23764) with labels: “♂ Entrerios, Uruguay, Claraz & Heusser” [handwritten on ruled white card]; “Blaptica, claraziana, ♂ Sauss.” [handwritten on green paper]; “Syntypus” [printed on red paper]. Specimen directly pinned with wings roughly spread. Most of both antennae are missing.

A ♂ specimen (MHNG-ARTO-22780) with the same label data. Specimen directly pinned with wings roughly folded. Part of the right antenna and the entire right front leg are lost.

A ♂ specimen (MHNG-ARTO-22779) with labels: “♂ San José Entrerios, R. Argent., envoi Claraz” [handwritten on ruled white card], “Blaptica, claraziana, ♂ Sauss.” [handwritten on green paper]; “Syntypus” [printed on red paper]. Specimen directly pinned with left wings spread and right wings folded. The ends of the antennae, the tibia and tarsi of the right hind leg and the tarsi of the left hind leg are missing.

A ♀ specimen (MHNG-ARTO-22781) with labels: “♀ San José Entrerios, R. Argent., envoi Claraz” [handwritten on ruled white card]; “Blaptica, claraziana, ♀ Sauss.” [handwritten on green paper]; “Syntypus” [printed on red paper]. Specimen directly pinned. The ends of the antennae and three tarsal segments of the left hind leg are lost.

A ♀ specimen (MHNG-ARTO-22782) with labels: “♀ San José Entrerios, Uruguay, R. Argent., envoi Claraz” [handwritten on ruled white card]; “Blaptica, claraziana, ♀ Sauss.” [handwritten on green paper]; “Syntypus” [printed on red paper]. Specimen directly pinned. The ends of the antennae, two tarsal segments of the right front leg, the claw of the left hind leg and the tarsi of the right hind leg are lost. There is insect feeding damage to the underside of the specimen.

A ♀ specimen (MHNG-ARTO-22784) with labels: “♀ Buenos Ayres, Rep. Argent., envoi Claraz” [handwritten on ruled white card]; “Blaptica, claraziana, ♀ Sauss.” [handwritten on green paper]. Specimen directly pinned. The ends of the antennae, the tarsi of the left front leg and the entire right middle leg are missing. This specimen is not a syntype because it is from a different locality, as are two immature specimens (MHNG-ARTO-22789 and 22794) sent by Claraz. Box Blattodea 55.

A junior synonym of *Blaptica dubia* (Serville, 1838)

*couloniana* Saussure, 1873: 124, fig. 42 [*Glyptopeltis*]
L'île de Java? Unspecified number of ♀.

The MHNG collection contains one specimen (MHNG-ARTO-22628) with labels: “♀ Java M H de Saussure” [handwritten on white card]; “Couloniana Sss., (Glyptopeltis), Java Type” [handwritten on white card]; “Syntypus” [printed on red paper]. Specimen directly pinned. Both antennae, the claw of the left front leg, the tarsi of the left middle leg, the entire right middle leg and right hind leg are lost. The type series is unspecified

and so the specimen is regarded as a syntype. Box
Blattodea 49.

*Glyptopeltis couloniana* Saussure, 1873

*cubensis* Saussure, 1864a: 347 [*Blabera*]

Cuba. Unspecified series.

Saussure (1864b) gave measurements of both sexes. The MHNG collection contains four specimens. A ♂ specimen (MHNG-ARTO-22736) with labels: “♂ Cuba, Mr H de Saussure” [handwritten on ruled white card]; “Blabera, cubensis, ♂ Sauss.” [handwritten on green paper]; “Syntypus” [printed on red paper]. Specimen directly pinned with wings folded. The end of the left antenna, the tarsi of the left middle leg and the claw of the left hind leg are missing.

A ♂ specimen (MHNG-ARTO-22738) with labels: “♂ Cuba, Mr H de Saussure” [handwritten on ruled white card]; “Blabera, cubensis, ♂ Sauss.” [handwritten on green paper]; “Syntypus” [printed on red paper]. Specimen directly pinned with wings folded. Both antennae, the tarsi of both front legs and the tarsi of the right hind leg are lost. There is insect feeding damage to the underside of the abdomen, thorax, and right hind leg.

A ♀ specimen (MHNG-ARTO-22739) with labels: “♀ Cuba, Mr H de Saussure” [handwritten on ruled white card]; “Blabera, cubensis, ♀ Sauss.” [handwritten on green paper]; “La Bl. aequatoriana de Bolivar [illegible] paraît être une variété de Cubensis, à élytres et ailes plus transparents” [handwritten on green paper], “Syntypus” [printed on red paper]. Specimen directly pinned with right wings spread and left wings folded. Both antennae, the tarsi of the right front leg, the tarsi of the left hind leg and the claw of the right hind leg are lost.

A ♂ specimen (MHNG-ARTO-22737) with labels: “♂ Puerto Rico, Antilles, coll. Guérin” [handwritten on ruled white card]; “Blabera, Cubensis Sauss, Antilles ♂” [handwritten on green card]; “Blabera, cubensis, ♂ Sauss.” [handwritten on green paper]. Specimen directly pinned with wings spread. The ends of both antennae, the right middle leg and the last tarsal segment of both hind legs are lost. There is insect feeding damage to the thorax and the specimen has been repaired with glue. Although the locality does not match that given in the original description, Saussure described species using specimens from Guérin-Méneville’s collection early in his career and so would have had this individual available for this description. Box Blattodea 54.

A junior synonym of *Blaberus discoidalis* Serville, 1838

*deplanata* Saussure, 1864a: 348 [*Blabera*]

Antillae [sic]. Unspecified series.

Saussure (1864b) mentioned only the ♀ and gave the locality as Cuba. Saussure (1870) stated that the type series was in the MNHN. The MHNG collection contains one ♂ specimen (MHNG-ARTO-22680) with labels “♂ Cuba, Dr Krug” [handwritten on ruled white
card]; “Archimandrita, deplanata Sauss. ♂” [handwritten on green paper]; “Aspiduchus ♂, deplanatus (Sauss.) 1864, Det. E. Gutiérrez IX.1999” [handwritten on white card]. Specimen directly pinned. Parts of the antennae and legs are glued to card mounts on the original pin. The dissected genitalia are in a small tube secured through the stopper on the original pin. Given the sex of the specimen, and the provenance, it seems unlikely that it can be a syntype. Carl Wilhelm Leopold Krug (1833-1898) was a German businessman and naturalist based in Puerto Rico who funded collecting trips by Juan Gundlach in Cuba and Paul Sintenis in St Thomas, some of them after his return to Berlin in 1876. Box Blattodea 52. There is a ♀ specimen in the MNHN (MNHN-EP-EP1267) referred to as the holotype in their online database.

*Aspiduchus deplanatus* (Saussure, 1864)

**diabolus** Saussure, 1864a: 345 [*Brachycolla*]

Brasilia. Unspecified series.

Saussure (1864b) mentions the ♂ and states that the type series was collected by Saint-Hilaire and was in the MNHN. No specimens found in the MHNG collection. There is a specimen in the MNHN (MNHN-EP-EP1248) referred to as the holotype in their online database.

*Bionoblatta diabolus* (Saussure, 1864)

**ferruginea** Saussure, 1864b: 241 [*Blabera*]

Le Mexique.

Saussure attributed this name to Stoll and said he did not know this species.

A junior synonym of *Blaberus craniifer* Burmeister, 1838

**fraterna** Saussure, 1864b: 241-242 [*Blabera*]

L'Amérique du Sud (les Antilles; Cuba?) (Musée de Paris). Unspecified number of ♂.

No specimens found under this name in the MHNG collection. There is a series placed under this name in the MNHN, but these specimens are from Argentina and therefore unlikely to be the type series.

A junior synonym of *Blaberus minor* Saussure, 1864

**granulata** Saussure, 1893: 68 [*Hemiblabera*]

Mexico. Unspecified number of ♀.

The MHNG collection contains one ♀ specimen (MHNG-ARTO-22646) with labels: “♀ Antigua M H de Saussure” [handwritten on ruled white card]; “Hemiblabera granulata Sauss. ♀” [handwritten on green paper]; “Possible syntype of H. granulata Saussure, J. Hollier 2022” [handwritten on red paper]. Specimen directly pinned. The right front wing is detached and glued to a card mount secured on the original pin. The locality on the label does not match that of the description, but it does match the distribution attributed to this species by CSF. Box Blattodea 47.

*Hemiblabera granulata* Saussure, 1893

**immacula** Saussure & Zehntner, 1894: 120, pl. 5 fig. 23 [*Blabera*]

Brazil; Pernambuco (Mus. Genavense). Unspecified number of ♂ and ♀.

The MHNG collection contains three specimens. A ♂ specimen (MHNG-ARTO-22688) with labels; “2 25 Pernamb., Brésil, M De Lessert” [handwritten on ruled white card]; “Blabera, imacula [sic], ♂ Sss. et Z.” [handwritten on green paper]; “Syntypus” [printed on red paper]. Specimen directly pinned with wings roughly folded. Most of the left antenna is lost.

A ♀ specimen (MHNG-ARTO-22687) with labels: “2 25 Pernamb., Brésil, M De Lessert” [handwritten on ruled white card]; “Blabera, immacula, ♀ Sss. et Z.” [handwritten on green paper]; “Syntypus” [printed on red paper]. Specimen directly pinned with wings spread. The last tarsal segment of the right middle leg is missing. An immature specimen (MHNG-ARTO-22686) with the same label data. Specimen directly pinned. The right front leg is missing, and there is a build-up of verdigris on the pin which is starting to split the specimen where the pin is inserted. Box Blattodea 52.

*Eublaberus immaculus* (Saussure & Zehntner, 1894)

**manca** Saussure, 1893: 68 [*Hemiblabera*]

Portorico [sic]. Unspecified number of ♂.

The MHNG collection contains one ♂ specimen (MHNG-ARTO-22644) with labels: “Porto Rico ♂ M H de Saussure” [handwritten on white card]; “Hemiblabera manca Sss. ♂” [handwritten on green paper]; “Syntypus” [printed on red paper]. Specimen directly pinned, with the left front wing spread. The ends of both antennae and the claws of the left hind leg are missing. The type series was unspecified and so the specimen is considered a syntype. Blattodea 49.

A junior synonym of *Hemiblabera brunneri* (Saussure, 1869)

**marmorata** Saussure, 1869a: 113 [*Dasyposoma*]

Venezuela. Unspecified number of ♀.

Saussure (1870) stated that the type series was in the MNHN and gave the locality as “Colombie”.

No specimens found in the MHNG collection. There is a specimen in the MNHN (MNHN-EP-EP8455) under the name *Dasyposoma marmorata* and with the locality label “Colombie” which is certainly a syntype.

*Hormetica marmorata* (Saussure, 1869)

**mexicana** Saussure, 1862: 233-234 [*Blabera*]

Mexico. Unspecified series.

Saussure (1864b) gave measurements of both sexes and listed the localities of Tampico, Tuxpan and Cordoba.

The MHNG collection contains five specimens that were collected by Saussure and are therefore probably syntypes. These have been placed under the name *Blabera trapezoideus* but the species name labels in the
insect box have “mexicana Sss.” handwritten on them. A ♂ specimen (MHNG-ARTO-22712) with labels: “♂ Cordoba t.c., Méxique, Mr H de Saussure” [handwritten on ruled white card]; “Blabera, trapezoideus, ♂ Burm.” [handwritten on green paper]; “Probable syntype, B. mexicana Sss., Hollier 2024” [handwritten on red paper]. Specimen directly pinned with wings folded. The ends of the antennae, the tarsi of the left middle leg and the last tarsal segment of the right hind leg are missing.

A ♂ specimen (MHNG-ARTO-22714) with the same label data. Specimen directly pinned with wings folded. The right antenna, the tibia and tarsi of the left middle leg and the entire right middle and hind legs are lost.

A ♀ specimen (MHNG-ARTO-22715) with labels: “♀ Tampico t.c., Méxique, Mr H de Saussure” [handwritten on ruled white card]; “Blabera, trapezoideus, ♀ Burm.” [handwritten on green paper]; “Probable syntype, B. mexicana Sss., Hollier 2024” [handwritten on red paper]. Specimen directly pinned with wings folded. The right antenna is missing and there is mould damage to the underside.

A ♀ specimen (MHNG-ARTO-22717) with the same label data. Specimen directly pinned with wings folded. The head, the left front leg, the last tarsal segment of the right front leg and the entire left hind leg are missing. There is insect feeding damage to the underside of the thorax.

A juvenile specimen (MHNG-ARTO-22710) with labels: “♀ Cordoba, Méxique, Mr H de Saussure” [handwritten on ruled white card]; “Blabera, trapezoideus” [handwritten on green paper]; “Probable syntype, B. mexicana Sss., Hollier 2024” [handwritten on red paper]. The specimen was directly pinned, but a build-up of verdigris has broken it in half where the pin was inserted. The two pieces have been pinned into place in the insect box. The tips of the antennae are lost and the tarsi of the right hind leg are detached and glued to a card mound on the original pin. Box Blattodea 53. A junior synonym of *Blaberus giganteus* (Linnaeus, 1758)

### minor Saussure, 1864a: 347-348 [Blabera]

Brasilia. Unspecified number of ♂ and ♀.

The MHNG collection contains one ♂ specimen (MHNG-ARTO-22690) with labels: “♂ Paraguay, M H de Saussure” [handwritten on ruled white card]; “Blabera, minor Sauss, Paraguay” [handwritten on green card]; “Blabera, minor, ♂ Sauss.” [handwritten on green paper]. Specimen directly pinned with wings spread. Both antennae, both front legs, the tibia and tarsi of the left middle leg, the tarsi of the right middle leg, the tibia and tarsi of the right hind leg and the tarsi of the left hind leg are lost. Although the locality on the labels does not match that of the original description, the green card label shows that this specimen entered Saussure's collection fairly early, and it is possible this specimen

is a syntype. The whereabouts of the other syntypes is unknown. Box Blattodea 52.

*Blaberus minor* Saussure, 1864

### obscura Saussure & Zehntner, 1894: 121 [Blaptica]

Brazil (Mus. Genavense). Unspecified number of ♂. The MHNG collection contains one specimen (MHNG-ARTO-22776) with labels: “♂ Brésil” [handwritten on ruled white card]; “obscura Sss & Z” [handwritten on green card]; “Blaptica, obscura, ♂ Sauss. & Z.” [handwritten on green paper]; “Syntypus” [printed on red paper]. Specimen directly pinned with wings roughly folded. The ends of the antennae are missing, the abdomen is detached, and the terminal part is glued to a card mount secured on the original pin. The type series is unspecified and so the specimen must be regarded as a syntype. Box Blattodea 55.

*Blaptica obscura* Saussure & Zehntner, 1894

### punctata Saussure, 1873: 145-146 [Parahormetica]

Le Brésil. More than one ♂.

No specimens found in the MHNG collection. The whereabouts of the type series is unknown.

*Parahormetica punctata* Saussure, 1873

### reflexa Saussure & Zehntner, 1893: 68-69, pl. 4 fig. 35 [Rhicnoda]

Nicaragua; Chontales (Janson). Unspecified number of ♂.

No specimens found in the MHNG collection. There is a specimen in the NHMUK (BMNH(E)878054) referred to as the holotype in their online database and in CSF, with labels: “Chontales, Nicaragua. Janson” [printed on white card]; “Brit. Mus. 1894-125.” [printed on white card]; “Rhicnoda reflexa, ♂ Sauss.” [handwritten on green paper]; “Holotype” [printed on red-bordered paper]. Specimen directly pinned with some verdigris ventrally. The right hind tibia and tarsus and left hind tarsus are lost. The type series is undefined, and notwithstanding the holotype label, this specimen should be regarded as a syntype.

*Hyporhicnoda reflexa* (Saussure & Zehntner, 1893)

### rufescens Saussure & Zehntner, 1894: 119, pl. 5 fig. 22 [Blabera]

Antilles; Cuba (Dr. Krug). Unspecified number of ♀.

The MHNG collection contains one ♀ specimen (MHNG-ARTO-22740) with labels: “♀ Cuba, Dr. Krug” [handwritten on ruled white card]; “Blabera, rufescens, ♀ Sss.” [handwritten on green paper]; “Syntypus” [printed on red paper]. Specimen directly pinned with wings spread. The end of the left antenna, the entire right antenna, and both front legs are lost. The type series is undefined and so the specimen must be regarded as a syntype. Box Blattodea 54.

A junior synonym of *Blaberus discoidalis* Serville, 1838
**scutata** Saussure & Zehntner, 1894: 119, pl. 5 fig. 24
[Blabera]

Brazil; Pernambuco (Mus. Genavense). Unspecified
number of ♂ and ♀.

The MHNG collection contains eight specimens and an
ootheca. A ♂ specimen (MHNG-ARTO-22741) with
labels: “♂ Pernamb., Brésil, Mr De Lessert” [handwritten
on ruled white card]; “Blabera, scutata, ♂ Sauss.”
[handwritten on green paper]; “Syntypus” [printed on
red paper]. Specimen directly pinned with wings spread.
The tips of the antennae and three tarsal segments of the
left hind leg are lost. The left antenna and left forewing
are detached and glued to a card mount on a separate pin.

A ♂ specimen (MHNG-ARTO-22742) with labels: “♂
Pernam, buco, Mr De Lessert” [handwritten on ruled
white card]; “Blabera, scutata, ♂ Sauss.” [handwritten
on green paper]; “Syntypus” [printed on red paper].
Specimen directly pinned with wings roughly folded. The
end of the right antenna is missing. The tarsi of the right
hind leg are detached and glued to a card mount secured
on the original pin.

A ♂ specimen (MHNG-ARTO-22744) with labels: “♂
Pernamb., Brésil, Mr De Lessert” [handwritten on ruled
white card]; “Blabera, scutata, var. ♂ Sss.” [handwritten
on green paper]; “Syntypus” [printed on red paper].
Specimen directly pinned with wings spread.

A ♀ specimen (MHNG-ARTO-22743) with labels: “♀
Pernam, buco, Mr De Lessert” [handwritten on ruled
white card]; “Blabera, scutata, ♀ Sauss.” [handwritten
on green paper]; “Syntypus” [printed on red paper].
Specimen directly pinned with wings folded. The tarsi
of the right middle leg and the claw of the right hind leg
are missing.

A ♀ specimen (MHNG-ARTO-22745) with labels: “♀
Pernamb., Brésil, Mr De Lessert” [handwritten on ruled
white card]; “Blabera, scutata, var. ♀ Sss.” [handwritten
on green paper]; “Syntypus” [printed on red paper].
Specimen directly pinned with wings spread. The ends of
both antennae are lost.

A ♂ specimen (MHNG-ARTO-22746) with the same
label data. Specimen directly pinned with wings folded.

A ♀ specimen (MHNG-ARTO-22748) with the same
label data. Specimen directly pinned with wings folded.
The ends of both antennae are lost.

A juvenile specimen (MHNG-ARTO-22747) with labels: "2 Pernamb., 25 Brésil, Mr De Lessert" [handwritten on ruled white card]; "Blabera, scutata, Sss. & Z., larve" [handwritten on green paper]; "Syntypus" [printed on red paper]. Specimen directly pinned. The claw of the right hind leg is lost.

The ootheca (MHNG-ARTO-22749) is glued to a card
mount and has the label "2 Pernamb., 25 Brésil, Mr De
Lessert" [handwritten on ruled card].

The “varietas obscura” is mentioned in the original
description, and the remark later in the description
that “when the black macula of the pronotum is of the

normal form...” indicated that this refers to intra-specific
variation, and that all of these specimens are syntypes.
Box Blattodea 54.

*Blaberus scutatus* Saussure & Zehntner, 1894

**semialata** Saussure, 1864b: 258 [*Monastria*]

L'Amérique méridionale. Unspecified number of ♀.

The MHNG collection contains one possible syntype.
A ♀ specimen (MHNG-ARTO-29133) with labels: “♀
Brésil” [handwritten on ruled white card]; “Monastria,
semialata Sauss., Brésil” [handwritten on green card];
“Monastria semialata, ♀ Sauss.” [handwritten on green
paper]; “MHNG” [printed on white paper]; “Possible
syntype, of M. semialata, Hollier 2024” [handwritten
on red paper]. Specimen directly pinned. Both antennae
and two tarsal segments of the left front leg are missing.
There is mould damage to the underside of the body. The
other specimens placed under this name were collected
by Hans Fruhstorfer (1866-1922) and must have been
collected after the original publication and so cannot be
syntypes. Box Blattodea 50.

A junior synonym of *Hiereoblatta cassidea* (Eschscholtz, 1822)

**strumosa** Saussure & Zehntner, 1895a: 56 [*Hormetica*]

Equator. Unspecified number of ♂.

The MHNG collection contains one ♂ specimen (MHNG-ARTO-22622) with labels: “♂ Ecuador” [handwritten on ruled white card]; “Hormetica strumosa ♂ S. et Z.” [handwritten on green paper]; “Syntypus” [printed on red paper]. Specimen set with left wings spread and right wings folded. The type series is undefined and so the specimen is regarded as a syntype. Box Blattodea 49.

*Hormetica strumosa* Saussure & Zehntner, 1895

**thoracica** Saussure & Zehntner, 1894: 120, pl. 5 fig. 25
[Blabera]

Panama. Colombia; Bogota (Mus. Genavense). More
than one ♀.

The MHNG collection contains three specimens. A ♀ specimen (MHNG-ARTO-22684) with labels: “♀ Bogota” [handwritten on ruled white card]; “Blabera, thoracica, ♀ Sauss.” [handwritten on green paper]; “Syntypus” [printed on red paper]. Specimen directly pinned with wings folded. The claw of the right middle leg is missing, and there is insect feeding damage to the underside of the abdomen.

A ♀ specimen (MHNG-ARTO-22685) with labels: “♀
Bogota” [handwritten on ruled white card]; “Blabera,
thoracica, ♀ Stoll [sic]” [handwritten on green paper];
“Syntypus” [printed on red paper]. Specimen directly
pinned with wings spread. The right antenna, the right
front leg, the tarsi of the right middle leg and the tarsi of
the left hind leg are missing.

A ♀ specimen (MHNG-ARTO-22683) with the same
label data. Specimen directly pinned with wings folded.
The abdomen is missing, and the head and prothorax are becoming detached. Box Blattodea 52. The whereabouts of the other syntypes is unknown.

A junior synonym of *Eublaberus posticus* (Erichson, 1848)

*thunbergi minor* Saussure & Zehntner, 1894: 120
[Byrsotria]

[Cuba]. Unspecified series.

The MHNG collection contains four specimens. A ♂ (MHNG-ARTO-22767) with labels: “♂ Cuba” [handwritten on ruled white card]; “Byrsotria, Thunbergi, var. minor, ♂ Sss & Z” [handwritten on green paper], “Syntypus” [printed on red paper]. Specimen directly pinned with wings roughly spread. The tips of the antennae, the tarsi of the right front leg, the tarsi of the right middle leg and the tibia and tarsi of the right hind leg are lost.

A ♂ (MHNG-ARTO-22770) with the same label data. Specimen directly pinned with wings folded. The right antenna and the tibia and tarsi of the right hind leg are missing.

A ♀ (MHNG-ARTO-22769) with labels: “Thunbergi, Guér. ♀, Cub.” [handwritten on white card]; “Byrsotria, Thunbergi, var. minor, ♂ Sss & Z” [handwritten on green paper], “Syntypus” [printed on red paper]. Specimen directly pinned. The right antenna and the tarsi of the left front leg are missing.

A ♀ (MHNG-ARTO-22768) with labels: “♀ Cuba” [handwritten on ruled white card]; “Byrsotria, Thunbergi, var. minor, ♂ Sss & Z” [handwritten on green paper], “Syntypus” [printed on red paper]. Specimen directly pinned. Most of both antennae, the tarsi of the right front leg and the tibia and tarsi of the right hind leg are lost. The left front wing is displaced forward, and the specimen is starting to break apart where the pin is inserted. Box Blattodea 55.

A junior synonym of *Byrsotria fumigata* (Guérin-Méneville, 1857)

*trilobita* Saussure, 1862: 233 [*Hormetica*]

Brasilia. Unspecified series.

Saussure (1864b) only mentions the ♀. The MHNG collection contains two specimens. A ♀ specimen (MHNG-ARTO-22605) with labels: “Brésil, coll. Guérin” [handwritten on ruled white card]; “monticollis Burm., (trilobata Sauss.), Brésil Parahormetica” [handwritten on green card]; “Parahormetica monticollis, ♀ Burm.” [handwritten on green paper]; “Syntypus” [printed on red paper]. Specimen directly pinned. The left antenna, the tibia and tarsus of the right middle leg and the tarsi of both hind legs are missing. The abdomen is split ventrally. The type series is unspecified and so the specimen is regarded as a syntype.

A ♀ specimen (MHNG-ARTO-22604) with labels: “Rio Janeiro, Mr H de Saussure” [handwritten on

green paper]; “Parahormetica monticollis ♀ Burm.” [handwritten on green paper]. Specimen directly pinned. The ends of both antennae are missing. This specimen is probably not a syntype. Box Blattodea 49.

A junior synonym of *Parahormetica monticollis* (Burmeister, 1838)

The collection also contains a paratype of *Hyporhicnoda metae* Hebard, 1921 (MHNG-ARTO-22632) in box Blattodea 49, 7 syntypes of *Richnoda carinata* Biolley, 1900 (a junior synonym of *Hyporhicnoda reflexa* Saussure & Zehntner, 1893) (MHNG-ARTO-22633-22639) in box Blattodea 49, and a specimen of *Eublaberus sulzeri* (Guérin-Méneville, 1857) (MHNG-ARTO-22681) labelled as a type in box Blattodea 52.

## GEOSCAPHEINAE

*castanea* Saussure, 1869b: 285 [*Dasyposoma*]

La nouvelle Hollande. Unspecified number of ♂.

No specimens found in the MHNG collection. There is a ♂ specimen in the MNHN (MNHN-EP-EP1205) referred to as the holotype in their online database.

*Geoscapheus castaneus* (Saussure, 1869)

*dilatata* Saussure, 1864a: 346-347 [*Panesthia*]

Nova Hollandia. Unspecified number of ♂ and juveniles. The MHNG collection contains two specimens. A ♂ specimen (MHNG-ARTO-22909) with labels: “♂ Nouv. Holl., ach. de Biederman” [handwritten on ruled white card]; “Plana, dilatata, ♂ Sss.” [handwritten on purple paper]; “Geoscapheus ♂, dilatatus (Sauss.), det L.M. Roth 1974 ♂ Paralectotype” [handwritten on white paper with “det L.M. Roth 19” printed]; “Syntypus” [printed on red paper]. Specimen directly pinned. The tarsi of both hind legs are lost.

A ♂ specimen (MHNG-ARTO-22910) with labels: “603 34 Queensland, Nouv. Holl., Mus. D'Aukland” [handwritten on ruled white card]; “603 34 Darling Downs, Queensland” [handwritten on white paper]; “11” [handwritten on white paper]; “86” [printed on white card]; “Plana, deplanata, ♂ Sss, dilatata” [handwritten on purple paper]; Geoscapheus, dilatatus (Saussure), det L.M. Roth 1974, ♂ Lectotype” [handwritten on white paper with “det L.M. Roth 19” printed]; “Syntypus” [printed on red paper]. Specimen directly pinned. Both antennae, the tarsi of both front legs, the tibia and tarsi of the left middle leg, the entire right middle leg, the entire left hind leg and the tibia and tarsi of the right hind leg are lost. There is insect feeding damage to the underside of the thorax.

The second, larger, specimen is almost certainly a syntype of the subspecies *G. dilatatus major* (Saussure 1895) rather than part of the original type series. Box Blattodea 57.

*Geoscapheus dilatatus* (Saussure, 1864)
**dilatata major** Saussure, 1895: 327 [*Plana*]
Queensland. Unspecified number of ♂.

See specimen MHNG-ARTO-22910 above. Box
Blattodea 57.

A junior synonym of *Geoscaphus dilatatus* (Saussure, 1864)

*kraussiana* Saussure, 1873: 150-151 [*Panesthia*]

La Nouvelle-Hollande; Melbourne. Unspecified number
of ♀ and one damaged juvenile.

The MHNG collection contains one ♀ specimen (MHNG-ARTO-22902) with labels: “♀ Melbourne, Nouv. Holl., M H de Saussure” [handwritten on ruled white card]; “85” [printed on white card]; “Hemipanesthia, Kraussiana, Sauss” [handwritten on yellow paper]; “♀ Lectotype, det L.M. Roth 1974” [handwritten on white paper with “det L.M. Roth 19” printed]; “Syntypus” [printed on red paper]. Specimen directly pinned. Both antennae and the tarsi of all legs are lost. The whereabouts of the other syntypes is unknown. Box Blattodea 57.

*Macropanesthia kraussiana* (Saussure, 1873)

*muelleri* Saussure, 1895: 329-330, fig. 5 [*Macropanesthia*]

Centrali-Australia (Mus. Stuttgardiense, Bn von Müller).
More than one ♂.

The MHNG collection contains one ♂ specimen (MHNG-ARTO-22915) with labels: “C. Austr., v. Mull. 93” [printed on white card with “v.” handwritten]; “98” [printed on white card]; “1” [handwritten on white paper]; “Macropanesthia, (Muelleri Sauss.), melolontha” [handwritten on purple paper]; “♂ Lectotype, det L.M. Roth 1974” [handwritten on white paper with “det L.M. Roth 19” printed]; “= Macropanesthia, gigantea (Tepper), det L.M. Roth 1974” [handwritten on white paper with “det L.M. Roth 19” printed]; “Syntypus” [printed on red paper]. Specimen directly pinned. Both antennae, the tarsi of the right front leg and of the other coxae and legs are lost. There is insect feeding damage to the remaining femur and the underside of the thorax, and the specimen has been strengthened with glue. The is a specimen labelled as the holotype in the SMNS. Box Blattodea 57. A junior synonym of *Parapanesthia gigantea* (Tepper, 1894)

*rhinoceros* Saussure, 1895: 329, fig. 4 [*Macropanesthia*]
Australia orientalis; Bowen (Godeffroy). Unspecified
number of ♂.

There are no specimens under this name in the MHNG
collection, but there are two specimens placed under
“*Macropanesthia Godeffredi*”, the smaller of which
(MHNG-ARTO-22914) matches the measurements
given in the original description of *M. rhinoceros*. A ♂
specimen with labels: “♂, Nouv. Holl., Mus. God. (ach)”
[handwritten on ruled white card]; “Godeffredi Sss, Nouv.
Holl. (Panesthia)” [handwritten on white card]; “88”

[printed on white card]; “Macropanesthia, Godeffredi, ♂ Sauss.” [handwritten on purple paper]; “Possible syntype of M. rhinoceros Sauss., Hollier 2024” [handwritten on red paper]. There is considerable insect feeding damage to the underside of the thorax. Both antennae, the tarsi of the left front leg, the tarsi of both middle legs and the entire hind legs are missing. The type series is undefined and so this specimen must be regarded as a syntype. The other specimen is much larger and so not a syntype. Box Blattodea 57.

*Macropanesthia rhinoceros* Saussure, 1895

The MHNG collection has three specimens of
*Geoscaphus robustus* Tepper, 1893 (MHNG-
ARTO-22911, 22912 and 22913) labelled as types in box
Blattodea 57.

PANESTHIINAE

*bigibbosa* Saussure, 1895: 324-325, fig. 1 [*Miopanesthia*]
India orientalis; Sikkim, Khasi Hills. More than one ♂.
The MHNG collection contains three specimens. A ♂
specimen (MHNG-ARTO-22905) with labels: “333,
Sikkim” [handwritten on white card]; “Sikkim, 6701,
8” [printed on white card with “Sikkim” handwritten];
“Miopanesthia, bigibbosa, Sauss.” [handwritten on
yellow paper]; “Syntypus” [printed on red paper].
Specimen directly pinned with wings folded. The right
antenna is lost.

A ♂ specimen (MHNG-ARTO-22907) with labels: “381, Khasi Hill” [handwritten on white card]; “Khasi, M Godwin-Austin” [handwritten on a white card disk]; “Miopanesthia, bigibbosa, Sauss.” [handwritten on yellow paper]; “Syntypus” [printed on red paper]. Specimen directly pinned with wings roughly folded. The right antenna, the tarsi of the right front leg, the tarsi of both middle legs, the tibia and tarsi of the left hind leg and the tarsi of the right hind leg are lost.
The third specimen (MHNG-ARTO-22906) with labels: “600 81, Indes. Or., Anc. Coll.” [handwritten on ruled white card]; “Miopanesthia, bigibbosa, ♂ Sauss.” [handwritten on yellow paper]. Specimen directly pinned with wings spread. Most of both antennae, the tarsi of both front legs, the tarsi of the left middle leg and the entire left hind leg are lost. It is not clear whether this specimen is a syntype. Box Blattodea 57.

*Miopanesthia bigibbosa* Saussure, 1895

*biglumis* Saussure, 1895: 308, 319-20 [*Panesthia*]

India; Sikkim (Mus. Calcuttae). More than one ♀.

The MHNG collection contains one ♀ specimen (MHNG-ARTO-22801) with labels: “Sikkim” [printed on white card]; “308, Sikkim” [handwritten on white paper]; “Salganea, biglumis, ♀ Sauss.” [handwritten on yellow paper]; “Syntypus” [printed on red paper]. Specimen directly pinned. The right antenna, the tarsi of both middle legs and all of both hind legs are missing.
The original description gave measurement ranges and so this specimen is a syntype. Box Blattodea 56.
*Salganea biglumis* (Saussure, 1895)

**bramina** [sic] Saussure, 1895: 310, 322 [*Panesthia*]
India; Johore. Unspecified number of ♂ and ♀.

The MHNG collection has two specimens. A ♀ specimen (MHNG-ARTO-22893) with labels: “Johore” [handwritten on white card disk]; “Panesthia, brahmina, ♀ Sauss.” [handwritten on yellow paper]; “Panesthia, brahmina Sauss., det L.M. Roth 1974, ♀ Lectotype” [handwritten on white paper with “det L.M. Roth 19” printed]; Syntypus” [printed on red paper]. Specimen directly pinned with wings roughly spread. The right antenna, the tibia and tarsi of the left front leg, the tarsi of the right hind leg and the entire left hind leg are lost.
A juvenile specimen (MHNG-ARTO-22894) with labels: “Johore” [handwritten on white card disk]; “297, Johore” [handwritten on white card]; “Panesthia, brahmina, ♀ Sauss.” [handwritten on yellow paper]; “Panesthia ♀ nymph, brahmina Sauss., det L.M. Roth 1974, Paralectotype” [handwritten on white paper with “det L.M. Roth 19” printed]; Syntypus” [printed on red paper]. Specimen directly pinned. Both antennae, the tarsi of both middle legs, the tarsi of the right hind leg and two tarsal segments of the left hind leg are missing.

The whereabouts of the ♂ syntype(s) is unknown. Box
Blattodea 57.

A junior synonym of *Panesthia transversa* Burmeister, 1838

**cribrata** Saussure, 1864a: 346 [*Panesthia*]

Nova Hollandia. Unspecified number of ♀.

The MHNG collection contains four specimens. A ♂ specimen (MHNG-ARTO-22863) with labels: “♀ [sic], Nouv. Holl., Musée de Paris” [handwritten on ruled white card]; “Panesthia, cribrata, ♂ Sss” [handwritten on purple paper]; “♂ Paralectotype, det L.M. Roth 1974” [handwritten on white paper with “det L.M. Roth 19” printed]; “Syntypus” [printed on red paper]. Specimen directly pinned. Both antennae, the right front leg and the tarsi of both hind legs are lost.

A ♂ specimen (MHNG-ARTO-22864) with labels: “N.S. Wales, Nouv. Holl., Mr H de Saussure” [handwritten on ruled white card]; “Panesthia, cribrata, ♂ Sss” [handwritten on purple paper]; “♂ Paralectotype, det L.M. Roth 1974” [handwritten on white paper with “det L.M. Roth 19” printed]. Specimen directly pinned. Both antennae, the tarsi of the right front leg, the tarsi of the left middle leg, the tarsi of the right hind leg and the entire left hind leg are missing.

A ♀ specimen (MHNG-ARTO-22865) with labels: “♀,
Nouv. Holl., Musée de Paris” [handwritten on ruled
white card]; “Panesthia, cribrata, ♀ Sss” [handwritten
on purple paper]; “Panesthia, integra, n. sp., N.S. Wales”
[handwritten in pencil on white paper]; “♀ Paralectotype,

det L.M. Roth 1974” [handwritten on white paper with
“det L.M. Roth 19” printed]; “Syntypus” [printed on
red paper]. Specimen directly pinned. The head, both
front legs, the tarsi of the left middle leg, the entire right
middle leg, the tibia and tarsi of the left hind leg and the
entire right hind leg are lost.

A juvenile specimen (MHNG-ARTO-22866) with labels: “Nouv. Holl.” [handwritten on ruled white card]; “Panesthia, cribrata Sauss.” [printed on white card]; “Panesthia ♂ nymph, cribrata Sauss., det L.M. Roth 1974” [handwritten on white paper with “det L.M. Roth 19” printed]. Box Blattodea 57.

*Panesthia cribrata* Saussure, 1864

**discoidalis** Saussure, 1895: 324, 326 [*Miopanesthia*]

Java. India; Pondichéri. More than one ♀.

The MHNG collection contains two specimens. A juvenile specimen (MHNG-ARTO-22903) with labels: “Pondichery” [handwritten on ruled white card]; “Miopanesthia, discoidalis Sss.” [handwritten on yellow paper]; “Syntypus” [printed on red paper]. Specimen directly pinned. Both antennae, the tibia and tarsi of the left front leg and the tarsi of the right hind leg are lost.

An apparently juvenile specimen (MHNG-ARTO-22904) with labels: “♀ var. minor, Java, M H de Saussure” [handwritten on ruled white card]; “Miopanesthia, discoidalis Sss., ♀ larva?” [handwritten on yellow paper]; “Syntypus” [printed on red paper]. Specimen directly pinned. Both antennae, the tibia and tarsi of the right front leg and the tarsi of the right middle leg are missing. Box Blattodea 57.

*Miopanesthia discoidalis* Saussure, 1895

**forceps** Saussure, 1869b: 287-288, fig. 24 [*Panesthia*]

Les Indes orientales; Pondichéri. Unspecified number of ♀.

No specimens found in the MHNG collection. The
whereabouts of the type series is unknown.

*Microdina forceps* (Saussure, 1869)

**foveolata** Saussure, 1895: 303-305 [*Salganea*]

Macassar (Mus. Genavense). One damaged ♀.

The MHNG collection contains one ♀ specimen (MHNG-ARTO-22824) with labels: “4” [handwritten on white card]; “Bonthein, Macassar” [handwritten on white paper]; “Musée de, Schaffhausen” [handwritten on white paper]; “Salganea, foveolata, ♀ Sauss.” [handwritten on yellow paper]; “Holotypus” [printed on red card]. Specimen directly pinned but splitting apart where the pin is inserted. The ends of the antennae are lost. Box Blattodea 56.

*Salganea foveolata* Saussure, 1895

**laevicollis** Saussure, 1873: 151-152 [*Panesthia*]

La Nouvelle-Hollande. Unspecified number of ♀.

The MHNG collection contains four specimens. A ♀
specimen (MHNG-ARTO-22867) with labels: "603 34 N. S. Wales, Nouv. Holl., Mus d'Aukland" [handwritten on ruled white card]; "603 34 New S. Wales" [handwritten on white paper]; "25" [handwritten on white paper]; "82" [printed on white card]; "Panesthia, laevicollis ♀ Sauss." [handwritten on purple paper]; "♀ Lectotype, det L.M. Roth 1974" [handwritten on white paper with "det L.M. Roth 19" printed]; "Synonym of, Panesthia, cribrata Sauss., det L.M. Roth 1974" [handwritten on white paper with "det. L.M. Roth 19" printed]; "Syntypus" [printed on red paper]. Specimen directly pinned. Both antennae, the left front leg, the tarsi of the right front leg, the tarsi of the left middle leg, the tarsi of the right hind leg and the tibia and tarsi of the left hind leg are lost.

A juvenile specimen (MHNG-ARTO-22868) with labels: "L ♂, Nouv. Holl., M H de Saussure" [handwritten on ruled white card]; "Panesthia, laevicollis, ♂ Sauss." [handwritten on purple paper]; "♂ nymph, Paraelotype, det L.M. Roth 1974" [handwritten on white paper with "det L.M. Roth 19" printed]; "Synonym of, Panesthia, cribrata Sauss., det L.M. Roth 1974" [handwritten on white paper with "det L.M. Roth 19" printed]; "Syntypus" [printed on red paper]. Specimen directly pinned. Both antennae, the right front coxa and leg, the tarsi of the left middle leg, the right middle coxa and leg, the tarsi of the left hind leg and the right hind coxa and leg are missing. The specimen has been repaired with glue and a card mound added on the original pin.

A ♀ specimen (MHNG-ARTO-22869) with labels: "♀ Australie, M H de Saussure" [handwritten on ruled white card]; "Panesthia, laevicollis Sauss, Nouv. Holl." [handwritten on white card]; "Paraelotype, det L.M. Roth 1974" [handwritten on white paper with "det L.M. Roth 19" printed]; "Synonym of, Panesthia, cribrata Sauss., det L.M. Roth 1974" [handwritten on white paper with "det L.M. Roth 19" printed]; "Syntypus" [printed on red paper]. Specimen directly pinned. Both antennae, the tarsi of the left front leg and the entire right hind leg are missing.

A juvenile specimen (MHNG-ARTO-22870) with labels: "♀ Australie, M H de Saussure" [handwritten on ruled white card]; "Panesthia, laevicollis, ♀ Sauss." [handwritten on purple paper]; "♂ nymph, Paraelotype, det L.M. Roth 1974" [handwritten on white paper with "det L.M. Roth 19" printed]; "Synonym of, Panesthia, cribrata Sauss., det L.M. Roth 1974" [handwritten on white paper with "det L.M. Roth 19" printed]; "Syntypus" [printed on red paper]. Specimen directly pinned. The left antenna, two tarsal segments of the right middle leg, the tarsi of the left hind leg and two tarsal segments of the right hind leg are lost. Box Blattodea 57.

A junior synonym of *Panesthia cribrata* Saussure, 1864

*mandarinea* Saussure, 1863: 168-169, fig. 25 [*Panesthia*]
La Chine (Musée de Neuchâtel). One specimen (without
abdomen).

The MHNG collection contains one specimen under this name. A ♀ specimen (MHNG-ARTO-22895) with labels: "Panesthia, mandarinea Sauss." [handwritten on yellow paper]; "Panesthia, mandarinea Sauss.m det L.M Roth 1975, ♀ Lectotype" [handwritten on white paper with "det L.M. Roth 19" printed]. The species name label in the insect box has the handwritten locality "China". Specimen directly pinned, with front wings spread and hind wings folded. The antennae, the tarsi of the left front leg, the tarsi of the left middle leg, all of both hind legs and the abdomen are missing. Part of an antenna and a set of tarsi, presumably from this specimen, are glued to a card mount on the original pin.

It appears that Roth thought this was the second of the
two species described by Saussure as *P. mandarinea*
because the first was based on a single specimen (the
holotype). However, there is a handwritten note on a
separate pin reading "Mandarinea, Sauss. Mel. Orth.
I 1er fasc., No 37 pl I fig. 25, nec 2e fasc. p106 pl 3,
fig 23. La dernière fig, [illegible] Caeparia, Saussurei
W.-Mas., J.A.S.B 1876, 190" which implies that this is
the holotype of the first species. No specimens could be
found in the MHNN.

A junior synonym of *Panesthia transversa* Burmeister,
1838

*mandarinea* Saussure, 1869b: 286-287, fig. 23 [*Panesthia*]

Les Indes orientales. Unspecified series of ♀.

Saussure thought that he was redescribing *P. mandarinea*
Saussure, 1863, which had been based on one damaged
specimen. However, this type series was not conspecific
with *P. mandarinea* Saussure, 1863, and the taxon was
given the replacement name *P. saussurei* Wood-Mason,
1876.

No specimens found in the MHNG collection.

Replaced by *Caeparia saussurei* (Wood-Mason, 1876)

*nicobarensis* Saussure, 1895: 308, 316 [*Panesthia*]

Insulae Nicobarense. Unspecified number of ♂ and ♀.
The MHNG collection contains two specimens. A ♂ specimen (MHNG-ARTO-22807) with labels: "106, Nicobar I" [handwritten on white card]; "Nicobar Is" [handwritten on white paper]; "Panesthia, nicobarensis, ♂ Sauss" [handwritten on yellow paper]; "♂ gen. slide #1, Panesthia, nicobarensis Sauss., det L.M. Roth 1975, ♂ Lectotype" [handwritten on white paper with "det L.M. Roth 19" printed]; "Syntypus" [printed on red paper]. Specimen directly pinned. The right antenna, the tarsi of the left front leg, the tarsi of both middle legs and the tarsi of both hind legs are missing. The dissected genitalia are on a microscope slide with labels "1 Geneva, Panesthia, Nicobarensis, Nicobar Islands" and "♂ genitalia. Lectotype, L. Roth 1975" pinned in the insect box.

A juvenile specimen (MHNG-ARTO-22806) with labels:
"110, Nicobar Is" [handwritten on white paper]; "Nicobars F.W.M" [handwritten on white card disc]; "Panesthia, nicobarensis, ♂ Sauss." [handwritten on yellow paper]; "Panesthia ♂ nymph, nicobarensis Sauss, det L.M. Roth 1975, Paralectotype" [handwritten on white paper with "det L.M. Roth 19" printed]; "Syntypus" [printed on red paper]. Specimen directly pinned. The tibia and tarsi of the right hind leg and the entire left hind leg are lost. The whereabouts of the ♀ specimen(s) is unknown. Box Blattodea 56.

A junior synonym of *Panesthia angustipennis angustipennis* (Illiger, 1801)

*nigrita* Saussure, 1895: 308, 317-318 [*Panesthia*]

Macassar (Mus. Genavense). Unspecified number of ♂ and ♀.

The MHNG collection contains two specimens. A ♀ specimen (MHNG-ARTO-22808) with labels: "Bontheim, Macassar" [handwritten on white paper]; "Panesthia, nigrita Sss., ♀" [handwritten on yellow paper]; "♀ Lectotype, Panesthia, nigrita Saussure, det. L.M. Roth 1974" [handwritten on white paper with "det. L.M. Roth 19" printed]; "= Panesthia, angustipennis (Ill.), det. L.M. Roth 1975" [handwritten on white paper with "det. L.M. Roth 19" printed]; "Syntypus" [printed on red paper]. Specimen directly pinned with left wings spread and right wings folded].

A ♂ specimen (MHNG-ARTO-22809) with labels: "620 67 Sumatra, Iles de Sonde, Mr C. Pictet" [handwritten on ruled white card]; "DELI SUMATRA, C. PICTET" [printed on yellow paper] "Panesthia, nigrita, ♂ Sauss." [handwritten on yellow paper]; "Panesthia slide # 2, nigrita Sauss., det. L.M. Roth 1974" [handwritten on white paper with "det. L.M. Roth 19" printed]; "= Panesthia, angustipennis (Ill.), det. L.M. Roth 1975" [handwritten on white paper with "det. L.M. Roth 19" printed]. Specimen directly pinned with wings roughly folded. The left antenna, the tibia and tarsi of the left front leg, the tarsi of the right front leg, the claw of the right middle leg and the tarsi of the left hind leg are lost. The dissected genitalia are on a microscope slide with labels "2 Geneva, Panesthia, angustipennis, Sumatra (Ill.), L. Roth 1975" and "♂ Paralectotype of, Panesthia, nigrita Sauss." pinned in the insect box. The identification of this specimen as part of the type series is very doubtful, given the provenance. The whereabouts of the ♂ syntype(s) is unknown. Box Blattodea 56.

A junior synonym of *Panesthia angustipennis angustipennis* (Illiger, 1801)

*obscura* Saussure, 1873: 146-147 [*Parahormetica*]

Suivant l'étiquette, Java. Unspecified number of ♀.

The MHNG collection contains one ♀ specimen (MHNG-ARTO-22627) with labels: "Java, Mr H de Saussure" [handwritten on ruled white card]; "♀ Java, Mr H de Saussure" [handwritten on white card];

"Parahormetica obscura Sss. ♀" [handwritten on yellow paper]; "Syntypus" [printed on red paper]. Specimen directly pinned. The ends of both antennae and the tarsi of the right hind leg are lost. Box Blattodea 49.

*Panesthia obscura* (Saussure, 1873)

*ornata* Saussure, 1873: 152-153, fig. 54 [*Panesthia*]

Java (La Chine?). Unspecified number of ♂.

The MHNG collection contains one specimen (MHNG-ARTO-22888) with labels: "♂ Java, M H de Saussure" [handwritten on ruled white card]; "ornata Sauss., (Panesthia), Java" [handwritten on white card]; "Panesthia, ornata, ♀ Sauss." [handwritten on yellow paper]; "Syntypus" [printed on red paper]. Specimen directly pinned with wings spread. The tarsi of the right middle leg are lost. There is mould damage to the underside. The type series was undefined and so the specimen must be regarded as a syntype. Box Blattodea 57.

*Panesthia ornata* Saussure, 1873

*quadriglumis* Saussure, 1895: 308, 316-317, fig. 3 [*Panesthia*]

Australia; Gayndah (Godeffroy). More than one ♀.

The MHNG collection contains two specimens. A ♀ specimen (MHNG-ARTO-22873) with labels: "602 33 Gayndah, Nouv. Holl., Mus. Godeffroy" [handwritten on ruled white card]; "603 33 Panesthia sp.?, Gayndah M. Gdf." [handwritten on white card]; "15002" [handwritten on white card]; "C. Australia, [illegible], 25.9.93" [handwritten on white paper]; "84" [printed on white card]; "Panesthia, 4-glumis, ♀ Sss." [handwritten on purple paper]; "Syntypus" [printed on red paper]. Specimen directly pinned. Both antennae, the right front leg, the tarsi of the right middle leg and the tarsi of both hind legs are missing.

A ♀ specimen (MHNG-ARTO-22874) with labels: "602 33 Gayndah, Nouv. Holl., Mus. Godeffroy" [handwritten on ruled white card]; "603 33 Panesthia sp.?, Gayndah M. Gdf." [handwritten on white card]; "15002" [handwritten on white card]; "Panesthia, 4-glumis, ♀ Sss." [handwritten on purple paper]; "Syntypus" [printed on red paper]. Specimen directly pinned. The tarsi of the left hind leg are lost. Box blattodea 57.

A junior synonym of *Panesthia cribrata* Saussure, 1864

*regina* Saussure, 1863: 167-168, fig. 24 [*Panesthia*]

Java (Musée de Neuchâtel). Unspecified series.

The MHNG collection has no specimen under this name. There are three specimens (MHNG-ARTO-22811, 22812 and 22815) from Java placed under the name *Salganea morio* but no indication that they are part of the type series. Box Blattodea 56.

There is a ♂ specimen in the MHNN (MHNN-65-3307) which is here considered to be a syntype.

A junior synonym of *Salganea morio* (Burmeister, 1838)
**rugulata** Saussure, 1895: 303, 304 [*Salganea*]

Java. Unspecified number of ♀.

The MHNG collection contains four specimens. A ♀ specimen (MHNG-ARTO-22803) with labels: “♀ Java, M H de Saussure” [handwritten on ruled white card]; “Salganea, rugulata, ♀ Sss.” [handwritten on yellow paper]; “Syntypus” [printed on red paper]. Specimen directly pinned with wings spread. The left antenna, the tarsi of both middle legs and most of the tibia and the tarsi of the left hind leg are lost.

A ♀ specimen (MHNG-ARTO-22802) with the same label data. Specimen directly pinned with wings spread. The tarsi of both front legs, the tarsi of the left middle leg, the tarsi of the right hind leg and most of the left hind wing are lost.

A juvenile specimen (MHNG-ARTO-22804) with the same label data. Specimen directly pinned. The body has broken where the pin was inserted, and the abdomen and part of the thorax have been secured on a separate pin. Most of both antennae, the tarsi of the right middle leg and the entire left hind leg are lost.

A juvenile specimen (MHNG-ARTO-22805) with labels: “larva, Java, M H de Saussure” [handwritten on ruled white card]; “Salganea, rugulata, ♀ Sss.” [handwritten on yellow paper]; “Syntypus” [printed on red paper]. Specimen directly pinned. Box Blattodea 56.

*Salganea rugulata* Saussure, 1895

**serrata** Saussure, 1895: 307, 315 [*Panesthia*]

India; Sink [sic]. Unspecified number of ♀.

No specimens found in the MHNG collection. The whereabouts of the type series is unknown.

*Panesthia serrata* Saussure, 1895

**sinuata** Saussure, 1895: 318-319 [*Panesthia*]

Singapore. Unspecified number of ♂.

No specimens found in the MHNG collection. The whereabouts of the type series is unknown.

*Panesthia sinuata* Saussure, 1895

**stellata** Saussure, 1895: 307, 313-314 [*Panesthia*]

India orientalis; Sikkim. Unspecified number of ♂ and ♀.

The MHNG collection contains two specimens. A ♂ specimen (MHNG-ARTO-22875) with labels: “160, Assam” [handwritten on white card]; “Seebsaugor, S.E. Peal” [handwritten on a greenish card disc]; “Panesthia, stellata, ♂ Sss.” [handwritten on yellow paper]; “Syntypus” [printed on red paper]. Specimen directly pinned. Both antennae, the tarsi of both middle legs and the tarsi of both hind legs are missing.

A ♂ specimen (MHNG-ARTO-22876) with labels: “Sikkim, 3212, 8” [printed on white card with “Sikkim” handwritten]; “312, Sikkim” [handwritten on white card]; “Panesthia, stellata, ♂ Sss.” [handwritten on yellow paper]; “Syntypus” [printed on red paper].

Specimen directly pinned. Both antennae, the right front leg and the tarsi of the right middle leg are lost.

The whereabouts of the ♀ syntype(s) is unknown. Box Blattodea 57.

*Panesthia stellata stellata* Saussure, 1895

**stenotarsis** Saussure, 1895: 324, 325, fig. 2 [*Miopanesthia*]

Java. Unspecified number of ♂.

The MHNG collection contains one ♂ specimen (MHNG-ARTO-22908) with labels: “121 19, Java” [handwritten on white card]; “Miopanesthia, stenotarsis, ♂ Sss.” [handwritten on yellow paper]; “Syntypus” [printed on red paper]. Specimen directly pinned with wings spread. The ends of the antennae and the claw of the right hind leg are missing. The type series was undefined and so the specimen must be regarded as a syntype. Box Blattodea 57.

*Miopanesthia stenotarsis* Saussure, 1895

## ACKNOWLEDGEMENTS

Thanks are due to Anita Hollier and Peter Schwendinger for comments on the manuscript. Special thanks are due to Emmanuelle Werlen for entering the MHNG specimens into the MHNG database. We are grateful to Ben Price for exporting data from the BMNH database to facilitate searching, to Joachim Holstein for information about the holdings of the SMNS and to Jessica Litman for information about the holdings of the MHNN.

## REFERENCES

Beccaloni G.W. 2014. Cockroach species file online. Version 5.0/5.0. Available at: http://Cockroach.SpeciesFile.org (accessed n.n.2024).

Biolley P. 1900. IV. Ortópteros recogidos en Costa Rica desde 1890 a 1900. *Informe Museo Nacional de Costa Rica* 1899-1900: 40-49.

Burmeister H. 1838. Kakerlaken, Schaben. Blattina (pp. 469-517). In: Burmeister H. (ed.). *Handbuch der Entomologie* Volume 2, Part 1. Berlin, Reimer, pp. 397-571.

Erichson W.F. 1848. Insecten (pp. 553-617). In: Schomburgk R. (ed.). *Versuch einer Fauna und Flora von Britisch-Guiana*. Weber, Leipzig, 726 pp.

Eschscholtz F. 1822. *Entomographien*. Erste Lieferung. Mit zwei illuminierten Kupfertafeln. G. Reimer, Berlin, 128 pp.

Guérin-Méneville M.F.E. 1857. *Histoire physique, politique et naturelle de l'Ile de Cuba*. Partie 7. Animaux articulés à pieds articulés. Arthus Bertrand, Paris, lxxxvii + 868 pp.

Hebard M. 1921. Studies in the Dermaptera and Orthoptera of Colombia. Second paper. Dermaptera and Orthopterous families Blattidae, Mantidae and Phasmidae. *Transactions of the American Entomological Society* 47: 107-168, pl. 8-10.

Hollier A. & Hollier J. 2012. Henri de Saussure's expedition to Mexico and the West Indies (1854-56). *Antenna* 36(4): 231-238.
Hollier A. & Hollier J. 2013. A re-evaluation of the nineteenth
century naturalist Henri de Saussure. *Archives of Natural
History* 40: 302-319.

Hollier J. & Hollier A. 2018. Leo Zehntner, Swiss pioneer of
tropical applied entomology. *Antenna* 42 (2): 56-60.

Hollier J., Marshall J. & Legendre F. 2020. An annotated list of
the Blattodea (Insecta) described by Henri de Saussure. Part
1: the Corydiidae. *Revue suisse de Zoologie* 127 (2):
341-248.

Illiger J.K.W. 1801. Neue Insekten. *Magazin für Insektenkunde* 1: 163-208.

Linnaeus C. 1758. *Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentis synonymis, locis.* 10th edition. Holmiae, Laurentii Salvi, 823 pp.

Saussure H. de 1862. *Orthoptera nova Americana (Diagnoses praeliminares). Revue et Magasin de Zoologie pure et appliquée* 14: 163-171, 227-234.

Saussure H. de 1863. Mélanges Orthoptérologiques. Fascicule I Blattidae. *Mémoires de la Société de physique et d'histoire naturelle de Genève* 17: 129-172.

Saussure H. de 1864a. *Blattarum novarum species aliquot. Revue et Magasin de Zoologie pure et appliquée* 16: 305-326, 341-348.

Saussure H. de 1864b. *Orthoptères de l'Amérique moyenne*. Ramboz & Schuchardt, Genève, 271 pp.

Saussure H. de 1869a. *Blattarum novarum species aliquot. Revue et Magasin de zoologie pure et appliquée* 21: 109-113.

Saussure H. de 1869b. Mélanges Orthoptérologiques. Fascicule II. Blattides et Phasmides. *Mémoires de la Société de physique et d'histoire naturelle de Genève* 20: 227-328.

Saussure H. de 1870. Famille des Blattides (pp. 1-123, plates 1-2). *In: Saussure H. de, 1870-1874. Mission scientifique au Mexique et dans l'Amérique Centrale.* Recherches zoologiques 6 (section 1): 1-533, 8 pl.

Saussure H. de 1873. Mélanges Orthoptérologiques. Fascicule IV. Mantides et Blattides. *Mémoires de la Société de physique et d'histoire Naturelle de Genève* 23: 1-163.

Saussure H. de 1893. De quelques genres de Blattes. *Societas Entomologica* 8: 57-58, 67-68.

Saussure H. de 1895. Révision de la tribu des Panesthiens et de celle des Epilampriens, Orthoptères de la famille des Blattes. *Revue suisse de Zoologie* 3: 299-364.

Saussure H. de & Zehntner L. 1893. *Blattidae (part)* (pp. 13-104). *In: Godman F.D. & Salvin O. (eds).* *Biologia Centrali-Americana*. Insecta Orthoptera (Orthoptera Genuina) Volume 1. 458 pp., 22 pl.

Saussure H. de, Zehntner L. 1894. *Blattidae (part)* (pp. 105-123). *In: Godman F.D. & Salvin O. (eds).* *Biologia Centrali-Americana*. Insecta Orthoptera (Orthoptera Genuina) Volume 1. 458 pp., 22 pl.

Saussure, H. de & Zehntner L. 1895. Revision de la tribu des Périsphaeriens (Insectes Orthoptères de la famille des Blattides). *Revue suisse de Zoologie* 3: 1-59.

Serville J.G.A. 1838. *Histoire naturelle des insectes*. Orthoptères. Roret, Paris, XVIII + 776 pp., 14 pls.

Tepper J.G.O. 1893. The Blattariae of Australia and Polynesia. *Transactions of the Royal Society of South Australia* 17: 25-126.

Tepper J.G.O. 1894. Supplemental notes on the Blattariae of Australia. *Transactions of the Royal Society of South Australia* 18: 169-189.

Wood-Mason J. 1876. Descriptions of new species of Blattidae belonging to the genus *Panesthia*. *Journal of the Asiatic Society of Bengal* 45(2): 189-190.