*Tovomita lanceolata* Cuatrec., Anales Inst. Biol. Univ. Nac. México 20: 102. (Cuatrecasas 1950). — *Arawakia lanceolata* (Cuatrec.) L.Marinho, *Molec. Phylogen. Evol.* 134: 147 [publ. online Feb. 2019] (Marinho 2019).

*Tovomita glossophylla* Cuatrec., *Revista Acad. Colomb. Ci. Exact.* 8 (29): 62 [Nov. 1950] (Cuatrecasas 1950). — *Arawakia glossophylla* (Cuatrec.) L.Marinho, *Molec. Phylogen. Evol.* 134: 147 [publ. online Feb. 2019] (Marinho 2019).

*Clusia pithecobia* Standl. & L.O.Williams, *Ceiba* 1 (4): 244 (Standley & Williams 1951). — *Tovomita pithecobia* (Standl. & L.O.Williams) Gahagen, *Syst. Bot.* 40 (4): 985 [15 Dec. 2015] (Gahagen 2015). — *Arawakia pithecobia* (Standl. & L.O.Williams) L.Marinho, *Molec. Phylogen. Evol.* 134: 148 [publ. online Feb. 2019] (Marinho 2019).

*Tovomita angustata* Steyerm., *Fieldiana*, Bot. 28 (2): 399 (Steyermark 1952). — *Arawakia angustata* (Steyerm.) L.Marinho, *Molec. Phylogen. Evol.* 134: 147 [publ. online Feb. 2019] (Marinho 2019).

*Tovomita rileyi* Cuatrec., *Brittonia* 14 (1): 52 (Cuatrecasas 1962). — *Arawakia rileyi* (Cuatrec.) L.Marinho, *Molec. Phylogen. Evol.* 134: 148 [publ. online Feb. 2019] (Marinho 2019).

*Tovomita coriacea* Maguire, *Phytologia* 36 (4): 406 [22 Aug. 1977] (Maguire 1977). — *Arawakia coriacea* (Maguire) L.Marinho, *Molec. Phylogen. Evol.* 134: 147 [publ. online Feb. 2019] (Marinho 2019).

*Tovomita caputmonsis* Gahagen, *Syst. Bot.* 40 (4): 975 [15 Dec. 2015] (Gahagen 2015). — *Arawakia caputmonsis* (Gahagen) L.Marinho, *Molec. Phylogen. Evol.* 134: 147 [publ. online Feb. 2019] (Marinho 2019).

*Tovomita divesora* Gahagen, *Syst. Bot.* 40 (4): 975 [15 Dec. 2015] (Gahagen 2015). — *Arawakia divesora* (Gahagen) L.Marinho, *Molec. Phylogen. Evol.* 134: 147 [publ. online Feb. 2019] (Marinho 2019).

*Tovomita manchamancha* Gahagen, *Syst. Bot.* 40 (4): 982 [15 Dec. 2015] (Gahagen 2015). — *Arawakia manchamancha* (Gahagen) L.Marinho, *Molec. Phylogen. Evol.* 134: 148 [publ. online Feb. 2019] (Marinho 2019).

*Tovomita panamaea* Gahagen, *Syst. Bot.* 40 (4): 983 [15 Dec. 2015] (Gahagen 2015). — *Arawakia panamaea* (Gahagen) L.Marinho, *Molec. Phylogen. Evol.* 134: 148 [publ. online Feb. 2019] (Marinho 2019).

*Tovomita parvifolia* Gahagen, *Syst. Bot.* 40 (4): 983 [15 Dec. 2015] (Gahagen 2015). — *Arawakia parvifolia* (Gahagen) L.Marinho, *Molec. Phylogen. Evol.* 134: 148 [publ. online Feb. 2019] (Marinho 2019).

NOTES. — Gahagen, Terbush & Ballard (2016) resurrected several names previously considered synonyms of *T. weddelliana*, and segregated 5 new species from what they called the “*T. weddelliana* species complex”. Their study relied on “quantitative” analyses of morphological characters, based on two to 11 herbarium specimens per species. In our opinion, the number of studied specimens is too low to capture the natural variability of the morphological characters (e.g. twig diameter, pedicel length or leaf blade size). Moreover, the results of their statistical analyses are unconvincing and do not clearly distinguish their 11 so-called species, whose morphological differences could simply reflect the plasticity of a single species found from the Pacific coast to the Tepuis of the Guyana and slopes of the Andes, from 0 to 1300 m altitude. Subsequently, Marinho *et al.* (2019) segregated the “*T. weddelliana* complex” in a new genus, *Arawakia*, on the basis of plastid and nuclear ITS genome analyses of 30 species out of the c. 500 that form the tribe Clusieae. However, their splitting of *Tovomita* into two main groups (one being *Arawakia*) appears much less supported (85-86 bootstrap values and 0.51-

0.66 posterior probabilities) than the monophyly of other genera (99-100 bootstrap/0.85-1 pp). Further, two of the five diagnostic characters that Marinho *et al.* (2019) claim to distinguish *Arawakia* from *Tovomita* do not hold: - white latex, said to be characteristic of *Arawakia* (vs yellow in *Tovomita*), has been observed by the first two authors of the present publication in *T. umbellata* (J.-F. Molino 731 and J.-F. Molino 936, CAY), *T. schomburgkii* (D. Sabatier & M.-F. Prévost 1345, D. Sabatier & M.-F. Prévost 3354 and D. Sabatier & J.-F. Molino 4679, CAY) and *T. caloneura* (D. Sabatier & M.-F. Prévost 1363, CAY); - the sepals, said to be always persistant and adpressed on fruit in *Arawakia* (vs not adpressed in *Tovomita*) are actually not adpressed on fruits of the types of *A. lingulata*, *A. macrocarpa* and *A. rileyi*. The remaining characters Marinho *et al.* (2019) present do not warrant the creation of a new genus: two are purely vegetative (leaf shape and colour of the interpretiolar line, black in *Arawakia* vs not black in *Tovomita*), and the third is epicarp colour (red in *Arawakia* vs green to brown in *Tovomita*).

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: *D. Sabatier* 5799.

SIZE. — Up to 30 m tall (Cuello 1998).

[397] *Tovomita* sp. A

HERBARIUM DATA (FG). — A single collection, *D. Sabatier* 4872.

INVENTORY DATA (FG). — 2 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 18$ cm.

Family COMBRETACEAE R.Br.
Genus *Conocarpus* L.

[398] *Conocarpus erectus* L.
(Fig. 19A)

*Sp. Pl.* 1: 176 [1 May 1753] (Linnaeus 1753), “*erecta*”. — *Terminalia erecta* (L.) Baill., *Hist. Pl. [Baillon]* 6: 266, 275 [Jan.-May 1876] (Baillon 1876). — *Conocarpus erectus* var. arboreus DC., *Prodr.* [*A. P. de Candolle*] 3: 16 [mid Mar. 1828] (Candolle 1828), “*erecta* var. arborea”, nom. illeg. *superfl.* (based on the type of *Conocarpus erectus*).

*Conocarpus procumbens* L., *Sp. Pl.* 1: 177 [1 May 1753] (Linnaeus 1753). — *Conocarpus supinus* Crantz, *Inst. Rei Herb.* 1: 355 (Crantz 1766), nom. illeg. *superfl.* (based on *Conocarpus procumbens*). — *Conocarpus erectus* var. *procumbens* (L.) DC., *Prodr.* [*A. P. de Candolle*] 3: 16 [mid Mar. 1828] (Candolle 1828). — *Terminalia erecta* var. *procumbens* (L.) M.Gómez, *Dicc. Bot. Nombres Vulg. Cub. Puerto-Riq.*: 73 (Gómez 1889).

*Conocarpus acutifolius* Humb. & Bonpl. ex Roem. & Schult., *Syst. Veg. [Roemer & Schultes]* 5: 574 [Dec. 1819] (Roemer & Schultes 1819).

*Conocarpus pubescens* Schumach., *Beskr. Guin. Pl.*: 115 (Schumacher 1827).

NOTE. — A species restricted to mangrove forests.

VERNACULAR NAMES. — Br: mangue-de-botão, mangue-negro.

HERBARIUM DATA (FG). — 5 collections at CAY. Sel. exs.: *M.-F. Prévost & C. Proisy* 4550.

SIZE. — Up to 20 m tall (Stace 2010).
Genus *Laguncularia* C.F.Gaertn.

[399] *Laguncularia racemosa* (L.) C.F.Gaertn.

*Suppl. Carp.*: 209 (Gaertner 1807). — *Conocarpus racemosus* L., *Syst. Nat.*, ed. 10, 2: 930 [7 June 1759] (Linnaeus 1759). — *Horau racemosus* (L.) M.Gómez, *Fl. Cuba* [Gómez & Roig]: 75 (Gómez 1914).

*Schousboea commutata* Spreng., *Syst. Veg. [Sprengel]* 2: 332 [Jan.-May 1825] (Sprengel 1825), *nom. illeg. superfl.* (based on *Conocarpus racemosus*).

*Rhizaeris alba* Raf., *Sylva Tellur.*: 90 (Rafinesque 1838), *nom. illeg. superfl.* (based on *Conocarpus racemosus*).

*Laguncularia obovata* Miq., *Linnaea* 18: 752 ["1844" publ. Aug.-Oct. 1845] (Miquel 1845).

*Laguncularia racemosa* f. longifolia J.F.Macbr., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 8 (2): 125 (Macbride 1930).

**NOTES.** — A species restricted to mangrove forests. Rafinesque (1838: 90) described *Rhizaeris* as a new genus "very diff. from Conocarpus", then "*R. alba* Raf., *Conocarpus racemosa* L. auct. The white Mangrove of Antilles". He apparently thought that *C. racemosus* L. was indeed a *Conocarpus*, that this name had been wrongly used for the white mangrove, and accordingly he proposed a new name. He did not realized that *C. racemosus* is actually the white mangrove and had already been transferred to *Laguncularia* (Gaertner 1807).

**VERNACULAR NAMES.** — Ka: akila'i, ele'u, elewu • Cr: pativyé-gri • Fr: palétuvier gris • Br: falso-mangue, mangue-amarelo, mangue-branco.

**HERBARIUM DATA** (FG). — 37 collections at CAY. Sel. exs.: *D. Sabatier* 2259, dbh 10 cm.

Genus *Terminalia* L.

[400] *Terminalia amazonia* (J.F.Gmel.) Exell
(Fig. 19B)

*Fl. Suriname* 3 (1): 173 (Exell 1935). — *Chuncoa amazonia* J.F.Gmel., *Syst. Nat.*, ed. 13[bis], 2 (1): 702 [late Sep.-Nov. 1791] (Gmelin 1791).

*Gimbernatia obovata* Ruiz & Pav., *Syst. Veg. Fl. Peruv. Chil.* 1: 274 [late Dec. 1798] (Ruiz & Pavón 1798). — *Chuncoa obovata* (Ruiz & Pav.) Pers., *Syn. Pl. [Persoon]* 1: 486 [1 Apr.-15 June 1805] (Persoon 1805), "Chunchoa". — *Terminalia obovata* (Ruiz & Pav.) Steud., *Nomencl. Bot. [Steudel]*, ed. 2, 2: 668 (Steudel 1841), *nom. illeg. hom.*, non Cambess. (Cambessèdes 1830). — *Myrobalanus obovata* (Ruiz & Pav.) Kuntze, *Revis. Gen. Pl.* 1: 237 [5 Nov. 1891] (Kuntze 1891).

*Bucida angustifolia* DC., *Prodr. [A. P. de Candolle]* 3: 10 [mid Mar. 1828] (Candolle 1828). — *Bucida buceras* L. var. angustifolia (DC.) Eichler, *Fl. Bras. [Martius]* 14 (2): 95 [17 Apr. 1867] (Eichler 1867).

*Terminalia odontoptera* Van Heurck & Müll.Arg., *Observ. Bot. [Van Heurck]* 2: 217 [10 Sep. 1871] (Van Heurck & Müller 1871).

*Terminalia excelsa* Liebm. ex Hemsl., *Biol. Cent.-Amer., Bot.* 1 (5): 402 [June 1880] (Hemsley 1880), *nom. nud*.

*Myrobalanus excelsa* Kuntze, *Revis. Gen. Pl.* 1: 237 [5 Nov. 1891] (Kuntze 1891).

**VERNACULAR NAMES.** — Pa: katauma, katumã • Ka: ityutano kwai, katulimya, kwai • Wp: alala munuwí sili, tatulimã • Nt: anangosi, kwata pepee • Cr: angouchi, bwa-blanché, grènn-oko • Br: cuiarana, mirindiba-branca.

**HERBARIUM DATA** (FG). — 49 collections at CAY. Sel. exs.: *G.S. Per-röttet s.n.* (original material of *Bucida angustifolia*: G[G00236066]).

**INVENTORY DATA** (FG). — 61 trees in 24 plots; $F_{max} = 1.9\%$; dbh$_{inv} = 71$ cm.

[401] *Terminalia aubletii* Gere & Boatwr.
(Fig. 19C)

*Bot. J. Linn. Soc.* 184 (3): 319 [epubl. 22 June 2017] (Gere & Boatwright 2017). — *Pamea guianensis* Aubl., *Hist. Pl. Guiane* 2: 946 [Jun.-Dec. 1775] (Aublet 1775). — *Terminalia pamea* DC., *Prodr. [A. P. de Candolle]* 3: 13 [mid Mar. 1828] (Candolle 1828), *nom. illeg. superfl.* (based on *Pamea guianensis*). — *Myrobalanus pamea* Kuntze, *Revis. Gen. Pl.* 1: 237 [5 Nov. 1891] (Kuntze 1891), "Pamaea". — *Buchenavia guianensis* (Aubl.) Alwan & Stace, *Nordic J. Bot.* 5 (5): 447 (Alwan & Stace 1985).

**NOTES.** — *Myrobalanus pamea* Kuntze is based on the illegitimate *Terminalia pamea* DC., but is legitimate as a replacement name for *Pamea guianensis* Aubl. (which is indirectly cited through *T. pamea*), *non M. guianensis* (Aubl.) Kuntze (based on *Tanibouca guianensis* Aubl., and synonym of *Terminalia dichotoma* [1818: 177]).

**VERNACULAR NAMES.** — Ka: katulimya • Wp: kwata kaya, tatulimã sili • Nt: anangosi • Cr: zanmann-sovaj • Fr: amandier sauvage • Br: tanimbucá.

**HERBARIUM DATA** (FG). — 6 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material of *Pamea guianensis*: BM[BM000953713]).

**INVENTORY DATA** (FG). — 17 trees in 15 plots; $F_{max} < 1\%$; dbh$_{inv} = 101$ cm.

[402] *Terminalia carinata* Sabatier & J.Engel
(Fig. 19D)

*Adansonia*, sér. 3, 42 (16): 262 [2 Nov. 2020] (Engel & Sabatier 2020).

**VERNACULAR NAMES.** — Pa: karevru-danô • Ka: kalalawa akunepili • Nt: anangosi, kwata pepe, kwata pepee • Cr: angouchi, bwa-blanché, grènn-oko • Fr: amandier sauvage • Br: mirindiba-branca.

**HERBARIUM DATA** (FG). — 27 collections at CAY. Sel. exs.: *D. Sabatier et al. 4891* (holo-, CAY[CAY182887]; iso-, K, P[P01155925]).

**INVENTORY DATA** (FG). — 56 trees in 32 plots; $F_{max} < 1\%$; dbh$_{inv} = 120$ cm.

[403] *Terminalia catappa* L.

*Mant. Pl.* 1: 128 (Linnaeus 1767). — *Juglans catappa* (L.) Lour., *Fl. Cochinch.* 2: 573 [Sep. 1790] (Loureiro 1790). — *Myrobalanus catappa* (L.) Kuntze, *Revis. Gen. Pl.* 1: 237 [5 Nov. 1891] (Kuntze 1891). — *Buceras catappa* (L.) Hitchc., *Rep. (Annual) Missouri Bot. Gard.* 4: 85 (Hitchcock 1893).

*Phytolacca javanica* Osbeck, *Dagb. Ostind. Resa*: 276 (Osbeck 1757).
*Terminalia moluccana* Lam., *Encycl. [J. Lamarck et al.]* 1 (2): 349
[1 Aug. 1785] (Lamarck 1785).

*Terminalia ovatifolia* Noronha, *Verh. Batav. Genootsch. Kunst.* 5 (4):
27 (Noronha 1790).

*Badamia commersonii* Gaertn., *Fruct. Sem. Pl.* 2: 90 [Sep.-Nov.
1790] (Gaertner 1790), "*Commersoni*".

*Terminalia subcordata* Humb. & Bonpl. ex Willd., *Sp. Pl.*, ed. 4 4
(2): 968 [Apr. 1806] (Willdenow 1806). — *Terminalia catappa*
var. *subcordata* (Humb. & Bonpl. ex Willd.) DC., *Prodr. [A. P. de
Candolle]* 3: 11 [mid Mar. 1828] (Candolle 1828).

*Myrobalanus badamia* Poir., *Encycl. [J. Lamarck et al.]* Suppl. 3:
707 [3 Sep. 1814] (Poiret 1814), "Mirobolanus". — *Terminalia
badamia* (Poir.) DC., *Prodr. [A. P. de Candolle]* 3: 12 [mid Mar.
1828] (Candolle 1828).

*Myrobalanus terminalia* Poir., *Encycl. [J. Lamarck et al.]* Suppl. 3:
707 [3 Sep. 1814] (Poiret 1814), "*Mirobolanus*".

*Terminalia myrobalana* Roth, *Nov. Pl.* *Sp.* 378 [Apr. 1821] (Roth
1821).

*Terminalia intermedia* Bertero ex Spreng., *Syst. Veg. [Sprengel]* 2:
359 [Jan.-May 1825] (Sprengel 1825).

*Terminalia kydiana* Roxb. ex Wall., *Numer. List* no. 3974 (Wallich
1831).

*Terminalia latifolia* Blanco, *Fl. Filip. [F.M. Blanco]* 376 (Blanco 1837).

*Terminalia paraensis* Mart., *Flora* 24 (2, Beibl.): 24 (Martius 1841).

*Terminalia mauritiana* Blanco, *Fl. Filip. [F.M. Blanco]*, ed. 2: 264
(Blanco 1845).

*Terminalia rubrigemmis* Tul., *Ann. Sci. Nat., Bot.* sér. 4, 6: 102
(Tulasne 1856). — *Myrobalanus rubrigemmis* (Tul.) Kuntze, *Revis.
Gen. Pl.* 1: 237 [5 Nov. 1891] (Kuntze 1891).

*Myrobalanus commersonii* Kuntze, *Revis. Gen. Pl.* 1: 237 [5 Nov.
1891] (Kuntze 1891).

NOTE. — A species native to South-West Asia, introduced and naturalised in most tropical regions.

VERNACULAR NAMES. — Pa: zamâd, zaman • Ka: amandra • Nt: amanda, amandaa • Cr: zanmann • Fr: amandier pays • Br: amendoira-da-praia.

HERBARIUM DATA (FG). — 13 collections at CAY. Sel. exs.: *S.A. Mori et al.* 25515.

SIZE. — Up to 35 m tall (Stace 2010).

[404] *Terminalia dichotoma* G.Mey.

*Prim. Fl. Esseq.* 177 [Nov. 1818] (Meyer 1818). — *Terminalia latifolia* Sw. var. *dichotoma* (G.Mey.) DC., *Prodr. [A. P. de Candolle]* 3: 12 [mid Mar. 1828] (Candolle 1828).

*Tanibouca guianensis* Aubl., *Hist. Pl. Guiane* 1: 448 [Jun.-Dec. 1775)
(Aublet 1775). — *Myrobalanus guianensis* (Aubl.) Kuntze, *Revis.
Gen. Pl.* 1: 237 [5 Nov. 1891] (Kuntze 1891). — *Terminalia
tanibouca* Rich., *Actes Soc. Hist. Nat. Paris* 1: 109 [Oct. 1792]
(Richard 1792), *nom. illeg. superfl.* (based on *Tanibouca guianensis*).

*Catappa guianensis* C.F.Gaertn., *Suppl. Carp.* 207 (Gaertner 1807).

NOTE. — According to Stace (2010), *Catappa guianensis* Gaertn.
is not based on *Tanibouca guianensis* Aubl.

VERNACULAR NAMES. — Ka: kalalawa akunepili • Wp: alala munuwi
• Cr: angouchi, grènn-oko • Br: cinzeiro, tanimbucá.

HERBARIUM DATA (FG). — 26 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material of *Tanibouca guianensis*: BM[BM000953723]).

INVENTORY DATA (FG). — 13 trees in 5 plots; Fmax < 1%; dbhinv = 59 cm.

[405] *Terminalia grandis* (Ducke) Gere & Boatwr.

Bot. *J. Linn. Soc.* 184 (3): 320 [epubl. 22 June 2017] (Gere & Boatwright 2017). — *Buchenavia grandis* Ducke, *Arch. Jard. Bot. Rio de Janeiro* 4: 148 (Ducke 1925).

*Buchenavia huberi* Ducke, *Bol. Tècn. Inst. Agron. N.* 4: 24 [31 Mar. 1945] (Ducke 1945).

VERNACULAR NAMES. — Nt: anangosi • Cr: zanmann-sovaj • Fr: amandier sauvage • Br: marindiba, tanimbucá.

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 4829*.

INVENTORY DATA (FG). — 23 trees in 19 plots; Fmax < 1%; dbhinv = 60.5 cm.

[406] *Terminalia guyanensis* Eichler

*Fl. Bras. [Martius]* 14 (2): 88 [17 Apr. 1867] (Eichler 1867).

*Myrobalanus eichleriana* Kuntze, *Revis. Gen. Pl.* 1: 237 [5 Nov. 1891] (Kuntze 1891).

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: *P.A. Poiteau s.n.* (holo-, B[not seen, photo F neg. N° 14333]; iso-, GH[GH00068628], K[K000640657], P[P01901252, P01901253, P01901254], U[U0001197]).

INVENTORY DATA (FG). — 9 trees in 7 plots; Fmax < 1%; dbhinv = 80 cm.

[407] *Terminalia lucida* Hoffmanns. ex Mart. & Zucc.

*Flora* 7 (1, Beibl.): 130 [25 Apr. 1824] (Martius & Zuccarini 1824). — *Myrobalanus lucida* (Hoffmanns. ex Mart. & Zucc.) Kuntze, *Revis. Gen. Pl.* 1: 237 [5 Nov. 1891] (Kuntze 1891).

*Terminalia firma* Mart. & Zucc., *Flora* 7 (1, Beibl.): 130 [25 Apr. 1824] (Martius & Zuccarini 1824).

*Terminalia eriantha* Benth., *Hooker's J. Bot. Kew Gard. Misc.* 2: 240
(Bentham 1850).

*Terminalia scutifera* Planch. ex M.A.Lawson, *Fl. Trop. Afr.* 2: 417
(Lawson 1871). — *Myrobalanus scutifera* (Planch. ex M.A.Lawson)
Kuntze, *Revis. Gen. Pl.* 1: 237 [5 Nov. 1891] (Kuntze 1891).

*Terminalia hayesii* Pittier, *Contr. U.S. Natl. Herb.* 18 (6): 239 [22 Sep. 1917] (Pittier 1917).

*Terminalia nyssifolia* Britton, *Bull. Torrey Bot. Club* 48 (12): 333
["1921" publ. 1922] (Britton 1922), "*nyssaefolia*".
<figure><img src="image_30.png" /><figcaption>FIG. 19. — Combretaceae: **A**, *Conocarpus erectus* L. (M.-F. Prévost & S. Gonzalez 5313); **B**, *Terminalia amazonia* (J.F.Gmel.) Exell; **C**, *Terminalia aubletii* Gere & Boatwr. (D. Sabatier 2283); **D**, *Terminalia carinata* Sabatier & J.Engel (D. Sabatier et al. 4891). © D. Sabatier/IRD.</figcaption></figure>

NOTES. — Martius (1824: 44) cited the collection locality of *T. firma* Mart. & Zucc. as "*Habitat in editis montis Serra de Araracoara ad fluvium Japurá, Provinciae flum. nigri.*" Stace (2010) cited the locality as being situated in Brazil (Amazonas), but in January 1820 Martius collected the original gathering in Araracuara, on the Rio Caquetá, which is in Colombia.

VERNACULAR NAMES. — Wp: alala munuwi • Cr: angouchi, bwa-blanché, grènn-oko • Br: cinzeiro, tanimbucá.

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *M.-F. Prévost 1221*; *M.-F. Prévost & D. Sabatier 4895*, dbh 12 cm.

[408] *Terminalia macrophylla* (Eichler) Gere & Boatwr.

Bot. J. Linn. Soc. 184 (3): 320 [epubl. 22 June 2017] (Gere & Boatwright 2017). — *Buchenavia macrophylla* Eichler, *Flora* 49 (11): 166 (Eichler 1866). — *Terminalia macrophylla* Spruce ex Eichler, *Fl. Bras. [Martius]* 14 (2): 98 [17 Apr. 1867] (Eichler 1867), *nom. nud. pro syn.*

Buchenavia stellae Cuatrec., Fieldiana, Bot. 27 (1): 109 (Cuatrecasas 1950).

VERNACULAR NAMES. — Ka: katulimya • Wp: kwata kaya, tatulimá sili • Wn: alimi huhu, ekesima, ekesimë • Nt: kwata bobi • Cr: tapouliman, zanmann-sovaj • Br: tanimbucá.

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 5017*, dbh 43 cm.

[409] *Terminalia megalophylla* (Van Heurck & Müll.Arg.) Gere & Boatwr.

Bot. J. Linn. Soc. 184 (3): 320 [epubl. 22 June 2017] (Gere & Boatwright 2017). — *Buchenavia megalophylla* Van Heurck & Müll.Arg., *Observ. Bot. [Van Heurck]* 2: 211 [10 Sep. 1871] (Van Heurck & Müller 1871).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — A single collection, *J.-J. de Granville B-4792*.

SIZE. — Up to 45 m tall (Stace 2009).

[410] *Terminalia nitidissima* Rich.

Actes Soc. Hist. Nat. Paris 1: 109 [Oct. 1792] (Richard 1792). — Myrobalanus nitidissima (Rich.) Kuntze, *Revis. Gen. Pl.* 1: 237 [5 Nov. 1891] (Kuntze 1891). — *Buchenavia nitidissima* (Rich.) Alwan & Stace, *Nordic J. Bot.* 5 (5): 449 (Alwan & Stace 1985).

NOTE. — Known only from French Guiana.
HERBARIUM DATA (FG). — 17 collections at CAY. Sel. exs.: *J.B. Leblond s.n.* (original material G[G00177930]).

INVENTORY DATA (FG). — 5 trees in 5 plots; $F_{max} < 1\%$; $dbh_{inv} = 32.9$ cm.

[411] *Terminalia oblonga* (Ruiz & Pav.) Steud.

Nomencl. Bot. [Steudel], ed. 2, 2: 668 (Steudel 1841). — *Gimbernatia oblonga* Ruiz & Pav., Syst. Veg. Fl. Peruv. Chil. 1: 275 [late Dec. 1798] (Ruiz & Pavón 1798). — *Chuncoa oblonga* (Ruiz & Pav.) Pers., Syn. Pl. [Persoon] 1: 486 [1 Apr.-15 June 1805] (Persoon 1805), “Chunchoa”. — *Myrobalanus oblonga* (Ruiz & Pav.) Kuntze, Revis. Gen. Pl. 1: 237 [5 Nov. 1891] (Kuntze 1891). — *Chuncoa diptera* F.Dietr., Nachtr. Vollst. Lex. Gärtn. 2: 244 [May 1816] (Dietrich 1816), nom. illeg. superfl. (based on *Gimbernatia oblonga*).

*Terminalia tarapotensis* Van Heurck & Müll.Arg., Observ. Bot. [Van Heurck] 2: 213 [10 Sep. 1871] (Van Heurck & Müller 1871).

*Terminalia chiriquensis* Pittier, Contr. U.S. Natl. Herb. 18 (6): 238 [22 Sep. 1917] (Pittier 1917).

*Terminalia obidensis* Ducke, Arch. Jard. Bot. Rio de Janeiro 4: 147 (Ducke 1925).

VERNACULAR NAMES. — Wp: alala munuwi • Cr: angouchi, grènn-oko • Br: cinzeiro, tanimbucá.

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *P. Grenand 1303*.

SIZE. — Brazil, Pará. *D.C. Daly* 1645 (MO), 20 m × 40 cm.

[412] *Terminalia ochroprumna* (Eichler) Gere & Boatwr.

Bot. J. Linn. Soc. 184 (3): 321 [epubl. 22 June 2017] (Gere & Boatwright 2017). — *Buchenavia ochroprumna* Eichler, *Flora* 49 (11): 165 (Eichler 1866).

*Buchenavia discolor* Diels, Verh. Bot. Vereins Prov. Brandenburg 48: 192 [“1906” publ. 8 Mar. 1907] (Diels 1907).

VERNACULAR NAMES. — Br: tanimbucá-do-igapó, tanimbucá-rana.

HERBARIUM DATA (FG). — A single collection, *P. Béna* 1313.

SIZE. — Up to 30 m tall (Stace 2009).

[413] *Terminalia parvifolia* (Ducke) Gere & Boatwr.

Bot. J. Linn. Soc. 184 (3): 321 [epubl. 22 June 2017] (Gere & Boatwright 2017). — *Buchenavia parvifolia* Ducke, Arch. Jard. Bot. Rio de Janeiro 4: 150 (Ducke 1925).

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *D. Sabatier* 2300.

SIZE. — Brazil, Rondônia. *G.T. Prance et al.* 8551 (P), 15 m × 10 cm.

[414] *Terminalia tetraphylla* (Aubl.) Gere & Boatwr.

Bot. J. Linn. Soc. 184 (3): 322 [epubl. 22 June 2017] (Gere & Boatwright 2017). — *Cordia tetraphylla* Aubl., Hist. Pl. Guiane 1: 224 [Jun.-Dec. 1775] (Aublet 1775). — *Firensia lutea* Raf., Sylva Tellur.: 40 (Rafinesque 1838), nom. illeg. superfl. (based on *Cordia tetraphylla*). — *Lithocardium tetraphyllum* (Aubl.) Kuntze, Revis. Gen. Pl. 2: 977 [5 Nov. 1891] (Kuntze 1891). — *Buchenavia tetraphylla* (Aubl.) R.A.Howard, *J. Arnold Arbor* 64 (2): 266 [Apr. 1983] (Howard 1983). — *Gerascanthus tetraphyllus* (Aubl.) Borhidi, *Acta Bot. Hung.* 34 (3-4): 402 (Borhidi 1988).

Bucida capitata Vahl, *Eclog. Amer.* 1: 50 (Vahl 1797). — *Buchenavia capitata* (Vahl) Eichler, *Flora* 49 (11): 165 (Eichler 1866). — *Terminalia capitata* (Vahl) C.Wright, *Anales Acad. Ci. Med. Habana* 5: 410 (Wright 1869).

Hudsonia arborea A.Rob. ex Lunan, *Hort. Jamaic.* 2: 310 (Lunan 1814).

*Terminalia obovata* Cambess., *Fl. Bras. Merid.* [*A. St.-Hil.*] (quarto ed.) 2 (17): 241 [“1829” publ. 4 Dec. 1830] (Cambessèdes 1830). — *Terminalia hilariana* Steud., *Nomencl. Bot. [Steudel]*, ed. 2, 2: 668 (Steudel 1841), nom. illeg. superfl. (based on *Terminalia obovata*).

Bucida angustifolia Spruce ex Eichler, *Flora* 49 (11): 165 (Eichler 1866), nom. illeg. hom., non DC. (Candolle 1828).

Pseudolmedia bucidifolia Bello, *Anales Soc. Esp. Hist. Nat.* 12: 109 (Bello 1883), “bucidaefolia”.

*Buchenavia gracilis* Glaz., *Bull. Soc. Bot. France* 54 (Mém. 3c): 203 [“1907” publ. 22 Feb. 1908] (Glaziou 1908), nom. nud.

*Buchenavia macahensis* Glaz., *Bull. Soc. Bot. France* 54 (Mém. 3c): 203 [“1907” publ. 22 Feb. 1908] (Glaziou 1908), nom. nud.

*Buchenavia vaupesana* Cuatrec., *Fieldiana*, *Bot.* 27 (1): 108 (Cuatrecas 1950).

*Buchenavia ptariensis* Steyerm., *Fieldiana*, *Bot.* 28 (2): 423 (Steyermark 1952).

VERNACULAR NAMES. — Ka: katulimya • Wp: kwata kaya, tatulimā sili • Cr: tapouliman, zanmann-sovaj • Br: cuiarana, mirindiba.

HERBARIUM DATA (FG). — 16 collections at CAY. Sel. exs.: *D. Loubry* 1934.

INVENTORY DATA (FG). — 10 trees in 10 plots; $F_{max} < 1\%$; $dbh_{inv} = 75$ cm.

[415] *Terminalia viridiflora* (Ducke) Gere & Boatwr.

Bot. J. Linn. Soc. 184 (3): 322 [epubl. 22 June 2017] (Gere & Boatwright 2017). — *Buchenavia viridiflora* Ducke, *Arch. Inst. Biol. Veg.* 2 (1): 63 [Sep. 1935] (Ducke 1935).

*Buchenavia amazonia* Alwan & Stace, *Ann. Missouri Bot. Gard.* 76 (4): 1127 (Alwan & Stace 1989).

VERNACULAR NAMES. — Pa: wakaba • Wp: tatulimā • Cr: zanmann-dan-bwa.

HERBARIUM DATA (FG). — 5 collections at CAY. Sel. exs.: *P. Grenand* 1776.

INVENTORY DATA (FG). — 2 trees in 2 plots; $dh_{inv} = 90$ cm.
Family CORDIACEAE R.Br. ex Dumort.
Genus *Cordia* L.

[416] *Cordia alliodora* (Ruiz & Pav.) Oken

Allg. Naturgesch. 3 (2): 1098 (Oken 1841). — *Cerdana alliodora* Ruiz & Pav., *Flora Peruviana* 2: 47 (Ruiz & Pavón 1799). — *Cordia alliodora* (Ruiz & Pav.) Cham., *Linnaea* 8 (2): 121 (Chamisso 1833), *nom. inval.* (genus name and specific epithet not associated [Turland et al. 2018: Art. 35.2]). — *Lithocardium alliodorum* (Ruiz & Pav.) Kuntze, *Revis. Gen. Pl.* 2: 976 [5 Nov. 1891] (Kuntze 1891). — *Gerascanthus alliodorus* (Ruiz & Pav.) Kuhlm. & Mattos, *Loefgrenia* 47: 1 (Kuhlmann & Mattos 1970), “*alliodora*”.

VERNACULAR NAMES. — Nt: wataa kanu • Cr: sèd-sam • Fr: cèdre sam • Br: freijó, freijó-branco, louro-alho.

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: S.A. *Mori et al.* 24798.

SIZE. — Up to 40 m tall (Gaviria 1997).

[417] *Cordia bicolor* A.DC.

Prodr. [A. P. de Candolle] 9: 485 [1 Jan. 1845] (Candolle 1845). — Lithocardium bicolor (A.DC.) Kuntze, *Revis. Gen. Pl.* 2: 976 [5 Nov. 1891] (Kuntze 1891). — *Gerascanthus bicolor* (A.DC.) Borhidi, *Acta Bot. Hung.* 34 (3-4): 399 (Borhidi 1988).

Lithocardium lockhartii Kuntze, *Revis. Gen. Pl.* 2: 438 [5 Nov. 1891] (Kuntze 1891). — *Cordia lockhartii* Kuntze, *Revis. Gen. Pl.* 2: 438 [5 Nov. 1891] (Kuntze 1891), “*lockartii*”, *nom. nud. pro syn.*

*Cordia trichostyla* Pittier, *Contr. U.S. Natl. Herb.* 18 (6): 252 [22 Sep. 1917] (Pittier 1917).

*Cordia carnosa* Rusby, *Descr. S. Amer. Pl.* 104 [20 Dec. 1920] (Rusby 1920).

*Cordia belizensis* Lundell, *Amer. Midl. Naturalist* 29 (2): 488 [5 Apr. 1943] (Lundell 1943).

NOTE. — Although the “Borragineae” of the *Prodromus* were treated by A. P. de Candolle himself, note 1 on p. 466 states that his son Alphonse provided annotations and additions, among them several species whose names and descriptions are explicitly ascribed to “Alph. DC.”

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: M.-F. *Prévost 169*.

INVENTORY DATA (FG). — 40 trees in 21 plots; $F_{max} < 1\%$; $dbh_{inv} = 83$ cm.

[418] *Cordia dentata* Poir.

Encycl. [J. Lamarck et al.] 7: 48 [6 July 1806] (Poiret 1806). — *Carpiphea dentata* (Poir.) Raf., *Sylva Tellur.*: 39 (Rafinesque 1838).

*Cordia calyptrata* Bertero ex Spreng., *Syst. Veg. [Sprengel]* 1: 649 [“1825” publ. late 1824] (Sprengel 1824). — *Varronia calyptrata* (Bertero ex Spreng.) DC., *Prodr. [A. P. de Candolle]* 9: 469 [1 Jan. 1845] (Candolle 1845).

Varronia calyptrata var. hartweggii DC., *Prodr. [A. P. de Candolle]* 9: 469 [1 Jan. 1845] (Candolle 1845).

*Cordia tenuifolia* Bertol., *Misc. Bot. [Bertol.]* 21: 13 (Bertoloni 1860). — *Lithocardium tenuifolium* (Bertol.) Kuntze, *Revis. Gen. Pl.* 2: 977 [5 Nov. 1891] (Kuntze 1891).

*Cordia leptopoda* K.Krause, *Bot. Jahrb. Syst.* 37 (5): 628 [30 Oct. 1906] (Krause 1906).

*Cordia ovata* Brandegee, *Univ. Calif. Publ. Bot.* 10: 187 (Brandegee 1922).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: M.-F. *Prévost 4476*.

SIZE. — Up to 25 m tall (Nowicke 1969).

[419] *Cordia exaltata* Lam.

Tabl. Encycl. 1[2 (1)]: 422 [30 July 1792] (Lamarck 1792). — *Lithocardium exaltatum* (Lam.) Kuntze, *Revis. Gen. Pl.* 2: 977 [5 Nov. 1891] (Kuntze 1891). — *Gerascanthus exaltatus* (Lam.) Borhidi, *Acta Bot. Hung.* 34 (3-4): 399 (Borhidi 1988).

*Cordia melanoneura* Klotzsch ex M.R.Schomb., *Reis. Br.-Guiana [Ri. Schomburgk]* 3: 960 [“1848” publ. 7-10 Mar. 1849] (Schomburgk 1849), *nom. nud.*

*Cordia scabrida* Mart. ex Fresen., *Fl. Bras. [Martius]* 8 (1): 11 [28 Feb. 1857] (Fresenius 1857). — *Lithocardium scabridum* (Mart. ex Fresen.) Kuntze, *Revis. Gen. Pl.* 2: 439 [5 Nov. 1891] (Kuntze 1891). — *Gerascanthus scabridus* (Mart. ex Fresen.) Borhidi, *Acta Bot. Hung.* 34 (3-4): 402 (Borhidi 1988).

*Cordia exaltata* var. melanoneura I.M.Johnst., *J. Arnold Arbor.* 16 (1): 30 [25 Jan. 1935] (Johnston 1935).

VERNACULAR NAMES. — Wp: kuluwa'i, kuluwa'i sili • Wn: mojoi • Nt: busi kiikii • Cr: bwa-sip • Br: cauarú-caá, chapéu-de-sol.

HERBARIUM DATA (FG). — 23 collections at CAY. Sel. exs.: L. C. *Richard s.n.* (holo-, P[P00633485]; iso-, P[P00633484, P03892075, P03892077]).

INVENTORY DATA (FG). — 4 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 37$ cm.

[420] *Cordia fallax* I.M.Johnst.

*J. Arnold Arbor.* 16 (1): 10 [25 Jan. 1935] (Johnston 1935).

*Cordia guianensis* Klotzsch, *Reis. Br.-Guiana [Ri. Schomburgk]* 3: 960 [“1848” publ. 7-10 Mar. 1849] (Klotzsch 1849), *nom. illeg. hom., non* (Desv.) Roem. & Schult. (Roemer & Schultes 1819).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: G. Cremers 8385.

SIZE. — Up to 35 m tall (Gaviria 1997).

[421] *Cordia fulva* I.M.Johnst.

*J. Arnold Arbor.* 16 (1): 20 [25 Jan. 1935] (Johnston 1935).

NOTE. — Known only from the Guiana Shield.
HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *E.M. Mélinon* 137, 1877 (holo-, GH[GH00095576]; iso-, GH[GH00095577]).

INVENTORY DATA (FG). — 1 tree, dbh = 27.7 cm.

[422] *Cordia goeldiana* Huber

*Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 6: 89 (Huber 1910). — *Ger-ascanthus goeldianus* (Huber) M.Kuhlm. & Mattos, *Loefgrenia* 47: 2 (Kuhlmann & Mattos 1970).

VERNACULAR NAMES. — Wp: ka'i tuli (?) • Br: freijó, frei-jorge.

HERBARIUM DATA (FG). — 5 collections at CAY. Sel. exs.: *D. Larpin* 946.

INVENTORY DATA (FG). — 22 trees in 17 plots; Fmax < 1%; dbhinv = 85 cm.

[423] *Cordia laevifrons* I.M.Johnst.
(Fig. 20A)

J. Arnold Arbor. 16 (1): 16 [25 Jan. 1935] (Johnston 1935).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — 19 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino* 5647.

INVENTORY DATA (FG). — 7 trees in 5 plots; Fmax < 1%; dbhinv = 15.1 cm.

[424] *Cordia lomatoloba* I.M.Johnst.

J. Arnold Arbor. 18 (1): 12 [22 Jan. 1937] (Johnston 1937). — *Gerascanthus lomatolobus* (I.M.Johnst.) Borhidi, *Acta Bot. Hung.* 34 (3-4): 400 (Borhidi 1988).

HERBARIUM DATA (FG). — 16 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 4221.

INVENTORY DATA (FG). — 35 trees in 18 plots; Fmax < 1%; dbhinv = 83 cm.

[425] *Cordia naidophila* I.M.Johnst.

J. Arnold Arbor. 17 (1): 32 [24 Jan. 1936] (Johnston 1936). — *Gerascanthus naidophilus* (I.M.Johnst.) Borhidi, *Acta Bot. Hung.* 34 (3-4): 400 (Borhidi 1988).

HERBARIUM DATA (FG). — 5 collections at CAY. Sel. exs.: *M.-F. Prévost* 178.

SIZE. — Up to 15 m tall (Gaviria 1997).

[426] *Cordia nervosa* Lam.

Tabl. Encycl. 1[2 (1)]: 422 [30 July 1792] (Lamarck 1792). — *Lithocardium nervosum* (Lam.) Kuntze, *Revis. Gen. Pl.* 2: 977 [5 Nov. 1891] (Kuntze 1891).

*Cordia calophylla* Vahl, *Eclog. Amer.* 3: 5 (Vahl 1807). — *Gerascanthus calophyllus* (Vahl) Borhidi, *Acta Bot. Hung.* 34 (3-4): 399 (Borhidi 1988).

1988). — *Lithocardium calophyllum* (Vahl) Kuntze, *Revis. Gen. Pl.* 2: 976 [5 Nov. 1891] (Kuntze 1891).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — 18 collections at CAY. Sel. exs.: *L.C. Richard* s.n. (original material P[P03892659]).

INVENTORY DATA (FG). — 2 trees in 2 plots; Fmax < 1%; dbhinv = 18.8 cm.

[427] *Cordia nodosa* Lam.

Tabl. Encycl. 1[2 (1)]: 422 [30 July 1792] (Lamarck 1792). — *Cordia collococa* Aubl., *Hist. Pl. Guiane* 1: 219 [Jun.-Dec. 1775] (Aublet 1775), “*collo-cocus*” on plate, nom. illeg. hom., non *Cordia collococca* L. (Linnaeus 1759). — *Cordia hirsuta* Willd., *Sp. Pl.*, ed. 4 1 (2): 1076 [July 1798] (Willdenow 1798), nom. illeg. superfl. (based on the illegitimate *Cordia collococa*, for which the replacement name *C. nodosa* already existed). — *Cordia formicarum* Roem. & Schult., *Syst. Veg. [Roemer & Schultes]* 4: 800 [Mar.-June 1819] (Roemer & Schultes 1819). — *Cordia nodosa* var. hispidissima Fresen., *Fl. Bras. [Martius]* 8 (1): 17 [28 Feb. 1857] (Fresenius 1857). — *Cordia nodosa* var. glabrior Fresen., *Fl. Bras. [Martius]* 8 (1): 16 [28 Feb. 1857] (Fresenius 1857). — *Lithocardium nodosum* (Lam.) Kuntze, *Revis. Gen. Pl.* 2: 977 [5 Nov. 1891] (Kuntze 1891).

Firensia hirsuta Raf., *Sylva Tellur.:* 40 (Rafinesque 1838), nom. illeg. superfl. (based on *C. hirsuta* and *Cordia collococa*, for which the replacement name *C. nodosa* already existed).

*Cordia hispidissima* DC., *Prodr. [A. P. de Candolle]* 9: 475 [1 Jan. 1845] (Candolle 1845). — *Lithocardium hispidissimum* (DC.) Kuntze, *Revis. Gen. Pl.* 2: 977 [5 Nov. 1891] (Kuntze 1891).

*Cordia miranda* DC., *Prodr. [A. P. de Candolle]* 9: 475 [1 Jan. 1845] (Candolle 1845). — *Lithocardium mirandum* (DC.) Kuntze, *Revis. Gen. Pl.* 2: 977 [5 Nov. 1891] (Kuntze 1891).

*Cordia nodosa* var. *angustifolia* Fresen., *Fl. Bras. [Martius]* 8 (1): 17 [28 Feb. 1857] (Fresenius 1857).

*Cordia umbrosa* Spruce ex Rusby, *Bull. Torrey Bot. Club* 26 (3): 147 [18 Mar. 1899] (Rusby 1899).

*Cordia volubilis* Pittier, *J. Wash. Acad. Sci.* 19: 184 (Pittier 1929).

NOTE. — Var. *glabrior* is based on *C. formicarum*, and var. *hispidissima* is based on *C. collococa*.

VERNACULAR NAMES. — Pa: kasiu-van, wiwis-kasiuvan • Ka: aluko wonale, awale emulutano, awale mulu, awata epi • Wp: yawatai • Wn: jenëkapo, siktukahmit • Nt: wasiwasi uwii, wasiwasi wiwii • Cr: bwa-fronmi, lanmoussé-fronmi • Br: arua-felpudo, graó-de-galo, pau-de-formiga.

HERBARIUM DATA (FG). — 145 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material P-JJR, P00777834]).

INVENTORY DATA (FG). — 4 trees in 4 plots; Fmax < 1%; dbhinv = 11.5 cm.

[428] *Cordia panicularis* Rudge

*Pl. Guian. [Rudge]* 1 (4): 30 [Apr.-May 1806] (Rudge 1806). — *Lithocardium paniculare* (Rudge) Kuntze, *Revis. Gen. Pl.* 2: 976 [5 Nov. 1891] (Kuntze 1891). — *Gerascanthus panicularis* (Rudge)
Borhidi, *Acta Bot. Hung.* 34 (3-4): 401 (Borhidi 1988), nom. inval.
(basionym citation incomplete).

VERNACULAR NAMES. — Te: tapi’ilupabi hun.

HERBARIUM DATA (FG). — 5 collections at CAY. Sel. exs.: *J. Martin* 151 (holo-, K[K000583350]; iso-, BM[BM000953118], GH[GH00095780], P[P00634041, P00634042]).

INVENTORY DATA (FG). — 13 trees in 6 plots; Fmax = 1.1 %;
dbhinv = 19.4 cm.

[429] *Cordia sagotii* I.M.Johnst.
(Fig. 20B)

*J. Arnold Arbor.* 16 (1): 27 [25 Jan. 1935] (Johnston 1935). — *Gerascanthus sagotii* (I.M.Johnst.) Borhidi, *Acta Bot. Hung.* 34 (3-4): 401 (Borhidi 1988).

*Cordia coriacea* Sagot ex Benoist, *Arch. Bot.* 5 (Mém. 1): 257 [27 Mar. 1933] (Benoist 1933), *nom. illeg. hom.*, *non* Killip (1927).

*Cordia hirta* I.M.Johnst., *J. Arnold Arbor.* 16 (1): 29 [25 Jan. 1935] (Johnston 1935). — *Gerascanthus hirtus* (I.M.Johnst.) Borhidi, *Acta Bot. Hung.* 34 (3-4): 401 (Borhidi 1988).

NOTE. — A label affixed by J. Gaviria in 1987 on a K specimen, K000583352, wrongly claims that it is the holotype of *Cordia sagotii*. Actually, Johnston (1935: 28) wrote: “Acarouany [...], Dec. 1856, *Sagot* 447 (TYPE of *C. coriacea* Sagot and *C. Sagotii*, Paris)”. This unambiguously designates as holotype the specimen P00634059. This is furthermore confirmed by two labels in Johnston’s hand on P00634059. All other Sagot material bearing the same number 447, but dated “1855” or “1857” (including K000583352)), are probably syntypes.

VERNACULAR NAMES. — Pa: imavui-kamwi, wiwis-ahavukunõ, wiwis-priyo, wiwis-purubumna • Ka: alatuluka, kulebogo tikalayi, tagulewe itudyano • Wp: kuluwa’i • Nt: busi kiikii • Cr: bwa-parasòl • Fr: arbre parasol, bois parasol.

HERBARIUM DATA (FG). — 59 collections at CAY. Sel. exs.: *P.A. Sagot* 447, Dec. 1856 (holo-, P[P00634059]; iso-, BM[BM000906178], K[K000583351], P[P03860113]).

INVENTORY DATA (FG). — 170 trees in 93 plots; Fmax = 2.6 %;
dbhinv = 53.7 cm.

[430] *Cordia sericicalyx* A.DC.

*Prodr. [A. P. de Candolle]* 9: 485 [1 Jan. 1845] (Candolle 1845). — *Lithocardium sericicalyx* (A.DC.) Kuntze, *Revis. Gen. Pl.* 2: 977 [5 Nov. 1891] (Kuntze 1891). — *Gerascanthus sericicalyx* (A.DC.) Borhidi, *Acta Bot. Hung.* 34 (3-4): 402 (Borhidi 1988).

*Cordia sericicalyx* A.DC. var. *latifolia* Miq., *Stirp. Surinam. Select.*: 140 [1850 publ. Mar. 1851] (Miquel 1851).

*Cordia opaca* Rusby, *Descr. S. Amer. Pl.* 104 [20 Dec. 1920] (Rusby 1920). — *Gerascanthus opacus* (Rusby) Borhidi, *Acta Bot. Hung.* 34 (3-4): 401 (Borhidi 1988).

*Bourreria viridis* Rusby, *Descr. S. Amer. Pl.* 106 [20 Dec. 1920] (Rusby 1920). — *Gerascanthus viridis* (Rusby) Borhidi, *Acta Bot. Hung.* 34 (3-4): 403 (Borhidi 1988). — *Cordia viridis* (Rusby) I.M.Johnst., *J. Arnold Arbor.* 33 (1): 64 [Jan. 1952] (Johnston 1952).

*Cordia ierensis* Britton, *Bull. Torrey Bot. Club* 50 (1): 54 [Jan. 1923] (Britton 1923).

*Cordia coriacea* Killip, *J. Wash. Acad. Sci.* 17: 329 (Killip 1927).

VERNACULAR NAMES. — Wp: kuluwa’i, kuluwa’i sili.

HERBARIUM DATA (FG). — 1 collections at CAY. Sel. exs.: *P. Gre- nand* 237.

SIZE. — Guyana. *P. Mutchnick* 773, 15 m.

[431] *Cordia sprucei* Mez

*Bot. Jahrb. Syst.* 12: 549 [23 Dec. 1890] (Mez 1890). — *Lithocard- ium sprucei* (Mez) Kuntze, *Revis. Gen. Pl.* 2: 977 [5 Nov. 1891] (Kuntze 1891). — *Gerascanthus sprucei* (Mez) Borhidi, *Acta Bot. Hung.* 34 (3-4): 402 (Borhidi 1988).

VERNACULAR NAMES. — Wp: kuluwa’i.

HERBARIUM DATA (FG). — 24 collections at CAY. Sel. exs.: *M.-F. Prévost & C. Feuillet* 3968.

INVENTORY DATA (FG). — 8 trees in 7 plots; Fmax < 1 %; dbhinv = 25.1 cm.

[432] *Cordia tetrandra* Aubl.
(Fig. 20C)

*Hist. Pl. Guiane* 1: 222 [Jun.-Dec. 1775] (Aublet 1775). — *Lithocardium tetrandrum* (Aubl.) Kuntze, *Revis. Gen. Pl.* 2: 977 [5 Nov. 1891] (Kuntze 1891). — *Borellia aspera* Raf., *Sylva Téllur.:* 41 (Rafinesque 1838), *nom. illeg. superfl.* (based on *Cordia tetran- dra*). — *Gerascanthus tetrandrus* (Aubl.) Borhidi, *Acta Bot. Hung.* 34 (3-4): 402 (Borhidi 1988).

*Cordia cordifolia* Kunth, *Nova genera et species plantarum [H.B.K.]* 3: 70 [Late Sep. 1818] (Kunth 1818). — *Lithocardium cordifolium* (Kunth) Kuntze, *Revis. Gen. Pl.* 2: 976 [5 Nov. 1891] (Kuntze 1891). — *Gerascanthus cordifolius* (Kunth) Borhidi, *Acta Bot. Hung.* 34 (3-4): 399 (Borhidi 1988).

*Cordia muneco* Kunth, *Nova genera et species plantarum [H.B.K.]* 7: 207 [21 Feb. 1825] (Kunth 1825). — *Lithocardium muneco* (Kunth) Kuntze, *Revis. Gen. Pl.* 2: 977 [5 Nov. 1891] (Kuntze 1891). — *Gerascanthus muneco* (Kunth) Borhidi, *Acta Bot. Hung.* 34 (3-4): 401 (Borhidi 1988).

*Cordia umbraculifera* DC., *Prodr. [A. P. de Candolle]* 9: 484 [1 Jan. 1845] (Candolle 1845).

VERNACULAR NAMES. — Ka: alatuluka • Wp: kuluwa’i u • Br: chapéu-de-sol.

HERBARIUM DATA (FG). — 26 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JJR[P00777832] designated by Lanjouw & Uittien [1940: 150]).

SIZE. — Up to 18 m tall (Gaviria 1997).

[433] *Cordia toqueve* Aubl.

*Hist. Pl. Guiane* 1: 228 [Jun.-Dec. 1775] (Aublet 1775). — *Lithocardium toqueve* (Aubl.) Kuntze, *Revis. Gen. Pl.* 2: 977 [5 Nov. 1923].
<figure><img src="image_31.png" /><figcaption>FIG. 20. — Cordiaceae: **A**, *Cordia laevifrons* I.M.Johnst. (D. Sabatier & J.-F. Molino 5647); **B**, *Cordia sagotii* I.M.Johnst. (M.-F. Prévost & D. Sabatier 4804); **C**, *Cordia tetrandra* Aubl. (M.-F. Prévost 1500). Dichapetalaceae: **D**, *Tapura capitulifera* Baill. A, D © D. Sabatier/IRD; B, C, © M.-F. Prévost/IRD.</figcaption></figure>

1891] (Kuntze 1891). — *Gerascanthus toqueve* (Aubl.) Borhidi, *Acta Bot. Hung.* 34 (3-4): 402 (Borhidi 1988).

*Cordia heterophylla* Poir., *Dict. Sci. Nat., ed. 2* [F. Cuvier], 10: 409 (Poiret 1818). — *Lithocardium heterophyllum* (Poir.) Kuntze, *Revis. Gen. Pl.* 2: 977 [5 Nov. 1891] (Kuntze 1891). — *Gerascanthus heterophyllus* (Poir.) Borhidi, *Acta Bot. Hung.* 34 (3-4): 400 (Borhidi 1988).

*Toquera tomentosa* Raf., *Sylva Tellur.*: 40 (Rafinesque 1838).

*Cordia hebecarpa* DC., *Prodr.* [A. P. de Candolle] 9: 488 [1 Jan. 1845] (Candolle 1845). — *Lithocardium hebecarpum* (DC.) Kuntze, *Revis. Gen. Pl.* 2: 977 [5 Nov. 1891] (Kuntze 1891).

*Cordia pubescens* Roem. & Schult., *Syst. Veg. [Roemer & Schultes]* 4: 800 (Mar.-June 1819) (Roemer & Schultes 1819). — *Gerascanthus pubescens* (Roem. & Schult.) Borhidi, *Acta Bot. Hung.* 34 (3-4): 401 (Borhidi 1988). — *Lithocardium pubescens* (Roem. & Schult.) Kuntze, *Revis. Gen. Pl.* 2: 977 [5 Nov. 1891] (Kuntze 1891).

*Cordia expansa* Lingelsh., *Repert. Spec. Nov. Regni Veg.* 7: 244 (Lingelsheim 1909).

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JJR[P00777830] designated by Lanjouw & Uittien [1940: 150]).

INVENTORY DATA (FG). — 1 tree, dbh = 15.5 cm.

Family DICHAPETALACEAE Baill.
Genus *Tapura* Aubl.

[434] *Tapura amazonica* Poepp.

*Nova genera ac species plantarum* [Poeppig & Endlicher] 3: 41 [8-11 Mar. 1843] (Poeppig 1843).

*Tapura ciliata* Gardner, *Icon. Pl.* 5 [n.s., 1]: t. 466 [Jan.-Oct. 1842] (Gardner 1842). — *Tapura amazonica* var. *ciliata* (Gardner) Baill., *Fl. Bras.* [*Martius*] 12 (1): 375 [1 Apr. 1886] (Baillon 1886).

*Tapura amazonica* var. *cuspidata* Baill., *Fl. Bras.* [*Martius*] 12 (1): 375 [1 Apr. 1886] (Baillon 1886).

*Tapura amazonica* var. *dasyphylla* Baill., *Fl. Bras.* [*Martius*] 12 (1): 375 [1 Apr. 1886] (Baillon 1886).

*Tapura amazonica* var. *sublanceolata* Baill., *Fl. Bras.* [*Martius*] 12 (1): 375 [1 Apr. 1886] (Baillon 1886).

VERNACULAR NAMES. — Pa: á-kasiuminio.

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: *E.M. Mélinon 170* (P[P04764399], syntype of *Tapura amazonica* var. *dasyphylla*.

INVENTORY DATA (FG). — 20 trees in 15 plots; $F_{max} < 1\%$; $dbh_{inv} = 27.4$ cm.
[435] *Tapura capitulifera* Baill.
(Fig. 20D)

*Adansonia* [Baillon] 11: 112 (Baillon 1873). — *Chailletia capitulifera* Spruce ex Baill., *Adansonia* [Baillon] 11: 112 (Baillon 1873), *nom. nud. pro syn.*

HERBARIUM DATA (FG). — 47 collections at CAY. Sel. exs.: *D. Sabatier* 5792.

INVENTORY DATA (FG). — 455 trees in 76 plots; Fmax = 5.5 %; dbhinv = 95.5 cm.

[436] *Tapura guianensis* Aubl.

*Hist. Pl. Guiane* 1: 126 [Jun.-Dec. 1775] (Aublet 1775). — *Rohria tapura* J.F.Gmel., *Syst. Nat.*, ed. 13[bis], 2 (1): 93 [late Sep.-Nov. 1791] (Gmelin 1791), *nom. illeg. superfl.* (based on *Tapura guianensis*). — *Rohria petioliflora* Willd., *Sp. Pl.*, ed. 4 1 (1): 186 [June 1797] (Willdenow 1797), *nom. illeg. superfl.* (based on *Tapura guianensis*).

*Rohria schreberi* J.F.Gmel., *Syst. Nat.*, ed. 13[bis], 2 (1): 93 [late Sep.-Nov. 1791] (Gmelin 1791).

*Chailletia sessiliflora* DC., *Nouv. Bull. Sci. Soc. Philom. Paris* 2 (40): 205 [Jan. 1811] (Candolle 1811).

*Tapura cucullata* Benth., *Hooker's J. Bot. Kew Gard. Misc.* 5: 292 (Bentham 1853).

*Tapura negrensis* Suess., *Repert. Spec. Nov. Regni Veg.* 51: 199 (Sues-senguth 1942).

VERNACULAR NAMES. — Ka: wasakau • Wp: awalapuna sili, ka'a sili, paa lo'o, wila towakape sī.

HERBARIUM DATA (FG). — 144 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JJR[P00777925] designated by Lanjouw & Uittien [1940: 156]).

INVENTORY DATA (FG). — 117 trees in 31 plots; Fmax = 2.5 %; dbhinv = 22.9 cm.

[437] *Tapura singularis* Ducke

*Trop. Woods* 90: 21 (Ducke 1947).

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: *D. Sabatier* 2530.

INVENTORY DATA (FG). — 7 trees in 6 plots; Fmax < 1 %; dbhinv = 38.2 cm.

Family DILLENIACEAE Salisb.
Genus *Curatella* Loefl.

[438] *Curatella americana* L.

*Syst. Nat.*, ed. 10, 2: 1079 [7 June 1759] (Linnaeus 1759), “*americ*.”

*Curatella cambaiba* A.St.-Hil., *Fl. Bras. Merid. [A. St.-Hil.]* (quarto ed.) 1 (1): 22 [23 Feb. 1824] (Saint-Hilaire 1824), “*Çambaiba*.”

*Curatella grisebachiana* Eichler, *Fl. Bras. [Martius]* 13 (1): 69 [15 Jan. 1863] (Eichler 1863).

*Curatella americana* var. *pentagyna* Donn.Sm., *Bot. Gaz.* 46 (2): 109 [22 Aug. 1908] (Donnell Smith 1908).

NOTE. — Restricted to savannas.

VERNACULAR NAMES. — Pa: sawu-anen-kamwi • Ka: kulata • Cr: féy-sab • Br: caimbé, lixeira.

HERBARIUM DATA (FG). — 19 collections at CAY. Sel. exs.: *M.-F. Prévost* 1636.

SIZE. — Up to 16 m tall (Aymard C. 1998).

Family EBENACEAE Gürke
Genus *Diospyros* L.

[439] *Diospyros capreifolia* Mart. ex Hiern

*Trans. Cambridge Philos. Soc.* 12 (1): 254 (Hiern 1873), “*capreafolia*”.

*Maba melinonii* Hiern, *Trans. Cambridge Philos. Soc.* 12 (1): 143 (Hiern 1873), “*Melinonii*”. — *Diospyros melinonii* (Hiern) A.C.Sm., *Bull. Torrey Bot. Club* 60 (6): 390 [1 June 1933] (Smith 1933), “*Melinonii*”.

VERNACULAR NAMES. — Pa: á-seiminio-priyo, á-seiminio-priyu, miret • Nt: bita tiki • Cr: bwa-krayon • Br: altinha.

HERBARIUM DATA (FG). — 34 collections at CAY. Sel. exs.: *E.M. Mélinon s.n.* (original material of *Maba melinonii*: F[V0063306F, V0063307F, V0063308F]).

INVENTORY DATA (FG). — 31 trees in 25 plots; Fmax < 1 %; dbhinv = 28.6 cm.

[440] *Diospyros carbonaria* Benoist

*Arch. Bot.* 5 (Mém. 1): 249 [27 Mar. 1933] (Benoist 1933).

*Diospyros duckei* Sandwith, *Kew Bull.* 4 (4): 488 [“1949” publ. 16 Feb. 1950] (Sandwith 1950).

*Diospyros capimnensis* Pires & Cavalcante, *Bol. Mus. Paraense Emílio Goeldi, N.S.*, *Bot.* 9: 1 (Pires & Cavalcante 1960).

VERNACULAR NAMES. — Pa: miret • Nt: bita tiki • Cr: bwa-charbon, bwa-krayon.

HERBARIUM DATA (FG). — 29 collections at CAY. Sel. exs.: *R. Benoist* 448 (original material P[P00721344, P00721346]).

INVENTORY DATA (FG). — 137 trees in 73 plots; Fmax = 2 %; dbhinv = 35.3 cm.

[441] *Diospyros cavalcantei* Sothers

*Kew Bull.* 55 (2): 471 (Sothers 2000).

VERNACULAR NAMES. — Pa: miret • Cr: bwa-krayon.

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: *M.-F. Prévost et al.* 4671.

INVENTORY DATA (FG). — 13 trees in 10 plots; Fmax < 1 %; dbhinv = 76.1 cm.
[442] *Diospyros cayennensis* A.DC.

Prodr. [A. P. de Candolle] 8: 224 [mid Mar. 1844] (Candolle 1844).

*Diospyros ierensis* Britton, *Bull. Torrey Bot. Club* 48 (12): 336 ["1921" publ. 1922] (Britton 1922).

VERNACULAR NAMES. — Ka: palala, simyali epi, talala.

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *J. Martin s.n.* (lecto-, G-DC[G00142182] designated by Wallnöfer [2001: 888]; isolecto-, P[P00721345, P00721347]).

INVENTORY DATA (FG). — 30 trees in 24 plots; $F_{max} < 1\%$; $dbh_{inv} = 48.7$ cm.

[443] *Diospyros dichroa* Sandwith
(Fig. 21A)

*Bull. Misc. Inform. Kew* 1931 (10): 482 [31 Dec. 1931] (Sandwith 1931).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Pa: á-kabebetye, miret, pukuu-kamwi • Ka: yukutuna • Nt: bita tiki • Cr: bwa-krayon, bwa-zoranj.

HERBARIUM DATA (FG). — 28 collections at CAY. Sel. exs.: *D. Sabatier et al. 4457*.

INVENTORY DATA (FG). — 35 trees in 9 plots; $F_{max} = 3.5\%$; $dbh_{inv} = 45$ cm.

[444] *Diospyros guianensis* (Aubl.) Gürke
(Fig. 21B)

*Nat. Pflanzenfam. [Engler & Pranti] 4* (1): 163 (Gürke 1891). — *Paralea guianensis* Aubl., *Hist. Pl. Guiane* 1: 576 [Jun.-Dec. 1775] (Aublet 1775).

VERNACULAR NAMES. — Pa: miret • Ka: palala, simyali epi, talala • Wp: miku lapi'a u • Wn: meikolo muhunu, meikolo muhupu, mekolo nohunu • Nt: baaka tiki • Cr: bwa-krayon.

HERBARIUM DATA (FG). — 34 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material FI[FI012223]).

INVENTORY DATA (FG). — 5 trees in 5 plots; $F_{max} < 1\%$; $dbh_{inv} = 39.6$ cm.

[445] *Diospyros martinii* Benoist

*Arch. Bot.* 5 (Mém. 1): 248 [27 Mar. 1933] (Benoist 1933), "Martini".

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 5705*.

INVENTORY DATA (FG). — 6 trees in 4 plots; $F_{max} < 1\%$; $dbh_{inv} = 48.5$ cm.

[446] *Diospyros sericea* A.DC.

Prodr. [A. P. de Candolle] 8: 236 [mid Mar. 1844] (Candolle 1844). — *Maba sericea* (A.DC.) Hiern, *Trans. Cambridge Philos. Soc.* 12 (1): 140 (Hiern 1873).

*Diospyros vestita* Benoist, *Arch. Bot.* 5 (Mém. 1): 248 [27 Mar. 1933] (Benoist 1933), *in clavi*; non Bakh. [May 1933] (Bakhuizen 1933).

*Diospyros praetermissa* Sandwith, *Kew Bull.* 4 (4): 483 ["1949" publ. 16 Feb. 1950] (Sandwith 1950).

HERBARIUM DATA (FG). — 13 collections at CAY. Sel. exs.: *R. Benoist 520* (original material of *Diospyros vestita*: P[P00721401]).

INVENTORY DATA (FG). — 10 trees in 10 plots; $F_{max} < 1\%$; $dbh_{inv} = 51.9$ cm.

[447] *Diospyros tetrandra* Hiern

*Trans. Cambridge Philos. Soc.* 12 (1): 210 (Hiern 1873), *non Span.* (1841, *nom. nud.*).

VERNACULAR NAMES. — Ka: palala, simyali epi, talala • Wp: miku lapi'a sówī • Nt: baaka tiki.

HERBARIUM DATA (FG). — 21 collections at CAY. Sel. exs.: *J. Martin s.n.* (lecto-, K[K000644356] designated by Wallnöfer [2020: 231]).

INVENTORY DATA (FG). — 4 trees in 3 plots; $F_{max} < 1\%$; $dbh_{inv} = 23$ cm.

[448] *Diospyros* sp. A

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *D. Sabatier et al. 5803*.

INVENTORY DATA (FG). — 2 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 47.7$ cm.

Family ELAEOCARPACEAE Juss.

NOTE

The presence of *Sloanea usurpatrix* Sprague & Riley in French Guiana seems plausible given its known trans-Amazonian distribution. According to Pennington & Wise (2017), it is attested by a single sterile collection (Nouragues Nature Reserve, 40 km SW of Regina, *Mori et al. 26678*, CAY, K). However, the leaf morphology of this specimen clearly differs from that of *S. usurpatrix*, while fitting perfectly with *S. synandra*.

Genus *Sloanea* L.

[449] *Sloanea acutiflora* Uittien
(Fig. 22A)

*Recueil Trav. Bot. Néerl.* 22: 357 ["1925" publ. Jan. 1926] (Uitten 1926).
NOTES. — Known only from the Guiana Shield. Pennington & Wise (2017: 77) placed *S. acutiflora* in synonymy under *S. latifolia* (Rich.) K.Schum., arguing that they have the same floral morphology, and that there are intermediate states throughout the range of the species (i.e. most of Amazonia) in fruit spine development and leaf shape; the typical form of *S. latifolia* has long petiole and large broad leaves with rounded bases while other “forms”, typified by *S. acutiflora* in the Guiana Shield and *S. inermis* Ducke in Southern and Western Amazonia, have smaller leaves with acute or cuneate base and short petioles. Our observations contradict these statements. In French Guiana, all specimens that correspond to the Pennington and Wise concept of *S. latifolia* can easily be separated into two very distinct groups. In the first group, which includes the original material (see Notes under *S. latifolia*), trees have both buttresses and stilt roots, petioles of very variable length on the same branchlet, the longest 3-15 cm, and large leaves drying yellowish brown, of variable size but frequently 20 cm long, with rounded to obtuse, rarely cuneate base and a densely reticulate venation. Inflorescences are large, well-developed many-flowered corymbs, flower buds are slender, the base gradually attenuated on the pedicel, and covered with a dense yellow tomentum. Fruit is up to 4.5 cm long, brown, smooth, and seeds are surrounded by a white to pink aril (Fig. 22D). Trees of the second group have buttresses but no stilt roots. Petioles are shorter (usually < 1 cm, rarely 1-1.5 cm). Leaves dry to a light green, and are much smaller (rarely up to 15 cm long), with an acute-attenuated base, never round or obtuse, and a much looser reticulum. Inflorescences are short, more or less cymose, bearing 1-3 (-5) flowers. Buds are subglobose to ovoid, the base abruptly reduced at the junction with the pedicel, the whitish, much less dense pilosity making them appear greyish rather than yellow. Fruit is red or pink, less than 2.5 cm long, sometimes covered with sparse, short conic spines, and the aril is orange (Fig. 22A). This last group corresponds with the type of *S. acutiflora*. Pennington & Wise (2017: 77) designated the lectotype of *S. acutiflora* as "[...] BW 6298 (lectotype, U, chosen here; isolectotypes, K, L)". At U there are three sheets of C.J. Zaandam-Boschwezen Suriname 6298; these sheets (barcodes U0001682, U0001683, U0001684) are all kept together, and according to Art. 8.3 of the ICN (Turland *et al.* 2018), they are here treated as a single specimen with multiple preparations. Isolectotypes are at K(not seen), L(L0013589), RB(RB00538334) and US (00098537).

Although the distributions of these two groups overlap throughout much of French Guiana (west, center and south), no intermediate states have yet been found between the character syndromes defining these two groups. It is also interesting to note that, except in the extreme south of the territory (Tumuc-Humac Mounts), the "typical" *S. latifolia* form has never been found within 50 km of the border with Suriname. In view of the above, the reinstatement of *S. acutiflora* appears necessary.

VERNACULAR NAMES. — Pa: waaduk-seine.

HERBARIUM DATA (FG). — 27 collections at CAY. Sel. exs.: *D. Sabatier* 1234.

INVENTORY DATA (FG). — 39 trees in 31 plots; Fmax < 1%; dbhinv = 65 cm.

[450] *Sloanea brachytepala* Ducke

Trop. Woods 76: 22 (Ducke 1943).

*Sloanea calva* Pal.-Duque & Fern.Alonso, *Revista Acad. Colomb. Ci. Exact.* 29 (111): 180 [June 2005] (Palacios-Duque & Fernández Alonso 2005).

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *D. Sabatier* & M.-F. Prévost 3568.

INVENTORY DATA (FG). — 16 trees in 15 plots; Fmax < 1%; dbhinv = 54.4 cm.

[451] *Sloanea brevipes* Benth.

J. Proc. Linn. Soc., Bot. 5 (Suppl. 2): 68 [May 1861] (Bentham 1861).

VERNACULAR NAMES. — Pa: ihap-kamwi-duwê • Ka: kusewelan • Wn: inuuman • Nt: busi kusuwe • Cr: chatengn, roukou-gran-bwa, roukou-sovaj • Fr: roucou grand bois • Br: urucurana.

HERBARIUM DATA (FG). — 26 collections at CAY. Sel. exs.: *D. Sabatier* 844.

INVENTORY DATA (FG). — 104 trees in 55 plots; Fmax < 1%; dbhinv = 80 cm.

[452] *Sloanea conferta* T.D.Penn.

Opusc. Neotrop. 2: 6 [13 July 2016] (Pennington 2016).

NOTE. — Known only from French Guiana.

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *D. Sabatier* 4897 (holo-, K, not seen; iso-, CAY[CAY065610], US[00943331]).

INVENTORY DATA (FG). — 8 trees in 8 plots; Fmax < 1%; dbhinv = 27.1 cm.

[453] *Sloanea echinocarpa* Uittien

Recueil Trav. Bot. Néerl. 22: 354 ["1925" publ. Jan. 1926] (Uittien 1926).

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *D. Sabatier* & M.-F. Prévost 2268.

SIZE. — Up to 65 cm dbh (Pennington & Wise 2017).

[454] *Sloanea eichleri* K.Schum.

Fl. Bras. [Martius] 12 (3): 183 [1 Nov. 1886] (Schumann 1886).

*Sloanea sinemariensis* var. *melinonii* Sagot, *Ann. Sci. Nat.*, Bot. sér. 6, 11: 156 (Sagot 1881), "Melinonis".

NOTE. — The epithet "*melinonis*", which honours the French botanist E. Mélinon, is to be automatically corrected to "*melinonii*" (Turland *et al.* 2018: Art. 60.8).

VERNACULAR NAMES. — Pa: ihap-kamwi, ihap-kamwi-seine • Wp: uluku panali sili, ulukupalani sili • Cr: roukou-gran-bwa, roukou-sovaj • Br: urucurana.

HERBARIUM DATA (FG). — 25 collections at CAY. Sel. exs.: *J.-F. Molino* & *D. Sabatier* 2693.

INVENTORY DATA (FG). — 36 trees in 28 plots; Fmax = 1%; dbhinv = 32 cm.
<figure><img src="image_32.png" /><figcaption>FIG. 21. — Ebenaceae: **A**, *Diospyros dichroa* Sandwith (D. Sabatier & M.-F. Prévost 5246); **B**, *Diospyros guianensis* (Aubl.) Gürke. Erythroxylaceae: **C**, *Erythroxylum fimbriatum* Peyr. (D. Sabatier & J.-F. Molino 5229); **D**, *Erythroxylum suberosum* A.St.-Hil. et al. (D. Sabatier & J.-F. Molino 5198). © D. Sabatier/IRD.</figcaption></figure>

[455] *Sloanea erythrocarpa* T.D.Penn.
(Fig. 22B)

*Opusc. Neotrop.* 2: 10 [13 July 2016] (Pennington 2016).

NOTE. — Known only from French Guiana.

HERBARIUM DATA (FG). — 13 collections at CAY. Sel. exs.: *D. Sabatier 3540*.

INVENTORY DATA (FG). — 21 trees in 21 plots; $F_{max} < 1\%$; $dbh_{inv} = 45$ cm.

[456] *Sloanea floribunda* Spruce ex Benth.

J. Proc. Linn. Soc., Bot. 5 (Suppl. 2): 66 [May 1861] (Bentham 1861).

*Sloanea paniculata* Spruce ex Benth., J. Proc. Linn. Soc., Bot. 5 (Suppl. 2): 66 [May 1861] (Bentham 1861).

*Sloanea longipes* Ducke, Arch. Inst. Biol. Veg. 2 (2): 166 [Dec. 1935] (Ducke 1935).

*Sloanea maroana* Steyerm., Pittieria 7: 14 (Steyermark 1978).

*Sloanea sipapoana* Steyerm., Ann. Missouri Bot. Gard. 75 (4): 1580 ["1988" publ. 8 Mar. 1989] (Steyermark 1989).

HERBARIUM DATA (FG). — No collections at CAY. Sel. exs.: S.A. Mori 26549 (K, fide Pennington & Wise [2017], not seen).

SIZE. — Up to 120 cm dbh (Pennington & Wise 2017).

[457] *Sloanea garckeana* K.Schum.

Fl. Bras. [Martius] 12 (3): 177 [1 Nov. 1886] (Schumann 1886).

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: M.-F. Prévost 1893.

INVENTORY DATA (FG). — 15 trees in 10 plots; $F_{max} < 1\%$; $dbh_{inv} = 73.5$ cm.
[458] *Sloanea grandiflora* Sm.

Cycl. [Rees] 33: (Sloanea no. 2) [17 May 1816] (Smith 1816).

*Sloanea amplifrons* I.M.Johnst., *Contr. Gray Herb.*, n.s., 75: 29 (Johnston 1925).

*Sloanea linderi* I.M.Johnst. ex Kribs, *Trop. Woods* 13: 16 (Kribs 1928), *nom. nud*.

VERNACULAR NAMES. — Pa: ihap-kamwi • Ka: kusewelan, polo ali • Te: tokolo-pipinatsi • Wp: tawisowiso, uluku panali, ulukupalani • Wn: inuuman • Nt: busi kusuwe • Cr: chatengn, roukou-gran-bwa, roukou-sovaj • Fr: roucou grand bois • Br: ururucana.

HERBARIUM DATA (FG). — 55 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 4341*.

INVENTORY DATA (FG). — 28 trees in 16 plots; Fmax < 1%; dbhinv = 73 cm.

[459] *Sloanea granulosa* Ducke

Bol. Tecn. Inst. Agron. N. 19: 13 (Ducke 1950).

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 4126*.

INVENTORY DATA (FG). — 13 trees in 12 plots; Fmax < 1%; dbhinv = 100 cm.

[460] *Sloanea guianensis* (Aubl.) Benth. subsp. *guianensis*

J. Proc. Linn. Soc., Bot. 5 (Suppl. 2): 69 [May 1861] (Bentham 1861). — *Ablania guianensis* Aubl., Hist. Pl. Guiane 1: 585 [Jun.-Dec. 1775] (Aublet 1775). — *Trichocarpus guianensis* (Aubl.) Raeusch., Nomencl. Bot. [Raeusch.], ed. 3, 158 (Raeuschel 1797), *nom. illeg. superfl.* (genus name superfluous, based on the type of *Ablania* Aubl.). — *Trichocarpus laurifolius* Willd., Sp. Pl., ed. 4 2 (2): 1224 [Dec. 1799] (Willldenow 1799), "*laurifolia*", *nom. illeg. superfl.* (genus name superfluous, based on the type of *Ablania* Aubl.)

*Ablania laurifolia* Pers., Syn. Pl. [Persoon] 2 (1): 81 [Nov. 1806] (Persoon 1806).

*Dasynema pubescens* Schott, Syst. Veg. [Sprengel] 4 (2, Cur. Post.): 408 [Jan.-June 1827] (Schott 1827). — *Sloanea pubescens* (Schott) Radlk., Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 12: 329 (Radlkofer 1882), *nom. illeg. hom.*, *non* Benth. (Bentham 1861).

*Sloanea alnifolia* Mart., *Flora* 20 (2, Beibl.): 94 (Martius 1837). — *Dasynema alnifolium* (Mart.) Walp., Repert. Bot. Syst. [Walpers] 1 (2): 352 [18-20 Sep. 1842] (Walpers 1842). — *Dasynema cuneifolium* (Mart.) Walp., Repert. Bot. Syst. [Walpers] 1 (2): 352 [18-20 Sep. 1842] (Walpers 1842).

*Sloanea cuneifolia* Mart., *Flora* 20 (2, Beibl.): 94 (Martius 1837).

*Dasynema pubescens* Turcz., Bull. Soc. Imp. Naturalistes Moscou 31 (1): 224 (Turczaninow 1858), *nom. illeg. hom.*, *non* Schott (1827) nec Poepp. (Poeppig 1845).

*Sloanea microcarpa* Planch. ex Benth., *J. Proc. Linn. Soc., Bot.* 5 (Suppl. 2): 69 [May 1861] (Bentham 1861). — *Sloanea guianensis* var. *microcarpa* (Planch. ex Benth.) K.Schum., *Fl. Bras. [Martius]* 12 (3): 191 [1 Nov. 1886] (Schumann 1886).

*Sloanea regelii* K.Schum., *Fl. Bras. [Martius]* 12 (3): 189 [1 Nov. 1886] (Schumann 1886).

*Sloanea alnifolia* var. *lancea* K.Schum., *Fl. Bras. [Martius]* 12 (3): 194 [1 Nov. 1886] (Schumann 1886).

*Sloanea alnifolia* var. *ovalis* K.Schum., *Fl. Bras. [Martius]* 12 (3): 194 [1 Nov. 1886] (Schumann 1886).

*Sloanea ptariana* Steyerm., *Fieldiana*, Bot. 28 (2): 360 (Steyermark 1952).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: ihap-kamwi • Ka: kusewelan • Wp: uluku panali sili, ulukupalani sili • Wn: inuuman • Nt: busi kusuwe • Cr: chatengn, roukou-gran-bwa, roukou-sovaj • Fr: roucou grand bois • Br: ururucana.

HERBARIUM DATA (FG). — 13 collections at CAY. Sel. exs.: *D. Sabatier 1237*.

INVENTORY DATA (FG). — 42 trees in 28 plots; Fmax = 1.2%; dbhinv = 60 cm.

[461] *Sloanea guianensis* subsp. *purdiei*
(Griseb. ex R.O.Williams & Cheesman) T.D.Penn.

Opusc. Neotrop. 2: 13 [13 July 2016] (Pennington 2016), "purdiaei". — *Sloanea purdiei* Griseb. ex Benth. & Hook.f., *Gen. Pl. [Bentham & Hooker f.]* 2 (2): 653 [May 1876] (Bentham & Hooker 1876), "Purdiaei", in obs., *nom. nud.* — *Sloanea purdiei* Griseb. ex R.O.Williams & Cheesman, *Fl. Trinidad & Tobago* 1 (2): 109 [Nov. 1929] (Williams & Cheesman 1929), "purdiaei".

*Sloanea breviseta* Steyerm., *Fieldiana*, Bot. 28 (2): 357 (Steyermark 1952).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *J.-J. de Granville B-5392*.

SIZE. — Up to 25 m tall (Pennington & Wise 2017).

[462] *Sloanea guianensis* subsp. *stipitata*
(Spruce ex Benth.) T.D.Penn.

Opusc. Neotrop. 2: 6 [13 July 2016] (Pennington 2016). — *Sloanea stipitata* Spruce ex Benth., *J. Proc. Linn. Soc., Bot.* 5 (Suppl. 2): 68 [May 1861] (Bentham 1861).

*Sloanea maximowicziana* K.Schum., *Fl. Bras. [Martius]* 12 (3): 192 [1 Nov. 1886] (Schumann 1886).

HERBARIUM DATA (FG). — A single collection, S.A. Mori & Y. Veyret 8972.

SIZE. — Up to 80 cm dhb (Pennington & Wise 2017).

[463] *Sloanea latifolia* (Rich.) K.Schum.
(Fig. 22D)

*Fl. Bras. [Martius]* 12 (3): 173 [1 Nov. 1886] (Schumann 1886). — *Blondea latifolia* Rich., *Actes Soc. Hist. Nat. Paris* 1: 110 [Oct. 1792] (Richard 1792).
*Sloanea corymbiflora* DC., *Prodr.* [A. P. de Candolle] 1: 516 [mid Jan. 1824] (Candolle 1824).

*Sloanea inermis* Ducke, *Arch. Inst. Biol. Veg.* 4 (1): 48 [June 1938] (Ducke 1938).

*Sloanea gentryi* Pal.-Duque & C.M.Baeza, *Novon* 23 (1): 70 [8 Apr. 2014] (Palacios-Duque & Baeza 2014).

**NOTE.** — Richard (1792: 110) along with the description of *Blondea latifolia* did not cite any specimen. Pennington (2016: 15) and Pennington and Wise (2017: 76) cited the type of this name as “French Guiana (Cayenne), fl., *LeBlond 219* (holotype, P; isotype, G)”. These citations cannot be treated as an inadvertent lectotypification because, being published after 2001, they should have been accompanied by “here designated” or a similar expression. In addition, there are three specimens of *Blondea latifolia* at P that were collected by Leblond, with the locality “*ad amnem Kourou*” (Kourou River), each with a different label, no collector number, and with barcodes P02440486, P02440487, and P02440488. Another specimen of *Blondea latifolia* collected by Leblond, with the collector number 209 (not “219”) is at G (G00104361). Therefore, a lectotype is hereafter designated.

**VERNACULAR NAMES.** — Pa: ihap-kamwi-purubumna, kunau, sitrukamwi • Ka: tokuhsa • Wp: tapi'ilo wapa'a, wila kalayulu, wila kayulu • Nt: busi kusuwe • Cr: chatengn, roukou-gran-bwa, roukou-sovaj • Fr: roucou grand bois • Br: urucurana.

**HERBARIUM DATA (FG).** — 30 collections at CAY. Sel. exs.: *J.B. Leblond 209* (lecto-, P[P02440487], here designated; isolecto-, G[G00104361], P[P02440486, P02440488]).

**INVENTORY DATA (FG).** — 25 trees in 20 plots; $F_{max} < 1\%$; $dbh_{inv} = 54$ cm.

[464] *Sloanea morii* T.D.Penn.

*Opusc. Neotrop.* 2: 17 [13 July 2016] (Pennington 2016).

**NOTE.** — Known only from French Guiana.

**HERBARIUM DATA (FG).** — 4 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 2263*.

**INVENTORY DATA (FG).** — 1 tree, $dbh = 11.1$ cm.

[465] *Sloanea nitida* G.Don

*Gen. Hist.* 1: 555 [early Aug. 1831] (Don 1831).

*Sloanea bracteosa* Ducke, *Arch. Inst. Biol. Veg.* 2 (2): 168 [Dec. 1935] (Ducke 1935).

**HERBARIUM DATA (FG).** — 4 collections at CAY. Sel. exs.: *J. Martin s.n.* (holo-, BM[BM000907458, BM000907459]; iso-, K[K000380070]).

**SIZE.** — Up to 60 cm $dbh$ (Pennington & Wise 2017).

[466] *Sloanea obtusifolia* (Moric.) K.Schum.

*Fl. Bras. [Martius]* 12 (3): 181 [1 Nov. 1886] (Schumann 1886). — *Adenobasium obtusifolium* Moric., *Pl. Nouv. Amér.* 83 (Moricand 1840). — *Dasynema obtusifolium* (Moric.) Walp., *Ann. Bot. Syst.*

[Walpers] 1 (1): 110 [6-7 Nov. 1848] (Walpers 1848), “obtusum”. — *Sloanea obtusifolia* (Moric.) Planch. ex Benth., *J. Proc. Linn. Soc., Bot.* 5 (Suppl. 2): 63, 68 [May 1861] (Bentham 1861), “obtusa”.

*Sloanea fernando-costae* Hoehne, *Arq. Bot. Estado São Paulo* 1 (4): 93 (Hoehne 1942).

**NOTES.** — *Dasynema “obtusum”* (Walpers 1848: 110) is treated here as a misspelling of *obtusifolium*, since Walpers unambiguously referenced *Adenobasium obtusifolium* Moric.; therefore it is not an illegitimate homonym of *D. obtusum* Splitg. Similarly, as Bentham (1861: 63) explicitely based his *Sloanea “obtusa”* on *A. obtusifolium*, the epithet is also to be corrected.

**HERBARIUM DATA (FG).** — 5 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier 4662*.

**INVENTORY DATA (FG).** — 11 trees in 11 plots; $F_{max} < 1\%$; $dbh_{inv} = 38.2$ cm.

[467] *Sloanea parviflora* Planch. ex Benth.

*J. Proc. Linn. Soc., Bot.* 5 (Suppl. 2): 63, 68 [May 1861] (Bentham 1861).

*Dasynema obtusum* Splitg., *Tijdschr. Nat. Geschied. Physiol.* 9: 98 [Aug.-Sep. 1842] (Splitgerber 1842). — *Sloanea obtusa* (Splitg.) K.Schum., *Fl. Bras. [Martius]* 12 (3): 181 [1 Nov. 1886] (Schumann 1886), *nom. illeg. hom., non* Planch. ex Benth. (1861) [synonym of *S. obtusifolia* (Moric.) K.Schum.]

*Sloanea parviflora* var. *pedicellata* Sagot, *Ann. Sci. Nat., Bot.* sér. 6, 11: 157 (Sagot 1881).

*Sloanea kappleriana* Pulle, *Enumer. Vasc. Pl. Surinam* 279 (Pulle 1906).

*Sloanea obidensis* Ducke, *Arch. Inst. Biol. Veg.* 2 (2): 160 [Dec. 1935] (Ducke 1935).

**HERBARIUM DATA (FG).** — 10 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 4429*.

**INVENTORY DATA (FG).** — 9 trees in 8 plots; $F_{max} < 1\%$; $dbh_{inv} = 26.4$ cm.

[468] *Sloanea pubescens* Benth.

*J. Proc. Linn. Soc., Bot.* 5 (Suppl. 2): 69 [May 1861] (Bentham 1861). — *Dasynema pubescens* Poepp., *Nova genera ac species plantarum [Poeppig & Endlicher]* 3: 74 [23-25 Jan. 1845] (Poeppig 1845), *nom. illeg. hom., non* Schott (1827) *nec* Turcz. (Turczaninow 1858). — *Sloanea egensis* Radlk., *Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 12: 329 (Radlkofer 1882).

*Sloanea gracilis* Uittien, *Recueil Trav. Bot. Néerl.* 22: 356 [“1925” publ. Jan. 1926] (Uittien 1926).

*Sloanea pseudodentata* Ducke, *Arch. Inst. Biol. Veg.* 2 (2): 159 [Dec. 1935] (Ducke 1935).

**NOTES.** — Although based on *Dasynema pubescens* Poepp., *S. pubescens* Benth. is to be treated as a new name because the former is illegitimate.

**VERNACULAR NAMES.** — Pa: ihap-kamwi-wašiune • Wp: uluku panali, ulukupalani • Cr: roukou-gran-bwa, roukou-sovaj • Fr: roucou grand bois • Br: urucurana.
HERBARIUM DATA (FG). — 24 collections at CAY. Sel. exs.: S. *Barrier* 5018.

INVENTORY DATA (FG). — 12 trees in 10 plots; $F_{max} < 1\%$; $dbh_{inv} = 40$ cm.

[469] *Sloanea rojasiae* Vásquez

*Arnaldoa* 16 (2): 57 [“2009” publ. Feb. 2010] (Vásquez 2010).

HERBARIUM DATA (FG). — A single collection, *D. Loubry* 833.

SIZE. — Up to 25 cm dbh (Pennington & Wise 2017).

[470] *Sloanea rufa* Planch. ex Benth.

J. Proc. Linn. Soc., Bot. 5 (Suppl. 2): 68 [May 1861] (Bentham 1861).

*Sloanea longicaudata* Ducke, Bol. Técn. Inst. Agron. N. 19: 14 (Ducke 1950).

*Sloanea filiformis* D.Samp. & V.C.Souza, *Phytotaxa* 16: 45 [4 Feb. 2011] (Sampaio & Souza 2011).

HERBARIUM DATA (FG). — 19 collections at CAY. Sel. exs.: *J. Martin s.n.* (holo-, K[K000380085, K000380086, K000380087]).

INVENTORY DATA (FG). — 19 trees in 18 plots; $F_{max} < 1\%$; $dbh_{inv} = 34.3$ cm.

[471] *Sloanea sinemariensis* Aubl.

Hist. Pl. Guiane 1: 534 [Jun.-Dec. 1775] (Aublet 1775). — *Sloanea aubletii* Sw., Fl. Ind. Occid. 2: 940 [late 1800] (Swartz 1800), nom. illeg. superfl. (based on *Sloanea sinemariensis*).

VERNACULAR NAMES. — Pa: ihap-kamwi-priye • Cr: roukou-gran-bwa, roukou-sovaj • Fr: roucou grand bois • Br: urucurana.

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 4421.

INVENTORY DATA (FG). — 13 trees in 7 plots; $F_{max} < 1\%$; $dbh_{inv} = 63$ cm.

[472] *Sloanea synandra* Spruce ex Benth. (Fig. 22C)

J. Proc. Linn. Soc., Bot. 5 (Suppl. 2): 66 [May 1861] (Bentham 1861).

*Sloanea macrantha* Ducke, Arch. Inst. Biol. Veg. 2 (2): 162 [Dec. 1935] (Ducke 1935).

*Sloanea megacarpa* Steyerm. & Marc.-Berti, Bol. Soc. Venez. Ci. Nat. 26: 467 (Steyermark & Marcano-Berti 1966).

VERNACULAR NAMES. — Wp: wila kalayulu, wilakayulu • Cr: roukou-gran-bwa, roukou-sovaj • Fr: roucou grand bois • Br: urucurana.

HERBARIUM DATA (FG). — 24 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 4848.

INVENTORY DATA (FG). — 22 trees in 16 plots; $F_{max} < 1\%$; $dbh_{inv} = 100$ cm.

[473] *Sloanea trichosticha* R.O.Williams & Sandwith

Fl. Trinidad & Tobago 1 (2): 110 [Nov. 1929] (Williams & Sandwith 1929).

HERBARIUM DATA (FG). — 21 collections at CAY. Sel. exs.: *J.-F. Molino & M.-F. Prévost* 2531.

INVENTORY DATA (FG). — 31 trees in 23 plots; $F_{max} < 1\%$; $dbh_{inv} = 62.9$ cm.

[474] *Sloanea wurdackii* Steyerm.

Ann. Missouri Bot. Gard. 75 (4): 1583 [“1988” publ. 8 Mar. 1989] (Steyermark 1989).

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *B. Riéra* 1257.

SIZE. — Up to 45 cm dbh (Pennington & Wise 2017).

[475] *Sloanea* sp. A

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier* 4792.

INVENTORY DATA (FG). — 17 trees in 15 plots; $F_{max} < 1\%$; $dbh_{inv} = 80$ cm.

[476] *Sloanea* sp. B

HERBARIUM DATA (FG). — 16 collections at CAY. Sel. exs.: *S.A. Mori & J.J. Pipoly* 15498.

INVENTORY DATA (FG). — 28 trees in 24 plots; $F_{max} < 1\%$; $dbh_{inv} = 100$ cm.

[477] *Sloanea* sp. C

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: *M.-F. Prévost et al.* 5014.

INVENTORY DATA (FG). — 4 trees in 4 plots; $F_{max} < 1\%$; $dbh_{inv} = 50$ cm.

[478] *Sloanea* sp. D

NOTE. — This is possibly *S. caribaea* Krug & Urb. ex Duss.

HERBARIUM DATA (FG). — A single collection, *H. Richard & Bagadi* 674, dbh = 35 cm.

[479] *Sloanea* sp. E

HERBARIUM DATA (FG). — A single collection, *D. Sabatier et al.* 5806, dbh = 35 cm.
[480] *Sloanea* sp. F

NOTES. — According to Pennington & Wise (2017), this fruiting collection could belong to *S. ferruginea* T.D.Penn., a species whose fruit is not known. However, the leaves of this collection are quite different from those of *S. ferruginea*. This specimen does not fit in any other French Guianan species.

HERBARIUM DATA (FG). — A single collection, *S.A. Mori et al. 25669*, 35 m × 75 cm.

[481] *Sloanea* sp. G

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier 2990*.

INVENTORY DATA (FG). — 51 trees in 36 plots; $F_{max} < 1\%$; $dbh_{inv} = 70$ cm.

[482] *Sloanea* sp. H

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 2947*.

INVENTORY DATA (FG). — 15 trees in 10 plots; $F_{max} < 1\%$; $dbh_{inv} = 70$ cm.

Family ERYTHROXYLACEAE Kunth
Genus *Erythroxylum* P.Browne

[483] *Erythroxylum amazonicum* Peyr.

Fl. Bras. [Martius] 12 (1): 167 [1 Dec. 1878] (Peyritsch 1878).

*Erythroxylum mucronatum* var. *majus* Sagot, *Ann. Sci. Nat., Bot.* sér. 6, 11: 179 (Sagot 1881), "major".

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: *P.A. Sagot s.n.*, 1858 (P[P05483039]).

INVENTORY DATA (FG). — 5 trees in 4 plots; $F_{max} < 1\%$; $dbh_{inv} = 15.7$ cm.

[484] *Erythroxylum citrifolium* A.St.-Hil.

Fl. Bras. Merid. [A. St.-Hil.] (quarto ed.) 2 (13): 94 [10 Oct. 1829] (Saint-Hilaire 1829).

*Erythroxylum gomphioides* Planch. & Linden ex Triana & Planch., *Ann. Sci. Nat., Bot.* sér. 4, 18: 341 (Triana & Planchon 1862).

*Erythroxylum micranthum* Bong. ex Peyr., *Fl. Bras. [Martius] 12* (1): 164 [1 Dec. 1878] (Peyritsch 1878).

*Erythroxylum paraense* Peyr., *Fl. Bras. [Martius] 12* (1): 164 [1 Dec. 1878] (Peyritsch 1878).

*Erythroxylum acutifolium* Steud. ex Peyr., *Fl. Bras. [Martius] 12* (1): 166 [1 Dec. 1878] (Peyritsch 1878).

*Erythroxylum citrifolium* var. *latifolium* O.E.Schulz, *Pflanzenr. [Engler] IV.134* (Heft 29): 37 [13 Aug. 1907] (Schulz 1907).

*Erythroxylum citrifolium* var. *minus* O.E.Schulz, *Pflanzenr. [Engler] IV.134* (Heft 29): 37 [13 Aug. 1907] (Schulz 1907).

*Erythroxylum duckei* Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr. 5* (2): 416 (Huber 1909).

VERNACULAR NAMES. — Pa: agagut, ararut • Wp: pikau ki'ìy, pikau lemiù'i, tuká nákù sili • Br: pimenta-de-nambú.

HERBARIUM DATA (FG). — 50 collections at CAY. Sel. exs.: *S.A. Mori et al. 15394*.

INVENTORY DATA (FG). — 5 trees in 4 plots; $F_{max} < 1\%$; $dbh_{inv} = 13$ cm.

[485] *Erythroxylum fimbriatum* Peyr.
(Fig. 21C)

Fl. Bras. [Martius] 12 (1): 162 [1 Dec. 1878] (Peyritsch 1878).

HERBARIUM DATA (FG). — 13 collections at CAY. Sel. exs.: *R.A.A. Oldeman B-2424*, 12 m × 25 cm.

[486] *Erythroxylum kapplerianum* Peyr.

Fl. Bras. [Martius] 12 (1): 159 [1 Dec. 1878] (Peyritsch 1878).

*Erythroxylum mapuerae* Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr. 5* (2): 421 (Huber 1909).

NOTE. — *E. grisebachii* Peyr and *E. coelophlebium* Mart. var. *grisebachii* (Peyr.) O.E.Schulz are possible synonyms of this taxon (Plowman & Hensold 2004).

HERBARIUM DATA (FG). — 23 collections at CAY. Sel. exs.: *R.A.A. Oldeman T-663*.

SIZE. — Up to 10 m tall (Plowman & Berry 1999).

[487] *Erythroxylum leptoneurum* O.E.Schulz

*Pflanzenr. [Engler] IV.134* (Heft 29): 101 [13 Aug. 1907] (Schulz 1907).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *J.-J. de Granville et al. 6480*, 6 m × 10 cm.

[488] *Erythroxylum ligustrinum* DC.

Prodr. [A. P. de Candolle] 1: 574 [mid Jan. 1824] (Candolle 1824).

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: *J.-B. Patris s.n.* (holo-, G-DC[G00211017]; iso-, F[V0055692F], G[G00353856], P[P00723895]).

INVENTORY DATA (FG). — 5 trees in 5 plots; $F_{max} < 1\%$; $dbh_{inv} = 20.1$ cm.

[489] *Erythroxylum lineolatum* DC.

Prodr. [A. P. de Candolle] 1: 575 [mid Jan. 1824] (Candolle 1824).
<figure><img src="image_33.png" /><figcaption>FIG. 22. — Elaeocarpaceae: **A**, *Sloanea acutiflora* Uittien (D. Sabatier 1234); **B**, *Sloanea erythrocarpa* T.D.Penn. (D. Sabatier 3540); **C**, *Sloanea synandra* Spruce ex Benth. (D. Sabatier & M.-F. Prévost 4964); **D**, *Sloanea latifolia* (Rich.) K.Schum. (D. Sabatier 2088). © D. Sabatier/IRD.</figcaption></figure>

**HERBARIUM DATA (FG).** — A single collection, *J.-B. Patris s.n.* (holo-, G-DC[G00211384]; iso-, G[G00353858]; probable iso-, P[P00723900]).

**INVENTORY DATA (FG).** — 1 tree, dbh = 11.1 cm.

[490] *Erythroxylum macrophyllum* Cav.

Diss. 10: 401 (Cavanilles 1789).

*Erythroxylum lucidum* Kunth, *Nova genera et species plantarum* [*H.B.K.] 5: 179 [25 Feb. 1822] (Kunth 1822).

*Erythroxylum floribundum* Mart., *Beitr. Erythroxylon*: 118 (Martius 1840).

*Erythroxylum laurinum* Planch. & Linden ex Triana & Planch., *Ann. Sci. Nat., Bot. sér. 4*, 18: 341 (Triana & Planchon 1862), *nom. illeg. hom., non* A.Massal. (Massalongo & Scarabelli 1859).

*Erythroxylum costaricense* Donn.Sm., *Bot. Gaz.* 23 (4): 240 [22 Apr. 1897] (Donnell Smith 1897). — *Erythroxylum lucidum* var. *costaricense* (Donn.Sm.) O.E.Schulz, *Pflanzenr. [Engler]* IV.134 (Heft 29): 25 [13 Aug. 1907] (Schulz 1907).

*Erythroxylum ellipticum* Ramírez, *Anales Inst. Méd.-Nac. México* 3: 37 (Ramírez 1897), "*Erythroxylon*", *nom. illeg. hom., non* R.Br. ex Benth. (Bentham 1863) *nec* Peyr. (Peyritsch 1878).

*Erythroxylum tabascense* Britton, *N. Amer. Fl.* 25 (1): 66 (Britton 1907).

*Erythroxylum filipes* Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (2): 415 (Huber 1909).

**VERNACULAR NAMES.** — Pa: yauknabwi-kiabumna • Ka: akale tapulala kili • Wp: tuká nákŭ, tuká nákŭ sili • Wn: okajpu epelï • Br: pimenta-de-nambú.

**HERBARIUM DATA (FG).** — 74 collections at CAY. Sel. exs.: *D. Stoupy s.n.* (original material MA[MA475675?]); *M.-F. Prévost & D. Sabatier 2839*, dbh = 10 cm.
[491] *Erythroxylum mucronatum* Benth.

London J. Bot. 2: 372 (Bentham 1843).

*Erythroxylum comosum* O.E.Schulz, *Pflanzenr.* [Engler] IV.134 (Heft 29): 32 [13 Aug. 1907] (Schulz 1907).

*Erythroxylum kirkianum* O.E.Schulz, *Repert. Spec. Nov. Regni Veg.* 30: 179 (Schulz 1932).

*Erythroxylum venezuelense* Steyerm., *Fieldiana*, Bot. 28 (2): 271 (Steyermark 1952).

*Erythroxylum albertianum* Kuhl.m. & W.A.Rodrigues, *Publ. Inst. Nac. Pesq. Amaz.*, Bot. 5: 3 (Kuhlmann & Rodrigues 1957).

HERBARIUM DATA (FG). — 22 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier* 2230.

INVENTORY DATA (FG). — 2 trees in 2 plots; Fmax < 1%; dbhinv = 15 cm.

[492] *Erythroxylum roraimae* Klotzsch ex O.E.Schulz

*Pflanzenr.* [Engler] IV.134 (Heft 29): 104 [13 Aug. 1907] (Schulz 1907).

HERBARIUM DATA (FG). — A single collection, *R.A.A. Oldeman* 2238.

SIZE. — Up to 10 m tall (Plowman & Berry 1999).

[493] *Erythroxylum squamatum* Sw.

Prodr. [Swartz]: 75 [20 Jun.-29 July 1788] (Swartz 1788).

*Erythroxylum aristigerum* Peyr., *Fl. Bras.* [Martius] 12 (1): 157 [1 Dec. 1878] (Peyritsch 1878).

*Erythroxylum bahiense* Peyr., *Fl. Bras.* [Martius] 12 (1): 160 [1 Dec. 1878] (Peyritsch 1878). — *Erythroxylum aristigerum* var. bahiense (Peyr.) O.E.Schulz, *Pflanzenr.* [Engler] IV.134 (Heft 29): 103 [13 Aug. 1907] (Schulz 1907).

*Erythroxylum squamatum* var. microcarpum O.E.Schulz, *Symb. Antill. [Urban]* 5 (2): 192 [20 May 1907] (Schulz 1907).

*Erythroxylum trinerve* Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (2): 417 (Huber 1909).

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *P.A. Sagot* 1229, Mar. 1859 (holotype of *Erythroxylum squamatum* var. microcarpum: B? [not seen]; iso-, K[K000407437], P[P00723997, P00723998, P00723999], U[U0099038]).

INVENTORY DATA (FG). — 4 trees in 2 plots; Fmax < 1%; dbhinv = 12.1 cm.

[494] *Erythroxylum suberosum* A.St.-Hil. (Fig. 21D)

A.Juss. & Cambess., *Pl. Usuel. Bras.* 14: t. 69, fig. A (Saint-Hilaire et al. 1828).

*Erythroxylum areolatum* Vell., *Fl. Flumin.* 194 [“1825” publ. 7 Sep.-28 Nov. 1829] (Vellozo 1829), “Erythroxylon”, nom. illeg. hom., non L. (Linnaeus 1759).

*Erythroxylum testaceum* Peyr., *Fl. Bras.* [Martius] 12 (1): 170 [1 Dec. 1878] (Peyritsch 1878).

*Erythroxylum suberosum* f. brevipetiolatum O.E.Schulz, *Pflanzenr.* [Engler] IV.134 (Heft 29): 27 [13 Aug. 1907] (Schulz 1907).

VERNACULAR NAMES. — Br: cabelo-de-negro, galinha-choca.

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino* 5198.

INVENTORY DATA (FG). — 3 trees in 2 plots; Fmax < 1%; dbhinv = 65.4 cm.

Family EUPHORBIACEAE Juss.
Genus *Adenophaedra* (Müll.Arg.) Müll.Arg.

[495] *Adenophaedra grandifolia* (Klotzsch) Müll.Arg.

Fl. Bras. [Martius] 11 (2): 385 [1 May 1874] (Müller 1874).

*Tragia grandifolia* Klotzsch, *London J. Bot.* 2: 46 (Klotzsch 1843). — *Bernardia grandifolia* (Klotzsch) Müll.Arg., *Linnaea* 34: 173 (Müller 1865).

*Cleidion denticulatum* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 4 (8): 218 [24 Oct. 1929] (Standley 1929). — *Bernardia denticulata* (Standl.) G.L.Webster, *Ann. Missouri Bot. Gard.* 54 (2): 200 [27 Oct. 1967] (Webster 1967).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *J.-J. de Granville et al.* 10656.

SIZE. — Guyana. H.D. Clarke et al. 9046 (MO), 12 m.

Genus *Alchornea* Sw.

[496] *Alchornea discolor* Poepp.

*Nova genera ac species plantarum* [Poeppig & Endlicher] 3: 19 [15-21 Aug. 1841] (Poeppig 1841).

*Alchornea schomburgkii* Klotzsch, *London J. Bot.* 2: 46 (Klotzsch 1843).

*Alchornea gardneri* Müll.Arg., *Flora* 47: 435 (Müller 1864).

*Alchornea glaziovii* Pax & K.Hoffm., *Pflanzenr. [Engler]* IV.147.VII (Heft 63): 238 [10 Nov. 1914] (Pax & Hoffmann 1914), in nota.

*Alchornea brachygyne* Pax & K.Hoffm., *Pflanzenr. [Engler]* IV.147. XIV (Heft 68): 20 [6 June 1919] (Pax & Hoffmann 1919).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: wakavu-kamwi, waravru-kamwi • Wp: pesupi • Wn: kunui, uluimë • Nt: afatu adeeten • Br: amor-seco, sardinheiro, supiárana, tapiá, tartaruginha.

HERBARIUM DATA (FG). — 16 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 4830.

INVENTORY DATA (FG). — 5 trees in 5 plots; Fmax < 1%; dbhinv = 52.4 cm.
[497] *Alchornea triplinervia* (Spreng.) Müll.Arg.

Prodr. [A. P. de Candolle] 15 (2.2): 909 [late Aug. 1866] (Müller 1866). — *Antidesma triplinervium* Spreng., *Neue Entdeck. Pflanzenk.* 2: 116 (Sprengel 1821). — *Alchornea triplinervia* var. *genuina* Müll. Arg., *Prodr. [A. P. de Candolle] 15 (2.2): 909* [late Aug. 1866] (Müller 1866), *nom. inval.* (Turland et al. 2018: Art. 24.3).

*Alchornea nemoralis* Mart., *Flora* 24 (2, Beibl.): 31 (Martius 1841). — *Alchornea triplinervia* var. *nemoralis* (Mart.) Pax & K.Hoffm., *Pflanzenr.* [Engler] IV.147.VII (Heft 63): 228 [10 Nov. 1914] (Pax & Hoffmann 1914).

*Alchornea janeirensis* Casar., *Atti Riunione Sci. Ital.* 3: 515 [“1841”] publ. June 1842] (Casaretto 1842). — *Alchornea nemoralis* var. *janeirensis* (Casar.) Baill., *Adansonia [Baillon]* 5: 239 (Baillon 1865). — *Alchornea triplinervia* var. *janeirensis* (Casar.) Müll. Arg., *Prodr. [A. P. de Candolle] 15 (2.2): 909* [late Aug. 1866] (Müller 1866).

*Alchornea parvifolia* Miq., *Linnaea* 22: 797 [“1849”] publ. May 1850] (Miquel 1850). — *Alchornea triplinervia* var. *parvifolia* (Miq.) Müll.Arg., *Prodr. [A. P. de Candolle] 15 (2.2): 910* [late Aug. 1866] (Müller 1866).

*Alchornea glandulosa* Poepp. var. *parvifolia* Benth., *Hooker's J. Bot. Kew Gard. Misc.* 6: 330 (Bentham 1854). — *Alchornea triplinervia* var. *laevigata* Müll.Arg., *Prodr. [A. P. de Candolle] 15 (2.2): 910* [late Aug. 1866] (Müller 1866), *nom. illeg. superfl.* (*A. glandulosa* var. *parvifolia* in synonymy).

*Alchornea intermedia* Klotzsch ex Benth., *Hooker's J. Bot. Kew Gard. Misc.* 6: 331 (Bentham 1854), *nom. nud.*

*Alchornea parvifolia* Klotzsch ex Benth., *Hooker's J. Bot. Kew Gard. Misc.* 6: 331 (Bentham 1854), *nom. nud.*

*Alchornea psilorhachis* Klotzsch ex Benth., *Hooker's J. Bot. Kew Gard. Misc.* 6: 331 (Bentham 1854), *nom. nud.*

*Alchornea rotundifolia* Moric. ex Baill., *Étude Euphorb.*: 447 (Baillon 1858), *nom. inval.* (not accepted by the author).

*Alchornea nemoralis* var. *lanceolata* Baill., *Adansonia [Baillon]* 5: 239 (Baillon 1865). — *Alchornea triplinervia* var. *lanceolata* (Baill.) Müll.Arg., *Prodr. [A. P. de Candolle] 15 (2.2): 910* [late Aug. 1866] (Müller 1866).

*Alchornea nemoralis* var. *parvifolia* Baill., *Adansonia [Baillon]* 5: 239 (Baillon 1865), *nom. nud.*

*Alchornea nemoralis* var. *psilorhachis* Baill., *Adansonia [Baillon]* 5: 239 (Baillon 1865), *nom. nud.*

*Alchornea nemoralis* var. *rotundifolia* Baill., *Adansonia [Baillon]* 5: 239 (Baillon 1865), *nom. nud.*

*Alchornea triplinervia f. intermedia* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 909 [late Aug. 1866] (Müller 1866). — *Alchornea nemoralis* var. *intermedia* Baill. ex Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 909 [late Aug. 1866] (Müller 1866), *nom. nud. pro syn.*

*Alchornea triplinervia f. psilorhachis* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 909 [late Aug. 1866] (Müller 1866).

*Alchornea triplinervia* var. *crassifolia* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 909 [late Aug. 1866] (Müller 1866).

*Alchornea triplinervia* var. *tomentella* Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 380 [1 May 1874] (Müller 1874).

*Alchornea triplinervia* var. *iricuranoides* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 5: 603 (Chodat & Hassler 1905).

*Alchornea acroneura* Pax & K.Hoffm., *Pflanzenr.* [Engler] IV.147. VII (Heft 63): 229 [10 Nov. 1914] (Pax & Hoffmann 1914).

*Alchornea brevistyla* Pax & K.Hoffm., *Pflanzenr.* [Engler] IV.147. VII (Heft 63): 227 [10 Nov. 1914] (Pax & Hoffmann 1914).

*Alchornea obovata* Pax & K.Hoffm., *Pflanzenr.* [Engler] IV.147.VII (Heft 63): 223 [10 Nov. 1914] (Pax & Hoffmann 1914).

*Alchornea triplinervia* var. *boliviana* Pax & K.Hoffm., *Pflanzenr.* [Engler] IV.147.VII (Heft 63): 227 [10 Nov. 1914] (Pax & Hoffmann 1914).

*Alchornea triplinervia* var. *trinitatis* L.Riley, *Bull. Misc. Inform. Kew* 1925 (3): 141 [17 Apr. 1925] (Riley 1925).

VERNACULAR NAMES. — Pa: wakavu-kamwi, waravru-kamwi • Ka: alepa weli, kiyelapolan • Wp: yāsī leaʿīy • Wn: pakutpë • Br: tapiá, tinteiro.

HERBARIUM DATA (FG). — 20 collections at CAY. Sel. exs.: *D. Sabatier* 2346.

INVENTORY DATA (FG). — 35 trees in 11 plots; $F_{max}$ = 3.4 %; $dbh_{inv}$ = 66.8 cm.

## Genus *Alchorneopsis* Müll.Arg.

[498] *Alchorneopsis floribunda* (Benth.) Müll.Arg.

*Linnaea* 34: 156 (Müller 1865). — *Alchornea glandulosa* var. *floribunda* Benth., *Hooker's J. Bot. Kew Gard. Misc.* 6: 331 (Bentham 1854). — *Alchornea nemoralis* var. *floribunda* (Benth.) Baill., *Adansonia [Baillon]* 5: 239 (Baillon 1865).

*Alchorneopsis* trimera Lanj., *Euphorb. Surinam* 23 (Lanjouw 1931).

VERNACULAR NAMES. — Pa: wakavu-kamwi, waravru-kamwi • Ka: atapilipyo, wala ekone • Wp: mulei • Wn: pakutpë • Br: canelarana.

HERBARIUM DATA (FG). — 38 collections at CAY. Sel. exs.: *M.-F. Prévost* 3456.

INVENTORY DATA (FG). — 24 trees in 19 plots; $F_{max}$ < 1 %; $dbh_{inv}$ = 45 cm.

## Genus *Aparisthmium* Endl.

[499] *Aparisthmium cordatum* (A.Juss.) Baill.

*Adansonia [Baillon]* 5: 307 (Baillon 1865). — *Conceveibum* cor-datum A.Juss., *Euphorb. Gen.*: 43 [21 Feb. 1824] (Jussieu 1824), “Conceveibum”. — *Alchornea cordata* (A.Juss.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 901 [late Aug. 1866] (Müller 1866), *nom. illeg. hom.*, *non* Benth. (Bentham 1849).

*Alchornea macrophylla* Mart., *Flora* 24 (2, Beibl.): 31 (Martius 1841). — *Aparisthmium macrophyllum* (Mart.) Benth., *Hooker's J. Bot. Kew Gard. Misc.* 6: 333 (Bentham 1854). — *Conceveiba macrophylla* Klotzsch ex Benth., *Hooker's J. Bot. Kew Gard. Misc.* 6: 333 (Bentham 1854), *nom. nud. pro syn.*
*Alchornea latifolia* Klotzsch, *London J. Bot.* 2: 46 (Klotzsch 1843), *nom. illeg. hom.*, *non Sw.* (Swartz 1788).

*Styloceras macrostachyum* Poepp. ex Baill., *Monog. Bux.*: 81 (Baillon 1859).

*Aparishmium spruceanum* Baill., *Adansonia [Baillon]* 5: 307 (Baillon 1865).

*Alchornea orinocensis* Croizat, *J. Arnold Arbor.* 26 (2): 191 [16 Apr. 1945] (Croizat 1945).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: ahakyu-kamwi, ihap-kamwi, waikwavia-kamwi, wakavu-kamwi, waravru-kamwi • Ka: saulanani • Te: a’i mididju • Wp: a’i meyu • Nt: busi kusuwe • Br: mameleiro, morocototó, urucurana.

HERBARIUM DATA (FG). — 70 collections at CAY. Sel. exs.: *L. C. Richard s.n.* (original material P[P05574226]).

INVENTORY DATA (FG). — 58 trees in 22 plots; Fmax = 2.4%; dbhinv = 17.5 cm.

Genus *Caryodendron* H.Karst.

[500] *Caryodendron amazonicum* Ducke

*Trop. Woods* 76: 18 (Ducke 1943).

VERNACULAR NAMES. — Br: castanha-de-porco.

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: *D. Sabatier 4707*.

INVENTORY DATA (FG). — 3 trees in 2 plots; Fmax < 1%; dbhinv = 33.9 cm.

Genus *Conceveiba* Aubl.

[501] *Conceveiba guianensis* Aubl.

*Hist. Pl. Guiane* 2: 924 [Jun.-Dec. 1775] (Aublet 1775).

*Conceveibum ovatum* A.Juss., *Euphorb. Gen.*: 43 [21 Feb. 1824] (Jussieu 1824), "*Conceveibum*".

*Conceveiba trigonocarpa* Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 372 [1 May 1874] (Müller 1874).

*Conceveiba simulata* Steyerm., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (5): 416 [27 May 1938] (Steyermark 1938).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: ihap-kamwi, wakavu-kamwi, waravru-kamwi • Ka: alemenango wewe, mapiwalan, yawalelan • Te: a’i mididju • Wp: a’i miniyu, aminiyu lâ, miniyu lâ • Wn: palulumuli • Nt: kusisi, lebi udu • Cr: oubarouna • Br: arara-seringa, arraieira, tapichó.

HERBARIUM DATA (FG). — 142 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000947432]).

INVENTORY DATA (FG). — 575 trees in 139 plots; Fmax = 4.5%; dbhinv = 50 cm.

[502] *Conceveiba krukoffii* Steyerm.

*Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (5): 414 [27 May 1938] (Steyermark 1938).

HERBARIUM DATA (FG). — A single collection, *F. Billiet & B. Jadin 1230* (BR) (not seen, *fide* Secco 2004).

SIZE. — Up to 25 cm dhb (Secco 2004).

[503] *Conceveiba martiana* Baill.
(Fig. 23A)

*Adansonia [Baillon]* 5: 221 (Baillon 1865). — *Conceveibastrum martianum* (Baill.) Pax & K.Hoffm., *Pflanzenr. [Engler]* IV.147. VII (Heft 63): 217 [10 Nov. 1914] (Pax & Hoffmann 1914). — *Alchornea martiana* (Baill.) Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 375 [1 May 1874] (Müller 1874).

*Conceveiba megalophylla* Müll.Arg., *Linnaea* 34: 167 (Müller 1865).

VERNACULAR NAMES. — Wp: a’i meyu • Br: arraieira-branca.

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: *P. Grenand 3502*.

INVENTORY DATA (FG). — 4 trees in 2 plots; Fmax < 1%; dbhinv = 40.7 cm.

Genus *Croton* L.

[504] *Croton cajucara* Benth.

*Hooker's J. Bot. Kew Gard. Misc.* 6: 376 (Bentham 1854). — *Oxydectes cajucara* (Benth.) Kuntze, *Revis. Gen. Pl.* 2: 611 [5 Nov. 1891] (Kuntze 1891).

*Croton seputubensis* Hoehne, *Exped. Roosevelt-Rondon. Bot.*: 53 (Hoehne 1914).

*Croton motilonorum* Croizat, *Mem. Soc. Ci. Nat. La Salle* 10: 126 (Croizat 1950).

VERNACULAR NAMES. — Br: sacaca.

HERBARIUM DATA (FG). — 5 collections at CAY. Sel. exs.: *H. Richard & J. Ateni* 432.

SIZE. — Brazil, Maranhão. *J. Jangoux* 1051 (MO), 10 m × 12 cm.

[505] *Croton cuneatus* Mart. ex Klotzsch

*London J. Bot.* 2: 49 (Klotzsch 1843). — *Macrocroton cuneatus* (Mart. ex Klotzsch) Klotzsch, *Reis. Br.-Guiana [Ri. Schomburgk]* 3: 1186 [“1848” publ. 7-10 Mar. 1849] (Klotzsch 1849). — *Oxydectes cuneatus* (Mart. ex Klotzsch) Kuntze, *Revis. Gen. Pl.* 2: 611 [5 Nov. 1891] (Kuntze 1891).

*Croton surinamensis* Müll.Arg., *Linnaea* 34 (1): 82 [pre-Mar. 1865] (Müller 1865). — *Oxydectes surinamensis* (Müll.Arg.) Kuntze, *Revis. Gen. Pl.* 2: 613 [5 Nov. 1891] (Kuntze 1891).

*Croton martii* Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 88 [1 Feb. 1873] (Müller 1873). — *Oxydectes martii* (Müll.Arg.) Kuntze, *Revis. Gen. Pl.* 2: 613 [5 Nov. 1891] (Kuntze 1891).
*Croton martii* var. *latifolius* Müll.Arg., *Fl. Bras.* [*Martius* 11 (2): 88 [1 Feb. 1873] (Müller 1873).

*Croton martii* var. *longifolius* Müll.Arg., *Fl. Bras.* [*Martius* 11 (2): 88 [1 Feb. 1873] (Müller 1873).

*Croton mimeticus* S.Moore, *Trans. Linn. Soc. London, Bot.* 4 (3): 453 [“1894-96” publ. Dec. 1895] (Moore 1895).

*Croton kaieteuri* Jabl., *Mem. New York Bot. Gard.* 12 (3): 155 (Jablonszky 1965).

*Croton subcoriaceus* Jabl., *Mem. New York Bot. Gard.* 12 (3): 156 (Jablonszky 1965).

*Croton monachinoensis* Jabl., *Mem. New York Bot. Gard.* 12 (3): 157 (Jablonszky 1965).

*Croton bilocularis* J.Murillo, *Caldasia* 21 (2): 156 (Murillo 1999).

NOTES. — *C. surinamensis* Müll.Arg. is based on *C. cuneatus sensu* Miquel (1848: 477); hence it is not a replacement name (see Turland *et al.* 2018: Art. 41.7, note 3). However, it is validly published since it refers to Miquel’s description and types.

VERNACULAR NAMES. — Ka: tasi • Wp: mayawa'i • Wn: ehnaijap, enajop, mojoi • Nt: ingi udu • Br: catinga-de-porco.

HERBARIUM DATA (FG). — 35 collections at CAY. Sel. exs.: *M.-F. Prévost 3847*.

INVENTORY DATA (FG). — 1 tree, dbh = 24.8 cm.

[506] *Croton draconoides* Müll.Arg.
(Fig. 23B)

*Linnaea* 34 (1): 90 [pre-Mar. 1865] (Müller 1865). — *Oxydectes draconoides* (Müll.Arg.) Kuntze, *Revis. Gen. Pl.* 2: 611 [5 Nov. 1891] (Kuntze 1891), *“draconodes”*.

HERBARIUM DATA (FG). — 14 collections at CAY. Sel. exs.: *M.-F. Prévost 185*.

INVENTORY DATA (FG). — 3 trees in 1 plot; Fmax < 1%; dbhinv = 20 cm.

[507] *Croton icabarui* Jabl.

*Mem. New York Bot. Gard.* 12 (3): 158 (Jablonszky 1965).

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *J. Engel et al.* 14, dbh = 10 cm.

[508] *Croton matourensis* Aubl.

*Hist. Pl. Guiane* 2: 879 [Jun.-Dec. 1775] (Aublet 1775), *“Matourense”*. — *Croton sericeus* Lam., *Encycl. [J. Lamarck et al.]* 2 (1): 210 [16 Oct. 1786] (Lamarck 1786), *nom. illeg. superfl.* (based on *Croton matourensis*). — *Oxydectes matourensis* (Aubl.) Kuntze, *Revis. Gen. Pl.* 2: 612 [5 Nov. 1891] (Kuntze 1891).

*Croton caryophyllus* Benth., *Hooker's J. Bot. Kew Gard. Misc.* 6: 374 (Bentham 1854). — *Oxydectes caryophyllus* (Benth.) Kuntze, *Revis. Gen. Pl.* 2: 611 [5 Nov. 1891] (Kuntze 1891).

*Croton matourensis* var. *benthamianus* Müll.Arg., *Linnaea* 34 (1): 95 [pre-Mar. 1865] (Müller 1865).

*Croton matourensis* var. *poeppigianus* Müll.Arg., *Linnaea* 34 (1): 95 [pre-Mar. 1865] (Müller 1865).

*Croton matourensis* var. *sericeus* Müll.Arg., *Linnaea* 34 (1): 95 [pre-Mar. 1865] (Müller 1865).

*Croton kavanayensis* Steyerm., *Fieldiana, Bot.* 28 (2): 313 (Steyermark 1952).

*Croton lanjouwianus* Jabl., *Mem. New York Bot. Gard.* 12 (3): 158 (Jablonszky 1965), *“lanjouwensis”*.

*Croton pakaraimae* Jabl., *Mem. New York Bot. Gard.* 12 (3): 159 (Jablonszky 1965).

VERNACULAR NAMES. — Pa: á-seiminio-kamwi, á-seiminio-kamwipriyu, mariuvra-wašiunó • Ka: kunapolan, mulewabo • Wp: mayawa'i • Wn: enajop, mojoi • Nt: ingi udu • Cr: bwa-ranmié • Fr: bois indien • Br: dima, maravuvuiá.

HERBARIUM DATA (FG). — 49 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material S, S-R-1308).

INVENTORY DATA (FG). — 57 trees in 11 plots; Fmax = 2.7%; dbhinv = 38.8 cm.

[509] *Croton palanostigma* Klotzsch

*London J. Bot.* 2: 48 (Klotzsch 1843). — *Oxydectes palanostigma* (Klotzsch) Kuntze, *Revis. Gen. Pl.* 2: 612 [5 Nov. 1891] (Kuntze 1891).

*Croton benthamianus* Müll.Arg., *Fl. Bras.* [*Martius* 11 (2): 105 [1 May 1874] (Müller 1874).

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *S.A. Mori et al.* 24778.

SIZE. — Up to 15 m tall (Gillespie 2002).

[510] *Croton schiedeanus* Schltdl.

*Linnaea* 19: 243 [“1847” publ. Feb. 1846) (Schlechtendal 1846). — *Oxydectes schiedeana* (Schltdl.) Kuntze, *Revis. Gen. Pl.* 2: 613 [5 Nov. 1891] (Kuntze 1891).

*Croton perobtusus* Lundell, *Phytologia* 1 (12): 405 [Oct. 1940] (Lundell 1940).

VERNACULAR NAMES. — Wp: pesupi.

HERBARIUM DATA (FG). — 32 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 4423*.

INVENTORY DATA (FG). — 58 trees in 9 plots; Fmax < 1%; dbhinv = 22.7 cm.

[511] *Croton* sp. A

NOTE. — This entity regroups specimens that are wrongly identified at CAY as *Croton tricolor* Klotzsch ex Baill., a species not present in French Guiana (Ricarda Riina, personal communication to JE, 2019).

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *J.-J. de Granville 13403*, dbh = 20 cm.
<figure><img src="image_34.png" /><figcaption>FIG. 23. — Euphorbiaceae: **A**, Conceveiba martiana Baill. (J.-F. Molino & D. Sabatier 2365); **B**, Croton draconoides Müll.Arg. (D. Sabatier 6013); **C**, Glycydendron amazonicum Ducke (D. Sabatier & E. Fonty 5760); **D**, Sapium glandulosum (L.) Morong (J.-F. Molino & D. Sabatier 2255). A, D, © J.-F. Molino/IRD; B, C, © D. Sabatier/IRD.</figcaption></figure>

## Genus *Dodecastigma* Ducke

[512] *Dodecastigma integrifolium* (Lanj.) Lanj. & Sandwith

Kew Bull. 5 (1): 134 (Lanjouw & Sandwith 1950). — *Pausandra integrifolia* Lanj., *Bull. Misc. Inform. Kew* 1932 (4): 183 [13 June 1932] (Lanjouw 1932).

*Dodecastigma mazarunense* Croizat, *Bull. Torrey Bot. Club* 75 (4): 404 [Jul.-Aug. 1948] (Croizat 1948).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: S.A. Mori et al. 22811.

INVENTORY DATA (FG). — 7 trees in 2 plots; Fmax = 1 %; dbhinv = 23.4 cm.
[513] *Dodecastigma uleanum* (Pax & K.Hoffm.) G.L.Webster

*Contr. Univ. Michigan Herb.* 25: 237 [13 Aug. 2007] (Webster 2007). — *Cunuria uleana* Pax & K.Hoffm., *Pflanzenr. [Engler]* IV.147.XIV (Heft 68): 51 [6 June 1919] (Pax & Hoffmann 1919). — *Anomalocalyx uleanus* (Pax & K.Hoffm.) Ducke, *Notizbl. Bot. Gart. Berlin-Dahlem* 11: 344 [30 Mar. 1932] (Ducke 1932).

HERBARIUM DATA (FG). — A single collection, R.A.A. Oldeman B-3139.

SIZE. — Up to 25 cm dbh (Secco 2005).

## Genus *Glycydendron* Ducke

[514] *Glycydendron amazonicum* Ducke (Fig. 23C)

*Arch. Jard. Bot. Rio de Janeiro* 3: 199 (Ducke 1922).

VERNACULAR NAMES. — Pa: paig-seine, pairi-seine • Nt: wandekole • Br: castanha-de-porco, mirindiba-doce, pau-doce.

HERBARIUM DATA (FG). — 30 collections at CAY. Sel. exs.: *D. Sabatier & E. Fonty* 5760.

INVENTORY DATA (FG). — 106 trees in 82 plots; Fmax < 1%; dbhinv = 92.3 cm.

## Genus *Hevea* Aubl.

[515] *Hevea guianensis* Aubl.

*Hist. Pl. Guiane* 2: 871 [Jun.-Dec. 1775] (Aublet 1775), "*peruviana*" on plate. — *Siphonia guianensis* (Aubl.) Juss. ex Baill., *Étude Euphorb.* 326 (Baillon 1858). — *Caoutchoua guianensis* (Aubl.) O.F.Cook, *J. Wash. Acad. Sci.* 31: 58 (Cook 1941).

*Hevea peruviana* Aubl., *Hist. Pl. Guiane* 4: t. 335 [Jun.-Dec. 1775] (Aublet 1775).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: akabdat, katatu-avain • Ka: apalaba, mapalaba • Nt: afasi boyti, efaa • Cr: évéa • Fr: hévéa • Br: seringueira-itaúba.

HERBARIUM DATA (FG). — 43 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000812056]).

INVENTORY DATA (FG). — 214 trees in 70 plots; Fmax = 1.5%; dbhinv = 63.9 cm.

## Genus *Hura* L.

[516] *Hura crepitans* L.

*Sp. Pl.* 2: 1008 [1 May 1753] (Linnaeus 1753).

*Hura brasiliensis* Willd., * Enum. Pl. [Willdenow] * 2: 997 [June 1809] (Willdenow 1809).

*Hura strepens* Willd., * Enum. Pl. [Willdenow] * 2: 997 [June 1809] (Willdenow 1809). — *Hura crepitans* var. strepens (Willd.) Müll. Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1230 [late Aug. 1866] (Müller 1866).

*Hura senegalensis* Baill., *Adansonia [Baillon]* 1: 77 (Baillon 1860). — *Hura crepitans* var. senegalensis (Baill.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1230 [late Aug. 1866] (Müller 1866).

*Hura crepitans* f. oblongifolia Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1230 [late Aug. 1866] (Müller 1866).

*Hura crepitans* f. orbicularis Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1229 [late Aug. 1866] (Müller 1866).

*Hura crepitans* f. ovata Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1229 [late Aug. 1866] (Müller 1866).

*Hura crepitans* var. genuina Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1229 [late Aug. 1866] (Müller 1866), *nom. inval.* (Türland et al. 2018: Art. 24.3).

*Hura crepitans* var. membranacea Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1229 [late Aug. 1866] (Müller 1866).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: tenway • Ka: asiwakala, kumakalan • Wp: wasaku • Wn: mutata • Nt: posenti, postii • Cr: bwa-dyab • Fr: bois diable, pet du diable, sablier • Br: açacu.

HERBARIUM DATA (FG). — 18 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 4857.

INVENTORY DATA (FG). — 1 tree, dbh = 10.4 cm.

## Genus *Mabea* Aubl.

[517] *Mabea piriri* Aubl.

*Hist. Pl. Guiane* 2: 867 [Jun.-Dec. 1775] (Aublet 1775).

*Mabea maynensis* Spruce ex Benth., *J. Proc. Linn. Soc.*, *Bot.* 5: 7 ["1861" publ. June 1860] (Bentham 1860).

VERNACULAR NAMES. — Ka: yukuyapoi • Wp: awalapuna • Wn: isoimë watki • Nt: busi bisangula • Cr: bwa-lélé • Fr: bois à calumet • Br: seringaí, taquari.

HERBARIUM DATA (FG). — 122 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000799363]).

INVENTORY DATA (FG). — 143 trees in 43 plots; Fmax = 3.9%; dbhinv = 28.3 cm.

[518] *Mabea salicoides* Esser

*Novon* 3 (4): 349 [winter 1993] (Esser 1993).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *S.A. Mori & B.M. Boom* 15146.

SIZE. — Up to 18 cm dbh (Gillespie 2002).

[519] *Mabea speciosa* Müll.Arg.

*Fl. Bras. [Martius]* 11 (2): 520 [1 May 1874] (Müller 1874).
*Mabea caudata* Pax & K.Hoffm., *Pflanzenr. [Engler] IV.147.IV* (Heft 52): 282 [13 Feb. 1912] (Pax & Hoffmann 1912).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Wp: awalapuna, awalapuna sili, paa lo'o • Wn: tihstili epir • Cr: bwa-lélé • Br: seringaí, taquari.

HERBARIUM DATA (FG). — 17 collections at CAY. Sel. exs.: *M.-F. Prévost et al. 4470*.

INVENTORY DATA (FG). — 48 trees in 20 plots; Fmax = 2.2%; dbhinv = 29 cm.

[520] *Mabea subsessilis* Pax & K.Hoffm.

*Pflanzenr. [Engler] IV.147.IV* (Heft 52): 282 [13 Feb. 1912] (Pax & Hoffmann 1912).

*Mabea argutissima* Croizat, *Bull. Torrey Bot. Club 67* (4): 288 [Apr. 1940] (Croizat 1940).

HERBARIUM DATA (FG). — 18 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier 2307*.

INVENTORY DATA (FG). — 14 trees in 4 plots; Fmax = 1.4%; dbhinv = 28 cm.

[521] *Mabea taquari* Aubl.

Hist. Pl. Guiane 2: 870 [Jun.-Dec. 1775] (Aublet 1775).

*Mabea schomburgkii* Benth., *Hooker's J. Bot. Kew Gard. Misc.* 6: 365 (Bentham 1854).

*Mabea taquari* var. *angustifolia* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1150 [late Aug. 1866] (Müller 1866).

VERNACULAR NAMES. — Ka: akale loway • Wn: isoimë watkï.

HERBARIUM DATA (FG). — 43 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material A[A00055264], BM[BM000799374], F[V0056865F], P[P00716889]; possible original material LINN[LINN-HS 1470.2], MPU[MPU015282]).

SIZE. — Guyana. *H.D. Clarke 3146*, 12 m.

Genus *Maprounea* Aubl.

[522] *Maprounea guianensis* Aubl.

Hist. Pl. Guiane 2: 895 [Jun.-Dec. 1775] (Aublet 1775), "*Mapronnea Guyannensis*" on plate. — *Stillingia guianensis* (Aubl.) Baill., *Adansonia [Baillon]* 5: 332 (Baillon 1865). — *Excoecaria guianensis* (Aubl.) Baill., *Hist. Pl. [Baillon]* 5: 133 [Jan.-Apr. 1874] (Baillon 1874).

*Aegopricum betulinum* L.f., *Suppl. Pl.*: 413 ["1781" publ. Apr. 1782] (Linnaeus 1782), "*Aegopricon*".

VERNACULAR NAMES. — Pa: iduras-veiti, iduras-vey, kagegut, karegut • Ka: alisigii, pilapisi • Wp: ka'a ki, kasila, yalakasila, yalakasilo • Nt: ayee wiwii, kusangula • Cr: radjé-chank • Br: vaquinha.

HERBARIUM DATA (FG). — 59 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000073601, BM000073602], S[S-R-10787]).

INVENTORY DATA (FG). — 14 trees in 11 plots; Fmax < 1%; dbhinv = 54.1 cm.

Genus *Micrandra* R.Br.

[523] *Micrandra elata* (Didr.) Müll.Arg.

Linnaea 34: 142 (Müller 1865). — *Pogonophyllum elatum* Didr., *Vidensk. Meddel. Dansk Naturhist. Foren. Kjøbenhavn* 1857: 145 (Didrichsen 1857).

*Micrandra bracteosa* Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 290 [1 Feb. 1873] (Müller 1873).

*Micrandra glaziovii* Pax, *Pflanzenr. [Engler] IV.147.1* (Heft 42): 20 [3 May 1910] (Pax 1910).

*Micrandra brownsbergensis* Lanj., *Euphorb. Surinam* 34 (Lanjouw 1931).

*Micrandra santanderensis* Croizat, *J. Arnold Arbor.* 24 (2): 169 [15 Apr. 1943] (Croizat 1943).

NOTE. — *Micrandra bracteosa* Müll.Arg. is not the basionym of *Cunuria bracteosa* Ducke nor that of *Cunuria spruceana* var. *bracteosa* (Ducke) R.E.Schult. Both names are synonyms of *Micrandra spruceana* (Baill.) R.E.Schult.

VERNACULAR NAMES. — Ka: mule'ilan, tipulu alome.

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *D. Sabatier et al. 4781*.

INVENTORY DATA (FG). — 151 trees in 13 plots; Fmax = 1.4%; dbhinv = 47.1 cm.

[524] *Micrandra rossiana* R.E.Schult.

Bot. Mus. Leafl. 15 (8): 211 [29 May 1952] (Schultes 1952).

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: *D. Sabatier et al. 5802*.

INVENTORY DATA (FG). — 16 trees in 2 plots; Fmax = 1.9%; dbhinv = 38.2 cm.

Genus *Pausandra* Radlk.

[525] *Pausandra fordii* Secco

Bol. Mus. Paraense Emilio Goeldi, N.S., Bot. 3 (1): 60 (Secco 1987).

VERNACULAR NAMES. — Pa: á-agag.

HERBARIUM DATA (FG). — 20 collections at CAY. Sel. exs.: *R.S. Cowan 38370* (holo-, NY[00272986]; iso-, NY[00272985], ).

INVENTORY DATA (FG). — 256 trees in 31 plots; Fmax = 4.6%; dbhinv = 27.5 cm.

[526] *Pausandra martinii* Baill.

*Adansonia [Baillon]* 11: 92 (Baillon 1873), "*Martini*".

*Pausandra flagellorhachis* Lanj., *Euphorb. Surinam* 30 (Lanjouw 1931).

VERNACULAR NAMES. — Pa: kawap.

HERBARIUM DATA (FG). — 35 collections at CAY. Sel. exs.: *J. Martin s.n.* (holo-, P[P00648720]; iso-, P[P00648721]).

SIZE. — Up to 20 m tall (Gillespie 2002).
[527] *Pausandra trianae* (Müll.Arg.) Baill.

Adansonia [Baillon] 11: 92 (Baillon 1873). — *Pogonophora trianae*
Müll.Arg., *Flora* 47: 434 (Müller 1864).

*Pausandra quadriglandulosa* Pax & K.Hoffm., *Pflanzenr.* [Engler]
IV.147.XIV (Heft 68): 43 [6 June 1919] (Pax & Hoffmann 1919).

*Pausandra extorris* Standl., *Trop. Woods* 17: 24 [Mar. 1929] (Stand-
ley 1929).

*Pausandra densiflora* Lanj., *Recueil Trav. Bot. Néerl.* 33: 766 (Lan-
jouw 1936).

*Pausandra sericea* Lanj., *Recueil Trav. Bot. Néerl.* 33: 767 (Lanjouw 1936).

*Clavija septentrionalis* L.O.Williams, *Fieldiana, Bot.* 32 (12): 205
(Williams 1970).

VERNACULAR NAMES. — Br: orelha-de-burro.

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *H. Jac-quemin* 2435.

SIZE. — Up to 30 m tall (Webster & Huft 1988).

Genus *Sagotia* Baill.

[528] *Sagotia brachysepala* (Müll.Arg.) Secco

*Acta Amazonica Supl.* 15 (1-2): 81, 85 (Secco 1985). — *Sagotia
racemosa* var. *brachysepala* Müll.Arg., *Flora* 47: 516 (Müller 1864).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020), but
not yet found in inventories of trees above 10 cm dbh in French
Guiana.

HERBARIUM DATA (FG). — 5 collections at CAY. Sel. exs.: *M.-F. Prévost 2719*.

SIZE. — Up to 35 m tall (Secco 1999).

[529] *Sagotia racemosa* Baill.

Adansonia [Baillon] 1: 54 (Baillon 1860).

*Sagotia racemosa* var. *genuina* Müll.Arg., *Flora* 47: 516 (Müller 1864),
*nom. inval.* (Turland *et al.* 2018: Art. 24.3).

*Sagotia racemosa* var. *ligularis* Müll.Arg., *Flora* 47: 516 (Müller 1864).

*Sagotia racemosa* var. *microsepala* Müll.Arg., *Flora* 47: 517 (Müller 1864).

*Sagotia tafelbergii* Croizat, *Bull. Torrey Bot. Club* 75 (4): 404 [Jul.-Aug. 1948] (Croizat 1948).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Wp: tatu mila • Nt: makwebo.

HERBARIUM DATA (FG). — 48 collections at CAY. Sel. exs.: *J. Martin*
27 (original material P[P00648674]).

INVENTORY DATA (FG). — 251 trees in 52 plots; Fmax = 4.2 %;
dbhinv = 38.2 cm.

Genus *Sandwithia* Lanj.

[530] *Sandwithia guyanensis* Lanj.

Bull. Misc. Inform. Kew 1932 (4): 184 [13 June 1932] (Lanjouw 1932).

VERNACULAR NAMES. — Wp: wila tuwi • Br: ururcurana-branca.

HERBARIUM DATA (FG). — 59 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 3589*.

INVENTORY DATA (FG). — 677 trees in 75 plots; Fmax = 6.2 %;
dbhinv = 40.7 cm.

Genus *Sapium* Jacq.

[531] *Sapium argutum* (Müll.Arg.) Huber

Bull. Herb. Boissier, sér. 2, 6: 439 (Huber 1906). — *Excoecaria arguta* Müll.Arg., *Fl. Bras.* [*Martius* 11 (2): 614 [1 May 1874] (Müller 1874), “*Excaecaria*”.

*Sapium montanum* Lanj., *Euphorb. Surinam* 47 (Lanjouw 1931).

VERNACULAR NAMES. — Pa: itey-avain, itey-avan.

HERBARIUM DATA (FG). — 35 collections at CAY. Sel. exs.: *S.A. Mori et al.* 25536.

SIZE. — Up to 10 m tall (Santos & Sales 2009).

[532] *Sapium ciliatum* Hemsl.

Hooker's Icon. Pl. 27 [ser. 4, 7]: t. 2683 [Feb. 1901] (Hemsley 1901).

VERNACULAR NAMES. — Wp: melekene, melekene sili • Br: murupita.

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *M.-F. Prévost & P. Grenand 1964*.

SIZE. — Brazil, Pará. *B.V. Rabelo 3741* (MO), 15 m.

[533] *Sapium glandulosum* (L.) Morong
(Fig. 23D)

Ann. New York Acad. Sci. 7: 227 (Morong 1893). — *Hippomane
glandulosa* L., *Sp. Pl.* 2: 1191 [1 May 1753] (Linnaeus 1753). —
*Sapium jamaicense* Sw., *Adnot. Bot.*: 62 (Swartz 1829), *nom. illeg.
superfl.* (includes the type of *Hippomane glandulosa*).

*Sapium aucuparium* Jacq., * Enum. Syst. Pl. *: 21 [Aug.-Sep. 1760] (Jacquin 1760). — *Hippomane aucuparia* (Jacq.) Crantz, *Inst. Rei Herb.* 1: 169 (Crantz 1766). — *Stillingia aucuparia* (Jacq.) Oken, *Allg. Naturgesch.* 3 (3): 1607 (Oken 1841). — *Sapium biglandulosum* var. *aucuparium* (Jacq.) Müll.Arg., *Linnaea* 32: 119 (Müller 1863). — *Excoecaria biglandulosa* var. *aucuparia* (Jacq.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1206 [late Aug. 1866] (Müller 1866), “*Excaecaria*”.

*Hippomane biglandulosa* L., *Sp. Pl.*, ed. 2, 2: 1431 [Aug. 1763] (Linnaeus 1763), *nom. illeg. superfl.* (based on *Sapium aucuparium*). — *Sapium hippomane* G.Mey., *Prim. Fl. Esseq.* 275 [Nov. 1818] (Meyer 1818), *nom. illeg. superfl.* (based on *Hippomane*
biglandulosa). — *Sapium biglandulosum* Müll.Arg., *Linnaea* 32: 116 (Müller 1863). — *Stillingia biglandulosa* Baill., *Adansonia [Baillon]* 5: 320 (Baillon 1865). — *Sapium montevidense* Klotzsch ex Baill., *Adansonia [Baillon]* 5: 320 (Baillon 1865), nom. nud. pro syn. — *Excoecaria biglandulosa* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1204 [late Aug. 1866] (Müller 1866), “*Excaecaria*”.

*Sapium salicifolium* Kunth, *Nova genera et species plantarum* [H.B.K.] 2: 65 [28 Apr. 1817] (Kunth 1817). — *Sapium biglandulosum* var. salicifolium (Kunth) Müll.Arg., *Linnaea* 32: 119 (Müller 1863). — *Stillingia salicifolia* (Kunth) Klotzsch ex Baill., *Adansonia [Baillon]* 5: 320 (Baillon 1865), nom. illeg. hom., non (Torr.) Raf. (Rafinesque 1832). — *Excoecaria biglandulosa* var. salicifolia (Kunth) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1207 [late Aug. 1866] (Müller 1866), “*Excaecaria*”.

*Omphalea glandulata* Vell., *Fl. Flumin. Icon.* 10: t. 14 [“1827” publ. 29 Oct. 1831] (Vellozo 1831). — *Sapium glandulatum* (Vell.) Pax, *Pflanzenr.* [Engler] IV.147.IV (Heft 52): 229 [13 Feb. 1912] (Pax 1912). — *Excoecaria biglandulosa* var. glandulata (Vell.) Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 621 [1 May 1874] (Müller 1874), “*Excaecaria*”.

*Sapium prunifolium* Klotzsch, *London J. Bot.* 2: 45 (Klotzsch 1843). — *Stillingia prunifolia* (Klotzsch) Baill., *Étude Euphorb.* 513 (Baillon 1858). — *Excoecaria biglandulosa* var. prunifolia (Klotzsch) Müll. Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1205 [late Aug. 1866] (Müller 1866), “*Excaecaria*”.

*Sapium moritzianum* Klotzsch, *Bot. Voy. Herald [Seemann]* 3: 100 [Nov. 1853] (Klotzsch 1853). — *Sapium biglandulosum* var. moritzianum (Klotzsch) Müll.Arg., *Linnaea* 32: 119 (Müller 1863). — *Excoecaria biglandulosa* var. moritziana (Klotzsch) Müll. Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1206 [late Aug. 1866] (Müller 1866), “*Excaecaria*”, “*Moritzianum*”. — *Sapium aucuparium* subsp. *moritzianum* (Klotzsch) Pittier, *Contr. U.S. Natl. Herb.* 20 (3): 128 [18 June 1918] (Pittier 1918).

*Stillingia cremostachya* Baill., *Étude Euphorb.* 512 (Baillon 1858). — *Sapium cremostachyum* (Baill.) I.M.Johnst., *Contr. Gray Herb.* 68: 91 (Johnston 1923).

*Stillingia hippomane* Baill., *Étude Euphorb.* 513 (Baillon 1858).

*Sapium aucuparium* var. hippomane Griseb., *Fl. Brit. W.I. [Grisebach]* 49 [Dec. 1859] (Grisebach 1859).

*Sapium aereum* Klotzsch ex Müll.Arg., *Linnaea* 32: 119 (Müller 1863). — *Excoecaria aerea* (Klotzsch ex Müll.Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1207 [late Aug. 1866] (Müller 1866), “*Excaecaria*”. — *Sapium aureum* H.Buek, *Gen. Sp. Synon. Cand.* 4: 364 [Jul.-Aug. 1874] (Buek 1874), sphalm.

*Sapium biglandulosum* f. obovata Müll.Arg., *Linnaea* 32: 116 (Müller 1863). — *Excoecaria biglandulosa* f. obovata (Müll.Arg.) Müll. Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1205 [late Aug. 1866] (Müller 1866), “*Excaecaria*”.

*Sapium biglandulosum* var. aubletianum Müll.Arg., *Linnaea* 32: 116 (Müller 1863). — *Excoecaria biglandulosa* var. aubletiana (Müll.Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1205 [late Aug. 1866] (Müller 1866), “*Excaecaria*”, “*Aubletianum*”. — *Sapium aubletianum* (Müll.Arg.) Huber, *Bull. Herb. Boissier*, sér. 2, 6: 362 (Huber 1906).

*Sapium biglandulosum* var. hamatum Müll.Arg., *Linnaea* 32: 116 (Müller 1863). — *Excoecaria biglandulosa* var. hamata (Müll.Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1204 [late Aug. 1866] (Müller 1866). — *Sapium hamatum* (Müll.Arg.) Pax &

K.Hoffm., *Pflanzenr.* [Engler] IV.147.IV (Heft 52): 229 [13 Feb. 1912] (Pax & Hoffmann 1912).

*Sapium biglandulosum* f. pavonianum Müll.Arg., *Linnaea* 32: 116 (Müller 1863), “*Peruvianum*”. — *Excoecaria biglandulosa* var. pavoniana (Müll.Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1204 [late Aug. 1866] (Müller 1866), “*Excaecaria*”, “*Pavoniana*”. — *Sapium pavonianum* (Müll.Arg.) Huber, *Bull. Herb. Boissier*, sér. 2, 6: 356 (Huber 1906).

*Sapium obtusilobum* Müll.Arg., *Linnaea* 32: 116 (Müller 1863). — *Excoecaria obtusiloba* (Müll.Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1203 [late Aug. 1866] (Müller 1866), “*Excaecaria*”.

*Sapium biglandulosum* var. meyerianum Müll.Arg., *Linnaea* 32: 116 (Müller 1863).

*Sapium biglandulosum* f. oblongatum Müll.Arg., *Linnaea* 32: 117 (Müller 1863). — *Excoecaria biglandulosa* f. oblongata (Müll. Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1205 [late Aug. 1866] (Müller 1866), “*Excaecaria*”.

*Sapium biglandulosum* var. cuneatum Müll.Arg., *Linnaea* 32: 117 (Müller 1863). — *Excoecaria biglandulosa* var. cuneata (Müll. Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1206 [late Aug. 1866] (Müller 1866), “*Excaecaria*”.

*Sapium biglandulosum* var. klotzschianum Müll.Arg., *Linnaea* 32: 117 (Müller 1863). — *Excoecaria biglandulosa* var. klotzschiana (Müll.Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1205 [late Aug. 1866] (Müller 1866), “*Excaecaria*”, “*Klotzschianum*”. — *Sapium klotzschianum* (Müll.Arg.) Huber, *Bull. Herb. Boissier*, sér. 2, 6: 438 (Huber 1906).

*Sapium biglandulosum* f. minus Müll.Arg., *Linnaea* 32: 117 (Müller 1863).

*Sapium biglandulosum* var. lanceolatum Müll.Arg., *Linnaea* 32: 118 (Müller 1863). — *Excoecaria biglandulosa* var. lanceolata (Müll. Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1206 [late Aug. 1866] (Müller 1866), “*Excaecaria*”. — *Excoecaria marginata* var. lanceolata (Müll.Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1208 [late Aug. 1866] (Müller 1866), “*Excaecaria*”. — *Sapium lanceolatum* (Müll.Arg.) Huber, *Bull. Herb. Boissier*, sér. 2, 6: 441 (Huber 1906).

*Sapium biglandulosum* var. serratum Müll.Arg., *Linnaea* 32: 118 (Müller 1863). — *Sapium serratum* (Müll.Arg.) Klotzsch ex Baill., *Adansonia [Baillon]* 5: 320 (Baillon 1865). — *Excoecaria biglandulosa* var. serrata (Müll.Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1206 [late Aug. 1866] (Müller 1866), “*Excaecaria*”.

*Sapium marginatum* Müll.Arg., *Linnaea* 32: 120 (Müller 1863). — *Excoecaria marginata* (Müll.Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1208 [late Aug. 1866] (Müller 1866), nom. illeg. hom., non Kunze ex Baill. (Baillon 1858). — *Stillingia marginata* (Müll.Arg.) Baill., *Adansonia [Baillon]* 5: 321 (Baillon 1865).

*Sapium marginatum* f. majus Müll.Arg., *Linnaea* 32: 120 (Müller 1863). — *Excoecaria marginata* f. major (Müll.Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1208 [late Aug. 1866] (Müller 1866), “*Excaecaria*”.

*Sapium marginatum* var. spathulatum Müll.Arg., *Linnaea* 32: 120 (Müller 1863). — *Excoecaria marginata* var. spathulata (Müll. Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1208 [late Aug. 1866] (Müller 1866), “*Excaecaria*”.

*Sapium marginatum* var. lanceolatum Müll.Arg., *Linnaea* 32: 120 (Müller 1863).
*Stillingia dracunculoides* Baill., *Adansonia* [Baillon] 5: 321 (Baillon 1865), in obs. — *Excoecaria biglandulosa* var. *dracunculoides* (Baill.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1206 [late Aug. 1866] (Müller 1866), "*Excaecaria*".

*Sapium suberosum* Müll.Arg., *Linnaea* 34: 217 (Müller 1865). — *Excoecaria suberosa* (Müll.Arg.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1202 [late Aug. 1866] (Müller 1866).

*Excoecaria biglandulosa* var. *hippomane* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1204 [late Aug. 1866] (Müller 1866), "*Excaecaria*".

*Excoecaria biglandulosa* var. *clausseniana* Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 618 [1 May 1874] (Müller 1874), "*Excaecaria*", "*Clausseniana*". — *Sapium claussenianum* (Müll.Arg.) Huber, *Bull. Herb. Boissier*, sér. 2, 6: 436 (Huber 1906).

*Excoecaria biglandulosa* var. *intercedens* Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 620 [1 May 1874] (Müller 1874). — *Sapium intercedens* (Müll.Arg.) Huber, *Bull. Herb. Boissier*, sér. 2, 6: 437 (Huber 1906).

*Excoecaria biglandulosa* var. *grandifolia* Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 622 [1 May 1874] (Müller 1874), "*Excaecaria*".

*Excoecaria biglandulosa* var. *leptadenia* Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 620 [1 May 1874] (Müller 1874), "*Excaecaria*". — *Sapium leptadenium* (Müll.Arg.) Huber, *Bull. Herb. Boissier*, sér. 2, 6: 436 (Huber 1906).

*Excoecaria biglandulosa* var. *montevidensis* Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 621 [1 May 1874] (Müller 1874).

*Excoecaria biglandulosa* var. *petiolaris* Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 621 [1 May 1874] (Müller 1874), "*Excaecaria*". — *Sapium aucuparium* var. *petiolare* (Müll.Arg.) Griseb., *Abh. Königl. Ges. Wiss. Göttingen* 24: 62 (Grisebach 1879). — *Sapium petiolare* (Müll. Arg.) Huber, *Bull. Herb. Boissier*, sér. 2, 6: 434 (Huber 1906).

*Excoecaria marginata* var. *conjungens* Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 617 [1 May 1874] (Müller 1874), "*Excaecaria*". — *Sapium marginatum* var. *conjungens* (Müll.Arg.) Müll.Arg., *Pflanzenr. [Engler]* IV.147.IV (Heft 52): 224 [13 Feb. 1912] (Müller 1912).

*Excoecaria marginata* var. *grandifolia* Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 618 [1 May 1874] (Müller 1874), "*Excaecaria*". — *Sapium marginatum* var. *grandifolium* (Müll.Arg.) Müll.Arg., *Pflanzenr. [Engler]* IV.147.IV (Heft 52): 224 [13 Feb. 1912] (Müller 1912).

*Excoecaria marginata* var. *longifolia* Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 618 [1 May 1874] (Müller 1874), "*Excaecaria*". — *Sapium marginatum* var. *longifolium* (Müll.Arg.) Müll.Arg., *Pflanzenr. [Engler]* IV.147.IV (Heft 52): 224 [13 Feb. 1912] (Müller 1912).

*Excoecaria occidentalis* Müll.Arg., *Fl. Bras. [Martius]* 11 (2): 615 [1 May 1874] (Müller 1874), "*Excaecaria*". — *Sapium occidentale* (Müll.Arg.) Huber, *Bull. Herb. Boissier*, sér. 2, 6: 441 (Huber 1906).

*Sapium poeppigii* Hemsl., *Hooker's Icon. Pl.* 27 [ser. 4, 7]: t. 2678 [Feb. 1901] (Hemsley 1901).

*Sapium caribaeum* Urb., *Symb. Antill. [Urban]* 3 (2): 308 [15 Aug. 1902] (Urban 1902).

*Sapium taburu* Ule, *Bot. Jahrb. Syst.* 35 (5): 671 [18 Apr. 1905] (Ule 1905).

*Sapium pittieri* Huber, *Bull. Herb. Boissier*, sér. 2, 6: 350 (Huber 1906).

*Sapium bogotense* Huber, *Bull. Herb. Boissier*, sér. 2, 6: 355 (Huber 1906). — *Sapium biglandulosum* var. *bogotense* (Huber) Monach., *Bull. Torrey Bot. Club* 67 (9): 772 [Dec. 1940] (Monachino 1940).

*Sapium hemsleyanum* Huber, *Bull. Herb. Boissier*, sér. 2, 6: 362 (Huber 1906).

*Sapium submarginatum* Huber, *Bull. Herb. Boissier*, sér. 2, 6: 443 (Huber 1906).

*Sapium oligoneurum* K.Schum. & Pittier, *Contr. U.S. Natl. Herb.* 12: 168 (Schumann & Pittier 1908). — *Sapium biglandulosum* var. *oligoneurum* (K.Schum. & Pittier) Monach., *Bull. Torrey Bot. Club* 67 (9): 772 [Dec. 1940] (Monachino 1940).

*Sapium sulciferum* Pittier, *Contr. U.S. Natl. Herb.* 12: 169 (Pittier 1908). — *Sapium biglandulosum* var. *sulciferum* (Pittier) Monach., *Bull. Torrey Bot. Club* 67 (9): 772 [Dec. 1940] (Monachino 1940).

*Sapium fendleri* Hemsl., *Hooker's Icon. Pl.* 29 [ser. 4, 9]: sub t. 2888 [June 1909] (Hemsley 1909).

*Sapium muelleri* Hemsl., *Hooker's Icon. Pl.* 29 [ser. 4, 9]: t. 2884 [June 1909] (Hemsley 1909).

*Hippomane zeocca* L. ex B.D.Jacks., *Index Linn. Herb.*: 86 (Jackson 1912), *nom. nud.*

*Sapium endlicherianum* Klotzsch ex Pax, *Pflanzenr. [Engler]* IV.147. IV (Heft 52): 228 [13 Feb. 1912] (Pax 1912), *nom. nud. pro syn.*

*Sapium obtusatum* Klotzsch ex Pax, *Pflanzenr. [Engler]* IV.147.IV (Heft 52): 224 [13 Feb. 1912] (Pax 1912), *nom. nud. pro syn.*

*Sapium pohlianum* Klotzsch ex Pax, *Pflanzenr. [Engler]* IV.147.IV (Heft 52): 228 [13 Feb. 1912] (Pax 1912), *nom. nud. pro syn.*

*Sapium punctatum* Klotzsch ex Pax, *Pflanzenr. [Engler]* IV.147.IV (Heft 52): 223 [13 Feb. 1912] (Pax 1912), *nom. nud. pro syn.*

*Sapium pycnostachys* K.Schum. ex Pax, *Pflanzenr. [Engler]* IV.147. IV (Heft 52): 208 [13 Feb. 1912] (Pax 1912), *nom. nud. pro syn.*

*Sapium subserratum* Klotzsch ex Pax, *Pflanzenr. [Engler]* IV.147.IV (Heft 52): 226 [13 Feb. 1912] (Pax 1912), *nom. nud. pro syn.*

*Sapium caudatum* Pittier, *Contr. U.S. Natl. Herb.* 20 (3): 127 [18 June 1918] (Pittier 1918).

*Sapium giganteum* Pittier, *Contr. U.S. Natl. Herb.* 20 (3): 128 [18 June 1918] (Pittier 1918).

*Sapium albomarginatum* Pax & K.Hoffm., *Pflanzenr. [Engler]* IV.147. XVII (Heft 85): 203 [16 May 1924] (Pax & Hoffmann 1924).

*Sapium fragile* Pax & K.Hoffm., *Pflanzenr. [Engler]* IV.147.XVII (Heft 85): 202 [16 May 1924] (Pax & Hoffmann 1924).

*Sapium paranaense* Pax & K.Hoffm., *Pflanzenr. [Engler]* IV.147. XVII (Heft 85): 203 [16 May 1924] (Pax & Hoffmann 1924).

*Sapium guaricense* Pittier, *J. Wash. Acad. Sci.* 19: 355 (Pittier 1929).

*Sapium naiguatense* Pittier, *J. Wash. Acad. Sci.* 19: 355 (Pittier 1929).

*Sapium paucistamineum* Pittier, *J. Wash. Acad. Sci.* 19: 356 (Pittier 1929).

*Sapium integrifolium* Splitg. ex Lanj., *Euphorb. Surinam* 176 (Lanjouw 1931), *nom. nud. pro syn.*

*Sapium klotzschianum* var. *glaziovii* Lanj., *Euphorb. Surinam* 46 (Lanjouw 1931).

*Stillingia haematantha* Standl., *Ann. Missouri Bot. Gard.* 27: 314 (Standley 1940).
*Sapium haematospermum* Müll.Arg. var. saltense O'Donell & Lourteig, *Lilloa* 8: 586 (O'Donell & Lourteig 1942), "saltensis". — *Sapium saltense* (O'Donell & Lourteig) Jabl., *Phytologia* 14 (7): 451 [15 May 1967] (Jablonszky 1967).

*Sapium schippii* Croizat, *Amer. Midl. Naturalist* 29 (2): 477 [5 Apr. 1943] (Croizat 1943).

*Sapium contortum* Croizat, *J. Arnold Arbor*. 26 (2): 193 [16 Apr. 1945] (Croizat 1945).

*Sapium nitidum* Alain, *Contr. Ocas. Mus. Hist. Nat. Colegio De La Salle* 11: 8 (Alain 1952), *nom. illeg. hom., non* (Monach.) Lundell (1943) [synonym of *S. lateriflorum* Hemsl.]

*Sapium moaense* Alain, *Revista Soc. Cub. Bot.* 10: 27 (Alain 1953).

*Sapium ixiamasense* Jabl., *Phytologia* 14 (7): 450 [15 May 1967] (Jablonszky 1967).

*Sapium itzanum* Lundell, *Wrightia* 5 (4): 77 (Lundell 1975).

*Sapium izabalense* Lundell, *Wrightia* 5 (9): 346 (Lundell 1977).

*Sapium alainianum* P.T.Li, *Guihaia* 14 (2): 130 (Li 1994).

NOTES. — *Excoecaria biglandulosa* Müll.Arg., *Sapium biglandulosum* Müll.Arg. and *Stillingia biglandulosa* Baill. are based on the illegitimate *Hippomane biglandulosa* L. *S. biglandulosum* is itself illegitimate, because it includes the type of *H. glandulosa*.

VERNACULAR NAMES. — Pa: itey-avain, itey-avan • Ka: alekosin, kumakalan • Nt: meliki udu • Cr: bwa-dilèt • Br: burra-leiteira, murupita.

HERBARIUM DATA (FG). — 30 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier* 2255.

INVENTORY DATA (FG). — 1 tree, dbh = 28.5 cm.

[534] *Sapium paucinervium* Hemsl.

*Hooker's Icon. Pl.* 27 [ser. 4, 7]: t. 2648 ["1901" publ. May 1900] (Hemsley 1900).

*Sapium microdentatum* Lanj., *Euphorb. Surinam* 46 (Lanjouw 1931).

VERNACULAR NAMES. — Br: murupita.

HERBARIUM DATA (FG). — 30 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier* 2970.

INVENTORY DATA (FG). — 21 trees in 12 plots; Fmax = 1%; dbhinv = 43 cm.

Family GOUPIACEAE Miers
Genus *Goupia* Aubl.

[535] *Goupia glabra* Aubl.
(Fig. 24A)

*Hist. Pl. Guiane* 1: 296 [Jun.-Dec. 1775] (Aublet 1775). — *Glossopetalum glabrum* (Aubl.) J.F.Gmel., *Syst. Nat.*, ed. 13[bis], 2 (1): 510 [late Sep.-Nov. 1791] (Gmelin 1791).

*Goupia tomentosa* Aubl., *Hist. Pl. Guiane* 1: 296 [Jun.-Dec. 1775] (Aublet 1775). — *Glossopetalum tomentosum* (Aubl.) J.F.Gmel., *Syst. Nat.*, ed. 13[bis], 2 (1): 510 [late Sep.-Nov. 1791] (Gmelin 1791).

*Goupia paraensis* Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 6: 86 (Huber 1910).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: pasis • Ka: kubi'i, kupi'i • Te: patsitsi • Wp: pasisi • Wn: walima • Nt: kopi • Cr: bwa-kaka, goupi • Fr: bois caca, goupi • Br: cupiúba.

HERBARIUM DATA (FG). — 151 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JJR[P00778564] designated by Lanjouw & Uittien [1940: 151]).

INVENTORY DATA (FG). — 382 trees in 134 plots; Fmax = 26.2%; dbhinv = 158 cm.

Family HERNANDIACEAE Blume
Genus *Hernandia* Plum. ex L.

[536] *Hernandia guianensis* Aubl.
(Fig. 24B)

*Hist. Pl. Guiane* 2: 848 [Jun.-Dec. 1775] (Aublet 1775).

VERNACULAR NAMES. — Pa: maoksi-adudu • Ka: ayowo, tokolo enulu • Wp: wali wowo • Nt: popolika • Cr: bwa-amadou, bwa-bannan • Br: ventosa.

HERBARIUM DATA (FG). — 24 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000993866], G[G00355749]).

INVENTORY DATA (FG). — 11 trees censused, 1 in 1 plot; dbhinv = 57 cm.

Family HUMIRIACEAE A.Juss.
Genus *Humiria* J.St.-Hil.

[537] *Humiria balsamifera* Aubl.

*Hist. Pl. Guiane* 1: 564 [Jun.-Dec. 1775] (Aublet 1775), "Houmiri". — *Myrodendrum balsamiferum* (Aubl.) J.F.Gmel., *Syst. Nat.*, ed. 13[bis], 2 (1): 817 [late Sep.-Nov. 1791] (Gmelin 1791).

*Myrodendrum amplexicaule* Willd., *Sp. Pl.*, ed. 4 2 (2): 1171 [Dec. 1799] (Willdenow 1799), *nom. illeg. superfl.* (based on *Humiria balsamifera*). — *Humiria amplexicaulis* Mart. ex Urb., *Fl. Bras. [Martius]* 12 (2): 440 [1 Dec. 1877] (Urban 1877), "*Humirium amplexicaule*", *nom. nud. pro syn.*

*Humiria arenaria* Guill. ex Baill., *Adansonia* [Baillon] 1: 209 (Bail-lon 1860), "*Humirium arenarium*".

*Humiria balsamifera* f. attenuata Cuatrec., *Contr. U.S. Natl. Herb.* 35 (2): 97 [14 Apr. 1961] (Cuatrecasas 1961).

NOTE. — In order to limit the inflation of binomials, we have chosen not to list here the orthographic variants "*Humirium*" and "*Myrodendron*", as well as all binomial based on them.

VERNACULAR NAMES. — Pa: psum, umeg • Ka: meli • Wp: weli • Nt: aneisi udu • Cr: bwa-rouj • Fr: houmiri baumier • Br: turamira, umiri.

HERBARIUM DATA (FG). — 44 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000795994]).

INVENTORY DATA (FG). — 28 trees in 16 plots; Fmax < 1%; dbhinv = 144.2 cm.
Genus *Humiriastrum* (Urb.) Cuatrec.

[538] *Humiriastrum excelsum* (Ducke) Cuatrec.

Contr. U.S. Natl. Herb. 35 (2): 133 [14 Apr. 1961] (Cuatrecasas 1961). — *Sacoglottis excelsa* Ducke, *Arch. Jard. Bot. Rio de Janeiro* 3: 178 (Ducke 1922).

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 4066*.

INVENTORY DATA (FG). — 5 trees in 3 plots; Fmax < 1%; dbhinv = 63.3 cm.

[539] *Humiriastrum subcrenatum* (Benth.) Cuatrec.

Contr. U.S. Natl. Herb. 35 (2): 138 [14 Apr. 1961] (Cuatrecasas 1961). — *Humiria subcrenata* Benth., *London J. Bot.* 2: 374 (Bentham 1843), "*Humirium subcrenatum*". — *Sacoglottis subcrenata* (Benth.) Urb., *Sitzungsber. Ges. Naturf. Freunde Berlin* 5 (Urban 1878).

NOTE. — Known only from French Guiana.

VERNACULAR NAMES. — Pa: ašiu, ašiwa, tumau-atamwa • Nt: lebi kookoo.

HERBARIUM DATA (FG). — 21 collections at CAY. Sel. exs.: *J. Martin s.n.* (type K[K000407329]).

INVENTORY DATA (FG). — 112 trees in 53 plots; Fmax = 1%; dbhinv = 75.8 cm.

Genus *Sacoglottis* Mart.

[540] *Sacoglottis cydonioides* Cuatrec.
(Fig. 24C)

Contr. U.S. Natl. Herb. 35 (2): 183 [14 Apr. 1961] (Cuatrecasas 1961).

VERNACULAR NAMES. — Pa: ašiu, ašiwa, tumau-atamwa • Ka: pelulu • Wp: asiwa • Cr: bwa-kochon, bwa-maypouri.

HERBARIUM DATA (FG). — 30 collections at CAY. Sel. exs.: *D. Sabatier & J.-L. Smock 5782*.

INVENTORY DATA (FG). — 31 trees in 22 plots; Fmax = 1.1%; dbhinv = 73 cm.

[541] *Sacoglottis guianensis* Benth.

Hooker's J. Bot. Kew Gard. Misc. 5: 104 (Bentham 1853).

Sacoglottis amazonica Benth., Hooker's J. Bot. Kew Gard. Misc. 5: 104 (Bentham 1853), nom. illeg. hom., non Mart. (Martius 1827).

Sacoglottis guianensis f. dolichocarpa Ducke, Arch. Jard. Bot. Rio de Janeiro 3: 179 (Ducke 1922).

Sacoglottis guianensis var. maior Ducke, Arch. Inst. Biol. Veg. 4 (1): 27 [June 1938] (Ducke 1938).

Sacoglottis guianensis var. hispidula Cuatrec., Contr. U.S. Natl. Herb. 35 (2): 180 [14 Apr. 1961] (Cuatrecasas 1961).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: ašiu, ašiwa, tumau-atamwa • Ka: apesiya, yapopali • Nt: bofo udu, bofoo udu • Cr: bwa-kochon, bwa-maypouri • Br: achuá, uchirana.

HERBARIUM DATA (FG). — 22 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 5202*.

INVENTORY DATA (FG). — 182 trees in 72 plots; Fmax = 2.5%; dbhinv = 61.3 cm.

Genus *Schistostemon* (Urb.) Cuatrec.

[542] *Schistostemon dichotomus* (Urb.) Cuatrec.

Contr. U.S. Natl. Herb. 35 (2): 159 [14 Apr. 1961] (Cuatrecasas 1961). — *Sacoglottis dichotoma* Urb., *Fl. Bras. [Martius]* 12 (2): 446 [1 Dec. 1877] (Urban 1877), "*Saccoglottis*".

NOTE. — The Greek root “stemon” is masculine, hence *dichotomus*.

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: *C. Sastre et al.* 4125.

SIZE. — > 10 cm dbh (Cardoso *et al.* 2017).

[543] *Schistostemon sylvaticus* Sabatier

Proc. Kon. Ned. Akad. Wetensch. C 90 (2): 206 (Sabatier 1987).

NOTES. — Known only from French Guiana. The Greek root “*stemon*” is masculine, hence *sylvaticus*.

VERNACULAR NAMES. — Nt: bofo udu, bofoo udu.

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: *D. Sabatier 842* (holo-, P[P00077220]; iso-, P[not seen], U[U0008171]).

INVENTORY DATA (FG). — 17 trees in 9 plots; Fmax < 1%; dbhinv = 56 cm.

Genus *Vantanea* Aubl.

[544] *Vantanea guianensis* Aubl.

Hist. Pl. Guiane 1: 572 [Jun.-Dec. 1775] (Aublet 1775). — *Lemniscia guianensis* (Aubl.) J.F.Gmel., *Syst. Nat.*, ed. 13[bis], 2 (1): 817 [late Sep.-Nov. 1791] (Gmelin 1791). — *Lemniscia floribunda* Willd., *Sp. Pl.*, ed. 4 2 (2): 1172 [Dec. 1799] (Willdenow 1799), nom. illeg. *superfl.* (based on *Vantanea guianensis*).

VERNACULAR NAMES. — Pa: ašiu-kamwi • Te: tepotsi ātā, wila pikikit • Nt: bofo udu, bofoo udu • Br: uchirana.

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000795993], S[S-R-9886]).

INVENTORY DATA (FG). — 34 trees in 17 plots; Fmax = 2.3%; dbhinv = 73 cm.
<figure><img src="image_35.png" /><figcaption>FIG. 24. — Goupiaceae: **A**, *Goupia glabra* Aubl. Hernandiaceae: **B**, *Hernandia guianensis* Aubl. (M.-F. Prévost 3500). Humiriaceae: **C**, *Sacoglottis cydonioides* Cuatrec. (D. Sabatier & J.-L. Smock 5782); **D**, *Vantanea maculicarpa* Sabatier & J.Engel (D. Sabatier 5574). © D. Sabatier/IRD.</figcaption></figure>

[545] *Vantanea maculicarpa* Sabatier & J.Engel
(Fig. 24D)

*Phytotaxa* 338 (1): 130 [09 Feb. 2018] (Sabatier & Engel 2018).

NOTE. — Known only from French Guiana.

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *D. Sabatier* 5574 (holo-, CAY[CAY111685]; iso-, CAY[CAY109439], P[P01156374]).

INVENTORY DATA (FG). — 4 trees in 3 plots; Fmax < 1%; dbhinv = 66.5 cm.

[546] *Vantanea ovicarpa* Sabatier

*Brittonia* 54 (4): 233 ["2002" publ. 16 Apr. 2003] (Sabatier 2003).

NOTE. — Known only from French Guiana.

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: S.A. Mori & N.P. Smith 25047 (holo-, NY[00718019, 00718020]; iso-, CAY[CAY015333], P[P00710334], US[00810841]).

SIZE. — Up to 50 cm dbh (Sabatier 2002).
[547] *Vantanea parviflora* Lam.

J. Hist. Nat. 1: 145 (Lamarck 1792).

*Vantanea cupularis* Huber, Bol. Mus. Goeldi Hist. Nat. Ethnogr. 6: 83 (Huber 1910).

VERNACULAR NAMES. — Pa: ahavuiye-wašiune • Nt: adugwe • Br: macucá-murici, quebra-machado, uchirana.

HERBARIUM DATA (FG). — 32 collections at CAY. Sel. exs.: *J.B. Leblond s.n.* (holo-, P-LA?, not seen; iso-, G[G00368596]).

INVENTORY DATA (FG). — 87 trees in 49 plots; Fmax = 3.1 %; dbhinv = 88 cm.

Family HYPERICACEAE Juss.
Genus *Vismia* Vand.

[548] *Vismia cayennensis* (Jacq.) Pers.

Syn. Pl. [Persoon] 2 (1): 86 [Nov. 1806] (Persoon 1806). — *Hypericum cayennense* Jacq., Enum. Syst. Pl.: 28 [Aug.-Sep. 1760] (Jacquin 1760). — *Caopia cayennensis* (Jacq.) Kuntze, Revis. Gen. Pl. 1: 58 [5 Nov. 1891] (Kuntze 1891).

*Hypericum eugeniifolium* Willd. ex Spreng., Syst. Veg. [Sprengel] 3: 350 [Jan.-Mar. 1826] (Sprengel 1826), “eugeniaefolium”, nom. nud. pro syn.

*Vismia floribunda* Sprague, Trans. & Proc. Bot. Soc. Edinburgh 22 (4): 428 (Sprague 1905).

VERNACULAR NAMES. — Pa: suwimba, suwimpa • Ka: siwinyani, suwinyani enekan • Wp: suwilani • Wn: osi epīt • Nt: lebi pindya udu, pindya udu • Cr: bwa-dat • Br: lacre-branco.

HERBARIUM DATA (FG). — 108 collections at CAY. Sel. exs.: The type from French Guiana (not traced); *W.J. Hahn* 3534.

INVENTORY DATA (FG). — 3 trees in 2 plots; Fmax < 1%; dbhinv = 25.6 cm.

[549] *Vismia gracilis* Hieron.

*Bot. Jahrb. Syst.* 20 (3, Beibl. 49): 52 [9 Apr. 1895] (Hieronymus 1895). — *Caopia gracilis* Hieron., *Bot. Jahrb. Syst.* 20 (3, Beibl. 49): 52 [9 Apr. 1895] (Hieronymus 1895).

*Vismia glaziovii* Ruhland, *Bot. Jahrb. Syst.* 30 (Beibl. 67): 27 [12 Mar. 1901] (Ruhland 1901).

*Vismia amazonica* Ewan, *Contr. U.S. Natl. Herb.* 35 (5): 355 [5 June 1962] (Ewan 1962).

*Vismia buchtienii* Ewan, *Contr. U.S. Natl. Herb.* 35 (5): 353 [5 June 1962] (Ewan 1962).

NOTE. — *Vismia gracilis* Hieron. and *Caopia gracilis* Hieron. were published simultaneously, but before 1953. Although alternative, both names are valid.

VERNACULAR NAMES. — Pa: suwimba, suwimpa • Wp: suwilani • Cr: bwa-dat • Br: lacre, pau-lacre.

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: *D. Sabatier* 2317, dbh 10 cm.

[550] *Vismia guianensis* (Aubl.) Choisy

Prodr. Monogr. Hypéric.: 34 [9 Mar. 1821] (Choisy 1821), “guyan-
ensis”. — *Hypericum guianense* Aubl., Hist. Pl. Guiane 2: 784 [Jun.-
Dec. 1775] (Aublet 1775), “Guianense”, “Hipericum Guyannense” on
plate. — *Vismia guttifera* Pers. var. guianensis (Aubl.) Pers., Syn. Pl.
[Persoon] 2 (1): 86 [Nov. 1806] (Persoon 1806). — *Caopia guianensis*
(Aubl.) A.Lyons, Pl. Nam., ed. 2: 94 (Lyons 1907), “guaianensis”.

*Hypericum acuminatum* Lam., *Encycl. [J. Lamarck et al.]* 4 (1): 150 [9 Feb. 1797] (Lamarck 1797). — *Vismia acuminata* (Lam.) Pers., *Syn. Pl. [Persoon]* 2 (1): 86 [Nov. 1806] (Persoon 1806). — *Caopia acuminata* (Lam.) Kuntze, *Revis. Gen. Pl.* 1: 59 [5 Nov. 1891] (Kuntze 1891). — *Vismia guianensis* var. *acuminata* (Lam.) M.E.Berg, *Bol. Mus. Paraense Emílio Goeldi, N.S., Bot.* 40: 7 (Berg 1971).

*Vismia caparosa* Kunth, *Nova genera et species plantarum* [H.B.K.] 5: 182 [25 Feb. 1822] (Kunth 1822).

*Vismia ferruginea* Kunth, *Nova genera et species plantarum* [H.B.K.] 5: 183 [25 Feb. 1822] (Kunth 1822).

VERNACULAR NAMES. — Ka: siwinyani, suwinyani, tamunen su-
winyani • Wn: osi epīt • Nt: pindya udu, weti pindya udu • Cr:
bwa-dat • Fr: bois baptiste, bois dartre, bois de sang (fide Aublet
1775) • Br: lacre-vermelho, uchirana.

HERBARIUM DATA (FG). — 73 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000624909]).

INVENTORY DATA (FG). — 6 trees in 6 plots; Fmax = 5.8 %; dbhinv = 38.8 cm.

[551] *Vismia latifolia* (Aubl.) Choisy
(Fig. 25A)

Prodr. Monogr. Hypéric. 36 [9 Mar. 1821] (Choisy 1821). — *Hypericum latifolium* Aubl., Hist. Pl. Guiane 2: 787 [Jun.-Dec. 1775] (Aublet 1775). — *Caopia latifolia* (Aubl.) Kuntze, Revis. Gen. Pl. 1: 58 [5 Nov. 1891] (Kuntze 1891).

VERNACULAR NAMES. — Ka: saipyalala • Wp: suwilani pilā • Wn: osi epīt • Nt: lebi pindya udu, pindya udu • Cr: bwa-dat • Br: lacre, pau-lacre.

HERBARIUM DATA (FG). — 84 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier* 4822.

INVENTORY DATA (FG). — 12 trees in 5 plots; Fmax < 1%; dbhinv = 31.2 cm.

[552] *Vismia macrophylla* Kunth

*Nova genera et species plantarum* [H.B.K.] 5: 184 [25 Feb. 1822] (Kunth 1822).

*Hypericum reticulatum* Poir., *Encycl. [J. Lamarck et al.]* Suppl. 3: 694 [3 Sep. 1814) (Poiret 1814). — *Vismia reticulata* (Poir.) Choisy, *Prodr. Monogr. Hypéric.* 34 [9 Mar. 1821] (Choisy 1821).

*Vismia angusta* Miq., *Linnaea* 18: 27 [24-26 Oct. 1844] (Miquel 1844).

*Vismia macrophylla* var. *glabrescens* Hochr., *Annuaire Conserv. Jard. Bot. Genève* 21: 53 (Hochreutiner 1919).
VERNACULAR NAMES. — Pa: suwimba-purubumna, suwimpa-purubumna • Ka: saipyalala • Cr: bwa-dat • Br: lacre, pau-lacre.

HERBARIUM DATA (FG). — 23 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 4068*, 18 m × 15 cm.

[553] *Vismia ramuliflora* Miq.

Stirp. *Surinam. Select.*: 88 [“1850” publ. Mar. 1851] (Miquel 1851).

VERNACULAR NAMES. — Wn: pëmulimë.

HERBARIUM DATA (FG). — 43 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier 2810*.

INVENTORY DATA (FG). — 3 trees in 3 plots; $F_{max} < 1\%$; $dbh_{inv} = 32.8$ cm.

[554] *Vismia sandwithii* Ewan

Contr. U.S. Natl. Herb. 35 (5): 309 [5 June 1962] (Ewan 1962).

VERNACULAR NAMES. — Pa: suwimba-ahavukune, suwimpa-ahau-wukune • Wp: suwilani, suwilani wu • Wn: osi épit • Br: lacre-vermelho-do-mato.

HERBARIUM DATA (FG). — 38 collections at CAY. Sel. exs.: *M.-F. Prévost & P. Grenand 890*.

SIZE. — Up to 10 m tall (Pipoly *et al.* 1998).

[555] *Vismia sessilifolia* (Aubl.) Choisy

Prodr. [*A. P. de Candolle*] 1: 542 [mid Jan. 1824] (Choisy 1824). — *Hypericum sessilifolium* Aubl., Hist. Pl. Guiane 2: 787 [Jun.-Dec. 1775] (Aublet 1775). — *Caopia sessilifolia* (Aubl.) Kuntze, Revis. Gen. Pl. 1: 59 [5 Nov. 1891] (Kuntze 1891). — *Vismia cayennensis* var. sessilifolia (Aubl.) M.E.Berg, *Acta Amazonica* 4 (3): 16 (Berg 1974).

*Hypericum rufescens* Lam., *Encycl. [J. Lamarck et al.]* 4 (1): 150 [9 Feb. 1797] (Lamarck 1797). — *Vismia rufescens* (Lam.) Pers., *Syn. Pl. [Persoon]* 2 (1): 86 [Nov. 1806] (Persoon 1806).

VERNACULAR NAMES. — Pa: suwimba-kamwi, suwimpa-kamwi • Ka: talukuwa epi • Wp: kuya là • Fr: bois baptiste, bois dartre, bois de sang (*fide* Aublet 1775).

HERBARIUM DATA (FG). — 63 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000624924], G-DC[G00209682]).

INVENTORY DATA (FG). — 21 trees in 14 plots; $F_{max} = 3.8\%$; $dbh_{inv} = 33.7$ cm.

Family IXONANTHACEAE Planch. ex Miq.
Genus *Cyrillopsis* Kuhlm.

[556] *Cyrillopsis paraensis* Kuhlm.
(Fig. 25B)

Arch. Jard. Bot. Rio de Janeiro 4: 357 (Kuhlmann 1925).

HERBARIUM DATA (FG). — 13 collections at CAY. Sel. exs.: *J.-F. Molino 3429*.

INVENTORY DATA (FG). — 53 trees in 20 plots; $F_{max} = 1.3\%$; $dbh_{inv} = 63.4$ cm.

Family LACISTEMATACEAE Mart.
Genus *Lacistema* Sw.

[557] *Lacistema aggregatum* (P.J.Bergius) Rusby

Bull. New York Bot. Gard. 4 (14): 447 [7 Dec. 1907] (Rusby 1907). — *Piper aggregatum* P.J.Bergius, *Acta Helv. Phys.-Math. 7*: 131 (Bergius 1772). — *Lacistema myricoides* Sw., *Prodr. [Swartz]* 12 [20 Jun.-29 July 1788] (Swartz 1788), *nom. illeg. superfl.* (based on *Piper aggregatum*).

*Nematospermum laevigatum* Rich., *Actes Soc. Hist. Nat. Paris* 1: 105 [Oct. 1792] (Richard 1792).

*Synzyganthera purpurea* Ruiz & Pav., *Fl. Peruv. Prodr.*: 137 [early Oct. 1794] (Ruiz & Pavón 1794). — *Didymandra purpurea* (Ruiz & Pav.) Willd., *Sp. Pl.*, ed. 4 4 (2): 971 [Apr. 1806] (Willdenow 1806), *nom. illeg. superfl.* (genus name superfluous, based on the type of *Synzyganthera Ruiz & Pav.*). — *Lacistema purpureum* (Ruiz & Pav.) Á.DC., *Prodr. [A. P. de Candolle]* 16 (2.2): 593 [mid July 1868] (Candolle 1868).

*Piper fasciculare* Rudge, *Pl. Guian. [Rudge]* 1 (1): t. 4 [Apr. 1805] (Rudge 1805).

*Lacistema floribundum* Miq., *Linnaea* 18: 24 [24-26 Oct. 1844] (Miquel 1844), “floribunda”.

*Lacistema elongatum* Schnizl., *Fl. Bras. [Martius]* 4 (1): 282 [28 Feb. 1857] (Schnizlein 1857).

*Lacistema angustum* Schnizl., *Fl. Bras. [Martius]* 4 (1): 283 [28 Feb. 1857] (Schnizlein 1857).

*Lacistema recurvum* Schnizl., *Fl. Bras. [Martius]* 4 (1): 283 [28 Feb. 1857] (Schnizlein 1857).

*Lacistema myricoides* var. *stipitatum* A.DC., *Prodr. [A. P. de Candolle]* 16 (2.2): 592 [mid July 1868] (Candolle 1868).

*Lacistema coriaceum* A.DC., *Prodr. [A. P. de Candolle]* 16 (2.2): 593 [mid July 1868] (Candolle 1868).

*Lacistema poeppigii* A.DC., *Prodr. [A. P. de Candolle]* 16 (2.2): 593 [mid July 1868] (Candolle 1868).

*Lacistema bolivianum* Gand., *Bull. Soc. Bot. France* 66 (7): 288 [“1919” publ. 1920] (Gandoger 1920).

*Lacistema guyanense* Gand., *Bull. Soc. Bot. France* 66 (7): 288 [“1919” publ. 1920] (Gandoger 1920).

*Lacistema rosidiscum* J.F.Macbr., *Candollea* 5: 392 (Macbride 1934).

*Lacistema curtum* J.F.Macbr., *Candollea* 5: 393 (Macbride 1934).

*Lacistema orinocense* Baehni, *Candollea* 8: 43 (Baehni 1940).

*Lacistema weberbaueri* Baehni, *Candollea* 8: 46 (Baehni 1940).

*Lacistema aggregatum* var. *elongatum* Maguire, *Bull. Torrey Bot. Club* 75 (3): 293 [May-June 1948] (Maguire 1948).
*Lacistema occidentale* Cuatrec., *Fieldiana*, Bot. 27 (1): 93 (Cuatrecas 1950).

*Lacistema pacificum* Cuatrec., *Fieldiana*, Bot. 27 (1): 93 (Cuatrecas 1950).

NOTE. — The type of *Didymandra* Willld. is that of *Synzyganthera* Ruiz & Pav., thus both *Didymandra* and *D. purpurea* (Ruiz & Pav.) Willld. are superfluous.

VERNACULAR NAMES. — Pa: agagut, ararut • Wp: inámu sī, inámusī wila, tatumila sili, yape'a pitā sili.

HERBARIUM DATA (FG). — 57 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier* 2419.

INVENTORY DATA (FG). — 41 trees in 18 plots; $F_{max} = 1\%$; $dbh_{inv} = 24.6$ cm.

[558] *Lacistema grandifolium* Schnizl.

Fl. Bras. [Martius] 4 (1): 284 [28 Feb. 1857] (Schnizlein 1857).

VERNACULAR NAMES. — Pa: tahuma • Wp: mulei sili.

HERBARIUM DATA (FG). — 56 collections at CAY. Sel. exs.: *P.A. Poiteau s.n.* (original material G[G00165292, G00202465]).

INVENTORY DATA (FG). — 41 trees in 28 plots; $F_{max} < 1\%$; $dbh_{inv} = 18.5$ cm.

[559] *Lacistema polystachyum* Schnizl.
(Fig. 25C)

Fl. Bras. [Martius] 4 (1): 284 [28 Feb. 1857] (Schnizlein 1857).

HERBARIUM DATA (FG). — 19 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier* 2367.

INVENTORY DATA (FG). — 1 tree, dbh = 11.6 cm.

[560] *Lacistema pubescens* Mart.

*Nova genera et species plantarum* [Martius] 1 (4): 155 ["1824" publ. Jan.-Mar. 1826] (Martius 1826).

*Lacistema ellipticum* Schnizl., *Fl. Bras. [Martius]* 4 (1): 285 [28 Feb. 1857] (Schnizlein 1857).

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier* 4891.

INVENTORY DATA (FG). — 2 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 15.8$ cm.

Family LAMIACEAE Martinov
Genus *Aegiphila* Jacq.

[561] *Aegiphila integrifolia* (Jacq.) B.D.Jacks.

*Index Kew.* 1 (1): 46 [6 Sep. 1893] (Jackson 1893). — *Callicarpa integrifolia* Jacq., * Enum. Syst. Pl.*: 12 [Aug.-Sep. 1760] (Jacquin 1760).

Manabea arborescens Aubl., *Hist. Pl. Guiane* 1: 64 [Jun.-Dec. 1775] (Aublet 1775). — *Aegiphila arborescens* (Aubl.) J.F.Gmel., *Syst. Nat.*, ed. 13[bis], 2 (1): 259 [late Sep.-Nov. 1791] (Gmelin 1791).

Callicarpa globiflora Ruiz & Pav., *Flora Peruviana* 1: 49 (Ruiz & Pavón 1798).

Callicarpa discolor Willld. ex Steud., *Nomencl. Bot.* [Steudel], ed. 2, 1: 257 (Steudel 1840), *nom. nud. pro syn.*

*Aegiphila arborescens* var. *longiflora* Schauer, *Prodr.* [A. P. de Candolle] 11: 649 [25 Nov. 1847] (Schauer 1847).

*Aegiphila arborescens* var. *breviflora* Schauer, *Prodr.* [A. P. de Candolle] 11: 649 [25 Nov. 1847] (Schauer 1847).

*Aegiphila guianensis* Moldenke, *Repert. Spec. Nov. Regni Veg.* 33: 125 (Moldenke 1933).

*Aegiphila parviflora* Moldenke, *Repert. Spec. Nov. Regni Veg.* 33: 135 (Moldenke 1933).

*Aegiphila arborea* Spruce ex Moldenke, *Phytologia* 1 (5): 206 [1 Mar. 1937] (Moldenke 1937), *nom. nud. pro syn.*

*Aegiphila amazonica* Moldenke, *Phytologia* 1 (11): 378 [6 Sept 1940] (Moldenke 1940).

*Aegiphila cowanii* Moldenke, *Phytologia* 27 (1): 63 [Oct. 1973] (Moldenke 1973).

*Aegiphila integrifolia* var. *lopez-palacii* Moldenke, *Phytologia* 36: 53 [June 1977] (Moldenke 1977).

VERNACULAR NAMES. — Pa: aigvan-kamwi • Cr: bwa-tabak, féy-tabak.

HERBARIUM DATA (FG). — 19 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lectotype of *Manabea arborescens* P-JJR[P00777561] designated by Lanjouw & Uittien [1940: 152]).

SIZE. — Venezuela. *L. Marcano-Berti* 244-981 (MO), 12 m.

[562] *Aegiphila villosa* (Aubl.) J.F.Gmel.

*Syst. Nat.*, ed. 13[bis], 2 (1): 259 [late Sep.-Nov. 1791] (Gmelin 1791). — *Manabea villosa* Aubl., *Hist. Pl. Guiane* 1: 61 [Jun.-Dec. 1775] (Aublet 1775).

VERNACULAR NAMES. — Pa: aigvan, audika-retni-seine • Wp: a'i makule • Cr: bwa-tabak, féy-tabak • Br: camaá, cambara.

HERBARIUM DATA (FG). — 52 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000598566]); *R.A.A. Olde-man B-2398*, 11 m × 25 cm

Genus *Vitex* L.

[563] *Vitex compressa* Turcz.

*Bull. Soc. Imp. Naturalistes Moscou* 36 (3): 224 (Turczaninow 1863).

*Vitex brittoniana* Moldenke, *Torreya* 33: 67 (Moldenke 1933).

*Vitex compressa* f. *angustifolia* Moldenke, *Phytologia* 47 (1): 17 [13 Nov. 1980] (Moldenke 1980).

VERNACULAR NAMES. — Ka: kwa'iyan, kwailan • Nt: pakila udu.

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: *M. Fleury* 567.

SIZE. — Up to 30 m tall (Jansen-Jacobs 1988).
<figure><img src="image_36.png" /></figure>

FIG. 25. — Hypericaceae: **A, Vismia latifolia** (Aubl.) Choisy. Ixonanthaceae: **B, Cyrillopsis paraensis** Kuhl.m. (J.-F. Molino 3429). Lacistemataceae: **C, Lacistema poly-stachyum** Schnizl. (J.-F. Molino & D. Sabatier 2367). Lamiaceae: **D, Vitex guianensis** Moldenke (D. Sabatier 2332). A, D, © D. Sabatier/IRD; B, C, © J.-F. Molino/IRD.

[564] *Vitex guianensis* Moldenke
(Fig. 25D)

Phytologia 1 (15): 486 [Jan. 1941] (Moldenke 1941).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *D. Sabatier 1220*.

INVENTORY DATA (FG). — 19 trees in 13 plots; Fmax < 1%; dbhinv = 86.6 cm.

[565] *Vitex orinocensis* Kunth var. *multiflora* (Miq.) Huber

Bol. Mus. Goeldi Hist. Nat. Ethnogr. 5 (1): 217 (Huber 1908). — *Vitex multiflora* Miq., *Linnaea* 18: 739 [“1844” publ. Aug.-Oct. 1845] (Miquel 1845).

HERBARIUM DATA (FG). — A single collection, M.-F. Prévost 1460.

SIZE. — Up to 20 m tall (Jansen-Jacobs 1988).

[566] *Vitex stahelii* Moldenke

Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., Sect. 2, 30: 310 (Moldenke 1940).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Wp: tayi lā.

HERBARIUM DATA (FG). — 5 collections at CAY. Sel. exs.: *F. Billiet & B. Jadin 1431*.

SIZE. — Up to 40 m tall (Jansen-Jacobs 1988).

[567] *Vitex triflora* Vahl

Eclog. Amer. 2: 49 (Vahl 1798).

Pyrostoma ternatum G.Mey., *Prim. Fl. Esseq.* 220 [Nov. 1818] (Meyer 1818), “ternata”.
*Casarettoa diversifolia* Walp., *Repert. Bot. Syst.* [Walpers] 4 (1): 92 [25-28 June 1845] (Walpers 1847), *nom. illeg. superfl.* (based on *Vitex triflora*).

*Macrostegia ruiziana* Nees, *Prodr.* [A. P. de Candolle] 11: 218 [25 Nov. 1847] (Nees 1847).

*Ruellia macrocalyx* Ruiz ex Nees, *Prodr.* [A. P. de Candolle] 11: 218 [25 Nov. 1847] (Nees 1847), *nom. nud. pro syn.*

*Vitex triflora* var. *angustiloba* Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (1): 215 (Huber 1908).

*Vitex triflora* var. *coriacea* Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (1): 215 (Huber 1908).

*Vitex triflora* var. *floribunda* Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (1): 215 (Huber 1908).

*Vitex triflora* var. *kraatzii* Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (1): 216 (Huber 1908).

*Vitex triflora* var. *tenuifolia* Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (1): 214 (Huber 1908).

*Vitex triflora* var. *quinquefoliolata* Moldenke, *Phytologia* 1 (2): 104 [4 Aug. 1934] (Moldenke 1934). — *Vitex triflora* f. *quinquefoliolata* (Moldenke) Moldenke, *Phytologia* 44 (6): 384 [11 Dec. 1979] (Moldenke 1979).

*Vitex triflora* var. *hirsuta* Moldenke, *Phytologia* 23 (3): 315 [31 May 1972] (Moldenke 1972).

**VERNACULAR NAMES.** — Pa: kwairu-kamwi • Wp: tayi lā • Wn: wilimë • Nt: madyo uman wiwii, madyoma uwii • Br: pau-d'arcorana, tarumā.

**HERBARIUM DATA (FG).** — 51 collections at CAY. Sel. exs.: *J.P.B. von Rohr s.n.* (original material B, B-W 11701-02 0).

**INVENTORY DATA (FG).** — 30 trees in 28 plots; $F_{max} < 1\%$; $dbh_{inv} = 17$ cm.

Family LAURACEAE Juss.
Genus *Aiouea* Aubl.

[568] *Aiouea guianensis* Aubl.
(Fig. 26A)

*Hist. Pl. Guiane* 1: 311 [Jun.-Dec. 1775] (Aublet 1775). — *Laurus hexandra* Willd., *Sp. Pl.*, ed. 4 2 (1): 482 [Mar. 1799] (Willdenow 1799), *nom. illeg. superfl.* (based on *Aiouea guianensis*). — *Douglasia laurina* Sm., *Cycl. [Rees]* 12: (Douglassia no. 1) [14 Feb. 1809] (Smith 1809), *nom. illeg. superfl.* (based on *Aiouea guianensis*).

*Laurus difformis* Rich., *Actes Soc. Hist. Nat. Paris* 1: 108 [Oct. 1792] (Richard 1792).

*Aiouea tenella* Nees, *Linnaea* 21: 512 (Nees 1848).

*Aiouea schomburgkii* Meisn., *Prodr.* [A. P. de Candolle] 15 (1): 82 [May 1864] (Meisner 1864).

*Aiouea rubra* A.C.Sm., *Phytologia* 1 (3): 115 [21 Jan. 1935] (Smith 1935).

*Aiouea demerarensis* Kosterm., *Recueil Trav. Bot. Néerl.* 35: 77 (Kostermans 1938).

**NOTES.** — *L.C. Richard s.n.* (B-W, B-W 07804-00 0; LE[LE00012694; P[P02004886]) was used by Willdenow (1799: 482) to illustrate his *Laurus hexandra*. It has a label with exactly the same description as in the protologue ("*Laurus hexandra foliis lanceolatis, racemis axillaribus, floribus hexandris. Sp. pl. 2 p. 482*"). Nonetheless, this is a specimen of *Aniba guianensis* Aubl., not *Aiouea guianensis* Aubl. (see notes under *Aniba guianensis*).

**VERNACULAR NAMES.** — Pa: panauna-puvemna.

**HERBARIUM DATA (FG).** — 33 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material P[P00128374, P00662841]).

**INVENTORY DATA (FG).** — 7 trees in 6 plots; $F_{max} < 1\%$; $dbh_{inv} = 46.6$ cm.

[569] *Aiouea impressa* (Meisn.) Kosterm.

*Recueil Trav. Bot. Néerl.* 35: 75 (Kostermans 1938). — *Phoebe impressa* Meisn., *Prodr.* [A. P. de Candolle] 15 (1): 33 [May 1864] (Meisner 1864). — *Endlicheria impressa* (Meisn.) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 132 (Mez 1889). — *Cinnamomum impressum* (Meisn.) Kosterm., *Reinwardtia* 10 (5): 444 (Kostermans 1988).

*Aiouea minutiflora* Coe-Teix., *Bol. Mus. Paraense Emilio Goeldi, N.S., Bot.* 34: 1 (Coe-Teixeira 1970).

**HERBARIUM DATA (FG).** — 2 collections at CAY. Sel. exs.: *J.-B. Patris s.n.* "Cayenne 31" (original material G[G00368709], L[L0035523]).

**SIZE.** — Up to 20 m tall (Renner 1982).

[570] *Aiouea longipetiolata* van der Werff

*Novon* 4 (1): 61 [spring 1994] (van der Werff 1994).

**NOTE.** — Known only from the Guiana Shield.

**VERNACULAR NAMES.** — Pa: panauna-kamwi • Wp: yakami ka'a, yakami'i.

**HERBARIUM DATA (FG).** — 28 collections at CAY. Sel. exs.: *S.A. Mori et al.* 20758 (holo-, MO[MO-247078]; iso-, CAY[CAY027837], NY[00022599], P[P00745701], U[U0002590], US[00754392]).

**INVENTORY DATA (FG).** — 26 trees in 16 plots; $F_{max} = 1\%$; $dbh_{inv} = 55.4$ cm.

[571] *Aiouea montana* (Sw.) R.Rohde

*Taxon* 66 (5): 1102 [epubl. 24 Oct. 2017] (Rohde 2017). — *Laurus montana* Sw., *Prodr.* [Swartz] 65 [20 Jun.-29 July 1788] (Swartz 1788). — *Cinnamomum montanum* (Sw.) J.Presl, *Přír. Rostlin* 2: 36 (Presl 1825). — *Persea montana* (Sw.) Spreng., *Syst. Veg.* [*Sprengel*] 2: 268 [Jan.-May 1825] (Sprengel 1825). — *Phoebe montana* (Sw.) Griseb., *Pl. Wright.* [*Grisebach*] 1: 187 [Dec. 1860] (Grisebach 1860).

*Laurus triplinervis* Ruiz & Pav., *Flora Peruviana* 4: t. 363 [1804-1830] (Ruiz & Pavón 1804). — *Phoebe peruviana* Meisn., *Prodr.* [A. P. de Candolle] 15 (1): 32 [May 1864] (Meisner 1864), *nom. illeg. superfl.* (based on *Laurus triplinervis*). — *Phoebe triplinervis* (Ruiz & Pav.) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 211 (Mez 1889), *nom. illeg. hom.*, *non Griseb.* (Grisebach 1860) [synonym of *Cinnamomum grisebachii* Lorea-Hern.]. — *Cinnamomum trip-***
*linerve* (Ruiz & Pav.) Kosterm., *Reinwardtia* 6 (1): 24 (Kostermans 1961), “*triplinervis*”.

*Persea cinnamomifolia* Kunth, *Nova genera et species plantarum* [H.B.K.] 2: 160 [8 Dec. 1817] (Kunth 1818). — *Phoebe cinnamomifolia* (Kunth) Nees, *Linnaea* 21: 488 (Nees 1848). — *Phoebe granatensis* Meisn., *Prodr. [A. P. de Candolle]* 15 (1): 32 [May 1864] (Meisner 1864), *nom. illeg. superfl.* (based on *Persea cinnamomifolia*). — *Cinnamomum cinnamomifolium* (Kunth) Kosterm., *Reinwardtia* 6 (1): 20 (Kostermans 1961).

*Phoebe elongata* Nees, *Syst. Laur.*: 116 [30 Oct.-5 Nov. 1836] (Nees 1836). — *Laurus elongata* Vahl ex Nees, *Syst. Laur.*: 116 [30 Oct.-5 Nov. 1836] (Nees 1836), *nom. nud. pro syn.* — *Cinnamomum elongatum* (Nees) Kosterm., *Reinwardtia* 6 (1): 21 (Kostermans 1961), *nom. illeg. hom.*, *non Saporta* (1889) *nec Usnadze* (1949).

*Phoebe maynensis* Nees, *Syst. Laur.*: 118 [30 Oct.-5 Nov. 1836] (Nees 1836). — *Phoebe peruviana* var. *glabriflora* Meisn., *Prodr. [A. P. de Candolle]* 15 (1): 32 [May 1864] (Meisner 1864), *nom. illeg. superfl.* (based on *Phoebe maynensis*). — *Cinnamomum maynense* (Nees) Kosterm., *Reinwardtia* 6 (1): 22 (Kostermans 1961).

*Phoebe cubensis* Nees, *Syst. Laur.*: 120 [30 Oct.-5 Nov. 1836] (Nees 1836). — *Phoebe antillana* Meisn. var. *cubensis* (Nees) Meisn., *Prodr. [A. P. de Candolle]* 15 (1): 31 [May 1864] (Meisner 1864). — *Phoebe triplinervis* var. *cubensis* (Nees) C.Wright, *Anales Acad. Ci. Med. Habana* 7: 347 [15 Nov. 1870] (Wright 1870). — *Cinnamomum cubense* (Nees) Kosterm., *Reinwardtia* 6 (1): 21 (Kostermans 1961), “*cubensis*”.

*Phoebe valenzuelana* A.Rich., *Hist. Fís. Cuba, Bot.* 11: 185 (Richard 1850). — *Persea triplinervis* (Griseb.) M.Gómez var. *valenzuelana* (A.Rich.) M.Gómez, *Dicc. Bot. Nombres Vulg. Cub. Puerto-Riq.*: 25 (Gómez 1889).

*Oreodaphne alba* A.Rich., *Hist. Fís. Cuba, Bot.* 11: 189 (Richard 1850).

*Phoebe mexicana* Meisn., *Prodr. [A. P. de Candolle]* 15 (1): 31 [May 1864] (Meisner 1864). — *Persea mexicana* (Meisn.) Hemsl., *Biol. Cent.-Amer., Bot.* 3 (14): 72 [Oct. 1882] (Hemsley 1882). — *Cinnamomum mexicanum* (Meisn.) Kosterm., *Reinwardtia* 6 (1): 22 (Kostermans 1961).

*Phoebe poeppigii* Meisn., *Prodr. [A. P. de Candolle]* 15 (1): 33 [May 1864] (Meisner 1864).

*Phoebe brasiliensis* Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 198 (Mez 1889).

*Phoebe mexicana* var. *bourgeauana* Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 214 (Mez 1889). — *Cinnamomum brasiliense* (Mez) Kosterm., *Reinwardtia* 6 (1): 20 (Kostermans 1961), “*brasiliensis*”.

*Phoebe heterotepala* Mez, *Repert. Spec. Nov. Regni Veg.* 3: 67 (Mez 1906). — *Cinnamomum heterotepalum* (Mez) Kosterm., *Reinwardtia* 6 (1): 21 (Kostermans 1961).

*Phoebe paraguariensis* Hassl., *Annuaire Conserv. Jard. Bot. Genève* 21: 79 (Hassler 1919). — *Cinnamomum paraguariense* (Hassl.) Kosterm., *Reinwardtia* 6 (1): 22 (Kostermans 1961), “*paraguariensis*”.

*Ocotea flavescens* Rusby, *Descr. S. Amer. Pl.*: 20 [20 Dec. 1920] (Rusby 1920).

*Phoebe pichisensis* A.C.Sm., *Bull. Torrey Bot. Club* 58 (2): 103 [Feb. 1931] (Smith 1931). — *Cinnamomum pichisense* (A.C.Sm.) Kosterm., *Reinwardtia* 6 (1): 23 (Kostermans 1961), “*pichisensis*”.

*Phoebe johnstonii* C.K.Allen, *J. Arnold Arbor.* 26 (4): 433 [15 Oct. 1945] (Allen 1945). — *Cinnamomum johnstonii* (C.K.Allen) Kosterm., *Reinwardtia* 6 (1): 21 (Kostermans 1961).

*Cinnamomum chana* Vattimo-Gil, *Arch. Jard. Bot. Rio de Janeiro* 17: 223 (Vattimo-Gil 1962).

*Cinnamomum australe* Vattimo-Gil, *Arch. Jard. Bot. Rio de Janeiro* 17: 224 (Vattimo-Gil 1962).

*Cinnamomum xinguense* Vattimo-Gil, *Arch. Jard. Bot. Rio de Janeiro* 17: 224 (Vattimo-Gil 1962).

*Phoebe filamentosa* C.K.Allen, *Mem. New York Bot. Gard.* 15: 69 (Allen 1966). — *Cinnamomum filamentosum* (C.K.Allen) Kosterm., *Reinwardtia* 7 (5): 461 (Kostermans 1969).

*Cinnamomum portosecurianum* Vattimo-Gil, *Anais* 15 *Congr. Soc. Bot. Brasil.* 170 (Vattimo-Gil 1967).

*Phoebe pickellii* Coe-Teix., Hoehnea 1: 187 (Coe-Teixeira 1971), “*pickelli*”. — *Cinnamomum pickellii* (Coe-Teix.) Kosterm., *Reinwardtia* 10 (5): 448 (Kostermans 1988).

*Phoebe fruticosa* Lundell, *Wrightia* 5 (9): 342 (Lundell 1977). — *Cinnamomum fruticosum* (Lundell) Kosterm., *Reinwardtia* 10 (5): 443 (Kostermans 1988).

*Cinnamomum peruvianum* Kosterm., *Reinwardtia* 10 (5): 448 (Kostermans 1988).

*Cinnamomum phoebe* Doweld, *Phytotaxa* 243 (2): 191 [epubl. 12 Jan. 2016] (Doweld 2016).

*HERBARIUM DATA* (FG). — 14 collections at CAY. Sel. exs.: S.A. *Mori* et al. 15024.

*INVENTORY DATA* (FG). — 1 tree, dbh = 15.4 cm.

[572] *Aiouea opaca* van der Werff

Brittonia 47 (4): 372 (van der Werff 1995).

NOTE. — Known only from French Guiana.

*HERBARIUM DATA* (FG). — 4 collections at CAY. Sel. exs.: S.A. *Mori* et al. 20928 (holo., MO[MO-247076]; iso., CAY[CAY027842], NY[00214355], P[P00745702], U[U0002592], US[00478967]).

*INVENTORY DATA* (FG). — 16 trees in 5 plots; $F_{max} < 1\%$; $dbh_{inv} = 37.7$ cm.

Genus *Aniba* Aubl.

[573] *Aniba affinis* (Meisn.) Mez

*Jahrb. Königl. Bot. Gart. Berlin* 5: 63 (Mez 1889). — *Aydendron affine* Meisn., *Prodr. [A. P. de Candolle]* 15 (1): 92 [May 1864] (Meisner 1864).

*HERBARIUM DATA* (FG). — A single collection, *J.-J. de Granville* et al. 10796.

SIZE. — Up to 15 m tall (Kubitzki 1982).
<figure><img src="image_37.png" /><figcaption>FIG. 26. — Lauraceae: **A**, *Aiouea guianensis* Aubl. (J.-F. Molino et al. 3371); **B**, *Aniba megaphylla* Mez (J.-F. Molino & D. Sabatier 2834); **C**, *Aniba williamsii* O.C.Schmidt (D. Sabatier & E. Fonty 5662); **D-F**, *Nectandra matogrossensis* Coe-Teix. (J.-F. Molino et al. 3406). A, B, D-F, © J.-F. Molino/IRD; C, © D. Sabatier/IRD.</figcaption></figure>

[574] *Aniba citrifolia* (Nees) Mez

Jahrb. Königl. Bot. Gart. Berlin 5: 74 (Mez 1889). — *Aydendron citrifolium* Nees, Syst. Laur. 257 [30 Oct.-5 Nov. 1836] (Nees 1836).

*Aydendron trinitatis* Meisn., Prodr. [A. P. de Candolle] 15 (1): 91 [May 1864] (Meisner 1864). — *Aniba trinitatis* (Meisn.) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 68 (Mez 1889).

*Aniba muelleriana* Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 72 (Mez 1889), “Mülleriana”.

*Cryptocarya pachycarpa* Gleason, *Bull. Torrey Bot. Club* 54 (8): 607 [Nov. 1927] (Gleason 1927).

*Aniba opaca* A.C.Sm., *Bull. Torrey Bot. Club* 61 (4): 191 [Apr. 1934] (Smith 1934).

**VERNACULAR NAMES.** — Pa: kudagman, kudagman-purubumna
• Te: uwahi pitāg • Wp: āyū'i tawa, iwa'e • Nt: bamba apisi • Cr:
sèd • Fr: cèdre.

**HERBARIUM DATA (FG).** — 42 collections at CAY. Sel. exs.: *FM.R. Leprieur 224* (original material of *Aniba muelleriana*: G[G00368670], P[P00745591, P00745592]).

**INVENTORY DATA (FG).** — 33 trees in 27 plots; $F_{max} < 1\%$; $dbh_{inv} = 33.4$ cm.

[575] *Aniba guianensis* Aubl.

*Hist. Pl. Guiane* 1: 327 [Jun.-Dec. 1775] (Aublet 1775). — *Cedrota guianensis* (Aubl.) Raeusch., *Nomencl. Bot. [Raeusch.]* ed. 3: 111 (Raeuschel 1797). — *Cedrota longifolia* Willd., *Sp. Pl.*, ed. 4 2 (1): 338 [Mar. 1799] (Willdenow 1799), *nom. illeg. superfl.* (based on *Aniba guianensis*).

*Aniba salicifolia* Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 71 (Mez 1889).

*Aniba tessmannii* O.C.Schmidt, *Notizbl. Bot. Gart. Berlin-Dahlem* 10: 226 [30 Mar. 1928] (Schmidt 1928).

**NOTES.** — Excluded synonyms: *Aydendron salicifolium* (Sw.) Nees [syn. nov. of *Licaria parvifolia* (Lam.) Kosterm.] and its basionym *Laurus salicifolia* Sw. Kubitzki (1982: 45) placed “*Aydendron salicifolium* Nees” and “*Aniba salicifolia* (Nees) Mez” in synonymy under *Aniba guianensis* Aubl. Actually, although Nees misinterpreted *L. salicifolia* (synonym of *Licaria parvifolia*, a Caribbean taxon), and based his description of *Aydendron salicifolium* on a French Guianan collection of *Aniba guianensis* (*L.C. Richard s.n.* [B-W, B-W07804-000; LE[LE00012694; P[P02004886], see also notes under *Aiouea guianensis*), he explicitly referred to *L. salicifolia* and to its type. Therefore, he effectively transferred *L. salicifolia* into *Aydendron*. Later, when transferring *Aydendron salicifolium* into
*Aniba*, Mez (1889: 71) excluded *L. salicifolia* from synonymy. He thereby created a new name for the Guianan species.

VERNACULAR NAMES. — Wp: áyū'i sili, áyū'i tawa • Cr: sèd-kannel.

HERBARIUM DATA (FG). — 25 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000617483]).

INVENTORY DATA (FG). — 39 trees in 30 plots; Fmax < 1%; dbhinv = 31.8 cm.

[576] *Aniba hostmanniana* (Nees) Mez

Jahrb. Königl. Bot. Gart. Berlin 5: 67 (Mez 1889). — *Aydendron hostmannianum* Nees, Linnaea 21: 499 (Nees 1848).

*Aniba gigantifolia* O.C.Schmidt, Notizbl. Bot. Gart. Berlin-Dahlem 10: 225 [30 Mar. 1928] (Schmidt 1928).

*Aniba Pittieri* O.C.Schmidt, Repert. Spec. Nov. Regni Veg. 31: 168 (Schmidt 1933).

VERNACULAR NAMES. — Ka: apotono ali, siduwapali, sipilupipyo, waikyala • Wp: kule'i sī, pilapuku'i • Wn: kaikui amoman • Nt: bamba apisi • Cr: sèd-marikaj • Fr: cèdre marécage • Br: louro-capitiu.

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: *Service Forestier 7816*.

SIZE. — Up to 20 m tall (Kubitzki 1982).

[577] *Aniba jenmanii* Mez

Jahrb. Königl. Bot. Gart. Berlin 5: 70 (Mez 1889), “Jenmani”.

*Aniba castanea* C.K.Allen, Mem. New York Bot. Gard. 10 (5): 45 (Allen 1964).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — 14 collections at CAY. Sel. exs.: *D. Sabatier 4370*.

INVENTORY DATA (FG). — 5 trees in 5 plots; Fmax < 1%; dbhinv = 15.9 cm.

[578] *Aniba kappleri* Mez

Jahrb. Königl. Bot. Gart. Berlin 5: 52 (Mez 1889).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Ka: siduwapali.

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *J.-F. Molino et al. 3333*.

INVENTORY DATA (FG). — 15 trees in 10 plots; Fmax < 1%; dbhinv = 19.4 cm.

[579] *Aniba megaphylla* Mez
(Fig. 26B)

Jahrb. Königl. Bot. Gart. Berlin 5: 67 (Mez 1889).

*Aniba anisosepala* Sandwith, Bull. Misc. Inform. Kew 1933 (7): 336 [25 Sep. 1933] (Sandwith 1933).

*Aniba koumaroucapa* Kosterm., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 25: 24 (Kostermans 1936). — *Laurus koumaroucapa* Rich. ex Kosterm., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 25: 24 (Kostermans 1936), nom. nud. pro syn.

HERBARIUM DATA (FG). — 16 collections at CAY. Sel. exs.: *F.M.R. Leprieur 225* (holo-, P[P00745605]; iso-, F[V0061345F], G[G00368662], P[P00745604]).

INVENTORY DATA (FG). — 1 tree, dbh = 10.5 cm.

[580] *Aniba panurensis* (Meisn.) Mez

Jahrb. Königl. Bot. Gart. Berlin 5: 58 (Mez 1889). — *Aydendron panurensis* Meisn., Prodr. [A. P. de Candolle] 15 (1): 89 [May 1864] (Meisner 1864).

*Aniba gonggrijpii* Kosterm., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 25: 31 (Kostermans 1936).

*Aniba mas* Kosterm., Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 25: 31 (Kostermans 1936).

VERNACULAR NAMES. — Ka: sipilulan.

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *D. Sabatier et al. 4604*.

INVENTORY DATA (FG). — 19 trees in 15 plots; Fmax < 1%; dbhinv = 43 cm.

[581] *Aniba parviflora* (Meisn.) Mez

Jahrb. Königl. Bot. Gart. Berlin 5: 56 (Mez 1889). — *Aydendron parviflorum* Meisn., Prodr. [A. P. de Candolle] 15 (1): 90 [May 1864] (Meisner 1864).

VERNACULAR NAMES. — Pa: sedri-kamwi-seine • Nt: bwadiwosu • Br: louro-puxuri.

HERBARIUM DATA (FG). — 25 collections at CAY. Sel. exs.: *J.-F. Molino et al. 3352*.

INVENTORY DATA (FG). — 33 trees in 21 plots; Fmax = 1%; dbhinv = 50.3 cm.

[582] *Aniba riparia* (Nees) Mez

Jahrb. Königl. Bot. Gart. Berlin 5: 62 (Mez 1889). — *Aydendron riparium* Nees, Linnaea 21: 497 (Nees 1848).

VERNACULAR NAMES. — Ka: siduwapali.

HERBARIUM DATA (FG). — A single collection, *C. Feuillet 10338* (US).

SIZE. — Up to 16 m tall (Kubitzki 1982).

[583] *Aniba rosodora* Ducke

Rev. Bot. Appl. Agric. Colon. 88: 845 [Dec. 1928] (Ducke 1928), “rosaeodora”.

*Aniba rosodora* var. *amazonica* Ducke, Arch. Jard. Bot. Rio de Janeiro 5: 83 (Ducke 1930), “rosaeodora”.
*Aniba duckei* Kosterm., *Recueil Trav. Bot. Néerl.* 35: 924 (Kostermans 1938).

NOTE. — Epithet corrected according to Art. 60.10 and Rec. 60G.1 (a), of the Shenzhen Code (Turland *et al.* 2018).

VERNACULAR NAMES. — Pa: sedri-eminyo • Wp: áyǔ'i tawa, pauteloso • Wn: matolosi, mijetaimé • Nt: bwadiwosu • Cr: bwa-dé-ròz • Fr: bois de rose • Br: pau-rosa.

HERBARIUM DATA (FG). — 13 collections at CAY. Sel. exs.: *M.-F. Prévost 3301*.

INVENTORY DATA (FG). — 15 trees in 13 plots; $F_{max} < 1\%$; $dbh_{inv} = 36.8$ cm.

[584] *Aniba taubertiana* Mez

Jahrb. Königl. Bot. Gart. Berlin 5: 65 (Mez 1889).

*Aniba simulans* C.K.Allen, *Mem. New York Bot. Gard.* 10 (5): 51 (Allen 1964).

VERNACULAR NAMES. — Ka: tamunen wonu • Wp: áyǔ'i piye.

HERBARIUM DATA (FG). — 13 collections at CAY. Sel. exs.: *E.M. Mélinon s.n.* (holo-, B[not seen, photo F neg. N° 3809]; iso-, BM[BM000617490], CAS[CAS0026940], F[V0061352F], G[G00368692], GH[GH00056912], K[K000601899, K000601900], NY[00354852, 00354853], P[P00745619, P00745620, P00745621], US[00099455]).

INVENTORY DATA (FG). — 67 trees in 54 plots; $F_{max} < 1\%$; $dbh_{inv} = 32.9$ cm.

[585] *Aniba terminalis* Ducke

*Rev. Bot. Appl. Agric. Colon.* 88: 846 [Dec. 1928] (Ducke 1928).

VERNACULAR NAMES. — Ka: siduwapali.

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 5099*.

INVENTORY DATA (FG). — 1 tree, $dbh = 12.3$ cm.

[586] *Aniba williamsii* O.C.Schmidt
(Fig. 26C)

*Repert. Spec. Nov. Regni Veg.* 31: 169 (Schmidt 1933).

*Aniba murcana* C.K.Allen, *Mem. New York Bot. Gard.* 15: 57 (Allen 1966).

VERNACULAR NAMES. — Wp: kule'i • Br: louro-abacate.

HERBARIUM DATA (FG). — 17 collections at CAY. Sel. exs.: *D. Sabatier & E. Fonty 5662*.

INVENTORY DATA (FG). — 20 trees in 17 plots; $F_{max} < 1\%$; $dbh_{inv} = 25.1$ cm.

[587] *Aniba* sp. A

HERBARIUM DATA (FG). — A single collection, *D. Sabatier & E. Fonty 5591*.

INVENTORY DATA (FG). — 3 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 27.3$ cm.

[588] *Aniba* sp. B

HERBARIUM DATA (FG). — A single collection, *D. Sabatier & M.-F. Prévost 1917*.

INVENTORY DATA (FG). — 3 trees in 3 plots; $F_{max} < 1\%$; $dbh_{inv} = 15.5$ cm.

[589] *Aniba* sp. C

HERBARIUM DATA (FG). — A single collection, *S.A. Mori & T.D. Pennington 18052*.

SIZE. — Up to 18 m tall (Werff 2002, as *Rhodostemonodaphne* sp. 1).

Genus *Beilschmiedia* Nees

[590] *Beilschmiedia hexanthera* van der Werff

Brittonia 47 (4): 374 (van der Werff 1995).

NOTE. — Known only from French Guiana.

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *H. van der Werff et al. 12951* (holo-, MO[MO-276584]; iso-, B[B100247471], BM[BM000588156, BM000588157], CAY[CAY184925], MO[MO-276583], NY[00214380], U[U0008155], XAL[XAL0091566]).

INVENTORY DATA (FG). — 1 tree, $dbh = 37.6$ cm.

Genus *Cryptocarya* R.Br.

[591] *Cryptocarya guianensis* Meisn.

*Prodr. [A. P. de Candolle]* 15 (1): 75 [May 1864] (Meisner 1864).

*Cryptocarya maroniensis* Benoist, *Bull. Mus. Natl. Hist. Nat.* 30: 510 (Benoist 1924).

*Cryptocarya nigropunctata* Vattimo-Gil, *Rodriguésia* 25: 222 (Vattimo-Gil 1966).

VERNACULAR NAMES. — Pa: wen-kamwi • Wp: iwa pane • Cr: sèdmarikaj • Br: caa-xió, caiaxió.

HERBARIUM DATA (FG). — 20 collections at CAY. Sel. exs.: *M.E. Moricand 113* (lecto-, G-DC[not seen], designated by Moraes [2005: 791]; isolecto-, G[G00418990, G00418997]).

INVENTORY DATA (FG). — 1 tree, $dbh = 22.4$ cm.

Genus *Damburneya* Raf.

[592] *Damburneya purpurea* (Ruiz & Pav.) Trofimov

*Taxon* 65 (5): 991 [27 Oct. 2016] (Trofimov 2016). — *Laurus purpurea* Ruiz & Pav., *Flora Peruviana* 4: t. 351 [1804-1830] (Ruiz & Pavón 1804). — *Nectandra purpurea* (Ruiz & Pav.) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 443 (Mez 1889).

*Ocotea latifolia* Kunth, *Nova genera et species plantarum [H.B.K.]* 2: 165 [8 Dec. 1817] (Kunth 1818). — *Persea latifolia* (Kunth) Spreng., *Syst. Veg. [Sprengel]* 2: 270 [Jan.-May 1825] (Sprengel 1825). — *Nectandra latifolia* (Kunth) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 454 (Mez 1889).
*Nectandra polita* Nees & Mart., *Syst. Laur.* 325 [30 Oct.-5 Nov. 1836] (Nees & Martius 1836).

*Nectandra polita* var. oerstedii Meisn., *Prodr. [A. P. de Candolle]* 15 (1): 164 [May 1864] (Meisner 1864).

*Ocotea flexuosa* Rusby, *Descr. S. Amer. Pl.* 21 [20 Dec. 1920] (Rusby 1920).

**VERNACULAR NAMES.** — Pa: akamna, akamnâ, akapna.

**HERBARIUM DATA (FG).** — 2 collections at CAY. Sel. exs.: *S.A. Mori et al. 24258*.

**SIZE.** — Peru, Amazonas. *T.D. Pennington & A. Daza Yomona 17839* (MO), 10 m × 20 cm.

## Genus *Endlicheria* Nees

### [593] *Endlicheria bracteolata* (Meisn.) C.K.Allen

*Mem. New York Bot. Gard.* 10 (5): 64 (Allen 1964). — *Goeppertia sericea* (Nees) Nees var. *bracteolata* Meisn., *Prodr. [A. P. de Candolle]* 15 (1): 174 [May 1864] (Meisner 1864).

**VERNACULAR NAMES.** — Wp: âyû'i alapitâ milâ • Br: louro-peludo.

**HERBARIUM DATA (FG).** — 12 collections at CAY. Sel. exs.: *R.A.A. Oldeman B-3580*.

**SIZE.** — Up to 15 m tall (Chanderbali 2004).

### [594] *Endlicheria canescens* Chanderb.

*Novon* 6 (4): 328 [winter 1996] (Chanderbali 1996).

**HERBARIUM DATA (FG).** — 7 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier 2438*.

**INVENTORY DATA (FG).** — 6 trees in 6 plots; $F_{max} < 1\%$; $dbh_{inv} = 29.6$ cm.

### [595] *Endlicheria chalisea* Chanderb.

*Novon* 6 (4): 329 [winter 1996] (Chanderbali 1996).

**HERBARIUM DATA (FG).** — 4 collections at CAY. Sel. exs.: *S. Barrier 4026*.

**INVENTORY DATA (FG).** — 5 trees in 3 plots; $F_{max} < 1\%$; $dbh_{inv} = 25.9$ cm.

### [596] *Endlicheria melinonii* Benoist

*Arch. Bot.* 5 (Mém. 1): 63 [27 Mar. 1933] (Benoist 1933), "Melinonii".

**NOTE.** — Known only from the Guiana Shield.

**VERNACULAR NAMES.** — Pa: migukat, mihukat, mirukat.

**HERBARIUM DATA (FG).** — 20 collections at CAY. Sel. exs.: *E.M. Melinon s.n.* (holo-, P[P00077245]; iso-, P[P00745669]).

**INVENTORY DATA (FG).** — 51 trees in 43 plots; $F_{max} < 1\%$; $dbh_{inv} = 67.8$ cm.

### [597] *Endlicheria punctulata* (Mez) C.K.Allen

*Mem. New York Bot. Gard.* 15: 68 (Allen 1966). — *Ocotea punctulata* Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 379 (Mez 1889).

**NOTE.** — Known only from the Guiana Shield.

**VERNACULAR NAMES.** — Wp: âyû'i alapitâ milâ.

**HERBARIUM DATA (FG).** — 24 collections at CAY. Sel. exs.: *E.M. Melinon 204*, 1842 (lecto-, P[P00745657], designated by Allen [1966: 68]; isolecto-, P[P00745658]).

**INVENTORY DATA (FG).** — 10 trees in 5 plots; $F_{max} < 1\%$; $dbh_{inv} = 46.7$ cm.

### [598] *Endlicheria pyriformis* (Nees) Mez

*Jahrb. Königl. Bot. Gart. Berlin* 5: 116 (Mez 1889), "*piriformis*". — *Cryptocarya pyriformis* Nees, *Syst. Laur.*: 220 [30 Oct.-5 Nov. 1836] (Nees 1836). — *Mespilodaphne pyriformis* (Nees) Meisn., *Prodr. [A. P. de Candolle]* 15 (1): 108 [May 1864] (Meisner 1864).

**Endlicheria glaberrima** Mez, *Bull. Herb. Boissier*, sér. 2, 5: 236 (Mez 1905).

*Aniba flexuosa* A.C.Sm., *Phytologia* 1 (3): 117 [21 Jan. 1935] (Smith 1935).

**VERNACULAR NAMES.** — Pa: kudagman • Ka: woko mapili • Wp: amâkiya sili, âyû'i • Wn: alupkamata • Cr: sèd • Br: louro.

**HERBARIUM DATA (FG).** — 69 collections at CAY. Sel. exs.: *P.A. Poiteau s.n.* (holo-, B[not seen, photo F neg. N° 3820]; iso-, BM[BM000993875], G-DC[G00201070], LE[LE00012615, LE00012616, LE00012728], NY[00099497], P[P00745654]).

**SIZE.** — Up to 10 m tall (Chanderbali 2004).

### [599] *Endlicheria rubriflora* Mez

*Jahrb. Königl. Bot. Gart. Berlin* 5: 494 (Mez 1889).

*Aniba reticulata* A.C.Sm., *Bull. Torrey Bot. Club* 58 (2): 99 [Feb. 1931] (Smith 1931).

**Endlicheria trianae** O.C.Schmidt, *Repert. Spec. Nov. Regni Veg.* 31: 175 (Schmidt 1933).

**Endlicheria wurdackiana** C.K.Allen, *Mem. New York Bot. Gard.* 10 (5): 67 (Allen 1964).

**HERBARIUM DATA (FG).** — 2 collections at CAY. Sel. exs.: *S.A. Mori et al. 23320*.

**SIZE.** — Brazil, Amazonas. *G. T. Prance et al. 14498* (NY), 10 m × 15 cm.

### [600] *Endlicheria szyszylowiczii* Mez

*Jahrb. Königl. Bot. Gart. Berlin* 5: 121 (Mez 1889).

**HERBARIUM DATA (FG).** — A single collection, *R.A.A. Oldeman B-2312*, 12 m × 35 cm.
[601] *Endlicheria* sp. A

NOTE. — This is possibly *E. glomerata* Mez.

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier 2803*.

INVENTORY DATA (FG). — 2 trees in 2 plots; dbh<sub>inv</sub> = 19.5 cm.

## Genus *Kubitzkia* van der Werff

[602] *Kubitzkia mezii* (Kosterm.) van der Werff

*Taxon* 35 (1): 165 [Feb. 1986] (van der Werff 1986). — *Systemonodaphne mezii* Kosterm., *Recueil Trav. Bot. Néerl.* 33: 756 (Kostermans 1936), in obs.

VERNACULAR NAMES. — Wp: kwata pili.

HERBARIUM DATA (FG). — 29 collections at CAY. Sel. exs.: *J. Martin 45* (holotype not traced; iso-, K[K000601987], P[P00745651, P00745652, P00745653], U[U0002743]).

INVENTORY DATA (FG). — 7 trees in 6 plots; F<sub>max</sub> < 1%; dbh<sub>inv</sub> = 16.6 cm.

## Genus *Licaria* Aubl.

[603] *Licaria cannella* (Meisn.) Kosterm.

*Recueil Trav. Bot. Néerl.* 34: 583 (Kostermans 1937), "*Canella*". — *Aydendron cannella* Meisn., *Prodr. [A. P. de Candolle]* 15 (1): 90 [May 1864] (Meisner 1864). — *Acrodiclidium cannella* (Meisn.) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 90 (Mez 1889), "*Canella*". — *Misanteca cannella* (Meisn.) Lundell, *Wrightia* 4 (3): 99 (Lundell 1969), "*Canella*".

*Aniba megacarpa* Hemsl., *Hooker's Icon. Pl.* 28 [ser. 4, 8]: tt. 2751, 2752 [Nov. 1903] (Hemsley 1903).

VERNACULAR NAMES. — Ka: ayiwi, ayu'i, ikalikanali, sitaipyo, wa'elan, wayaka • Wp: āyū'i tawa • Wn: mijetaimë, watau pokan • Nt: kaneli apisi • Cr: bwa-kannel, sèd-jonn, sèd-kannel • Fr: cèdre cannelle • Br: louro-chumbo, louro-pirarucu, louro-preto.

HERBARIUM DATA (FG). — 71 collections at CAY. Sel. exs.: *P.A. Sagot 1190*, Aug. 1858 (holo-, B[not seen, photo F neg. N° 3837]; iso-, BM[BM000993917], K[K000602022], P[P00128461, P00128462], NY[00354870]).

INVENTORY DATA (FG). — 197 trees in 96 plots; F<sub>max</sub> = 3.8%; dbh<sub>inv</sub> = 92 cm.

[604] *Licaria chrysophylla* (Meisn.) Kosterm.

*Recueil Trav. Bot. Néerl.* 34: 601 (Kostermans 1937). — *Acrodiclidium chrysophyllum* Meisn., *Prodr. [A. P. de Candolle]* 15 (1): 87 [May 1864] (Meisner 1864). — *Misanteca chrysophylla* (Meisn.) Lundell, *Wrightia* 4 (3): 99 (Lundell 1969).

*Acrodiclidium guianense* var. *oppositifolium* Meisn., *Prodr. [A. P. de Candolle]* 15 (1): 85 [May 1864] (Meisner 1864).

*Acrodiclidium rigidum* Kosterm., *Meded. Bot. Mus. Herb. Rijks Univ. Utrecht* 25: 38 (Kostermans 1936). — *Licaria rigida* (Kosterm.) Kosterm., *Recueil Trav. Bot. Néerl.* 34: 584 (Kostermans 1937). —

*Misanteca rigida* (Kosterm.) Lundell, *Wrightia* 4 (3): 101 (Lundell 1969).

HERBARIUM DATA (FG). — 20 collections at CAY. Sel. exs.: *P.A. Sagot 956*, 1858 (holo-, B[not seen, photo F neg. N° 3838]; iso-, MEL[MEL2386590]; possible iso-, P[P00128467, P00128468]).

INVENTORY DATA (FG). — 58 trees in 33 plots; F<sub>max</sub> < 1%; dbh<sub>inv</sub> = 73 cm.

[605] *Licaria crassifolia* (Poir.) P.L.R.Moraes

Komarovia 6 (1): 55 (Moraes 2008). — *Laurus crassifolia* Poir., *Encycl. [J. Lamarck et al.]* Suppl. 3: 323 [24 Sep. 1813] (Poiret 1813).

*Aydendron cayennense* Meisn., *Prodr. [A. P. de Candolle]* 15 (1): 95 [May 1864] (Meisner 1864). — *Acrodiclidium cayennense* (Meisn.) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 91 (Mez 1889). — *Licaria cayennensis* (Meisn.) Kosterm., *Recueil Trav. Bot. Néerl.* 34: 583 (Kostermans 1937). — *Misanteca cayennensis* (Meisn.) Lundell, *Wrightia* 4 (3): 99 (Lundell 1969).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *L.C. Richards.n.* (holo-, P[P00128465]; iso-, P[P00128463, P00128464, P00662838]).

SIZE. — > 10 cm dbh (Cardoso *et al.* 2017).

[606] *Licaria debilis* (Mez) Kosterm.

*Recueil Trav. Bot. Néerl.* 34: 596 (Kostermans 1937). — *Acrodiclidium debile* Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 99 (Mez 1889). — *Misanteca debilis* (Mez) Lundell, *Wrightia* 4 (3): 100 (Lundell 1969).

VERNACULAR NAMES. — Wp: ka'i piye.

HERBARIUM DATA (FG). — 22 collections at CAY. Sel. exs.: *J.-F. Molino 907*.

INVENTORY DATA (FG). — 3 trees in 3 plots; F<sub>max</sub> < 1%; dbh<sub>inv</sub> = 14.3 cm.

[607] *Licaria guianensis* Aubl.

*Hist. Pl. Guiane* 1: 313 [Jun.-Dec. 1775] (Aublet 1775), "*Guyanensis*" on plate. — *Acrodiclidium aubletii* Kosterm., *Meded. Bot. Mus. Herb. Rijks Univ. Utrecht* 25: 34 (Kostermans 1936).

*Acrodiclidium appelii* Mez, *Bot. Jahrb. Syst.* 17 (5): 519 [7 Nov. 1893] (Mez 1893). — *Licaria appelii* (Mez) Kosterm., *Recueil Trav. Bot. Néerl.* 34: 601 (Kostermans 1937). — *Misanteca appelii* (Mez) Lundell, *Wrightia* 4 (3): 99 (Lundell 1969).

VERNACULAR NAMES. — Br: louro-aritu, louro-cravo.

HERBARIUM DATA (FG). — 23 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material P[P00128472, P00128471]).

INVENTORY DATA (FG). — 25 trees in 17 plots; F<sub>max</sub> < 1%; dbh<sub>inv</sub> = 19.4 cm.
[608] *Licaria martiniana* (Mez) Kosterm.

Recueil Trav. Bot. Néerl. 34: 601 (Kostermans 1937). — *Acrodi-
clidium martinianum* Mez, Jahrb. Königl. Bot. Gart. Berlin 5: 93
(Mez 1889). — *Misanteca martiniana* (Mez) Lundell, Wrightia 4
(3): 100 (Lundell 1969).

*Licaria foveolata* Lemée, Fl. Guyane Franç. 1: 648 (Lemée 1955).

VERNACULAR NAMES. — Pa: migukat, mihukat, mirukat, sedri-kamwi, sedri-priyo • Ka: yolokan pomiidyi • Te: uwae pitág • Nt: geli apisi, mal bwadiwosu.

HERBARIUM DATA (FG). — 31 collections at CAY. Sel. exs.: *D. Sabatier* 2306.

INVENTORY DATA (FG). — 100 trees in 53 plots; Fmax = 2%; dbhinv = 69.8 cm.

[609] *Licaria pachycarpa* (Meisn.) Kosterm.

Recueil Trav. Bot. Néerl. 34: 604 (Kostermans 1937). — *Aydendron pachycarpum* Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 95 [May 1864] (Meisner 1864). — *Acrodiclidium pachycarpum* (Meisn.) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 91 (Mez 1889). — *Misanteca pachycarpa* (Meisn.) Lundell, *Wrightia* 4 (3): 101 (Lundell 1969).

*Acrodiclidium amarum* Mez, Repert. Spec. Nov. Regni Veg. 16: 305 [31 Apr. 1920] (Mez 1920). — *Licaria amara* (Mez) Kosterm., *Recueil Trav. Bot. Néerl.* 34: 583 (Kostermans 1937). — *Misanteca amara* (Mez) Lundell, Wrightia 4 (3): 99 (Lundell 1969).

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 3780.

INVENTORY DATA (FG). — 11 trees in 9 plots; Fmax < 1%; dbhinv = 25.6 cm.

[610] *Licaria polyphylla* (Nees) Kosterm.

Recueil Trav. Bot. Néerl. 34: 584 (Kostermans 1937). — *Nectandra polyphylla* Nees, Syst. Laur.: 332 [30 Oct.-5 Nov. 1836] (Nees 1836). — *Misanteca polyphylla* (Nees) Lundell, Wrightia 4 (3): 101 (Lundell 1969).

*Acrodiclidium guianense* Nees, Syst. Laur.: 268 [30 Oct.-5 Nov. 1836] (Nees 1836).

*Acrodiclidium guianense* var. *caudatum* Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 85 [May 1864] (Meisner 1864). — *Nectandra caudata* Nees & Mart. ex Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 85 [May 1864] (Meisner 1864), nom. nud. pro syn.

*Acrodiclidium guianense* var. *reticulatum* Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 85 [May 1864] (Meisner 1864).

*Acrodiclidium meissneri* Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 98 (Mez 1889). — *Licaria meissneri* (Mez) Kosterm., *Recueil Trav. Bot. Néerl.* 34: 597 (Kostermans 1937). — *Misanteca meissneri* (Mez) Lundell, Wrightia 4 (3): 100 (Lundell 1969).

NOTE. — *Acrodiclidium guianense* var. *caudatum* is based on the type of *Acrodiclidium meissneri*.

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: R.A.A. Oldeman B-1205, 15 m × 35 cm.

[611] *Licaria rufotomentosa* van der Werff

Novon 13 (3): 342 [autumn 2003] (van der Werff 2003).

NOTE. — Known only from French Guiana.

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *B. Du-
trève & F. Crozier* 7 (holo-, MO[MO-104256, MO-104257]; iso-,
CAY[CAY014160, CAY014161]).

INVENTORY DATA (FG). — 6 trees in 5 plots; Fmax < 1%; dbhinv = 32.5 cm.

[612] *Licaria subbullata* Kosterm.

Reinwardtia 6 (3): 286 (Kostermans 1962). — *Misanteca subbullata* (Kosterm.) Lundell, Wrightia 4 (3): 101 (Lundell 1969).

*Licaria wilhelminensis* C.K.Allen, Mem. New York Bot. Gard. 15: 65 (Allen 1966). — *Misanteca wilhelminensis* (C.K.Allen) Lundell, Wrightia 4 (3): 102 (Lundell 1969).

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 2559, 15 m × 12 cm.

[613] *Licaria vernicosa* (Mez) Kosterm.

Recueil Trav. Bot. Néerl. 34: 604 (Kostermans 1937). — *Ocotea vernicosa* Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 331 (Mez 1889). — *Acrodiclidium vernicosum* (Mez) Kosterm., *Recueil Trav. Bot. Néerl.* 33: 753 (Kostermans 1936). — *Misanteca vernicosa* (Mez) Lundell, Wrightia 4 (3): 102 (Lundell 1969).

*Licaria simulans* C.K.Allen, Mem. New York Bot. Gard. 10 (5): 55 (Allen 1964).

HERBARIUM DATA (FG). — 21 collections at CAY. Sel. exs.: *J.-F. Molino* 1363.

INVENTORY DATA (FG). — 1 tree, dbh = 13.5 cm.

Genus *Mezilaurus* Kuntze ex Taub.

[614] *Mezilaurus itauba* (Meisn.) Taub. ex Mez

Arbeiten Königl. Bot. Gart. Breslau 1: 112 (Mez 1892), "Ita-Uba". — *Acrodiclidium itauba* Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 86 [May 1864] (Meisner 1864), "Ita-uba". — *Endiandra itauba* (Meisn.) Benth. & Hook.f., *Gen. Pl.* [*Bentham & Hooker f.*] 3 (1): 154 [7 Feb. 1880] (Bentham & Hooker 1880), "Ita-uba". — *Silvia itauba* (Meisn.) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 107 (Mez 1889), "Ita-Uba". — *Mezia itauba* (Meisn.) Kuntze, *Revis. Gen. Pl.* 2: 574 [5 Nov. 1891] (Kuntze 1891), "Ita-Uba".

*Acrodiclidium anacardioides* Spruce ex Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 86 [May 1864] (Meisner 1864). — *Misanteca anacardioides* (Spruce ex Meisn.) Benth., Hooker's Icon. Pl. 13 [ser. 3, 3](2): 47 [June 1878] (Bentham 1878). — *Silvia anacardioides* (Spruce ex Meisn.) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 108 (Mez 1889). — *Mezia anacardioides* (Spruce ex Meisn.) Kuntze, *Revis. Gen. Pl.* 2: 574 [5 Nov. 1891] (Kuntze 1891). — *Mezilaurus anacardioides* (Spruce ex Meisn.) Taub. ex Mez, *Arbeiten Königl. Bot. Gart. Breslau* 1: 112 (Mez 1892).
*Oreodaphne hookeriana* Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 131 [May 1864] (Meisner 1864).

NOTES. — Hyperdominant in Amazonia (ter Steege *et al.* 2020). The specific epithet refers to the Tupi-Guarani word *itaúba* meaning “forest on rock”, from *ita* (rock) and *úba* (forest). The hyphen was artificially introduced (“Ita-úba”) by Meissner (in Candolle 1864: 86) and has therefore to be suppressed according to Art. 60.9 of the Shenzhen Code (Turland *et al.* 2018).

VERNACULAR NAMES. — Ka: ikalikanali • Br: itaúba, louro-itaúba.

HERBARIUM DATA (FG). — 19 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier* 2716.

INVENTORY DATA (FG). — 33 trees in 22 plots; $F_{max} < 1\%$; $dbh_{inv} = 55.7$ cm.

Genus *Nectandra* Rol. ex Rottb.

[615] *Nectandra amazonum* Nees

Syst. Laur.: 282 [30 Oct.-5 Nov. 1836] (Nees 1836).

*Nectandra pallida* Nees, *Linnaea* 21: 510 (Nees 1848).

*Nectandra urophylla* Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 157 [May 1864] (Meisner 1864).

*Nectandra ambigua* Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 158 [May 1864] (Meisner 1864).

*Nectandra bombycina* S.Moore, *Trans. Linn. Soc. London, Bot.* 4 (3): 449 ["1894-96" publ. Dec. 1895] (Moore 1895).

VERNACULAR NAMES. — Br: louro-do-igapó.

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *O. Poncy et al.* 1680.

INVENTORY DATA (FG). — 2 trees in 2 plots; $dbh_{inv} = 33.6$ cm.

[616] *Nectandra cissiflora* Nees

Syst. Laur.: 296 [30 Oct.-5 Nov. 1836] (Nees 1836).

*Nectandra myriantha* Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 163 [May 1864] (Meisner 1864).

*Nectandra myriantha* var. attenuata Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 163 [May 1864] (Meisner 1864).

*Nectandra myriantha* var. glabrata Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 163 [May 1864] (Meisner 1864).

*Nectandra surinamensis* Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 454 (Mez 1889), *nom. illeg. hom., non* Rottb. ex Bercht. & J.Presl (Berchtold & Presl 1825).

*Nectandra kuntzeana* Mez in Kuntze, *Revis. Gen. Pl.* 3 (3): 277 [28 Sep. 1898] (Mez 1898).

*Nectandra capanahuensis* O.C.Schmidt, *Notizbl. Bot. Gart. Berlin-Dahlem* 10: 228 [30 Mar. 1928] (Schmidt 1928).

*Nectandra steinbachii* O.C.Schmidt, *Repert. Spec. Nov. Regni Veg.* 31: 186 (Schmidt 1933).

*Nectandra paulii* C.K.Allen, *J. Arnold Arbor.* 26 (4): 400 [15 Oct. 1945] (Allen 1945).

VERNACULAR NAMES. — Br: canela-fedida, louro-babão.

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier* 2484.

INVENTORY DATA (FG). — 9 trees in 8 plots; $F_{max} < 1\%$; $dbh_{inv} = 44$ cm.

[617] *Nectandra globosa* (Aubl.) Mez

Jahrb. Königl. Bot. Gart. Berlin 5: 415 (Mez 1889). — *Laurus globosa* Aubl., *Hist. Pl. Guiane* 1: 364 [Jun.-Dec. 1775] (Aublet 1775). — *Persea globosa* (Aubl.) Spreng., *Syst. Veg. [Sprengel]* 2: 269 [Jan.-May 1825] (Sprengel 1825). — *Ocotea globosa* (Aubl.) Schltdl. & Cham., *Linnaea* 6: 366 (Schlechtendal & Chamisso 1831).

*Nectandra pisi* Miq., *Stirp. Surinam. Select.*: 199 ["1850" publ. Mar. 1851] (Miquel 1851).

*Nectandra vaga* Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 153 [May 1864] (Meisner 1864).

*Nectandra vaga* var. major Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 154 [May 1864] (Meisner 1864).

*Nectandra vaga* var. sprucei Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 154 [May 1864] (Meisner 1864).

*Nectandra vaga* var. vulgaris Meisn., Prodr. [*A. P. de Candolle*] 15 (1): 154 [May 1864] (Meisner 1864).

*Nectandra globosa* var. barbeyana Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 416 (Mez 1889).

VERNACULAR NAMES. — Wp: mulei.

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000947248]).

INVENTORY DATA (FG). — 8 trees in 4 plots; $F_{max} < 1\%$; $dbh_{inv} = 23.4$ cm.

[618] *Nectandra hihua* (Ruiz & Pav.) Rohwer

Fl. Neotrop. Monogr. 60: 196 [13 Aug. 1993] (Rohwer 1993). — *Laurus hihua* Ruiz & Pav., *Flora Peruviana* 4: t. 366 [1804-1830] (Ruiz & Pavón 1804).

*Nectandra lucida* Nees, Syst. Laur.: 334 [30 Oct.-5 Nov. 1836] (Nees 1836), *nom. illeg. hom., non* Nees (*Linnaea* 8: 47. 1833), *nec* Nees (1836: 334).

*Nectandra willdenoviana* Nees, Syst. Laur.: 290 [30 Oct.-5 Nov. 1836] (Nees 1836), "*Nectandra (Pomatia) willdenoviana*", *non* Nees (1836: 321).

*Nectandra glabrescens* Benth., *Bot. Voy. Sulphur* [Bentham]: 161 [8 May 1846] (Bentham 1846).

*Nectandra bredemeyeriana* Nees, *Linnaea* 21: 505 (Nees 1848).

*Nectandra magnolifolia* A.Rich., *Hist. Fís. Cuba, Bot.* 11: 188 (Richard 1850), "*magnoliaefolia*".
*Nectandra leucantha* Nees & Mart. var. *attenuata* Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 151 [May 1864] (Meisner 1864).

*Nectandra leucantha* var. *guianensis* Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 151 [May 1864] (Meisner 1864).

*Nectandra leucantha* var. *peruviana* Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 151 [May 1864] (Meisner 1864).

*Nectandra schomburgkii* Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 151 [May 1864] (Meisner 1864).

*Nectandra grandiflora* Nees & Mart. var. *latifolia* Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 152 [May 1864] (Meisner 1864), *nom. illeg. hom.*, *non* Nees (1836: 323).

*Nectandra antillana* Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 153 [May 1864] (Meisner 1864).

*Sassafridium macrophyllum* Rose, *Contr. U.S. Natl. Herb.* 1 (9): 355 [31 Jan. 1895] (Rose 1895).

*Nectandra guanaiensis* Rusby, *Bull. New York Bot. Gard.* 6 (22): 508 [30 Nov. 1910] (Rusby 1910).

*Nectandra megaphylla* Hassl., *Annuaire Conserv. Jard. Bot. Genève* 21: 92 (Hassler 1919).

*Nectandra maranonensis* O.C.Schmidt, *Notizbl. Bot. Gart. Berlin-Dahlem* 10: 229 [30 Mar. 1928] (Schmidt 1928).

*Nectandra tessmannii* O.C.Schmidt, *Notizbl. Bot. Gart. Berlin-Dahlem* 10: 229 [30 Mar. 1928] (Schmidt 1928).

*Nectandra albiflora* Lundell, *Wrightia* 1 (2): 147 (Lundell 1946).

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: S.A. Mori & C. Gracie 24212.

SIZE. — Up to 35 m tall (Rohwer 1993).

[619] *Nectandra matogrossensis* Coe-Teix.
(Fig. 26D-F)

*Acta Amazonica* 5 (2): 158 (Coe-Teixeira 1975).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *J.-F. Molino et al.* 3406.

INVENTORY DATA (FG). — 4 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 14.4$ cm.

[620] *Nectandra pulverulenta* Nees

*Syst. Laur.*: 283 [30 Oct.-5 Nov. 1836] (Nees 1836).

*Nectandra purusensis* Coe-Teix., *Acta Amazonica* 5 (2): 158 (Coe-Teixeira 1975).

VERNACULAR NAMES. — Wp: áyǔ'i sili, áyǔ'i tawa, áyǔ'i witowa • Br: louro-abacate.

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: *M.-F. Prévost & P. Grenand* 1000.

SIZE. — Up to 30 m tall (Rohwer 1993).

[621] *Nectandra reticulata* (Ruiz & Pav.) Mez

Jahrb. Königl. Bot. Gart. Berlin 5: 404 (Mez 1889). — *Laurus reticulata* Ruiz & Pav., *Flora Peruviana* 4: t. 348 [1804-1830] (Ruiz & Pavón 1804).

*Ocotea mollis* Kunth, *Nova genera et species plantarum* [H.B.K.] 2: 164 [8 Dec. 1817] (Kunth 1818). — *Persea mollis* (Kunth) Spreng., *Syst. Veg.* [Sprengel] 2: 270 [Jan.-May 1825] (Sprengel 1825). — *Nectandra mollis* (Kunth) Nees, *Hufeland*. III: 14 (Nees 1833).

*Persea incana* Schott, *Syst. Veg.* [*Sprengel*] 4 (2, Cur. Post.): 405 [Jan.-June 1827] (Schott 1827).

*Laurus aestivalis* Vell., *Fl. Flumin.*: 164 [“1825” publ. 7 Sep.-28 Nov. 1829] (Vellozo 1829), *nom. illeg. hom.*, *non* L. (Linnaeus 1753).

*Nectandra villosa* Nees & Mart., *Linnaea* 8: 47 (Nees & Martius 1833). — *Nectandra mollis* var. *villosa* (Nees & Mart.) Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 148 [May 1864] (Meisner 1864).

*Nectandra villosa* var. *venosa* Nees, *Syst. Laur.*: 291 [30 Oct.-5 Nov. 1836] (Nees 1836), “*Venosa*”. — *Nectandra mollis* var. *venosa* (Nees) Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 149 [May 1864] (Meisner 1864).

*Nectandra discolor* (Kunth) Nees var. *subvenosa* Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 157 [May 1864] (Meisner 1864).

*Nectandra laurel* Klotzsch ex Nees var. *triquetra* Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 147 [May 1864] (Meisner 1864).

*Nectandra mollis* var. *attenuata* Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 148 [May 1864] (Meisner 1864).

*Nectandra mollis* var. *intermedia* Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 149 [May 1864] (Meisner 1864).

*Nectandra pittieri* Lasser, *Bol. Técn. Minist. Agric.* 3: 13 (Lasser 1942).

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *J.-F. Molino et al.* 3309.

INVENTORY DATA (FG). — 6 trees in 4 plots; $F_{max} < 1\%$; $dbh_{inv} = 55.5$ cm.

[622] *Nectandra* sp. A

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier* 2816.

INVENTORY DATA (FG). — 2 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 28.7$ cm.

Genus *Ocotea* Aubl.

[623] *Ocotea amazonica* (Meisn.) Mez

Jahrb. Königl. Bot. Gart. Berlin 5: 359 (Mez 1889). — *Oreodaphne amazonica* Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 123 [May 1864] (Meisner 1864).

VERNACULAR NAMES. — Nt: *fiiman* • Br: *louro-preto*.

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 3593.

INVENTORY DATA (FG). — 15 trees in 12 plots; $F_{max} < 1\%$; $dbh_{inv} = 67$ cm.
[624] *Ocotea argyrophylla* Ducke

Arch. Jard. Bot. Rio de Janeiro 4: 193 (Ducke 1925).

VERNACULAR NAMES. — Wp: áyǔ'i alapitā milā • Wn: taihem • Nt: weti apisi, weti sikin apisi • Cr: sèd-nwé • Fr: cèdre argenté • Br: folha-de-prata.

HERBARIUM DATA (FG). — 26 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 2147*.

INVENTORY DATA (FG). — 93 trees in 43 plots; Fmax = 2.4%; dbhinv = 64.1 cm.

[625] *Ocotea canaliculata* (Rich.) Mez

Jahrb. Königl. Bot. Gart. Berlin 5: 361 (Mez 1889). — *Laurus canaliculata* Rich., *Actes Soc. Hist. Nat. Paris* 1: 108 [Oct. 1792] (Richard 1792).

*Oreodaphne cayennensis* Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 133 [May 1864] (Meisner 1864), *syn. nov*.

*Ocotea scrobiculata* Benoist, *Bull. Soc. Bot. France* 75 (5): 979 ["1928" publ. 1929] (Benoist 1929), *nom. nud*.

VERNACULAR NAMES. — Wp: áyǔ'i witowa • Br: louro-pimenta.

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: *J.B. Leblond s.n.* (holo-, P[P00756974]; P[P00756975, P00756976]).

INVENTORY DATA (FG). — 10 trees in 6 plots; Fmax < 1%; dbhinv = 47.4 cm.

[626] *Ocotea ceanothifolia* (Nees) Mez

Jahrb. Königl. Bot. Gart. Berlin 5: 365 (Mez 1889). — *Mespilodaphne ceanothifolia* Nees, *Syst. Laur.*: 236 [30 Oct.-5 Nov. 1836] (Nees 1836). — *Persea ceanothifolia* Mart. ex Nees, *Syst. Laur.*: 236 [30 Oct.-5 Nov. 1836] (Nees 1836), *nom. nud. pro syn*.

VERNACULAR NAMES. — Wp: áyǔ'i tawa • Nt: likanau apisi.

HERBARIUM DATA (FG). — 24 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 4200*.

INVENTORY DATA (FG). — 25 trees in 16 plots; Fmax < 1%; dbhinv = 103 cm.

[627] *Ocotea cernua* (Nees) Mez

Mitt. Bot. Vereins Kreis Freiburg 47-48: 422 (Mez 1888). — *Oreodaphne cernua* Nees, *Syst. Laur.*: 424 [30 Oct.-5 Nov. 1836] (Nees 1836).

*Laurus nitida* Ruiz & Pav., *Flora Peruviana* 4: t. 353 [1804-1830] (Ruiz & Pavón 1804). — *Mespilodaphne nitida* (Ruiz & Pav.) Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 106 [May 1864] (Meisner 1864).

*Laurus leptobotra* Ruiz & Pav., *Flora Peruviana* 4: t. 368 [1804-1830] (Ruiz & Pavón 1804). — *Oreodaphne leptobotra* (Ruiz & Pav.) Nees, *Linnaea* 21: 520 (Nees 1848). — *Ocotea leptobotra* (Ruiz & Pav.) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 376 (Mez 1889).

*Leptodaphne tenuiflora* Nees, *Syst. Laur.*: 359 [30 Oct.-5 Nov. 1836] (Nees 1836). — *Persea tenuiflora* Mart. ex Nees, *Syst. Laur.*: 359 [30 Oct.-5 Nov. 1836] (Nees 1836), *nom. nud. pro syn*.

*Leptodaphne subalpina* Nees, *Syst. Laur.*: 361 [30 Oct.-5 Nov. 1836] (Nees 1836). — *Oreodaphne subalpina* (Nees) Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 137 [May 1864] (Meisner 1864). — *Ocotea subalpina* (Nees) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 375 (Mez 1889).

*Oreodaphne caudata* Nees, *Linnaea* 21: 519 (Nees 1848). — *Camphoromoea surinamensis* Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 145 [May 1864] (Meisner 1864), *nom. illeg. superfl.* (one of the syntypes [Hostmann 964] is a syntype of *Oreodaphne caudata*). — *Ocotea caudata* (Nees) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 378 (Mez 1889). — *Camphoromoea tenuiflora* (Nees) Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 144 [May 1864] (Meisner 1864).

*Oreodaphne marowynensis* Miq., *Stirp. Surinam. Select.*: 201 ["1850" publ. Mar. 1851] (Miquel 1851). — *Ocotea marowynensis* (Miq.) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 380 (Mez 1889).

*Oreodaphne urophylla* Meisn., *Prodr.* [*A. P. de Candolle*] 15 (1): 145 [May 1864] (Meisner 1864). — *Ocotea urophylla* (Meisn.) Mez, *Jahrb. Königl. Bot. Gart. Berlin* 5: 379 (Mez 1889).

*Ocotea marmellensis* Mez var. *acrensis* J.F.Macbr., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 13 (2.3): 903 [31 Oct. 1938] (Macbride 1938).

NOTES. — Excluded synonym: *Nectandra revoluta* Miq. ex Rohwer (1993: 302), *nom. nud*. Duplicates of Pl. *Surinam. 629bb* distributed under A. Kappler's name (M, MPU, P[P00756833], S, U[U0002941]) bear on the original label "*Nectandra revoluta* Miq." (This is not the case for the duplicates distributed under Hostmann's name). A label by C.K. Allen dated 1970 affixed to the S duplicate wrongly states that this specimen is an isotype of *N. revoluta*, and that this name is a synonym of *Oreodaphne marowynensis* Miquel (1851: 201). With respect to the latter name, Miquel originally wrote (1851: 201): "*Nectandra revoluta* a me perperam dicta" ("which I wrongly called *N. revoluta*"). Therefore, *A. Kappler* 629bb is not the type of *N. revoluta*, which is a *nomen nudum* and is not a synonym of *O. marowynensis*.

VERNACULAR NAMES. — Ka: wa'elan, wayaka • Te: uwae hun • Wp: áyǔ'i tawa, áyǔ'i witowa.

HERBARIUM DATA (FG). — 19 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier 2719*.

INVENTORY DATA (FG). — 22 trees in 15 plots; Fmax < 1%; dbhinv = 23.9 cm.

[628] *Ocotea cinerea* van der Werff

Novon 10 (3): 268 [autumn 2000] (van der Werff 2000).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: migukat, mihukat, mirukat • Wp: kwata pili • Nt: baaka sikin apisi.

HERBARIUM DATA (FG). — 33 collections at CAY. Sel. exs.: S.A. *Mori et al.* 23898.

INVENTORY DATA (FG). — 159 trees in 58 plots; Fmax = 4.9%; dbhinv = 98.7 cm.