*Psychotria ernestii* K.Krause, *Verh. Bot. Vereins Prov. Brandenburg* 50: 109 [30 Sep. 1908] (Krause 1908), "*Ernesti*".

*Psychotria mathewsii* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 4 (8): 342 [24 Oct. 1929] (Standley 1929).

**VERNACULAR NAMES.** — Pa: pune-etni-seine • Ka: kuyaken elepali • Te: tapi'i ka'a lilu • Wp: tapi'i ka'alulu • Wn: talepëpë • Nt: man agu siton • Br: apuruí.

**HERBARIUM DATA (FG).** — 154 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JJR[P00778114] designated by Lanjouw & Uittien [1940: 153]).

**SIZE.** — Up to 25 m tall (Boom & Delprete 2002).

[1495] *Psychotria pedunculosa* Rich.

*Actes Soc. Hist. Nat. Paris* 1: 107 [Oct. 1792] (Richard 1792). — *Uragoga pedunculosa* (Rich.) Kuntze, *Revis. Gen. Pl.* 2: 962 [5 Nov. 1891] (Kuntze 1891).

*Psychotria mapourioides* DC., *Prodr.* [*A. P. de Candolle*] 4: 509 [late Sep. 1830] (Candolle 1830).

*Palicourea pedunculosa* DC., *Prodr.* [*A. P. de Candolle*] 4: 526 [late Sep. 1830] (Candolle 1830).

*Palicourea chionantha* DC., *Prodr.* [*A. P. de Candolle*] 4: 526 [late Sep. 1830] (Candolle 1830). — *Mapouria chionantha* (DC.) Müll.Arg., *Fl. Bras.* [*Martius*] 6 (5): 387 [1 July 1881] (Müller 1881). — *Psychotria chionantha* (DC.) Britton, *Bull. Torrey Bot. Club* 18 (4): 109 (Britton 1891). — *Uragoga chionantha* (DC.) Kuntze, *Revis. Gen. Pl.* 2: 959 [5 Nov. 1891] (Kuntze 1891). — *Psychotria mapourioides* var. *chionantha* (DC.) Steyerm., *Mem. New York Bot. Gard.* 23: 462 (Steyermark 1972).

**HERBARIUM DATA (FG).** — 47 collections at CAY. Sel. exs.: *J.B. Leblond* 319 (original material G[G00341845]).

**SIZE.** — Brazil. *S.A. Mori* & *T.S. dos Santos* 10131 (MO), 16 m × 20 cm.

Genus *Randia* L.

[1496] *Randia armata* (Sw.) DC.

*Prodr.* [*A. P. de Candolle*] 4: 387 [late Sep. 1830] (Candolle 1830). — *Gardenia armata* Sw., *Prodr.* [Swartz]; 51 [20 Jun.-29 July 1788] (Swartz 1788). — *Solena armata* (Sw.) D.Dietr., *Syn. Pl.* [*D. Dietrich*] 1: 800 [July 1839] (Dietrich 1839).

*Mussaenda spinosa* Jacq., * Enum. Syst. Pl. *: 16 [Aug.-Sep. 1760] (Jacquin 1760). — *Gardenia tetracantha* Lam., *Encycl.* [*J. Lamarck et al.*] 2 (2): 609 [14 Apr. 1788] (Lamarck 1788), *nom. illeg. superfl.* (based on *Mussaenda spinosa*). — *Randia spinosa* (Jacq.) H.Karst., *Fl. Columb.* [*H. Karsten*] 2 (4): 128 [28 Nov. 1866] (Karsten 1866), *nom. illeg. hom.*, *non* (Thunb.) Poir. (Poiret 1812). — *Basanacantha spinosa* (Jacq.) K.Schum., *Fl. Bras.* [*Martius*] 6 (6): 376 [15 June 1889] (Schumann 1889). — *Basanacantha spinosa* var. *typica* K.Schum., *Fl. Bras.* [*Martius*] 6 (6): 377 [15 June 1889] (Schumann 1889), *nom. inval.* (Turland *et al.* 2018: Art. 24.3).

*Randia ovata* Duchass. ex Griseb., *Bonplandia* 6 (1): 8 (Grisebach 1858).

*Randia guianensis* Sagot ex K.Schum., *Fl. Bras.* [*Martius*] 6 (6): 377 [15 June 1889] (Schumann 1889), *nom. nud. pro syn.*

*Basanacantha spinosa* var. *polyantha* K.Schum., *Fl. Bras.* [*Martius*] 6 (6): 377 [15 June 1889] (Schumann 1889).

*Basanacantha spinosa* f. *pubiloba* K.Schum., *Fl. Bras.* [*Martius*] 6 (6): 377 [15 June 1889] (Schumann 1889).

*Basanacantha spinosa* var. *longipedunculata* Kuntze, *Revis. Gen. Pl.* 3 (3): 118 [28 Sep. 1898] (Kuntze 1898).

*Basanacantha spinosa* f. *grandiflora* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 173 (Chodat & Hassler 1904).

*Basanacantha spinosa* f. *puberula* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 173 (Chodat & Hassler 1904).

*Basanacantha spinosa* var. *macrocalyx* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 173 (Chodat & Hassler 1904).

*Basanacantha phyllosepala* Sprague & R.O.Williams ex R.O.Williams & Cheesman, *Fl. Trinidad & Tobago* 2 (1): 22 [Aug. 1928] (Williams & Cheesman 1928).

**VERNACULAR NAMES.** — Ka: alemikilan • Wp: alamakulu alami • Br: espinho-de-judeu, esporão-de-galo.

**HERBARIUM DATA (FG).** — 36 collections at CAY. Sel. exs.: *J.-F. Molino* & *D. Sabatier* 2250.

**INVENTORY DATA (FG).** — 3 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 11.5$ cm.

Genus *Rudgea* Salisb.

[1497] *Rudgea crassiloba* (Benth.) B.L.Rob.

*Proc. Amer. Acad. Arts* 45: 408 (Robinson 1910). — *Coffea crassiloba* Benth., *J. Bot.* [*Hooker*] 3: 233 (Bentham 1841).

*Rudgea schomburgkiana* Benth., *Linnaea* 23: 459 (Bentham 1850), *nom. illeg. superfl.* (based on *Coffea crassiloba* Benth.)

**HERBARIUM DATA (FG).** — 7 collections at CAY. Sel. exs.: *J.-J. de Granville* 5909.

**SIZE.** — Brazil, Pará. *T.C. Plowman* 9069 (MO), 7 m × 10 cm.

[1498] *Rudgea graciliflora* Standl.

*Publ. Field Mus. Nat. Hist., Bot. Ser.* 11 (5): 262 (Standley 1936).

*Rudgea duckei* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 11 (5): 261 (Standley 1936).

*Rudgea standleyana* Steyerm., *Mem. New York Bot. Gard.* 17 (1): 409 (Steyermark 1967).

**HERBARIUM DATA (FG).** — 2 collections at CAY. Sel. exs.: *S. Gonzalez* 2334.

**SIZE.** — Brazil, Amazonas. *S.A. Mori* 20612 (MO), 20 m × 10 cm.
[1499] *Rudgea lanceifolia* Salisb.
(Fig. 50C)

*Trans. Linn. Soc. London* 8: 327 (Salisbury 1807), "lanceaefolia". — *Uragoga lanceifolia* (Salisb.) Kuntze, *Revis. Gen. Pl.* 2: 961 [5 Nov. 1891] (Kuntze 1891), "lanceaefolia".

*Rudgea ovalifolia* Salisb., *Trans. Linn. Soc. London* 8: 328 (Salisbury 1807). — *Uragoga ovalifolia* (Salisb.) Kuntze, *Revis. Gen. Pl.* 2: 961 [5 Nov. 1891] (Kuntze 1891).

*Rudgea fissistipula* Müll.Arg., *Flora* 59: 449 (Müller 1876). — *Uragoga fissistipula* Müll.Arg.) Kuntze, *Revis. Gen. Pl.* 2: 960 [5 Nov. 1891] (Kuntze 1891).

*Rudgea carolina* Standl. & Steyerm., *Fieldiana*, Bot. 28 (3): 614 (Standley & Steyermark 1953).

*Rudgea dasyantha* Bremek., *Brittonia* 8 (4): 244 [Jan. 1957] (Bremekamp 1957).

*Rudgea sipapoensis* Steyerm., *Mem. New York Bot. Gard.* 17 (1): 411 (Steyermark 1967).

*Rudgea oldemanii* Steyerm., *Brittonia* 33 (3): 397 (Steyermark 1981).

*Rudgea prancei* Steyerm., *Brittonia* 33 (3): 398 (Steyermark 1981).

VERNACULAR NAMES. — Pa: pune-etni-kamwi.

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: *J. Martin s.n.* (original material BM[BM000582180, BM000582181]).

INVENTORY DATA (FG). — 9 trees in 3 plots; Fmax < 1%; dbhinv = 13.8 cm.

Genus *Simira* Aubl.

[1500] *Simira tinctoria* Aubl.

*Hist. Pl. Guiane* 1: 170 [Jun.-Dec. 1775] (Aublet 1775). — *Psychotria tinctoria* (Aubl.) Raeusch., *Nomencl. Bot. [Raeusch.]*, ed. 3: 55 (Raeuschel 1797). — *Psychotria parviflora* Willd., *Sp. Pl.*, ed. 4 1 (2): 962 [July 1798] (Willdenow 1798), *nom. illeg. superfl.* (based on *Simira tinctoria*). — *Psychotria simira* Roem. & Schult., *Syst. Veg. [Roemer & Schultes]* 5: 187 [Dec. 1819] (Roemer & Schultes 1819), *nom. illeg. superfl.* (based on *Simira tinctoria*). — *Mapouria simira* A.Rich., *Mém. Rubiac.*: 94 [Dec. 1830] (Richard 1830), *nom. illeg. superfl.* (based on *Simira tinctoria*). — *Uragoga simira* Baill., *Hist. Pl. [Baillon]* 7: 374 [Feb. 1880] (Baillon 1880), *nom. illeg. superfl.* (based on *Simira tinctoria*). — *Uragoga tinctoria* (Aubl.) Kuntze, *Revis. Gen. Pl.* 2: 958 [5 Nov. 1891] (Kuntze 1891). — *Sickingia tinctoria* (Aubl.) Lemée, *Fl. Guyane Franç.* 3: 505 (Lemée 1954), *nom. illeg. hom.*, *non* (Willd. ex Schult.) K.Schum. (Schumann 1888).

*Chimarrhis clausicorollata* J.H.Kirkbr., *BioLlania*, ed. *espec*. 6: 394 (Kirkbride 1997).

VERNACULAR NAMES. — Pa: adamna • Ka: simira (fide Aublet 1775) • Wn: kujeimë, kusi wewe • Nt: kenkii udu • Br: arariba.

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JJR[P00778112] designated by Lanjouw & Uitten [1940: 156]); *D. Sabatier & M.-F. Prévost 3400*, dbh = 35 cm.

Genus *Stenostomum* C.F.Gaertn.

[1501] *Stenostomum acreanum*
(K.Krause) Achille & Delprete

*Blumea* 55 (2): 167 [17 Aug. 2010] (Achille & Delprete 2010). — *Stenostomum acreanum* (K.Krause) C.M.Taylor, *Novon* 20 (3): 360 [13 Sep. 2010] (Taylor 2010), comb. superfl. — *Guettarda acreana* K.Krause, *Notizbl. Königl. Bot. Gart. Berlin* 6: 204 [15 May 1914] (Krause 1914).

*Antirhea panamensis* Standl., *N. Amer. Fl.* 32 (4): 264 [21 Dec. 1934] (Standley 1934). — *Chomelia panamensis* (Standl.) Dwyer, *Ann. Missouri Bot. Gard.* 67 (1): 100 (Dwyer 1980).

*Antirhea surinamensis* Bremek., *Acta Bot. Neerl.* 8: 479 (Bremekamp 1959).

*Chomelia chambersii* Dwyer & M.V.Hayden, *Ann. Missouri Bot. Gard.* 54 (2): 138 [27 Oct. 1967] (Dwyer & Hayden 1967). — *Chione chambersii* Dwyer, *Ann. Missouri Bot. Gard.* 67 (1): 100 (Dwyer 1980).

*Guettarda leiantha* Steyerm., *Ann. Missouri Bot. Gard.* 71 (4): 1175 (Steyermark 1984).

NOTE. — *Chione chambersii* is a mispell of *Chomelia chambersii* in the synonymy of *Chomelia panamensis*.

HERBARIUM DATA (FG). — 23 collections at CAY. Sel. exs.: *S.A. Mori & B.M. Boom 14691*.

INVENTORY DATA (FG). — 39 trees in 12 plots; Fmax = 1.6%; dbhinv = 50 cm.

[1502] *Stenostomum guianensis*
(Bremek.) Delprete & Achille

*Blumea* 55 (2): 169 [17 Aug. 2010] (Delprete & Achille 2010). — *Antirhea guianensis* Bremek., *Kew Bull.* 7 (2): 260 [25 July 1952] (Bremekamp 1952).

*Guettarda fanshawei* Steyerm., *Mem. New York Bot. Gard.* 23: 359 (Steyermark 1972).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — A single collection, *R.A.A. Oldeman B-1936*.

SIZE. — Up to 50 cm dbh (Delprete et al. 2010).

Genus *Tocoyena* Aubl.

[1503] *Tocoyena guianensis* K.Schum.

*Fl. Bras. [Martius]* 6 (6): 346 [15 June 1889] (Schumann 1889).

*Tocoyena guianensis* var. *communis* Steyerm., *Mem. New York Bot. Gard.* 12 (3): 194 (Steyermark 1965).

*Tocoyena guianensis* var. *glabriuscula* Steyerm., *Mem. New York Bot. Gard.* 12 (3): 194 (Steyermark 1965).

VERNACULAR NAMES. — Pa: amuwan, tukuyuy-kamwi • Wp: yanipa lali u • Wn: kuwajimë, piilimokju • Br: jenipapo-do-campo.
HERBARIUM DATA (FG). — 42 collections at CAY. Sel. exs.: *R.A.A.*
*Oldeman 2756*.

INVENTORY DATA (FG). — 8 trees in 1 plot; Fmax< 1 %; dbh<sub>inv</sub> =
33 cm.

Family RUTACEAE Juss.
Genus *Erythrochiton* Nees & Mart.

[1504] *Erythrochiton brasiliensis* Nees & Mart.

*Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur.* 11: 166 (Nees & Martius 1823).

Pentamorpha graveolens Scheidw., *Bull. Acad. Roy. Sci. Bruxelles* 9 (1): 21 (Scheidweiler 1842).

VERNACULAR NAMES. — Wn: kiapoku, pakutpë, pija pëloman.

HERBARIUM DATA (FG). — 17 collections at CAY. Sel. exs.: *M. Fleury 2053*.

SIZE. — Up to 12 m tall (Kallunki 1992).

Genus *Esenbeckia* Kunth

[1505] *Esenbeckia cowanii* Kaastra

*Acta Bot. Neerl.* 26: 481 (Kaastra 1977).

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *R.S. Cowan 38757* (holo-, US[00101521]; iso-, F[V0071245F], NY[00051857]).

INVENTORY DATA (FG). — 34 trees in 8 plots; Fmax < 1 %; dbh<sub>inv</sub> =
35 cm.

[1506] *Esenbeckia grandiflora* Mart.

*Nova genera et species plantarum* [Martius] 3 (2): 85 ["1829" publ. Jan.-Mar (?) 1831] (Martius 1831). — *Colythrum grandiflorum* (Mart.) Steud., *Nomencl. Bot. [Steudel]*, ed. 2, 1: 399 (Steudel 1840).

Polembryum castanocarpum A.Juss., *Mém. Mus. Hist. Nat.* 12: 541 (Jussieu 1825).

Polembryum jussieui Schott, *Rutac.* t. 6 (Schott 1834).

*Esenbeckia attenuata* Griseb., *Fl. Brit. W.I. [Grisebach]*: 135 [June 1859] (Grisebach 1859).

*Esenbeckia fasciculata* Barb.Rodr., *Rev. Engenh.* 5: 1 (Barbosa Rodrigues 1883).

*Esenbeckia grandiflora* var. *macrophylla* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 1285 (Chodat & Hassler 1904).

*Esenbeckia obovalifolia* Pittier, *Bol. Soc. Venez. Ci. Nat.* 9: 122 (Pittier 1944).

*Esenbeckia grandiflora* var. *peruviana* J.F.Macbr., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 13 (3.2): 673 (Macbride 1949).

*Esenbeckia rigida* R.S.Cowan, *Bol. Mus. Nac. Rio de Janeiro, Bot.* 27: 1 (Cowan 1961).

VERNACULAR NAMES. — Br: chupa-ferro, guaxupita, pitaguará-amarelo.

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: *M.-F. Prévost 3423*.

SIZE. — Up to 20 cm dbh (Kallunki 2005).

Genus *Galipea* Aubl.

[1507] *Galipea davisii* Sandwith

*Bull. Misc. Inform. Kew* 1935 (3): 118 [20 May 1935] (Sandwith 1935).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *C. Sastre & C. Moretti 3852* (P).

SIZE. — Up to 15 m tall (Kallunki 2005).

[1508] *Galipea trifoliata* Aubl.
(Fig. 51A)

Hist. Pl. Guiane 2: 662 [Jun.-Dec. 1775] (Aublet 1775).

*Galipea fissa* Miq., *Stirp. Surinam. Select.*: 64 ["1850" publ. Mar. 1851] (Miquel 1851).

VERNACULAR NAMES. — Ka: waikyara • Br: amarelinho, canela-develho, folha-de-couro.

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000630633]).

SIZE. — Up to 15 m tall (Pirani & Kallunki 2007).

Genus *Hortia* Vand.

[1509] *Hortia excelsa* Ducke
(Fig. 51B)

*Arch. Jard. Bot. Rio de Janeiro* 3: 182 (Ducke 1922).

VERNACULAR NAMES. — Nt: gaan busi tobitutu • Br: cachaceiro.

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *M.-F. Prévost & P. Grenand 3753*.

INVENTORY DATA (FG). — 1 tree, dbh = 10.8 cm.

[1510] *Hortia superba* Ducke

*Arch. Inst. Biol. Veg.* 1 (3): 207 [Aug. 1935] (Ducke 1935).

HERBARIUM DATA (FG). — A single collection, *J.-J. de Granville 17272*.

SIZE. — Up to 30 cm dbh (Groppo & Pirani 2012).

Genus *Neoraputia* Emmerich ex Kallunki

[1511] *Neoraputia paraensis* (Ducke) Emmerich ex Kallunki

*Brittonia* 61 (1): 29 [2 Mar. 2009] (Kallunki 2009). — *Raputia paraensis* Ducke, *Arch. Jard. Bot. Rio de Janeiro* 3: 184 (Ducke
1922). — *Neoraputia paraensis* (Ducke) Emmerich, *Rodriguésia* 30 (45): 243 (Emmerich 1978), *nom. inval.* (genus not validly published).

*Neoraputia cowanii* Emmerich, *Rodriguésia* 30 (45): 259 (Emmerich 1978), *nom. inval.* (genus not validly published).

VERNACULAR NAMES. — Br: capança, caporé.

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *C. Moretti 354*.

SIZE. — Up to 10 m tall (Kallunki 2005).

## Genus *Pilocarpus* Vahl

[1512] *Pilocarpus racemosus* Vahl

*Eclog. Amer.* 1: 29 (Vahl 1797).

*Pilocarpus latifolius* A.St.-Hil. ex Tul., *Ann. Sci. Nat., Bot.* sér. 3, 7: 285 (Tulasne 1847), “latifolia”.

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: *M.-F. Prévost et al. 4604*.

INVENTORY DATA (FG). — 25 trees in 3 plots; $F_{max} < 1\%$; $dbh_{inv} = 27.2$ cm.

## Genus *Raputia* Aubl.

[1513] *Raputia* sp. A

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier 2174*.

INVENTORY DATA (FG). — 19 trees in 6 plots; $F_{max} = 1.5\%$; $dbh_{inv} = 50$ cm.

## Genus *Ticorea* Aubl.

[1514] *Ticorea foetida* Aubl.
(Fig. 51C)

*Hist. Pl. Guiane* 2: 689 [Jun.-Dec. 1775] (Aublet 1775).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — 18 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000570872]).

SIZE. — Up to 14 m tall (Kallunki 1998).

[1515] *Ticorea longiflora* DC.

*Mém. Mus. Hist. Nat.* 9: 146 (Candolle 1822).

VERNACULAR NAMES. — Pa: kwik-wašiune, waravru-kamwi • Wp: yapukuliwa sili • Wn: malihaliimë.

HERBARIUM DATA (FG). — 18 collections at CAY. Sel. exs.: *J. Martin s.n.* (original material G-DC, G00310774]).

SIZE. — Up to 33 cm dbh (Kallunki 1998).

## Genus *Zanthoxylum* L.

[1516] *Zanthoxylum acuminatum* (Sw.) Sw. subsp. *juniperinum* (Poepp.) Reynel

*Phytoneuron* 2015-22: 6 (Reynel 2015).

*Zanthoxylum juniperinum* Poepp., *Nova genera ac species plantarum* [Poeppig & Endlicher] 3: 77 [23-25 Jan. 1845] (Poeppig 1845). — *Fagara juniperina* (Poepp.) Engl., *Nat. Pflanzenfam.* [*Engler & Prantl*] 3 (4): 117 (Engler 1896).

*Zanthoxylum minutiflorum* Tul., *Ann. Sci. Nat., Bot.* sér. 3, 7: 278 (Tulasne 1847). — *Fagara minutiflora* (Tul.) Engl., *Nat. Pflanzenfam.* [*Engler & Prantl*] 3 (4): 118 (Engler 1896).

*Zanthoxylum pringlei* S.Watson, *Proc. Amer. Acad. Arts* 26: 134 (Watson 1891), “Xanthoxylum”. — *Fagara pringlei* (S.Watson) Engl., *Nat. Pflanzenfam.* [*Engler & Prantl*] 3 (4): 117 (Engler 1896).

*Fagara warmingiana* Engl., *Nat. Pflanzenfam.* [*Engler & Prantl*] 3 (4): 117 (Engler 1896).

*Zanthoxylum procerum* Donn.Sm., *Bot. Gaz.* 23 (1): 4 [20 Jan. 1897] (Donnell Smith 1897). — *Fagara procera* (Donn.Sm.) Engl., *Nat. Pflanzenfam.* [*Engler & Prantl*], ed. 2, 19a: 223 (Engler 1931).

*Fagara acrea* K.Krause, *Notizbl. Königl. Bot. Gart. Berlin* 6: 143 [4 Apr. 1914] (Krause 1914). — *Zanthoxylum acreanum* (K.Krause) J.F.Macbr., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 13 (3.2): 664 (Macbride 1949).

*Fagara ocumarensis* Pittier, *Arb. Arbust. Venez.* 9-10: 120 [Dec. 1929] (Pittier 1929). — *Zanthoxylum ocumarensis* (Pittier) Steyerm., *Fieldiana*, *Bot.* 28 (2): 273 (Steyermark 1952).

VERNACULAR NAMES. — Ka: awaladala, awalatala.

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *S.A. Mori et al. 21661*.

INVENTORY DATA (FG). — 7 trees in 7 plots; $F_{max} < 1\%$; $dbh_{inv} = 45.2$ cm.

[1517] *Zanthoxylum amapaense* (Albuq.) P.G.Waterman

*Taxon* 24 (2/3): 363 [6 June 1975] (Waterman 1975). — *Fagara amapaense* Albuq., *Publ. Inst. Nac. Pesq. Amaz., Bot.* 28: 11 (Albu-querque 1968).

*Fagara idae* Albuq., *Publ. Inst. Nac. Pesq. Amaz., Bot.* 28: 8 (Albu-querque 1968). — *Zanthoxylum idae* (Albuq.) P.G.Waterman, *Taxon* 24 (2/3): 364 [6 June 1975] (Waterman 1975).

*Fagara machadoi* Albuq., *Acta Amazonica* 1 (1): 24 (Albuquerque 1971). — *Zanthoxylum machadoi* (Albuq.) Albuq., *Acta Amazonica* 6 (3, Suppl.): 55 [Sep. 1976] (Albuquerque 1976).

HERBARIUM DATA (FG). — A single collection, *J.-F. Molino & D. Sabatier 2349*.

INVENTORY DATA (FG). — 1 tree, dbh = 56.7 cm.
[1518] *Zanthoxylum apiculatum* (Sandwith) P.G.Waterman

Taxon 24 (2/3): 363 [6 June 1975] (Waterman 1975). — *Fagara apiculata* Sandwith, *Bull. Misc. Inform. Kew* 1935 (3): 120 [20 May 1935] (Sandwith 1935).

Fagara dellomei Albuq., *Publ. Inst. Nac. Pesq. Amaz.*, Bot. 28: 3 (Albu-
querque 1968). — *Zanthoxylum dellomei* (Albuq.) P.G.Waterman,
*Taxon* 24 (2/3): 363 [6 June 1975] (Waterman 1975).

Zanthoxylum achidek Sanoja, *Acta Bot. Venez.* 32 (2): 304 (Sanoja 2009).

HERBARIUM DATA (FG). — 14 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 2901*.

INVENTORY DATA (FG). — 14 trees in 12 plots; Fmax < 1%; dbhinv = 33.3 cm.

[1519] *Zanthoxylum caribaeum* Lam.

Encycl. [J. Lamarck et al.] 2 (1): 39 [16 Oct. 1786] (Lamarck 1786). — *Fagara caribaea* (Lam.) Krug & Urb., *Bot. Jahrb. Syst.* 21 (5): 562 [12 May 1896] (Krug & Urban 1896).

Zanthoxylum elephantiasis Macfad., *Fl. Jamaica* [Macfadyen] 1: 193 (Macfadyen 1837). — *Fagara elephantiasis* (Macfad.) Krug & Urb., *Bot. Jahrb. Syst.* 21 (5): 564 [12 May 1896] (Krug & Urban 1896).

Zanthoxylum horridum Sessé & Moc., *Fl. Mexic.*, ed. 2, 231 (Sessé & Mociño 1894), "Horrida".

Zanthoxylum occidentale Rose, *Contr. U.S. Natl. Herb.* 5 (4): 164 [31 Oct. 1899] (Rose 1899). — *Fagara occidentalis* (Rose) Engl., *Nat. Pflanzenfam.* [*Engler & Prantl*], ed. 2, 19a: 219 (Engler 1931).

Zanthoxylum gentlei Lundell, *Lloydia* 2 (2): 92 (Lundell 1939).

HERBARIUM DATA (FG). — A single collection, *H. de Foresta 125ter*.

SIZE. — Brazil, Pará. *G. T. Prance* 1822 (MO), 10 m × 12 cm.

[1520] *Zanthoxylum ekmanii* (Urb.) Alain
(Fig. 51D)

Contr. Ocas. Mus. Hist. Nat. Colegio De La Salle 9: 24 (Alain 1950). — *Fagara ekmanii* Urb., *Repert. Spec. Nov. Regni Veg.* 20: 302 (Urban 1924).

Zanthoxylum belicense Lundell, *Contr. Univ. Michigan Herb.* 6: 35 (Lundell 1941).

Zanthoxylum sobrevielae D.R.Simpson, *Phytologia* 51 (5): 316 [July 1982] (Simpson 1982).

VERNACULAR NAMES. — Wp: sala’i, sala’i sili • Cr: bwa-pidjan, zépini-gran-fey • Br: limãozinho, maminha-de-porco, tamanqueira.

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: S.A. Mori & J.J. Pipoly 15603.

INVENTORY DATA (FG). — 3 trees in 3 plots; Fmax < 1%; dbhinv = 27.2 cm.

[1521] *Zanthoxylum paulae* (Albuq.) P.G.Waterman

Taxon 24 (2/3): 364 [6 June 1975] (Waterman 1975). — *Fagara paulae* Albuq., *Publ. Inst. Nac. Pesq. Amaz.*, Bot. 27: 4 (Albuquerque 1968).

Fagara luizii Albuq., *Publ. Inst. Nac. Pesq. Amaz.*, Bot. 27: 5 (Albu-
querque 1968). — *Zanthoxylum luizii* (Albuq.) P.G.Waterman,
*Taxon* 24 (2/3): 364 [6 June 1975] (Waterman 1975).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *V. Hequet 3018bis*.

INVENTORY DATA (FG). — 1 tree, dbh = 66.9 cm.

[1522] *Zanthoxylum pentandrum* (Aubl.) R.A.Howard

J. Arnold Arbor. 64 (2): 269 [Apr. 1983] (Howard 1983). — *Fagara pentandra* Aubl., *Hist. Pl. Guiane* 1: 78 [Jun.-Dec. 1775] (Aublet 1775). — *Fagara guianensis* Lam., *Encycl. [J. Lamarck et al.]* 2 (2): 446 [14 Apr. 1788] (Lamarck 1788), *nom. illeg. superfl.* (based on *Fagara pentandra*). — *Zanthoxylum hermaphroditum* Willd., *Sp. Pl.*, ed. 4 4 (2): 756 [Apr. 1806] (Willdenow 1806), *nom. illeg. superfl.* (based on *Fagara pentandra*).

Zanthoxylum trinitense R.O.Williams, *Fl. Trinidad & Tobago* 1 (2): 148 [Nov. 1929] (Williams 1929). — *Fagara trinitensis* (R.Ö.Williams) Beard, *Empire For. J.* 21: 14 (Beard 1942).

VERNACULAR NAMES. — Pa: puduku-wakäu-purubumna, wahinhu
• Ka: alemikilan, awaladala, awalatala • Wp: sala'i u • Nt: kee udu,
kee maka udu • Cr: bwa-pidjan, zépini-gran-fey • Fr: bois piquant
• Br: limãozinho, maminha-de-porco, tamanqueira.

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: J.B. Aublet s.n. (lecto-, P-JJR[P00777615] designated by Lanjouw & Uittien [1940: 151]).

INVENTORY DATA (FG). — 4 trees in 4 plots; Fmax < 1%; dbhinv = 31.7 cm.

[1523] *Zanthoxylum rhoifolium* Lam.

Encycl. [J. Lamarck et al.] 2 (1): 39 [16 Oct. 1786] (Lamarck 1786). — *Fagara rhoifolia* (Lam.) Engl., *Nat. Pflanzenfam.* [*Engler & Prantl*] 3 (4): 118 (Engler 1896).

Langsdorfia instrumentaria Leandro, *Denkschr. Königl. Akad. Wiss. München* 7: 242 (Leandro 1819). — *Pohlana langsdorffii* Nees & Mart., *Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur.* 12: 17 (Nees & Martius 1824), *nom. illeg. superfl.* (based on *Langsdorfia instrumentaria*). — *Zanthoxylum langsdorffii* A.St.-Hil., *Fl. Bras. Merid. [A. St.-Hil.]* (quarto ed.) 1 (2): 76 [26 Mar. 1825] (Saint-Hilaire 1825), *nom. illeg. superfl.* (based on *Pohlana langsdorffii*, thus indirectly on *Langsdorfia instrumentaria*).

Zanthoxylum perrottetii DC., *Prodr. [A. P. de Candolle]* 1: 726 [mid Jan. 1824] (Candolle 1824), “perrotetii”. — *Fagara perrottetii* (DC.) Lemée, *Fl. Guyane Franç.* 2: 181 (Lemée 1952).

Zanthoxylum sorbifolium A.St.-Hil., *Fl. Bras. Merid. [A. St.-Hil.]* (quarto ed.) 1 (2): 75 [26 Mar. 1825] (Saint-Hilaire 1825).

Pohlana instrumentaria Mart. ex Steudel, *Nomencl. Bot. [Steudel]*, ed. 2, 2: 366, 796, (Steudel 1841) *nom. nud*.

Zanthoxylum pubescens A.St.-Hil. & Tul., *Ann. Sci. Nat.*, Bot. sér. 2, 17: 141 (Saint-Hilaire & Tulasne 1842). — *Schinus pubescens*
<figure><img src="image_73.png" /><figcaption>FIG. 50. — Rubiaceae: **A, Ferdinandusa** sp. A (D. Sabatier & M.-F. Prévost 4928); **B, Posoqueria longiflora** Aubl.; **C, Rudgea lanceifolia** Salisb.; **D, Kutchubaea insignis** Fisch. ex DC. (M.-F. Prévost et al. 4734). A, © M.-F. Prévost/IRD; B, © P. G. Delprete/IRD; C, D, © D. Sabatier/IRD.</figcaption></figure>

(A.St.-Hil. & Tul.) Spreng. ex Mart., *Fl. Bras.* [Martius] 12 (2): 176 [1 Sep. 1874] (Martius 1874), *nom. nud. pro syn.* — *Zanthoxylum rhoifolium* var. *pubescens* (A.St.-Hil. & Tul.) Engl., *Fl. Bras.* [Martius] 12 (2): 176 [1 Sep. 1874] (Engler 1874). — *Fagara rhoifolia* subsp. *pubescens* (A.St.-Hil. & Tul.) Engl., *Nat. Pflanzenfam.* [Engler & Prantl] 3 (4): 118 (Engler 1896). — *Fagara pubescens* (A.St.-Hil. & Tul.) Herzog, *Meded. Rijks-Herb.* 27: 80 (Herzog 1915), *nom. illeg. hom.*, *non A.Chev*. (Chevalier 1912).

*Zanthoxylum peltophorum* Turcz., *Bull. Soc. Imp. Naturalistes Moscou* 31 (1): 442 (Turczaninow 1858). — *Zanthoxylum rhoifolium* var. *peltophorum* (Turcz.) Engl., *Fl. Bras.* [Martius] 12 (2): 175 [1 Sep. 1874] (Engler 1874). — *Fagara rhoifolia* var. *peltophora* (Turcz.) Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 1283 (Chodat & Hassler 1904), “*peltophorum*”.

*Zanthoxylum microcarpum* Griseb., *Fl. Brit. W.I. (Grisebach)*: 138 [June 1859] (Grisebach 1859). — *Fagara microcarpa* (Griseb.) Krug & Urb., *Bot. Jahrb. Syst.* 21 (5): 570 [12 May 1896] (Krug & Urban 1896).

*Zanthoxylum obscurum* Engl., *Fl. Bras.* [Martius] 12 (2): 169 [1 Sep. 1874] (Engler 1874). — *Fagara obscura* (Engl.) Engl., *Nat. Pflanzenfam.* [Engler & Prantl] 3 (4): 117 (Engler 1896).

*Zanthoxylum acutifolium* Engl., *Fl. Bras.* [Martius] 12 (2): 170 [1 Sep. 1874] (Engler 1874). — *Fagara acutifolia* (Engl.) Engl., *Nat. Pflanzenfam.* [Engler & Prantl] 3 (4): 117 (Engler 1896).

*Zanthoxylum acutifolium* var. *petiolulatum* Engl., *Fl. Bras.* [Martius] 12 (2): 170 [1 Sep. 1874] (Engler 1874). — *Fagara rhoifolia* var. *petiolulata* (Engl.) Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 1283 (Chodat & Hassler 1904), “*petiolulatum*”.

*Zanthoxylum obscurum* var. *ruizianum* Klotzsch ex Engl., *Fl. Bras.* [Martius] 12 (2): 170 [1 Sep. 1874] (Engler 1874). — *Fagara ruiziana* (Klotzsch ex Engl.) Engl., *Nat. Pflanzenfam.* [Engler & Prantl] 3 (4): 117 (Engler 1896). — *Zanthoxylum ruizianum* (Klotzsch ex Engl.) J.F.Macbr., *Publ. Field Mus. Nat. Hist., Bot. Ser. 13* (3.2): 667 (Macbride 1949).
*Zanthoxylum regnellianum* Engl., *Fl. Bras.* [*Martius*] 12 (2): 174 [1 Sep. 1874] (Engler 1874). — *Fagara regnelliana* (Engl.) Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 1283 (Chodat & Hassler 1904).

*Zanthoxylum rhoifolium* var. *petiolulatum* Engl., *Fl. Bras.* [*Martius*] 12 (2): 175 [1 Sep. 1874] (Engler 1874).

*Zanthoxylum rhoifolium* var. *sessilifolium* Engl., *Fl. Bras.* [*Martius*] 12 (2): 176 [1 Sep. 1874] (Engler 1874).

*Fagara regnelliana* var. *calvata* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 1283 (Chodat & Hassler 1904).

*Fagara rhoifolia* f. *paucijuga* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 1283 (Chodat & Hassler 1904).

*Fagara rhoifolia* f. *angustifolia* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 1284vt (Chodat & Hassler 1904).

*Fagara rhoifolia* f. *intermedia* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 1284 (Chodat & Hassler 1904).

*Fagara rhoifolia* f. *latifolia* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 1284 (Chodat & Hassler 1904).

*Fagara coco* (Gillies ex Hook.f. & Arn.) Engl. var. *formosana* Lillo, *Contr. Conoc. Arb. Argent.*: 89 (Lillo 1910). — *Zanthoxylum rhoifolium* var. *formosanum* (Lillo) P.G.Waterman, *Taxon* 25 (5/6): 594 [26 Nov. 1976] (Waterman 1976).

*Fagara rothschuhii* Loes., *Bot. Jahrb. Syst.* 60 (4): 366 [1 Aug. 1926] (Loesener 1926).

*Fagara astrigera* R.S.Cowan, *Sellowia* 12: 81 (Cowan 1960). — *Zanthoxylum astrigerum* (R.S.Cowan) P.G.Waterman, *Taxon* 24 (2/3): 363 [6 June 1975] (Waterman 1975).

*Fagara rhoifolia* var. *surparanaensis* Najera, Galdeano & Escal., *Bot. Soc. Argent. Bot.* 14 (3): 244 (Najera et al. 1972). — *Zanthoxylum rhoifolium* var. *surparanaense* (Najera, Galdeano & Escal.) P.G.Waterman, *Taxon* 25 (5/6): 594 [26 Nov. 1976] (Waterman 1976).

*Fagara rhoifolia* var. *intermedia* R.S.Cowan & L.B.Sm., *Fl. Ilustr. Catar.* (Ruta.): 13 (Cowan & Smith 1973).

**VERNACULAR NAMES.** — Pa: puduku-wakáu • Ka: awaladala, awalatala • Te: wila atsi • Wn: kanike • Nt: kee udu, kee maka udu • Cr: bwa-pidjan, zépini-ti'fey • Fr: bois piquant • Br: mamiqueira, tamanqueira, tembetaru.

**HERBARIUM DATA (FG).** — 44 collections at CAY. Sel. exs.: *M.-F. Prévost 3781*, dbh 10 cm.

Family SABIACEAE Blume
Genus Meliosma Blume

[1524] *Meliosma* sp. A
(Fig. 51E)

**HERBARIUM DATA (FG).** — 4 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 5672*.

**INVENTORY DATA (FG).** — 5 trees in 4 plots; $F_{max} < 1\%$; dbh$_{inv}$ = 31.3 cm.

Genus *Ophiocaryon* Endl.

[1525] *Ophiocaryon chironectes* Barneby

Mem. New York Bot. Gard. 23: 117 (Barneby 1972).

VERNACULAR NAME. — Wp: tāpē lemiū’i.

HERBARIUM DATA (FG). — A single collection, *P. Grenand 1543*. The type is from Clevelandia, Amapá, on the Brazilian border of the Oyapock River: *L.Y.T. Westra 47298* (NY[00387482]).

INVENTORY DATA (FG). — 1 tree, dbh = 10.8 cm.

[1526] *Ophiocaryon paradoxum* R.H.Schomb.

London *J. Bot.* 4: 377 (Schomburgk 1845).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — A single collection, *C. Sastre 4704*.

SIZE. — Guyana. *B. Maguire 22937* (MO), dbh 30 cm.

Family SALICACEAE Mirb.
Genus *Banara* Aubl.

[1527] *Banara guianensis* Aubl.

*Hist. Pl. Guiane* 1: 548 [Jun.-Dec. 1775] (Aublet 1775), “*Guyannensis*” on plate.

*Banara fagifolia* Vahl, *Symb. Bot.* [Vahl] 3: 65 (Vahl 1794), *nom. illeg. superfl.* (based on *Banara guianensis*).

*Xyladenius glandulosus* Desv. ex Ham., *Prodr. Pl. Ind. Occid.* [*Hamilton*]: 41 [Oct. 1825] (Hamilton 1825). — *Banara glandulosa* (Desv. ex Ham.) Speg., *Revista Argent. Bot.* 1: 210 (Spegazzini 1926).

*Kuhlia mollis* Poepp., *Nova genera ac species plantarum* [*Poeppig & Endlicher*] 3: 74 [23-25 Jan. 1845] (Poeppig 1845). — *Banara mollis* (Poepp.) Tul., *Ann. Sci. Nat.*, *Bot.* sér. 3, 7: 288 (Tulasne 1847). — *Banara guianensis* var. *mollis* (Poepp.) Eichler, *Fl. Bras.* [*Martius*] 13 (1): 501 [1 Oct. 1871] (Eichler 1871).

*Laetia glandulosa* Poepp. ex Tul., *Ann. Sci. Nat.*, *Bot.* sér. 3, 7: 288 (Tulasne 1847), *nom. nud. pro syn.*

*Trilix glandulosa* Dombey ex Tul., *Ann. Sci. Nat.*, *Bot.* sér. 3, 7: 288 (Tulasne 1847), “*Trilin*”, *nom. nud. pro syn.*

*Banara pubescens* Spruce ex Benth., *J. Proc. Linn. Soc.*, *Bot.* 5 (Suppl. 2): 92 [May 1861] (Bentham 1861).

*Banara pyramidata* Rusby, *Mem. Torrey Bot. Club* 3: 33 (Rusby 1893).

*Banara guianensis* var. *spruceana* Briq., *Annuaire Conserv. Jard. Bot. Genève* 2: 49 (Briquet 1898).

*Banara tulasnei* J.F.Macbr., *Candollea* 5: 389 (Macbride 1934).

*Trilix macrobotrys* Ruiz & Pav., *Anales Inst. Bot. Cavanilles* 16: 374 (Ruiz & Pavón 1959).

**VERNACULAR NAMES.** — Pa: kuuku-ariut-priye, tahuma • Wp: yape'a pitā sili • Wn: kumidan • Nt: bita tiki, boni udu • Cr: mavévé-sikriyé • Br: cavaqueiro-do-baixo, lacre-branco.

**HERBARIUM DATA (FG).** — 109 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material P-JU, P00672058]).

INVENTORY DATA (FG). — 6 trees in 3 plots; dbh$_{inv}$ = 16.4 cm.
## Genus *Casearia* Jacq.

[1528] *Casearia arborea* (Rich.) Urb.

Symb. Antill. [Urban] 4 (3): 421 [15 May 1910] (Urban 1910). — *Samyda arborea* Rich., *Actes Soc. Hist. Nat. Paris* 1: 109 [Oct. 1792] (Richard 1792). — *Casearia stipularis* Vent., *Choix Pl.* 1: 46 [Apr. 1804] (Ventenat 1804), *nom. illeg. superfl.* (based on *Samyda arborea*). — *Samyda stipularis* Poir., *Encycl. [J. Lamarck et al.]* Suppl. 5: 32 [1 Nov. 1817] (Poiret 1817), *nom. illeg. superfl.* (based on *Samyda arborea*). — *Casearia stipularis* Poir. ex Spreng., *Syst. Veg. [Sprengel]* 2: 355 [Jan.-May 1825] (Sprengel 1825), *nom. illeg. superfl.* (based on *Samyda stipularis*, thus on *Samyda arborea*). — *Guidonia stipularis* M.Gómez, *Dicc. Bot. Nombres Vulg. Cub. Puerto-Riq.* 54 (Gómez 1889), *nom. illeg. superfl.* (based on *Casearia stipularis* Vent., thus on *Samyda arborea*).

*Samyda viridiflora* Aubl., *Hist. Pl. Guiane* 1: 402 [Jun.-Dec. 1775] (Aublet 1775).

*Chaetocrater capitatum* Ruiz & Pav., *Syst. Veg. Fl. Peruv. Chil.* 1: 108 [late Dec. 1798] (Ruiz & Pavón 1798). — *Casearia capitata* (Ruiz & Pav.) Pers., *Syn. Pl. [Persoon]* 1: 485 [1 Apr.-15 June 1805] (Persoon 1805).

*Samyda niviana* Poir., *Encycl. [J. Lamarck et al.]* 6 (2): 493 [28 Aug. 1805] (Poiret 1805).

*Casearia incana* Bertero ex Spreng., *Syst. Veg. [Sprengel]* 2: 355 [Jan.-May 1825] (Sprengel 1825).

*Casearia serrata* Macfad., *Fl. Jamaica [Macfadyen]* 1: 216 (Macfadyen 1837).

*Casearia lanceolata* Miq., *Linnaea* 18: 753 ["1844" publ. Aug.-Oct. 1845] (Miquel 1845).

*Casearia hostmanniana* Steud. ex Griseb., *Fl. Brit. W.I. [Grisebach]* 23 [Dec. 1859] (Grisebach 1859), *nom. nud. pro syn.*

*Casearia poeppigii* Eichler, *Fl. Bras. [Martius]* 13 (1): 475 [1 Oct. 1871] (Eichler 1871).

*Casearia cambessedesii* var. *angustifolia* Eichler, *Fl. Bras. [Martius]* 13 (1): 476 [1 Oct. 1871] (Eichler 1871).

*Casearia cambessedesii* var. *parvifolia* Eichler, *Fl. Bras. [Martius]* 13 (1): 476 [1 Oct. 1871] (Eichler 1871).

*Casearia oligantha* Eichler, *Fl. Bras. [Martius]* 13 (1): 476 [1 Oct. 1871] (Eichler 1871).

*Casearia brasiliensis* Eichler, *Fl. Bras. [Martius]* 13 (1): 477 [1 Oct. 1871] (Eichler 1871).

*Casearia bangii* Rusby, *Mem. Torrey Bot. Club* 3: 34 (Rusby 1893).

*Samyda virgata* Sessé & Moc., *Fl. Mexic.*, ed. 2, 113 (Sessé & Mociño 1894).

*Casearia glaziovii* Briq., *Annuaire Conserv. Jard. Bot. Genève* 2: 72 (Briquet 1898).

*Casearia umbellifera* Benoist, *Bull. Mus. Natl. Hist. Nat.* 26: 353 (Benoist 1920).

*Casearia glaberrima* Uittien, *Recueil Trav. Bot. Néerl.* 22: 371 ["1925" publ. Jan. 1926] (Uittien 1926).

NOTE. — *Samyda virgata* Sessé & Moc. appears also in the first edition of *Flora mexicana* (p. 123, as "*Uirgata*"), but edition 2 has

priority, probably from p. 49 onward (Stafleu & Cowan, 1985: TL-2-11756)].

VERNACULAR NAMES. — Ka: akukuwa elepali, pilitalaipyo • Wp: yape’a pitā sili • Wn: luweluwe • Br: capança-braba, carniceiro, erva-de-lagarto.

HERBARIUM DATA (FG). — 46 collections at CAY. Sel. exs.: *J.B. Leblond s.n.* (in herb. L.C. Richard) (type P[P00371634]).

INVENTORY DATA (FG). — 1 tree, dbh = 14 cm.

[1529] *Casearia bicolor* Urb.

Symb. Antill. [Urban] 1 (2): 372 [10 Apr. 1899] (Urban 1899).

*Laetia procera* (Poepp.) Eichler, *Fl. Bras. [Martius]* 13 (1): 453 [1 Oct. 1871] (Eichler 1871). — *Samyda procera* Poepp., *Nova genera ac species plantarum [Poeppig & Endlicher]* 3: 67 [23-25 Jan. 1845] (Poeppig 1845). — *Casinga procera* (Poepp.) Griseb., *Fl. Brit. W.I. [Grisebach]* 710 [Oct. 1864] (Grisebach 1864). — *Guidonia procera* (Poepp.) Kuntze, *Revis. Gen. Pl.* 1: 44 [5 Nov. 1891] (Kuntze 1891).

*Laetia casearioides* Sagot ex Benth., *J. Proc. Linn. Soc.*, Bot. 5 (Suppl. 2): 85 [May 1861] (Bentham 1861).

*Samyda obtusifolia* Rich. ex Eichler, *Fl. Bras. [Martius]* 13 (1): 454 [1 Oct. 1871] (Eichler 1871), in obs.

*Laetia obtusifolia* Eichler ex Benoist, *Bull. Mus. Natl. Hist. Nat.* 26: 351 (Benoist 1920), *nom. nud.*

*Casearia belizensis* Standl., *Publ. Field Mus. Nat. Hist.*, Bot. Ser. 12: 412 (Standley 1936).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020). The genus *Laetia* Loefl. ex L. was included in *Casearia* Jacq. by Samarakoon & Alford (2019).

VERNACULAR NAMES. — Pa: ahakyu, ahikyu • Ka: akale wewe, alokoyulu, auluko yulu, kayman udu, mainyapo • Te: tale’i li • Wp: miyǔ’i pepo, muyǔ’i pepo, tale’i li • Wn: mijelemişi • Nt: kayman udu • Cr: bwa-jako, bwa-zanmand • Fr: bois caïman • Br: apijó, casinga-cheirosa, muira-pucu, pau-jacaré, tuchauá.

HERBARIUM DATA (FG). — 79 collections at CAY. Sel. exs.: *P.A. Sagot 1137, 1857* (original material of *Laetia casearioides*: K[K000471402, K000471403], P[P02442044, P02442045, P02442046, P02442047]).

INVENTORY DATA (FG). — 153 trees in 88 plots; $F_{max} = 1.2\%$; dbh$_{inv} = 94.2$ cm.

[1530] *Casearia decandra* Jacq.

* Enum. Syst. Pl.*: 21 [Aug.-Sep. 1760] (Jacquin 1760). — *Casearia parviflora* Jacq., *Select. Stirp. Amer. Hist.*, ed. 1780-1781, 66 (Jacquin 1780), *nom. illeg. superfl.* (based on the illustration of *Casearia decandra*). — *Casearia parvifolia* Willd., *Sp. Pl.*, ed. 4 2 (1): 627 [Mar. 1799] (Willdenow 1799), *nom. illeg. superfl.* (based on *Casearia decandra*). — *Samyda parvifolia* Poir., *Encycl. [J. Lamarck et al.]* 6 (2): 492 [28 Aug. 1805] (Poiret 1805), *nom. illeg. superfl.* (based on *Casearia decandra*). — *Anavinga parvifolia* (Poir.) Lam., *Tabl. Encycl.* 2[5 (2)]: 484 [31 Oct. 1819] (Lamarck 1819), *nom. illeg. superfl.* (based on *Samyda parvifolia*, thus on *Casearia decandra*). — *Guidonia parvifolia* M.Gómez, *Dicc. Bot. Nombres Vulg.*
*Cub. Puerto-Rig.*: 93 (Gómez 1889), *nom. illeg. superfl.* (based on *Casearia parvifolia*, thus on *Casearia decandra*).

*Casearia adamantium* Cambess., *Fl. Bras. Merid. [A. St.-Hil.]* (quarto ed.) 2 (16): 230 [“1829” publ. Oct.-Nov. 1830] (Cambessèdes 1830).

*Casearia adstringens* Mart., *Syst. Mat. Med. Veg. Bras.* 51 (Martius 1843), *nom. nud.*

*Casearia nitida* Sieber ex Griseb., *Fl. Brit. W.I. [Grisebach]* 23 [Dec. 1859] (Grisebach 1859), *nom. nud. pro syn.*

*Casearia serrulata* Sieber ex Griseb., *Fl. Brit. W.I. [Grisebach]* 23 [Dec. 1859] (Grisebach 1859), *nom. nud. pro syn.*

*Casearia parvifolia* var. microcarpa Eggers, *Bull. U.S. Natl. Mus.* 13: 26 (Eggers 1879).

*Guidonia adstringens* Baill., *Traité Bot. Méd. Phan.* 2: 827 [Mar. 1884] (Baillon 1884).

*Samyda lancifolia* Sessé & Moc., *Fl. Mexic.*, ed. 2, 113 (Sessé & Mociño 1894).

*Casearia floribunda* Briq., *Bull. Herb. Boissier*, sér. 2, 7: 672 (Briquet 1907).

*Casearia parvifolia* var. paraguariensis Briq., *Bull. Herb. Boissier*, sér. 2, 7: 672 (Briquet 1907).

*Casearia albicaulis* Rusby, *Mem. New York Bot. Gard.* 7: 307 (Rusby 1927).

*Casearia pavoniana* Sleumer, *Notizbl. Bot. Gart. Berlin-Dahlem* 11: 958 [20 Jan. 1934] (Sleumer 1934).

*Casearia reflexa* Sleumer, *Notizbl. Bot. Gart. Berlin-Dahlem* 11: 959 [20 Jan. 1934] (Sleumer 1934).

*Chaetocrater reflexus* Ruiz & Pav., *Anales Inst. Bot. Cavanilles* 15: 130 (Ruiz & Pavón 1958), “reflexum”.

NOTES. — Jacquin (1780: 66) cited *Samyda parviflora* L. “Syst. 340”, in synonymy under *Casearia parviflora*. Jacq., but his plate 127 is the same as plate 85 in the first edition of *Selectarum stirpium Americannarum historia* (Jacquin 1763), which illustrates *C. decandra* Jacq. (p. 133). Although *Samyda lancifolia* Sessé & Moc. already appeared in the first edition of *Flora mexicana* (Sessé & Mociño 1892-1898: 123), edition 2 has priority, probably from p. 49 onward (Stafleu & Cowan, 1985: TL-2-11756). *Guidonia adstringens* Baill. is based on the invalid *Casearia adstringens* Mart.

VERNACULAR NAMES. — Br: caferana, café-do-mato.

HERBARIUM DATA (FG). — 17 collections at CAY. Sel. exs.: *M.-F. Prévost 3064*.

INVENTORY DATA (FG). — 6 trees in 6 plots; $F_{max} = 1.3\%$; $dbh_{inv} = 16.9$ cm.

[1531] *Casearia grandiflora* Cambess.
(Fig. 52A)

*Fl. Bras. Merid. [A. St.-Hil.]* (quarto ed.) 2 (16): 232 [“1829” publ. Oct.-Nov. 1830] (Cambessèdes 1830).

*Casearia hypoleuca* Mart., *Flora* 20 (2, Beibl.): 128 (Martius 1837). — *Casearia grandiflora* var. hypoleuca (Mart.) Eichler, *Fl. Bras. [Martius]* 13 (1): 479 [1 Oct. 1871] (Eichler 1871).

*Casearia grandiflora* var. obtrusifolia Eichler, *Fl. Bras. [Martius]* 13 (1): 479 [1 Oct. 1871] (Eichler 1871).

*Casearia grandiflora* var. pauciflora Eichler, *Fl. Bras. [Martius]* 13 (1): 479 [1 Oct. 1871] (Eichler 1871).

*Casearia anavinga* A.St.-Hil. ex Sagot, *Ann. Sci. Nat., Bot.* sér. 6, 13: 285 (Sagot 1882).

HERBARIUM DATA (FG). — 25 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 5187*.

INVENTORY DATA (FG). — 5 trees in 4 plots; $F_{max} < 1\%$; $dbh_{inv} = 21.7$ cm.

[1532] *Casearia guianensis* (Aubl.) Urb.

*Symb. Antill. [Urban]* 3 (2): 322 [15 Aug. 1902] (Urban 1902). — *Iroucana guianensis* Aubl., *Hist. Pl. Guiane* 1: 329 [Jun.-Dec. 1775] (Aublet 1775), “Guyannensis” on plate. — *Athenaea guianensis* (Aubl.) J.F.Gmel., *Syst. Nat.*, ed. 13[bis], 2 (1): 629 [late Sep.-Nov. 1791] (Gmelin 1791). — *Casearia ramiflora* Vahl, *Symb. Bot. [Vahl]* 2: 50 [Jul.-Dec. 1791] (Vahl 1791), *nom. illeg. superfl.* (based on *Iroucana guianensis*). — *Samyda iroucana* Rich., *Actes Soc. Hist. Nat. Paris* 1: 109 [Oct. 1792] (Richard 1792), *nom. illeg. superfl.* (based on *Iroucana guianensis*). — *Guidonia ramiflora* M.Gómez, *Dicc. Bot. Nombres Vulg. Cub. Puerto-Rig.* 64 (Gómez 1889), *nom. illeg. superfl.* (based on *Casearia ramiflora*, thus indirectly on *Iroucana guianensis*).

*Casearia fallax* Miq., *Linnaea* 19: 128 [“1847” publ. Feb. 1846] (Miquel 1846).

*Samyda octandra* Sessé & Moc., *Fl. Mexic.*, ed. 2, 113 (Sessé & Mociño 1894).

NOTE. — *Samyda octandra* Sessé & Moc. appears also in the first edition of *Flora mexicana* (Sessé & Mociño 1892-1898: 123), but edition 2 has priority, probably from p. 49 onward (Stafleu & Cowan, 1985: TL-2-11756)].

VERNACULAR NAMES. — Ka: alekikolan.

HERBARIUM DATA (FG). — 30 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material P[P00689737]).

INVENTORY DATA (FG). — 1 tree, $dbh = 16.2$ cm.

[1533] *Casearia mariquitensis* Kunth

*Nova genera et species plantarum [H.B.K.]* 5: 363 [24 Mar. 1823] (Kunth 1823).

*Casearia fockeana* Miq., *Ann. Sci. Nat., Bot.* sér. 3, 1: 39 (Miquel 1844).

*Casearia pubiflora* Benth., *Bot. Voy. Sulphur [Bentham]*: 66 [16 Aug. 1844] (Bentham 1844).

*Casearia platyphylla* Briq., *Annuaire Conserv. Jard. Bot. Genève* 2: 68 (Briquet 1898), *pro parte quoad floram tantum* (Sleumer 1980).

*Casearia camporum* Sprague, *Trans. & Proc. Bot. Soc. Edinburgh* 22 (4): 427 (Sprague 1905).

*Casearia tarapotina* Pilg., *Verh. Bot. Vereins Prov. Brandenburg* 47: 161 [1 Oct. 1905] (Pilger 1905).
<figure><img src="image_74.png" /><figcaption>FIG. 51. — Rutaceae: **A**, Galipea trifoliata Aubl. (J.-F. Molino & D. Sabatier 2164); **B**, Hortia excelsa Ducke (D. Sabatier & M.-F. Prévost 4935); **C**, Ticorea foetida Aubl. (J.-F. Molino et al. 2041); **D**, Zanthoxylum ekmanii (Urb.) Alain (D. Sabatier & J.-F. Molino 5747). Sabiaceae: **E**, Meliosma sp. A (D. Sabatier & J.-F. Molino 5672). A, © J.-F. Molino/IRD; B-E, © D. Sabatier/IRD.</figcaption></figure>

**HERBARIUM DATA (FG).** — A single collection *fide* Sleumer (1980): *L. C. Richard s.n.* (P). This collection could not be located with certainty, but could correspond to P04821776 or P04821777.

**SIZE.** — Up to 60 cm dhb (Sleumer 1980).

[1534] *Casearia negrensis* Eichler

Fl. Bras. [Martius] 13 (1): 466 [1 Oct. 1871] (Eichler 1871).

**VERNACULAR NAMES.** — Pa: waaduk.

**HERBARIUM DATA (FG).** — 27 collections at CAY. Sel. exs.: *R.A.A. Oldeman* 3191.

**INVENTORY DATA (FG).** — 2 trees in 2 plots; $F_{max} < 1\%$; $dhb_{inv}$ 14.3 cm.

[1535] *Casearia pitumba* Sleumer

*Blumea* 24: 118 (Sleumer 1978). — *Pitumba guianensis* Aubl., *Hist. Pl. Guiane* 2 (Suppl.): 29 [Jun.-Dec. 1775] (Aublet 1775). — *Samyda pitumba* Poir., *Encycl. [J. Lamarck et al.]* 6 (2): 492 [28 Aug. 1805] (Poiret 1805), *nom. illeg. superfl.* (based on *Pitumba guianensis*). — *Casearia macrophylla* Vahl, *Eclog. Amer.* 2: 32 (Vahl 1798), *nom. illeg. superfl.* (based on *Pitumba guianensis*). — *Casearia microphylla* Dennst., *Nomencl. Bot. [Dennstedt]* 1: 94 (Dennstedt 1810), *nom. illeg. superfl.* (based on *Pitumba guianensis*).

*Pitumba edulis* A.Rich. ex Eichler, *Fl. Bras. [Martius]* 13 (1): 470 [1 Oct. 1871] (Eichler 1871), *nom. nud. pro syn.*

*Casearia timbuchi* J.F.Macbr., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 15: 358 (Macbride 1936), *nom. inval.* (anglice).

*Casearia macrophylla* var. *barbatula* J.F.Macbr., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 13 (4.1): 45 (Macbride 1941).
NOTE. — *Casearia microphylla* is an orthographic variant that possibly originated from a typographical error (*macrophylla* intended).

VERNACULAR NAMES. — Pa: agagut-aška, ararut-aška, tahuma-purubumna, tahuma-seine • Ka: pelepele apotokon • Wp: yape'a pitā, yape'a pitā u • Wn: luweluwe • Nt: kwasikwasi tiki • Cr: grënn-oko, grënn-toti • Br : cabelo-de-cotia.

HERBARIUM DATA (FG). — 156 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000611049]).

INVENTORY DATA (FG). — 12 trees in 10 plots; Fmax < 1%; dbhinv = 22.3 cm.

[1536] *Casearia prunifolia* Kunth

*Nova genera et species plantarum [H.B.K.]* 5: 364 [24 Mar. 1823] (Kunth 1823).

*Casearia mathewsii* Turcz., *Bull. Soc. Imp. Naturalistes Moscou* 36 (1): 608 (Turczaninow 1863).

HERBARIUM DATA (FG). — A single collection, *C. Feuillet et al. 10319*.

SIZE. — Up to 20 m tall (Olson *et al.* 1999).

[1537] *Casearia rusbyana* Briq.

*Annuaire Conserv. Jard. Bot. Genève* 2: 73 (Briquet 1898).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Pa: ahakyu, ahikyu.

HERBARIUM DATA (FG). — 25 collections at CAY. Sel. exs.: *D. Sabatier 2321*.

INVENTORY DATA (FG). — 6 trees in 5 plots; Fmax < 1%; dbhinv = 29.4 cm.

[1538] *Casearia singularis* Eichler

*Fl. Bras. [Martius]* 13 (1): 473 [1 Oct. 1871] (Eichler 1871).

HERBARIUM DATA (FG). — 26 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 5642*.

INVENTORY DATA (FG). — 3 trees in 3 plots; Fmax < 1%; dbhinv = 11.5 cm.

[1539] *Casearia sylvestris* Sw.

*Fl. Ind. Occid.* 2: 752 [Jan.-June 1798] (Swartz 1798). — *Samyda sylvestris* (Sw.) Poir., *Encycl. [J. Lamarck et al.]* 6 (2): 492 [28 Aug. 1805) (Poiret 1805), "silvestris". — *Guidonia sylvestris* (Sw.) M.Gómez, *Dicc. Bot. Nombres Vulg. Cub. Puerto-Rig.* 95 (Gómez 1889).

*Samyda parviflora* L., *Syst. Nat.*, ed. 10, 2: 1025 [7 June 1759] (Linnaeus 1759), *nom. illeg. hom.*, *non* Loeffl. (Loefling 1758, *nom. rej.*, *nec* Sessé & Moc. (Sessé & Mociño 1894).

*Casearia parviflora* J.F.Gmel., *Syst. Nat.*, ed. 13[bis], 2 (1): 700 [late Sep.-Nov. 1791] (Gmelin 1791), *nom. illeg. hom.*, *non* Jacq. (Jacquin 1780).

*Casearia serrulata* Sw., *Fl. Ind. Occid.* 2: 754 [Jan.-June 1798] (Swartz 1798).

*Casearia parviflora* Willd., *Sp. Pl.*, ed. 4 2 (1): 627 [Mar. 1799] (Willdenow 1799), *nom. illeg. hom.*, *non* Jacq. (Jacquin 1780).

*Anavinga samyda* C.F.Gaertn., *Suppl. Carp.* 241 (Gaertner 1807). — *Casearia samyda* (C.F.Gaertn.) DC., *Prodr. [A. P. de Candolle]* 1: 51 [mid Jan. 1824] (Candolle 1824).

*Casearia punctata* Spreng., *Neue Entdeck. Pflanzenk.* 2: 154 (Sprengel 1821).

*Casearia sylvestris* var. *platyphylla* DC., *Prodr. [A. P. de Candolle]* 2: 49 [mid Nov. 1825] (Candolle 1825).

*Casearia parviflora* var. *microphylla* Schltdl., *Linnaea* 4: 90 (Schlechtendal 1829).

*Casearia affinis* Gardner, *London J. Bot.* 1: 529 (Gardner 1842).

*Casearia benthamiana* Miq., *Ann. Sci. Nat., Bot.* sér. 3, 1: 38 (Miquel 1844). — *Casearia sylvestris* var. *benthamiana* (Miq.) Uittien, *Recueil Trav. Bot. Néerl.* 22: 373 ["1925" publ. Jan. 1926] (Uittien 1926).

*Casearia sylvestris* var. *myricoides* Griseb., *Cat. Pl. Cub. [Grisebach]* 9 [May-Aug. 1866] (Grisebach 1866).

*Samyda parviflora* Sessé & Moc., *Fl. Mexic.*, ed. 2, 112 (Sessé & Mociño 1894), *nom. illeg. hom.*, *non* Loeffl. (Loefling 1758, *nom. rej.*, *nec* L. (Linnaeus 1759).

*Casearia attenuata* Rusby, *Mem. Torrey Bot. Club* 6 (1): 41 (Rusby 1896).

*Casearia sylvestris* var. *tomentella* Rusby, *Mem. Torrey Bot. Club* 6 (1): 41 (Rusby 1896).

*Casearia sylvestris* var. *wydleri* Briq., *Annuaire Conserv. Jard. Bot. Genève* 2: 74 (Briquet 1898).

*Casearia sylvestris* var. *eichleri* Briq., *Bull. Herb. Boissier*, sér. 2, 7: 672 (Briquet 1907).

*Casearia onacaensis* Rusby, *Descr. S. Amer. Pl.* 62 [20 Dec. 1920] (Rusby 1920).

*Casearia chlorophoroidea* Rusby, *Descr. S. Amer. Pl.* 63 [20 Dec. 1920] (Rusby 1920). — *Casearia sylvestris* var. *chlorophoroidea* (Rusby) Sleumer, *Fl. Suriname* 3 (1): 298 (Sleumer 1935).

*Casearia herbert-smithii* Rusby, *Descr. S. Amer. Pl.* 63 [20 Dec. 1920] (Rusby 1920).

*Casearia formosa* Urb., *Repert. Spec. Nov. Regni Veg.* 22: 91 (Urban 1925).

*Casearia lindeniana* Urb., *Repert. Spec. Nov. Regni Veg.* 22: 92 (Urban 1925).

*Casearia caudata* Uittien, *Recueil Trav. Bot. Néerl.* 22: 373 ["1925" publ. Jan. 1926] (Uittien 1926).

*Casearia sylvestris* var. *angustifolia* Uittien, *Recueil Trav. Bot. Néerl.* 22: 373 ["1925" publ. Jan. 1926] (Uittien 1926).

*Casearia sylvestris* var. *paraensis* Uittien, *Recueil Trav. Bot. Néerl.* 22: 373 ["1925" publ. Jan. 1926] (Uittien 1926).

*Casearia schulziana* O.C.Schmidt, *Repert. Spec. Nov. Regni Veg.* 32: 87 (Schmidt 1933).
*Casearia ekmanii* Sleumer, *Notizbl. Bot. Gart. Berlin-Dahlem* 11: 954 [20 Jan. 1934] (Sleumer 1934).

*Casearia ovoidea* Sleumer, *Notizbl. Bot. Gart. Berlin-Dahlem* 11: 958 [20 Jan. 1934] (Sleumer 1934).

*Casearia sylvestris* var. *martinicensis* J.F.Macbr. ex Ll.Williams, *Publ. Field Mus. Nat. Hist., Bot. Ser.* 15: 357 (Williams 1936), *nom. inval. (anglice)*.

*Casearia subsessiliflora* Lundell, *Contr. Univ. Michigan Herb.* 6: 50 (Lundell 1941).

NOTE. — *Samyda parviflora* Sessé & Moc. also appears in the first edition of *Flora mexicana* (Sessé & Mociño 1892-1898: 123, as “*Paruiflora*”), but edition 2 has priority, probably from p. 49 onward (Stafleu & Cowan, 1985: TL-2-11756)].

VERNACULAR NAMES. — Wp: yape'a pitá, yape'a pitá sili • Nt: kwasikwasi tiki • Br: cafézeiro-do-mato, erva-de-bugre, erva-de-lagarto, guaçatunga.

HERBARIUM DATA (FG). — 16 collections at CAY. Sel. exs.: *D. Sabatier 2100*.

INVENTORY DATA (FG). — 35 trees in 22 plots; $F_{max} < 1\%$; $dbh_{inv} = 22$ cm.

[1540] *Casearia ulmifolia* Vahl ex Vent.
(Fig. 52C)

*Choix Pl.* 1: 46, verso [Apr. 1804] (Ventenat 1804). — *Guidonia ulmifolia* (Vahl ex Vent.) Baill., *Traité Bot. Méd. Phan.* 2: 827 [Mar. 1884] (Baillon 1884).

*Casearia celtidifolia* Kunth, *Nova genera et species plantarum [H.B.K.]* 5: 363 [24 Mar. 1823] (Kunth 1823).

*Casearia petraea* Benth., *J. Bot. [Hooker]* 4: 111 (Bentham 1842).

*Casearia blanchetiana* Miq., *Linnaea* 22: 801 [“1849” publ. May 1850] (Miquel 1850).

*Casearia cambessedesii* Eichler, *Fl. Bras. [Martius]* 13 (1): 475 [1 Oct. 1871] (Eichler 1871).

*Casearia celtidifolia* Poepp. ex Eichler, *Fl. Bras. [Martius]* 13 (1): 477 [1 Oct. 1871] (Eichler 1871), *nom. illeg. hom., non* Kunth (1823).

*Casearia cotticensis* Uittien, *Recueil Trav. Bot. Néerl.* 22: 369 [“1925” publ. Jan. 1926] (Uittien 1926).

*Casearia reginae* J.F.Macbr. ex Ll.Williams, *Publ. Field Mus. Nat. Hist., Bot. Ser.* 15: 356 (Williams 1936), *nom. inval. (anglice)*.

*Casearia tremifolia* J.F.Macbr., *Candollea* 8: 23 (Macbride 1940).

VERNACULAR NAMES. — Wp: yape'a pitá sili • Br: caferana.

HERBARIUM DATA (FG). — 21 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 5006bis*.

INVENTORY DATA (FG). — 16 trees in 13 plots; $F_{max} < 1\%$; $dbh_{inv} = 23.3$ cm.

[1541] *Casearia zizyphoides* Kunth

*Nova genera et species plantarum [H.B.K.]* 5: 362 [24 Mar. 1823] (Kunth 1823).

*Casearia celastroides* Klotzsch, *Reis. Br.-Guiana [Ri. Schomburgk]* 3: 1167 [“1848” publ. 7-10 Mar. 1849] (Klotzsch 1849), *nom. nud.*

*Casearia laevigata* Willd. ex Klotzsch, *Reis. Br.-Guiana [Ri. Schomburgk]* 3: 1167 [“1848” publ. 7-10 Mar. 1849] (Klotzsch 1849), *nom. nud.*

HERBARIUM DATA (FG). — A single collection *fide* Sleumer (1980): *J. Martin s.n.* (K, not seen).

SIZE. — Up to 10 m tall (Sleumer 1980).

[1542] *Casearia* sp. A

NOTE. — This species is also the “*Casearia* sp. A” in Mori & Fischer (2002).

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: *S.A. Mori & B.M. Boom 14721*.

INVENTORY DATA (FG). — 13 trees in 13 plots; $F_{max} < 1\%$; $dbh_{inv} = 16.4$ cm.

[1543] *Casearia* sp. B
(Fig. 52B)

NOTE. — This species is also the “*Casearia* sp. B” in Mori & Fischer (2002).

HERBARIUM DATA (FG). — 14 collections at CAY. Sel. exs.: *S.A. Mori & T.D. Pennington 18124*.

INVENTORY DATA (FG). — 25 trees in 20 plots; $F_{max} < 1\%$; $dbh_{inv} = 28.2$ cm.

[1544] *Casearia* sp. C

NOTES. — This species, represented at CAY by specimens formerly identified as *Casearia acuminata* DC., differs from the latter in: 1) habit (trees vs treelets/shrubs of 5 cm dbh maximum in *C. acuminata*), 2) leaf pilosity (hairs on midvein on abaxial side vs glabrous), 3) leaf colour and size (brownish when dried and relatively large vs yellowish and smaller), 4) fruit pilosity (very short white hairs on the lower half of fruit vs glabrous except for a few long hairs at apex), 5) fruit colour (dull black vs shiny reddish).

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 4064*.

INVENTORY DATA (FG). — 4 trees in 4 plots; $F_{max} < 1\%$; $dbh_{inv} = 22.3$ cm.

[1545] *Casearia* sp. D

HERBARIUM DATA (FG). — A single collection, *M.-F. Prévost & D. Sabatier 2280*.

INVENTORY DATA (FG). — 1 tree, dbh = 10 cm.

Genus *Hasseltia* Kunth

[1546] *Hasseltia floribunda* Kunth

*Nova genera et species plantarum [H.B.K.]* 7: 232 [25 Apr. 1825] (Kunth 1825).
*Hasseltia pubescens* Benth., *Pl. Hartw. [Bentham]*: 164 [before mid-Nov. 1845] (Bentham 1845).

*Banara laxiflora* Benth., *J. Proc. Linn. Soc., Bot.* 5 (Suppl. 2): 91 [May 1861] (Bentham 1861). — *Hasseltia laxiflora* (Benth.) Eichler, *Fl. Bras. [Martius]* 13 (1): 502 [1 Oct. 1871] (Eichler 1871).

*Hasseltia peruviana* Pilg., *Verh. Bot. Vereins Prov. Brandenburg* 47: 160 [1 Oct. 1905] (Pilger 1905).

*Hasseltia tomentulosa* Cuatrec., *Fieldiana, Bot.* 27 (1): 98 (Cuatrecasas 1950).

*Hasseltia monagensis* Steyerm., *Fieldiana, Bot.* 28 (2): 407 (Steyermark 1952).

*Hasseltia panamensis* Standl. ex Dayton, *Phytologia* 4 (4): 253 [June 1953] (Dayton 1953), *nom. nud.*

*Hasseltia micrantha* L.O.Williams, *Fieldiana, Bot.* 29 (6): 361 (Williams 1961).

*Hasseltia psittacarum* L.O.Williams, *Fieldiana, Bot.* 31 (1-2): 28 [25 Nov. 1964] (Williams 1964).

*Hasseltia rigida* Woodson ex A.Robyns, *Ann. Missouri Bot. Gard.* 55 (1): 77 (Robyns 1968).

*HERBARIUM DATA* (FG). — 9 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino* 5683.

*INVENTORY DATA* (FG). — 28 trees in 9 plots; Fmax = 3 %; dbhinv = 47.7 cm.

## Genus *Homalium* Jacq.

[1547] *Homalium guianense* (Aubl.) Oken

*Allg. Naturgesch.* 3 (2): 810 (Oken 1841). — *Racoubea guianensis* Aubl., *Hist. Pl. Guiane* 1: 590 [Jun.-Dec. 1775] (Aublet 1775), "Guyannensis" on plate. — *Homalium spicatum* Lam., *Encycl. [J. Lamarck et al.]* 1 (1): 32 [2 Dec. 1783] (Lamarck 1783), *nom. illeg. superfl.* (based on *Racoubea guianensis*). — *Homalium racoubea* Sw., *Prodr. [Swartz]* 86 [20 Jun.-29 July 1788] (Swartz 1788), *nom. illeg. superfl.* (based on *Racoubea guianensis*).

*Napimoga guianensis* Aubl., *Hist. Pl. Guiane* 1: 592 [Jun.-Dec. 1775] (Aublet 1775), "Guyannensis" on plate. — *Homalium napimoga* Spreng., *Syst. Veg. [Sprengel]* 4 (2, Cur. Post.): 210 [Jan.-June 1827] (Sprengel 1827), *nom. illeg. superfl.* (based on *Napimoga guianensis*).

*Homalium surinamense* Steud., *Flora* 26 (45): 756 [17 Dec. 1843] (Steudel 1843).

*Homalium densiflorum* Spruce ex Benth., *J. Proc. Linn. Soc., Bot.* 4: 36 ["1860" publ. July 1859] (Bentham 1859).

*Homalium puberulum* Klotzsch ex Eichler, *Fl. Bras. [Martius]* 13 (1): 507 [1 Oct. 1871] (Eichler 1871).

*Homalium mattogrossense* Malme, *Ark. Bot.* 23A(4): 20 (Malme 1930).

*Homalium chocoense* Cuatrec., *Fieldiana, Bot.* 27 (1): 99 (Cuatrecasas 1950).

*VERNACULAR NAMES.* — Pa: tahuma • Ka: alekikolan, masalaipyo • Wp: yape'a pitá sili, yawa luway • Cr: bwa-anmé.

*HERBARIUM DATA* (FG). — 46 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material of *Racoubea guianensis*: BM[BM000624532]).

*SIZE.* — Up to 12 cm dbh (Sleumer 1980).

[1548] *Homalium racemosum* Jacq.

* Enum. Syst. Pl.*: 24 [Aug.-Sep. 1760] (Jacquin 1760).

*Homalium senarium* Sessé & Moc. ex DC., *Prodr. [A. P. de Candolle]* 2: 54 [mid Nov. 1825] (Candolle 1825).

*Homalium obtusatum* Turcz., *Bull. Soc. Imp. Naturalistes Moscou* 31 (1): 465 (Turczaninow 1858).

*Homalium pedicellatum* Spruce ex Benth., *J. Proc. Linn. Soc., Bot.* 4: 36 ["1860" publ. July 1859] (Bentham 1859).

*Homalium hondurense* Donn.Sm., *Bot. Gaz.* 20 (1): 4 [18 Jan. 1895] (Donnell Smith 1895).

*Homalium integrifolium* Britton, *Bull. Torrey Bot. Club* 37 (7): 354 (Britton 1910), *nom. illeg. hom., non* (Lam.) Baill. (Baillon 1886).

*Homalium trichostemon* S.F.Blake, *Contr. Gray Herb.* 53: 60 [26 Feb. 1918] (Blake 1918).

*Homalium nicaraguense* S.F.Blake, *Contr. U.S. Natl. Herb.* 20 (7): 225 [9 Sep. 1919] (Blake 1919).

*Homalium mollicellum* S.F.Blake, *Contr. U.S. Natl. Herb.* 20 (7): 226 [9 Sep. 1919] (Blake 1919).

*Homalium pleiandrum* S.F.Blake, *Contr. U.S. Natl. Herb.* 20 (7): 227 [9 Sep. 1919] (Blake 1919).

*Homalium hemistylum* S.F.Blake, *Contr. U.S. Natl. Herb.* 20 (7): 228 [9 Sep. 1919] (Blake 1919).

*Homalium leiogynum* S.F.Blake, *Contr. U.S. Natl. Herb.* 20 (7): 228 [9 Sep. 1919] (Blake 1919).

*Homalium racemosum* subsp. *barbellatum* S.F.Blake, *Contr. U.S. Natl. Herb.* 20 (7): 229 [9 Sep. 1919] (Blake 1919).

*Homalium pittieri* S.F.Blake, *Contr. U.S. Natl. Herb.* 20 (7): 230 [9 Sep. 1919] (Blake 1919).

*Homalium trichocladum* S.F.Blake, *Contr. U.S. Natl. Herb.* 20 (7): 231 [9 Sep. 1919] (Blake 1919).

*Homalium eleutherostylum* S.F.Blake, *Contr. U.S. Natl. Herb.* 20 (7): 232 [9 Sep. 1919] (Blake 1919).

*Homalium columbianum* S.F.Blake, *Contr. U.S. Natl. Herb.* 20 (7): 233 [9 Sep. 1919] (Blake 1919).

*Homalium eurypetalum* S.F.Blake, *Contr. U.S. Natl. Herb.* 20 (7): 234 [9 Sep. 1919] (Blake 1919).

*Homalium stenosepalum* S.F.Blake, *Contr. U.S. Natl. Herb.* 20 (7): 234 [9 Sep. 1919] (Blake 1919).

*Homalium riparium* Standl., *Publ. Carnegie Inst. Wash.* 461: 74 [26 Nov. 1935] (Standley 1935), *nom. illeg. hom., non* Gilg (1908).

*Homalium schippii* Standl., *Trop. Woods* 52: 27 (Standley 1937).

*Homalium anzoateguiense* Steyerm., *Fieldiana, Bot.* 28 (2): 410 (Steyermark 1952).

*Homalium mituense* Cuatrec., *Trop. Woods* 101: 19 (Cuatrecasas 1955).

*HERBARIUM DATA* (FG). — 6 collections at CAY. Sel. exs.: *J. Martin s.n.* (P[P04822073]).

*SIZE.* — Brazil, Maranhão. *D.C. Daly D649* (MO), 15 m × 25 cm.
Genus *Piparea* Aubl.

[1549] *Piparea dentata* Aubl.

Hist. Pl. Guiane 2 (Suppl.): 31 [Jun.-Dec. 1775] (Aublet 1775). — *Casearia dentata* (Aubl.) Eichler, Fl. Bras. [Martius] 13 (1): 484 [1 Oct. 1871] (Eichler 1871), nom. illeg. hom., non Sessé & Moc. ex DC. (Candolle 1825). — Alsodeia piparea Spreng., Syst. Veg. [Sprengel] 1: 807 [1825 publ. late 1824] (Sprengel 1824), nom. illeg. superfl. (based on *Piparea dentata*). — Conohoria aubletii D.Dietr., Syn. Pl. [D. Dietrich] 1: 831 [July 1839] (Dietrich 1839), nom. illeg. superfl. (based on *Piparea dentata*).

Casearia commersoniana Cambess., Fl. Bras. Merid. [A. St.-Hil.] (quarto ed.) 2 (16): 235 ["1829" publ. Oct.-Nov. 1830] (Cambessèdes 1830).

Casearia densiflora Benth., J. Bot. [Hooker] 4: 113 (Bentham 1842).

Casearia laurifolia Benth., J. Bot. [Hooker] 4: 113 (Bentham 1842).

Casearia lasiosperma Triana & Planch., Ann. Sci. Nat., Bot. sér. 4, 17: 113 (Triana & Planchon 1862).

Casearia congestiflora Turcz., Bull. Soc. Imp. Naturalistes Moscou 36 (1): 608 (Turczaninow 1863).

Casearia myriantha Turcz., Bull. Soc. Imp. Naturalistes Moscou 36 (1): 609 (Turczaninow 1863). — *Casearia javitensis* var. *myriantha* (Turcz.) L.O.Williams, *Fieldiana*, Bot. 29 (6): 359 (Williams 1961).

Casearia densiflora var. parviflora Eichler, Fl. Bras. [Martius] 13 (1): 487 [1 Oct. 1871] (Eichler 1871).

Casearia maximilianii Eichler, Fl. Bras. [Martius] 13 (1): 487 [1 Oct. 1871] (Eichler 1871), "maximiliani".

Casearia miradorensis Eichler, Fl. Bras. [Martius] 13 (1): 487 [1 Oct. 1871] (Eichler 1871).

Casearia brighamii Watson, Proc. Amer. Acad. Arts 21: 459 (Watson 1886), "Brighami".

Casearia piparea Sleumer, Lilloa 23: 249 (Sleumer 1950).

NOTE. — The genus *Piparea* Aubl., once sinkted into *Casearia* Jacq., has been resurrected by Samarakoon & Alford (Samarakoon & Alford 2019).

VERNACULAR NAMES. — Pa: tahuma • Wp: kasilikita'i sili, kasilikita'i sili, kwasiilikita'i sili • Br: cafezinho-do-mato, canela-de-veado.

HERBARIUM DATA (FG). — 87 collections at CAY. Sel. exs.: *J.-B. Aublet s.n.* (original material of *Piparea dentata*: BM[BM000624376]).

SIZE. — Up to 20 cm dbh (Sleumer 1980).

[1550] *Piparea multiflora* C.F.Gaertn.

Suppl. Carp. 231 (Gaertner 1807). — *Casearia gaertneri* Donn. Sm., Enum. Pl. Guatem.: 25 (Donnell Smith 1891), nom. illeg. superfl. (based on *Piparea multiflora*, excl. specim. cit. Donn. Sm., which is *Casearia commersoniana* Cambess.)

Casearia javitensis Kunth, Nova genera et species plantarum [H.B.K.] 5: 366 [24 Mar. 1823] (Kunth 1823), in nota, under *Lindleya glabra*. — *Chaetocrater javitensis* (Kunth) Raf., Sylva Tellur.: 149 (Rafinesque 1838).

Casearia iquitosensis J.F.Macbr., Candollea 8: 22 (Macbride 1940).

NOTE. — The genus *Piparea* Aubl., once sinkted into *Casearia* Jacq., has been resurrected by Samarakoon & Alford (Samarakoon & Alford 2019).

VERNACULAR NAMES. — Pa: kanegma, yawu-seine • Ka: alawata muleli • Wp: kasilikita'i, kasilikita'i, kwasiilikita'i • Wn: sieuju • Nt: kwasikwasi tiki • Cr: kafé-sovaj • Br: capança.

HERBARIUM DATA (FG). — 47 collections at CAY. Sel. exs.: *J.-B. Patris s.n.* (original material of *Piparea multiflora*: G-DC, G00208434)).

INVENTORY DATA (FG). — 162 trees in 100 plots; Fmax = 1.7 %; dbhinv = 29.6 cm.

Genus *Ryania* Vahl

[1551] *Ryania pyrifera* (Rich.) Uittien & Sleumer

Fl. Suriname 3 (1): 286 (Uittien & Sleumer 1935). — *Patrisia pyrifera* Rich., *Actes Soc. Hist. Nat. Paris* 1: 110 [Oct. 1792] (Richard 1792).

Ryania sagotiana Eichler, Fl. Bras. [Martius] 13 (1): 491 [1 Oct. 1871] (Eichler 1871). — *Patrisia sagotiana* (Eichler) Kuntze, *Revis. Gen. Pl.* 1: 45 [5 Nov. 1891] (Kuntze 1891).

NOTE. — Excluded synonym: *Ryania speciosa* Vahl.

VERNACULAR NAMES. — Pa: tahuma seine.

HERBARIUM DATA (FG). — 27 collections at CAY. Sel. exs.: *J.B. Leblond s.n.* (original material P[P02442081]).

SIZE. — Up to 10 cm dbh (Sleumer 1980).

[1552] *Ryania speciosa* Vahl var. *bicolor* (DC.) Monach. (Fig. 52D)

Lloydia 12: 19 (Monachino 1949). — *Patrisia bicolor* DC., *Prodr. A. P. de Candolle* 1: 256 [mid Jan. 1824] (Candolle 1824). — *Ryania candollei* Miq., *Ann. Mag. Nat. Hist.* 11: 16 (Miquel 1843), nom. illeg. superfl. (based on *Patrisia bicolor*). — *Ryania bicolor* (DC.) Eichler, *Fl. Bras. [Martius]* 13 (1): 491 [1 Oct. 1871] (Eichler 1871).

VERNACULAR NAMES. — Pa: kuuku-ariut-seine • Ka: tomoipo • Wp: iwikwisī, mukwisī.

HERBARIUM DATA (FG). — 19 collections at CAY. Sel. exs.: *J.-B. Patris s.n.* (original material G-DC, G00207421)).

INVENTORY DATA (FG). — 1 tree, dbh = 10.2 cm.

[1553] *Ryania speciosa* var. *subuliflora* (Sandwith) Monach.

Lloydia 12: 14 (Monachino 1949). — *Ryania pyrifera* var. *subuliflora* Sandwith, *J. Arnold Arbor*. 24 (2): 219 [15 Apr. 1943] (Sandwith 1943).

VERNACULAR NAMES. — Pa: kuuku-ariut • Wp: kasilikita'i, kasilikita'i, kwasiilikita'i • Br: capança, mata-calado.

HERBARIUM DATA (FG). — 28 collections at CAY. Sel. exs.: *R.A.A. Oldeman 2386*.

SIZE. — Brazil, Amazonas. *L.O. Adão Teixeira 943* (MO), 8 m × 10 cm.
[1554] *Ryania speciosa* var. *tomentosa* (Miq.) Monach.

*Lloydia* 12: 16 (Monachino 1949). — *Ryania tomentosa* Miq., *Ann. Mag. Nat. Hist.* 11: 15 (Miquel 1843). — *Patrisia tomentosa* (Miq.) M.Roem., *Fam. Nat. Syn. Monogr.* 2: 136 (Roemer 1846). — *Ryania pyrifera* var. *tomentosa* (Miq.) Sleumer ex Sleumer & Uittien, *Fl. Suriname* 3 (1): 286 (Sleumer & Uittien 1935).

*Patrisia parviflora* DC., *Prodr. [A. P. de Candolle]* 1: 256 [mid Jan. 1824] (Candolle 1824). — *Ryania patrisii* Miq., *Ann. Mag. Nat. Hist.* 11: 16 (Miquel 1843), *nom. illeg. superfl.* (based on *Patrisia parviflora*). — *Ryania parviflora* (DC.) Griseb., *Fl. Brit. W.I.* [Grisebach]: 296 [late 1860] (Grisebach 1860).

*Ryania casiquiarensis* Steyerm., *Fieldiana*, Bot. 28 (2): 411 (Steyermark 1952).

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *J.-B. Patris s.n.* (original material of *Patrisia parviflora*: P[P02442059]).

SIZE. — Colombia, Amazonas. *A. Rudas L.* 3391 (MO), 10 m × 15.9 cm.

## Genus *Xylosma* G. Forst.

[1555] *Xylosma benthamii* (Tul.) Triana & Planch.

*Ann. Sci. Nat.*, Bot. sér. 4, 17: 99 (Triana & Planchon 1862). — *Flacourtia benthamii* Tul., *Ann. Sci. Nat.*, Bot. sér. 3, 7: 291 (Tulasne 1847), “*benthami*”. — *Hisingera benthamii* (Tul.) Clos, *Ann. Sci. Nat.*, Bot. sér. 4, 8: 225 (Clos 1857). — *Myroxylon benthamii* (Tul.) Kuntze, *Revis. Gen. Pl.* 1: 44 [5 Nov. 1891] (Kuntze 1891).

*Flacourtia nitida* Benth., *Hooker's J. Bot. Kew Gard. Misc.* 3: 119 (Bentham 1851). — *Hisingera lucens* Clos, *Ann. Sci. Nat.*, Bot. sér. 4, 8: 225 (Clos 1857), *nom. illeg. superfl.* (based on *Flacourtia nitida*). — *Myroxylon lucens* Kuntze, *Revis. Gen. Pl.* 1: 44 [5 Nov. 1891] (Kuntze 1891).

*Xylosma digyna* Eichler, *Fl. Bras. [Martius]* 13 (1): 447 [1 Oct. 1871] (Eichler 1871), “*digynum*”. — *Flacourtia digyna* Benth. ex Eichler, *Fl. Bras. [Martius]* 13 (1): 447 [1 Oct. 1871] (Eichler 1871), *nom. nud. pro syn.*

*Myroxylon digynum* Kuntze, *Revis. Gen. Pl.* 1: 44 [5 Nov. 1891] (Kuntze 1891).

*Xylosma armata* J.F.Macbr., *Candollea* 5: 391 (Macbride 1934).

*Xylosma pilosa* J.F.Macbr., *Candollea* 5: 391 (Macbride 1934).

*Xylosma pallidifolia* Sleumer ex Steyerm. & O.Huber, *Fl. Avila*: 425 (Steyermark & Huber 1978), “*pallidifolium*”, *nom. nud. pro syn.*

NOTES. — Being based on *Flacourtia nitida*, *Hisingera lucens* is superfluous and illegitimate. Therefore it cannot serve as the basionym of *Myroxylon lucens*.

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *L.C. Richard s.n.* (P[P04734493]).

INVENTORY DATA (FG). — 3 trees in 3 plots; $F_{max} < 1\%$; $dbh_{inv} = 13.5$ cm.

[1556] *Xylosma ciliatifolia* (Clos) Eichler

*Fl. Bras. [Martius]* 13 (1): 449 [1 Oct. 1871] (Eichler 1871), “*ciliatifolium*”. — *Hisingera ciliatifolia* Clos, *Ann. Sci. Nat.*, Bot. sér. 4, 8: 223 (Clos 1857). — *Myroxylon ciliatifolium* (Clos) Kuntze, *Revis. Gen. Pl.* 1: 44 [5 Nov. 1891] (Kuntze 1891).

*Xylosma pou-orfilae* Herter, *Revista Sudamer. Bot.* 7: 214 (Herter 1943), *nom. nud.*

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *P. Gremand & M.-F. Prévost 1959*.

INVENTORY DATA (FG). — 1 tree, dbh = 10.2 cm.

[1557] *Xylosma tessmannii* Sleumer

*Notizbl. Bot. Gart. Berlin-Dahlem* 12: 477 [30 June 1935] (Sleumer 1935).

NOTE. — According to Sleumer (1980), this species is close to *X. benthamii*, perhaps only a luxuriant form of it.

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *M.-F. Prévost 3302*.

SIZE. — Up to 40 cm dbh (Sleumer 1980).

## Family SAPINDACEAE Juss.

### Genus *Allophylus* L.

[1558] *Allophylus edulis* (A.St.-Hil., A.Juss. & Cambess.)

### Hieron. ex Niederl.

### (Fig. 53A)

*Bol. Mens. Mus. Prod. Argent.* 3 (29): 180 [Oct. 1890] (Niederlein 1890). — *Schmidelia edulis* A.St.-Hil., A.Juss. & Cambess., *Pl. Usuel. Bras.* 14: t. 67 [4 Aug. 1828] (Saint-Hilaire *et al.* 1828).

*Allophylus edulis* var. *gracilis* Radlk., *Fl. Bras. [Martius]* 13 (3): 485 [1 Apr. 1900] (Radlkofer 1900).

*Allophylus edulis* var. *subsessilis* Huber, *Bull. Soc. Bot. Genève*, sér. 2, 6: 185 [“1914” publ. 1915] (Huber 1915).

NOTES. — Acevedo-Rodríguez stated (pers. comm. 2015) that he mistakenly treated this taxon as *A. angustatus* (Triana & Planch.) Radlk. in (Acevedo-Rodríguez 2012). Á. Jussieu and Cambessèdes are co-authors of parts 9-14 (pl. 41-70) of *Plantes usuelles des Brasiliens* (Saint-Hilaire *et al.* 1828).

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier 2356*.

INVENTORY DATA (FG). — 19 trees in 12 plots; $F_{max} < 1\%$; $dbh_{inv} = 22.3$ cm.

[1559] *Allophylus latifolius* Huber

*Bull. Soc. Bot. Genève*, sér. 2, 6: 186 [“1914” publ. 1915] (Huber 1915).

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *S.A. Mori & B.M. Boom 14752*.

INVENTORY DATA (FG). — 4 trees in 4 plots; $F_{max} < 1\%$; $dbh_{inv} = 13.5$ cm.
<figure><img src="image_75.png" /><figcaption>A</figcaption></figure>

<figure><img src="image_76.png" /><figcaption>B</figcaption></figure>

<figure><img src="image_77.png" /><figcaption>C</figcaption></figure>

<figure><img src="image_78.png" /><figcaption>D</figcaption></figure>

FIG. 52. — Salicaceae: **A**, *Casearia grandiflora* Cambess. (*D. Sabatier & J.-F. Molino* 5187); **B**, *Casearia* sp. B (J.-F. Molino & D. Sabatier 2469); **C**, *Casearia ulmifolia* Vahl ex Vent. (*D. Sabatier & J.-F. Molino* 5006bis); **D**, *Ryania speciosa* Vahl var. bicolor (DC.) Monach. (*J.-F. Molino et al.* 1852). A, D, © D. Sabatier/IRD; B, C, © J.-F. Molino/IRD.

[1560] *Allophylus leucoclados* Radlk.

Fl. Bras. [Martius] 13 (3): 473 [1 Apr. 1900] (Radlkofer 1900).

VERNACULAR NAMES. — Wp: tapi'i lo wapa'a.

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: *C. Sastre* 1621, height 15 m.

[1561] *Allophylus robustus* Radlk.

Monogr. Paullinia: 185 (Radlkofer 1895).

HERBARIUM DATA (FG). — 27 collections at CAY. Sel. exs.: *P.A. Poiteau s.n.* (original material G-DC, G00008481).

INVENTORY DATA (FG). — 4 trees in 4 plots; Fmax < 1%; dbhinv = 17 cm.

Genus *Cupania* L.

[1562] *Cupania dipphylla* Vahl

Eclog. Amer. 3: 9 (Vahl 1807).

*Cupania geminata* Poir., *Encycl. [J. Lamarck et al.]* Suppl. 2: 419 [3 July 1812] (Poiret 1812).

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *Unknown coll. s.n.* (“Herb. Poir.”) (probable original material of *Cupania geminata*: P[P06671081]).

INVENTORY DATA (FG). — 10 trees in 9 plots; Fmax < 1%; dbhinv = 14.6 cm.

[1563] *Cupania hirsuta* Radlk.

Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 9: 565 (Radlkofer 1879).
*Cupania velutina* Klotzsch, *Reis. Br.-Guiana* [*Ri. Schomburgk*] 3: 1180 ["1848" publ. 7-10 Mar. 1849] (Klotzsch 1849), *nom. nud.*

*Cupania lanuginosa* Sagot ex Radlk., *Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 9: 566 (Radlkofer 1879).

**HERBARIUM DATA (FG).** — 9 collections at CAY. Sel. exs.: *P.A. Sagot 1214*, Jan. 1859 (holotype of *Cupania lanuginosa*: P[P06671233]; iso-, P[P06671232, P06671234, P06671235]; possible iso-, F[V0071577F, V0071578F], NY[00337808]).

**INVENTORY DATA (FG).** — 27 trees in 18 plots; $F_{max} = 1\%$; $dbh_{inv} = 19$ cm.

[1564] *Cupania macrostylis* (Radlk.) Acev.-Rodr.

*Fl. Guianas*, ser. A, 29: 25 (Acevedo-Rodríguez 2012). — *Matayba macrostylis* Radlk., *Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 9: 625 (Radlkofer 1879).

*Matayba macrolepis* Radlk., *Fl. Bras. [Martius]* 13 (3): 605 [1 Apr. 1900] (Radlkofer 1900).

*Trichilia caudata* Killip & Cuatrec., *Revista Acad. Colomb. Ci. Exact.* 5: 36 [Jan.-June 1942] (Killip & Cuatrecasas 1942).

**HERBARIUM DATA (FG).** — 3 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier* 2401.

**INVENTORY DATA (FG).** — 4 trees in 3 plots; $F_{max} < 1\%$; $dbh_{inv} = 15.5$ cm.

[1565] *Cupania rubiginosa* (Poir.) Radlk.

*Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 9: 566 (Radlkofer 1879). — *Robinia rubiginosa* Poir., *Encycl. [J. Lamarck et al.]* 6 (1): 227 [2 Oct. 1804] (Poiret 1804).

*Cupania subrepanda* Mart., *Flora* 21 (2, Beibl.): 70 (Martius 1838).

**VERNACULAR NAMES.** — Pa: mbagwi, mbagwi-purubumna, mbarui, mbarui-purubumna • Ka: tohmopala • Wp: kalima'i, kalima'i sili, kalima'i yowa • Nt: singabusu • Cr: gangouti • Br: pau-de-arapuce.

**HERBARIUM DATA (FG).** — 70 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier* 2792.

**INVENTORY DATA (FG).** — 24 trees in 19 plots; $F_{max} < 1\%$; $dbh_{inv} = 19.9$ cm.

[1566] *Cupania scrobiculata* Rich.
(Fig. 53B)

*Actes Soc. Hist. Nat. Paris* 1: 109 [Oct. 1792] (Richard 1792).

*Cupania reticulata* Cambess., *Mém. Mus. Hist. Nat.* 18: 41 (Cambessèdes 1829). — *Cupania scrobiculata* f. reticulata (Cambess.) Radlk., *Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 9: 565 (Radlkofer 1879). — *Cupania scrobiculata* var. reticulata (Miq.) Uittien, *Fl. Suriname* 2 (1): 389 (Uittien 1937).

*Cupania frondosa* Benth., *Hooker's J. Bot. Kew Gard. Misc.* 3: 199 (Bentham 1851). — *Cupania scrobiculata* f. frondosa (Benth.) Radlk., *Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss.*

München 9: 565 (Radlkofer 1879). — *Cupania scrobiculata* var. frondosa (Benth.) Uittien, *Fl. Suriname* 2 (1): 390 (Uittien 1937).

*Cupania porosa* Miq., *Stirp. Surinam. Select.*: 70 ["1850" publ. Mar. 1851] (Miquel 1851).

*Cupania guianensis* Miq., *Stirp. Surinam. Select.*: 71 ["1850" publ. Mar. 1851] (Miquel 1851). — *Cupania scrobiculata* f. guianensis (Miq.) Radlk., *Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 9: 565 (Radlkofer 1879).

*Cupania obovata* Gleason, *Bull. Torrey Bot. Club* 54 (8): 616 [Nov. 1927] (Gleason 1927).

*Cupania auriculata* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 8 (1): 20 (Standley 1930).

**NOTE.** — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

**VERNACULAR NAMES.** — Pa: mbagwi, mbagwi-seine, mbarui, mbarui-seine • Ka: tohmopala, tonolo ipyo, tonolo polipyo • Te: tononipö • Wp: kalima'i, kalimai'si • Wn: kabuman, kaikui wapa • Nt: singabusu • Cr: gangouti-blan • Br: pau-de-arapuce.

**HERBARIUM DATA (FG).** — 113 collections at CAY. Sel. exs.: *L. C. Richard s.n.* (holo-, P[P00117101; iso-, P00117102, P00117103, P00117104, P00117105, P00117106]).

**INVENTORY DATA (FG).** — 331 trees in 120 plots; $F_{max} = 9.7\%$; $dbh_{inv} = 38$ cm.

## Genus *Matayba* Aubl.

[1567] *Matayba arborescens* (Aubl.) Radlk.

*Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 9: 353 (Radlkofer 1879). — *Sapindus arborescens* Aubl., *Hist. Pl. Guiane* 1: 357 [Jun.-Dec. 1775] (Aublet 1775).

*Cupania laevigata* Rich., *Actes Soc. Hist. Nat. Paris* 1: 109 [Oct. 1792] (Richard 1792), "Laevigata".

*Cupania aubletii* Miq., *Stirp. Surinam. Select.*: 67 ["1850" publ. Mar. 1851] (Miquel 1851).

*Matayba reducta* Steyerm., *Fieldiana*, *Bot.* 28 (2): 351 (Steyermark 1952).

**NOTE.** — The original material of *Sapindus arborescens* at BM (BM000884118)) is labelled "TYPE SPECIMEN *Sapindus frutescens* Aubl. Pl. Gui. 1: 355. T 138", while that of *S. frutescens* Aubl. [synonym of *Pseudima frutescens* (Aubl.) Radlk.] (BM000884091)) is labelled "TYPE SPECIMEN *Sapindus arborescens* Aubl. Pl. Gui. 1: 357. T 137" (Acevedo-Rodríguez 2012: 38).

**VERNACULAR NAMES.** — Ka: tonolo ipyo, tonolo polipyo • Te: tononipö • Wp: kalima'i, kalimai'si, yatoa'i sī • Nt: singabusu • Cr: sanvonié • Br: breu-de-tucano.

**HERBARIUM DATA (FG).** — 44 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000884118]).

**INVENTORY DATA (FG).** — 10 trees in 10 plots; $F_{max} < 1\%$; $dbh_{inv} = 27.1$ cm.
[1568] *Matayba guianensis* Aubl.

Hist. Pl. Guiane 1: 331 [Jun.-Dec. 1775] (Aublet 1775). — *Matayba guianensis* f. genuina Radlk., Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 9: 631 (Radlkofer 1879), nom. inval. (Turland et al. 2018: Art. 24.3).

VERNACULAR NAMES. — Wn: asisimë, sihkëimëimë.

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000884102]).

INVENTORY DATA (FG). — 9 trees in 7 plots; Fmax < 1%; dbhinv = 30.1 cm.

[1569] *Matayba inelegans* Radlk.

Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 9: 535 (Radlkofer 1879). — *Cupania inelegans* Spruce ex Radlk., Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 9: 517 (Radlkofer 1879), nom. nud. pro syn.

Matayba sororopaniana Steyerm., Fieldiana, Bot. 28 (2): 348 (Steyermark 1952).

NOTE. — Although Spruce's label on type reads "*Cupania inelegans* Spruce", Spruce never made the combination *Matayba inelegans*. Authorship is thus Radlk., not "Spruce ex Radlk."

VERNACULAR NAMES. — Pa: kwepan-wašiunu • Ka: tonolo ipyo, tonolo polipyo • Nt: geli udu.

HERBARIUM DATA (FG). — 25 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 3348*.

INVENTORY DATA (FG). — 126 trees in 27 plots; Fmax = 10.9%; dbhinv = 33.9 cm.

[1570] *Matayba laevigata* Radlk.

Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 9: 628 (Radlkofer 1879). — *Cupania laevigata* Miq., Stirp. Surinam. Select.: 72 ["1850" publ. Mar. 1851] (Miquel 1851), nom. illeg. hom., non Rich. (Richard 1792) [synonym of *Matayba guianensis*].

Matayba guianensis subf. subovalis Radlk., Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 9: 632 (Radlkofer 1879).

Matayba miquelii Uittien, Fl. Suriname 2 (1): 384 (Uittien 1937).

NOTES. — *Cupania laevigata* Miq. is illegitimate, thus cannot be the basionym of *Matayba laevigata* Radlk. Although published as a subforma of the invalid forma genuina, *M. guianensis* subf. subovalis Radlk. is legitimate.

HERBARIUM DATA (FG). — 18 collections at CAY. Sel. exs.: *FM.R. Leprieur 333* (original material of *Matayba guianensis* subf. subovalis: L[L0014432, L0014433]).

INVENTORY DATA (FG). — 9 trees in 9 plots; Fmax < 1%; dbhinv = 32.7 cm.

[1571] *Matayba opaca* Radlk.
(Fig. 53C)

Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 9: 628 (Radlkofer 1879).

Matayba fallax Radlk., Fl. Bras. [Martius] 13 (3): 612 [1 Apr. 1900] (Radlkofer 1900). — *Matayba opaca* var. fallax (Radlk.) Uittien, *Fl. Suriname* 2 (1): 384 (Uittien 1937).

Matayba affinis Steyerm., Ann. Missouri Bot. Gard. 75 (3): 1067 [19 Oct. 1988] (Steyermark 1988).

VERNACULAR NAMES. — Ka: tikalayen tonoloipyo • Wp: wila pisi'u • Wn: jala.

HERBARIUM DATA (FG). — 17 collections at CAY. Sel. exs.: *D. Sabatier 2102*.

INVENTORY DATA (FG). — 50 trees in 8 plots; Fmax = 3.4%; dbhinv = 28.3 cm.

[1572] *Matayba peruviana* Radlk. subsp. *oligandra* (Sandwith) T.D.Penn. ex Acev.-Rodr.

Fl. Guianas, ser. A, 29: 46 (Acevedo-Rodríguez 2012). — *Matayba oligandra* Sandwith, *Bull. Misc. Inform. Kew* 1935 (3): 123 [20 May 1935] (Sandwith 1935).

Trichilia ptariana Steyerm., Fieldiana, Bot. 28 (2): 278 (Steyermark 1952).

Matayba oligandra var. ptariana (Steyerm.) Steyerm., Bol. Soc. Venez. Ci. Nat. 26: 426 (Steyermark 1966).

Matayba chimantensis Steyerm. & Maguire, Mem. New York Bot. Gard. 17 (1): 447 (Steyermark & Maguire 1967).

Matayba jauaensis Steyerm., Bol. Soc. Venez. Ci. Nat. 33: 347 (Steyermark 1976).

Matayba oligandra var. occidentalis Steyerm., Ann. Missouri Bot. Gard. 75 (3): 1069 [19 Oct. 1988] (Steyermark 1988).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *S.A. Mori & T.D. Pennington 17937*.

SIZE. — Up to 20 m tall (Acevedo-Rodríguez 2012).

[1573] *Matayba stenodictya* Radlk.

Fl. Bras. [Martius] 13 (3): 629 [1 Apr. 1900] (Radlkofer 1900).

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *D. Sabatier et al. 4712*.

INVENTORY DATA (FG). — 12 trees in 5 plots; Fmax < 1%; dbhinv = 44.6 cm.

Genus *Melicoccus* P.Browne

[1574] *Melicoccus pedicellaris* (Sagot ex Radlk.) Acev.-Rodr.

Fl. Neotrop. Monogr. 87: 30 [22 May 2003] (Acevedo-Rodríguez 2003). — *Talisia pedicellaris* Sagot ex Radlk., Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 8: 342 (Radlkofer 1878).

Talisia pulverulenta Radlk., Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 8: 342 (Radlkofer 1878).
NOTE. — Radlkofer (1878) did ascribe the binomial *Talisia pedicellaris* to Sagot, by stating: "Der speciesbeiname ist aus dem von Sagot der Pflanze beigelegten Namen [...]$^{)}$".

VERNACULAR NAMES. — Wp: tuli âtá u.

HERBARIUM DATA (FG). — 33 collections at CAY. Sel. exs.: *P.A. Sagot* 1188, Aug. 1858 (lecto-, P, designated by Acevedo-Rodríguez [2003: 31], not seen, fragm. M[M-0224883]; isolecto-, BM[BM000884044], BR[BR0000005417018], F[V0071843F, V0071844F, V0071845F], GOET[GOET010791], K[K000035428, K000035429], MPU[MPU010928], P[P00117093, P00117094, P00117095, P00117096, P00117097, P00117098], U[U0006556]; possible isolecto-, P[P02428274, P02428275]).

INVENTORY DATA (FG). — 80 trees in 52 plots; F<sub>max</sub> = 1.2%; dbh<sub>inv</sub> = 27.7 cm.

Genus *Pentascyphus* Radlk.

[1575] *Pentascyphus thyrsisflorus* Radlk.

Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München
9: 539 (Radlkofer 1879).

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: *FM.R. Leprieur* 335 (holo-, P[P00214699]; iso-, L[L0014517]).

INVENTORY DATA (FG). — 3 trees in 3 plots; F<sub>max</sub> < 1%; dbh<sub>inv</sub> = 24.8 cm.

Genus *Pseudima* Radlk.

[1576] *Pseudima frutescens* (Aubl.) Radlk.

Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München
8: 300 (Radlkofer 1878). — *Sapindus frutescens* Aubl., *Hist. Pl. Guiane* 1: 355 [Jun.-Dec. 1775] (Aublet 1775).

NOTE. — The original material of *Sapindus frutescens* at BM (BM000884091) is labelled "TYPE SPECIMEN *Sapindus arborescens* Aubl. Pl. Gui. 1: 357. T 137", while that of *S. arborescens* Aubl. [synonym of *Matayba arborescens* (Aubl.) Radlk.] (BM000884118)] is labelled "TYPE SPECIMEN *Sapindus frutescens* Aubl. Pl. Gui. 1: 355. T 138" (Acevedo-Rodríguez 2012: 38).

VERNACULAR NAMES. — Pa: tuu-ariut, tuu-kamwi-seine • Ka: tiyawasisyen • Wp: yikisǐǐ, yikisǐi lo • Cr: sanvonié • Br: camaá, fruta-de-anel, pau-de-gafanhoto, pitomba-de-macaco.

HERBARIUM DATA (FG). — 50 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000884091]).

INVENTORY DATA (FG). — 2 trees in 2 plots; F<sub>max</sub> < 1%; dbh<sub>inv</sub> = 44.1 cm.

Genus *Scyphonychium* Radlk.

[1577] *Scyphonychium multiflorum* (Mart.) Radlk.

Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München
9: 519 (Radlkofer 1879). — *Cupania multiflora* Mart., *Flora* 21 (2,
Beibl.): 75 (Martius 1838).

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier* 3479.

INVENTORY DATA (FG). — 5 trees in 4 plots; F<sub>max</sub> < 1%; dbh<sub>inv</sub> = 19.3 cm.

Genus *Talisia* Aubl.

[1578] *Talisia clathrata* Radlk. subsp. *canescens* Acev.-Rodr.

Fl. Neotrop. Monogr. 87: 56 [22 May 2003] (Acevedo-Rodríguez 2003).

VERNACULAR NAMES. — Nt: baala wiwii.

HERBARIUM DATA (FG). — 18 collections at CAY. Sel. exs.: *P. Acevedo-Rodríguez et al. 5012* (holo-, US[00614239]; iso-, A, CAY[CAY001980, CAY001981], CTES, F, HUA, K, NY, P[P04857357], L[exU]).

INVENTORY DATA (FG). — 16 trees in 11 plots; F<sub>max</sub> < 1%; dbh<sub>inv</sub> = 25.6 cm.

[1579] *Talisia furfuracea* Sandwith

Bull. Misc. Inform. Kew 1935 (3): 121 [20 May 1935] (Sandwith 1935).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 3663.

INVENTORY DATA (FG). — 46 trees in 39 plots; F<sub>max</sub> < 1%; dbh<sub>inv</sub> = 47.1 cm.

[1580] *Talisia guianensis* Aubl.

Hist. Pl. Guiane 1: 349 [Jun.-Dec. 1775] (Aublet 1775), "Guy-
annensis" on plate. — *Talisia rosea* Vahl, *Eclog. Amer.* 2: 30 (Vahl
1798), *nom. illeg. superfl.* (based on *Talisia guianensis*).

*Talisia glabra* DC., *Prodr.* [A. P. de Candolle] 1: 609 [mid Jan. 1824] (Candolle 1824).

HERBARIUM DATA (FG). — 31 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000838014]); *P. Acevedo-Rodríguez et al. 4871* (MO), 10 m tall.

[1581] *Talisia hemidasya* Radlk.

Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München
8: 349 (Radlkofer 1878).

*Talisia glandulifera* Steyerm., *Ann. Missouri Bot. Gard.* 75 (3): 1072
[19 Oct. 1988] (Steyermark 1988).

VERNACULAR NAMES. — Ka: sepupi, supupi, tonolo ipyo • Wp: tuli átá • Cr: bwa-flanbo, payakousa • Br: pitomba-do-norte, pitombeira.

HERBARIUM DATA (FG). — 24 collections at CAY. Sel. exs.: S.A. Mori et al. 15027 (holotype of *Talisia glandulifera*: MO[MO-256425]; iso-, NY[00039423]).

INVENTORY DATA (FG). — 9 trees in 4 plots; F<sub>max</sub> = 1%; dbh<sub>inv</sub> = 26.5 cm.
[1582] *Talisia hexaphylla* Vahl

Eclog. Amer. 2: 29 (Vahl 1798).

*Talisia panamensis* Pittier, *Contr. U.S. Natl. Herb.* 20 (3): 129 [18 June 1918] (Pittier 1918).

*Talisia cararensis* Cuatrec., *Revista Acad. Colomb. Ci. Exact.* 8 (31): 306 [Dec. 1951] (Cuatrecasas 1951).

**VERNACULAR NAMES.** — Pa: miumiu-asiru-wašiunu, tuu-ariut • Ka: wa:u • Te: tuli • Wn: kělěpukaju • Nt: gaan tatu tiki, tatu, tatu udu • Cr: bwa-flanbo, payakousa • Br: pitomba-do-norte, pitombeira.

**HERBARIUM DATA (FG).** — 39 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 4631*.

**INVENTORY DATA (FG).** — 198 trees in 107 plots; $F_{max} = 1\%$; $dbh_{inv} = 46.5$ cm.

[1583] *Talisia longifolia* (Benth.) Radlk.

*Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 8: 348 (Radlkofer 1878). — *Cupania longifolia* Benth., *Hooker's J. Bot. Kew Gard. Misc.* 2: 211 (Bentham 1850).

**VERNACULAR NAMES.** — Wp: tuli âtā • Wn: tëpu • Nt: tatu • Cr: bwa-flanbo, payakousa • Br: pitomba-do-norte, pitombeira.

**HERBARIUM DATA (FG).** — 15 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 2872*.

**INVENTORY DATA (FG).** — 7 trees in 6 plots; $F_{max} < 1\%$; $dbh_{inv} = 14.5$ cm.

[1584] *Talisia macrophylla* (Mart.) Radlk.

*Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 8: 347 (Radlkofer 1878). — *Cupania macrophylla* Mart., *Flora* 21 (2, Beibl.): 74 (Martius 1838).

*Talisia elephantipes* Sandwith, *J. Bot.* 78: 256 (Sandwith 1940).

*Talisia allenii* Croat, *Ann. Missouri Bot. Gard.* 63: 527 (Croat 1976).

*Talisia pentantha* Steyerm., *Ann. Missouri Bot. Gard.* 75 (3): 1075 [19 Oct. 1988] (Steyermark 1988).

*Talisia sancarlosiana* Steyerm., *Ann. Missouri Bot. Gard.* 75 (3): 1075 [19 Oct. 1988] (Steyermark 1988).

**VERNACULAR NAMES.** — Te: tuli • Wp: tuli âtā u • Cr: bwa-flanbo, payakousa.

**HERBARIUM DATA (FG).** — 14 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 3253*.

**INVENTORY DATA (FG).** — 8 trees in 5 plots; $F_{max} < 1\%$; $dbh_{inv} = 15$ cm.

[1585] *Talisia megaphylla* Sagot ex Radlk.

*Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 8: 350 (Radlkofer 1878).

**VERNACULAR NAMES.** — Pa: tuu-aška, tuu-kamwi • Ka: malay wokili, tulisi, tulusili.

**HERBARIUM DATA (FG).** — 33 collections at CAY. Sel. exs.: *P.A. Sagot* 1194, 1857 (lecto-, P[P06694733], designated by Acevedo-Rodríguez [2003: 136]; isolecto-, BR[BR0000005417100], K[K000035531, K000035532]).

**INVENTORY DATA (FG).** — 8 trees in 8 plots; $F_{max} < 1\%$; $dbh_{inv} = 15.5$ cm.

[1586] *Talisia microphylla* Uittien

*Recueil Trav. Bot. Néerl.* 34: 483 (Uittien 1937).

**HERBARIUM DATA (FG).** — 25 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 3855*.

**INVENTORY DATA (FG).** — 46 trees in 34 plots; $F_{max} < 1\%$; $dbh_{inv} = 41.1$ cm.

[1587] *Talisia mollis* Kunth ex Cambess.

*Mém. Mus. Hist. Nat.* 18: 48 (Cambessèdes 1829).

*Tapirocarpus talisia* Sagot, *Ann. Sci. Nat., Bot.* sér. 6, 13: 292 (Sagot 1882), *pro parte vegetativa*.

**VERNACULAR NAMES.** — Wp: tuli âtā, tuli âtā u, tuli âtā yowa • Nt: tatu • Cr: bwa-flanbo, payakousa • Br: pitomba-brava.

**HERBARIUM DATA (FG).** — 52 collections at CAY. Sel. exs.: *R.A.A. Oldeman B-4183*.

**INVENTORY DATA (FG).** — 3 trees in 3 plots; $F_{max} < 1\%$; $dbh_{inv} = 12.4$ cm.

[1588] *Talisia pachycarpa* Radlk.

*Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 8: 350 (Radlkofer 1878).

**HERBARIUM DATA (FG).** — 2 collections at CAY. Sel. exs.: *P.A. Poiteau s.n.* (holo-, G[G00008241, photo F neg. 23635]; iso-, F[V0071842F], possible iso-, K[K000542729]).

**SIZE.** — Up to 15 cm dbh (Acevedo-Rodríguez 2003).

[1589] *Talisia praealta* Sagot ex Radlk.

*Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 8: 345 (Radlkofer 1878). — *Cupania praealta* (Sagot ex Radlk.) Sagot, *Ann. Sci. Nat., Bot.* sér. 6, 12: 199 (Sagot 1881).

**VERNACULAR NAMES.** — Pa: bukutru-ateu • Ka: tulisi, tulusili • Te: tuli • Wn: jala, tëpuimë • Cr: bwa-flanbo, bwa-golèt, payakousa • Br: pitomba-do-norte, pitombeira.

**HERBARIUM DATA (FG).** — 40 collections at CAY. Sel. exs.: *P.A. Sagot* 1047, 1857 (holo-, B[not seen, photo F neg 5679]; lecto-, P[P04857336], designated by Acevedo-Rodríguez [2003: 76]; isolecto-, BM[BM001042336], BR[BR0000005416639], F[V0071846F, V0071847F], GOET[GOET010870], K[K000035414, K000035415], M[M0224877], MPU[MPU010929], NY[00658344], P[P04857336], U[U0006555]).

**INVENTORY DATA (FG).** — 114 trees in 74 plots; $F_{max} < 1\%$; $dbh_{inv} = 79.5$ cm.
[1590] *Talisia simaboides* K.U.Kramer

*Acta Bot. Neerl.* 21: 676 (Kramer 1972).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Nt: gaan tatu tiki, tatu udu • Cr: bwa-flanbo, payakousa.

HERBARIUM DATA (FG). — 35 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 3658*.

INVENTORY DATA (FG). — 132 trees in 93 plots; Fmax < 1%; dbhinv = 70 cm.

[1591] *Talisia squarrosa* Radlk.

*Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 8: 346 (Radlkofer 1878).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — 13 collections at CAY. Sel. exs.: *M.-F. Prévost 2170*.

INVENTORY DATA (FG). — 36 trees in 4 plots; Fmax = 2.4%; dbhinv = 54.7 cm.

[1592] *Talisia sylvatica* (Aubl.) Radlk.

*Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 8: 341 (Radlkofer 1878). — *Racaria sylvatica* Aubl., *Hist. Pl. Guiane* 2 (Suppl.): 24 [Jun.-Dec. 1775] (Aublet 1775).

*Talisia carinata* Radlk. f. acutisepala Radlk., *Fl. Bras. [Martius]* 13 (3): 549 [1 Apr. 1900] (Radlkofer 1900).

*Talisia micrantha* Radlk., *Repert. Spec. Nov. Regni Veg.* 9: 375 (Radlkofer 1911).

*Talisia reticulata* Radlk., *Repert. Spec. Nov. Regni Veg.* 9: 376 (Radlkofer 1911).

VERNACULAR NAMES. — Pa: tuu-aška • Te: tuli • Wp: tuli átā • Wn: tëpu.

HERBARIUM DATA (FG). — 64 collections at CAY. Sel. exs.: *P. Acevedo-Rodríguez et al. 4911*.

INVENTORY DATA (FG). — 1 tree, dbh = 25.6 cm.

Genus *Toulicia* Aubl.

[1593] *Toulicia elliptica* Radlk.
(Fig. 53D)

*Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 8: 371 (Radlkofer 1878).

VERNACULAR NAMES. — Pa: tuu-kamwi.

HERBARIUM DATA (FG). — A single collection, *D. Sabatier & J.-F. Molino 5076*.

INVENTORY DATA (FG). — 41 trees in 7 plots; Fmax = 2.5%; dbhinv = 47.4 cm.

[1594] *Toulicia guianensis* Aubl.

*Hist. Pl. Guiane* 1: 359 [Jun.-Dec. 1775] (Aublet 1775).

*Toulicia pulvinata* Radlk., *Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 8: 371 (Radlkofer 1878).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020), but not yet found in inventories of trees above 10 cm dbh in French Guiana.

VERNACULAR NAMES. — Ka: tulisi, tulusili • Wp: tuli sī.

HERBARIUM DATA (FG). — 21 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000629523]).

SIZE. — Up to 30 m tall (Acevedo-Rodríguez 2012).

Genus *Vouarana* Aubl.

[1595] *Vouarana guianensis* Aubl.

*Hist. Pl. Guiane* 2 (Suppl.): 12 [Jun.-Dec. 1775] (Aublet 1775).

VERNACULAR NAMES. — Wp: tápě lemiŭ’i • Br: maraximbé.

HERBARIUM DATA (FG). — 28 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000645706]); D. Sabatier 1538 (epitype US[00725189] ; isoepitypes CAY[CAY015974], P[P06671280]).

INVENTORY DATA (FG). — 47 trees in 30 plots; Fmax < 1%; dbhinv = 30.9 cm.

Family SAPOTACEAE Juss.
Genus *Chromolucuma* Ducke

[1596] *Chromolucuma congestifolia*
(Pilz) Alves-Araújo & M.Alves

*Brittonia* 64 (1): 28 (Alves-Araújo & Alves 2012). — *Pouteria congestifolia* Pilz, *Ann. Missouri Bot. Gard.* 68 (1): 191 [17 Nov. 1981] (Pilz 1981).

HERBARIUM DATA (FG). — A single collection, *T.D. Pennington et al. 13840* (K).

SIZE. — Costa Rica. *T.D. Pennington 11539* (MO), 12 m × 30 cm.

Genus *Chrysophyllum* L.

[1597] *Chrysophyllum argenteum* Jacq.
subsp. *auratum* (Miq.) T.D.Penn.

*Fl. Neotrop. Monogr.* 52: 545 [26 Apr. 1990] (Pennington 1990). — *Chrysophyllum auratum* Miq., *Fl. Bras. [Martius]* 7: 97 [15 Jan. 1863] (Miquel 1863). — *Cynodendron auratum* (Miq.) Baehni, *Boissiera* 11: 143 (Baehni 1965).

*Chrysophyllum sericeum* A.DC., *Prodr. [A. P. de Candolle]* 8: 158 [mid Mar. 1844] (Candolle 1844), *non* Salisb. (1796, nom. nud.)

*Chrysophyllum richardii* Klotzsch ex Miq., *Fl. Bras. [Martius]* 7: 97 [15 Jan. 1863] (Miquel 1863), *nom. nud. pro syn.*
*Chrysophyllum auratum* var. *majus* Miq., *Fl. Bras.* [*Martius*] 7: 98 [15 Jan. 1863] (Miquel 1863).

*Chrysophyllum guyanense* Klotzsch ex Miq., *Fl. Bras.* [*Martius*] 7: 98 [15 Jan. 1863] (Miquel 1863), *nom. nud. pro syn.*

*Chrysophyllum hostmannianum* Klotzsch ex Miq., *Fl. Bras.* [*Martius*] 7: 98 [15 Jan. 1863] (Miquel 1863), *nom. nud. pro syn.*

*Chrysophyllum nitidissimum* Klotzsch ex Miq., *Fl. Bras.* [*Martius*] 7: 98 [15 Jan. 1863] (Miquel 1863), *nom. nud. pro syn.*

*Chrysophyllum sericeum* var. *acutifolium* Miq., *Fl. Bras.* [*Martius*] 7: 98 [15 Jan. 1863] (Miquel 1863).

*Chrysophyllum sericeum* var. *obtusifolium* Miq., *Fl. Bras.* [*Martius*] 7: 98 [15 Jan. 1863] (Miquel 1863).

**VERNACULAR NAMES.** — Ka: kalu melei, kameli • Wp: kalamūlī sili, wai piyū, wai piyū sili, wai to wilā • Cr: zoliv • Br: ajará, guajará, uajara-branco.

**HERBARIUM DATA (FG).** — 60 collections at CAY. Sel. exs.: *A. Gabriel s.n.* (holotype of *Chrysophyllum sericeum*: G-DC[G00139532]; iso-, F[V0071948F], G[G00434796, G00434797]).

**INVENTORY DATA (FG).** — 28 trees in 13 plots; $F_{max} < 1\%$; $dbh_{inv} = 61.4$ cm.

[1598] *Chrysophyllum argenteum*
subsp. *nitidum* (G.Mey.) T.D.Penn.

*Fl. Neotrop. Monogr.* 52: 548 [26 Apr. 1990] (Pennington 1990). — *Chrysophyllum nitidum* G.Mey., *Prim. Fl. Esseq.:* 116 [Nov. 1818] (Meyer 1818). — *Cynodendron nitidum* (G.Mey.) Aubrév., *Adansonia*, n.s., 7 (4): 466 [Dec. 1967] (Aubréville 1967).

*Chrysophyllum psilophyllum* A.DC., *Prodr. [A. P. de Candolle]* 8: 160 [mid Mar. 1844] (Candolle 1844).

**NOTE.** — Known only from the Guiana Shield.

**VERNACULAR NAMES.** — Ka: kalu melei, kameli • Wp: wai • Wn: wapolimë • Nt: kwata bobi, nyanboka • Cr: zoliv • Br: ajará, guajará.

**HERBARIUM DATA (FG).** — 25 collections at CAY. Sel. exs.: *J. Martin s.n.* (holotype of *Chrysophyllum psilophyllum*: P[P00649300]; iso-, F[V0071943F], P[P00649301, P00649302, P00649303]; possible iso-, US[00323625]).

**INVENTORY DATA (FG).** — 250 trees in 108 plots; $F_{max} = 1.9\%$; $dbh_{inv} = 43.9$ cm.

[1599] *Chrysophyllum cuneifolium* (Rudge) A.DC.

*Prodr. [A. P. de Candolle]* 8: 160 [mid Mar. 1844] (Candolle 1844). — *Bumelia cuneifolia* Rudge, *Pl. Guian. [Rudge]* 1 (4): 30 [Apr.-May 1806] (Rudge 1806). — *Prieurella cuneifolia* (Rudge) Pierre ex Aubrév., *Adansonia*, n.s., 1 (1): 19 [Nov. 1961] (Aubréville 1961), *nom. inval.* — *Ecclinusa cuneifolia* (Rudge) Aubrév., *Adansonia*, n.s., 1 (1): 20 [Nov. 1961] (Aubréville 1961), *nom. inval.*

**NOTES.** — The new combinations *Prieurella cuneifolia* and *Ecclinusa cuneifolia* were not validly published because the basionym was not explicitly cited (Turland *et al.* 2018: Art. 41.3). *Prieurella cuneifolia* (Rudge) Pierre ex Aubrév. appeared on p. 69 of Pierre's *Notes Botaniques: Sapotaceae*, but pp. 69-83 of that work were never published (they exist only as proofs at P).

**VERNACULAR NAMES.** — Pa: uu-kamwi-wahuyo, wakukwa-adudu • Ka: atakamala, kusili palatali • Te: wila pulu'a • Wp: wila pele • Nt: kwata bobi • Cr: tété-kwata • Br: abiurana, maçarandúbarana.

**HERBARIUM DATA (FG).** — 47 collections at CAY. Sel. exs.: *J. Martin s.n.* (original material BM[BM000952593]).

**INVENTORY DATA (FG).** — 51 trees in 36 plots; $F_{max} < 1\%$; $dbh_{inv} = 22.1$ cm.

[1600] *Chrysophyllum durifructum* (W.A.Rodrigues)
T.D.Penn.

*Fl. Neotrop. Monogr.* 52: 604 [26 Apr. 1990] (Pennington 1990). — *Achrouteria durifructa* W.A.Rodrigues, *Acta Amazonica* 4 (3): 15 (Rodrigues 1974).

**HERBARIUM DATA (FG).** — 4 collections at CAY. Sel. exs.: *S.A. Mori et al.* 24828.

**INVENTORY DATA (FG).** — 16 trees in 4 plots; $F_{max} = 1.3\%$; $dbh_{inv} = 84$ cm.

[1601] *Chrysophyllum eximium* Ducke

*Bull. Mus. Natl. Hist. Nat.*, sér. 2, 4 (6): 744 (Ducke 1932). — *Ecclinusa eximia* (Ducke) Cronquist, *Bull. Torrey Bot. Club* 73 (3): 310 [June 1946] (Cronquist 1946).

*Chrysophyllum rufocupreum* Ducke, *Arch. Inst. Biol. Veg.* 2 (1): 67 [Sep. 1935] (Ducke 1935). — *Ecclinusa rufocuprea* (Ducke) Cronquist, *Bull. Torrey Bot. Club* 73 (3): 311 [June 1946] (Cronquist 1946).

**VERNACULAR NAMES.** — Nt: busi apaa.

**HERBARIUM DATA (FG).** — 9 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 2568.*

**INVENTORY DATA (FG).** — 107 trees in 30 plots; $F_{max} = 1.5\%$; $dbh_{inv} = 66.3$ cm.

[1602] *Chrysophyllum lucentifolium* Cronquist
subsp. *pachycarpum* Pires & T.D.Penn.

*Fl. Neotrop. Monogr.* 52: 606 [26 Apr. 1990] (Pires & Pennington 1990).

**HERBARIUM DATA (FG).** — 43 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 2696.*

**INVENTORY DATA (FG).** — 109 trees in 46 plots; $F_{max} = 2.4\%$; $dbh_{inv} = 125$ cm.

[1603] *Chrysophyllum manaosense* (Aubrév.) T.D.Penn.

*Fl. Neotrop. Monogr.* 52: 598 [26 Apr. 1990] (Pennington 1990). — *Prieurella manaosensis* Aubrév., *Adansonia*, n.s., 4 (3): 370 [Dec. 1964] (Aubréville 1964).

**HERBARIUM DATA (FG).** — A single collection, *D. Sabatier & J.-F. Molino 5134.*

**INVENTORY DATA (FG).** — 6 trees in 6 plots; $F_{max} = 1.2\%$; $dbh_{inv} = 34$ cm.
[1604] *Chrysophyllum pomiferum* (Eyma) T.D.Penn.

Fl. Neotrop. Monogr. 52: 602 [26 Apr. 1990] (Pennington 1990). — *Achrouteria pomifera* Eyma, *Recueil Trav. Bot. Néerl.* 33: 193 (Eyma 1936). — *Pouteria pomifera* (Eyma) Baehni, *Candollea* 9: 353 (Baehni 1942).

*Planchonella guianensis* P.Royen, *Blumea* 8: 395 (Royen 1957).

VERNACULAR NAMES. — Pa: aku-aiwut • Wn: tèpepuluimè • Nt: mongii soke • Cr: balata-ponm • Fr: jaune d'œuf à grandes feuilles • Br: abiurana.

HERBARIUM DATA (FG). — 22 collections at CAY. Sel. exs.: *D. Sabatier* 984.

INVENTORY DATA (FG). — 81 trees in 58 plots; Fmax < 1%; dbhinv = 79.1 cm.

[1605] *Chrysophyllum prieurii* A.DC.
(Fig. 54A)

Prodr. [A. P. de Candolle] 8: 161 [mid Mar. 1844] (Candolle 1844), "Prieurei", — *Ecclinusa prieurii* (A.DC.) Aubrév., *Adansonia*, n.s., 1 (1): 20 [Nov. 1961] (Aubréville 1961). — *Prieurella prieurii* (A.DC.) Aubrév., *Adansonia*, n.s., 4 (3): 370 [Dec. 1964] (Aubréville 1964).

*Chrysophyllum cyanogenum* Ducke, *Trop. Woods* 71: 18 (Ducke 1942). — *Ecclinusa cyanogena* (Ducke) Aubrév., *Adansonia*, n.s., 1 (1): 20 [Nov. 1961] (Aubréville 1961).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: panauna-priye, uu-kamwi • Wp: takamala • Nt: pepee boyti • Cr: apisi-jonn, balata-jonndèf • Fr: balata indien, balata jaune d'œuf • Br: abiurana-mocambo, abiurana-vermelha, maçarandúbarana.

HERBARIUM DATA (FG). — 44 collections at CAY. Sel. exs.: *FM.R. Leprieur s.n.* (holo-, G-DC[G00139685]; iso-, GH[GH00075582], P[P00649386, P00649387]).

INVENTORY DATA (FG). — 350 trees in 138 plots; Fmax = 1.9%; dbhinv = 71.9 cm.

[1606] *Chrysophyllum sanguinolentum* (Pierre) Baehni

Boissiera 11: 74 (Baehni 1965). — *Ragala sanguinolenta* Pierre, *Notes Bot. Sapot.* 2: 60 (Pierre 1891). — *Ecclinusa sanguinolenta* Pierre, *Notes Bot. Sapot.* 2: 58 (Pierre 1891), *nom. nud. pro syn.* — *Ecclinusa sanguinolenta* (Pierre) Engl., *Nat. Pflanzenfam.* [*Engler & Prantl*] *Nachtr.* 1: 278 (Engler 1897).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Nt: switi amini • Cr: balata-senj-rouj • Fr: balata pommier • Br: balata-brava, ucuquirana.

HERBARIUM DATA (FG). — 42 collections at CAY. Sel. exs.: *E.M. Mélinon* 42, 1862 (holo-, P[P00649392]; iso-, BM[BM000952597], F[V0072216F, V0072217F], US[00067565, 00067566, 00067567]).

INVENTORY DATA (FG). — 399 trees in 123 plots; Fmax = 5.3%; dbhinv = 76.9 cm.

[1607] *Chrysophyllum sparsiflorum* Klotzsch ex Miq.

Fl. Bras. [Martius] 7: 90 [15 Jan. 1863] (Miquel 1863).

*Chrysophyllum sparsiflorum* var. *fagifolium* Miq., *Fl. Bras.* [Martius] 7: 90 [15 Jan. 1863] (Miquel 1863).

*Chrysophyllum steyermarkii* Monach., *Fieldiana*, *Bot.* 28 (3): 480 (Monachino 1953).

VERNACULAR NAMES. — Br: mangabarana.

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *C. Feuille* 3936.

INVENTORY DATA (FG). — 9 trees in 7 plots; Fmax < 1%; dbhinv = 15.4 cm.

[1608] *Chrysophyllum venezuelanense* (Pierre) T.D.Penn.

Fl. Neotrop. Monogr. 52: 607 [26 Apr. 1990] (Pennington 1990). — *Cornuella venezuelanensis* Pierre, *Notes Bot. Sapot.* 2: 68 (Pierre 1891).

*Chrysophyllum excelsum* Huber, *Bol. Mus. Paraense Hist. Nat. Ethnogr.* 3: 55 (Huber 1900). — *Gambeya excelsa* (Huber) Aubrév., *Notul. Syst. (Paris)* 16: 248 (Aubréville 1961).

*Lucuma lucentifolia* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 4 (8): 251 [24 Oct. 1929] (Standley 1929). — *Pouteria lucentifolia* (Standl.) Baehni, *Candollea* 9: 424 (Baehni 1942).

*Lucuma pentasperma* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 4 (8): 251 [24 Oct. 1929] (Standley 1929). — *Pouteria pentasperma* (Standl.) Baehni, *Candollea* 9: 353 (Baehni 1942).

*Pouteria petenensis* Lundell, *Wrightia* 5 (1): 6 (Lundell 1972).

*Pouteria dibrachiata* Lundell, *Wrightia* 5 (4): 95 (Lundell 1975).

*Pouteria mayana* Lundell, *Wrightia* 5 (4): 100 (Lundell 1975). — *Achras mayana* (Lundell) Lundell, *Wrightia* 5 (9): 350 (Lundell 1977).

NOTE. — Incipiently domesticated by pre-Columbian Amerindians (Levis *et al.* 2017).

VERNACULAR NAMES. — Wp: wila kea.

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *M.-F. Prévost* 4166.

INVENTORY DATA (FG). — 1 tree, dbh = 12.6 cm.

[1609] *Chrysophyllum* sp. A

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino* 5332.

INVENTORY DATA (FG). — 15 trees in 13 plots; Fmax < 1%; dbhinv = 20.7 cm.

[1610] *Chrysophyllum* sp. B

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: *D. Sabatier* 1464.

INVENTORY DATA (FG). — 24 trees in 17 plots; Fmax < 1%; dbhinv = 92 cm.
<figure><img src="image_79.png" /></figure>

<figure><img src="image_80.png" /></figure>

<figure><img src="image_81.png" /></figure>

<figure><img src="image_82.png" /><figcaption>FIG. 53. — Sapindaceae: **A**, *Allophylus edulis* (A.St.-Hil., A.Juss. & Cambess.) Hieron. ex Niederl. (*D. Sabatier & M.-F. Prévost 1648*); **B**, *Cupania scrobiculata* Rich. (*J.-F. Molino & D. Sabatier 2484*); **C**, *Matayba opaca* Radlk. (*D. Sabatier 2102*); **D**, *Toulicia elliptica* Radlk. (*D. Sabatier & J.-F. Molino 5076*). A, C, D, © D. Sabatier/IRD; B, © J.-F. Molino/IRD.</figcaption></figure>

[1611] *Chrysophyllum* sp. C

HERBARIUM DATA (FG). — A single collection, *D. Sabatier 3535*, dbh = 35 cm.

[1612] *Chrysophyllum* sp. D

HERBARIUM DATA (FG). — A single collection, *D. Sabatier et al. 5262*.

INVENTORY DATA (FG). — 24 trees in 5 plots; Fmax = 1.9%; dbhinv = 38.5 cm.

[1613] *Chrysophyllum* sp. E

HERBARIUM DATA (FG). — 5 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 5651*.

INVENTORY DATA (FG). — 5 trees in 5 plots; Fmax < 1%; dbhinv = 79.3 cm.

[1614] *Chrysophyllum* sp. F

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier 2815*.

INVENTORY DATA (FG). — 1 tree, dbh = 11 cm.

## Genus *Diploön* Cronquist

[1615] *Diploön cuspidatum* (Hoehne) Cronquist

Bull. Torrey Bot. Club 73 (5): 466 [Oct. 1946] (Cronquist 1946). — *Chrysophyllum cuspidatum* Hoehne, *Ostenia*: 302 (Hoehne 1933).

*Diploön venezuelanum* Aubrév., *Mem. New York Bot. Gard.* 23: 225 (Aubréville 1972), “venezuelana”.

HERBARIUM DATA (FG). — 17 collections at CAY. Sel. exs.: *J.-F. Molino et al. 2076*.

INVENTORY DATA (FG). — 68 trees in 30 plots; Fmax = 1.4%; dbhinv = 57.2 cm.

## Genus *Ecclinusa* Mart.

[1616] *Ecclinusa guianensis* Eyma

Recueil Trav. Bot. Néerl. 33: 203 (Eyma 1936). — *Chrysophyllum guianense* (Eyma) Baehni, *Boissiera* 11: 74 (Baehni 1965), “guyanense”.

*Ecclinusa bacuri* Aubrév. & Pellegr., *Adansonia*, n.s., 1 (1): 21 [Nov. 1961] (Aubréville & Pellegrin 1961).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).
VERNACULAR NAMES. — Ka: balatabali, kulumoto, matamata wewe, palatabali • Wp: kalamüli, kalamüli sili, tamanuwa • Nt: busi apaa, malobi • Cr: balata-blan, bataballi • Fr: balata bali • Br: abiurana-abiu, abiurana-bacuri, abiurana-caju, caramuri.

HERBARIUM DATA (FG). — 43 collections at CAY. Sel. exs.: *R.A.A.*
*Oldeman T-762*.

INVENTORY DATA (FG). — 329 trees in 87 plots; Fmax = 24.1 %; dbhinv = 71 cm.

[1617] *Ecclinusa lanceolata* (Mart. & Eichler) Pierre

*Notes Bot. Sapot.* 2: 57 (Pierre 1891). — *Passaveria lanceolata*
Mart. & Eichler, *Fl. Bras. [Martius]* 7: 86 [15 Jan. 1863] (Mart-
tius & Eichler 1863).

VERNACULAR NAMES. — Nt: malobi.

HERBARIUM DATA (FG). — 17 collections at CAY. Sel. exs.: *D.*
*Sabatier 3519*.

INVENTORY DATA (FG). — 8 trees in 5 plots; Fmax < 1%; dbhinv = 22 cm.

[1618] *Ecclinusa ramiflora* Mart.

*Flora* 22 (1, Beibl.): 2 (Martius 1839). — *Chrysophyllum ramiflorum* (Mart.) A.DC., *Prodr. [A. P. de Candolle]* 8: 158 [mid Mar. 1844] (Candolle 1844). — *Passaveria obovata* Mart. & Eichler, *Fl. Bras. [Martius]* 7: 87 [15 Jan. 1863] (Martius & Eichler 1863), *nom. illeg. superfl.* (based on *Ecclinusa ramiflora*).

*Passaveria obovata* var. *tomentosa* Miq., *Fl. Bras. [Martius]* 7: 87 [15 Jan. 1863] (Miquel 1863).

*Lucuma tomentosa* Poepp. ex Miq., *Fl. Bras. [Martius]* 7: 87 [15 Jan. 1863] (Miquel 1863), *nom. nud. pro syn.*

*Ecclinusa costata* Pierre, *Notes Bot. Sapot.* 2: 56 (Pierre 1891).

*Ecclinusa abbreviata* Ducke, *Bull. Mus. Natl. Hist. Nat.*, sér. 2, 4 (6): 743 (Ducke 1932). — *Chrysophyllum abbreviatum* (Ducke) Baehni, *Boissiera* 11: 75 (Baehni 1965).

VERNACULAR NAMES. — Wp: miku lapi'a u • Nt: malobi • Br: abiurofo, coquirana-brava.

HERBARIUM DATA (FG). — 39 collections at CAY. Sel. exs.: *D.*
*Sabatier 1027*.

INVENTORY DATA (FG). — 74 trees in 54 plots; Fmax < 1%; dbhinv = 31.8 cm.

Genus *Elaeoluma* Baill.

[1619] *Elaeoluma nuda* (Baehni) Aubrév.

*Mem. New York Bot. Gard.* 23: 224 (Aubréville 1972). — *Pouteria nuda* Baehni, *Candollea* 14: 72 (Baehni 1952).

*Pouteria auyantepuiensis* Steyerm., *Acta Bot. Venez.* 2 (5-8): 278 (Steyermark 1967).

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: *D.*
*Sabatier & M.-F. Prévost 2889*.

INVENTORY DATA (FG). — 34 trees in 23 plots; Fmax = 1.2%; dbhinv = 53.5 cm.

[1620] *Elaeoluma* sp. A

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *M.-F.*
*Prévost & D. Sabatier 4647*.

INVENTORY DATA (FG). — 9 trees in 6 plots; Fmax < 1%; dbhinv = 29.6 cm.

Genus *Manilkara* Adans.

[1621] *Manilkara bidentata* (A.DC.) A.Chev.
subsp. *bidentata*
*(Fig. 54B)*

*Rev. Bot. Appl. Agric. Trop.* 12: 270 (Chevalier 1932). — *Mimusops bidentata* A.DC., *Prodr. [A. P. de Candolle]* 8: 204 [mid Mar. 1844] (Candolle 1844). — *Manilkara balata* Dubard, *Ann. Mus. Colon. Marseille*, sér. 3, 3: 19 (Dubard 1915), *nom. illeg. superfl.* (*Mimusops bidentata* in synonymy).

*Mimusops sieberi* A.DC., *Prodr. [A. P. de Candolle]* 8: 204 [mid Mar. 1844] (Candolle 1844). — *Mimusops balata* (Aubl.) C.F.Gaertn. var. sieberi (A.DC.) Pierre, *Bull. Mens. Soc. Linn. Paris* 1 (64): 508 (Pierre 1885). — *Manilkara balata* var. sieberi (A.DC.) Dubard, *Ann. Mus. Colon. Marseille*, sér. 3, 3: 20 (Dubard 1915).

*Sapota mulleri* Blume, *Ann. Sci. Nat.*, Bot. sér. 4, 7: 225 (Blume 1857).

*Mimusops balata* var. gutta Pierre, *Bull. Mens. Soc. Linn. Paris* 1 (64): 507 (Pierre 1885). — *Manilkara balata* var. gutta (Pierre) Dubard, *Ann. Mus. Colon. Marseille*, sér. 3, 3: 21 (Dubard 1915).

*Mimusops balata* var. melinonii Pierre, *Bull. Mens. Soc. Linn. Paris* 1 (64): 507 (Pierre 1885), “Melinonis”. — *Manilkara balata* var. melinonii (Pierre) Dubard, *Ann. Mus. Colon. Marseille*, sér. 3, 3: 21 (Dubard 1915), “Melinonis”.

*Mimusops balata* var. schomburgkii Pierre, *Bull. Mens. Soc. Linn. Paris* 1 (64): 507 (Pierre 1885). — *Manilkara balata* var. schomburgkii (Pierre) Dubard, *Ann. Mus. Colon. Marseille*, sér. 3, 3: 20 (Dubard 1915).

*Mimusops balata* var. cruegeri Pierre, *Symb. Antill. [Urban]* 5 (1): 165 [20 May 1904] (Pierre 1904). — *Manilkara balata* var. cruegeri (Pierre) Dubard, *Ann. Mus. Colon. Marseille*, sér. 3, 3: 20 (Dubard 1915). — *Manilkara bidentata* var. cruegeri (Pierre) A.Chev., *Rev. Bot. Appl. Agric. Trop.* 12: 275 (Chevalier 1932).

*Mimusops balata* var. hartii Pierre, *Symb. Antill. [Urban]* 5 (1): 166 [20 May 1904] (Pierre 1904). — *Manilkara balata* var. hartii (Pierre) Dubard, *Ann. Mus. Colon. Marseille*, sér. 3, 3: 21 (Dubard 1915).

*Mimusops darienensis* Pittier, *Contr. U.S. Natl. Herb.* 18 (6): 249 [22 Sep. 1917] (Pittier 1917). — *Manilkara darienensis* (Pittier) Standl., *Trop. Woods* 31: 45 (Standley 1932).

*Manilkara williamsii* Standl., *Publ. Field Mus. Nat. Hist.*, Bot. Ser. 22 (3): 165 (Standley 1940).

NOTES. — Hyperdominant in Amazonia (ter Steege *et al.* 2020). *Mimusops balata* sensu Pierre, *non* (Aubl.) C.F.Gaertn., is a misinterpretation of *Achras balata* Aubl. The latter was described from a tree imported from the Mauritius Island (“Île de France”) and cultivated at Cayenne (Aublet 1775). The type of *A. balata* is in P-JU, and was obtained from a plant growing in the Mascarene Islands (Chevalier 1932, Pennington 1990). Although Aublet in his *Histoire des plantes* did not mention the Guianan “Balata”, he
collected specimens of this species, which are in P and P-JU (Chevalier 1932). Pierre (1885) correctly stated that these 2 specimens were conspecific with the type of *Mimusops bidentata* A.DC., but wrongly concluded that they were type material of *A. balata*. Although aware of the previous transfer of *A. balata* to *Mimusops L.* by Gaertner (1807: 133), Pierre reproposed the same combination, as "*Mimusops balata* non Gaertner". Notwithstanding, all varieties created by Pierre (1885) under *Mimusops balata* are legitimate under *Mimusops balata* (Aubl.) C.F.Gaertn. *Manilkara balata* is similarly confused: it is based on "*Mimusops balata* Pierre" and *Mimusops bidentata* is a synonym, which accords with the Guianan plant, even though *A. balata* was also cited in synonymy. When the latter is excluded from synonymy, then *Manilkara balata* (Aubl.) Dubard is superfluous (because *Mimusops bidentata* is in synonymy) and is therefore illegitimate. Bleekrode (1857) ascribed both the name and description of *Sapota mulleri* to Blume, hence authorship is "Blume", not "Blume ex Bleekrode"; by the way, although the author of the "*Notice sur la gutta-percha de Surinam*", as it appears on the first page, is "M. le Professeur Bleekrod de l'Académie de Delft", he is actually Salomon Abraham Bleekrode (not "Bleekrod", see https://nl.wikipedia.org/wiki/Salomon_Abraham_Bleekrode). The epithet "*melinonis*", which honours the French botanist E. Mélinon, is to be automatically corrected to "*melinonii*" (Turland *et al.* 2018: Art. 60.8); hence *Manilkara balata* var. *melinonii* and *Mimusops balata* var. *melinonii*.

**VERNACULAR NAMES.** — Pa: balata • Ka: palata, polata • Te: balataï • Wp: iwïi, iwilâ, masalâni • Wn: palakta, palaktaimë • Nt: boyti • Cr: balata-fran, balata-gonm • Fr: balata franc • Br: balateira, maçarandúba, maparajúba.

**HERBARIUM DATA (FG).** — 53 collections at CAY. Sel. exs.: *L. C. Richard s.n.* (holo-, P[P00645496]; iso-, P[P00645494, P00645495]).

**INVENTORY DATA (FG).** — 649 trees in 139 plots; $F_{max} = 7.4\%$; $dbh_{inv} = 111.4$ cm.

[1622] *Manilkara bidentata*
subsp. *surinamensis* (Miq.) T.D.Penn.

Fl. Neotrop. Monogr. 52: 61 [26 Apr. 1990] (Pennington 1990). — *Mimusops surinamensis* Miq., Fl. Bras. [Martius] 7: 43 [15 Jan. 1863] (Miquel 1863). — *Kaukenia surinamensis* (Miq.) Kuntze, Revis. Gen. Pl. 2: 406 [5 Nov. 1891] (Kuntze 1891). — *Manilkara surinamensis* (Miq.) Dubard, Ann. Mus. Colon. Marseille, sér. 3, 3: 22 (Dubard 1915).

*Mimusops balata* var. *domingensis* Pierre, Bull. Mens. Soc. Linn. Paris 1 (64): 508 (Pierre 1885). — *Manilkara balata* var. *domingensis* (Pierre) Dubard, Ann. Mus. Colon. Marseille, sér. 3, 3: 21 (Dubard 1915). — *Mimusops domingensis* (Pierre) Moscoso, Cat. Fl. Domingensis 1: 484 (Moscoso 1943).

*Mimusops riedeliana* Pierre ex Baill., Bull. Mens. Soc. Linn. Paris 2 (116): 922 (Baillon 1891).

*Achras nitida* Sessé & Moc., Fl. Mexic., ed. 2: 85 (Sessé & Mociño 1894). — *Mimusops nitida* (Sessé & Moc.) Urb., Symb. Antill. [Urban] 5 (1): 167 [20 May 1904] (Urban 1904). — *Manilkara nitida* (Sessé & Moc.) Dubard, Ann. Mus. Colon. Marseille, sér. 3, 3: 18 (Dubard 1915).

*Mimusops riedleana* Pierre ex Duss, Ann. Inst. Colon. Marseille 3: 387 ["1896" publ. 1897] (Duss 1897). — *Manilkara riedleana* (Pierre ex Duss) Dubard, Ann. Mus. Colon. Marseille, sér. 3, 3: 17 (Dubard 1915).

*Mimusops amazonica* Huber, Bol. Mus. Goeldi Hist. Nat. Ethnogr. 4: 433 (Huber 1906). — *Manilkara amazonica* (Huber) Standl., Trop. Woods 34: 41 (Standley 1933).

*Mimusops maparajuba* Huber, Bol. Mus. Goeldi Hist. Nat. Ethnogr. 4: 434 (Huber 1906).

*Manilkara longiciliata* Ducke, Trop. Woods 71: 22 (Ducke 1942).

*Mimusops longiciliata* Ducke, Trop. Woods 71: 22 (Ducke 1942).

*Manilkara siqueiraei* Ducke, Trop. Woods 71: 24 (Ducke 1942).

*Mimusops siqueiraei* Ducke, Trop. Woods 71: 24 (Ducke 1942).

**NOTES.** — Ducke (1942: 22) simultaneously published *Manilkara longiciliata* and *Mimusops longiciliata* as alternative names, but both are nevertheless valid, because they were published before 1953 (Turland *et al.* 2018: Art. 36.3). The same applies to *Manilkara siqueiraei* and *Mimusops siqueiraei* (Ducke 1942: 24).

**VERNACULAR NAMES.** — Wp: iwilâ, iwilâ pilâ.

**HERBARIUM DATA (FG).** — 13 collections at CAY. Sel. exs.: *P. Grenand 1488*.

**SIZE.** — Up to 10 cm dbh (Pennington 1990).

[1623] *Manilkara huberi* (Ducke) A.Chev.

Rev. Bot. Appl. Agric. Trop. 12: 276 (Chevalier 1932). — *Mimusops huberi* Ducke, Arch. Jard. Bot. Rio de Janeiro 2: 14 (Ducke 1917).

**NOTE.** — Incipiently domesticated by pre-Columbian Amerindians (Clement 1999; Levis *et al.* 2017).

**VERNACULAR NAMES.** — Pa: balata-duwê • Nt: boyti, kayman buba boyti • Cr: balata • Fr: balata franc • Br: maçarandúba-da-terrafirme, maparajúba.

**HERBARIUM DATA (FG).** — 25 collections at CAY. Sel. exs.: *B. Riéra & D. Sabatier 1904*.

**INVENTORY DATA (FG).** — 76 trees in 34 plots; $F_{max} = 1.9\%$; $dbh_{inv} = 123.3$ cm.

[1624] *Manilkara paraensis* (Huber) Standl.

Trop. Woods 34: 41 (Standley 1933). — *Mimusops paraensis* Huber, Bol. Mus. Goeldi Hist. Nat. Ethnogr. 4: 435 (Huber 1906).

*Mimusops paraensis* var. *densiflora* Huber, Bol. Mus. Goeldi Hist. Nat. Ethnogr. 4: 435 (Huber 1906).

*Mimusops paraensis* var. *discolor* Huber, Bol. Mus. Goeldi Hist. Nat. Ethnogr. 4: 435 (Huber 1906).

**VERNACULAR NAMES.** — Pa: balata-puvemna • Wp: masalâni • Br: maçarandúba-de-folha miuda, maçarandúba-jacaré.

**HERBARIUM DATA (FG).** — 11 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 3422*.

**INVENTORY DATA (FG).** — 1 tree, $dbh = 48.7$ cm.
## Genus *Micropholis* (Griseb.) Pierre

[1625] *Micropholis acutangula* (Ducke) Eyma
(Fig. 54C)

Recueil Trav. Bot. Néerl. 33: 198 (Eyma 1936). — *Sideroxylon acutangulum* Ducke, Arch. Jard. Bot. Rio de Janeiro 4: 159 (Ducke 1925). — *Pouteria acutangula* (Ducke) Baehni, *Candollea* 9: 246 (Baehni 1942). — *Paramicropholis acutangula* (Ducke) Aubrév. & Pellegr., *Adansonia*, n.s., 1 (2): 171 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville & Pellegrin 1962). — *Paralabatia acutangula* (Ducke) Baehni, *Boissiera* 11: 141 (Baehni 1965).

*Micropholis paraensis* Eyma, Recueil Trav. Bot. Néerl. 33: 198 (Eyma 1936), nom. nud. — *Sideroxylon paraense* Huber ex Eyma, Recueil Trav. Bot. Néerl. 33: 198 (Eyma 1936), nom. nud. pro syn.

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: *D. Sabatier & S. Gonzalez* 5371.

INVENTORY DATA (FG). — 3 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 46.7$ cm.

[1626] *Micropholis cayennensis* T.D.Penn.

Fl. Neotrop. Monogr. 52: 220 [26 Apr. 1990] (Pennington 1990).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Pa: uu-kamwi, uu-kamwi-purubumna, yauknabwi-wahuye • Nt: bakuuman, nyanboka • Cr: zoliv.

HERBARIUM DATA (FG). — 24 collections at CAY. Sel. exs.: *B. Maguire et al.* 47074 (holo-, NY[01199690]; iso-, G[G00439289], GH[GH00589970], K[K000641500], R[R000112723], S[S05-10441], SP[SP001602], US[00323728]).

INVENTORY DATA (FG). — 100 trees in 36 plots; $F_{max} = 1.3\%$; $dbh_{inv} = 89.1$ cm.

[1627] *Micropholis egensis* (A.DC.) Pierre

*Symb. Antill. [Urban]* 5 (1): 127 [20 May 1904] (Pierre 1904). — *Sideroxylon egense* A.DC., *Prodr. [A. P. de Candolle]* 8: 182 [mid Mar. 1844] (Candolle 1844). — *Bumelia egensis* Poepp. ex A.DC., *Prodr. [A. P. de Candolle]* 8: 182 [mid Mar. 1844] (Candolle 1844), *nom. nud. pro syn.* — *Sideroxylon rugosum* (Sw.) Roem. & Schult. var. *egense* (A.DC.) Miq., *Fl. Bras. [Martius]* 7: 51 [15 Jan. 1863] (Miquel 1863), “*Sideroxylum*”. — *Pouteria egensis* (A.DC.) Baehni, *Candollea* 9: 207 (Baehni 1942).

*Micropholis martiana* Pierre, *Symb. Antill. [Urban]* 5 (1): 126 [20 May 1904] (Pierre 1904).

*Sideroxylon ulei* K.Krause, Verh. Bot. Vereins Prov. Brandenburg 50: 95 [10 June 1908] (Krause 1908). — *Micropholis ulei* (K.Krause) Eyma, *Recueil Trav. Bot. Néerl.* 33: 198 (Eyma 1936). — *Pouteria ulei* (K.Krause) Baehni, *Candollea* 9: 217 (Baehni 1942).

*Sideroxylon quinilla* Standl. ex Ll.Williams, *Publ. Field Mus. Nat. Hist., Bot. Ser.* 15: 414 (Williams 1936), *nom. inval.* (anglice).

*Pouteria setosa* Baehni, *Candollea* 18: 168 (Baehni 1962).

*Micropholis wurdackii* Aubrév., *Mem. New York Bot. Gard.* 23: 212 (Aubréville 1972).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: paivan • Wp: waa • Wn: tamaniwa • Nt: mamanten • Cr: zoliv • Br: mangabarana.

HERBARIUM DATA (FG). — 34 collections at CAY. Sel. exs.: *D. Sabatier* 1267.

INVENTORY DATA (FG). — 162 trees in 84 plots; $F_{max} = 1\%$; $dbh_{inv} = 85$ cm.

[1628] *Micropholis guyanensis* (A.DC.) Pierre subsp. *guyanensis*

Notes Bot. Sapot. 2: 40 (Pierre 1891), “*Guyanensis*”.

*Sideroxylon guyanense* A.DC., *Prodr. [A. P. de Candolle]* 8: 182 [mid Mar. 1844] (Candolle 1844), “*Guyanense*”.

*Sideroxylon rufum* Mart. & Eichler, *Fl. Bras. [Martius]* 7: 52 [15 Jan. 1863] (Martius & Eichler 1863). — *Micropholis rufa* (Mart. & Eichler) Pierre, *Symb. Antill. [Urban]* 5 (1): 130 [20 May 1904] (Pierre 1904). — *Pouteria rufa* (Mart. & Eichler) Baehni, *Candollea* 9: 199 (Baehni 1942).

*Sideroxylon cyrtobotryum* Mart. ex Miq., *Fl. Bras. [Martius]* 7: 57 [15 Jan. 1863] (Miquel 1863). — *Sprucella cyrtobotrya* (Mart. ex Miq.) Pierre, *Notes Bot. Sapot.* 1: 27 (Pierre 1890). — *Micropholis cyrtobotrya* (Mart. ex Miq.) Baill., *Hist. Pl. [Baillon]* 11: 282 [Sep.-Oct. 1891] (Baillon 1891), *nom. inval.* (genus name and epithet not associated). — *Pouteria cyrtobotrya* (Mart. ex Miq.) Baehni, *Candollea* 9: 199 (Baehni 1942).

*Chrysophyllum melinonii* Engl., *Bot. Jahrb. Syst.* 12: 521 [23 Dec. 1890] (Engler 1890), “*Melinoni*”. — *Pouteria melinonii* (Engl.) Baehni, *Candollea* 9: 200 (Baehni 1942). — *Planchonella melinonii* (Engl.) Baehni, *Boissiera* 11: 67 (Baehni 1965).

*Micropholis chrysophylloides* Pierre, *Notes Bot. Sapot.* 2: 38 (Pierre 1891). — *Pouteria chrysophylloides* (Pierre) Stehlé, Caribbean Forest. 4 (3): 118 (Stehlé 1943).

*Micropholis cruegeriana* Pierre, *Notes Bot. Sapot.* 2: 38 (Pierre 1891). — *Pouteria cruegeriana* (Pierre) Baehni, *Candollea* 9: 206 (Baehni 1942).

*Micropholis portoricensis* Pierre, *Notes Bot. Sapot.* 2: 38 (Pierre 1891).

*Micropholis imrayana* Pierre, *Notes Bot. Sapot.* 2: 39 (Pierre 1891).

*Micropholis portoricensis* var. *curvata* Pierre, *Notes Bot. Sapot.* 2: 39 (Pierre 1891). — *Micropholis curvata* (Pierre) Urb., *Symb. Antill. [Urban]* 5 (1): 124 [20 May 1904] (Urban 1904), in obs.

*Micropholis portoricensis* var. *mesuifolia* Pierre, *Notes Bot. Sapot.* 2: 39 (Pierre 1891).

*Micropholis achradoformis* Pierre, *Symb. Antill. [Urban]* 5 (1): 118 [20 May 1904] (Pierre 1904). — *Pouteria achradoformis* (Pierre) Baehni, *Candollea* 9: 207 (Baehni 1942), “*achradiformis*”.

*Micropholis eggersiana* Pierre, *Symb. Antill. [Urban]* 5 (1): 118 [20 May 1904] (Pierre 1904).

*Micropholis truncata* Pierre, *Symb. Antill. [Urban]* 5 (1): 119 [20 May 1904] (Pierre 1904). — *Pouteria truncata* (Pierre) Baehni, *Candollea* 9: 216 (Baehni 1942).

*Micropholis chrysophylloides* var. *truncata* Pierre, *Symb. Antill. [Urban]* 5 (1): 119 [20 May 1904] (Pierre 1904), *nom. nud. pro syn.*
*Micropholis balata* Pierre, *Symb. Antill. [Urban]* 5 (1): 120 [20 May 1904] (Pierre 1904). — *Pouteria balata* (Pierre) Baehni, *Candollea* 9: 351 (Baehni 1942).

*Micropholis discolor* Pierre, *Symb. Antill. [Urban]* 5 (1): 121 [20 May 1904] (Pierre 1904). — *Chrysophyllum discolor* Walp. & Duchass. ex Pierre, *Symb. Antill. [Urban]* 5 (1): 121 [20 May 1904] (Pierre 1904), *nom. nud. pro syn.* — *Pouteria discolor* (Pierre) Stehlé, Caribbean Forest. 4 (3): 118 (Stehlé 1943), *nom. illeg. hom.*, non (Baill.) Baehni (1942).

*Micropholis dominicensis* Pierre, *Symb. Antill. [Urban]* 5 (1): 122 [20 May 1904] (Pierre 1904). — *Pouteria dominensis* (Pierre) Stehlé, *Fl. Guadeloupe* [Stehlé] 4: 121 (Stehlé 1943).

*Micropholis grandifolia* Pierre ex Glaz., *Bull. Soc. Bot. France* 57 (Mém. 3e): 441 (Glaziou 1910), *nom. nud.*

*Sideroxylon williamsii* Baehni, *Candollea* 7: 135 (Baehni 1936). — *Mastichodendron williamsii* (Baehni) Baehni ex Bernardi, *Candollea* 22 (2): 231 (Bernardi 1967).

*Pouteria aeranea* Baehni, *Candollea* 9: 205 (Baehni 1942).

*Pouteria silvicola* Baehni, *Candollea* 18: 161 (Baehni 1962).

*Pouteria bopiensis* Baehni, *Candollea* 18: 164 (Baehni 1962).

*Pouteria piresii* Baehni, *Candollea* 18: 168 (Baehni 1962).

*Micropholis cowanii* Aubrév., *Mem. New York Bot. Gard.* 23: 211 (Aubréville 1972).

**NOTES.** — Hyperdominant in Amazonia (ter Steege *et al.* 2020). *Micropholis curvata* is a valid recombination, by Urban alone (not "Pierre & Urban" as in Pennington, 1990), of *M. portoricensis* var. *curvata* Pierre.

**VERNACULAR NAMES.** — Pa: aku-aiwut, uu-kamwi, waaduk-seine, yauknabui • Ka: asepoko, asepuku, kusili palatali, wesepuku, woko wokulu • Wp: wamisalu, watuliya • Wn: kanawa popalin, palakta jalan • Nt: bakuuman, gaan wiwii bakuuman, tapooka mapa • Cr: balata-blan, balata-endjen • Fr: balata blanc à grandes feuilles • Br: balata-rosada, chicle-bravo, maparajúba.

**HERBARIUM DATA (FG).** — 69 collections at CAY. Sel. exs.: *J. Martin s.n.* (holo-, P[P00649234]).

**INVENTORY DATA (FG).** — 382 trees in 129 plots; $F_{max} = 2.3\%$; $dbh_{inv} = 96$ cm.

[1629] *Micropholis guyanensis* subsp. *duckeana* (Baehni)
T.D.Penn.

*Fl. Neotrop. Monogr.* 52: 180 [26 Apr. 1990] (Pennington 1990). — *Pouteria duckeana* Baehni, *Candollea* 18: 161 (Baehni 1962).

*Micropholis rosadinha-brava* Aubrév. & Pellegr., *Adansonia*, n.s., 1 (2): 178 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville & Pellegrin 1962).

**VERNACULAR NAMES.** — Pa: yauknabui • Br: abiu-balatarana, balata-rosadinha.

**HERBARIUM DATA (FG).** — 19 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier* 2973.

**INVENTORY DATA (FG).** — 73 trees in 29 plots; $F_{max} < 1\%$; $dbh_{inv} = 53$ cm.

[1630] *Micropholis longipedicellata* Aubrév.

*Mem. New York Bot. Gard.* 23: 209 (Aubréville 1972).

**NOTE.** — Known only from the Guiana Shield.

**VERNACULAR NAMES.** — Pa: impitit-ahavukune, mpitit-ahavukune.

**HERBARIUM DATA (FG).** — 16 collections at CAY. Sel. exs.: *D. Sabatier* 2241.

**INVENTORY DATA (FG).** — 55 trees in 26 plots; $F_{max} = 1.6\%$; $dbh_{inv} = 37.6$ cm.

[1631] *Micropholis melinoniana* Pierre

*Notes Bot. Sapot.* 2: 40 (Pierre 1891). — *Pouteria melinoniana* (Pierre) Baehni, *Candollea* 9: 212 (Baehni 1942).

*Sideroxylon rugosum* var. *parviflorum* Miq., *Fl. Bras.* [Martius] 7: 51 [15 Jan. 1863] (Miquel 1863).

*Stephanoluma rugosa* Baill., *Hist. Pl. [Baillon]* 11: 283 [Sep.-Oct. 1891] (Baillon 1891).

*Sideroxylon calophylloides* Lundell, *Contr. Univ. Michigan Herb.* 6: 56 (Lundell 1941), *nom. illeg. hom.*, non (Pierre) Engl. (Engler 1897).

*Micropholis mexicana* Gilly ex Cronquist, *Lloydia* 9 (4): 257 [“1945” publ. 20 Dec. 1946] (Cronquist 1946).

*Micropholis guatemalensis* Lundell, *Wrightia* 5 (4): 92 (Lundell 1975).

**NOTE.** — Baillon based his *Stephanoluma rugosa* on "*Sideroxylon rugosum* A.DC. [...] (non Roem. & Sch.)". Actually, "*Sideroxylon rugosum* (Sw.) Roem. & Schult. by Candolle. Therefore, *Stephanoluma rugosa* is a new name, and its authorship is Baill., not "(A.D.C.) Baill."

**VERNACULAR NAMES.** — Pa: muhut-wašiunó • Wp: waa • Wn: kanawa popalin, wopoli • Nt: mamanten • Cr: balata-blan, zoliv-maypouri • Br: bacuri-de-anta, balata-rosada, caramuri-da-várzea, corrupixá.

**HERBARIUM DATA (FG).** — 45 collections at CAY. Sel. exs.: *E.M. Mélinon s.n.* (holo-, P[P00649246]).

**INVENTORY DATA (FG).** — 142 trees in 59 plots; $F_{max} = 2.9\%$; $dbh_{inv} = 155$ cm.

[1632] *Micropholis mensalis* (Baehni) Aubrév.

*Adansonia*, n.s., 3 (1): 21 [Jan.-Mar. 1963] (Aubréville 1963). — *Pouteria mensalis* Baehni, *Candollea* 14: 64 (Baehni 1952).

**VERNACULAR NAMES.** — Br: abiurana-goiabinha, abiurana-roxa.

**HERBARIUM DATA (FG).** — 11 collections at CAY. Sel. exs.: *D. Sabatier et al.* 6122.

**INVENTORY DATA (FG).** — 24 trees in 11 plots; $F_{max} = 1.3\%$; $dbh_{inv} = 16.9$ cm.
[1633] *Micropholis obscura* T.D.Penn.
(Fig. 54D)

Fl. Neotrop. Monogr. 52: 227 [26 Apr. 1990] (Pennington 1990).

VERNACULAR NAMES. — Nt: akwansiba • Cr: balata-blan, balata-endjen, zoliv • Fr: balata blanc • Br: balata-rosada.

HERBARIUM DATA (FG). — 37 collections at CAY. Sel. exs.: S.A. Mori & B.M. Boom 14784 (holo-, K, not seen; iso-, MA[MA400083, MA400083-2], NY[00901963], U[U0006611]).

INVENTORY DATA (FG). — 152 trees in 62 plots; Fmax = 1.2%; dbhinv = 95 cm.

[1634] *Micropholis porphyrocarpa* (Baehni) Monach.

Brittonia 7: 409 [Oct. 1952] (Monachino 1952). — *Pouteria porphyrocarpa* Baehni, *Candollea* 9: 214 (Baehni 1942).

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: D. Sabatier & M.-F. Prévost 1853.

INVENTORY DATA (FG). — 24 trees in 13 plots; Fmax < 1%; dbhinv = 63.8 cm.

[1635] *Micropholis sanctae-rosae* (Baehni) T.D.Penn.

Fl. Neotrop. Monogr. 52: 215 [26 Apr. 1990] (Pennington 1990). — *Pouteria sanctae-rosae* Baehni, *Candollea* 9: 211 (Baehni 1942).

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: D. Sabatier et al. 4794.

INVENTORY DATA (FG). — 30 trees in 12 plots; Fmax < 1%; dbhinv = 21.6 cm.

[1636] *Micropholis trunciflora* Ducke

Bol. Técn. Inst. Agron. N. 19: 19 (Ducke 1950).

Pouteria klugii Baehni, *Candollea* 14: 76 (Baehni 1952).

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: D. Sabatier & M.-F. Prévost 3298.

INVENTORY DATA (FG). — 5 trees in 4 plots; Fmax < 1%; dbhinv = 44.2 cm.

[1637] *Micropholis venulosa* (Mart. & Eichler) Pierre

Notes Bot. Sapot. 2: 40 (Pierre 1891). — *Sideroxylon venulosum* Mart. & Eichler, *Fl. Bras. [Martius]* 7: 52 [15 Jan. 1863] (Martius & Eichler 1863). — *Lucuma venulosa* Spruce ex Mart. & Eichler, *Fl. Bras. [Martius]* 7: 52 [15 Jan. 1863] (Martius & Eichler 1863), *nom. nud. pro syn.* — *Pouteria venulosa* (Mart. & Eichler) Baehni, *Candollea* 9: 195 (Baehni 1942). — *Xantolis venulosa* (Mart. & Eichler) Baehni, *Boissiera* 11: 24 (Baehni 1965).

*Micropholis calophylloides* Pierre, *Notes Bot. Sapot.* 2: 40 (Pierre 1891). — *Platyluma calophylloides* (Pierre) Baill., *Hist. Pl. [Bail-lon]* 11: 282 [Sep.-Oct. 1891] (Baillon 1891). — *Sideroxylon calophylloides* (Pierre) Engl., *Nat. Pflanzenfam. [Engler & Prantl]* Nachtr. 1: 276 (Engler 1897).

Meioluma guianensis Baill., Hist. Pl. [Baillon] 11: 282 [Sep.-Oct. 1891] (Baillon 1891). — *Sideroxylon guianense* (Baill.) Engl., *Nat. Pflanzenfam. [Engler & Prantl]* Nachtr. 1: 276 (Engler 1897), *nom. illeg. hom., non A.DC.* (Candolle 1844).

*Micropholis mucronata* Pierre, *Symb. Antill. [Urban]* 5 (1): 112 [20 May 1904] (Pierre 1904).

*Micropholis rigida* Pierre, *Symb. Antill. [Urban]* 5 (1): 125 [20 May 1904] (Pierre 1904).

Pouteria polyneura Baehni, *Candollea* 7: 133 (Baehni 1936).

Pouteria rivularis Baehni, *Candollea* 9: 208 (Baehni 1942).

Pouteria flava Baehni, *Candollea* 18: 164 (Baehni 1962).

Pouteria cataractae Baehni, *Candollea* 18: 168 (Baehni 1962).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: inam-etni-seinô, muhut-wašiunô • Ka: tabuku, woko popi • Te: kutsili palatanan • Wp: wai tawa • Nt: bakuuman • Cr: zoliv • Br: abiurana-branca, mulungu-rosa-verde.

HERBARIUM DATA (FG). — 63 collections at CAY. Sel. exs.: *E.M. Mélinon s.n.* (original material of *Meioluma guianensis*: P[P00649260]).

INVENTORY DATA (FG). — 127 trees in 74 plots; Fmax < 1%; dbhinv = 61.6 cm.

[1638] *Micropholis* sp. A

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *D. Sabatier* 2339.

INVENTORY DATA (FG). — 7 trees in 4 plots; Fmax < 1%; dbhinv = 26.8 cm.

## Genus *Pouteria* Aubl.

[1639] *Pouteria ambelaniifolia* (Sandwith) T.D.Penn.

Fl. Neotrop. Monogr. 52: 362 [26 Apr. 1990] (Pennington 1990). — *Chrysophyllum ambelaniifolium* Sandwith, *Bull. Misc. Inform. Kew* 1931 (10): 476 [31 Dec. 1931] (Sandwith 1931). — *Oxythece ambelaniifolia* (Sandwith) Cronquist, *Bull. Torrey Bot. Club* 73 (3): 310 [June 1946] (Cronquist 1946). — *Pseudoxythece ambelaniifolia* (Sandwith) Aubrév., *Mem. New York Bot. Gard.* 23: 219 (Aubréville 1972).

VERNACULAR NAMES. — Pa: waaduk-seine.

HERBARIUM DATA (FG). — 20 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 2973.

INVENTORY DATA (FG). — 138 trees in 73 plots; Fmax = 1.1%; dbhinv = 57.9 cm.

[1640] *Pouteria anomala* (Pires) T.D.Penn.

Fl. Neotrop. Monogr. 52: 344 [26 Apr. 1990] (Pennington 1990). — *Chrysophyllum anomalum* Pires, *Bol. Técn. Inst. Agron. N.* 38: 34 (Pires 1960).

VERNACULAR NAMES. — Br: abiurana-roxa.
HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *D. Loubry* 621.

SIZE. — Up to 70 cm dbh (Pennington 1990).

[1641] *Pouteria aubrevillei* Bernardi

Candollea 22 (2): 231 (Bernardi 1967).

Eremoluma krukoffii Aubrév. & Pellegr., *Adansonia*, n.s., 1 (2): 169 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville & Pellegrin 1962), “Krukoffii”.

Eremoluma wurdackii Aubrév., *Adansonia*, n.s., 5 (2): 197 [July 1965] (Aubréville 1965).

VERNACULAR NAMES. — Pa: uu-kamwi • Wp: ei li • Cr: bwa-kay • Br: abiurana.

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino* 5130.

INVENTORY DATA (FG). — 30 trees in 17 plots; Fmax < 1%; dbhinv = 36.6 cm.

[1642] *Pouteria bangii* (Rusby) T.D.Penn.

Fl. Neotrop. Monogr. 52: 288 [26 Apr. 1990] (Pennington 1990). — *Sideroxylon bangii* Rusby, *Bull. New York Bot. Gard.* 4 (14): 407 [7 Dec. 1907] (Rusby 1907).

Pseudocladia melinonii Baill., *Hist. Pl. [Baillon]* 11: 291 [Sep.-Oct. 1891] (Baillon 1891), “Melinoni”, nom. nud. — *Lucuma melinonii* Engl., *Nat. Pflanzenfam. [Engler & Prantl]* Nachtr. 1: 275 (Engler 1897), “Melinoni”, nom. illeg., based on the illegitimate *Pseudocladia melinonii*.

Podoluma glaziovii Baill. ex Glaz., *Bull. Soc. Bot. France* 57 (Mém. 3e): 442 (Glaziou 1910).

Pouteria scytalophora Eyma, *Recueil Trav. Bot. Néerl.* 33: 181 (Eyma 1936). — *Pseudocladia scytalophora* (Eyma) Aubrév., *Adansonia*, n.s., 1 (2): 165 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville 1962).

Pouteria micans Baehni, *Candollea* 9: 236 (Baehni 1942).

Pseudocladia colombiana Aubrév., *Adansonia*, n.s., 7 (2): 148 [Jul.-Sep. 1967] (Aubréville 1967).

VERNACULAR NAMES. — Ka: lemu epi • Wp: wila tawa • Cr: balata-pwé • Br: abiu-rosadinha, maparajúba-branca.

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *D. Sabatier* 4462.

INVENTORY DATA (FG). — 86 trees in 49 plots; Fmax < 1%; dbhinv = 54.3 cm.

[1643] *Pouteria benai* (Aubrév. & Pellegr.) T.D.Penn. (Fig. 55A)

Fl. Neotrop. Monogr. 52: 274 [26 Apr. 1990] (Pennington 1990). — *Podoluma benai* Aubrév. & Pellegr., *Adansonia*, n.s., 1 (2): 182 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville & Pellegrin 1962), “Benai”.

NOTE. — Known only from French Guiana.

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: *Service Forestier M-207* (holo-, P[P00640533]; iso-, P[P00640534], U[U0006626]).

INVENTORY DATA (FG). — 60 trees in 30 plots; Fmax = 1.2%; dbhinv = 49.7 cm.

[1644] *Pouteria bilocularis* (H.J.P.Winkl.) Baehni

Candollea 9: 229 (Baehni 1942). — *Labatia bilocularis* H.J.P.Winkl., *Repert. Spec. Nov. Regni Veg.* 7: 112 (Winkler 1909).

Pseudocladia neblinaensis Aubrév., *Mem. New York Bot. Gard.* 23: 204 (Aubréville 1972).

VERNACULAR NAMES. — Pa: impitit-wašiunō, mpitit-wašiunō • Br: abiurana.

HERBARIUM DATA (FG). — 20 collections at CAY. Sel. exs.: *D. Sabatier* 3524.

INVENTORY DATA (FG). — 91 trees in 61 plots; Fmax < 1%; dbhinv = 43.6 cm.

[1645] *Pouteria brachyandra* (Aubrév. & Pellegr.) T.D.Penn.

Fl. Neotrop. Monogr. 52: 404 [26 Apr. 1990] (Pennington 1990). — *Radlkoferella brachyandra* Aubrév. & Pellegr., *Adansonia*, n.s., 1 (2): 186 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville & Pellegrin 1962).

VERNACULAR NAMES. — Wp: pakuku, yá laânga.

HERBARIUM DATA (FG). — 22 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino* 5005.

INVENTORY DATA (FG). — 16 trees in 10 plots; Fmax < 1%; dbhinv = 35.3 cm.

[1646] *Pouteria caimito* (Ruiz & Pav.) Radlk.

Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 12: 312 (Radlkofer 1882). — *Achras caimito* Ruiz & Pav., *Flora Peruviana* 3: 18 (Ruiz & Pavón 1802). — *Lucuma caimito* (Ruiz & Pav.) Roem. & Schult., *Syst. Veg. [Roemer & Schultes]* 4: 701 [Mar.-June 1819] (Roemer & Schultes 1819). — *Caleatia caimito* (Ruiz & Pav.) Mart. ex Steud., *Nomencl. Bot. [Steudel]*, ed. 2, 2: 76, (Steudel 1841) *nom. nud. pro syn.* — *Labatia caimito* (Ruiz & Pav.) Mart., *Flora* 21 (2, Beibl.): 90 (Martius 1838). — *Guapeba caimito* (Ruiz & Pav.) Pierre, *Notes Bot. Sapot.* 2: 42 (Pierre 1891).

Guapeba laurifolia Gomes, Mem. Math. Phis. Acad. Real Sci. Lisboa 3 (Mem. Corresp. 1): 21 (Gomes 1812). — *Lucuma laurifolia* (Gomes) A.DC., *Prodr. [A. P. de Candolle]* 8: 166 [mid Mar. 1844] (Candolle 1844). — *Pouteria laurifolia* (Gomes) Radlk., *Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 12: 333 (Radlkofer 1882). — *Pouteria caimito* var. *laurifolia* (Gomes) Baehni, *Candollea* 9: 260 (Baehni 1942).

Lucuma temare Kunth, *Nova genera et species plantarum [H.B.K.]* 3: 241 [9 July 1819] (Kunth 1819). — *Richardella temare* (Kunth) Pierre, *Notes Bot. Sapot.* 1: 20 (Pierre 1890). — *Pouteria temare* (Kunth) Aubrév., *Adansonia*, n.s., 1 (2): 157 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville 1962).

Labatia reticulata Mart., *Flora* 21 (2, Beibl.): 90 (Martius 1838). — *Lucuma laurifolia* var. *reticulata* (Mart.) A.DC., *Prodr. [A. P. de*
*Candolle* 8: 166 [mid Mar. 1844] (Candolle 1844). — *Guapeba brasiliensis* Steud., *Nomencl. Bot. [Steudel]*, ed. 2, 1: 708 (SteuDEL 1840), *nom. illeg. superfl.* (based on *Labatia reticulata*).

*Labatia lasiocarpa* Mart., *Flora* 21 (2, Beibl.): 91 (Martius 1838). — *Lucuma lasiocarpa* (Mart.) A.DC., *Prodr. [A. P. de Candolle]* 8: 166 [mid Mar. 1844] (Candolle 1844). — *Pouteria lasiocarpa* (Mart.) Radlk., *Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 12: 333 (Radlkofer 1882). — *Guapeba lasiocarpa* (Mart.) Pierre, *Notes Bot. Sapot.* 2: 42 (Pierre 1891).

*Achas guapeba* Casar., *Nov. Stirp. Bras.* 7: 61 [Sep ("Jul") 1843] (Casaretto 1843).

*Pouteria leucophaea* Baehni, *Candollea* 18: 176 (Baehni 1962).

**NOTES.** — Hyperdominant in Amazonia (ter Steege *et al.* 2020). Domesticated by pre-Columbian Amerindians (Clement 1999; Levis *et al.* 2017).

**VERNACULAR NAMES.** — Ka: atakamala, kusili palatali • Wp: wila kea • Wn: alimihmo • Cr: zoliv • Br: abiu, abiurana-acariquara.

**HERBARIUM DATA (FG).** — 19 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier* 2564.

**INVENTORY DATA (FG).** — 19 trees in 7 plots; $F_{max} = 1.2\%$; $dbh_{inv} = 32.8$ cm.

### [1647] *Pouteria cayennensis* (A.DC.) Eyma

*Recueil Trav. Bot. Néerl.* 33: 174 (Eyma 1936). — *Chrysophyllum cayennense* A.DC., *Prodr. [A. P. de Candolle]* 8: 160 [mid Mar. 1844] (Candolle 1844), "Cayennense". — *Richardella cayennensis* (A.DC.) Aubrév., *Adansonia*, sér. 2, 11 (2): 300 [July 1971] (Aubréville 1971).

*Chrysophyllum sessiliflorum* Poir., *Encycl. [J. Lamarck et al.]* Suppl. 2: 16 [23 Oct. 1811] (Poiret 1811).

*Lucuma pulverulenta* Mart. & Eichler, *Fl. Bras. [Martius]* 7: 70 [15 Jan. 1863] (Martius & Eichler 1863). — *Vitellaria pulverulenta* (Mart. & Eichler) Radlk., *Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 12: 326 (Radlkofer 1882).

**NOTE.** — Pennington (1990: 366) states that the holotype is at P, and that there is an isotype at G-DC. However, there is at G-DC a specimen (G00139668) bearing a label in the hand of Alphonse de Candolle, which reads "*Chrysophyllum Cayennense* A. DC.", as well as another which reads "Sapotaceae. Cayenne ou Guyane françoise. Museum de Paris 1821". There is no reason not to admit that it is the very specimen to which Candolle refers in the protologue of *Chrysophyllum cayennense*, in the form "*v. s. comm. a Mus. par.*" (i.e., "an exsiccatum that I saw, given by the Paris Museum"). In other words, the specimen at G-DC (G00139668) must be the holotype.

**HERBARIUM DATA (FG).** — 12 collections at CAY. Sel. exs.: *J. Martin s.n.* (holo-, G-DC[G00139668]; iso-, P[P00640548, P00640549, P00640550], US[00037030]).

**INVENTORY DATA (FG).** — 85 trees in 26 plots; $F_{max} = 3.8\%$; $dbh_{inv} = 87$ cm.

### [1648] *Pouteria cladantha* Sandwith

*Bull. Misc. Inform. Kew* 1931 (10): 480 [31 Dec. 1931] (Sandwith 1931). — *Neoxythece cladantha* (Sandwith) Aubrév., *Adansonia*, n.s., 1 (2): 183 ["Dec. 1961" publ. Jan.-Mar. 1962] (Aubréville 1962). — *Richardella cladantha* (Sandwith) Baehni, *Boissiera* 11: 97 (Baehni 1965).

*Pouteria loretensis* Baehni, *Candollea* 9: 235 (Baehni 1942).

**VERNACULAR NAMES.** — Ka: kusali yepo • Wp: wai piyū • Cr: zoliv • Br: abiurana-seca, caramuri-preto.

**HERBARIUM DATA (FG).** — 5 collections at CAY. Sel. exs.: *J.-J. de Granville et al.* 8099.

**INVENTORY DATA (FG).** — 36 trees in 11 plots; $F_{max} = 2.2\%$; $dbh_{inv} = 125$ cm.

### [1649] *Pouteria coriacea* (Pierre) Pierre

*Symb. Antill. [Urban]* 5 (1): 109 [20 May 1904] (Pierre 1904). — *Guapeba coriacea* Pierre, *Notes Bot. Sapot.* 2: 42 (Pierre 1891).

*Chrysophyllum minutiflorum* Britton, *Bull. Torrey Bot. Club* 48 (12): 337 ["1921" publ. 1922] (Britton 1922). — *Pouteria minutiflora* (Britton) Sandwith, *Bull. Misc. Inform. Kew* 1931 (10): 478 [31 Dec. 1931] (Sandwith 1931). — *Pseudocladia minutiflora* (Britton) Aubrév., *Adansonia*, n.s., 1 (2): 165 ["Dec. 1961" publ. Jan.-Mar. 1962] (Aubréville 1962).

**VERNACULAR NAMES.** — Pa: uu-kamwi-duwē • Wn: tëpuklem.

**HERBARIUM DATA (FG).** — 61 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 3886.

**INVENTORY DATA (FG).** — 70 trees in 28 plots; $F_{max} = 2.1\%$; $dbh_{inv} = 55.3$ cm.

### [1650] *Pouteria cuspidata* (A.DC.) Baehni subsp. *dura* (Eyma) T.D.Penn.

*Fl. Neotrop. Monogr.* 52: 352 [26 Apr. 1990] (Pennington 1990). — *Pouteria dura* Eyma, *Recueil Trav. Bot. Néerl.* 33: 187 (Eyma 1936). — *Neoxythece dura* (Eyma) Aubrév. & Pellegr., *Adansonia*, n.s., 1 (1): 17 [Nov. 1961] (Aubréville & Pellegrin 1961).

*Sideroxylon cuspidatum* A.DC. var. *crassifolium* Miq., *Fl. Bras. [Martius]* 7: 56 [15 Jan. 1863] (Miquel 1863).

*Sideroxylon durum* Klotzsch ex Miq., *Fl. Bras. [Martius]* 7: 56 [15 Jan. 1863] (Miquel 1863), *nom. nud. pro syn.*

*Neoxythece schulzii* Aubrév., *Adansonia*, n.s., 4 (2): 231 [July 1964] (Aubréville 1964).

*Neoxythece wurdackii* Aubrév., *Adansonia*, n.s., 5 (2): 201 [July 1965] (Aubréville 1965). — *Chrysophyllum wurdackii* (Aubrév.) Bernardi, *Candollea* 22 (2): 231 (Bernardi 1967).

**NOTE.** — *P. cuspidata* is hyperdominant in Amazonia (ter Steege *et al.* 2020).

**VERNACULAR NAMES.** — Nt: nyanboka • Br: abiurana-arana.

**HERBARIUM DATA (FG).** — 7 collections at CAY. Sel. exs.: *D. Sabatier* 1430.

**INVENTORY DATA (FG).** — 22 trees in 15 plots; $F_{max} < 1\%$; $dbh_{inv} = 43$ cm.
<figure><img src="image_83.png" /></figure>

<figure><img src="image_84.png" /></figure>

FIG. 54. — Sapotaceae: **A**, *Chrysophyllum prieurii* A.DC. (D. Sabatier et al. 4620); **B**, *Manilkara bidentata* (A.DC.) A.Chev. subsp. *bidentata* (D. Sabatier 3553); **C**, *Micropholis acutangula* (D. Sabatier & S. Gonzalez 5371); **D**, *Micropholis obscura* T.D.Penn. (D. Sabatier 1221). A-C, © D. Sabatier/IRD; D, © M.-F. Prévost/IRD.

[1651] *Pouteria cuspidata* subsp. robusta (Mart. & Eichler) T.D.Penn.

Fl. Neotrop. Monogr. 52: 351 [26 Apr. 1990] (Pennington 1990). — *Sideroxylon robustum* Mart. & Eichler, Fl. Bras. [Martius] 7: 56 [15 Jan. 1863] (Martius & Eichler 1863). — *Oxythece robusta* (Mart. & Eichler) Pierre, Symb. Antill. [Urban] 5 (1): 161 [20 May 1904] (Pierre 1904), “robustum”. — *Pouteria robusta* (Mart. & Eichler) Eyma, Recueil Trav. Bot. Néerl. 33: 188 (Eyma 1936). — *Neoxythece robusta* (Mart. & Eichler) Aubrév. & Pellegr., Adansonia, sér. 2, 1 (1): 17 [Nov. 1961] (Aubréville & Pellegrin 1961).

*Pouteria robusta* var. longifolia Eyma, Recueil Trav. Bot. Néerl. 33: 188 (Eyma 1936).

*Pouteria robusta* var. typica Baehni, Candollea 9: 213 (Baehni 1942), nom. inval. (Turland et al. 2018: Art. 24.3).

*Oxythece robusta* var. brevipetiolata Monach., Fieldiana, Bot. 28 (3): 486 (Monachino 1953).

NOTES. — *P. cuspidata*. is hyperdominant in Amazonia (ter Steege *et al.* 2020). Although Miquel provided the treatment of Sapotaceae for the *Flora Brasiliensis* (Miquel 1863), Martius clearly claims, in the footnote at the beginning of the treatment of this family (Miquel 1863: 37), his coauthorship with Eichler of some new species, among which is *Sideroxylon robustum* Mart. & Eichler.

VERNACULAR NAMES. — Ka: atakamala, kusili palatali.

<figure><img src="image_85.png" /></figure>

<figure><img src="image_86.png" /></figure>

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: *D. Sabatier & S. Gonzalez 5385*.

INVENTORY DATA (FG). — 4 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 29$ cm.

[1652] *Pouteria decorticans* T.D.Penn.

Fl. Neotrop. Monogr. 52: 489 [26 Apr. 1990] (Pennington 1990).

VERNACULAR NAMES. — Pa: kuyau-kamwi-duwē, kuyau-kamwi-seinō, uu-kamwi-wahuyo • Wp: yawayi lū • Nt: nyanboka • Cr: zoliv • Br: abiu-giboia.

HERBARIUM DATA (FG). — 33 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 2617*.

INVENTORY DATA (FG). — 147 trees in 38 plots; $F_{max} = 3.3\%$; $dbh_{inv} = 41.1$ cm.

[1653] *Pouteria deliciosa* T.D.Penn.

Fl. Neotrop. Monogr. 52: 446 [26 Apr. 1990] (Pennington 1990).

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: *D. Sabatier 1213*.

INVENTORY DATA (FG). — 24 trees in 15 plots; $F_{max} < 1\%$; $dbh_{inv} = 75$ cm.
[1654] *Pouteria durlandii* (Standl.) Baehni

*Candollea* 9: 422 (Baehni 1942). — *Lucuma durlandii* Standl., *Trop. Woods* 4: 5 (Standley 1925). — *Paralabatia durlandii* (Standl.) Aubrév., *Adansonia*, n.s., 3 (1): 21 [Jan.-Mar. 1963] (Aubréville 1963). — *Peteni-odendron durlandii* (Standl.) Lundell, *Wrightia* 5 (7): 255 (Lundell 1976).

*Pouteria suffulta* Baehni, *Candollea* 14: 70 (Baehni 1952).

*Pouteria potosina* Lundell, *Wrightia* 5 (4): 101 (Lundell 1975). — *Peteniodendron potosinum* (Lundell) Lundell, *Wrightia* 5 (7): 255 (Lundell 1976). — *Paralabatia potosina* (Lundell) Aubrév., *Wrightia* 5 (9): 350 (Aubréville 1977).

*Peteniodendron belizense* Lundell, *Wrightia* 5 (7): 254 (Lundell 1976). — *Paralabatia belizensis* (Lundell) Lundell, *Wrightia* 5 (9): 350 (Lundell 1977).

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: *T.D. Pennington & S.A. Mori* 12177.

INVENTORY DATA (FG). — 14 trees in 9 plots; $F_{max} < 1\%$; $dbh_{inv} = 19.4$ cm.

[1655] *Pouteria egregia* Sandwith

*Bull. Misc. Inform. Kew* 1931 (10): 479 [31 Dec. 1931] (Sandwith 1931). — *Sandwithiodoxa egregia* (Sandwith) Aubrév. & Pellegr., *Adansonia*, n.s., 1 (2): 163 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville & Pellegrin 1962).

VERNACULAR NAMES. — Ka: olomelan • Wp: wila náná, wila náná sili • Br: abiu-pitomba, abiurana, cururu.

HERBARIUM DATA (FG). — 23 collections at CAY. Sel. exs.: *D. Sabatier* 1219.

INVENTORY DATA (FG). — 74 trees in 38 plots; $F_{max} = 1.6\%$; $dbh_{inv} = 70$ cm.

[1656] *Pouteria engleri* Eyma

*Recueil Trav. Bot. Néerl.* 33: 178 (Eyma 1936). — *Nemaluma engleri* (Eyma) Aubrév. & Pellegr., *Adansonia*, n.s., 1 (1): 31 [Nov. 1961] (Aubréville & Pellegrin 1961).

*Chrysophyllum alnifolium* Engl., *Bot. Jahrb. Syst.* 12: 522 [23 Dec. 1890] (Engler 1890), *nom. illeg. hom., non* Baker (1877).

VERNACULAR NAMES. — Ka: atakamala, kusili palatali • Nt: nyanboka.

HERBARIUM DATA (FG). — 20 collections at CAY. Sel. exs.: *E.M. Mé-linon s.n.* (original material P[P00640595, P00640596, P00640597]).

INVENTORY DATA (FG). — 71 trees in 44 plots; $F_{max} = 1.2\%$; $dbh_{inv} = 72.9$ cm.

[1657] *Pouteria ephedrantha* (A.C.Sm.) T.D.Penn.

*Fl. Neotrop. Monogr.* 52: 508 [26 Apr. 1990] (Pennington 1990). — *Lucuma ephedrantha* A.C.Sm., *Brittonia* 2 (2): 158 (Smith 1936).

HERBARIUM DATA (FG). — A single collection, *D. Sabatier & J.-F. Molino* 5717.

INVENTORY DATA (FG). — 46 trees in 11 plots; $dbh_{inv} = 115$ cm.

[1658] *Pouteria eugeniifolia* (Pierre) Baehni
(Fig. 55B)

*Candollea* 9: 218 (Baehni 1942). — *Micropholis eugeniifolia* Pierre, *Notes Bot. Sapot.* 2: 40 (Pierre 1891), “eugeniaefolia”; — *Sideroxylon eugeniifolium* (Pierre) Engl., *Nat. Pflanzenfam. [Engler & Prantl]* *Nachr.* 1: 276 (Engler 1897). — *Myrtiluma eugeniifolia* (Pierre) Aubrév., *Adansonia*, n.s., 1 (2): 180 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville 1962), “eugeniaefolia”.

*Micropholis steyermarkii* Monach., *Fieldiana*, Bot. 28 (3): 481 (Monachino 1953), *nom. inval.* (simultaneously published with *Sideroxylon steyermarkii*).

*Pouteria steyermarkii* Monach., *Fieldiana*, Bot. 28 (3): 481 (Monachino 1953), *nom. inval.* (simultaneously published with *Micropholis steyermarkii*).

VERNACULAR NAMES. — Ka: mandasabay • Nt: nyanboka • Br: batinga.

HERBARIUM DATA (FG). — 26 collections at CAY. Sel. exs.: *E.M. Mé-linon s.n.* (holo-, P[P00647901]; iso-, G[G00439472], P[P00640598, P00640599, P00647902]).

INVENTORY DATA (FG). — 177 trees in 78 plots; $F_{max} = 1.2\%$; $dbh_{inv} = 98$ cm.

[1659] *Pouteria filipes* Eyma

*Recueil Trav. Bot. Néerl.* 33: 180 (Eyma 1936). — *Pseudolabatia filipes* (Eyma) Aubrév., *Adansonia*, n.s., 3 (1): 21 [Jan.-Mar. 1963] (Aubréville 1963).

VERNACULAR NAMES. — Pa: kuyau-kamwi-puvemna.

HERBARIUM DATA (FG). — 23 collections at CAY. Sel. exs.: *D. Sabatier* 1516.

INVENTORY DATA (FG). — 89 trees in 31 plots; $F_{max} = 3.5\%$; $dbh_{inv} = 87$ cm.

[1660] *Pouteria fimbriata* Baehni

*Candollea* 14: 68 (Baehni 1952).

*Pseudolabatia raoulantonia* Aubrév. & Pellegr., *Adansonia*, n.s., 1 (2): 168 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville & Pellegrin 1962).

HERBARIUM DATA (FG). — 14 collections at CAY. Sel. exs.: *D. Sabatier & B. Riéra* 1741.

INVENTORY DATA (FG). — 123 trees in 81 plots; $F_{max} < 1\%$; $dbh_{inv} = 63$ cm.

[1661] *Pouteria flavilatex* T.D.Penn.

*Rodriguésia* 57 (2): 308 (Pennington 2006).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino* 5128.

INVENTORY DATA (FG). — 41 trees in 33 plots; $F_{max} < 1\%$; $dbh_{inv} = 58.1$ cm.
[1662] *Pouteria franciscana* Baehni

Candollea 9: 262 (Baehni 1942).

VERNACULAR NAMES. — Wp: waa pitā, wapitā • Wn: palakwa • Nt: nyanboka • Cr: zoliv • Br: abiurana-branca.

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 1637*.

INVENTORY DATA (FG). — 24 trees in 7 plots; Fmax = 1.9%; dbhinv = 104 cm.

[1663] *Pouteria glomerata* (Miq.) Radlk.

Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 12: 333 (Radlkofer 1882). — *Lucuma glomerata* Miq., Fl. Bras. [Martius] 7: 81 [15 Jan. 1863] (Miquel 1863). — *Labatia glomerata* (Miq.) Radlk., Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 14: 451 (Radlkofer 1884). — *Guapeba glomerata* (Miq.) Pierre, *Notes Bot. Sapot.* 2: 43 (Pierre 1891). — *Richardella glomerata* (Miq.) Baehni, *Boissiera* 11: 97 (Baehni 1965). — *Neolabatia glomerata* (Miq.) Aubrév., *Mem. New York Bot. Gard.* 23: 203 (Aubréville 1972).

Labatia parinarioides Radlk., Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München 14: 451 (Radlkofer 1884). — *Guapeba parinarioides* (Radlk.) Pierre, *Notes Bot. Sapot.* 2: 43 (Pierre 1891).

Pouteria weddelliana Pierre, *Notes Bot. Sapot.* 2: 45 (Pierre 1891).

Pouteria glomerata var. glabrescens Huber, *Bull. Soc. Bot. Genève*, sér. 2, 6: 197 ["1914" publ. 1915] (Huber 1915), "Ponteria".

Labatia parviflora Pittier, *Arb. Arborn. Venez.* 2-3: 41 (Pittier 1923). — *Neolabatia parviflora* (Pittier) Aubrév., *Mem. New York Bot. Gard.* 23: 203 (Aubréville 1972).

Lucuma hypoglauca Standl., *Lista Pl. Salvador* [Standley & S. Calde-ron]: 170 (Standley 1925). — *Pouteria hypoglauca* (Standl.) Baehni, *Candollea* 9: 250 (Baehni 1942). — *Richardella hypoglauca* (Standl.) Baehni, *Boissiera* 11: 97 (Baehni 1965). — *Neolabatia hypoglauca* (Standl.) Aubrév., *Mem. New York Bot. Gard.* 23: 203 (Aubréville 1972).

Pouteria glomerata var. typica Baehni, *Candollea* 9: 243 (Baehni 1942), *nom. inval.* (Turland *et al.* 2018: Art. 24.3).

Pouteria pittieri Baehni, *Candollea* 9: 254 (Baehni 1942).

HERBARIUM DATA (FG). — 5 collections at CAY. Sel. exs.: *D. Sabatier 1488*.

INVENTORY DATA (FG). — 7 trees in 3 plots; Fmax = 1.1%; dbhinv = 43 cm.

[1664] *Pouteria gonggrijpii* Eyma

Recueil Trav. Bot. Néerl. 33: 185 (Eyma 1936), "Gongrijpii". — *Franchetella gonggrijpii* (Eyma) Aubrév., *Adansonia*, n.s., 1 (2): 183 ["Dec. 1961" publ. Jan.-Mar. 1962] (Aubréville 1962), "Gongrijpii".

NOTE. — This species is dedicated to the Dutch collector Justus Willem Gonggrijp. Eyma himself wrote "*Pouteria gonggrijpii*" on the type. The spelling "*gongrijpii*" should therefore be taken as an

unintentional typographical error, and corrected (Turland *et al.* 2018: Art. 60.1).

VERNACULAR NAMES. — Pa: kuyau-kamwi, tukuyuy-kamwi, uu-kamwi, uu-kamwi-duwē • Ka: ololome, olome • Te: wila pitāg tsing • Nt: nyanboka • Cr: balata-pwé • Fr: balata poirier • Br: abiurana-vermelha.

HERBARIUM DATA (FG). — 100 collections at CAY. Sel. exs.: *M.-F. Prévost & P. Grenand 4304*.

INVENTORY DATA (FG). — 611 trees in 126 plots; Fmax = 8.6%; dbhinv = 51 cm.

[1665] *Pouteria grandis* Eyma

Recueil Trav. Bot. Néerl. 33: 170 (Eyma 1936). — *Radlkoferella grandis* (Eyma) Aubrév., *Adansonia*, n.s., 1 (2): 183 ["Dec. 1961" publ. Jan.-Mar. 1962] (Aubréville 1962).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Pa: uu-aška.

HERBARIUM DATA (FG). — 26 collections at CAY. Sel. exs.: *D. Sabatier 2450*.

INVENTORY DATA (FG). — 101 trees in 41 plots; Fmax = 2%; dbhinv = 70.7 cm.

[1666] *Pouteria guianensis* Aubl.

Hist. Pl. Guiane 1: 85 [Jun.-Dec. 1775] (Aublet 1775). — *Labatia pedunculata* Willd., *Sp. Pl.*, ed. 4 1 (2): 624 [July 1798] (Willdenow 1798), *nom. illeg. superfl.* (based on *Pouteria guianensis*). — *Pouteria pedunculata* Poir., *Encycl. [J. Lamarck et al.]* Suppl. 4: 546 [29 June 1816] (Poiret 1816).

Lucuma psammophila (Mart.) A.DC. var. macrophylla Raunk., *Vidensk. Meddel. Dansk Naturhist. Foren. Kjøbenhavn* 1889: 6 (Raunkiaer 1889).

Labatia tovarensis Engl., *Bot. Jahrb. Syst.* 12: 515 [23 Dec. 1890] (Engler 1890). — *Pouteria tovarensis* (Engl.) Engl., *Nat. Pflanzenfam. [Engler & Prantl]* 4 (1): 142 (Engler 1891).

Guapeba glazioviana Pierre, *Notes Bot. Sapot.* 2: 43 (Pierre 1891), "Glazioveana", *nom. nud.* — *Pouteria glazioviana* Dubard, *Notul. Syst. (Paris)* 1: 380 (Dubard 1911), "Glazioveana", *nom. nov.*, based on *Lucuma psammophila* var. macrophylla.

Krugella hartii Pierre, *Notes Bot. Sapot.* 2: 52 (Pierre 1891). — *Pouteria hartii* (Pierre) Dubard, *Ann. Mus. Colon. Marseille*, sér. 2, 10: 34 (Dubard 1912).

Lucuma hartii Hemsl., *Hooker's Icon. Pl.* 26 [ser. 4, 6]: t. 2565 ["1899" publ. Apr. 1898] (Hemsley 1898).

Lucuma glazioviana Pierre ex Glaz., *Bull. Soc. Bot. France* 57 (Mem. 3e): 438 (Glaziou 1910), *nom. nud*.

Pouteria obidensis Huber, *Bull. Soc. Bot. Genève*, sér. 2, 6: 197 ["1914" publ. 1915] (Huber 1915).

Pouteria demerarae Sandwith, *Bull. Misc. Inform. Kew* 1931 (10): 477 [31 Dec. 1931] (Sandwith 1931).
*Lucuma huallagae* Standl. ex Ll.Williams, *Publ. Field Mus. Nat. Hist., Bot. Ser.* 15: 411 (Williams 1936).

*Pouteria caimito* var. *strigosa* Monach., *Fieldiana*, Bot. 28 (3): 487 (Monachino 1953).

*Pouteria cuatrecasasii* Aubrév., *Adansonia*, n.s., 7 (2): 141 [Jul.-Sep. 1967] (Aubréville 1967).

**NOTES.** — Hyperdominant in Amazonia (ter Steege *et al.* 2020). *Pouteria hartii* (Pierre) Dubard is based on both *Krugella hartii* Pierre and *Lucuma hartii* Hemsl. These names have distinct types, which are both cited by Dubard (1912: 35).

**VERNACULAR NAMES.** — Pa: balata-kamwi, uu-ahavukune, uu-kamwi-seine • Ka: asepoko, asepuku, kulumoto, wasepuku • Te: wila pitāg piñāg • Wp: wau, wawiyu • Wn: alimihmo • Nt: nyanboka • Cr: grënn-kwata, jonndëf • Fr: jaune d'œuf • Br: abiurana-abiu, abiurana-branca, abiurana-casca-fina.

**HERBARIUM DATA (FG).** — 79 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JJR[P00777520] designated by Lanjouw & Uittien [1940: 156]).

**INVENTORY DATA (FG).** — 411 trees in 147 plots; $F_{max} = 2.6\%$; $dbh_{inv} = 70$ cm.

[1667] *Pouteria hispida* Eyma

*Recueil Trav. Bot. Néerl.* 33: 177 (Eyma 1936).

*Pouteria trichopoda* Baehni, *Candollea* 9: 245 (Baehni 1942).

*Pouteria solimoesensis* Aubrév. & Pellegr., *Adansonia*, n.s., 1 (2): 157 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville & Pellegrin 1962).

*Pouteria benoistii* Aubrév., *Adansonia*, n.s., 4 (2): 228 [July 1964] (Aubréville 1964).

**VERNACULAR NAMES.** — Pa: uu-kamwi • Ka: asepuku, tuwonule, wasepuku • Wp: wawiyu • Nt: nyanboka • Br: abiurana-braba, abiurana-jarani.

**HERBARIUM DATA (FG).** — 45 collections at CAY. Sel. exs.: *E.M. Mélinon 108*, 1863 (holotype of *Pouteria benoistii*: P[P00647949]; iso-, P[P00647950, P00647951]).

**INVENTORY DATA (FG).** — 100 trees in 62 plots; $F_{max} = 1\%$; $dbh_{inv} = 110$ cm.

[1668] *Pouteria jariensis* Pires & T.D.Penn. (Fig. 55D)

*Fl. Neotrop. Monogr.* 52: 331 [26 Apr. 1990] (Pires & Pennington 1990).

**VERNACULAR NAMES.** — Pa: tukuyuy-kamwi • Cr: zoliv • Br: abiurana.

**HERBARIUM DATA (FG).** — 25 collections at CAY. Sel. exs.: *J.-F. Molino 1198*.

**INVENTORY DATA (FG).** — 95 trees in 64 plots; $F_{max} < 1\%$; $dbh_{inv} = 67$ cm.

[1669] *Pouteria laevigata* (Mart.) Radlk.

*Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 14: 457 (Radlkofer 1884). — *Labatia laevigata* Mart., *Flora* 21 (2, Beibl.): 92 (Martius 1838). — *Lucuma laevigata* (Mart.) A.DC., *Prodr. [A. P. de Candolle]* 8: 167 [mid Mar. 1844] (Candolle 1844). — *Guapeba laevigata* (Mart.) Pierre, *Notes Bot. Sapot.* 2: 42 (Pierre 1891).

*Pouteria bouffardiana* Bernardi, *Candollea* 22 (2): 228 (Bernardi 1967).

**VERNACULAR NAMES.** — Wp: wau âtâ • Br: abiurana-de-casca-grossa, abiurana-sapota.

**HERBARIUM DATA (FG).** — 12 collections at CAY. Sel. exs.: *D. Sabatier 1098*.

**INVENTORY DATA (FG).** — 18 trees in 15 plots; $F_{max} < 1\%$; $dbh_{inv} = 85.1$ cm.

[1670] *Pouteria macrocarpa* (Mart.) D.Dietr.

*Syn. Pl. [D. Dietrich]* 1: 431 [July 1839] (Dietrich 1839). — *Labatia macrocarpa* Mart., *Nova genera et species plantarum* [Martius] 2 (2): 71 [Jan.-June 1827] (Martius 1827). — *Richardella macrocarpa* (Mart.) Baehni, *Boissiera* 11: 97 (Baehni 1965). — *Neolabatia macrocarpa* (Mart.) Aubrév., *Mem. New York Bot. Gard.* 23: 203 (Aubréville 1972).

*Lucuma paraensis* Standl., *Trop. Woods* 33: 13 (Standley 1933). — *Pouteria paraensis* (Standl.) Baehni, *Candollea* 9: 424 (Baehni 1942).

**NOTE.** — Semi-domesticated by pre-Columbian Amerindians (Clement 1999; Levis *et al.* 2017).

**VERNACULAR NAMES.** — Pa: uu-kamwi-duwē, uu-purubumna • Wp: pakuku, yā laānga • Br: abiu-grande, cutite-grande.

**HERBARIUM DATA (FG).** — 4 collections at CAY. Sel. exs.: *P. Grenand 699*.

**INVENTORY DATA (FG).** — 2 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 30.2$ cm.

[1671] *Pouteria macrophylla* (Lam.) Eyma

*Recueil Trav. Bot. Néerl.* 33: 164 (Eyma 1936). — *Chrysophyllum macrophyllum* Lam., *Tabl. Encycl.* 2[5 (2)]: 44 [1 Sep. 1794] (Lamarck 1794). — *Richardella macrophylla* (Lam.) Aubrév., *Adansonia*, n.s., 1 (2): 175 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville 1962).

*Bumelia nervosa* Vahl, *Eclog. Amer.* 1: 28 (Vahl 1797).

*Lucuma rivicoa* C.F.Gaertn., *Suppl. Carp.* 130 (Gaertner 1807). — *Vitellaria rivicoa* (C.F.Gaertn.) Radlk., *Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 12: 326 (Radlkofer 1882). — *Richardella rivicoa* (C.F.Gaertn.) Pierre, *Notes Bot. Sapot.* 1: 19 (Pierre 1890).

*Achras glaucescens* Willd. ex Steud., *Nomencl. Bot. [Steudel]*, ed. 2, 1: 15 (Steudel 1840), *nom. nud*.

*Vitellaria glaucophylla* Engl., *Bot. Jahrb. Syst.* 12: 513 [23 Dec. 1890] (Engler 1890). — *Lucuma rivicoa* var. *glaucophylla* (Engl.) Dubard, *Ann. Mus. Colon. Marseille*, sér. 2, 10: 15 (Dubard 1912).

*Lucuma acreana* K.Krause, *Notizbl. Königl. Bot. Gart. Berlin* 6: 169 [4 Apr. 1914] (Krause 1914).
NOTES. — Hyperdominant in Amazonia (ter Steege *et al.* 2020), but not yet found in inventories of trees above 10 cm dbh in French Guiana. Semi-domesticated by pre-Columbian Amerindians (Clement 1999; Levis *et al.* 2017).

VERNACULAR NAMES. — Pa: uu • Wp: akusi yeti • Nt: sooke • Cr: jonndèf • Br: abiurana-cutiti, acará-uba, cutite, cutitiribá.

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: *J.P.B. von Rohr* 125 (original material of *Bumelia nervosa*: C[C10018776, C10018777]).

SIZE. — Brazil, Amazonas. *G.T. Prance et al.* 22998 (MO), 15 m × 15 cm.

[1672] *Pouteria maxima* T.D.Penn.

Rodriguésia 57 (2): 319 (Pennington 2006).

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *T.D. Pennington & S.A. Mori* 12137.

INVENTORY DATA (FG). — 5 trees in 4 plots; $F_{max} < 1\%$; $dbh_{inv} = 110$ cm.

[1673] *Pouteria melanopoda* Eyma

Recueil Trav. Bot. Néerl. 33: 174 (Eyma 1936).

VERNACULAR NAMES. — Ka: asepokolan • Br: abiurana-goiabinha.

HERBARIUM DATA (FG). — 29 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 1843.

INVENTORY DATA (FG). — 61 trees in 49 plots; $F_{max} < 1\%$; $dbh_{inv} = 61.3$ cm.

[1674] *Pouteria oblanceolata* Pires

Bol. Tècn. Inst. Agron. N. 38: 38 (Pires 1960).

VERNACULAR NAMES. — Nt: nyanboka • Br: abiurana.

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: *D. Sabatier* 1106.

INVENTORY DATA (FG). — 4 trees in 3 plots; $F_{max} < 1\%$; $dbh_{inv} = 47$ cm.

[1675] *Pouteria platyphylla* (A.C.Sm.) Baehni

Candollea 9: 274 (Baehni 1942). — *Lucuma platyphylla* A.C.Sm., *Bull. Torrey Bot. Club* 60 (6): 388 [1 June 1933] (Smith 1933). — *Franchetella platyphylla* (A.C.Sm.) Aubrév., *Adansonia*, n.s., 1 (2): 183 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville 1962).

VERNACULAR NAMES. — Pa: uu-kamwi • Br: abiurana-vermelha.

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *D. Sabatier* 5843.

INVENTORY DATA (FG). — 11 trees in 11 plots; $F_{max} < 1\%$; $dbh_{inv} = 40$ cm.

[1676] *Pouteria procera* (Mart.) K.Hammer

Verz. Landwirtsch. Gartn. KulturpfL., ed. 2, 2: 1046 (Hammer 1986). — *Lucuma procera* Mart., *Flora* 22 (1, Beibl.): 57 (Martius 1839). — *Vitellaria procera* (Mart.) Radlk., *Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 12: 325 (Radlkofer 1882). — *Urbanella procera* (Mart.) Pierre, *Notes Bot. Sapot.* 1: 25 (Pierre 1890). — *Calocarpum procerum* (Mart.) Dubard, *Ann. Mus. Colon. Marseille*, sér. 2, 10: 8 (Dubard 1912).

Lucuma procera var. cuspidata Miq., *Fl. Bras. [Martius]* 7: 72 [15 Jan. 1863] (Miquel 1863). — *Urbanella cuspidata* (Miq.) Pierre, *Notes Bot. Sapot.* 1: 25 (Pierre 1890). — *Calocarpum procerum* var. cuspidatum (Miq.) Dubard, *Ann. Mus. Colon. Marseille*, sér. 2, 10: 8 (Dubard 1912).

*Urbanella buchananiifolia* Pierre, *Notes Bot. Sapot.* 1: 25 (Pierre 1890). — *Lucuma buchananiifolia* Pierre, *Notes Bot. Sapot.* 1: 25 (Pierre 1890), “*buchananiaefolia*”, nom. nud. pro syn. — *Calocarpum buchananiifolium* (Pierre) Dubard, *Ann. Mus. Colon. Marseille*, sér. 2, 10: 9 (Dubard 1912), “*buchananiaefolium*”. — *Achras buchananiifolia* (Pierre) Bernardi, *Candollea* 22 (2): 230 (Bernardi 1967).

*Urbanella oblonga* Pierre, *Notes Bot. Sapot.* 1: 25 (Pierre 1890). — *Lucuma oblonga* Pierre, *Notes Bot. Sapot.* 1: 25 (Pierre 1890), nom. nud. pro syn. — *Calocarpum procerum* var. *oblongum* (Pierre) Dubard, *Ann. Mus. Colon. Marseille*, sér. 2, 10: 9 (Dubard 1912), “*oblongifolium*”.

*Lucuma excelsa* A.C.Sm., *Brittonia* 2 (2): 159 (Smith 1936). — *Pouteria excelsa* (A.C.Sm.) Baehni, *Candollea* 9: 350 (Baehni 1942). — *Urbanella excelsa* (A.C.Sm.) Aubrév., *Adansonia*, n.s., 3 (1): 21 [Jan.-Mar. 1963] (Aubréville 1963).

VERNACULAR NAMES. — Br: maparajúba-vermelha.

HERBARIUM DATA (FG). — A single collection, *J.-J. de Granville et al.* 7632 (P).

SIZE. — Up to 100 cm dbh (Pennington 1990).

[1677] *Pouteria putamen-ovi* T.D.Penn.

*Fl. Neotrop. Monogr.* 52: 270 [26 Apr. 1990] (Pennington 1990).

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 3407.

INVENTORY DATA (FG). — 1 tree, dbh = 44.4 cm.

[1678] *Pouteria reticulata* (Engl.) Eyma

Recueil Trav. Bot. Néerl. 33: 183 (Eyma 1936). — *Chrysophyllum reticulatum* Engl., *Bot. Jahrb. Syst.* 12: 522 [23 Dec. 1890] (Engler 1890). — *Franchetella reticulata* (Engl.) Aubrév., *Adansonia*, n.s., 1 (2): 183 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville 1962).

*Lucuma minutiflora* Allemão, Trab. Comm. Sci. Expl., Bot., Rio de Janeiro 1: 57 (Allemão 1866).

*Sideroxylon uniloculare* Donn.Sm., *Bot. Gaz.* 35 (1): 5 [20 Jan. 1903] (Donnell Smith 1903). — *Pouteria unilocularis* (Donn.Sm.) Baehni, *Candollea* 9: 273 (Baehni 1942). — *Franchetella unilocularis* (Donn.Sm.) Aubrév., *Adansonia*, n.s., 3 (1): 21 [Jan.-Mar. 1963] (Aubréville 1963).
*Sideroxylon hondurense* Pittier, *Contr. U.S. Natl. Herb.* 20 (12): 483 [9 Jan. 1922] (Pittier 1922). — *Pouteria hondurensis* (Pittier) Baehni, *Candollea* 9: 228 (Baehni 1942).

*Sideroxylon meyeri* Standl., *Trop. Woods* 31: 45 (Standley 1932). — *Pouteria meyeri* (Standl.) Baehni, *Candollea* 9: 272 (Baehni 1942).

*Lucuma anibifolia* A.C.Sm., *Bull. Torrey Bot. Club* 60 (6): 389 [1 June 1933] (Smith 1933), "anibaefolia". — *Pouteria anibifolia* (A.C.Sm.) Baehni, *Candollea* 9: 269 (Baehni 1942). — *Franchetella anibifolia* (A.C.Sm.) Aubrév., *Adansonia*, n.s., 1 (2): 183 ["Dec. 1961" publ. Jan.-Mar. 1962] (Aubréville 1962).

*Pouteria cearensis* Baehni, *Candollea* 9: 279 (Baehni 1942).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: kuyau-kamwi-duwě • Ka: olomelan • Br: abiurana-cascuda.

HERBARIUM DATA (FG). — 21 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino* 5322bis.

INVENTORY DATA (FG). — 86 trees in 27 plots; $F_{max}$ = 3.7 %; $dbh_{inv}$ = 53.8 cm.

[1679] *Pouteria retinervis* T.D.Penn.

Fl. Neotrop. Monogr. 52: 308 [26 Apr. 1990] (Pennington 1990).

VERNACULAR NAMES. — Br: abiurana-grande.

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: *S.A. Mori & B.M. Boom* 15239 (holo-, NY[00335312]; iso-, CAY[CAY044794], K, NO[NO0109877]).

INVENTORY DATA (FG). — 9 trees in 7 plots; $F_{max}$ < 1 %; $dbh_{inv}$ = 51.9 cm.

[1680] *Pouteria rodriguesiana* Pires & T.D.Penn.

Fl. Neotrop. Monogr. 52: 379 [26 Apr. 1990] (Pires & Pennington 1990).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Pa: uu-kamwi, uu-kamwi-seine • Te: wila pitāg hun • Nt: nyanboka • Br: abiu-maparajúba.

HERBARIUM DATA (FG). — 17 collections at CAY. Sel. exs.: *B. Maguire et al.* 47097 (holo-, K[K000994418, K000994419]; iso-, NY[01212902], US[00516534]).

INVENTORY DATA (FG). — 70 trees in 29 plots; $F_{max}$ = 2.4 %; $dbh_{inv}$ = 80 cm.

[1681] *Pouteria sagotiana* (Baill.) Eyma
(Fig. 55C)

Recueil Trav. Bot. Néerl. 33: 170 (Eyma 1936). — *Eremoluma sagotiana* Baill., *Hist. Pl. [Baillon]* 11: 292 [Sep.-Oct. 1891] (Baillon 1891). — *Lucuma sagotiana* (Baill.) Engl., *Nat. Pflanzenfam. [Engler & Prantl] Nachtr.* 1: 274 (Engler 1897).

VERNACULAR NAMES. — Pa: inam-etni-seinó, waaduk • Ka: asinadan.

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *P.A. Sagot* 1189, 1858 (holo-, P[P00648047]).

INVENTORY DATA (FG). — 26 trees in 16 plots; $F_{max}$ = 2 %; $dbh_{inv}$ = 30.9 cm.

[1682] *Pouteria singularis* T.D.Penn.

Fl. Neotrop. Monogr. 52: 459 [26 Apr. 1990] (Pennington 1990).

VERNACULAR NAMES. — Pa: kuyau-kamwi-duwō, uu-kamwi • Nt: nyanboka • Cr: jonndèf, zoliv • Fr: jaune d'œuf.

HERBARIUM DATA (FG). — 17 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 2645.

INVENTORY DATA (FG). — 149 trees in 62 plots; $F_{max}$ = 2.8 %; $dbh_{inv}$ = 60 cm.

[1683] *Pouteria speciosa* (Ducke) Baehni

*Candollea* 9: 398 (Baehni 1942). — *Lucuma speciosa* Ducke, *Arch. Mus. Nac. Rio de Janeiro* 22: 67 (Ducke 1919). — *Englerella speciosa* (Ducke) Ducke, *Arch. Jard. Bot. Rio de Janeiro* 3: 230 (Ducke 1922), *nom. inval.* (new combination not formally accepted by the author). — *Richardella speciosa* (Ducke) Aubrév., *Adansonia*, n.s., 1 (2): 175 ["Dec. 1961" publ. Jan.-Mar. 1962] (Aubréville 1962).

*Englerella macrocarpa* Pierre, *Notes Bot. Sapot.* 2: 47 (Pierre 1891).

VERNACULAR NAMES. — Wn: kanapupali • Nt: taka, tyaka • Br: pajurá, pajurá-de-obidos.

HERBARIUM DATA (FG). — 14 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino* 5117.

INVENTORY DATA (FG). — 30 trees in 9 plots; $F_{max}$ < 1 %; $dbh_{inv}$ = 88.4 cm.

[1684] *Pouteria stipulifera* T.D.Penn.

*Rodriguésia* 57 (2): 283 (Pennington 2006).

HERBARIUM DATA (FG). — A single collection, *J. Engel & P. Petronelli* 19 (CAY).

INVENTORY DATA (FG). — 1 tree, $dbh$ = 29.3 cm.

[1685] *Pouteria tenuisepala* Pires & T.D.Penn.

Fl. Neotrop. Monogr. 52: 435 [26 Apr. 1990] (Pires & Pennington 1990).

VERNACULAR NAMES. — Wp: ei li.

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: *W.A. Egler* 47673 (holo-, MG, not seen; iso-, K[K000641152]).

INVENTORY DATA (FG). — 24 trees in 7 plots; $F_{max}$ = 1.4 %; $dbh_{inv}$ = 38 cm.
[1686] *Pouteria torta* (Mart.) Radlk.
subsp. *glabra* T.D.Penn.

Fl. Neotrop. Monogr. 52: 484 [26 Apr. 1990] (Pennington 1990).

*Lucuma jenmanii* Pittier, *Contr. U.S. Natl. Herb.* 13: 458 (Pittier 1912). — *Pouteria jenmanii* (Pittier) Sandwith, *Bull. Misc. Inform. Kew* 1931 (10): 478 [31 Dec. 1931] (Sandwith 1931).

*Lucuma gutta* Ducke, *Ann. Acad. Brasil Sci.* 6: 208 (Ducke 1934). — *Pouteria gutta* Ducke, *Ann. Acad. Brasil Sci.* 6: 208; (Ducke 1934) in nota.

*Pouteria gutta* Baehni, *Candollea* 9: 261 (Baehni 1942), isonym.

*Lucuma dolichophylla* Standl. ex Ll.Williams, *Bot. Mus. Leafl.* 13 (9): 289 [18 Apr. 1949] (Williams 1949).

**NOTES.** — Schultes (1949: 289) stated that he only translated into latin Ll. Williams' English description of *Lucuma dolichophylla*, and he unequivocally ascribed authorship to Ll. Williams. *Lucuma gutta* Ducke and *Pouteria gutta* Ducke are alternative names (published simultaneously for the same taxon), but both are nevertheless valid, because they were published before 1953 (Turland *et al*. 2018: Art. 36.3). Therefore, the transfer of *Lucuma gutta* to *Pouteria* by Baehni is superfluous.

**VERNACULAR NAMES.** — Pa: uu-kamwi, uu-kamwi-wahuyo • Wp: wamisalu, wau • Nt: nyanboka • Cr: zoliv • Br: abiurana-camazal, curiola, grão-de-galo, guapeva.

**HERBARIUM DATA (FG).** — 35 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 1351*.

**INVENTORY DATA (FG).** — 203 trees in 92 plots; $F_{max}$ = 1.6 %; $dbh_{inv}$ = 55.4 cm.

[1687] *Pouteria torta* subsp. *tuberculata*
(Sleumer) T.D.Penn.

Fl. Neotrop. Monogr. 52: 483 [26 Apr. 1990] (Pennington 1990). — *Lucuma tuberculata* Sleumer, *Repert. Spec. Nov. Regni Veg.* 45: 18 (Sleumer 1938). — *Xantolis tuberculata* (Sleumer) Baehni, *Boissiera* 11: 22 (Baehni 1965).

*Pouteria neglecta* Cronquist, *Lloydia* 9 (4): 286 [“1945” publ. 20 Dec. 1946] (Cronquist 1946).

*Pouteria cooperi* Cronquist, *Lloydia* 9 (4): 291 [“1945” publ. 20 Dec. 1946] (Cronquist 1946). — *Calocarpum cooperi* (Cronquist) Lundell, *Wrightia* 5 (7): 252 (Lundell 1976).

*Pouteria wurdackii* Aubrév., *Adansonia*, n.s., 5 (2): 201 [July 1965] (Aubréville 1965).

*Pouteria echinocarpa* W.A.Rodrigues, *Acta Amazonica* 4 (3): 9 (Rodrigues 1974).

**VERNACULAR NAMES.** — Nt: nyanboka.

**HERBARIUM DATA (FG).** — 22 collections at CAY. Sel. exs.: *D. Larpin 885*.

**INVENTORY DATA (FG).** — 78 trees in 44 plots; $F_{max}$ < 1 %; $dbh_{inv}$ = 51.3 cm.

[1688] *Pouteria venosa* (Mart.) Baehni subsp. *amazonica*
T.D.Penn.

Fl. Neotrop. Monogr. 52: 399 [26 Apr. 1990] (Pennington 1990).

*Lucuma littoralis* Mart., *Fl. Bras. [Martius]* 7: 65 [15 Jan. 1863] (Martius 1863). — *Vitellaria littoralis* (Mart.) Radlk., *Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 12: 326 (Radlkofer 1882). — *Radlkoferella littoralis* (Mart.) Pierre, *Notes Bot. Sapot.* 1: 21 (Pierre 1890).

*Vitellaria dissepala* K.Krause, *Notizbl. Königl. Bot. Gart.* Berlin 6: 171 [4 Apr. 1914] (Krause 1914). — *Lucuma dissepala* (K.Krause) Ducke, *Arch. Jard. Bot. Rio de Janeiro* 3: 233 (Ducke 1922). — *Pouteria dissepala* (K.Krause) Rizzini, *Revista Brasil. Biol.* 33 (4): 599 (Rizzini 1973), *nom. inval.* (no reference to the basionym).

*Lucuma duckei* Huber, *Bull. Soc. Bot. Genève*, sér. 2, 6: 194 [“1914” publ. 1915] (Huber 1915).

*Pouteria marginata* (Mart. & Eichler) Rizzini var. *impressa* Rizzini, *Revista Brasil. Biol.* 33 (4): 599 (Rizzini 1973).

*Pouteria dasystyla* Rizzini, *Revista Brasil. Biol.* 33 (4): 599 (Rizzini 1973).

*Pouteria dasystyla* var. *abaetensis* Rizzini, *Revista Brasil. Biol.* 33 (4): 600 (Rizzini 1973).

**VERNACULAR NAMES.** — Ka: wasepuku • Wp: pakuku, wamisalu, yà laânga • Nt: nyanboka • Br: abiurana, cutitiriba-rana.

**HERBARIUM DATA (FG).** — 26 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier 1995*.

**INVENTORY DATA (FG).** — 55 trees in 30 plots; $F_{max}$ = 1.5 %; $dbh_{inv}$ = 44.4 cm.

[1689] *Pouteria virescens* Baehni

*Candollea* 14: 66 (Baehni 1952).

**HERBARIUM DATA (FG).** — 9 collections at CAY. Sel. exs.: *D. Sabatier 1528*.

**INVENTORY DATA (FG).** — 7 trees in 4 plots; $F_{max}$ < 1 %; $dbh_{inv}$ = 107 cm.

[1690] *Pouteria williamii* (Aubrév. & Pellegr.) T.D.Penn.

Fl. Neotrop. Monogr. 52: 338 [26 Apr. 1990] (Pennington 1990). — *Eremoluma williamii* Aubrév. & Pellegr., *Adansonia*, n.s., 1 (2): 169 [“Dec. 1961” publ. Jan.-Mar. 1962] (Aubréville & Pellegrin 1962).

**VERNACULAR NAMES.** — Pa: á-maruibika, balata-kamwi • Wp: kalamûli • Br: abiurana-balatatinha.

**HERBARIUM DATA (FG).** — 10 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier 2787*.

**INVENTORY DATA (FG).** — 2 trees in 2 plots; $dh_{inv}$ = 26.2 cm.

[1691] *Pouteria* sp. A

**HERBARIUM DATA (FG).** — 2 collections at CAY. Sel. exs.: *S.A. Mori & J.J. Pipoly 15515*.

**INVENTORY DATA (FG).** — 1 tree, $dh$ = 17 cm.
[1692] *Pouteria* sp. B

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 3651*.

INVENTORY DATA (FG). — 14 trees in 14 plots; $F_{max} < 1\%$; $dbh_{inv} = 54.6$ cm.

[1693] *Pouteria* sp. C

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 2343*.

INVENTORY DATA (FG). — 15 trees in 13 plots; $F_{max} < 1\%$; $dbh_{inv} = 33.5$ cm.

[1694] *Pouteria* sp. D

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 3648*.

INVENTORY DATA (FG). — 51 trees in 35 plots; $F_{max} < 1\%$; $dbh_{inv} = 53.3$ cm.

[1695] *Pouteria* sp. E

HERBARIUM DATA (FG). — 18 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 1266*.

INVENTORY DATA (FG). — 131 trees in 29 plots; $F_{max} = 6\%$; $dbh_{inv} = 52.2$ cm.

[1696] *Pouteria* sp. F

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *J.-F. Molino 1617*.

INVENTORY DATA (FG). — 12 trees in 10 plots; $F_{max} < 1\%$; $dbh_{inv} = 57.9$ cm.

[1697] *Pouteria* sp. G

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 5554*.

INVENTORY DATA (FG). — 5 trees in 3 plots; $F_{max} < 1\%$; $dbh_{inv} = 86$ cm.

[1698] *Pouteria* sp. H

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 5550*.

INVENTORY DATA (FG). — 1 tree, $dbh = 33.4$ cm.

[1699] *Pouteria* sp. I

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 5628*.

INVENTORY DATA (FG). — 14 trees in 3 plots; $F_{max} = 2.1\%$; $dbh_{inv} = 34.9$ cm.

## Genus *Pradosia* Liais

[1700] *Pradosia cochlearia* (Lecomte) T.D.Penn.

Fl. Neotrop. Monogr. 52: 656 [26 Apr. 1990] (Pennington 1990). — *Chrysophyllum cochlearium* Lecomte, *Notul. Syst. (Paris)* 4: 63 (Lecomte 1923). — *Ecclinusa cochlearia* (Lecomte) Aubrév., *Adansonia*, n.s., 1 (1): 20 [Nov. 1961] (Aubréville 1961).

VERNACULAR NAMES. — Pa: uu-kamwi-puvemna • Nt: gaan kinboto • Br: casca-doce, miraceem.

HERBARIUM DATA (FG). — 16 collections at CAY. Sel. exs.: *R. Benoist 224* (holo-, P[P00649438]; iso-, P[P00649439]).

INVENTORY DATA (FG). — 449 trees in 78 plots; $F_{max} = 4.1\%$; $dbh_{inv} = 129.9$ cm.

[1701] *Pradosia huberi* (Ducke) Ducke

Trop. Woods 71: 16 (Ducke 1942). — *Glycoxylon huberi* Ducke, *Arch. Jard. Bot. Rio de Janeiro* 3: 235 (Ducke 1922).

VERNACULAR NAMES. — Pa: á-maruibika • Br: casca-doce, pau-doce, prahuuba-de-leite.

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 2895*.

INVENTORY DATA (FG). — 7 trees in 4 plots; $F_{max} < 1\%$; $dbh_{inv} = 50$ cm.

[1702] *Pradosia ptychandra* (Eyma) T.D.Penn. (Fig. 55E)

Fl. Neotrop. Monogr. 52: 648 [26 Apr. 1990] (Pennington 1990). — *Pouteria ptychandra* Eyma, *Recueil Trav. Bot. Néerl.* 33: 189 (Eyma 1936). — *Neopometia ptychandra* (Eyma) Aubrév., *Adansonia*, n.s., 1 (1): 26 [Nov. 1961] (Aubréville 1961).

Voyara montana Aubl., *Hist. Pl. Guiane* 2 (Suppl.): 26 [Jun.-Dec. 1775] (Aublet 1775), *pro parte fruto excludo*. — *Capparis montana* (Aubl.) Lemée, *Fl. Guyane Franç.* 1: 666 (Lemée 1955).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Pa: inam-etni-seinô • Wp: tasiasiay, wila ê'ê • Wn: tamaniwa • Nt: busi amanda, busi amandaa, kinboto • Cr: balata-ponm, grënn-kwata • Fr: balata pommier • Br: casca-doce.

HERBARIUM DATA (FG). — 78 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material of *Voyara montana*: BM[BM000952572]).

INVENTORY DATA (FG). — 229 trees in 97 plots; $F_{max} = 2.2\%$; $dbh_{inv} = 58.6$ cm.
<figure><img src="image_87.png" /><figcaption>FIG. 55. — Sapotaceae: **A**, *Pouteria benai* (Aubrév. & Pellegr.) T.D.Penn. (*D. Sabatier & M.-F. Prévost 3533*); **B**, *Pouteria eugenifolia* (Pierre) Baehni (*D. Sabatier* 5045); **C**, *Pouteria sagotiana* (Baill.) Eyma; *Pradosia ptychandra* (Eyma) T.D.Penn. (*D. Sabatier* 823); **D**, *Pouteria jariensis* Pires & T.D.Penn. (*D. Sabatier & J.-F. Molino* 5541); **E**, *Pradosia ptychandra* (Eyma) T.D.Penn. (*D. Sabatier* 823). A, B, D, © D. Sabatier/IRD; C, © J.-F. Molino/IRD; E, © M.-F. Prévost/IRD.</figcaption></figure>

[1703] *Pradosia surinamensis* (Eyma) T.D.Penn.

Fl. Neotrop. Monogr. 52: 652 [26 Apr. 1990] (Pennington 1990). — *Pouteria surinamensis* Eyma, *Recueil Trav. Bot. Néerl.* 33: 189 (Eyma 1936).

VERNACULAR NAMES. — Ka: ololome, olome • Wn: tamaniwa • Nt: kinboto • Br: abiurana-branca.

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 3319*.

INVENTORY DATA (FG). — 3 trees in 1 plots; Fmax < 1%; dbhinv = 35 cm.

[1704] *Pradosia verticillata* Ducke

Trop. Woods 71: 12 (Ducke 1942).

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *T.D. Pennington & S.A. Mori 12134*.

INVENTORY DATA (FG). — 2 trees in 2 plots; Fmax < 1%; dbhinv = 14.2 cm.

[1705] *Pradosia* sp. A

NOTE. — A species resembling *P. subverticillata* Ducke.

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: *D. Sabatier 1772*, dbh = 80 cm.

Genus *Sarcaulus* Radlk.

[1706] *Sarcaulus brasiliensis* (A.DC.) Eyma

*Recueil Trav. Bot. Néerl.* 33: 192 (Eyma 1936). — *Chrysophyllum brasiliense* A.DC., *Prodr.* [*A. P. de Candolle*] 8: 156 [mid Mar. 1844] (Candolle 1844).

*Chrysophyllum macrophyllum* Mart., *Flora* 21 (2, Beibl.): 95 (Martius 1838), *nom. illeg. hom.*, *non* Lam. (Lamarck 1794) *nec* C.F.Gaertn. (Gartner 1806) *nec* Sabine (1824), *nec* Desf. (Desfontaines 1829). — *Achras brasiliensis* Willd. ex Steud., *Nomencl. Bot. [Steudel]*, ed. 2, 1: 15 (Steudel 1840), *nom. illeg. superfl.* (based on *Chrysophyllum macrophyllum*).

*Chrysophyllum brasiliense* var. minus Miq., *Fl. Bras. [Martius]* 7: 103 [15 Jan. 1863] (Miquel 1863).
*Sarcaulus macrophyllus Radlk., Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 12: 311 (Radlkofer 1882).

**VERNACULAR NAMES.** — Pa: ukum-kamwi, uu-kamwi-wahuyo • Wp: wai • Br: ajará, guajara-do-igapó, jaraí.

**HERBARIUM DATA (FG).** — 17 collections at CAY. Sel. exs.: *D. Sabatier* 2338.

**INVENTORY DATA (FG).** — 8 trees in 7 plots; $F_{max} < 1\%$; $dbh_{inv} = 31.5$ cm.

Family SIMAROUBACEAE DC.
Genus *Homalolepis* Turcz.

[1707] *Homalolepis cedron* (Planch.) Devecchi & Pirani

*Phytotaxa* 366 (1): 67 [epubl. 28 Aug. 2018] (Devecchi & Pirani 2018). — *Simaba cedron* Planch., *London J. Bot.* 5: 566 (Planchon 1846). — *Quassia cedron* (Planch.) Baill., *Hist. Pl. [Baillon]* 4: 406 [Jul.-Sep. 1873] (Baillon 1873). — *Aruba cedron* (Planch.) Kuntze, *Revis. Gen. Pl.* 1: 103 [5 Nov. 1891] (Kuntze 1891).

**VERNACULAR NAMES.** — Pa: timaa-avain-purubumna • Nt: wan ede • Cr: wan-édé • Br: manga-de-anta, pau-para-tudo.

**HERBARIUM DATA (FG).** — 9 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 2255.

**INVENTORY DATA (FG).** — 485 trees in 77 plots; $F_{max} = 6.9\%$; $dbh_{inv} = 29.8$ cm.

[1708] *Homalolepis morettii* (Feuillet) Devecchi & Pirani
(Fig. 56A)

*Phytotaxa* 366 (1): 43 [epubl. 28 Aug. 2018] (Devecchi & Pirani 2018). — *Simaba morettii* Feuillet, *Candollea* 38 (2): 746 (Feuillet 1983).

**NOTE.** — Known only from French Guiana.

**VERNACULAR NAMES.** — Nt: busi kananbuli • Cr: koubari-savann, mal-simarouba.

**HERBARIUM DATA (FG).** — 16 collections at CAY. Sel. exs.: *C. Moretti* 1027 (holo-, P, not seen; iso-, CAY[CAY024960, CAY024961, CAY024962], P[P01817225]).

**INVENTORY DATA (FG).** — 77 trees in 50 plots; $F_{max} = 1\%$; $dbh_{inv} = 86.3$ cm.

Genus *Picrolemma* Hook.f.

[1709] *Picrolemma sprucei* Hook.f.

*Gen. Pl. [Bentham & Hooker f.]* 1 (1): 312 [7 Aug. 1862] (Hooker 1862).

*Picrolemma pseudocoffea* Ducke, *Arch. Jard. Bot. Rio de Janeiro* 4: 196 (Ducke 1925).

**VERNACULAR NAMES.** — Pa: tuu-ariut, tuu-kamwi • Cr: kafé-lane • Br: caferana.

**HERBARIUM DATA (FG).** — 17 collections at CAY. Sel. exs.: *M.-F. Prévost et al.* 4440.

**SIZE.** — Up to 10 m tall (Thomas & Franceschinelli 2005).

Genus *Quassia* L.

[1710] *Quassia amara* L.

*Sp. Pl.*, ed. 2, 1: 553 [Sep. 1762] (Linnaeus 1762).

*Quassia amara* var. paniculata Engl., *Fl. Bras. [Martius]* 12 (2): 207 [1 Sep. 1874] (Engler 1874). — *Quassia amara* f. paniculata (Engl.) Cronquist, *Brittonia* 5: 146 (Cronquist 1944).

**NOTE.** — A species native to Central America and the Caraibes, probably introduced to French Guiana during the 18th century by French Jesuit priests (Odonne *et al.* 2020).

**VERNACULAR NAMES.** — Pa: kwašvan • Ka: ki'elepun, kuasi bita, kuwasi, pekui • Nt: bita kwasi, kwasi • Cr: bwa-kwachi, kwachi • Fr: couachi • Br: quina, quina-de-Caiena.

**HERBARIUM DATA (FG).** — 19 collections at CAY. Sel. exs.: *P. Grenand* 2118.

**SIZE.** — Costa Rica. *K. Martínez Sequeira* 280 (MO), 9 m × 10 cm.

Genus *Simaba* Aubl.

[1711] *Simaba guianensis* Aubl.

*Hist. Pl. Guiane* 1: 400 [Jun.-Dec. 1775] (Aublet 1775), “*Guyannensis*” on plate. — *Zwingera amara* Willd., *Sp. Pl.*, ed. 4 2 (1): 569 [Mar. 1799] (Willdenow 1799), *nom. illeg. superfl.* (based on *Simaba guianensis*). — *Quassia crocea* Vahl, *Eclog. Amer.* 3: 12 (Vahl 1807), *nom. illeg. superfl.* (based on *Simaba guianensis*). — *Quassia guianensis* (Aubl.) D.Dietr., *Syn. Pl. [D. Dietrich]* 2: 1416 [1-20 Dec. 1840] (Dietrich 1840).

*Aruba guianensis* Aubl., *Hist. Pl. Guiane* 1: 294 [Jun.-Dec. 1775] (Aublet 1775), “*Guyannensis*” on plate. — *Simaba guianensis* (Aubl.) Engl., *Fl. Bras. [Martius]* 12 (2): 211 [1 Sep. 1874] (Engler 1874), *nom. illeg. hom.*, *non* Aubl. (Aublet 1775).

*Zwingera aruba* Spreng., *Syst. Veg. [Sprengel]* 2: 319 [Jan.-May 1825] (Sprengel 1825). — *Quassia aruba* (Spreng.) D.Dietr., *Syn. Pl. [D. Dietrich]* 2: 1416 [1-20 Dec. 1840] (Dietrich 1840).

*Simaba cuspidata* Spruce ex Engl., *Fl. Bras. [Martius]* 12 (2): 212 [1 Sep. 1874] (Engler 1874). — *Quassia cuspidata* (Spruce ex Engl.) Noot., *Blumea* 11: 522 [15 Dec. 1962] (Nooteboom 1962).

**NOTE.** — *Zwingera amara* Willd. is based on *Simaba guianensis* Aubl., not on *Simarouba amara* Aubl.

**VERNACULAR NAMES.** — Wp: wasaku sili • Br: cajuirana, maruparana.

**HERBARIUM DATA (FG).** — 41 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lectotype of *Aruba guianensis* Aubl. [P-JJR, P00778459] designated by Lanjouw & Uittien [1940: 148]).

**INVENTORY DATA (FG).** — 17 trees in 12 plots; $F_{max} < 1\%$; $dbh_{inv} = 36.1$ cm.
[1712] *Simaba orinocensis* Kunth

*Nova genera et species plantarum [H.B.K.] 6: 18* [14 Apr. 1823] (Kunth 1823). — *Zwingera orinocensis* (Kunth) Spreng., *Syst. Veg. [Sprengel] 2: 319* [Jan.-May 1825] (Sprengel 1825). — *Quassia orinocensis* (Kunth) D.Dietr., *Syn. Pl. [D. Dietrich] 2: 1416* [1-20 Dec. 1840] (Dietrich 1840). — *Aruba orinocensis* (Kunth) Kuntze, *Revis. Gen. Pl. 1: 103* [5 Nov. 1891] (Kuntze 1891).

*Simaba multiflora* A.Juss., *Mém. Mus. Hist. Nat. 12: t. 27, f. 45a* (Jussieu 1825). — *Quassia multiflora* (A.Juss.) Noot., *Blumea 11: 522* [15 Dec. 1962] (Nootboom 1962).

*Simaba foetida* Benth., *Hooker's J. Bot. Kew Gard. Misc. 3: 370* (Bentham 1851).

*Simaba angustifolia* Benth., *Hooker's J. Bot. Kew Gard. Misc. 3: 371* (Bentham 1851).

*Simaba crustacea* Engl., *Fl. Bras. [Martius] 12* (2): 211 [1 Sep. 1874] (Engler 1874). — *Aruba crustacea* (Engl.) Kuntze, *Revis. Gen. Pl. 1: 103* [5 Nov. 1891] (Kuntze 1891). — *Quassia crustacea* (Engl.) Noot., *Blumea 11: 522* [15 Dec. 1962] (Nootboom 1962).

*Simaba alata* Maguire, *Bull. Torrey Bot. Club 75* (4): 397 [Jul.-Aug. 1948] (Maguire 1948).

**VERNACULAR NAMES.** — Pa: kwepan, kwepan-wašiunu • Ka: kuyali, pakoweni • Wp: paku akayu, sokolo • Wn: kanapali • Nt: kananbuli • Cr: kananbouli • Br: cajurana, papacuaió.

**HERBARIUM DATA (FG).** — 9 collections at CAY. Sel. exs.: *P. Gre- nand 1291*.

**SIZE.** — Up to 20 cm dbh (Cavalcante 1983).

[1713] *Simaba polyphylla* (Cavalcante) W.W.Thomas

*Acta Amazonica* 15 (1-2, Suppl.): 76 [“1985” publ. 1987] (Thomas 1987). — *Simaba guianensis* subsp. *polyphylla* Cavalcante, Publ. Avulsas Mus. Paraense Emílio Goeldi 37: 40 (Cavalcante 1983).

**VERNACULAR NAMES.** — Pa: timaa-avain • Br: marupazinho.

**HERBARIUM DATA (FG).** — 19 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier 2788*.

**INVENTORY DATA (FG).** — 57 trees in 40 plots; $F_{max} < 1\%$; $dbh_{inv} = 41.4$ cm.

Genus *Simarouba* Aubl.

[1714] *Simarouba amara* Aubl.

*Hist. Pl. Guiane* 2: 860 [Jun.-Dec. 1775] (Aublet 1775).

*Quassia dioica* P.J.Bergius, *Mater. Med.:* 355 (Bergius 1778).

*Quassia simaruba* L.f., *Suppl. Pl.:* 234 [“1781” publ. Apr. 1782] (Linnaeus 1782).

*Quassia officinalis* Rich., *Actes Soc. Hist. Nat. Paris* 1: 108 [Oct. 1792] (Richard 1792).

*Simarouba glauca* DC., *Ann. Mus. Natl. Hist. Nat.* 17: 424 (Candolle 1811), “*Simaruba*”. — *Quassia glauca* (DC.) Spreng., *Syst. Veg. [Sprengel] 2: 319* [Jan.-May 1825] (Sprengel 1825).

*Quassia alatifolia* Stokes, *Bot. Mat. Med.* 2: 491 (Stokes 1812).

*Simarouba amara* var. *opaca* Engl., *Nat. Pflanzenfam. [Engler & Prantl] 3* (4): 213 (Engler 1896). — *Simarouba opaca* (Engl.) Radlk. ex F. Boas, *Beih. Bot. Centralbl.* 29 (1): 332 (Boas 1913).

*Simarouba amara* var. *typica* Cronquist, *Bull. Torrey Bot. Club* 71 (3): 229 [May 1944] (Cronquist 1944), *nom. inval.* (Turland et al. 2018: Art. 24.3).

*Simarouba glauca* var. *latifolia* Cronquist, *Bull. Torrey Bot. Club* 71 (3): 231 [May 1944] (Cronquist 1944).

*Simarouba glauca* var. *typica* Cronquist, *Bull. Torrey Bot. Club* 71 (3): 231 [May 1944] (Cronquist 1944), *nom. inval.* (Turland et al. 2018): Art. 24.3).

*Simarouba amara* var. *puberula* Cuatrec., *Revista Acad. Colomb. Ci. Exact.* 8 (32): 466 [June 1952] (Cuatrecasas 1952).

**NOTE.** — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

**VERNACULAR NAMES.** — Pa: timaa-avain • Ka: šimaluba, simalupa, sumaliba • Wp: iwēʾi • Wn: pëunpë • Nt: sumaypa • Cr: akajoublan, simarouba • Fr: simarouba • Br: marupa, paraiba.

**HERBARIUM DATA (FG).** — 61 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material not traced); *R.A.A. Oldeman B-1925* (P).

**INVENTORY DATA (FG).** — 77 trees in 62 plots; $F_{max} < 1\%$; $dbh_{inv} = 90$ cm.

Family *SIPARUNACEAE* Schodde
Genus *Siparuna* Aubl.

[1715] *Siparuna cristata* (Poepp. & Endl.) A.DC.

*Prodr. [A. P. de Candolle] 16* (2): 655 [mid July 1868] (Candolle 1868). — *Citrosma cristata* Poepp. & Endl., *Nova genera ac species plantarum [Poeppig & Endlicher] 2: 47* [Jan.-Sep. 1838] (Poeppig & Endlicher 1838), “*cristatum*”.

*Siparuna cristata* var. *macrophylla* A.DC., *Prodr. [A. P. de Candolle] 16* (2): 655 [mid July 1868] (Candolle 1868).

*Siparuna cristata* var. *petiolaris* A.DC., *Prodr. [A. P. de Candolle] 16* (2): 655 [mid July 1868] (Candolle 1868).

*Siparuna williamsii* J.F.Macbr., *Candollea* 5: 353 (Macbride 1934).

*Siparuna monogyna* Jangoux, *Bol. Mus. Paraense Emílio Goeldi, N.S., Bot.* 7 (1): 133 (Jangoux 1991).

*Siparuna obconica* Jangoux, *Bol. Mus. Paraense Emílio Goeldi, N.S., Bot.* 7 (1): 135 (Jangoux 1991).

**VERNACULAR NAMES.** — Pa: yahiwemna-kamwi, yariwapna-kamwi • Wp: mila suku, wila suku.

**HERBARIUM DATA (FG).** — 18 collections at CAY. Sel. exs.: *J. Martin s.n.* (holotype of *Siparuna cristata* var. *petiolaris*: P[P016912]).

**INVENTORY DATA (FG).** — 31 trees in 13 plots; $F_{max} = 1.3\%$; $dbh_{inv} = 19.4$ cm.
[1716] *Siparuna cuspidata* (Tul.) A.DC.

Prodr. [A. P. de Candolle] 16 (2.2): 655 [mid July 1868] (Candolle 1868). — *Citrosma cuspidata* Tul., *Ann. Sci. Nat.*, Bot. sér. 4, 3: 40 [May 1855] (Tulasne 1855), "*Citriosma*".

*Citrosma myristicoidea* Spruce, *J. Proc. Linn. Soc.*, Bot. 5: 5 ["1861" publ. June 1860] (Spruce 1860). — *Siparuna crassiflora* Perkins, Bot. Jahrb. Syst. 28 (5): 702 [11 Jan. 1901] (Perkins 1901), nom. illeg. superfl. (based on the type of *Citrosma myristicoidea*).

*Siparuna argyrochrysea* Perkins, Bot. Jahrb. Syst. 28 (5): 702 [11 Jan. 1901] (Perkins 1901).

*Siparuna lepidiflora* Perkins, Notizbl. Bot. Gart. Berlin-Dahlem 10: 166 [20 Dec. 1927] (Perkins 1927).

*Siparuna ucayaliensis* Perkins, Notizbl. Bot. Gart. Berlin-Dahlem 10: 167 [20 Dec. 1927] (Perkins 1927).

HERBARIUM DATA (FG). — 80 collections at CAY. Sel. exs.: *E.M. Mélinon s.n.* (original material of *Siparuna argyrochrysea*: holo-, B (not seen, photo F neg 13478, fragm. GH[GH00046549]); possible iso-, P[P00053272, P00053273, P00053274]).

INVENTORY DATA (FG). — 36 trees in 19 plots; $F_{max}$ = 1.7%; $dbh_{inv}$ = 18.8 cm.

[1717] *Siparuna decipiens* (Tul.) A.DC.

Prodr. [A. P. de Candolle] 16 (2): 643 [mid July 1868] (Candolle 1868). — *Citrosma decipiens* Tul., *Arch. Mus. Hist. Nat.* 8: 368 (Tulasne 1855), "*Citriosma*".

*Conuleum guyannense* Rich. ex A.Rich., *Mém. Soc. Hist. Nat. Paris*, sér. 2, 1: 391 (Richard 1823).

*Siparuna lepidantha* Perkins, Bot. Jahrb. Syst. 28 (5): 701 [11 Jan. 1901] (Perkins 1901).

NOTE. — Hyperdominant in Amazonia (ter Steege et al. 2020).

VERNACULAR NAMES. — Pa: á-kamwi, kwatri-purubumna • Ka: piyawapan wewe • Wp: mila suku, wila suku • Wn: awalepokan • Nt: faya pao • Br: amarelinho, quariquara-branca.

HERBARIUM DATA (FG). — 122 collections at CAY. Sel. exs.: *J. Martin s.n.* (original material K[K000587966], MEL[MEL2384184], P[P016899, P00053052, P00053053], S[S-R-7098]).

INVENTORY DATA (FG). — 329 trees in 125 plots; $F_{max}$ = 3.3%; $dbh_{inv}$ = 73.5 cm.

[1718] *Siparuna guianensis* Aubl.
(Fig. 56B)

*Hist. Pl. Guiane* 2: 865 [Jun.-Dec. 1775] (Aublet 1775), "*Guyannensis*" on plate. — *Citrosma guianensis* (Aubl.) Tul., *Arch. Mus. Hist. Nat.* 8: 361 (Tulasne 1855), "*Citriosma*".

*Citrosma discolor* Poepp. & Endl., *Nova genera ac species plantarum* [*Poeppig & Endlicher*] 2: 47 [Jan.-Sep. 1838] (Poeppig & Endlicher 1838). — *Siparuna discolor* (Poepp. & Endl.) A.DC., *Prodr. [A. P. de Candolle]* 16 (2.2): 656 [mid July 1868] (Candolle 1868).

*Citrosma camporum* Tul., *Ann. Sci. Nat.*, Bot. sér. 4, 3: 39 [May 1855] (Tulasne 1855), "*Citriosma*". — *Siparuna camporum* (Tul.)

A.DC., *Prodr. [A. P. de Candolle]* 16 (2.2): 654 [mid July 1868] (Candolle 1868).

*Citrosma guianensis* var. *divergentifolia* Tul., *Arch. Mus. Hist. Nat.* 8: 362 (Tulasne 1855), "*Citriosma*". — *Siparuna guianensis* var. *divergentifolia* (Tul.) A.DC., *Prodr. [A. P. de Candolle]* 16 (2.2): 654 [mid July 1868] (Candolle 1868).

*Citrosma guianensis* var. nuda Tul., *Arch. Mus. Hist. Nat.* 8: 362 (Tulasne 1855), "*Citriosma*".

*Siparuna panamensis* A.DC., *J. Bot.* 3: 219 (Candolle 1865).

*Siparuna guianensis* var. *glabrescens* A.DC., *Prodr. [A. P. de Candolle]* 16 (2.2): 654 [mid July 1868] (Candolle 1868), *nom. illeg. superfl.* (one of the syntypes is also a syntype of *S. guianensis* var. nuda).

*Siparuna guianensis* var. *longifolia* A.DC., *Prodr. [A. P. de Candolle]* 16 (2.2): 654 [mid July 1868] (Candolle 1868).

*Siparuna foetida* Barb.Rodr., *Vellozia*, ed. 2, 1: 68 (Barbosa Rodrigues 1891).

*Siparuna guianensis* var. *nitens* Kuntze, *Revis. Gen. Pl.* 3 (3): 276 [28 Sep. 1898] (Kuntze 1898).

*Siparuna archeri* A.C.Sm., *Bull. Torrey Bot. Club* 59 (9): 517 [Dec. 1932] (Smith 1932).

*Siparuna arianeae* V.Pereira, *Bradea* 4 (36): 291 (Pereira 1986).

*Siparuna cavalcantei* Jangoux, *Bol. Mus. Paraense Emilio Goeldi*, *N.S.*, Bot. 7 (1): 120 (Jangoux 1991).

*Siparuna duckeana* Jangoux, *Bol. Mus. Paraense Emilio Goeldi*, *N.S.*, Bot. 7 (1): 125 (Jangoux 1991).

*Siparuna itacaiunensis* Jangoux, *Bol. Mus. Paraense Emilio Goeldi*, *N.S.*, Bot. 7 (1): 129 (Jangoux 1991).

*Siparuna savanicola* Jangoux, *Bol. Mus. Paraense Emilio Goeldi*, *N.S.*, Bot. 7 (1): 144 (Jangoux 1991).

VERNACULAR NAMES. — Pa: yahiwemna, yariwapna • Ka: idyakopi, ilakopi, yalakopi • Te: dzawat móböha • Wp: enëmi'o, nëmi'o, wainimi'o • Wn: jalëkë, jawilëkë • Nt: kapasi wiwii • Cr: vénéré, viniré • Br: caá-pitiú.

HERBARIUM DATA (FG). — 140 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000611207]).

INVENTORY DATA (FG). — 1 tree, $dbh$ = 25 cm.

[1719] *Siparuna pachyantha* A.C.Sm.

*J. Arnold Arbor.* 20 (3): 293 [19 July 1939] (Smith 1939).

*Siparuna emarginata* R.S.Cowan, *Brittonia* 8 (4): 237 [Jan. 1957] (Cowan 1957).

VERNACULAR NAMES. — Pa: avakni-avak, avatni-awak • Wp: a'i makule • Cr: féy-tabak • Br: caá-pitiú-de-folha-grande.

HERBARIUM DATA (FG). — 25 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino* 5552.

INVENTORY DATA (FG). — 24 trees in 16 plots; $F_{max}$ < 1%; $dbh_{inv}$ = 70 cm.
[1720] *Siparuna poeppigii* (Tul.) A.DC.

Prodr. [A. P. de Candolle] 16 (2): 653 [mid July 1868] (Candolle 1868). — *Citrosma poeppigii* Tul., *Ann. Sci. Nat., Bot.* sér. 4, 3: 39 [May 1855] (Tulasne 1855), “*Citriosma*”.

Tetratome lepidota Poepp. & Endl., *Nova genera ac species plantarum* [*Poeppig & Endlicher* 2: 47 [Jan.-Sep. 1838] (Poeppig & Endlicher 1838).

*Siparuna sprucei* A.DC., *J. Bot.* 3: 219 (Candolle 1865).

*Siparuna amazonica* Mart. ex A.DC., *Prodr. [A. P. de Candolle]* 16 (2.2): 653 [mid July 1868] (Candolle 1868).

*Siparuna espinhacensis* Jangoux, *Bol. Mus. Paraense Emílio Goeldi*, *N.S.*, *Bot.* 7 (1): 127 (Jangoux 1991).

VERNACULAR NAMES. — Pa: yahiwemna, yariwapna • Wp: eněmi'o, němi'o, wainimi'o • Cr: vénéré, viniré • Br: caá-pitiú.

HERBARIUM DATA (FG). — 71 collections at CAY. Sel. exs.: *R.A.A. Oldeman* 1916.

SIZE. — Up to 20 cm dbh (Renner & Hausner 2005).

Family SOLANACEAE Juss.
Genus *Solanum* L.

[1721] *Solanum crinitum* Lam.

Tabl. Encycl. 2[4 (1)]; 20 [21 Nov. 1796] (Lamarck 1796).

*Solanum formosum* Humb. & Bonpl. ex Dunal, *Solan. Syn.*: 35 [Oct. 1816] (Dunal 1816).

*Solanum jubatum* Willd., *Syst. Veg. [Roemer & Schultes]* 4: 669 [Mar.-June 1819] (Willdenow 1819). — *Solanum cyananthum* var. *jubatum* (Willd.) Dunal, *Prodr. [A. P. de Candolle]* 13 (1): 316 [10 May 1852] (Dunal 1852).

*Solanum cyananthum* Dunal, *Prodr. [A. P. de Candolle]* 13 (1): 316 [10 May 1852] (Dunal 1852).

NOTE. — *Solanum jubatum* Willd. is to be ascribed to Willdenow alone (Turland *et al.* 2018: Art. 46.3, Ex. 15; see also IPNI, 2020 and Taylor & Gereau, 2019).

VERNACULAR NAMES. — Pa: puduku • Ka: kwasisi, kwašiši, palemulu, poto: no palemulu • Wp: yū ásīsī • Nt: mantuga maka • Cr: ponm-sousouri, tonmat-sousouri • Br: jurubeba.

HERBARIUM DATA (FG). — 24 collections at CAY. Sel. exs.: *P. Grenand* 15.

SIZE. — Colombia, Nariño. *M.I. Barreto* 141 (MO), 14 m × 30 cm.

[1722] *Solanum endopogon* (Bitter) Bohs subsp. *guianense* (Bohs) Bohs

*J. Bot. Res. Inst. Texas* 3 (2): 509 [25 Nov. 2009] (Bohs 2009). — *Cyphomandra endopogon* Bitter subsp. *guianensis* Bohs, *Fl. Neotrop. Monogr.* 63: 90 [5 July 1994] (Bohs 1994).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Wp: awala kāsī, uluwu kāsī • Cr: mavévéchien • Br: pau-mirim.

HERBARIUM DATA (FG). — 25 collections at CAY. Sel. exs.: *S.A. Mori & B.M. Boom* 14709 (holo-, F[V0072836F]; iso-, CAY[CAY070876], NY[00721521]); *S.A. Mori et al.* 15028, 15 m × 20 cm.

[1723] *Solanum splendens* (Dunal) Bohs (Fig. 56C)

Monogr. Syst. Bot. Missouri Bot. Gard. 131: 320 (Bohs 2015). — *Cyphomandra splendens* Dunal, *Prodr. [A. P. de Candolle]* 13 (1): 395 (Dunal 1852).

*Pionandra hartwegii* Miers, *London J. Bot.* 4: 363 (Miers 1845). — *Cyphomandra hartwegii* (Miers) Sendtn. ex Walp., *Repert. Bot. Syst. [Walpers]* 6 (4): 579 [19-22 May 1847] (Walpers 1847).

*Cyphomandra artocarpophyllos* H.J.P.Winkl., *Repert. Spec. Nov. Regni Veg.* 7: 246 (Winkler 1909).

*Cyphomandra mollicella* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 4 (8): 322 [24 Oct. 1929] (Standley 1929).

*Solanum circinatum* Bohs, *Taxon* 44 (4): 585 [Nov. 1995] (Bohs 1995).

VERNACULAR NAMES. — Pa: isuu-aveya, isuu-vey • Wp: awala kāsī, uluwu kāsī • Cr: mavévé-chien.

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *M.-F. Prévost & C. Feuillet* 3964.

SIZE. — Up to 12 m tall (Bohs 1994).

Family STEMONURACEAE Kârehed
Genus *Discophora* Miers

[1724] *Discophora guianensis* Miers (Fig. 56D)

*Ann. Mag. Nat. Hist.*, ser. 2, 10 (56): 119 [Aug. 1852] (Miers 1852). — *Kummeria brasiliensis* Mart. ex Engl., *Fl. Bras. [Martius]* 12 (2): 52 [1 Dec. 1872] (Engler 1872), *nom. illeg. superfl.* (based on *Discophora guianensis*).

*Lasianthera amazonica* Barb.Rodr., *Vellovia*, ed. 2, 1: 12 (Barbosa Rodrigues 1891).

*Discophora panamensis* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 4 (8): 222 [24 Oct. 1929] (Standley 1929).

*Discophora froesii* Pires, *Bol. Técn. Inst. Agron. N.* 38: 28 (Pires 1960).

VERNACULAR NAMES. — Pa: kanegma • Wp: ināmu sī, ināmusī wila, mulei sī, yawasi pita • Nt: boni udu • Cr: bwa-léchèl • Br: caferana, ivaí, pombinho.

HERBARIUM DATA (FG). — 31 collections at CAY. Sel. exs.: *B. Riéra & D. Sabatier* 1808.

INVENTORY DATA (FG). — 37 trees in 25 plots; Fmax < 1%; dbhinv = 15.5 cm.