*Myrcia plicatocostata* O.Berg, *Linnaea* 27 (1): 114 ["1854" publ. Nov. 1855] (Berg 1855), "plicato-costata".

*Myrcia sericea* O.Berg, *Linnaea* 27 (1): 114 ["1854" publ. Nov. 1855] (Berg 1855), *nom. illeg. hom.*, *non* G.Don (1832).

*Eugenia mollis* Willd. ex O.Berg, *Linnaea* 27 (1): 121 ["1854" publ. Nov. 1855] (Berg 1855), *nom. nud. pro syn.*

*Aulomyrcia costata* O.Berg, *Fl. Bras. [Martius]* 14 (1): 79 [15 May 1857] (Berg 1857).

*Myrcia costata* var. *bahiensis* O.Berg, *Fl. Bras. [Martius]* 14 (1): 157 [15 May 1857] (Berg 1857).

*Myrcia costata* var. *minensis* O.Berg, *Fl. Bras. [Martius]* 14 (1): 157 [15 May 1857] (Berg 1857).

*Myrcia elongata* O.Berg, *Fl. Bras. [Martius]* 14 (1): 159 [15 May 1857] (Berg 1857).

*Myrcia martiana* O.Berg, *Fl. Bras. [Martius]* 14 (1): 159 [15 May 1857] (Berg 1857). — *Myrcia rufula* var. *martiana* (O.Berg) Kiaersk., * Enum. Myrt. Bras. * 55 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Myrcia elongata* var. *brunnea* O.Berg, *Fl. Bras. [Martius]* 14 (1): 160 [15 May 1857] (Berg 1857).

*Myrcia elongata* var. *grandifolia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 160 [15 May 1857] (Berg 1857).

*Myrcia elongata* var. *ochracea* O.Berg, *Fl. Bras. [Martius]* 14 (1): 160 [15 May 1857] (Berg 1857).

*Myrcia guajavifolia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 160 [15 May 1857] (Berg 1857), "guajavaefolia".

*Myrcia guajavifolia* var. *impunctata* O.Berg, *Fl. Bras. [Martius]* 14 (1): 160 [15 May 1857] (Berg 1857).

*Myrcia guajavifolia* var. *perforata* O.Berg, *Fl. Bras. [Martius]* 14 (1): 160 [15 May 1857] (Berg 1857).

*Myrcia friburgensis* O.Berg, *Fl. Bras. [Martius]* 14 (1): 161 [15 May 1857] (Berg 1857).

*Myrcia guajavifolia* var. *bullata* O.Berg, *Fl. Bras. [Martius]* 14 (1): 161 [15 May 1857] (Berg 1857).

*Myrcia magnoliifolia* var. *angustifolia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 162 [15 May 1857] (Berg 1857).

*Myrcia magnoliifolia* var. *latifolia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 162 [15 May 1857] (Berg 1857).

*Myrcia magnoliifolia* var. *parvifolia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 162 [15 May 1857] (Berg 1857).

*Myrcia brandamii* O.Berg, *Fl. Bras. [Martius]* 14 (1): 164 [15 May 1857] (Berg 1857), "Brandami".

*Myrcia alagoensis* O.Berg, *Fl. Bras. [Martius]* 14 (1): 165 [15 May 1857] (Berg 1857).

*Myrcia alagoensis* var. *intermedia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 165 [15 May 1857] (Berg 1857).

*Myrcia alagoensis* var. *oblongata* O.Berg, *Fl. Bras. [Martius]* 14 (1): 165 [15 May 1857] (Berg 1857).

*Myrcia alagoensis* var. *ovata* O.Berg, *Fl. Bras. [Martius]* 14 (1): 165 [15 May 1857] (Berg 1857).

*Myrcia spruceana* O.Berg, *Fl. Bras. [Martius]* 14 (1): 165 [15 May 1857] (Berg 1857).

*Myrcia phaeoclada* O.Berg, *Fl. Bras. [Martius]* 14 (1): 167 [15 May 1857] (Berg 1857).

*Myrcia phaeoclada* var. *alagoensis* O.Berg, *Fl. Bras. [Martius]* 14 (1): 167 [15 May 1857] (Berg 1857).

*Myrcia phaeoclada* var. *guyanensis* O.Berg, *Fl. Bras. [Martius]* 14 (1): 167 [15 May 1857] (Berg 1857).

*Myrcia kegeliana* O.Berg, *Fl. Bras. [Martius]* 14 (1): 168 [15 May 1857] (Berg 1857).

*Myrcia kegeliana* var. *angustifolia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 168 [15 May 1857] (Berg 1857).

*Myrcia kegeliana* var. *latifolia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 168 [15 May 1857] (Berg 1857).

*Myrcia kegeliana* var. *longifolia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 168 [15 May 1857] (Berg 1857).

*Myrcia kegeliana* var. *pendula* O.Berg, *Fl. Bras. [Martius]* 14 (1): 168 [15 May 1857] (Berg 1857).

*Myrcia kegeliana* var. *vulgaris* O.Berg, *Fl. Bras. [Martius]* 14 (1): 168 [15 May 1857] (Berg 1857).

*Myrcia klotzschiana* O.Berg, *Fl. Bras. [Martius]* 14 (1): 169 [15 May 1857] (Berg 1857).

*Myrcia micrantha* O.Berg, *Fl. Bras. [Martius]* 14 (1): 169 [15 May 1857] (Berg 1857).

*Myrcia berberis* var. *angustifolia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 170 [15 May 1857] (Berg 1857).

*Myrcia berberis* var. *latifolia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 170 [15 May 1857] (Berg 1857).

*Myrcia latifolia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 170 [15 May 1857] (Berg 1857).

*Myrcia erythroxylon* O.Berg, *Fl. Bras. [Martius]* 14 (1): 173 [15 May 1857] (Berg 1857).

*Myrcia erythroxylon* var. *caerulescens* O.Berg, *Fl. Bras. [Martius]* 14 (1): 173 [15 May 1857] (Berg 1857).

*Myrcia pellucida* O.Berg, *Fl. Bras. [Martius]* 14 (1): 173 [15 May 1857] (Berg 1857).

*Myrcia gracilis* O.Berg, *Fl. Bras. [Martius]* 14 (1): 174 [15 May 1857] (Berg 1857). — *Myrcia rostrata* f. *gracilis* (O.Berg) D.Legrand, *Fl. Ilustr. Catar. 1 (Mirt.)*: 240 (Legrand 1969).

*Myrcia gracilis* var. *prasina* O.Berg, *Fl. Bras. [Martius]* 14 (1): 174 [15 May 1857] (Berg 1857).

*Myrcia acutata* O.Berg, *Fl. Bras. [Martius]* 14 (1): 175 [15 May 1857] (Berg 1857).

*Myrcia gracilis* var. *opaca* O.Berg, *Fl. Bras. [Martius]* 14 (1): 175 [15 May 1857] (Berg 1857).

*Myrcia laevigata* O.Berg, *Fl. Bras. [Martius]* 14 (1): 175 [15 May 1857] (Berg 1857).

*Myrcia laevigata* var. *brunnea* O.Berg, *Fl. Bras. [Martius]* 14 (1): 175 [15 May 1857] (Berg 1857).
*Myrcia laevigata* var. canescens O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 175 [15 May 1857] (Berg 1857).

*Myrcia catharinae* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 176 [15 May 1857] (Berg 1857).

*Myrcia corcovadensis* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 177 [15 May 1857] (Berg 1857).

*Myrcia opaca* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 177 [15 May 1857] (Berg 1857).

*Myrcia opaca* var. angustifolia O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 177 [15 May 1857] (Berg 1857).

*Myrcia opaca* var. latifolia O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 177 [15 May 1857] (Berg 1857).

*Myrcia ciarensis* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 178 [15 May 1857] (Berg 1857).

*Myrcia nitens* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 178 [15 May 1857] (Berg 1857).

*Myrcia sericiflora* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 178 [15 May 1857] (Berg 1857). — *Myrcia rostrata* f. sericiflora (O.Berg) D.Legrand, *Fl. Ilustr. Catar. 1* (Mirt.): 243 (Legrand 1969).

*Myrcia riparia* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 179 [15 May 1857] (Berg 1857).

*Myrcia schuechiana* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 181 [15 May 1857] (Berg 1857), “*Schüchiana*”.

*Myrcia mikaniana* var. angustifolia O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 182 [15 May 1857] (Berg 1857).

*Myrcia mikaniana* var. latifolia O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 182 [15 May 1857] (Berg 1857).

*Myrcia velutina* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 182 [15 May 1857] (Berg 1857).

*Myrcia communis* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 183 [15 May 1857] (Berg 1857). — *Myrcia rostrata* f. communis (O.Berg) D.Legrand, *Fl. Ilustr. Catar. 1* (Mirt.): 243 (Legrand 1969).

*Myrcia communis* var. glabrata O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 183 [15 May 1857] (Berg 1857).

*Myrcia velutina* var. canescens O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 183 [15 May 1857] (Berg 1857).

*Myrcia velutina* var. ochracea O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 183 [15 May 1857] (Berg 1857).

*Myrcia communis* var. latifolia O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 183 [15 May 1857] (Berg 1857).

*Myrcia gardneriana* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 184 [15 May 1857] (Berg 1857).

*Myrcia impressa* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 184 [15 May 1857] (Berg 1857).

*Myrcia ypanemensis* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 186 [15 May 1857] (Berg 1857).

*Myrcia barrensis* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 187 [15 May 1857] (Berg 1857).

*Myrcia hayneana* var. paraensis O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 187 [15 May 1857] (Berg 1857).

*Myrcia negrensis* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 187 [15 May 1857] (Berg 1857).

*Myrcia acutiloba* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 189 [15 May 1857] (Berg 1857).

*Myrcia sellowiana* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 197 [15 May 1857] (Berg 1857).

*Myrcia sellowiana* var. bullata O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 197 [15 May 1857] (Berg 1857).

*Myrcia sellowiana* var. costata O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 197 [15 May 1857] (Berg 1857).

*Myrcia superba* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 198 [15 May 1857] (Berg 1857).

*Myrcia saxicola* var. grandifolia O.Berg, *Linnaea* 29 (2): 219 [June 1858] (Berg 1858).

*Myrcia sartoriana* O.Berg, *Linnaea* 29 (2): 220 [June 1858] (Berg 1858).

*Myrcia langsdorffii* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 562 [15 Jan. 1859] (Berg 1859).

*Myrcia klotzschiana* var. impellucida O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 563 [15 Jan. 1859] (Berg 1859).

*Myrcia gracilis* var. sessiliflora O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 564 [15 Jan. 1859] (Berg 1859).

*Myrcia tingens* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 564 [15 Jan. 1859] (Berg 1859).

*Myrcia riedeliana* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 565 [15 Jan. 1859] (Berg 1859).

*Myrcia splendens* var. robustior Kuntze, *Revis. Gen. Pl.* 1: 241 [5 Nov. 1891] (Kuntze 1891).

*Myrcia augustana* Kiaersk., * Enum. Myrt. Bras.* 52 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Myrcia guajavifolia* f. grandifolia Kiaersk., * Enum. Myrt. Bras.* 54 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Myrcia melanosticta* Kiaersk., * Enum. Myrt. Bras.* 54 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Myrcia oxyoentophylla* Kiaersk., * Enum. Myrt. Bras.* 57 [Oct.-Nov. 1893] (Kiaerskov 1893), “*oxyoëntophylla*”.

*Myrcia martinicensis* Krug & Urb., *Bot. Jahrb. Syst.* 19 (5): 586 [26 Feb. 1895] (Krug & Urban 1895).

*Myrcia dictyoneura* Diels, *Bot. Jahrb. Syst.* 37 (5): 594 [30 Oct. 1906] (Diels 1906).

*Myrcia brachylopadia* Diels, *Bot. Jahrb. Syst.* 37 (5): 595 [30 Oct. 1906] (Diels 1906).

*Myrcia lamprosericea* Diels, *Bot. Jahrb. Syst.* 37 (5): 596 [30 Oct. 1906] (Diels 1906).

*Myrcia coroicensis* Rusby, *Bull. New York Bot. Gard.* 4 (14): 354 [7 Dec. 1907] (Rusby 1907).

*Myrcia luetzelburgii* Burret ex Luetzelb., Estud. Bot. Nordeste Braz. 3: 201 (Luetzelburg 1923), *nom. nud.*
*Myrcia aguitensis* Gleason, *Bull. Torrey Bot. Club* 58 (7): 409 [Oct. 1931] (Gleason 1931).

*Myrcia compressa* Gleason, *Bull. Torrey Bot. Club* 58 (7): 410 [Oct. 1931] (Gleason 1931).

*Myrcia longicaudata* Lundell, *Amer. Midl. Naturalist* 29 (2): 481 [5 Apr. 1943] (Lundell 1943).

*Myrcia schippii* Lundell, *Amer. Midl. Naturalist* 29 (2): 482 [5 Apr. 1943] (Lundell 1943).

*Myrcia splendens* var. *chrysocoma* McVaugh, *Fieldiana*, Bot. 29 (3): 193 [30 Nov. 1956] (McVaugh 1956).

*Myrcia belizensis* Lundell, *Wrightia* 2 (4): 213 (Lundell 1961).

*Myrcia splendens* var. *guantanamana* Borhidi & O.Muñiz, Bot. Közlem. 64 (1): 20 (Borhidi & Muñiz 1977).

*Myrcia rostrata* f. *flexuosa* Soares-Silva, *Bradea* 8 (48): 323 [22 Oct. 2002] (Soares-Silva 2002).

NOTES. — *Aulomyrcia costata* O.Berg is not based on *Myrcia costata* DC., which is treated separately by Berg in the same work (1857: 157). The type of *Myrtus stoupyi* Spreng. (i.e. that of *Myrtus bracteolaris* Poir.) was given to "citoyen" Lamarck by "citoyen" Stoupy (Poiret 1798: 411), hence the automatic correction of "stoupi".

VERNACULAR NAMES. — Pa: kagegut, karegut • Ka: kumete, kumeti, meli lan • Wp: ināmu pita, pakea'i, pila lea, pila lea u, pila lea wate'e, wila imi'i wu, wila umi'i wu • Nt: baaka bee sii, bee sii • Br: frutinheira, guamirim, murta-grande, murta-parida.

HERBARIUM DATA (FG). — 94 collections at CAY. Sel. exs.: *J.B. Leblond* 114 (original material of *Eugenia fallax*: G).

INVENTORY DATA (FG). — 25 trees in 20 plots; $F_{max} < 1\%$; $dbh_{inv} = 24.7$ cm.

[1308] *Myrcia subobliqua* (Benth.) Nied.

*Nat. Pflanzenfam. [Engler & Prantl]* 3 (7): 76 [Jan. 1893] (Niedenzu 1893). — *Eugenia subobliqua* Benth., *J. Bot. [Hooker]* 2: 322 (Bentham 1840). — *Aulomyrcia subobliqua* (Benth.) O.Berg, *Linnaea* 27 (1): 57 ["1854" publ. Nov. 1855] (Berg 1855).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Wp: pakea'i sī, pila lea.

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: *P. Grenand* 1328.

INVENTORY DATA (FG). — 2 trees in 1 plot; $F_{max} < 1\%$; $dbh_{inv} = 12.9$ cm.

[1309] *Myrcia subsessilis* O.Berg

*Linnaea* 31: 251 (Berg 1861-1862).

*Myrcia subsessilis* var. *subcordata* O.Berg, *Linnaea* 31: 251 (Berg 1861-1862).

*Myrcia subsessilis* var. *ovalis* O.Berg, *Linnaea* 31: 252 (Berg 1861-1862).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *J.-J. de Granville* 3327.

SIZE. — Brazil, Amapá. *S.A. Mori* 17488 (MO), 20 m × 30 cm.

[1310] *Myrcia sylvatica* (G.Mey.) DC.

Prodr. [*A. P. de Candolle*] 3: 244 [mid Mar. 1828] (Candolle 1828). — *Myrtus sylvatica* G.Mey., *Prim. Fl. Esseq.* 191 [Nov. 1818] (Meyer 1818), "Myrthus". — *Myrcia ambigua* var. *sylvatica* (G.Mey.) O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 180 [15 May 1857] (Berg 1857).

*Myrtus lucida* L., *Syst. Nat.*, ed. 10, 2: 1056 [7 June 1759] (Linnaeus 1759).

*Myrcia ambigua* DC., Prodr. [*A. P. de Candolle*] 3: 252 [mid Mar. 1828] (Candolle 1828).

*Myrcia ambigua* var. *pauciflora* DC., Prodr. [*A. P. de Candolle*] 3: 252 [mid Mar. 1828] (Candolle 1828).

*Myrcia ambigua* var. *dives* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 180 [15 May 1857] (Berg 1857).

*Myrcia ambigua* var. *multiflora* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 180 [15 May 1857] (Berg 1857).

*Myrcia ambigua* var. *latifolia* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 181 [15 May 1857] (Berg 1857).

*Myrcia ambigua* var. *rostrata* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 181 [15 May 1857] (Berg 1857).

VERNACULAR NAMES. — Ka: pikililan • Wp: kasila, pila lea, pila lea sili.

HERBARIUM DATA (FG). — 49 collections at CAY. Sel. exs.: *R.A.A. Oldeman* 3064.

SIZE. — Brazil, Amazonas. *M. Pacheco et al.* 43 (MO), 14 m × 15.4 cm.

[1311] *Myrcia tomentosa* (Aubl.) DC.

Prodr. [*A. P. de Candolle*] 3: 245 [mid Mar. 1828] (Candolle 1828). — *Eugenia tomentosa* Aubl., *Hist. Pl. Guiane* 1: 504 [Jun.-Dec. 1775] (Aublet 1775). — *Myrtus aubletii* Spreng., *Syst. Veg. [Sprengel]* 2: 486 [Jan.-May 1825] (Sprengel 1825), *nom. illeg. superfl.* (based on *Eugenia tomentosa*). — *Aguava tomentosa* (Aubl.) Raf., *Sylva Tellur.*: 107 (Rafinesque 1838). — *Cumetea tomentosa* (Aubl.) Raf., *Sylva Tellur.*: 106 (Rafinesque 1838). — *Aulomyrcia tomentosa* (Aubl.) Amshoff, *Recueil Trav. Bot. Néerl.* 39: 153 (Amshoff 1942).

*Myrcia hirtiflora* DC., Prodr. [*A. P. de Candolle*] 3: 249 [mid Mar. 1828] (Candolle 1828). — *Aulomyrcia hirtiflora* (DC.) O.Berg, *Linnaea* 27 (1): 40 ["1854" publ. Nov. 1855] (Berg 1855).

*Myrcia curatellifolia* DC., Prodr. [*A. P. de Candolle*] 3: 253 [mid Mar. 1828] (Candolle 1828), "*curatellaefolia*". — *Myrtus curatellifolia* Mart. ex DC., Prodr. [*A. P. de Candolle*] 3: 253 [mid Mar. 1828] (Candolle 1828), "*curatellaefolia*", *nom. nud. pro syn.* — *Aulomyrcia curatellifolia* (DC.) O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 95 [15 May 1857] (Berg 1857), "*curatellaefolia*".

*Myrcia prunifolia* DC., Prodr. [*A. P. de Candolle*] 3: 253 [mid Mar. 1828] (Candolle 1828). — *Aulomyrcia prunifolia* (DC.) O.Berg, *Linnaea* 27 (1): 55 ["1854" publ. Nov. 1855] (Berg 1855).

*Myrcia pilosa* DC., Prodr. [*A. P. de Candolle*] 3: 253 [mid Mar. 1828] (Candolle 1828). — *Myrtus pilosa* Mart. ex DC., Prodr. [*A. P. de Candolle*] 3: 253 [mid Mar. 1828] (Candolle 1828), *nom. nud. pro syn.*

*Myrcia prunifolia* var. *angustior* DC., Prodr. [*A. P. de Candolle*] 3: 253 [mid Mar. 1828] (Candolle 1828).
*Myrcia prunifolia* var. *obovata* DC., *Prodr.* [A. P. de Candolle] 3:253
[mid Mar. 1828] (Candolle 1828).

*Myrcia prunifolia* var. *ovata* DC., *Prodr.* [A. P. de Candolle] 3:253
[mid Mar. 1828] (Candolle 1828).

*Myrcia puberula* Cambess., *Fl. Bras. Merid.* [*A. St.-Hil.*] (quarto ed.)
2 (18): 316 ["1829" publ. 25 Feb. 1832] (Cambessèdes 1832).
— *Aulomyrcia puberula* (Cambess.) O.Berg, *Fl. Bras.* [*Martius*] 14
(1): 94 [15 May 1857] (Berg 1857). — *Myrcianthes puberula*
(Cambess.) Mattos, *Loefgrenia* 128: 1 [Apr. 2009] (Mattos 2009).

*Myrcia capivarhyensis* Cambess., *Fl. Bras. Merid.* [*A. St.-Hil.*] (quarto ed.) 2 (19): 328 ["1829" publ. 27 Oct. 1832] (Cambessèdes 1832). — *Aulomyrcia capivarhyensis* (Cambess.) O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 63 [15 May 1857] (Berg 1857).

*Myrcia floribunda* Miq., *Linnaea* 22: 534 (Miquel 1849).

*Aulomyrcia aureolanata* O.Berg, *Linnaea* 27 (1): 52 ["1854" publ. Nov. 1855] (Berg 1855), "aureo-lanata".

*Aulomyrcia alloiota* O.Berg, *Linnaea* 27 (1): 54 ["1854" publ. Nov. 1855] (Berg 1855). — *Myrcia alloiota* (O.Berg) Kiaersk., * Enum. Myrt. Bras.* 70 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Aulomyrcia rosalans* O.Berg, *Linnaea* 27 (1): 54 ["1854" publ. Nov. 1855] (Berg 1855). — *Myrcia rosalans* (O.Berg) Kiaersk., * Enum. Myrt. Bras.* 72 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Aulomyrcia ottonis* O.Berg, *Linnaea* 27 (1): 55 ["1854" publ. Nov. 1855] (Berg 1855).

*Aulomyrcia confusa* O.Berg, *Linnaea* 27 (1): 56 ["1854" publ. Nov. 1855] (Berg 1855).

*Calyptromyrcia venosa* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 57 [15 May 1857] (Berg 1857). — *Myrcia venosa* (O.Berg) Mattos, *Loefgrenia* 128: 3 [Apr. 2009] (Mattos 2009).

*Aulomyrcia lancea* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 88 [15 May 1857] (Berg 1857). — *Myrcia lancea* (O.Berg) Mattos, *Arq. Bot. Estado São Paulo* 4 (2): 62 (Mattos 1966).

*Aulomyrcia spathulata* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 91 [15 May 1857] (Berg 1857). — *Myrcia spathulata* (O.Berg) Kiaersk., * Enum. Myrt. Bras.* 72 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Aulomyrcia alloiota* var. *cuneata* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 92 [15 May 1857] (Berg 1857).

*Aulomyrcia alloiota* var. *obovata* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 92 [15 May 1857] (Berg 1857). — *Myrcia alloiota* var. *obovata* (O.Berg) Kiaersk., * Enum. Myrt. Bras.* 70 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Aulomyrcia alloiota* var. *ovalis* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 92 [15 May 1857] (Berg 1857).

*Aulomyrcia alloiota* var. *pyramidalis* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 92 [15 May 1857] (Berg 1857).

*Aulomyrcia alloiota* var. *subcordata* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 92 [15 May 1857] (Berg 1857). — *Myrcia alloiota* var. *subcordata* (O.Berg) N.Silveira, *Loefgrenia* 86: 1 (Silveira 1985).

*Aulomyrcia leucantha* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 93 [15 May 1857] (Berg 1857). — *Myrcia leucantha* (O.Berg) N.Silveira, *Roessleria* 7 (1): 66 (Silveira 1985).

*Aulomyrcia longipes* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 94 [15 May 1857] (Berg 1857). — *Myrcia longipes* (O.Berg) Kiaersk., * Enum. Myrt. Bras.* 70 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Aulomyrcia longipes* var. *latifolia* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 94 [15 May 1857] (Berg 1857).

*Aulomyrcia longipes* var. *obovata* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 94 [15 May 1857] (Berg 1857). — *Myrcia longipes* f. *obovata* (O.Berg) Kiaersk., * Enum. Myrt. Bras.* 70 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Aulomyrcia longipes* var. *spathulata* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 94 [15 May 1857] (Berg 1857).

*Aulomyrcia curatellifolia* var. *grandifolia* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 95 [15 May 1857] (Berg 1857), "curatellaefolia". — *Myrcia curatellifolia* var. *grandifolia* (O.Berg) Kiaersk., * Enum. Myrt. Bras.* 70 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Aulomyrcia curatellifolia* var. *parvifolia* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 95 [15 May 1857] (Berg 1857), "curatellaefolia". — *Myrcia curatellifolia* var. *parvifolia* (O.Berg) Kiaersk., * Enum. Myrt. Bras.* 70 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Aulomyrcia prunifolia* var. *brevipes* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 95 [15 May 1857] (Berg 1857).

*Aulomyrcia prunifolia* var. *longipes* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 96 [15 May 1857] (Berg 1857).

*Aulomyrcia lanuginosa* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 102 [15 May 1857] (Berg 1857). — *Myrcia lanuginosa* (O.Berg) Nied., *Nat. Pflanzenfam.* [*Engler & Prantl*] 3 (7): 76 [Jan. 1893] (Niedenzu 1893), *nom. illeg. hom., non* O.Berg (1857).

*Myrcia piauhiensis* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 196 [15 May 1857] (Berg 1857).

*Aulomyrcia micrantha* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 517 [1 Feb. 1858] (Berg 1858).

*Aulomyrcia curatellifolia* var. *australis* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 547 [15 Jan. 1859] (Berg 1859), "curatellaefolia".

*Aulomyrcia lanuginosa* var. *pyramidata* O.Berg, *Fl. Bras.* [*Martius*] 14 (1): 548 [15 Jan. 1859] (Berg 1859).

*Myrcia marahanensis* Kiaersk., * Enum. Myrt. Bras.* 71 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Myrcia membranacea* Kiaersk., * Enum. Myrt. Bras.* 71 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Myrcia rhodeosepala* Kiaersk., * Enum. Myrt. Bras.* 75 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Myrcia verruculata* S.Moore, *Trans. Linn. Soc. London, Bot.* 4 (3): 355 ["1894-96" publ. Dec. 1895] (Moore 1895).

*Eugenia vallis* Standl., *Ann. Missouri Bot. Gard.* 27: 323 (Standley 1940).

*HERBARIUM DATA* (FG). — 20 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000953642]).

*SIZE.* — Up to 10 m tall (Holst 2003).
[1312] *Myrcia* sp. A

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *J.-J. de Granville 14611*.

INVENTORY DATA (FG). — 3 trees in 3 plots; $F_{max} < 1\%$; $dbh_{inv} = 11.6$ cm.

[1313] *Myrcia* sp. B

NOTE. — Collections at CAY are provisionally classified by B. K. Holst as "*Marlierea* FG-2"

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier 2329*, $dbh = 16.9$ cm.

Genus *Myrcianthes* O.Berg

[1314] *Myrcianthes prodigiosa* McVaugh

*Fieldiana*, Bot. 29 (8): 492 [31 May 1963] (McVaugh 1963).

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Barthélémy 3687*.

INVENTORY DATA (FG). — 14 trees in 6 plots; $F_{max} < 1\%$; $dbh_{inv} = 38.5$ cm.

Genus *Myrciaria* O.Berg

[1315] *Myrciaria floribunda* (H.West ex Willd.) O.Berg

*Linnaea* 27 (2-3): 330 ["1854" publ. Jan. 1856] (Berg 1856). — *Eugenia floribunda* H.West ex Willd., *Sp. Pl.*, ed. 4 2 (2): 960 [Dec. 1799] (Willdenow 1799). — *Myrtus floribunda* (H.West ex Willd.) Spreng., *Syst. Veg. [Sprengel]* 2: 484 [Jan.-May 1825] (Sprengel 1825).

*Myrtus micrantha* Nees & Mart., *Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur.* 12: 51 (Nees & Martius 1824), *nom. illeg. hom., non* Kunth (1823) [synonym of *Eugenia monticola* (Sw.) DC.]

*Eugenia maximiliana* DC., *Prodr. [A. P. de Candolle]* 3: 270 [mid Mar. 1828] (Candolle 1828). — *Myrciaria maximiliana* (DC.) O.Berg, *Linnaea* 27 (2-3): 329 ["1854" publ. Jan. 1856] (Berg 1856).

*Eugenia ciliolata* Cambess., *Fl. Bras. Merid. [A. St.-Hil.]* (quarto ed.) 2 (19): 344 ["1829" publ. 27 Oct. 1832] (Cambessèdes 1832). — *Myrciaria ciliolata* (Cambess.) O.Berg, *Linnaea* 27 (2-3): 327 ["1854" publ. Jan. 1856] (Berg 1856). — *Paramyrciaria ciliolata* (Cambess.) Rotman, *Bol. Soc. Argent. Bot.* 24 (3-4): 418 (Rotman 1986).

*Eugenia salzmannii* Benth., *J. Bot. [Hooker]* 2: 319 (Bentham 1840), "Salzmanni". — *Myrciaria salzmannii* (Benth.) O.Berg, *Linnaea* 27 (2-3): 331 ["1854" publ. Jan. 1856] (Berg 1856), "Salzmanni".

*Myrtus verticillata* Salzm. ex Benth., *J. Bot. [Hooker]* 2: 320 (Bentham 1840), *nom. illeg. hom., non* Vell. (Vellozo 1829), *nom. nud.*

*Eugenia protracta* Steud., *Flora* 26 (45): 762 [17 Dec. 1843] (Steudel 1843). — *Myrciaria protracta* (Steud.) O.Berg, *Linnaea* 27 (2-3): 330 ["1854" publ. Jan. 1856] (Berg 1856).

*Myrciaria uliginosa* O.Berg, *Linnaea* 27 (2-3): 329 ["1854" publ. Jan. 1856] (Berg 1856).

*Myrciaria verticillata* O.Berg, *Linnaea* 27 (2-3): 332 ["1854" publ. Jan. 1856] (Berg 1856).

*Myrciaria leucadendron* O.Berg, *Fl. Bras. [Martius]* 14 (1): 364 [15 May 1857] (Berg 1857).

*Myrciaria leucophloea* O.Berg, *Fl. Bras. [Martius]* 14 (1): 370 [15 May 1857] (Berg 1857). — *Eugenia leucophloea* (O.Berg) Kiaersk., * Enum. Myrt. Bras.* 183 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Myrciaria leucophloea* var. *conferta* O.Berg, *Fl. Bras. [Martius]* 14 (1): 370 [15 May 1857] (Berg 1857).

*Myrciaria leucophloea* var. *laxa* O.Berg, *Fl. Bras. [Martius]* 14 (1): 370 [15 May 1857] (Berg 1857).

*Myrciaria splendens* O.Berg, *Fl. Bras. [Martius]* 14 (1): 371 [15 May 1857] (Berg 1857).

*Myrciaria tenuiramis* O.Berg, *Fl. Bras. [Martius]* 14 (1): 371 [15 May 1857] (Berg 1857).

*Myrciaria maragnanensis* O.Berg, *Fl. Bras. [Martius]* 14 (1): 372 [15 May 1857] (Berg 1857). — *Myrciaria maranhensis* O.Berg, *Fl. Bras. [Martius]* 14 (1): 652 [15 Jan. 1859] (Berg 1859), sphalm. — *Eugenia maranhensis* (O.Berg) Kiaersk., * Enum. Myrt. Bras.* 183 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Myrciaria tolpantha* O.Berg, *Fl. Bras. [Martius]* 14 (1): 372 [15 May 1857] (Berg 1857).

*Myrciaria tolpantha* var. *angustifolia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 372 [15 May 1857] (Berg 1857).

*Myrciaria tolpantha* var. *latifolia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 372 [15 May 1857] (Berg 1857).

*Myrciaria axillaris* O.Berg, *Fl. Bras. [Martius]* 14 (1): 373 [15 May 1857] (Berg 1857).

*Myrciaria prasina* O.Berg, *Fl. Bras. [Martius]* 14 (1): 373 [15 May 1857] (Berg 1857).

*Myrciaria schuechiana* O.Berg, *Fl. Bras. [Martius]* 14 (1): 373 [15 May 1857] (Berg 1857), "*schüchiana*".

*Myrciaria amazonica* O.Berg, *Fl. Bras. [Martius]* 14 (1): 374 [15 May 1857] (Berg 1857).

*Myrciaria sellowiana* O.Berg, *Fl. Bras. [Martius]* 14 (1): 374 [15 May 1857] (Berg 1857).

*Myrciaria ferruginea* O.Berg, *Fl. Bras. [Martius]* 14 (1): 597 [15 Jan. 1859] (Berg 1859).

*Myrciaria longipes* O.Berg, *Fl. Bras. [Martius]* 14 (1): 599 [15 Jan. 1859] (Berg 1859).

*Myrciaria schuechiana* var. *deflexa* O.Berg, *Fl. Bras. [Martius]* 14 (1): 599 [15 Jan. 1859] (Berg 1859), "*schüchiana*".

*Myrciaria schuechiana* var. *latifolia* O.Berg, *Fl. Bras. [Martius]* 14 (1): 599 [15 Jan. 1859] (Berg 1859), "*schüchiana*".

*Myrciaria longipes* var. *opaca* O.Berg, *Fl. Bras. [Martius]* 14 (1): 600 [15 Jan. 1859] (Berg 1859).

*Myrciaria longipes* var. *pellucida* O.Berg, *Fl. Bras. [Martius]* 14 (1): 600 [15 Jan. 1859] (Berg 1859).
*Marlierea brachymischa* Kiaersk., * Enum. Myrt. Bras. * 47 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Eugenia leucophloea* var. *warmingiana* Kiaersk., * Enum. Myrt. Bras. * 183 [Oct.-Nov. 1893] (Kiaerskov 1893). — *Myrciaria ciliolata* var. *warmingiana* (Kiaersk.) Mattos, *Loefgrenia* 78: 3 (Mattos 1983). — *Myrciaria leucophloea* var. *warmingiana* (Kiaersk.) Mattos, *Loefgrenia* 64: 3 (Mattos 1975).

*Eugenia polyneura* Urb., * Symb. Antill. [Urban] * 5 (3): 446 [20 May 1908] (Urban 1908), * nom. illeg. hom. , non Koord. & Valeton* (Koorders & Valeton 1900).

*Eugenia pycnoneura* Urb., * Symb. Antill. [Urban] * 6 (1): 25 [15 July 1909] (Urban 1909).

*Plinia acutissima* Urb., * Ark. Bot. * 22A(10): 25 (Urban 1929).

*Eugenia oneillii* Lundell, *Bull. Torrey Bot. Club* 64 (8): 555 [Nov. 1937] (Lundell 1937), "O'Neillii". — *Myrciaria oneillii* (Lundell) I.M.Johnst., *Sargentia* 8: 228 (Johnston 1949), "O'Neillii".

*Myrciaria arborea* D.Legrand, *Sellowia* 13: 328 (Legrand 1961).

*Siphoneugena cantareirae* Mattos, * Ci. & Cult. * 19 (2): 332 (Mattos 1967).

*Myrciaria longicaudata* Lundell, * Wrightia* 4 (4): 144 (Lundell 1970).

*Siphoneugena micrantha* Kausel, * Lilloa* 36 (6): 111 (Kausel 1972).

*Myrciaria mexicana* Lundell, * Wrightia* 5 (2): 44 (Lundell 1974).

*Myrciaria arborea* var. *rostrata* Mattos, *Loefgrenia* 66: 3 (Mattos 1975).

**NOTE.** — Excluded synonyms: *Calypranthes floribunda* Blume and *Caryophyllus floribundus* (Blume) Blume (from Java).

**VERNACULAR NAMES.** — Pa: awao, awau, awau-seinô • Ka: asaikiabe, seityape, tamunen seityape • Wp: kumesi • Cr: gouyavié-montangn • Br: goiaba-do-mato, goiabinha.

**HERBARIUM DATA (FG).** — 55 collections at CAY. Sel. exs.: *J.-F Molino & D. Sabatier* 2218.

**INVENTORY DATA (FG).** — 39 trees in 32 plots; $F_{max} < 1\%$; $dbh_{inv} = 33.3$ cm.

[1316] *Myrciaria vismiifolia* (Benth.) O.Berg

*Linnaea* 27 (2-3): 336 ["1854" publ. Jan. 1856] (Berg 1856), "vismeaefolia". — *Eugenia vismiifolia* Benth., *J. Bot. [Hooker]* 2: 320 (Bentham 1840), "vismeaefolia".

**NOTE.** — The specific epithet refers to the genus *Vismia* Vand., hence it is corrected to *vismiifolia*.

**VERNACULAR NAMES.** — Wn: hapi, pëneju, takamalakma.

**HERBARIUM DATA (FG).** — 10 collections at CAY. Sel. exs.: *Service Forestier* 7911, $dbh = 20$ cm.

Genus *Plinia* L.

[1317] *Plinia rivularis* (Cambess.) Rotman

*Bol. Soc. Argent. Bot.* 24 (1-2): 195 (Rotman 1985). — *Eugenia rivularis* Cambess., *Fl. Bras. Merid. [A. St.-Hil.]* (quarto ed.) 2 (19):

337 ["1829" publ. 27 Oct. 1832] (Cambessèdes 1832). — *Myrciaria rivularis* (Cambess.) O.Berg, *Linnaea* 27 (2-3): 320 ["1854" publ. Jan. 1856] (Berg 1856).

*Myrciaria hagendorffii* O.Berg, *Fl. Bras. [Martius]* 14 (1): 360 [15 May 1857] (Berg 1857). — *Eugenia hagendorffii* (O.Berg) Kiaersk., * Enum. Myrt. Bras.* 179 [Oct.-Nov. 1893] (Kiaerskov 1893).

*Myrcia silvatica* Barb.Rodr. ex Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 7: 803 (Chodat & Hassler 1907), *nom. nud.*

*Eugenia variifolia* Barb.Rodr. ex Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 7: 806 (Chodat & Hassler 1907), *nom. nud.*

*Myrcia granulata* R.O.Williams, *Fl. Trinidad & Tobago* 1 (6): 340 [Oct. 1934] (Williams 1934).

*Myrciaria baporeti* D.Legrand, *Anales Mus. Hist. Nat. Montevideo* 4 (11): 63 (Legrand 1936). — *Myrciariopsis baporeti* (D.Legrand) Kausel, *Ark. Bot., n.s.*, 3: 509 (Kausel 1956). — *Siphoneugena baporeti* (D.Legrand) Kausel, *Lilloa* 32: 345 (Kausel 1966). — *Myrciaria rivularis* var. *baporeti* (D.Legrand) D.Legrand, *Fl. Ilustr. Catar. (Mirt.)*: 768 (Legrand 1978). — *Plinia baporeti* (D.Legrand) Rotman, *Darwiniana* 24 (1-4): 169 (Rotman 1982).

*Siphoneugena legrandii* Mattos & N.Silveira, *Loefgrenia* 87: 2 (Mattos & Silveira 1985).

**VERNACULAR NAMES.** — Pa: mpitit-kamwi (for Plinia sp.) • Br: guamirim, jaboticaba-rana.

**HERBARIUM DATA (FG).** — A single collection, *C. Baraloto et al.* 8056.

**SIZE.** — Up to 18 m tall (Holst 2003).

Genus *Psidium* L.

[1318] *Psidium acutangulum* DC.

*Prodr. [A. P. de Candolle]* 3: 233 [mid Mar. 1828] (Candolle 1828). — *Guajava acutangula* (DC.) Kuntze, *Revis. Gen. Pl.* 1: 239 [5 Nov. 1891] (Kuntze 1891).

*Psidium acutangulum* var. *acidum* DC., *Prodr. [A. P. de Candolle]* 3: 233 [mid Mar. 1828] (Candolle 1828). — *Britoa acida* (DC.) O.Berg, *Linnaea* 27 (4): 436 ["1854" publ. Feb. 1856] (Berg 1856). — *Psidium acidum* Mart. ex O.Berg, *Linnaea* 27 (4): 436 ["1854" publ. Feb. 1856] (Berg 1856), *nom. nud. pro syn.* — *Psidium acidum* (DC.) Landrum, *Brittonia* 68 (4): 411 [epubl. 28 Apr. 2016] (Landrum 2016).

*Psidium acutangulum* var. *crassirame* O.Berg, *Fl. Bras. [Martius]* 14 (1): 409 [15 May 1857] (Berg 1857).

*Psidium acutangulum* var. *tenuirame* O.Berg, *Fl. Bras. [Martius]* 14 (1): 409 [15 May 1857] (Berg 1857).

*Psidium grandiflorum* Ruiz & Pav., *Anales Inst. Bot. Cavanilles* 15: 228 (Ruiz & Pavón 1958), *nom. illeg. hom., non* Aubl. (Aublet 1775) [synobym of *Campomanesia grandiflora* (Aubl.) Sagot)].

*Psidium persoonii* McVaugh, *Mem. New York Bot. Gard.* 18 (2): 255 (McVaugh 1969).

*Psidium acutangulum* var. *oblongatum* Mattos, *Loefgrenia* 94: 12 (Mattos 1989), "*oblongata*".

**NOTE.** — Incipiently domesticated by pre-Columbian Amerindians (Clement 1999; Levis *et al.* 2017).
VERNACULAR NAMES. — Te: ala'a • Wp: alali • Wn: takamalaimë • Nt: liba gobaya • Cr: gouyave-so • Br: araçá-pera.

HERBARIUM DATA (FG). — 24 collections at CAY. Sel. exs.: *M. Fleury* 2176.

SIZE. — Up to 10 m tall (Holst 2003).

[1319] *Psidium oligospermum* Mart. ex DC.

*Prodr.* [A. P. de Candolle] 3: 236 [mid Mar. 1828] (Candolle 1828). — *Guajava oligosperma* (Mart. ex DC.) Kuntze, *Revis. Gen. Pl.* 1: 239 [5 Nov. 1891] (Kuntze 1891). — *Mitropsidium oligospermum* (Mart. ex DC.) Burret, *Notizbl. Bot. Gart. Berlin-Dahlem* 15: 486 [30 Mar. 1941] (Burret 1941).

*Calyptranthes eugenioides* Cambess., *Fl. Bras. Merid.* [A. St.-Hil.] (quarto ed.) 2 (19): 370 [“1829” publ. 27 Oct. 1832) (Cambessèdes 1832), “Eugenioides”. — *Mitranthes eugenioides* (Cambess.) O.Berg, *Linnaea* 27 (2-3): 317 [“1854” publ. Jan. 1856] (Berg 1856). — *Chytraculia eugenioides* (Cambess.) Kuntze, *Revis. Gen. Pl.* 1: 238 [5 Nov. 1891] (Kuntze 1891). — *Psidium eugenioides* (Cambess.) Nied., *Nat. Pflanzenfam. [Engler & Prantl]* 3 (7): 69 [Jan. 1893] (Niedenzu 1893). — *Mitropsidium eugenioides* (Cambess.) Burret, *Notizbl. Bot. Gart. Berlin-Dahlem* 15: 489 [30 Mar. 1941] (Burret 1941).

*Psidium galapageium* Hook.f., *Trans. Linn. Soc. London* 20 (2): 224 (Hooker 1847).

*Psidium ciliatum* O.Berg, *Linnaea* 27 (2-3): 353 [“1854” publ. Jan. 1856] (Berg 1856), *nom. illeg. hom.*, *non* Benth. (Bentham 1840) [synonym of *P. salutare* (Kunth) O.Berg].

*Mitranthes gardneriana* O.Berg, *Fl. Bras.* [*Martius* 14 (1): 354 [15 May 1857] (Berg 1857). — *Chytraculia gardneriana* (O.Berg) Kuntze, *Revis. Gen. Pl.* 1: 238 [5 Nov. 1891] (Kuntze 1891). — *Mitropsidium gardnerianum* (O.Berg) Burret, *Notizbl. Bot. Gart. Berlin-Dahlem* 15: 487 [30 Mar. 1941] (Burret 1941).

*Mitranthes eugenioides* var. *oblongifolia* O.Berg, *Fl. Bras.* [*Martius* 14 (1): 355 [15 May 1857] (Berg 1857), *nom. illeg.* (species basionym in synonymy).

*Mitranthes eugenioides* var. *ovata* O.Berg, *Fl. Bras.* [*Martius* 14 (1): 355 [15 May 1857] (Berg 1857).

*Mitranthes sartoriana* O.Berg, *Linnaea* 29 (2): 248 [June 1858] (Berg 1858). — *Chytraculia sartoriana* (O.Berg) Kuntze, *Revis. Gen. Pl.* 1: 238 [5 Nov. 1891] (Kuntze 1891). — *Psidium sartorianum* (O.Berg) Nied., *Nat. Pflanzenfam. [Engler & Prantl]* 3 (7): 69 [Jan. 1893] (Niedenzu 1893). — *Calyptropsis sartorianum* (O.Berg) Krug & Urb., *Bot. Jahrb. Syst.* 19 (4): 571 [28 Dec. 1894] (Krug & Urban 1894). — *Mitropsidium sartorianum* (O.Berg) Burret, *Notizbl. Bot. Gart. Berlin-Dahlem* 15: 487 [30 Mar. 1941] (Burret 1941).

*Calycorectes protractus* Griseb., *Cat. Pl. Cub. [Grisebach]* 284 [May-Aug. 1866] (Grisebach 1866). — *Psidium protractum* (Griseb.) Lundell, *Wrightia* 5 (3): 70 (Lundell 1974).

*Calyptranthes tonduzii* Donn.Sm., *Bot. Gaz.* 23 (4): 245 [22 Apr. 1897] (Donnell Smith 1897).

*Psidium paucinerve* Urb., *Symb. Antill. [Urban]* 9 (2): 82 [1 Jan. 1923] (Urban 1923). — *Myrtus paucinervia* (Urb.) Landrum, *Canotia* 13: 49 (Landrum 2017), “paucinerve”, *nom. inval. pro syn.*

*Psidium claraense* Urb., *Symb. Antill. [Urban]* 9 (4): 466 [15 Mar. 1928] (Urban 1928). — *Myrtus claraensis* (Urb.) Bisse, *Ciencias (Havana)*, ser. 10, 12: 10 [“1976” publ. 1977] (Bisse 1977).

*Psidium microphyllum* Britton, *Sci. Surv. Porto Rico & Virgin Islands* 6: 555 (Britton 1930).

*Psidium socorrense* I.M.Johnst., *Proc. Calif. Acad. Sci.*, ser. 4, 20: 81 (Johnston 1931).

*Mitropsidium oblanceolatum* Burret, *Notizbl. Bot. Gart. Berlin-Dahlem* 15: 487 [30 Mar. 1941] (Burret 1941).

*Mitropsidium pittieri* Burret, *Notizbl. Bot. Gart. Berlin-Dahlem* 15: 488 [30 Mar. 1941] (Burret 1941).

*Psidium yucatanense* Lundell, *Contr. Univ. Michigan Herb.* 7: 35 (Lundell 1942). — *Psidium sartorianum* var. yucatanense (Lundell) McVaugh, *Fieldiana*, *Bot.* 29 (8): 527 [31 May 1963] (McVaugh 1963).

*Psidium solisii* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 23 (3): 133 [14 Feb. 1944] (Standley 1944).

*Psidium minutiflorum* Amshoff, *Recueil Trav. Bot. Néerl.* 42: 19 (Amshoff 1950).

*Psidium molinae* Amshoff, *Acta Bot. Neerl.* 5 (3): 277 [Nov. 1956] (Amshoff 1956), “Molinae”.

*Psidium galapageium* var. *howellii* D.M.Porter, *Ann. Missouri Bot. Gard.* 55 (3): 370 (Porter 1969).

NOTE. — McVaugh based *Psidium sartorianum* var. yucatanense on *P. yucatanense* Lundell, but argued that the former name should not be treated as a new combination, on the mistaken belief that *P. yucatanense* was a taxonomic synonym of *Calycorectes protractus* Griseb. As *P. yucatanense* was validly published, all requirements were fulfilled for a new combination.

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino* 5029.

SIZE. — Brazil, Pará. *T.C. Plowman et al.* 8730 (MO), 15 m × 30 cm.

[1320] *Myrtaceae* sp. A

NOTE. — Collections at CAY are provisionally classified by B. K. Holst as “*Myrtaceae* FG-2”.

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: *D. Sabatier et al.* 5812.

INVENTORY DATA (FG). — 11 trees in 7 plots; $F_{max} < 1\%$; $dbh_{inv} = 48.5$ cm.

[1321] *Myrtaceae* sp. B

NOTE. — Collections at CAY are provisionally classified by B. K. Holst as “*Myrtaceae* FG-3”.

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 5411.

INVENTORY DATA (FG). — 3 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 10.8$ cm.

[1322] *Myrtaceae* sp. C

HERBARIUM DATA (FG). — A single collection, *D. Sabatier & M.-F. Prévost* 2956.

INVENTORY DATA (FG). — 2 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 15.3$ cm.
Family NYCTAGINACEAE Juss.
Genus *Guapira* Aubl.

[1323] *Guapira eggersiana* (Heimerl) Lundell

*Wrightia* 4 (2): 80 [31 Dec. 1968] (Lundell 1968). — *Pisonia eggersiana* Heimerl, *Bot. Jahrb. Syst.* 21 (5): 627 [12 May 1896] (Heimerl 1896). — *Torrubia eggersiana* (Heimerl) Standl., *Contr. U.S. Natl. Herb.* 18 (3): 100 [11 Feb. 1916] (Standley 1916).

*Pisonia broadwayana* Heimerl, *Repert. Spec. Nov. Regni Veg.* 17: 1 (Heimerl 1921). — *Torrubia broadwayana* (Heimerl) Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 8 (5): 308 (Standley 1931). — *Guapira broadwayana* (Heimerl) Lundell, *Wrightia* 4 (2): 80 [31 Dec. 1968] (Lundell 1968).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Pa: aaku-gawut • Ka: wayamu sasamili • Wp: inimopo'i piyūnga, pikau ki'īy, pikau meyu • Cr: grènn-malenbé • Br: carvoeiro.

HERBARIUM DATA (FG). — 55 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 3833.

INVENTORY DATA (FG). — 55 trees in 22 plots; Fmax = 1%; dbhinv = 51.4 cm.

[1324] *Guapira guianensis* Aubl.

*Hist. Pl. Guiane* 1: 308 [Jun.-Dec. 1775] (Aublet 1775), "*Guapira Guyannensis*" on plate. — *Pisonia guianensis* (Aubl.) R.A.Howard, *J. Arnold Arbor.* 64 (2): 270 [Apr. 1983] (Howard 1983).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — 1 collection from French Guiana: *J.B. Aublet s.n.* (original material P-JU, P00675567]).

SIZE. — Venezuela, Aragua. *O. Huber* 333 (MO), 7 m × 11 cm.

[1325] *Guapira salicifolia* (Heimerl) Lundell

*Wrightia* 4 (2): 83 [31 Dec. 1968] (Lundell 1968). — *Pisonia salicifolia* Heimerl, *Symb. Antill. [Urban]* 7 (2): 216 [15 June 1912] (Heimerl 1912). — *Torrubia salicifolia* (Heimerl) Standl., *Contr. U.S. Natl. Herb.* 18 (3): 101 [11 Feb. 1916] (Standley 1916).

*Pisonia albiflora* Heimerl, *Bull. Misc. Inform. Kew* 1932 (5): 219 [27 June 1932] (Heimerl 1932).

*Torrubia heimerliana* Standl., *Lloydia* 2 (3): 177 (Standley 1939). — *Guapira heimerliana* (Standl.) Lundell, *Wrightia* 4 (2): 81 [31 Dec. 1968] (Lundell 1968).

VERNACULAR NAMES. — Wp: inimopo'i piyūnga • Cr: grènn-malenbé • Br: carvoeiro.

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *P. Grenand* 2107.

INVENTORY DATA (FG). — 1 tree, dbh = 14.6 cm.

[1326] *Guapira* sp. A

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *J.-F. Molino et al.* 2071.

INVENTORY DATA (FG). — 19 trees in 9 plots; Fmax < 1%; dbhinv = 27.6 cm.

Genus *Neea* Ruiz & Pav.

[1327] *Neea constricta* Spruce ex J.A.Schmidt

Fl. Bras. [Martius] 14 (2): 368 [1 May 1872] (Schmidt 1872).

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *S.A. Mori & C. Gracie* 24205.

SIZE. — Up to 15 cm dbh (DeFilipps 2002).

[1328] *Neea floribunda* Poepp. & Endl.
(Fig. 43A)

*Nova genera ac species plantarum* [Poeppig & Endlicher] 2: 46 [Jan.-Sep. 1838] (Poeppig & Endlicher 1838), "*Neeaea*".

*Neea cauliflora* Heimerl, *Nat. Pflanzenfam.* [Engler & Prantl], ed. 2, 16c: 129 (Heimerl 1934).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: mihukat-kamwi, mirukat-kamwi • Ka: sakusaku • Wp: inimopo'i piyūnga.

HERBARIUM DATA (FG). — 48 collections at CAY. Sel. exs.: *S.A. Mori & B.M. Boom* 15333.

INVENTORY DATA (FG). — 90 trees in 35 plots; Fmax = 2.1%; dbhinv = 37.2 cm.

[1329] *Neea mollis* Spruce ex J.A.Schmidt

Fl. Bras. [Martius] 14 (2): 367 [1 May 1872] (Schmidt 1872).

VERNACULAR NAMES. — Pa: kwepan-wašiunu • Ka: sakusaku • Wp: alasiku, inimopo'i piyūnga, ka'a sala • Br: joāo-mole.

HERBARIUM DATA (FG). — 28 collections at CAY. Sel. exs.: *D. Sabatier et al.* 4623, dbh = 14.6 cm.

[1330] *Neea ovalifolia* Spruce ex J.A.Schmidt

Fl. Bras. [Martius] 14 (2): 368 [1 May 1872] (Schmidt 1872).

VERNACULAR NAMES. — Wp: kwapo'i sili.
HERBARIUM DATA (FG). — 45 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier 2249*.

SIZE. — Up to 35 cm dbh (DeFilipps 2002).

[1331] *Neea spruceana* Heimerl

Notizbl. Königl. Bot. Gart. Berlin 6: 131 [20 Mar. 1914] (Heimerl 1914).

*Neea weberbaueri* Heimerl, Bot. Jahrb. Syst. 54 (1, Beibl. 117): 38 [26 Apr. 1916] (Heimerl 1916).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *J.-J. de Granville 652*.

SIZE. — Up to 20 cm dbh (DeFilipps & Maina 2003).

[1332] *Neea* sp. A
(Fig. 43B)

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier 3385*.

INVENTORY DATA (FG). — 3 trees in 3 plots; Fmax < 1%; dbhinv = 26.9 cm.

[1333] *Neea* sp. B

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 2760*.

INVENTORY DATA (FG). — 4 trees in 3 plots; Fmax = 2.8%; dbhinv = 22.4 cm.

[1334] *Neea* sp. C

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 5082*.

INVENTORY DATA (FG). — 5 trees in 5 plots; Fmax < 1%; dbhinv = 26.3 cm.

[1335] Nyctaginaceae sp. A

HERBARIUM DATA (FG). — 13 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 5334*.

INVENTORY DATA (FG). — 74 trees in 29 plots; Fmax = 1.4%; dbhinv = 57.3 cm.

[1336] Nyctaginaceae sp. B

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 1877*.

INVENTORY DATA (FG). — 2 trees in 2 plots; Fmax < 1%; dbhinv = 60 cm.

[1337] Nyctaginaceae sp. C

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 3169*.

INVENTORY DATA (FG). — 15 trees in 11 plots; Fmax < 1%; dbhinv = 34.3 cm.

[1338] Nyctaginaceae sp. D

HERBARIUM DATA (FG). — A single collection, *D. Sabatier et al. 4880*.

INVENTORY DATA (FG). — 21 trees in 2 plots; Fmax = 3%; dbhinv = 23.6 cm.

Family OCHNACEAE DC.
Genus *Elvasia* DC.

[1339] *Elvasia elvasioides* (Planch.) Gilg

Nat. Pflanzenfam. [Engler & Prantl] 3 (6): 145 (Gilg 1893). — *Hostmannia elvasioides* Planch., Icon. Pl. 8 [n.s., 4]: t. 709 [May 1845] (Planchon 1845). — *Elvasia hostmannia* Planch., London *J. Bot.* 5: 648 (Planchon 1846), *nom. illeg. superfl.* (based on *Hostmannia elvasioides*).

*Hostmannia sagotii* Tiegh., Ann. Sci. Nat., Bot. sér. 8, 16: 414 (Tieghem 1902), “Sagoti”. — *Elvasia sagotii* (Tiegh.) Lemée, Fl. Guyane Franç. 3: 8 (Lemée 1954), “Sagoti”.

*Elvasia caurensis* Pittier, Bol. Soc. Venez. Ci. Nat. 6 (41): 17 [“1939” publ. 1940] (Pittier 1940).

VERNACULAR NAMES. — Pa: yauknabui • Ka: pakila eu.

HERBARIUM DATA (FG). — 17 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier 3996*, dbh = 30.3 cm.

INVENTORY DATA (FG). — 12 trees in 11 plots; Fmax < 1%; dbhinv = 27.9 cm.

[1340] *Elvasia macrostipularis* Sastre & Lescure
(Fig. 44A)

*Caldasia* 12 (57): 134 (Sastre & Lescure 1978).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — 16 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier 4664*.

INVENTORY DATA (FG). — 292 trees in 31 plots; Fmax = 6.5%; dbhinv = 26.6 cm.

Genus *Lacunaria* Ducke

[1341] *Lacunaria crenata* (Tul.) A.C.Sm.
(Fig. 44B)

Trop. Woods 58: 31 (Smith 1939). — *Quiina crenata* Tul., Ann. Sci. Nat., Bot. sér. 3, 11: 163 (Tulasne 1849).
*Quiina panamensis* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser. 4* (8): 236 [24 Oct. 1929] (Standley 1929). — *Lacunaria panamensis* (Standl.) Standl., *Ann. Missouri Bot. Gard.* 29 (4): 358 (Standley 1942).

*Lacunaria pauciflora* Ducke, *Arch. Jard. Bot. Rio de Janeiro* 5: 169 (Ducke 1930).

**VERNACULAR NAMES.** — Ka: woko molokotoli • Wp: tulumele • Wn: kulataju • Br: moella-de-mutum.

**HERBARIUM DATA (FG).** — 67 collections at CAY. Sel. exs.: G.S. *Perrottet s.n.* (type P[P05466483]).

**INVENTORY DATA (FG).** — 67 trees in 49 plots; $F_{max} < 1\%$; $dbh_{inv} = 21.8$ cm.

[1342] *Lacunaria jenmanii* (Oliv.) Ducke subsp. *jenmanii*

*Arch. Jard. Bot. Rio de Janeiro* 5: 171 (Ducke 1930). — *Touroulia jenmanii* Oliv., *Hooker's Icon. Pl.* 20 [ser. 3, 10]: t. 1998 [Aug. 1891] (Oliver 1891), "*Jenmani*".

*Quiina silvatica* Pulle, *Recueil Trav. Bot. Néerl.* 6: 277 (Pulle 1909). — *Lacunaria silvatica* (Pulle) A.C.Sm., *Trop. Woods* 58: 31 (Smith 1939).

*Lacunaria grandiflora* Ducke, *Arch. Jard. Bot. Rio de Janeiro* 4: 140 (Ducke 1925).

**VERNACULAR NAMES.** — Ka: woko molokotoli • Wp: mitũ'ay, tulumele u • Br: moella-de-mutum.

**HERBARIUM DATA (FG).** — 45 collections at CAY. Sel. exs.: *D. Sabatier* 1423.

**INVENTORY DATA (FG).** — 52 trees in 40 plots; $F_{max} < 1\%$; $dbh_{inv} = 31.8$ cm.

[1343] *Lacunaria jenmanii* subsp. *subsessilis*
J.V.Schneid. & Zizka

*Syst. Bot.* 37 (1): 179 [1 Jan. 2012] (Schneider & Zizka 2012).

**VERNACULAR NAMES.** — Pa: avakni-awak puvemna, avakni-awak-seine.

**HERBARIUM DATA (FG).** — 2 collections at CAY. Sel. exs.: *J.-F. Viliers & C. Feuillet* 1974.

**SIZE.** — Up to 25 m tall (Schneider & Zizka 2012).

[1344] *Lacunaria umbonata* Pires

*Bol. Tècn. Inst. Agron. N.* 28: 46 [1954] (Pires 1953).

**NOTE.** — Known only from the Guiana Shield.

**HERBARIUM DATA (FG).** — A single collection, *R.L. Frôes 26671* (type IAN[IAN059825]).

**INVENTORY DATA (FG).** — 1 tree, $dbh = 20.7$ cm.

Genus *Ouratea* Aubl.

[1345] *Ouratea candollei* (Planch.) Tiegh.

*J. Bot. [Morot]* 16: 186 [June 1902] (Tieghem 1902). — *Gomphia candollei* Planch., *London J. Bot.* 6: 4 (Planchon 1847).

**NOTE.** — The binomial *Ouratea candollei* was first invalidly published by Engler (1876: 314) in synonymy under *O. guianensis* Aubl.

**VERNACULAR NAMES.** — Pa: psuk-awak • Cr: malmani • Br: batiputá.

**HERBARIUM DATA (FG).** — 18 collections at CAY. Sel. exs.: *J. Martin* 148 (holo-, P[P00542206]; iso-, BR[BR0000005281756], K[K000382126, K000382127, K000382128], MO[MO-279755], P[P00542207]).

**INVENTORY DATA (FG).** — 15 trees in 3 plots; $F_{max} = 1.8\%$; $dbh_{inv} = 22.6$ cm.

[1346] *Ouratea cerebroidea* Sastre

*Bull. Mus. Natl. Hist. Nat.*, B, *Adansonia* 8 (4): 413 ["1986" publ. 1987] (Sastre 1987).

**NOTE.** — Known only from the Guiana Shield.

**VERNACULAR NAMES.** — Pa: yauknabui.

**HERBARIUM DATA (FG).** — 17 collections at CAY. Sel. exs.: *C. Sastre* 6405 (holo-, P[P00542192]; iso-, P[P00542193], U[U0005264]).

**INVENTORY DATA (FG).** — 2 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 18.9$ cm.

[1347] *Ouratea decagyna* Maguire

*Bull. Torrey Bot. Club* 75 (6): 644 [Dec. 1948] (Maguire 1948).

**HERBARIUM DATA (FG).** — 6 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 2944.

**INVENTORY DATA (FG).** — 31 trees in 25 plots; $F_{max} < 1\%$; $dbh_{inv} = 28$ cm.

[1348] *Ouratea francinae* Sastre

*Bull. Mus. Natl. Hist. Nat.*, B, *Adansonia* 8 (4): 417 ["1986" publ. 1987] (Sastre 1987).

**VERNACULAR NAMES.** — Wp: wai tawa.

**HERBARIUM DATA (FG).** — Known only from the type: *P. Grenand* 510.

**SIZE.** — Up to 10 m tall (Sastre 1986).

[1349] *Ouratea guianensis* Aubl.

*Hist. Pl. Guiane* 1: 397 [Jun.-Dec. 1775] (Aublet 1775), "*Guyannensis*" on plate.

**NOTE.** — Known only from the Guiana Shield.
VERNACULAR NAMES. — Pa: yauknabui • Ka: akale tapulala kili • Wp: tukā nākū • Wn: potīpotpiu • Nt: weti tiki • Cr: malmani • Br: batiputá.

HERBARIUM DATA (FG). — 39 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000797998]).

INVENTORY DATA (FG). — 17 trees in 15 plots; $F_{max} < 1\%$; $dbh_{inv} = 33.7$ cm.

[1350] *Ouratea leblondii* (Tiegh.) Lemée

Fl. Guyane Franç. 3: 8 (Lemée 1954), “*Leblondi*”. — *Camptouratea leblondii* Tiegh., *Ann. Sci. Nat., Bot.* sér. 8, 16: 205 (Tieghem 1902), “*Leblondi*”.

*Camptouratea sagotii* Tiegh., *Ann. Sci. Nat., Bot.* sér. 8, 16: 211 (Tieghem 1902), “*Sagoti*”. — *Ouratea sagotii* (Tiegh.) Lemée, *Fl. Guyane Franç.* 3: 9 (Lemée 1954).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Wp: ii.

HERBARIUM DATA (FG). — 94 collections at CAY. Sel. exs.: *J.B. Leblond s.n.* (lecto-, P[P00542279], designated by Sastre & Offroy [2016: 60]; isolecto-, K[not seen], P[P00582165]); *C. Sastre & C. Moretti 4020*, 15 m.

[1351] *Ouratea macrocarpa* Sastre

Bull. Mus. Natl. Hist. Nat., B, *Adansonia* 8 (4): 414 [“1986” publ. 1987] (Sastre 1987).

NOTE. — Known only from French Guiana.

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *J.-J. de Granville 44* (holo-, P[P00542348]; iso-, CAY[CAY024895, CAY024896], COL[COL000002626], K[K000382115], NY[00428959], P[P00542349], RB[RB00542384], SI[SI003068, SI003069], U[U0008103], US[00289224]).

SIZE. — Up to 17 m tall (Sastre 1986).

[1352] *Ouratea melinonii* (Tiegh.) Lemée

Fl. Guyane Franç. 3: 12 (Lemée 1954). — *Cercouratea melinonii* Tiegh., *Ann. Sci. Nat., Bot.* sér. 8, 16: 278 (Tieghem 1902), “*Melinoni*”.

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Wp: ii.

HERBARIUM DATA (FG). — 25 collections at CAY. Sel. exs.: *E.M. Mélinon s.n.* (holo-, P[P00542463]; iso-, P[P00542464]).

INVENTORY DATA (FG). — 40 trees in 21 plots; $F_{max} = 1.1\%$; $dbh_{inv} = 18.8$ cm.

[1353] *Ouratea retrorsa* Sastre

*Adansonia*, sér. 3, 29 (1): 89 (Sastre 2007).

HERBARIUM DATA (FG). — Known only from the type: *D. Larpin 1021* (holo-, P[P00542460]; iso-, CAY[CAY079504]).

SIZE. — Up to 12 m tall (Sastre 2007).

[1354] *Ouratea scottii* Sastre
(Fig. 44C)

Brittonia 46 (4): 309 (Sastre 1994).

VERNACULAR NAMES. — Wp: tukā nili • Cr: malmani • Br: batiputá.

HERBARIUM DATA (FG). — 25 collections at CAY. Sel. exs.: *S.A. Mori et al. 14927* (holo-, NY[00345875]; iso-, CAY[CAY002334], P[P00542208]).

INVENTORY DATA (FG). — 19 trees in 6 plots; $F_{max} = 1.3\%$; $dbh_{inv} = 20.7$ cm.

Genus *Quiina* Aubl.

[1355] *Quiina berryi* J.V.Schneid. & Zizka

Candollea 58 (2): 464 (Schneider & Zizka 2003).

HERBARIUM DATA (FG). — A single collection, *J.-J. de Granville 7755*.

SIZE. — Up to 20 m tall (Schneider & Zizka 2016).

[1356] *Quiina cruegeriana* Griseb.

Fl. Brit. W.I. [Grisebach] 106 [June 1859] (Grisebach 1859).

*Quiina guianensis* Crueg. ex Griseb., *Fl. Brit. W.I. [Grisebach]* 106 [June 1859] (Grisebach 1859), *nom. nud. pro syn.*

*Quiina peruviana* Engl., *Fl. Bras. [Martius]* 12 (1): 481 [1 Apr. 1888] (Engler 1888).

*Quiina tessmannii* Mildbr., *Notizbl. Bot. Gart. Berlin-Dahlem* 11: 298 [10 Nov. 1931] (Mildbraed 1931).

*Quiina cajambrensis* Cuatrec., *Trop. Woods* 96: 41 (Cuatrecasas 1950).

*Quiina congesta* R.S.Cowan, *Brittonia* 8 (4): 239 [Jan. 1957] (Cowan 1957).

*Quiina guaporensis* Pires, *Bol. Técn. Inst. Agron. N.* 38: 31 (Pires 1960).

*Lacunaria colonensis* D'Arcy, *Ann. Missouri Bot. Gard.* 67 (4): 967 ["1980" publ. 1981] (D'Arcy 1981). — *Quiina colonensis* (D'Arcy) D'Arcy, *Monogr. Syst. Bot. Missouri Bot. Gard.* 18: 534 (D'Arcy 1987).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *P. Grenand 1902*.

SIZE. — Up to 15 m tall (Schneider & Zizka 2012).

[1357] *Quiina guianensis* Aubl.

*Hist. Pl. Guiane* 2 (Suppl.): 19 [Jun.-Dec. 1775] (Aublet 1775), “*Guyannensis*” on plate.

VERNACULAR NAMES. — Ka: wokunse • Wp: iwaka'i sili, waka'i sili • Nt: dyabatee, pikin busi suku • Br: quinarana.

HERBARIUM DATA (FG). — 49 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000073181]).

INVENTORY DATA (FG). — 18 trees in 13 plots; $F_{max} < 1\%$; $dbh_{inv} = 14.2$ cm.
<figure><img src="image_66.png" /><figcaption>FIG. 43. — Nyctaginaceae: **A**, *Neea floribunda* Poepp. & Endl. (*D. Sabatier & J.-F. Molino* 5616); **B**, *Neea* sp. *A* (*M.-F. Prévost* & *D. Sabatier* 3385), Olacaceae: **C**, *Dulacia guianensis* (Engl.) Kuntze; **D**, *Heisteria ovata* Benth. (*D. Sabatier & J.-F. Molino* 5086). A, C, D, © D. Sabatier/IRD; B, © M.-F. Prévost/IRD.</figcaption></figure>

[1358] *Quiina integrifolia* Pulle

*Recueil Trav. Bot. Néerl.* 9: 153 (Pulle 1912).

*Quiina rigidifolia* Pires, *Bol. Tècn. Inst. Agron.* N. 20: 45 (Pires 1950).

VERNACULAR NAMES. — Pa: yauknabui-puvemna • Br: quinarana.

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino* 4977.

INVENTORY DATA (FG). — 58 trees in 35 plots; Fmax = 2.1%; dbhinv = 26.3 cm.

[1359] *Quiina leptoclada* Tul.

*Ann. Sci. Nat., Bot.* sér. 3, 11: 159 (Tulasne 1849).

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: *E.M. Mélinon* 147, 1845 (holo-, P[P02441362]; iso-, K[K000328614], P[P00071950, P02441363]).

INVENTORY DATA (FG). — 5 trees in 4 plots; dbhinv = 16.5 cm.

[1360] *Quiina obovata* Tul.

*Ann. Sci. Nat., Bot.* sér. 3, 11: 157 (Tulasne 1849).

*Quiina decaisneana* Planch. & Triana, *Ann. Sci. Nat., Bot.* sér. 4, 15: 315 (Planchon & Triana 1861).

*Quiina albiflora* A.C.Sm., *Trop. Woods* 58: 30 (Smith 1939).

*Quiina oblanceolata* Sandwith, *Bull. Misc. Inform. Kew* 1939 (10): 546 ["1939" publ. 6 Jan. 1940] (Sandwith 1940).

VERNACULAR NAMES. — Pa: kwatri-wašiunô-purubumna • Ka: wokunse • Wp: iwaka’i, waka’i, waka’i u • Nt: dyabatee • Br: quinarana.

HERBARIUM DATA (FG). — 50 collections at CAY. Sel. exs.: *J. Martin s.n.* (holo-, P[P00071947]; iso-, B[B 10 0294392], K[K000328613], P[P02441367, P02441368], SI[SI004194], U[U0282664]).

INVENTORY DATA (FG). — 61 trees in 49 plots; Fmax < 1%; dbhinv = 25.9 cm.

[1361] *Quiina oiapocensis* Pires

*Bol. Tècn. Inst. Agron.* N. 38: 32 (Pires 1960).

VERNACULAR NAMES. — Pa: kwatri-duwě • Br: quinarana.

HERBARIUM DATA (FG). — 25 collections at CAY. Sel. exs.: *D. Sabatier* 1541.

INVENTORY DATA (FG). — 40 trees in 22 plots; Fmax < 1%; dbhinv = 22.3 cm.
<figure><img src="image_67.png" /></figure>

FIG. 44. — Ochnaceae: **A**, *Elvasia macrostipularis* Sastre & Lescure (J.-F. Molino & D. Sabatier 2099); **B**, *Lacunaria crenata* (Tul.) A.C.Sm. (D. Sabatier et al. 4805); **C**, *Ouratea scottii* Sastre (D. Sabatier & J.-F. Molino 5071); **D**, *Quiina* sp. A (D. Sabatier & J.-F. Molino 5197). © D. Sabatier/IRD.

[1362] *Quiina pteridophylla* (Radlk.) Pires

Bol. Tècn. Inst. Agron. N. 20: 48 (Pires 1950). — *Touroulia pteridophylla* Radlk., *Sitzungsber. Math.-Phys. Cl. Köngl. Bayer. Akad. Wiss. München* 19: 218 (Radlkofer 1889).

*Quiina acutangula* Ducke, *Arch. Jard. Bot. Rio de Janeiro* 4: 143 (Ducke 1925).

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: S.A. Mori et al. 23207.

SIZE. — Up to 12 m tall (Schneider & Zizka 2004).

[1363] *Quiina sessilis* Choisy ex Planch. & Triana

*Ann. Sci. Nat., Bot.* sér. 4, 15: 312 (Planchon & Triana 1861).

NOTE. — Known only from French Guiana.

VERNACULAR NAMES. — Pa: kwatri-wašiunô, kwatri-wašiunô-duwê.

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: *J.-B. Patris s.n.* (holo-, G[G00389893]; iso-, G[G00389894]; possible iso-, MPU[MPU014417]).

INVENTORY DATA (FG). — 7 trees in 6 plots; Fmax < 1%; dbhinv = 12.7 cm.

[1364] *Quiina yatuensis* J.V.Schneid. & Zizka

Novon 7 (4): 406 [“1997” publ. 1998] (Schneider & Zizka 1998).

HERBARIUM DATA (FG). — A single collection, *M.-F. Prévost & D. Sabatier* 4623, dbh 12.9 cm.

[1365] *Quiina* sp. A
(Fig. 44D)

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier* 2661.

INVENTORY DATA (FG). — 9 trees in 6 plots; Fmax < 1%; dbhinv = 28.3 cm.
## Genus *Touroulia* Aubl.

### [1366] *Touroulia guianensis* Aubl.

*Hist. Pl. Guiane* 1: 492 [Jun.-Dec. 1775] (Aublet 1775), "*Guyanensis*" on plate. — *Robinsonia guianensis* J.F.Gmel., *Syst. Nat.*, ed. 13[bis], 2 (1): 796 [late Sep.-Nov. 1791] (Gmelin 1791), *nom. illeg. superfl.* (based on *Touroulia guianensis*). — *Robinsonia melianthifolia* Willd., *Sp. Pl.*, ed. 4 2 (2): 999 [Dec. 1799] (Willdenow 1799), *nom. illeg. superfl.* (based on *Touroulia guianensis*). — *Touroulia solitaria* Stokes, *Bot. Mat. Med.* 3: 105 (Stokes 1812), "*Tourolia*", *nom. illeg. superfl.* (based on *Robinsonia melanthifolia*).

**VERNACULAR NAMES.** — Pa: yit-āra • Wp: wila taiwa • Nt: yooka wiwii • Cr: bwa-flanbo.

**HERBARIUM DATA (FG).** — 45 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P[P00114408] designated by Zizka & Schneider [1999: 232]).

**INVENTORY DATA (FG).** — 44 trees in 31 plots; $F_{max} = 1\%$; $dbh_{inv} = 26.9$ cm.

## Family OLACACEAE R.Br.
Genus *Cathedra* Miers

### [1367] *Cathedra acuminata* (Benth.) Miers

*Ann. Mag. Nat. Hist.*, ser. 3, 4: 361 (Miers 1859). — *Diplocrater acuminatus* Benth., *Hooker's J. Bot. Kew Gard. Misc.* 3: 367 (Bentham 1851).

*Cathedra crassifolia* Benth. ex Miers, *Ann. Mag. Nat. Hist.*, ser. 3, 4: 361 (Miers 1859).

*Cathedra aestuaria* Sleumer, *Repert. Spec. Nov. Regni Veg.* 39: 279 (Sleumer 1936).

*Cathedra inaequilatera* Sleumer, *Repert. Spec. Nov. Regni Veg.* 39: 280 (Sleumer 1936).

*Cathedra oblonga* Sleumer, *Repert. Spec. Nov. Regni Veg.* 39: 280 (Sleumer 1936).

*Cathedra caurensis* Pittier, *Bol. Soc. Venez. Ci. Nat.* 6 (41): 5 ["1939" publ. 1940] (Pittier 1940).

*Cathedra guianensis* Sandwith, *Kew Bull.* 10 (3): 469 [20 Dec. 1955] (Sandwith 1955).

**HERBARIUM DATA (FG).** — 9 collections at CAY. Sel. exs.: *D. Sabatier* 5778.

**INVENTORY DATA (FG).** — 14 trees in 11 plots; $F_{max} < 1\%$; $dbh_{inv} = 28.8$ cm.

## Genus *Chaunochiton* Benth.

### [1368] *Chaunochiton kappleri* (Sagot ex Engl.) Ducke

*Arch. Jard. Bot. Rio de Janeiro* 3: 41 (Ducke 1922). — *Heisteria kappleri* Sagot ex Engl., *Fl. Bras. [Martius]* 12 (2): 14 [1 Dec. 1872] (Engler 1872). — *Sagotanthus kappleri* (Sagot ex Engl.) Dyer, *Index Kew. Suppl.* 2: 162 (Dyer 1904).

*Chaunochiton breviflorum* Ducke, *Arch. Jard. Bot. Rio de Janeiro* 3: 41 (Ducke 1922).

*Chaunochiton purpurascens* Rizzini, *Dusenia* 1 (5): 289 (Rizzini 1950).

**VERNACULAR NAMES.** — Ka: aluku elepali, apesiya • Wp: wila pile lu • Nt: auman pisi, gawenti, tu pikin weko • Br: pau-vermelho.

**HERBARIUM DATA (FG).** — 17 collections at CAY. Sel. exs.: *P.A. Sagot* 1198, 1858 (holo-, P[P02441958]; iso-, B[B 10 0248312], BM[BM000839877], BR[BR0000005280810], GH[GH00035955], GOET[GOET008319], K[K000580931], L[L0038980], P[P02441959, P02441960, P02441961, P02441962, P02441963], S[S-R-2779], U[U0005296]).

**INVENTORY DATA (FG).** — 127 trees in 71 plots; $F_{max} = 1.4\%$; $dbh_{inv} = 72.6$ cm.

## Genus *Dulacia* Vell.

### [1369] *Dulacia guianensis* (Engl.) Kuntze
(Fig. 43C)

*Revis. Gen. Pl.* 1: 111 [5 Nov. 1891] (Kuntze 1891). — *Liriosma guianensis* Engl., *Fl. Bras. [Martius]* 12 (2): 25 [1 Dec. 1872] (Engler 1872). — *Olax guianensis* (Engl.) Christenh. & Byng, *Global Fl.* 4: 143 [9 Feb. 2018] (Christenhusz & Byng 2018).

*Olax schomburgkii* Klotzsch, *Reis. Br.-Guiana* [Ri. Schomburgk] 3: 1177 ["1848" publ. 7-10 Mar. 1849] (Klotzsch 1849), *nom. nud.*

*Liriosma cerifera* A.C.Sm., *Brittonia* 2 (2): 146 (Smith 1936).

**NOTE.** — According to Malécot & Nickrent (2008), *Dulacia* Vell. is possibly embedded in *Olax* L.

**VERNACULAR NAMES.** — Pa: balata-kamwi.

**HERBARIUM DATA (FG).** — 11 collections at CAY. Sel. exs.: *J.-F. Molino et al.* 2108.

**INVENTORY DATA (FG).** — 18 trees in 14 plots; $F_{max} < 1\%$; $dbh_{inv} = 38$ cm.

## Genus *Heisteria* Jacq.

### [1370] *Heisteria barbata* Cuatrec.

*Trop. Woods* 101: 26 (Cuatrecasas 1955).

**HERBARIUM DATA (FG).** — 4 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 5403.

**INVENTORY DATA (FG).** — 8 trees in 8 plots; $F_{max} < 1\%$; $dbh_{inv} = 31.2$ cm.

### [1371] *Heisteria densifrons* Engl.

*Fl. Bras. [Martius]* 12 (2): 17 [1 Dec. 1872] (Engler 1872).

*Heisteria microcarpa* Spruce ex Engl., *Fl. Bras. [Martius]* 12 (2): 21 [1 Dec. 1872] (Engler 1872).

*Heisteria microcalyx* Sagot, *Ann. Sci. Nat.*, Bot. sér. 6, 11: 159 (Sagot 1881).
*Heisteria sessilis* Ducke, *Arch. Jard. Bot. Rio de Janeiro* 4: 8 (Ducke 1925).

*Heisteria parvicalyx* A.C.Sm., *Brittonia* 2 (2): 147 (Smith 1936).

VERNACULAR NAMES. — Pa: pun-kamwi, sitru-kamwi • Wp: wila munuwi • Wn: mekuhuluimë, mulekju • Cr: bwa-pedri • Br: cafezinho.

HERBARIUM DATA (FG). — 137 collections at CAY. Sel. exs.: *FM.R. Leprieur s.n.* (holotype of *Heisteria microcalyx*: P[P02441925]; iso-, L[L0038998], P[P02441926]).

INVENTORY DATA (FG). — 71 trees in 51 plots; Fmax = 1.1%; dbhinv = 22.3 cm.

[1372] *Heisteria ovata* Benth.
(Fig. 43D)

Hooker's *J. Bot*. Kew Gard. Misc. 3: 366 (Bentham 1851).

*Heisteria subsessilis* Benth., Hooker's *J. Bot*. Kew Gard. Misc. 3: 367 (Bentham 1851).

*Heisteria nitida* Engl., *Fl. Bras.* [*Martius*] 12 (2): 18 [1 Dec. 1872] (Engler 1872).

*Heisteria flexuosa* Engl., *Fl. Bras.* [*Martius*] 12 (2): 21 [1 Dec. 1872] (Engler 1872).

*Heisteria rubricalyx* S.Moore, *Trans. Linn. Soc. London, Bot.* 4 (3): 337 ["1894-96" publ. Dec. 1895] (Moore 1895).

*Heisteria micrantha* Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (2): 340 (Huber 1909).

*Heisteria vageleri* Burret, *Notizbl. Bot. Gart. Berlin-Dahlem* 9: 50 [1 June 1924] (Burret 1924).

*Heisteria krukovii* A.C.Sm., *Brittonia* 2 (2): 147 (Smith 1936).

*Heisteria surinamensis* Amshoff, *Recueil Trav. Bot. Néerl.* 34: 497 (Amshoff 1937).

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino* 5086.

INVENTORY DATA (FG). — 12 trees in 9 plots; Fmax < 1%; dbhinv = 38.8 cm.

Genus *Minquartia* Aubl.

[1373] *Minquartia guianensis* Aubl.

*Hist. Pl. Guiane* 2 (Suppl.): 4 [Jun.-Dec. 1775] (Aublet 1775), "Guyannensis" on plate.

*Secretania loranthacea* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 228 [late Aug. 1866] (Müller 1866).

*Endusa punctata* Radlk., *Sitzungsber. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. München* 16: 313 (Radlkofer 1886). — *Minquartia punctata* (Radlk.) Sleumer, *Repert. Spec. Nov. Regni Veg.* 39: 282 (Sleumer 1936).

*Eganthus poeppigii* Tiegh., *J. Bot.* [*Morot*] 13: 77 (Tieghem 1899).

*Minquartia macrophylla* Ducke, *Arch. Inst. Biol. Veg.* 2 (1): 33 [Sep. 1935] (Ducke 1935).

*Minquartia parvifolia* A.C.Sm., *Brittonia* 2 (2): 149 (Smith 1936).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: mekwa, yawu • Ka: alataweli, iwiiyu, wiyu • Te: iwidju • Wp: wakali'i • Wn: kunawa • Nt: alata udu • Cr: mékwa • Fr: mincouart • Br: acariquara, acariúba.

HERBARIUM DATA (FG). — 71 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000028175]).

INVENTORY DATA (FG). — 373 trees in 143 plots; Fmax = 4.4%; dbhinv = 112.4 cm.

Genus *Ptychopetalum* Benth.

[1374] *Ptychopetalum olacoides* Benth.

London *J. Bot.* 2: 377 (Bentham 1843).

VERNACULAR NAMES. — Pa: aneku, aneku-kamwi • Te: wilapilatã • Wp: wila pilatã, wila tai • Cr: bwa-bandé • Fr: bois bandé • Br: marapuama, muira-puama.

HERBARIUM DATA (FG). — 40 collections at CAY. Sel. exs.: *J. Martin s.n.* (type K[K000581314]).

INVENTORY DATA (FG). — 138 trees in 46 plots; Fmax = 2.6%; dbhinv = 33.7 cm.

Genus *Ximenia* Plum. ex L.

[1375] *Ximenia americana* L.

*Sp. Pl.* 2: 1193 [1 May 1753] (Linnaeus 1753). — *Ximenia spinosa* Salisb., *Prodr. Stirp. Chap. Allerton*: 276 [Nov.-Dec. 1796] (Salisbury 1796), *nom. illeg. superfl.* (based on *Ximenia americana*). — *Ximenia montana* Macfad., *Fl. Jamaica* [*Macfadyen*] 1: 121 (Macfadyen 1837), *nom. illeg. superfl.* (based on *Ximenia americana* and *X. multiflora*).

*Amyris arborescens* P.Browne, *Civ. Nat. Hist. Jamaica*.: 209 (Browne 1756). — *Ximenia inermis* L., *Sp. Pl.*, ed. 2, 1: 497 [Sep. 1762] (Linnaeus 1762), *nom. illeg. superfl.* (based on *Amyris arborescens*).

*Heymassoli inermis* Aubl., *Hist. Pl. Guiane* 1: 325 [Jun.-Dec. 1775] (Aublet 1775).

*Heymassoli spinosa* Aubl., *Hist. Pl. Guiane* 1: 324 [Jun.-Dec. 1775] (Aublet 1775). — *Ximenia americana* var. *oblonga* DC., *Prodr. [A. P. de Candolle]* 1: 533 [mid Jan. 1824] (Candolle 1824), *nom. illeg. superfl.* (based on *Heymassoli spinosa*).

*Pimecaria odorata* Raf., *Alsogr. Amer.*: 65 (Rafinesque 1838).

*Ximenia aculeata* Crantz, *Inst. Rei Herb.* 2: 381 (Crantz 1766).

*Ximenia americana* f. inermis (Aubl.) Engl., *Fl. Bras.* [*Martius*] 12 (2): 9 [1 Dec. 1872] (Engler 1872).

*Ximenia americana* f. inermis S.Moore, *Trans. Linn. Soc. London, Bot.* 4 (3): 337 ["1894-96" publ. Dec. 1895] (Moore 1895), *nom. illeg. hom.*, *non* (Aubl.) Engl. (Engler 1872).

*Ximenia arborescens* Tussac ex Walp., *Repert. Bot. Syst.* [*Walpers*] 1 (2): 377 [18-20 Sep. 1842] (Walpers 1842).

*Ximenia elliptica* G.Forst., Fl. Ins. Austr. 27 [Oct.-Nov. 1786] (Forster 1786). — *Ximenia verrucosa* M.Roem., *Fam. Nat. Syn.*
*Monogr.* 1 (Hesperides): 23 (Roemer 1846), *nom. illeg. superfl.*
(based on *Ximenia elliptica*).

*Ximenia fluminensis* M.Roem., *Fam. Nat. Syn. Monogr.* 1 (Hesperides): 22 (Roemer 1846).

*Ximenia laurina* Delile, *Ann. Sci. Nat., Bot.* sér. 2, 20: 89 (Delile 1843).

*Ximenia loranthifolia* Span., *Linnaea* 15: 177 (Spanoghe 1841),
*nom. nud. pro syn.*

*Ximenia multiflora* Jacq., * Enum. Syst. Pl.*: 19 [Aug.-Sep. 1760] (Jacquin 1760). — *Ximenia americana* var. *ovata* DC., *Prodr.* [*A. P. de Candolle*] 1: 533 [mid Jan. 1824] (Candolle 1824), *nom. illeg. superfl.* (based on *Ximenia multiflora*).

*Ximenia oblonga* Lam. ex Hemsl., *Biol. Cent.-Amer., Bot.* 1 (3): 185 [Feb. 1879] (Hemsley 1879).

NOTES. — Restricted to coastal sandridges. *Ximenia americana* f. *inermis* S.Moore is not based on *Heymassoli inermis* Aubl.

VERNACULAR NAMES. — Ka: heymassoli (fide Aublet 1775) • Fr: citron de mer.

HERBARIUM DATA (FG). — 13 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material of *Heymassoli spinosa*: P-JU, P00671912)).

SIZE. — Up to 12 m tall (Hiepko 1993).

Family OLEACEAE Hoffmanns. & Link
Genus *Chionanthus* D.Royen

[1376] *Chionanthus guianensis* (Aubl.) Pers.

*Syn. Pl. [Persoon]* 1: 9 [1 Apr.-15 June 1805) (Persoon 1805). — *Mayepea guianensis* Aubl., *Hist. Pl. Guiane* 1: 81 [Jun.-Dec. 1775] (Aublet 1775), “*Guyannensis*” on plate. — *Chionanthus mayepea* Vahl, *Symb. Bot. [Vahl]* 2: 1 [Jul.-Dec. 1791] (Vahl 1791), *nom. illeg. superfl.* (based on *Mayepea guianensis*). — *Chionanthus tetrandrus* Vahl, * Enum. Pl. [Vahl]* 1: 45 [Jul.-Dec. 1804] (Vahl 1804), “*tetrandra*”, *nom. illeg. superfl.* (based on *Mayepea guianensis*).

*Linociera tetrandra* R.Br., Prodr. Fl. Nov. Holland. 523 [27 Mar. 1810] (Brown 1810), *nom. inval.* (genus name and epithet not associated).

NOTE. — Known only from French Guiana.

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JJR[P00777522] designated by Lanjouw & Uittien [1940: 153]).

SIZE. — Aublet (1775): “Son tronc a [...] environ cinq pouces de diamètre”, i.e. c. 12.5 cm diameter.

Family OPILIACEAE Valeton
Genus *Agonandra* Miers ex Benth.

[1377] *Agonandra silvatica* Ducke
(Fig. 45A)

Arch. Jard. Bot. Rio de Janeiro 3: 41 (Ducke 1922).

HERBARIUM DATA (FG). — 13 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 2341*.

INVENTORY DATA (FG). — 33 trees in 24 plots; $F_{max} < 1\%$; $dbh_{inv} = 43.9$ cm.

Family PENTAPHYLACACEAE Engl.
Genus *Ternstroemia* Mutis ex L.f.

[1378] *Ternstroemia delicatula* Choisy

Mém. Soc. Phys. Genève 14 (1): 106 (Choisy 1855).

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *J. Martin s.n.* (original material P[P00780885, P00780886, P00780887]).

INVENTORY DATA (FG). — 38 trees in 4 plots; $F_{max} = 2.8\%$; $dbh_{inv} = 55.1$ cm.

[1379] *Ternstroemia dentata* (Aubl.) Sw.
(Fig. 45B)

*Prodr. [Swartz]* 81 [20 Jun.-29 July 1788] (Swartz 1788). — *Taonabo dentata* Aubl., *Hist. Pl. Guiane* 1: 569 [Jun.-Dec. 1775] (Aublet 1775).

VERNACULAR NAMES. — Ka: palipyali, pipi • Wp: walatiwā sili • Wn: maipuli apepiton.

HERBARIUM DATA (FG). — 13 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000041973]).

INVENTORY DATA (FG). — 3 trees in 3 plots; $F_{max} < 1\%$; $dbh_{inv} = 15.3$ cm.

[1380] *Ternstroemia punctata* (Aubl.) Sw.

*Prodr. [Swartz]* 81 [20 Jun.-29 July 1788] (Swartz 1788). — *Taonabo punctata* Aubl., *Hist. Pl. Guiane* 1: 571 [Jun.-Dec. 1775] (Aublet 1775).

*Ternstroemia revoluta* Splitg., *Tijdschr. Nat. Geschied. Physiol.* 9: 99 [Aug.-Sep. 1842] (Splitgerber 1842).

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000041974]).

SIZE. — Aublet (1775, 1: 571) wrote: “Cette espèce de palétuvier qui s’élève aussi haut que le précédent, et qui en a le port [...]”, i.e. similar in height and habit to the preceding. The preceding is *Taonabo dentata* Aubl., which is said to have a trunk “deux pieds de diamètre”, i.e. c. 60 cm diameter.

Family PERACEAE Klotzsch
Genus *Chaetocarpus* Thwaites

[1381] *Chaetocarpus schomburgkianus*
(Kuntze) Pax & K.Hoffm.

*Pflanzenr. [Engler]* IV.147.IV (Heft 52): 10 [13 Feb. 1912] (Pax & Hoffmann 1912). — *Gaedawakka schomburgkiana* Kuntze, *Revis. Gen. Pl.* 2: 606 [5 Nov. 1891] (Kuntze 1891).
*Drypetes spruceana* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 454 [late Aug. 1866] (Müller 1866).

*Discocarpus mazarunensis* Croizat, *Bull. Torrey Bot. Club* 75 (4): 400 [Jul.-Aug. 1948] (Croizat 1948).

*Drypetes maguireana* Monach., *Phytologia* 3 (1): 33 [30 Aug. 1948] (Monachino 1948).

*Chaetocarpus williamsii* Steyerm., *Fieldiana*, *Bot.* 28 (2): 306 (Steyermark 1952).

VERNACULAR NAMES. — Ka: kusali yepo, wiyekane • Nt: sipiki udu.

HERBARIUM DATA (FG). — 63 collections at CAY. Sel. exs.: *D. Sabatier* 2329.

INVENTORY DATA (FG). — 386 trees in 123 plots; Fmax = 1.9%; dbhinv = 76 cm.

[1382] *Chaetocarpus* sp. A

HERBARIUM DATA (FG). — 16 collections at CAY. Sel. exs.: *R.ENOIST* 539 (P[P05479470]).

INVENTORY DATA (FG). — 80 trees in 29 plots; Fmax < 1%; dbhinv = 82 cm.

## Genus *Pera* Mutis

[1383] *Pera bicolor* (Klotzsch) Müll.Arg.

*Prodr. [A. P. de Candolle]* 15 (2.2): 1028 [late Aug. 1866] (Müller 1866). — *Peridium bicolor* Klotzsch, *London J. Bot.* 2: 44 (Klotzsch 1843).

*Peridium schomburgkii* Klotzsch, *Reis. Br.-Guiana [Ri. Schomburgk]* 3: 1089 [“1848” publ. 7-10 Mar. 1849] (Klotzsch 1849), *nom. nud.*

*Peridium schomburgkianum* Klotzsch ex Benth., *Hooker's J. Bot. Kew Gard. Misc.* 6: 323 (Bentham 1854), in obs. — *Pera schomburgkiana* (Klotzsch ex Benth.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1027 [late Aug. 1866] (Müller 1866).

VERNACULAR NAMES. — Ka: boko.

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: *D. Loubry* 55.

INVENTORY DATA (FG). — 5 trees in 4 plots; Fmax < 1%; dbhinv = 46.5 cm.

[1384] *Pera glabrata* (Schott) Poepp. ex Baill.

*Étude Euphorb.*: 434 (Baillon 1858). — *Peridium glabratum* Schott, *Syst. Veg. [Sprengel]* 4 (2, Cur. Post.): 410 [Jan.-June 1827] (Schott 1827).

*Peridium ferrugineum* Schott, *Syst. Veg. [Sprengel]* 4 (2, Cur. Post.): 410 [Jan.-June 1827] (Schott 1827). — *Pera ferruginea* (Schott) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 1031 [late Aug. 1866] (Müller 1866).

*Peridium oblongifolium* Benth., *Hooker's J. Bot. Kew Gard. Misc.* 2: 243 (Bentham 1850).

NOTE. — “*Pera arborea* Baill.”, listed in IPNI and Tropicos, does not exist. Baillon (*Adansonia* [Baillon] 5: 224 [Baillon 1865]) misinterpreted *P. arborea* Mutis, to which he explicitly referred.

VERNACULAR NAMES. — Ka: pilityalapo, woko popi.

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *M.-F. Prévost* 649.

INVENTORY DATA (FG). — 2 trees in 2 plots; Fmax < 1%; dbhinv = 38 cm.

## Genus *Pogonophora* Miers ex Benth.

[1385] *Pogonophora schomburgkiana* Miers ex Benth. (Fig. 45C)

Hooker's *J. Bot. Kew Gard. Misc.* 6: 373 (Bentham 1854).

*Pogonophora schomburgkiana* var. longifolia Benth., *Hooker's J. Bot. Kew Gard. Misc.* 6: 373 (Bentham 1854).

*Pogonophora schomburgkiana* f. elliptica Pax, *Pflanzenr.* [Engler] IV.147.III (Heft 47): 108 [14 Feb. 1911] (Pax 1911).

*Poraesia anomala* Gleason, *Bull. Torrey Bot. Club* 58 (6): 385 [June 1931] (Gleason 1931).

VERNACULAR NAMES. — Ka: tapilen pipyo, tikilin wewe • Nt: geli udu • Br: amarelinho, miatoá.

HERBARIUM DATA (FG). — 40 collections at CAY. Sel. exs.: *D. Sabatier* 3556.

INVENTORY DATA (FG). — 1054 trees in 100 plots; Fmax = 8.8%; dbhinv = 38.2 cm.

## Family PHYLLANTHACEAE Martinov
Genus *Amanoa* Aubl.

[1386] *Amanoa congesta* W.J.Hayden

*Brittonia* 42 (4): 261 (Hayden 1990).

VERNACULAR NAMES. — Wp: wila tã.

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: *M.-F. Prévost* 3768.

INVENTORY DATA (FG). — 16 trees in 11 plots; Fmax < 1%; dbhinv = 51.7 cm.

[1387] *Amanoa grandiflora* (Müll.Arg.) Müll.Arg. (Fig. 46A)

*Flora* 55: 2 (Müller 1872). — *Amanoa guianensis* var. grandiflora Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 910 [late Aug. 1866] (Müller 1866).

*Amanoa potamophila* Croizat, *Amer. Midl. Naturalist* 29 (2): 475 [5 Apr. 1943] (Croizat 1943).

*Amanoa macrocarpa* Cuatrec., *Brittonia* 11: 164 (Cuatrecasas 1959).
NOTES. — Hayden (1990) showed that the Aublet collections, which could have served as the type of *Amanoa guianensis* Aubl., actually belong to two distinct species. One is known only from a few specimens from French Guiana, Amapá and Suriname; the other is widespread from Central America to northern South America (Hayden 1990). He therefore selected among these specimens a lectotype (W[W-1889-0012576]; isolectotypes G[not seen], F[V0055866F], A[not seen], MO[not seen]) which corresponds to the current usage of Aublet's name, that is to the widespread species, and described a new species, *Amanoa neglecta* W.J.Hayden, based on the second entity found in Aublet collections (holo-, P-JJR[P00777979], iso-, BM[BM000645699]) (Hayden 1990). This was a laudable choice to preserve the current usage of the name *A. guianensis*, but unfortunately Hayden (1990) overlooked the previous lectotypification of *A. guianensis* Aubl. by Lanjouw & Uittien (1940: 148), who designated the P-JJR specimen. Given the anteriority of this lectotypification (Delprete 2015), there is no choice but to change the name of the most widespread taxon, and to adopt the oldest available name, *Amanoa grandiflora* (Müll.Arg.) Müll.Arg.

VERNACULAR NAMES. — Ka: kuata mopeli • Wp: wila tî.

HERBARIUM DATA (FG). — 35 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (W[W-1889-0012576], F[V0055866F]).

INVENTORY DATA (FG). — 31 trees in 20 plots; Fmax < 1%; dbhinv = 66.8 cm.

[1388] *Amanoa guianensis* Aubl.

*Hist. Pl. Guiane* 1: 256 [Jun.-Dec. 1775] (Aublet 1775). — *Amanoa neglecta* W.J.Hayden, *Brittonia* 42 (4): 267 (Hayden 1990), *nom. illeg. superfl.* (based on the lectotype of *A. guianensis*).

NOTES. — Known only from the Guiana Shield. As stated above [see notes under *Amanoa grandiflora* (Müll.Arg.) Müll.Arg.], the lectotypification of *A. guianensis* Aubl. by Lanjouw & Uittien (1940: 148) predates that by Hayden (1990). As a consequence, *A. neglecta* W.J.Hayden, which is based on the lectotype of *A. guianensis*, is an illegitimate superfluous name.

HERBARIUM DATA (FG). — 2 collections known from FG, none at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JJR[P00777979] designated by Lanjouw & Uittien [1940: 148]).

SIZE. — Brazil, Amapá. *N.A. Rosa & M.R. dos Santos 1834* (MO), 20 m × 48 cm (“1,50 m de circunferencia”).

Genus *Hieronyma* Allemão

[1389] *Hieronyma alchorneoides* Allemão

Hieronima alchorneoides [shortly after 30 Apr. 1848] (Allemão 1848), “*Hyeronima*”.

*Stilaginella amazonica* Tul., *Ann. Sci. Nat.*, Bot. sér. 3, 15: 241 (Tulasne 1851).

*Stilaginella laxiflora* Tul., *Ann. Sci. Nat.*, Bot. sér. 3, 15: 244 (Tulasne 1851). — *Hieronyma laxiflora* (Tul.) Müll.Arg., *Linnaea* 34 (1): 67 [pre-Mar. 1865] (Müller 1865).

*Stilaginella ferruginea* Tul., *Ann. Sci. Nat.*, Bot. sér. 3, 15: 250 (Tulasne 1851). — *Hieronyma ferruginea* (Tul.) Tul., *Fl. Bras. [Martius]* 4 (1): 334 [15 Feb. 1861] (Tulasne 1861).

*Hieronyma mollis* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 269 [late Aug. 1866] (Müller 1866).

*Hieronyma caribaea* Urb., *Repert. Spec. Nov. Regni Veg.* 16: 139 [15 Nov. 1919] (Urban 1919).

*Hieronyma heterotricha* Pax & K.Hoffm., *Pflanzenr. [Engler]* IV.147. XV (Heft 81): 39 [19 Sep. 1922] (Pax & Hoffmann 1922).

*Hieronyma mattogrossensis* Pax & K.Hoffm., *Pflanzenr. [Engler]* IV.147.XV (Heft 81): 39 [19 Sep. 1922] (Pax & Hoffmann 1922).

*Hieronyma chocoensis* Cuatrec., *Revista Acad. Colomb. Ci. Exact.* 7: 52 [Dec. 1946] (Cuatrecasas 1946).

*Hieronyma tectissima* Standl. & L.O.Williams, *Rain Forests Golfo Dulce*: 222 (Standley & Williams 1956), *nom. nud.*

*Hieronyma ovatifolia* Lundell, *Wrightia* 4 (4): 134 (Lundell 1970).

*Hieronyma alchorneoides* var. *stipulosa* P.Franco, *Bot. Jahrb. Syst.* 111 (3): 321 [27 Apr. 1990] (Franco 1990).

VERNACULAR NAMES. — Pa: waikwavia • Ka: ako, kasili wewe, tapilen katulimya, tokolo enulu • Wp: minua'i • Wn: okomë okï, palulumïi • Nt: win udu • Cr: bwa-diven • Br: aricurqua, muira-gonçalo, ururcurana.

HERBARIUM DATA (FG). — 42 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 2703*.

INVENTORY DATA (FG). — 21 trees in 14 plots; Fmax < 1%; dbhinv = 63.8 cm.

[1390] *Hieronyma oblonga* (Tul.) Müll.Arg.

*Linnaea* 34 (1): 66 [pre-Mar. 1865] (Müller 1865). — *Stilaginella oblonga* Tul., *Ann. Sci. Nat.*, Bot. sér. 3, 15: 248 (Tulasne 1851).

*Stilaginella benthamii* Tul., *Ann. Sci. Nat.*, Bot. sér. 3, 15: 247 (Tulasne 1851). — *Hieronyma oblonga* var. *benthamii* (Tul.) Müll.Arg., *Linnaea* 34 (1): 66 [pre-Mar. 1865] (Müller 1865).

*Stilaginella blanchetiana* Tul., *Ann. Sci. Nat.*, Bot. sér. 3, 15: 249 (Tulasne 1851). — *Hieronyma blanchetiana* (Tul.) Tul., *Fl. Bras. [Martius]* 4 (1): 333 [15 Feb. 1861] (Tulasne 1861). — *Hieronyma oblonga* var. *blanchetiana* (Tul.) Müll.Arg., *Linnaea* 34 (1): 66 [pre-Mar. 1865] (Müller 1865).

*Hieronyma oblonga* var. *obtusa* Müll.Arg., *Linnaea* 34 (1): 66 [pre-Mar. 1865] (Müller 1865).

*Hieronyma guatemalensis* Donn.Sm., *Bot. Gaz.* 54 (3): 241 [21 Sep. 1912] (Donnell Smith 1912).

*Hieronyma andina* Pax & K.Hoffm., *Pflanzenr. [Engler]* IV.147.XV (Heft 81): 37 [19 Sep. 1922] (Pax & Hoffmann 1922).

*Hieronyma poasana* Standl., *Publ. Field Mus. Nat. Hist.*, Bot. Ser. 18 (2): 611 [20 Oct. 1937] (Standley 1937).

*Hieronyma oblonga* var. *crassifolia* Cuatrec., *Revista Acad. Colomb. Ci. Exact.* 8 (31): 300 [Dec. 1951] (Cuatrecasas 1951).

*Hieronyma oblonga* var. *nervata* Cuatrec., *Revista Acad. Colomb. Ci. Exact.* 8 (31): 301 [Dec. 1951] (Cuatrecasas 1951).

*Hieronyma oblonga* f. *glabra* Steyerm., *Fieldiana*, Bot. 28 (4): 951 (Steyermark 1957).
*Hieronyma gentlei* Lundell, *Wrightia* 5 (7): 248 (Lundell 1976).

HERBARIUM DATA (FG). — 17 collections at CAY. Sel. exs.: *B. Riéra & D. Sabatier 1925*.

INVENTORY DATA (FG). — 32 trees in 22 plots; $F_{max} < 1\%$; $dbh_{inv} = 169.7$ cm.

Genus *Margaritaria* L.f.

[1391] *Margaritaria nobilis* L.f.
(Fig. 46B)

Suppl. Pl.: 428 [“1781” publ. Apr. 1782] (Linnaeus 1782). — *Phyllanthus nobilis* (L.f.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 414 [late Aug. 1866] (Müller 1866).

*Cicca antillana* A.Juss., *Euphorb. Gen.* 108 [21 Feb. 1824] (Jussieu 1824). — *Phyllanthus antillanus* (A.Juss.) Müll.Arg., *Linnaea* 32: 51 (Müller 1863). — *Phyllanthus nobilis* var. *antillanus* (A.Juss.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 415 [late Aug. 1866] (Müller 1866). — *Diasperus antillanus* (A.Juss.) Kuntze, *Revis. Gen. Pl.* 2: 597 [5 Nov. 1891] (Kuntze 1891). — *Margaritaria nobilis* var. *antillana* (A.Juss.) Stehlé & Quentin, *Fl. Guadeloupe* [Stehlé*] 2 (1): 47 [May 1938] (Stehlé & Quentin 1938).

*Cicca surinamensis* Miq., *Linnaea* 21: 479 (Miquel 1848).

*Cicca pavoniana* Baill., *Étude Euphorb.* 618 (Baillon 1858). — *Phyllanthus nobilis* var. *pavonianus* (Baill.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 415 [late Aug. 1866] (Müller 1866).

*Margaritaria adelioides* Rich. ex Baill., *Étude Euphorb.* 618 (Baillon 1858), *nom. nud*.

*Cicca antillana* var. *pedicellaris* Griseb., *Pl. Wright.* [*Grisebach*] 1: 158 [Dec. 1860] (Grisebach 1860).

*Phyllanthus antillanus* var. *pedicellaris* Müll.Arg., *Linnaea* 32: 51 (Müller 1863).

*Phyllanthus nobilis* var. *guyanensis* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 414 [late Aug. 1866] (Müller 1866).

*Phyllanthus nobilis* var. *peruvianus* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 414 [late Aug. 1866] (Müller 1866).

*Phyllanthus nobilis* var. *brasiliensis* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 415 [late Aug. 1866] (Müller 1866).

*Phyllanthus nobilis* var. *riedelianus* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 415 [late Aug. 1866] (Müller 1866).

*Phyllanthus nobilis* var. *martii* Müll.Arg., *Fl. Bras.* [*Martius*] 11 (2): 70 [1 Feb. 1873] (Müller 1873).

*Phyllanthus nobilis* var. *panamensis* Müll.Arg., *Fl. Bras.* [*Martius*] 11 (2): 70 [1 Feb. 1873] (Müller 1873).

*Phyllanthus nobilis* var. *hypomalacus* Standl., *Publ. Carnegie Inst. Wash.* 461: 68 (Standley 1935). — *Phyllanthus antillanus* var. *hypomalacus* (Standl.) Lundell, *Phytologia* 1 (10): 337 [27 Nov. 1939] (Lundell 1939). — *Margaritaria nobilis* var. *hypomalaca* (Standl.) Dugand, *Phytologia* 13 (6): 387 [19 Sep. 1966] (Dugand 1966).

VERNACULAR NAMES. — Br: cabelo-de-cotia, fruto-de-jacamin.

HERBARIUM DATA (FG). — 17 collections at CAY. Sel. exs.: *D. Sabatier & J.-F. Molino 4951*.

INVENTORY DATA (FG). — 7 trees in 6 plots; $F_{max} < 1\%$; $dbh_{inv} = 37.5$ cm.

Genus *Phyllanthus* L.

[1392] *Phyllanthus attenuatus* Miq.

*Linnaea* 21: 479 (Miquel 1848).

*Meborea guianensis* Aubl., *Hist. Pl. Guiane* 2: 825 [Jun.-Dec. 1775] (Aublet 1775), “Guyannensis” on plate. — *Phyllanthus guianensis* (Aubl.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 376 [late Aug. 1866] (Müller 1866), *nom. illeg. hom., non* Klotzsch (1843). — *Diasperus guianensis* (Aubl.) Kuntze, *Revis. Gen. Pl.* 2: 599 [5 Nov. 1891] (Kuntze 1891), “guyanensis”.

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *D. Larpin 670*.

SIZE. — Up to 12 m tall (Webster 2004).

[1393] *Phyllanthus juglandifolius* Willd.

* Enum. Pl. Suppl. [Willdenow]*: 64 [Jul.-Dec. 1814] (Willdenow 1814).

HERBARIUM DATA (FG). — A single collection, *R.A.A. Oldeman B-3261*.

SIZE. — Brazil, Amazonas. *G. T. Prance 10376* (MO), 15 m × 20 cm.

Genus *Richeria* Vahl

[1394] *Richeria grandis* Vahl

*Eclog. Amer.* 1: 29 (Vahl 1797).

*Amanoa divaricata* Poepp., *Nova genera ac species plantarum* [*Poep-pig & Endlicher*] 3: 22 [15-21 Aug. 1841] (Poeppig 1841). — *Richeria grandis* var. *divaricata* (Poepp.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 467 [late Aug. 1866] (Müller 1866).

*Amanoa racemosa* Poepp., *Nova genera ac species plantarum* [*Poeppig & Endlicher*] 3: 23 [15-21 Aug. 1841] (Poeppig 1841). — *Richeria grandis* var. *racemosa* (Poepp.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 467 [late Aug. 1866] (Müller 1866). — *Richeria racemosa* (Poepp.) Pax & K.Hoffm., *Pflanzenr. [Engler]* IV.147.XV (Heft 81): 28 [19 Sep. 1922] (Pax & Hoffmann 1922).

*Guarania ramiflora* Wedd. ex Baill., *Étude Euphorb.*: 598 (Baillon 1858), *nom. nud*.

*Guarania laurifolia* Baill., *Adansonia* [Baillon] 5: 348 (Baillon 1865). — *Richeria laurifolia* (Baill.) Baill., *Adansonia* [Baillon] 6: 16 (Baillon 1865). — *Richeria grandis* var. *laurifolia* (Baill.) Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 468 [late Aug. 1866] (Müller 1866).

*Richeria grandis* var. *genuina* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 468 [late Aug. 1866] (Müller 1866), *nom. inval.* (Turland et al. 2018: Art. 24.3).

*Richeria grandis* var. *obovata* Müll.Arg., *Prodr. [A. P. de Candolle]* 15 (2.2): 468 [late Aug. 1866] (Müller 1866). — *Richeria obovata* (Müll.Arg.) Pax & K.Hoffm., *Pflanzenr. [Engler]* IV.147.XV (Heft 81): 29 [19 Sep. 1922] (Pax & Hoffmann 1922).

*Richeria australis* Müll.Arg., *Fl. Bras.* [*Martius*] 11 (2): 17 [1 Feb. 1873] (Müller 1873).
<figure><img src="image_68.png" /></figure>

FIG. 45. — Opiliaceae: *A, Agonandra silvatica* Ducke (D. Sabatier & M.-F. Prévost 3065). Pentaphylacaceae: *B, Ternstroemia dentata* (Aubl.) Sw. Peraceae: *C, Pogonophora schomburgkiana* Miers ex Benth. (D. Sabatier & J.-F. Molino 4841). Piperaceae: *D, Piper aduncum* L. (M.-F. Prévost 4081). A-C, © D. Sabatier/IRD; D, © M.-F. Prévost/IRD.

*Richeria submembranacea* Steyerm., *Publ. Field Mus. Nat. Hist., Bot.* Ser. 17 (5): 419 [27 May 1938] (Steyermark 1938).

HERBARIUM DATA (FG). — 16 collections at CAY. Sel. exs.: *M.-F. Prévost 4262*.

INVENTORY DATA (FG). — 10 trees in 10 plots; Fmax < 1%; dbhinv = 63 cm.

Family PICRAMNIACEAE Fernando & Quinn
Genus *Picramnia* Sw.

[1395] *Picramnia guianensis* (Aubl.) Jans.-Jac.

Fl. Suriname 5 (1): 329 (Jansen-Jacobs 1979). — *Tariri guianensis* Aubl., *Hist. Pl. Guiane* 2 (Suppl.): 37 [Jun.-Dec. 1775] (Aublet 1775), “*Guyannensis*” on plate. — *Picramnia tariri* DC., *Prodr. [A. P. de Candolle]* 2: 66 [mid Nov. 1825] (Candolle 1825), *nom. illeg. superfl.* (based on *Tariri guianensis*).

VERNACULAR NAMES. — Pa: tarii, tarit • Ka: tariri (*fide* Aublet 1775)
• Wp: paipayo namipay • Wn: talili • Br: caferana, tariri.

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JU, P00678733, here designated; isolectotypes *fide* Howard [1983: 287]: BM, P[not seen]).

SIZE. — Brazil, Pará. *N.T. Silva* 2389 (MO), 10 cm.

[1396] *Picramnia latifolia* Tul.

Ann. Sci. Nat., Bot. sér. 3, 7: 258 (Tulasne 1847).

*Picramnia longissima* Tul., *Ann. Sci. Nat., Bot.* sér. 3, 7: 257 (Tulasne 1847).

*Picramnia umbrosa* Seem., *Bot. Voy. Herald* [Seemann] 3: 97 [Nov. 1853] (Seemann 1853).

*Picramnia macrostachys* Klotzsch ex Engl., *Fl. Bras. [Martius]* 12 (2): 238 [1 Sep. 1874] (Engler 1874).
*Picramnia eosina* J.F.Macbr., *Publ. Field Mus. Nat. Hist., Bot. Ser.* **13** (3.2): 697 (Macbride 1949).

*Picramnia cooperi* D.M.Porter, *J. Arnold Arbor.* **54** (2): 317 [26 Apr. 1973] (Porter 1973).

VERNACULAR NAMES. — Wp: paipayo namipay • Br: caferana, tariri.

HERBARIUM DATA (FG). — 16 collections at CAY. Sel. exs.: *R.A.A. Oldeman B-4007*.

INVENTORY DATA (FG). — 2 trees in 1 plot; Fmax < 1 %; dbhinv = 10 cm.

[1397] *Picramnia spruceana* Engl.

*Fl. Bras.* [*Martius*] **12** (2): 238 [1 Sep. 1874] (Engler 1874). — *Picramnia sellowii* Planch. subsp. *spruceana* (Engl.) Pirani, *Bol. Bot. Univ. São Paulo* **12**: 132 [“1990” publ. 1991] (Pirani 1991).

*Picramnia tenuis* J.F.Macbr., *Candollea* **5**: 375 (Macbride 1934).

*Picramnia krukovii* A.C.Sm., *Brittonia* **2** (2): 153 (Smith 1936).

VERNACULAR NAMES. — Wp: paipayo namipay • Br: caferana, tariri.

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: *P. Grenand 543*.

SIZE. — Up to 20 m tall (Pirani 2005).

Family PIPERACEAE Giseke
Genus *Piper* L.

[1398] *Piper aduncum* L.
(Fig. 45D)

*Sp. Pl.* **1**: 29 [1 May 1753] (Linnaeus 1753). — *Artanthe adunca* (L.) Miq., *Comm. Phytogr.* **14** (2): 49 [16-21 Mar. 1840] (Miquel 1840). — *Steffensia adunca* (L.) Kunth, *Linnaea* **13**: 633 [“1839” publ. Mar.-June 1840] (Kunth 1840).

*Piper angustifolium* Ruiz & Pav., *Flora Peruviana* **1**: 38 (Ruiz & Pavón 1798), *nom. illeg. hom.*, *non* Lam. (Lamarck 1791) [synonym of *P. consanguineum* (Kunth) Steud.]

*Piper elongatum* Vahl, *Enumer. Pl. [Vahl]* **1**: 312 [Jul.-Dec. 1804] (Vahl 1804). — *Steffensia elongata* (Vahl) Kunth, *Linnaea* **13**: 637 [“1839” publ. Mar.-June 1840] (Kunth 1840). — *Artanthe elongata* (Vahl) Miq., *Syst. Piperac.* [*F.A. W. Miquel*]: 434 [“1843” publ. 4 Apr. 1844] (Miquel 1844).

*Piper celtidifolium* Kunth, *Nova genera et species plantarum [H.B.K.]* **1**: 50 [29 Jan. 1816] (Kunth 1816). — *Steffensia celtidifolia* (Kunth) Kunth, *Linnaea* **13**: 635 [“1839” publ. Mar.-June 1840] (Kunth 1840).

*Piper purpurascens* D.Dietr., *Syn. Pl. [D. Dietrich]* **1**: 119 [July 1839] (Dietrich 1839), *nom. illeg. hom.*, *non* Desf. (Desfontaines 1829).

*Piper multinervium* M.Martens & Galeotti, *Bull. Acad. Roy. Sci. Bruxelles* **10**: 130 (Martens & Galeotti 1843).

*Artanthe galleottii* Miq., *Syst. Piperac.* [*F.A. W. Miquel*]: 451 [“1843” publ. 4 Apr. 1844] (Miquel 1844).

*Artanthe adunca* f. angustifolia Miq., *London J. Bot.* **4**: 456 (Miquel 1845). — *Artanthe celtidifolia* (Kunth) Miq., *Syst. Piperac.* [*F.A. W. Miquel*]: 452 [“1843” publ. 4 Apr. 1844] (Miquel 1844).

*Piper angustifolium* var. *cordulatum* C.DC., *Prodr.* [*A. P. de Candolle*] **16** (1): 286 [mid Nov. 1869] (Candolle 1869). — *Piper elongatum* var. *cordulatum* (C.DC.) C.DC., *Repert. Spec. Nov. Regni Veg.* **9**: 230 (Candolle 1911). — *Piper aduncum* var. *cordulatum* (C.DC.) Yunck., *Lilloa* **27**: 129 (Yuncker 1955).

*Piper pseudovelutinum* C.DC. var. *flavescens* C.DC., *Bull. Soc. Roy. Bot. Belgique* **30** (1): 203 [“1891” publ. 25 May 1892] (Candolle 1892), “pseudo-velutinum”. — *Piper flavescens* (C.DC.) Trel., *Contr. U.S. Natl. Herb.* **26** (4): 184 [31 Dec. 1929] (Trelease 1929).

*Piper kuntzei* C.DC. in Kuntze, *Revis. Gen. Pl.* **3** (3): 274 [28 Sep. 1898] (Candolle 1898).

*Piper guanaianum* C.DC., *Repert. Spec. Nov. Regni Veg.* **9**: 232 (Candolle 1911).

*Piper herzogii* C.DC., *Meded. Rijks-Herb.* **27**: 7 (Candolle 1915).

*Piper fatoanum* C.DC., *Smithsonian Misc. Collect.* **71** (6): 7 [12 Feb. 1920] (Candolle 1920).

*Piper aduncum* var. *laevifolium* C.DC., *Smithsonian Misc. Collect.* **71** (6): 8 [12 Feb. 1920] (Candolle 1920). — *Piper elongatum* var. *laevifolium* (C.DC.) Trel., *Contr. U.S. Natl. Herb.* **26** (2): 37 [21 May 1927] (Trelease 1927).

*Piper cardenasii* Trel., *Mem. New York Bot. Gard.* **7**: 223 (Trelease 1927).

*Piper elongatum* var. *brachyarthrum* Trel., *Contr. U.S. Natl. Herb.* **26** (2): 37 [21 May 1927] (Trelease 1927). — *Piper aduncum* var. *brachyarthrum* (Trel.) Yunck., *Ann. Missouri Bot. Gard.* **37**: 32 [31 Mar. 1950] (Yuncker 1950).

*Piper nonconformans* Trel., *J. Wash. Acad. Sci.* **19**: 334 (Trelease 1929).

*Piper disparispicum* Trel., *Contr. U.S. Natl. Herb.* **26** (4): 170 [31 Dec. 1929] (Trelease 1929).

*Piper aduncifolium* Trel., *Contr. U.S. Natl. Herb.* **26** (4): 171 [31 Dec. 1929] (Trelease 1929).

*Piper anguillispicum* Trel., *Contr. U.S. Natl. Herb.* **26** (4): 175 [31 Dec. 1929] (Trelease 1929), “anguillaespicum”.

*Piper oblanceolatum* Trel. var. *fragilicaule* Trel., *Contr. U.S. Natl. Herb.* **26** (4): 175 [31 Dec. 1929] (Trelease 1929).

*Piper submolle* Trel., *Contr. U.S. Natl. Herb.* **26** (4): 178 [31 Dec. 1929] (Trelease 1929).

*Piper multinervium* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* **10**: 160 (Trelease 1931), *nom. illeg. hom.*, *non* M.Martens & Galeotti (1843).

*Piper stevensonii* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* **12**: 104 (Trelease 1936), *nom. inval.* (*anglice*).

*Piper elongatifolium* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* **13** (2.1): 161 [10 June 1936] (Trelease 1936).

*Piper elongatum* var. *pampayacusum* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* **13** (2.1): 162 [10 June 1936] (Trelease 1936).

*Piper lineatum* Ruiz & Pav. var. *hirtipetiolatum* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* **13** (2.1): 184 [10 June 1936] (Trelease 1936).
<figure><img src="image_69.png" /><figcaption>FIG. 46. — Phyllanthaceae: **A**, *Amanoa grandiflora* (Müll.Arg.) Müll.Arg. (D. Sabatier & M.-F. Prévost 3007); **B**, *Margaritaria nobilis* L.f. (D. Sabatier & J.-F. Molino 4951). Polygonaceae: **C**, *Coccoloba mollis* Casar. (M.-F. Prévost & D. Sabatier 3010). Primulaceae: **D**, *Cybianthus potiae* (Mez) G.Agostini (D. Sabatier & J.-F. Molino 5545). © D. Sabatier/IRD.</figcaption></figure>

*Piper cuatrecasasii* Trel., *Trab. Mus. Nac. Ci. Nat.*, Ser. Bot. 33: 48 [30 July 1936] (Trelease 1936), "*Cuatrecasasi*", nom. inval. (anglice).

*Piper cumbricola* Trel., *Trab. Mus. Nac. Ci. Nat.*, Ser. Bot. 33: 48 [30 July 1936] (Trelease 1936), nom. inval. (anglice).

*Piper illudens* Trel., *Trab. Mus. Nac. Ci. Nat.*, Ser. Bot. 33: 50 [30 July 1936] (Trelease 1936), nom. inval. (anglice).

*Piper multinervium* var. *amplum* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (4): 347 [31 Jan. 1938] (Trelease 1938).

*Piper multinervium* var. *cayoense* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (4): 347 [31 Jan. 1938] (Trelease 1938).

*Piper multinervium* var. *flavicans* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (4): 348 [31 Jan. 1938] (Trelease 1938).

*Piper multinervium* var. *hirsuticaule* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (4): 348 [31 Jan. 1938] (Trelease 1938).

*Piper multinervium* var. *kantelolense* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (4): 348 [31 Jan. 1938] (Trelease 1938).

*Piper multinervium* var. *paralense* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (4): 348 [31 Jan. 1938] (Trelease 1938).

*Piper multinervium* var. *peracutum* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (4): 349 [31 Jan. 1938] (Trelease 1938).

*Piper multinervium* var. *productipes* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (4): 349 [31 Jan. 1938] (Trelease 1938).

*Piper multinervium* var. *protractifolium* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (4): 349 [31 Jan. 1938] (Trelease 1938).

*Piper multinervium* var. *puberulipedunculum* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (4): 349 [31 Jan. 1938] (Trelease 1938).

*Piper multinervium* var. *pubescenticaule* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (4): 350 [31 Jan. 1938] (Trelease 1938).

*Piper multinervium* var. *skutchii* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (4): 350 [31 Jan. 1938] (Trelease 1938), "*Skutchii*".

*Piper multinervium* var. *tamashense* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (4): 350 [31 Jan. 1938] (Trelease 1938).

*Piper multinervium* var. *telanum* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (4): 350 [31 Jan. 1938] (Trelease 1938).

*Piper intersitum* Trel., *Caldasia* 1 (1): 86 (Trelease 1940).

*Piper intersitum* f. *porcecitense* Trel., *Caldasia* 1 (1): 87 (Trelease 1940).

*Piper reciprocum* Trel., *Ciencia (Mexico)* 2 (5): 207 [25 May 1941] (Trelease 1941).

VERNACULAR NAMES. — Wn: pëtpëjot • Nt: gaanman udu ana • Br: pimenta-longa.
HERBARIUM DATA (FG). — 36 collections at CAY. Sel. exs.: *M.-F. Prévost 3567*.

SIZE. — Up to 10 m tall (Steyermark & Callejas-Posada 2003).

[1399] *Piper cernuum* Vell.

Fl. Flumin. 25 [“1825” publ. 7 Sep.-28 Nov. 1829] (Vellozo 1829). — *Artanthe cernua* (Vell.) C.Presl, *Abh. Königl. Böhm. Ges. Wiss.*, ser. 5, 6: 585 (Presl 1851).

*Steffensia eximia* Kunth, *Linnaea* 13: 665 [“1839” publ. Mar.-June 1840] (Kunth 1840). — *Piper eximium* Kunth, *Linnaea* 13: 665 [“1839” publ. Mar.-June 1840] (Kunth 1840), *nom. nud. pro syn.* — *Artanthe eximia* (Kunth) Miq., *Syst. Piperac. [F.A. W. Miquel]*: 393 [“1843” publ. 4 Apr. 1844] (Miquel 1844). — *Piper obliquum* Ruiz & Pav. var. *eximium* (Kunth) C.DC., *Prodr. [A. P. de Candolle]* 16 (1): 307 [mid Nov. 1869] (Candolle 1869).

*Steffensia coccoloboides* Kunth, *Linnaea* 13: 666 [“1839” publ. Mar.-June 1840] (Kunth 1840). — *Piper coccoloboides* Kunth, *Linnaea* 13: 666 [“1839” publ. Mar.-June 1840] (Kunth 1840), *nom. nud. pro syn.*

*Steffensia richardiifolia* Kunth, *Linnaea* 13: 668 [“1839” publ. Mar.-June 1840] (Kunth 1840), “*richardiaefolia*”. — *Piper richardiifolium* Kunth, *Linnaea* 13: 668 [“1839” publ. Mar.-June 1840] (Kunth 1840), “*richardiaefolium*”, *nom. nud. pro syn.* — *Artanthe richardiifolia* (Kunth) Miq., *Syst. Piperac. [F.A. W. Miquel]* 395 [“1843” publ. 4 Apr. 1844] (Miquel 1844), “*richardiaefolia*”.

*Artanthe spectabilis* Miq., *Linnaea* 20: 138 [17-18 May 1847] (Miquel 1847).

*Artanthe spectabilis* var. *glabrior* Miq., *Linnaea* 20: 139 [17-18 May 1847] (Miquel 1847).

*Piper bolivianum* C.DC., *Prodr. [A. P. de Candolle]* 16 (1): 280 [mid Nov. 1869] (Candolle 1869).

*Piper ovalifolium* C.DC., *Prodr. [A. P. de Candolle]* 16 (1): 305 [mid Nov. 1869] (Candolle 1869).

*Piper cabellense* C.DC., *Prodr. [A. P. de Candolle]* 16 (1): 306 [mid Nov. 1869] (Candolle 1869).

*Piper gigantifolium* C.DC., *Prodr. [A. P. de Candolle]* 16 (1): 306 [mid Nov. 1869] (Candolle 1869).

*Piper araguense* Trel. & Yunck., *Piperac. N. South Amer.* 1: 138 (Trelease & Yuncker 1950). — *Piper cernuum* var. *araguense* (Trel. & Yunck.) Steyerm., *Fl. Venez.* 2 (2): 372 (Steyermark 1984).

*Piper arbelaezii* Trel. & Yunck. var. *subglabrirameum* Yunck., *Fiel-diana, Bot.* 28 (1): 204 (Yuncker 1951).

*Piper perlongispicum* Yunck., *Bol. Soc. Venez. Ci. Nat.* 23 (101): 89 (Yuncker 1962). — *Piper cernuum* var. *perlongispicum* (Yunck.) Steyerm., *Fl. Venez.* 2 (2): 372 (Steyermark 1984).

*Piper cernuum* var. *glabricaule* Yunck., *Bol. Inst. Bot. (São Paulo)* 3: 20 [23 Dec. 1966] (Yuncker 1966).

*Piper cernuum* var. *biformipilum* Yunck., *Bol. Inst. Bot. (São Paulo)* 3: 21 [23 Dec. 1966] (Yuncker 1966).

NOTES. — Kunth published simultaneously *Piper coccoloboides* and *Steffensia coccoloboides* (Kunth 1840: 666) as well as *P. eximium* and *S. eximia* (Kunth 1840: 665), and *P. richardiifolium* and *S. richardiifolia* (Kunth 1840: 668).

As these names were published before 1953, all of them are valid; hence both *Artanthe eximia* (Kunth) Miq. and *A. richardiifolia* (Kunth) Miq. have alternative basionyms, respectively, *P. eximium* Kunth and *S. eximia* Kunth, and *P. richardiifolium* Kunth and *S. richardiifolia* Kunth.

VERNACULAR NAMES. — Wp: *taīwī* lenipi’ā, yali taku’ā sili.

HERBARIUM DATA (FG). — 5 collections at CAY. Sel. exs.: *J.-J. de Granville B-5436*.

SIZE. — Up to 12 m tall (Steyermark & Callejas-Posada 2003).

[1400] *Piper reticulatum* L.

Sp. Pl. 1: 29 [1 May 1753] (Linnaeus 1753). — *Enckea reticulata* (L.) Miq., *Syst. Piperac. [F.A. W. Miquel]*: 365 [“1843” publ. 4 Apr. 1844] (Miquel 1844). — *Discipiper reticulatum* (L.) Trel. & Stehlé, *Candollea* 10: 283 (Trelease & Stehlé 1946).

*Piper smilacifolium* Kunth, *Nova genera et species plantarum [H.B.K.]* 1: 56 [29 Jan. 1816] (Kunth 1816). — *Enckea smilacifolia* (Kunth) Kunth, *Linnaea* 13: 605 [“1839” publ. Mar.-June 1840] (Kunth 1840), “*Enckia*”.

*Piper latum* Kunth, *Nova genera et species plantarum [H.B.K.]* 1: 57 [29 Jan. 1816] (Kunth 1816). — *Enckea lata* (Kunth) Kunth, *Linnaea* 13: 606 [“1839” publ. Mar.-June 1840] (Kunth 1840), “*Enckia*”. — *Macropiper latum* (Kunth) C.Presl, *Abh. Königl. Böhm. Ges. Wiss.*, ser. 5, 6: 583 (Presl 1851).

*Artanthe ruiziana* Miq., *London J. Bot.* 4: 444 (Miquel 1845).

*Piper tarapotianum* C.DC., *Prodr. [A. P. de Candolle]* 16 (1): 246 [mid Nov. 1869] (Candolle 1869).

*Piper duchassaingii* C.DC., *Prodr. [A. P. de Candolle]* 16 (1): 251 [mid Nov. 1869] (Candolle 1869).

*Piper pangoense* C.DC., *Prodr. [A. P. de Candolle]* 16 (1): 368 [mid Nov. 1869] (Candolle 1869).

*Piper discophorum* C.DC., *Bull. Soc. Roy. Bot. Belgique* 30 (1): 201 [“1891” publ. 25 May 1892] (Candolle 1892).

*Piper reticulatum* var. *santarosanum* Trel., *Publ. Field Mus. Nat. Hist., Bot. Ser. 13* (2.1): 223 [10 June 1936] (Trelease 1936), “*santa-rosanum*”.

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *C. Feuillet 616*.

SIZE. — Up to 15 m tall (Görts van Rijn 2007).

Family POLYGONACEAE Juss.
Genus *Coccoloba* P.Browne

[1401] *Coccoloba latifolia* Poir.

Encycl. [J. Lamarck et al.] 6 (1): 61 [2 Oct. 1804] (Poiret 1804). — *Uvifera latifolia* (Poir.) Kuntze, *Revis. Gen. Pl.* 2: 561 [5 Nov. 1891] (Kuntze 1891).

*Coccoloba rheifolia* Desf., *Tabl. École Bot.*, ed. 2, 46 (Desfontaines 1815).
*Coccoloba grandis* Benth., *London J. Bot.* 4: 624 (Bentham 1845).

VERNACULAR NAMES. — Ka: matula, patula • Br: cabeça-de-macaco.

HERBARIUM DATA (FG). — 14 collections at CAY. Sel. exs.: *M.-F. Prévost 1579*.

SIZE. — Up to 15 m tall (Aymard C. & Howard 2004).

[1402] *Coccoloba mollis* Casar.
(Fig. 46C)

*Nov. Stirp. Bras.* 8: 72 [June 1844] (Casaretto 1844). — *Coccoloba polystachya* var. *mollis* (Casar.) Meisn., *Prodr. [A. P. de Candolle]* 14 (1): 151 [mid Oct. 1856] (Meisner 1856).

*Coccoloba polystachya* Wedd., *Ann. Sci. Nat., Bot.* sér. 3, 13: 261 (Weddell 1849). — *Uvifera polystachya* (Wedd.) Kuntze, *Revis. Gen. Pl.* 2: 561 [5 Nov. 1891] (Kuntze 1891).

*Coccoloba paniculata* Meisn., *Fl. Bras. [Martius]* 5 (1): 43 [1 Jan. 1855] (Meisner 1855).

*Coccoloba polystachya* var. *glabra* Lindau, *Bot. Jahrb. Syst.* 13 (2): 133 [28 Oct. 1890] (Lindau 1890).

*Coccoloba polystachya* var. *pubescens* Lindau, *Bot. Jahrb. Syst.* 13 (2): 133 [28 Oct. 1890] (Lindau 1890).

*Coccoloba polystachya* var. *jamaicensis* Fawc. & Rendle, *J. Bot.* 51: 125 (Fawcett & Rendle 1913).

*Coccoloba dugandiana* A.Fernández, *Mutisia* 5: 1 (Fernández 1952).

*Coccoloba standleyana* P.H.Allen, *Rain Forests Golfo Dulce* 409 (Allen 1956).

VERNACULAR NAMES. — Pa: tivu-arigni • Ka: matula, patula • Nt: doon udu, doonudu.

HERBARIUM DATA (FG). — 14 collections at CAY. Sel. exs.: *M.-F. Prévost & D. Sabatier 3010*.

INVENTORY DATA (FG). — 53 trees in 30 plots; $F_{max} < 1\%$; $dbh_{inv} = 38.4$ cm.

[1403] *Coccoloba uvifera* (L.) L.

*Syst. Nat.*, ed. 10, 2: 1007 [7 June 1759] (Linnaeus 1759). — Polygonum uvifera L., *Sp. Pl.* 1: 365 [1 May 1753] (Linnaeus 1753). — *Guaiabara uvifera* (L.) House, *Amer. Midl. Naturalist* 8: 64 (House 1922).

NOTE. — Restricted to coastal sandy beaches.

VERNACULAR NAMES. — Cr: rézen-bòdlanmé.

HERBARIUM DATA (FG). — 5 collections at CAY. Sel. exs.: *H. de Foresta 760*.

SIZE. — Panama, Colón, 17 May 1974. *M.H. Nee 11694* (MO), 7 m × 25 cm.

Genus *Ruprechtia* C.A.Mey.

[1404] *Ruprechtia brachysepala* Meisn.

*Fl. Bras. [Martius]* 5 (1): 57 [1 Jan. 1855] (Meisner 1855). — *Magonia brachysepala* (Meisn.) Kuntze, *Revis. Gen. Pl.* 2: 553 [5 Nov. 1891] (Kuntze 1891).

*Ruprechtia amentacea* Meisn., *Fl. Bras. [Martius]* 5 (1): 56 [1 Jan. 1855] (Meisner 1855). — *Magonia amentacea* (Meisn.) Kuntze, *Revis. Gen. Pl.* 2: 553 [5 Nov. 1891] (Kuntze 1891).

*Ruprechtia latifolia* Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (2): 345 (Huber 1909).

*Ruprechtia marowynensis* Eyma, *Recueil Trav. Bot. Néerl.* 32: 224 (Eyma 1935).

*Ruprechtia nitida* Brandbyge, *Nordic J. Bot.* 10 (2): 158 (Brandbyge 1990).

VERNACULAR NAMES. — Nt: pindya.

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *R.A.A. Oldeman B-3358*.

SIZE. — Up to 12 m tall (Pendry 2004).

Genus *Triplaris* Loefl.

[1405] *Triplaris americana* L.

*Syst. Nat.*, ed. 10, 2: 881 [7 June 1759] (Linnaeus 1759), “americ.”

*Triplaris pyramidalis* Jacq., *Select. Stirp. Amer. Hist.* 13 [5 Jan. 1763] (Jacquin 1763).

*Triplaris brasiliana* Cham., *Linnaea* 8: 139 (Chamisso 1833).

*Triplaris schomburgkiana* Benth., *London J. Bot.* 4: 628 (Bentham 1845).

*Triplaris felipensis* Wedd., *Ann. Sci. Nat., Bot.* sér. 3, 13: 263 (Weddell 1849).

*Triplaris noli-tangere* Wedd., *Ann. Sci. Nat., Bot.* sér. 3, 13: 264 (Weddell 1849).

*Ruprechtia martii* Meisn., *Fl. Bras. [Martius]* 5 (1): 58 [1 Jan. 1855] (Meisner 1855).

*Triplaris pavonii* Meisn., *Prodr. [A. P. de Candolle]* 14 (1): 172 [mid Oct. 1856] (Meisner 1856).

*Triplaris formicana* S.Moore, *Trans. Linn. Soc. London, Bot.* 4 (3): 444 [“1894-96” publ. Dec. 1895] (Moore 1895).

*Triplaris guanaiensis* Rusby, *Mem. Torrey Bot. Club* 6 (1): 111 (Rusby 1896).

*Triplaris estriata* Kuntze, *Revis. Gen. Pl.* 3 (3): 271 [28 Sep. 1898] (Kuntze 1898).

*Triplaris boliviana* Britton, *Bull. Torrey Bot. Club* 27 (3): 130 (Britton 1900).

*Triplaris euryphylla* S.F.Blake, *Contr. U.S. Natl. Herb.* 20 (7): 239 [9 Sep. 1919] (Blake 1919).
*Triplaris laxa* S.F.Blake, *Contr. U.S. Natl. Herb.* 20 (7): 240 [9 Sep. 1919] (Blake 1919).

*Triplaris williamsii* Rusby, *Mem. New York Bot. Gard.* 7: 235 (Rusby 1927).

HERBARIUM DATA (FG). — Not yet collected, but the presence of this very widespread species is expected.

SIZE. — Up to 27 m tall (Aymard C. & Howard 2004).

[1406] *Triplaris weigeltiana* (Rchb.) Kuntze

*Revis. Gen. Pl.* 3 (3): 270 [28 Sep. 1898] (Kuntze 1898). — *Blochmannia weigeltiana* Rchb., *Surinam Exsic. [Weigelt] s.n.* (Reichenbach 1828).

*Triplaris surinamensis* Cham., *Linnaea* 8: 138 (Chamisso 1833).

*Triplaris vahliana* Fisch. & C.A.Mey. ex C.A.Mey., *Mém. Acad. Imp. Sci. Saint-Pétersbourg*, Sér. 6, *Sci. Math., Seconde Pt. Sci. Nat.* 6 (2, Bot.): 148 [Nov. 1840] (Meyer 1840).

*Triplaris surinamensis* var. *crassifolia* Benth., *London J. Bot.* 4: 628 (Bentham 1845).

*Triplaris martiana* Fisch. & C.A.Mey. ex C.A.Mey. var. *oblongifolia* Meisn., *Fl. Bras. [Martius]* 5 (1): 50 [1 Jan. 1855] (Meisner 1855).

*Triplaris surinamensis* var. *benthamiana* Meisn., *Fl. Bras. [Martius]* 5 (1): 50 [1 Jan. 1855] (Meisner 1855), "*Benthamiana*".

*Triplaris surinamensis* var. *chamissoana* Meisn., *Fl. Bras. [Martius]* 5 (1): 50 [1 Jan. 1855] (Meisner 1855).

*Triplaris siphonopetala* H.Gross, *Bot. Jahrb. Syst.* 49 (2): 347 [14 Jan. 1913] (Gross 1913).

NOTES. — Hyperdominant in Amazonia (ter Steege *et al.* 2020). Weigelt's *Surinam exscicatae* were sent to several herbaria in 1828, together with printed labels bearing full descriptions (Stafleu & Cowan 1988). Therefore, all requirement are met for the effective publication of *Blochmannia weigeltiana* Rchb. (see Turland *et al.* 2018: Art. 30.8, Ex. 13).

VERNACULAR NAMES. — Pa: ã-wareuni • Ka: tasi • Wp: moyu tasi • Wn: mekulak, mukulap • Nt: doon udu, doonudu • Cr: bwa-fronmi • Br: taxi-preto-da-várzea, taxiseiro.

HERBARIUM DATA (FG). — 26 collections at CAY. Sel. exs.: *M.-F. Prévost 750*.

INVENTORY DATA (FG). — 1 tree, dbh = 48 cm.

Family PRIMULACEAE Batsch ex Borkh.
Genus *Cybianthus* Mart.

[1407] *Cybianthus amplus* (Mez) G.Agostini

*Acta Biol. Venez.* 10 (2): 152 (Agostini 1980). — *Conomorpha ampla* Mez, *Pflanzenr.* [Engler] IV.236 (Heft 9): 257 [6 May 1902] (Mez 1902).

*Conomorpha macrophylla* Mart., *Flora* 24 (2, Beibl.): 20 [28 Aug. 1841] (Martius 1841).

*Conomorpha utiarityi* Hoehne, *Comm. Lin. Télegr., Bot.* 6: 64 [Sep. 1915] (Hoehne 1915).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *R.A.A. Oldeman T-909*.

SIZE. — Brazil, Amazonas. *C.A. Cid Ferreira 7600* (MO), 15 m × 25 cm.

[1408] *Cybianthus guyanensis* (A.DC.) Miq. subsp. *multipunctatus* (A.DC.) Pipoly

*Ann. Missouri Bot. Gard.* 79 (4): 944 (Pipoly 1992). — *Cybianthus multipunctatus* A.DC., *Ann. Sci. Nat., Bot.* sér. 2, 16: 94 (Candolle 1841). — *Conomorpha multipunctata* (A.DC.) Miq., *Stirp. Surinam. Select.*: 111 [“1850” publ. Mar. 1851] (Miquel 1851).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — 27 collections at CAY. Sel. exs.: *J. Martin s.n.* (original material G-DC, G00139364)].

INVENTORY DATA (FG). — 1 tree, dbh = 19.4 cm.

[1409] *Cybianthus leprieurii* G.Agostini

*Acta Biol. Venez.* 10 (2): 159 (Agostini 1980). — *Weigeltia parviflora* Mez, *Pflanzenr.* [Engler] IV.236 (Heft 9): 288 [6 May 1902] (Mez 1902).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — 5 collections at CAY. Sel. exs.: *F.M.R. Leprieur s.n.* (original material P[P00715205]); *R.A.A. Oldeman B-521*, height 11 m.

[1410] *Cybianthus microbotrys* A.DC.

*Ann. Sci. Nat., Bot.* sér. 2, 16: 94 (Candolle 1841). — *Peckia microbotrys* (A.DC.) Kuntze, *Revis. Gen. Pl.* 2: 402 [5 Nov. 1891] (Kuntze 1891). — *Weigeltia microbotrys* (A.DC.) Mez, *Pflanzenr.* [Engler] IV.236 (Heft 9): 285 [6 May 1902] (Mez 1902).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Pa: pune-etni-kamwi • Wp: ka'a pelele • Br: capororoquinha.

HERBARIUM DATA (FG). — 45 collections at CAY. Sel. exs.: *S.A. Mori & J.J. Pipoly 15528*.

INVENTORY DATA (FG). — 4 trees in 4 plots; $F_{max} < 1\%$; dbh$_{inv}$ = 14.5 cm.

[1411] *Cybianthus potiae* (Mez) G.Agostini
(Fig. 46D)

*Acta Biol. Venez.* 10 (2): 161 (Agostini 1980). — *Weigeltia potiae* Mez, *Pflanzenr.* [Engler] IV.236 (Heft 9): 285 [6 May 1902] (Mez 1902).

HERBARIUM DATA (FG). — 11 collections at CAY. Sel. exs.: *P.A. Poiteau s.n.* (original material P[P00715218]).

INVENTORY DATA (FG). — 1 tree, dbh = 11.8 cm.
[1412] *Cybianthus prieurii* A.DC.

Ann. Sci. Nat., Bot. sér. 2, 16: 93 (Candolle 1841), "Prieurei". — *Peckia prieurii* (A.DC.) Kuntze, Revis. Gen. Pl. 2: 402 [5 Nov. 1891] (Kuntze 1891).

*Cybianthus nitidus* Miq., Fl. Bras. [Martius] 10: 295 [15 Mar. 1856] (Miquel 1856). — *Peckia nitida* (Miq.) Kuntze, Revis. Gen. Pl. 2: 402 [5 Nov. 1891] (Kuntze 1891).

*Cybianthus subspicatus* Benth. ex Miq., Fl. Bras. [Martius] 10: 296 [15 Mar. 1856] (Miquel 1856). — *Peckia subspicata* (Benth. ex Miq.) Kuntze, Revis. Gen. Pl. 2: 402 [5 Nov. 1891] (Kuntze 1891).

*Cybianthus comatus* Mez, Pflanzenr. [Engler] IV.236 (Heft 9): 219 [6 May 1902] (Mez 1902).

*Cybianthus viridiflorus* A.C.Sm., Lloydia 2 (3): 204 (Smith 1939).

HERBARIUM DATA (FG). — 5 collections at CAY. Sel. exs.: N. Marshall & J. Rombold 146.

INVENTORY DATA (FG). — 2 trees in 1 plot; Fmax < 1%; dbhinv = 12.7 cm.

[1413] *Cybianthus resinosus* Mez

Pflanzenr. [Engler] IV.236 (Heft 9): 219 [6 May 1902] (Mez 1902).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: J.-J. de Granville et al. 8915.

SIZE. — Up to 15 m tall (Pipoly & Gustafsson 2002).

[1414] *Cybianthus surinamensis* (A.Spreng.) G.Agostini

Acta Biol. Venez. 10 (2): 161 (Agostini 1980). — *Salvadora surinamensis* A.Spreng., *Tent. Suppl.*: 7 [Sep. 1828] (Sprengel 1828). — *Peckia surinamensis* (A.Spreng.) Kuntze, *Revis. Gen. Pl.* 2: 402 [5 Nov. 1891] (Kuntze 1891). — *Weigeltia surinamensis* (A.Spreng.) Mez, *Pflanzenr.* [Engler] IV.236 (Heft 9): 289 [6 May 1902] (Mez 1902).

*Weigeltia myrianthos* A.DC., Trans. Linn. Soc. London 17 (2): 102 ["1837" publ. 26 Apr.-8 May 1834] (Candolle 1834). — *Wallenia myrianthos* Rchb. ex A.DC., Trans. Linn. Soc. London 17 (2): 102 ["1837" publ. 26 Apr.-8 May 1834] (Candolle 1834), nom. nud. pro syn. — *Cybianthus myrianthos* (A.DC.) Miq., Fl. Bras. [Martius] 10: 300 [15 Mar. 1856] (Miquel 1856).

*Weigeltia capitellata* Miq., Fl. Bras. [Martius] 10: 300 [15 Mar. 1856] (Miquel 1856), nom. nud. pro syn.

*Weigeltia schomburgkiana* Mez, Pflanzenr. [Engler] IV.236 (Heft 9): 285 [6 May 1902] (Mez 1902). — *Cybianthus schomburgkianus* (Mez) G.Agostini, *Acta Biol. Venez.* 10 (2): 165 (Agostini 1980).

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: J.-J. de Granville B-4467.

INVENTORY DATA (FG). — 3 trees in 3 plots; Fmax < 1%; dbhinv = 13.8 cm.

[1415] *Cybianthus venezuelanus* Mez

Pflanzenr. [Engler] IV.236 (Heft 9): 221 [6 May 1902] (Mez 1902).

*Cybianthus egensis* Mez, Pflanzenr. [Engler] IV.236 (Heft 9): 222 [6 May 1902] (Mez 1902).

*Peckia purpurea* Rusby, *Bull. New York Bot. Gard.* 4 (14): 405 [7 Dec. 1907] (Rusby 1907).

*Cybianthus brownii* Gleason, *Bull. Torrey Bot. Club* 53 (5): 293 [May 1926] (Gleason 1926).

VERNACULAR NAMES. — Wp: wila ka'alulu wipë'i.

HERBARIUM DATA (FG). — 35 collections at CAY. Sel. exs.: R.A.A. Oldeman B-3045.

SIZE. — Brazil, Acre. C.A. Cid Ferreira 10202 (MO), 10 m × 25 cm.

[1416] *Cybianthus* sp. A

HERBARIUM DATA (FG). — A single collection, J.-F. Molino et al. 2130.

INVENTORY DATA (FG). — 1 tree, dbh = 13.1 cm.

## Genus *Myrsine* L.

[1417] *Myrsine guianensis* (Aubl.) Kuntze

*Revis. Gen. Pl.* 2: 402 [5 Nov. 1891] (Kuntze 1891). — *Rapanea guianensis* Aubl., *Hist. Pl. Guiane* 1: 121 [Jun.-Dec. 1775] (Aublet 1775). — *Myrsine rapanea* Roem. & Schult., *Syst. Veg. [Roemer & Schultes]* 4: 509 [Mar.-June 1819] (Roemer & Schultes 1819), nom. illeg. superfl. (based on *Rapanea guianensis*). — *Myrsine rapanea* f. communis Miq., Fl. Bras. [Martius] 10: 308 [15 Mar. 1856] (Miquel 1856), nom. illeg. superfl. (based on *Rapanea guianensis*).

*Myrsine rapanea* var. major Hook. & Arn., *J. Bot. [Hooker]* 1: 283 (Hooker & Arnott 1834).

*Myrsine rapanea* var. minor Hook. & Arn., *J. Bot. [Hooker]* 1: 283 (Hooker & Arnott 1834).

*Myrsine rapanea* var. subacuta A.DC., *Prodr. [A. P. de Candolle]* 8: 98 [mid Mar. 1844] (Candolle 1844).

*Myrsine rapanea* f. angustata Miq., Fl. Bras. [Martius] 10: 308 [15 Mar. 1856] (Miquel 1856).

*Myrsine rapanea* f. paviflora Miq., Fl. Bras. [Martius] 10: 308 [15 Mar. 1856] (Miquel 1856).

*Myrsine rapanea* f. robusta Miq., Fl. Bras. [Martius] 10: 308 [15 Mar. 1856] (Miquel 1856). — *Rapanea oblonga* Pohl ex Miq., Fl. Bras. [Martius] 10: 308 [15 Mar. 1856] (Miquel 1856), nom. nud. pro syn.

*Myrsine ovalifolia* Miq., Fl. Bras. [Martius] 10: 313 [15 Mar. 1856] (Miquel 1856), nom. illeg. hom., non (A.DC.) D.Dietr. (Dietrich 1839).

*Rapanea guianensis* var. andicola Cuatrec., *Revista Acad. Colomb. Ci. Exact.* 8 (31): 325 [Dec. 1951] (Cuatrecasas 1951), "guyanensis".

*Rapanea guianensis* var. bogotensis Cuatrec., *Revista Acad. Colomb. Ci. Exact.* 8 (31): 325 [Dec. 1951] (Cuatrecasas 1951), "guyanensis".

*Rapanea guianensis* var. calensis Cuatrec., *Revista Acad. Colomb. Ci. Exact.* 8 (31): 325 [Dec. 1951] (Cuatrecasas 1951), "guyanensis".
NOTE. — Roemer and Schultes mistakenly ascribed the binomial *Myrsine rapanea* to R.Br., referencing Brown (1810: 533). There Brown simply (and incorrectly) synonymised *Rapanea* Aubl. (i.e. *R. guianensis*) under *Samara floribunda* Willd.

VERNACULAR NAMES. — Ka: kunapolan, kusapoi.

HERBARIUM DATA (FG). — 19 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JJR[P00777907] designated by Lanjouw & Uittien [1940: 156]).

INVENTORY DATA (FG). — 3 trees in 3 plots; $F_{max} < 1\%$; $dbh_{inv} = 15.4$ cm.

Genus *Stylogyne* A.DC.

[1418] *Stylogyne orinocensis* (Kunth) Mez

*Pflanzenr.* [Engler] IV.236 (Heft 9): 270 [6 May 1902] (Mez 1902).
*Ardisia orinocensis* Kunth, *Nova genera et species plantarum [H.B.K.]* 3: 244 [9 July 1819] (Kunth 1819).
*Tinus orinocensis* (Kunth) Kuntze, *Revis. Gen. Pl.* 2: 974 [5 Nov. 1891] (Kuntze 1891).

*Badula schomburgkiana* A.DC., *Ann. Sci. Nat., Bot.* sér. 2, 16: 90 (Candolle 1841).
*Ardisia schomburgkiana* (A.DC.) Miq., *Fl. Bras. [Martius]* 10: 289 [15 Mar. 1856] (Miquel 1856).
*Tinus schomburgkiana* (A.DC.) Kuntze, *Revis. Gen. Pl.* 2: 975 [5 Nov. 1891] (Kuntze 1891).
*Stylogyne schomburgkiana* (A.DC.) Mez, *Pflanzenr. [Engler]* IV.236 (Heft 9): 270 [6 May 1902] (Mez 1902).

*Badula schomburgkiana* var. *brasiliensis* A.DC., *Prodr.* [*A. P. de Candolle*] 8: 111 [mid Mar. 1844] (Candolle 1844).
*Stylogyne brasiliensis* (A.DC.) Mez, *Pflanzenr.* [Engler] IV.236 (Heft 9): 269 [6 May 1902] (Mez 1902).

*Ardisia hostmannii* Miq., *Fl. Bras. [Martius]* 10: 288 [15 Mar. 1856] (Miquel 1856), "hostmanni".
*Tinus hostmannii* (Miq.) Kuntze, *Revis. Gen. Pl.* 2: 974 [5 Nov. 1891] (Kuntze 1891).

*Ardisia surinamensis* Miq., *Fl. Bras. [Martius]* 10: 288 [15 Mar. 1856] (Miquel 1856).
*Tinus surinamensis* (Miq.) Kuntze, *Revis. Gen. Pl.* 2: 975 [5 Nov. 1891] (Kuntze 1891).
*Stylogyne surinamensis* (Miq.) Mez, *Pflanzenr.* [Engler] IV.236 (Heft 9): 271 [6 May 1902] (Mez 1902).

*Stylogyne micans* Mez, *Pflanzenr.* [Engler] IV.236 (Heft 9): 272 [6 May 1902] (Mez 1902).

*Stylogyne poeppigii* Mez, *Pflanzenr.* [Engler] IV.236 (Heft 9): 274 [6 May 1902] (Mez 1902).

*Stylogyne amazonica* Mez, *Pflanzenr.* [Engler] IV.236 (Heft 9): 275 [6 May 1902] (Mez 1902).

*Stylogyne koppleri* Mez, *Pflanzenr.* [Engler] IV.236 (Heft 9): 275 [6 May 1902] (Mez 1902).

VERNACULAR NAMES. — Pa: pune-etni-priye (for *Stylogyne* sp.) • Ka: payawalu.

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *C. Moretti 606*.

SIZE. — Up to 14 m tall (Ricketson & Pipoly 2010).

Family PROTEACEAE Juss.
Genus *Euplassa* Salisb. ex Knight

[1419] *Euplassa pinnata* (Lam.) I.M.Johnst.

*Contr. Gray Herb.* 73: 42 (Johnston 1924).
*Roupala pinnata* Lam., *Tabl. Encycl.* 1[1 (2)]: 243 [13 Feb. 1792] (Lamarck 1792).
*Adenostephanus guyanensis* Meisn., *Fl. Bras. [Martius]* 5 (1): 95 [1 Jan. 1855] (Meisner 1855), *nom. illeg. superfl.* (based on *Roupala pinnata*).

*Euplassa meridionalis* Salisb. ex Knight, Cult. Prot. 101 [Dec. 1809] (Knight 1809).

VERNACULAR NAMES. — Wp: tatu kāsī • Nt: liba apisi • Cr: bwa-graj • Fr: bois grage blanc • Br: faeira, louro-faia.

HERBARIUM DATA (FG). — 10 collections at CAY. Sel. exs.: *J. Martin s.n.* (original material BM[BM000080820, BM000080821, BM000080822]).

INVENTORY DATA (FG). — 17 trees in 16 plots; $F_{max} < 1\%$; $dbh_{inv} = 42$ cm.

Genus *Panopsis* Salisb. ex Knight

[1420] *Panopsis rubescens* (Pohl) Rusby

*Mem. Torrey Bot. Club* 6 (1): 116 (Rusby 1896).
*Andriapetalum rubescens* Pohl, *Pl. Bras. Icon. Descr.* 1 (4): 114 [May-Dec. 1828] (Pohl 1828).

*Andriapetalum rubescens* var. *acuminatum* Meisn., *Prodr.* [*A. P. de Candolle*] 14 (1): 346 [mid Oct. 1856] (Meisner 1856), "*Andri-petalum*".
*Panopsis acuminata* (Meisn.) J.F.Macbr., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 11 (2): 66 (Macbride 1931).

*Andriapetalum sprucei* Meisn., *Prodr.* [*A. P. de Candolle*] 14 (1): 347 [mid Oct. 1856] (Meisner 1856).
*Panopsis rubescens* var. *sprucei* (Meisn.) Sleumer, *Bot. Jahrb. Syst.* 76 (2): 180 [Jan. 1954] (Sleumer 1954).

*Roupala yauaperyensis* Barb.Rodr., *Vellovia*, ed. 2, 1: 66 (Barbosa Rodrigues 1891).

*Panopsis sprucei* Rusby, *Mem. Torrey Bot. Club* 6 (1): 116 (Rusby 1896), *nom. illeg. superfl.*

*Panopsis rubescens* var. *simulans* J.F.Macbr., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 11 (2): 67 (Macbride 1931).

*Panopsis cuaensis* Steyerm., *Ann. Missouri Bot. Gard.* 74 (3): 612 (Steyermark 1987).

NOTES. — Rusby (1896: 116) did not cite *Andriapetalum sprucei* Meisn. However, he ascribed the binomial "*Panopsis sprucei*" to "Meissn. Ms. in Herb. Kew", while stating that the original material (*Bang 1686* [NY, 01364973]) is "the same as Spruce's no. 1817". *Spruce 1817* is the type of *Andriapetalum sprucei*, and the K specimens (K000634341, K000634342)) bear the handwritten annotation (not in Meissner's hand) "*Andripetalum Sprucei* Meisn.", which is presumably what Rusby was referring to. As Meisner did not use *Panopsis sprucei*, the authorship of this name is "Rusby", not "Meisn. ex Rusby". Furthermore, *P. sprucei* is an illegitimate superfluous name because although independent from *A. sprucei*, the type of the latter is cited in the protologue.
HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: S.A. Mori et al. 20968.

INVENTORY DATA (FG). — 8 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 54.1$ cm.

[1421] *Panopsis sessilifolia* (Rich.) Sandwith (Fig. 47A)

Bull. Misc. Inform. Kew 1932 (5): 226 [27 June 1932] (Sandwith 1932). — *Roupala sessilifolia* Rich., *Actes Soc. Hist. Nat. Paris* 1: 106 [Oct. 1792] (Richard 1792). — *Andriapetalum sessilifolium* (Rich.) Klotzsch, *Linnaea* 15: 53 (Klotzsch 1841), “*Andripetalum sessilifolia*”.

Roupala hameliifolia Rudge, Pl. Guian. [Rudge] 1 (3): 22 [June 1805] (Rudge 1805), “hameliaefolia”. — *Panopsis hameliifolia* (Rudge) Knight, Cult. Prot.: 104 [Dec. 1809] (Knight 1809).

*Andriapetalum cayennense* Klotzsch ex Meisn., *Fl. Bras. [Martius]* 5 (1): 78 [1 Jan. 1855] (Meisner 1855), “*Andripetalum*”.

VERNACULAR NAMES. — Wp: kuya'i lá • Wn: meliju • Cr: bwa-grajroz • Br: faeira, louro-faia.

HERBARIUM DATA (FG). — 15 collections at CAY. Sel. exs.: J.B. Leblond 224 (original material P[P00750531]).

INVENTORY DATA (FG). — 6 trees in 5 plots; $F_{max} < 1\%$; $dbh_{inv} = 34.1$ cm.

Genus *Roupala* Aubl.

[1422] *Roupala montana* Aubl. (Fig. 47B)

Hist. Pl. Guiane 1: 83 [Jun.-Dec. 1775] (Aublet 1775).

*Embothrium pinnatum* Ruiz & Pav., *Flora Peruviana* 1: 63 (Ruiz & Pavón 1798). — *Roupala diversifolia* R.Br., *Trans. Linn. Soc. London* 10 (1): 193 [“1811” publ. 1810] (Brown 1810), “*Hopala*”, nom. illeg. *superfl.* (based on *Embothrium pinnatum*). — *Roupala pinnata* (Ruiz & Pav.) Diels ex J.F.Macbr., *Publ. Field Mus. Nat. Hist., Bot. Ser. 13* (2.2): 374 [15 Mar. 1937] (Macbride 1937), nom. illeg. *hom.*, non Lam. (Lamarck 1791) [synonym of *Euplassa pinnata*].

Roupala pyrifolia Knight, Cult. Prot.: 102 [Dec. 1809] (Knight 1809).

Roupala media R.Br., *Trans. Linn. Soc. London* 10 (1): 191 [“1811” publ. 1810] (Brown 1810), “*Hopala*”.

Roupala dentata R.Br., *Trans. Linn. Soc. London* 10 (1): 192 [“1811” publ. 1810] (Brown 1810).

Roupala complicata Kunth, *Nova genera et species plantarum* [H.B.K.] 2: 153 [8 Dec. 1817] (Kunth 1818), “*Hopala*”.

Roupala ovalis Pohl, *Pl. Bras. Icon. Descr.* 1 (4): 107 [May-Dec. 1828] (Pohl 1828), “*Hopala*”.

Roupala affinis Pohl, *Pl. Bras. Icon. Descr.* 1 (4): 112 [May-Dec. 1828] (Pohl 1828), “*Hopala*”.

Roupala macropoda Klotzsch & H.Karst., *Linnaea* 20: 473 [Oct. 1847] (Klotzsch & Karsten 1847), “*Hopala*”.

Roupala tomentosa Pohl var. *sellowii* Meisn., *Fl. Bras. [Martius]* 5 (1): 82 [1 Jan. 1855] (Meisner 1855), “*Hopala*”. — *Roupala ovalis*

Klotzsch ex Meisn., *Fl. Bras. [Martius]* 5 (1): 82 [1 Jan. 1855] (Meisner 1855), “*Hopala*”, nom. nud. pro syn.

Roupala gardneri Meisn., *Fl. Bras. [Martius]* 5 (1): 83 [1 Jan. 1855] (Meisner 1855), “*Hopala*”.

Roupala gardneri var. angustata Meisn., *Fl. Bras. [Martius]* 5 (1): 83 [1 Jan. 1855] (Meisner 1855), “*Hopala*”.

Roupala gardneri var. dentata Meisn., *Fl. Bras. [Martius]* 5 (1): 83 [1 Jan. 1855] (Meisner 1855), “*Hopala*”.

Roupala gardneri var. integrifolia Meisn., *Fl. Bras. [Martius]* 5 (1): 83 [1 Jan. 1855] (Meisner 1855), “*Hopala*”.

Roupala glabrata Klotzsch ex Meisn., *Fl. Bras. [Martius]* 5 (1): 84 [1 Jan. 1855] (Meisner 1855), “*Hopala*”.

Roupala brasiliensis Klotzsch var. macropoda Meisn., *Fl. Bras. [Martius]* 5 (1): 86 [1 Jan. 1855] (Meisner 1855), “*Hopala*”.

Roupala martii Meisn., *Fl. Bras. [Martius]* 5 (1): 87 [1 Jan. 1855] (Meisner 1855), “*Hopala*”.

Roupala martii var. pinnata Meisn., *Fl. Bras. [Martius]* 5 (1): 87 [1 Jan. 1855] (Meisner 1855), “*Hopala*”.

Roupala martii var. simplicifolia Meisn., *Fl. Bras. [Martius]* 5 (1): 87 [1 Jan. 1855] (Meisner 1855), “*Hopala*”. — *Roupala veraguensis* Klotzsch ex Meisn., *Fl. Bras. [Martius]* 5 (1): 87 [1 Jan. 1855] (Meisner 1855), “*Hopala*”, nom. nud. pro syn.

Roupala boissieriana Meisn., *Prodr. [A. P. de Candolle]* 14 (1): 430 [mid Oct. 1856] (Meisner 1856), “*Hopala*”.

Roupala montana var. complicata (Kunth) Griseb., *Fl. Brit. W.I. [Grisebach]*: 277 [late 1860] (Grisebach 1860).

Roupala montana var. heterophylla Griseb., *Fl. Brit. W.I. [Grisebach]*: 278 [late 1860] (Grisebach 1860).

Roupala borealis Hemsl., *Biol. Cent.-Amer., Bot.* 3 (14): 78 [Oct. 1882] (Hemsley 1882).

Roupala acuminata Glaz., *Bull. Soc. Bot. France* 59 (Mém. 3g): 602 [“1912” publ. 1913] (Glaziou 1913), nom. nud.

Roupala darienensis Pittier, *Contr. U.S. Natl. Herb.* 18 (6): 228 [22 Sep. 1917] (Pittier 1917).

Roupala panamensis Pittier, *Contr. U.S. Natl. Herb.* 18 (6): 229 [22 Sep. 1917] (Pittier 1917).

Roupala discolor Rusby, *Descr. S. Amer. Pl.* 12 [20 Dec. 1920] (Rusby 1920).

Roupala dissimilis Pittier, *Bol. Soc. Venez. Ci. Nat.* 5: 303 (Pittier 1939).

Roupala repanda Lundell, *Amer. Midl. Naturalist* 29 (2): 472 [5 Apr. 1943] (Lundell 1943).

Roupala sphenophylla Diels ex Sleumer, *Bot. Jahrb. Syst.* 76 (2): 166 [Jan. 1954] (Sleumer 1954).

Roupala montana var. dentata (R.Br.) Sleumer, *Bot. Jahrb. Syst.* 76 (2): 173 [Jan. 1954] (Sleumer 1954).

Roupala mayana Lundell, *Wrightia* 5 (4): 73 (Lundell 1975).

NOTES. — *Roupala brasiliensis* var. macropoda Meisn. is not based on *R. macropoda* Klotzsch & H.Karst. Prance (2009: 134) incorrectly cited the protologue of *R. tomentosa* var. *sellowii* Meisn.: “*Prodr. [A. P. de Candolle]* 14 (1): 28. 1856”. It is obvious that the “4” was
omitted, as on p. 428 of Candolle's Prodromus this variety is cited
in synonymy under *R. tomentosa* var. *dentata* Meisn.

VERNACULAR NAMES. — Wp: tatu kāsī • Nt: liba apisi • Cr: bwa-graj-
rouj • Fr: bois grage rouge • Br: carne-de-vaca, carvalho-vermelha,
faeira, louro-faia.

HERBARIUM DATA (FG). — 21 collections at CAY. Sel. exs.: *J.B.*
*Aublet s.n.* (lecto-, P-JJR[P00777521] designated by Lanjouw &
Uittien [1940: 156]).

INVENTORY DATA (FG). — 6 trees in 5 plots; Fmax < 1%; dbhinv = 23.2 cm.

[1423] *Roupala nitida* Rudge

Pl. Guian. [Rudge] 1 (3): 26 [June 1805] (Rudge 1805).

Roupala laurifolia Knight, Cult. Prot. 102 [Dec. 1809] (Knight
1809), nom. illeg. superfl. (based on *Roupala nitida*).

VERNACULAR NAMES. — Wp: tatu kāsī • Wn: awawaju.

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *J. Martin* 52 (original material BM[BM000839785]); *P. Grenand* 626, height 15 m.

Family PUTRANJIVACEAE Endl.
Genus *Drypetes* Vahl

[1424] *Drypetes fanshawei* Sandwith
(Fig. 47C)

Kew Bull. 7 (2): 258 [25 July 1952] (Sandwith 1952).

VERNACULAR NAMES. — Pa: wakukwa-ára • Nt: weti kookoo.

HERBARIUM DATA (FG). — 25 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 2831.

INVENTORY DATA (FG). — 295 trees in 89 plots; Fmax = 5%; dbhinv = 41.7 cm.

[1425] *Drypetes variabilis* Uittien
(Fig. 47D)

Recueil Trav. Bot. Néerl. 22: 348 [“1925” publ. Jan. 1926] (Uit-
tien 1926).

VERNACULAR NAMES. — Ka: pana akala, sekeleu • Nt: weti kookoo.

HERBARIUM DATA (FG). — 128 collections at CAY. Sel. exs.: *J.-F. Molino* 1769.

INVENTORY DATA (FG). — 369 trees in 126 plots; Fmax = 5.4%; dbhinv = 73 cm.

Family RHABDODENDRACEAE Prance
Genus *Rhabdodendron* Gilg & Pilg.

[1426] *Rhabdodendron amazonicum*
(Spruce ex Benth.) Huber
(Fig. 48A)

*Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (2): 427 (Huber 1909). — *Lecostemon amazonicum* Spruce ex Benth., *Hooker's J. Bot. Kew Gard. Misc.* 5: 295 (Bentham 1853).

Lecostemon crassipes Spruce ex Benth., *Hooker's J. Bot. Kew Gard.* *Misc.* 5: 295 (Bentham 1853). — *Rhabdodendron crassipes* (Spruce ex Benth.) Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (2): 428 (Huber 1909).

Lecostemon crassipes var. cayennense Benth., *Hooker's J. Bot. Kew Gard.* *Misc.* 5: 296 (Bentham 1853).

Rhabdodendron duckei Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (2): 428 (Huber 1909).

Rhabdodendron longifolium Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (2): 429 (Huber 1909).

Rhabdodendron paniculatum Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (2): 429 (Huber 1909).

Rhabdodendron arirambae Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 5 (2): 430 (Huber 1909).

Lecostemon sylvestre Gleason, *Bull. Torrey Bot. Club* 54 (2): 68 [Feb. 1927] (Gleason 1927). — *Rhabdodendron sylvestre* (Gleason) Maguire, *Bull. Torrey Bot. Club* 75 (4): 397 [Jul.-Aug. 1948] (Maguire 1948).

VERNACULAR NAMES. — Pa: batakivie • Wp: alala kā'i sili, wila to
poko • Br: batiputá.

HERBARIUM DATA (FG). — 54 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 3022.

INVENTORY DATA (FG). — 93 trees in 30 plots; Fmax = 4%; dbhinv = 25.5 cm.

Family RHAMNACEAE Juss.
Genus *Sarcomphalus* P.Browne

[1427] *Sarcomphalus cinnamomum*
(Triana & Planch.) Hauenschild
(Fig. 48B)

Taxon 65 (1): 55 (Hauenschild 2016). — *Ziziphus cinnamomum* Triana & Planch., *Ann. Sci. Nat., Bot.* sér. 5, 16: 380 (Triana & Planchon 1872).

Ziziphus itacaiunensis Fróes, *Bol. Técn. Inst. Agron.* N. 36: 151 (Fróes 1959).

HERBARIUM DATA (FG). — 14 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost* 2538.

INVENTORY DATA (FG). — 5 trees in 5 plots; Fmax < 1%; dbhinv = 70 cm.

Family RHIZOPHORACEAE Pers.
Genus *Cassipourea* Aubl.

[1428] *Cassipourea elliptica* (Sw.) Poir.

Encycl. [J. Lamarck et al.] Suppl. 2: 131 [23 Oct. 1811] (Poiret
1811). — *Legnotis elliptica* Sw., *Prodr. [Swartz]:* 84 [20 Jun.-29
July 1788] (Swartz 1788). — *Cassipourea guianensis* var. *elliptica*
(Sw.) M.Gómez, *Dicc. Bot. Nombres Vulg. Cub. Puerto-Rig.:* 84
(Gómez 1889). — *Cassipourea elliptica* var. *typica* Stehlé, *Bull. Soc.
Bot. France* 92 (9): 265 (Stehlé 1945), *nom. inval.* (Turland et al.
2018: Art. 24.3).
*Cassipourea alba* Griseb., *Abh. Königl. Ges. Wiss. Göttingen* 7: 223 (Grisebach 1857). — *Cassipourea elliptica* var. *alba* (Griseb.) Griseb., *Fl. Brit. W.I. [Grisebach]*: 274 [late 1860] (Grisebach 1860).

*Cassipourea elliptica* var. pauciserrata Griseb., *Fl. Brit. W.I. [Grisebach]*: 274 [late 1860] (Grisebach 1860).

*Endosteira oppositifolia* Turcz., *Bull. Soc. Imp. Naturalistes Moscou* 36 (1): 576 (Turczaninow 1863).

*Cassipourea elliptica* var. ovata Griseb., *Cat. Pl. Cub. [Grisebach]*: 108 [May-Aug. 1866] (Grisebach 1866).

*Cassipourea cubensis* Urb., *Symb. Antill. [Urban]* 7 (2): 293 [15 June 1912] (Urban 1912).

*Cassipourea brittoniana* Fawc. & Rendle, *J. Bot.* 64: 14 (Fawcett & Rendle 1926).

*Cassipourea podantha* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 4 (8): 241 [24 Oct. 1929] (Standley 1929).

*Cassipourea macrodonta* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 4 (8): 242 [24 Oct. 1929] (Standley 1929).

*Cassipourea guildingii* Briq., *Candollea* 4: 343 (Briquet 1931).

*Cassipourea belizensis* Lundell, *Bull. Torrey Bot. Club* 66 (9): 598 [Dec. 1939] (Lundell 1939).

*Cassipourea elliptica* var. *parvifolia* Stehlé, *Bull. Soc. Bot. France* 92 (9): 265 (Stehlé 1945).

**HERBARIUM DATA (FG).** — 10 collections at CAY. Sel. exs.: S.A. Mori & J.J. Pipoly 15425.

**SIZE.** — Venezuela, Táchira, 7 Jul. 1983. *H. van der Werff* 5021 (MO), 15 m.

[1429] *Cassipourea guianensis* Aubl.
(Fig. 48C)

*Hist. Pl. Guiane* 1: 529 [Jun.-Dec. 1775] (Aublet 1775). — *Legnotis cassipourea* Sw., *Prodr. [Swartz]* 84 [20 Jun.-29 July 1788] (Swartz 1788), *nom. illeg. superfl.* (based on *Cassipourea guianensis*).

*Cassipourea macrophylla* DC., *Prodr. [A. P. de Candolle]* 3: 34 [mid Mar. 1828] (Candolle 1828). — *Legnotis macrophylla* Mart. ex DC., *Prodr. [A. P. de Candolle]* 3: 34 [mid Mar. 1828] (Candolle 1828), *nom. nud. pro syn.*

*Cassipourea serrata* Benth., *J. Bot. [Hooker]* 2: 223 (Bentham 1840). — *Cassipourea guianensis* var. *serrata* (Benth.) Engl., *Fl. Bras. [Martius]* 12 (2): 430 [1 Sep. 1876] (Engler 1876).

*Cassipourea quadrilocularis* Spruce ex Benth., *Fl. Bras. [Martius]* 12 (2): 429 [1 Sep. 1876] (Bentham 1876), *nom. nud. pro syn.*

*Cassipourea latifolia* Alston, *Bull. Misc. Inform. Kew* 1925 (6): 266 [16 June 1925] (Alston 1925). — *Cassipourea elliptica* var. *latifolia* (Alston) Stehlé, *Bull. Soc. Bot. France* 92 (9): 265 (Stehlé 1945).

*Cassipourea broadwayi* Briq., *Candollea* 4: 342 (Briquet 1931).

*Cassipourea guianensis* var. *trichopoda* Briq., *Candollea* 4: 346 (Briquet 1931).

**VERNACULAR NAMES.** — Pa: wakāu • Ka: apali’ilan, apaliyulan • Wp: yawa poi li • Cr: pativyé-gri • Br: laranja-do-mato, mangued’agua-doce.

**HERBARIUM DATA (FG).** — 88 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000016972]).

**INVENTORY DATA (FG).** — 423 trees in 93 plots; $F_{max} = 4.5\%$; $dbh_{inv} = 23.1$ cm.

Genus *Rhizophora* L.

[1430] *Rhizophora mangle* L.

*Sp. Pl.* 1: 443 [1 May 1753] (Linnaeus 1753). — *Rhizophora americana* Nutt., *N. Amer. Sylv.* 1 (2): 112 [Jul.-Dec. 1842] (Nuttall 1842), *nom. illeg. superfl.* (based on *Rhizophora mangle*).

*Rhizophora mangle* var. *samoensis* Hochr., *Candollea* 2: 447 (Hochreutiner 1925).

*Rhizophora samoensis* (Hochr.) Salvoza, *Nat. Appl. Sci. Bull. Univ. Philipp.* 5: 220 (Salvoza 1936).

**NOTE.** — A species restricted to mangrove forests.

**VERNACULAR NAMES.** — Pa: kwatri • Ka: konopo, kunapo • Cr: pativyé-janm, pativyé-rouj • Fr: palétuvier rouge • Br: manguevermelho.

**HERBARIUM DATA (FG).** — 11 collections at CAY. Sel. exs.: *D. Loubry 2000*.

**SIZE.** — Up to 100 cm dbh (Prance 2018).

[1431] *Rhizophora racemosa* G.Mey.

*Prim. Fl. Esseq.* 185 [Nov. 1818] (Meyer 1818). — *Rhizophora mangle* var. *racemosa* (G.Mey.) Engl., *Fl. Bras. [Martius]* 12 (2): 427 [1 Sep. 1876] (Engler 1876).

**NOTE.** — A species restricted to mangrove forests.

**VERNACULAR NAMES.** — Pa: kwatri • Ka: konopo, kunapo • Cr: pativyé-janm, pativyé-rouj • Fr: palétuvier rouge • Br: manguevermelho.

**HERBARIUM DATA (FG).** — 20 collections at CAY. Sel. exs.: *M.-F. Prévost 4684*.

**SIZE.** — Up to 15 m tall (Steyermark 2004).

Family ROSACEAE Juss.
Genus *Prunus* L.

[1432] *Prunus accumulans* (Koehne) C.L.Li & Aymard

*BioLlania, ed. espec.* 6: 449 (Li & Aymard 1997). — *Prunus myrtifolia* var. *accumulans* Koehne, *Bot. Jahrb. Syst.* 52 (4-5): 320 [3 Aug. 1915] (Koehne 1915).

**HERBARIUM DATA (FG).** — A single collection, *G. Bourdy* 2941.

**INVENTORY DATA (FG).** — 13 trees in 11 plots; $F_{max} < 1\%$; $dbh_{inv} = 53$ cm.
[1433] *Prunus myrtifolia* (L.) Urb.
(Fig. 48D)

Bot. Jahrb. Syst. 52 (4-5): 320 [3 Aug. 1915] (Urban 1915). — *Celastrus myrtifolius* L., Sp. Pl. 1: 196 [1 May 1753] (Linnaeus 1753). — *Lauro-cerasus myrtifolia* (L.) Britton, N. Amer. Trees 510 [9 May 1908] (Britton 1908), "*Laurocerasus*".

*Prunus sphaerocarpa* Sw., Prodr. [Swartz] 80 [20 Jun.-29 July 1788] (Swartz 1788).

*Prunus tikalana* Lundell, *Wrightia* 4 (2): 88 [31 Dec. 1968] (Lundell 1968).

VERNACULAR NAMES. — Pa: kunau • Ka: okoipyo • Wp: kunawalu'i • Cr: bwa-noyo • Br: marmelo-bravo, virarú.

HERBARIUM DATA (FG). — 19 collections at CAY. Sel. exs.: *J.-J. de Granville et al. 10765*.

INVENTORY DATA (FG). — 7 trees in 6 plots; Fmax < 1%; dbhinv = 48.1 cm.

Family RUBIACEAE Juss.

Genus *Alibertia* A.Rich.

[1434] *Alibertia edulis* (Rich.) A.Rich.

Prodr. [*A. P. de Candolle*] 4: 443 [late Sep. 1830] (Richard 1830). — *Genipa edulis* Rich., *Actes Soc. Hist. Nat. Paris* 1: 107 [Oct. 1792] (Richard 1792). — *Gardenia edulis* (Rich.) Poir., *Encycl. [J. Lamarck et al.]* Suppl. 2: 708 [3 July 1812] (Poiret 1812). — *Alibertia utilis* A.Rich., *Mém. Rubiac.*, t. 21 [Dec. 1830] (Richard 1830), orthographical variant. — *Amaioua utilis* Baill., *Bull. Mens. Soc. Linn. Paris* 1 (28): 220 (Baillon 1879), *nom. illeg. superfl.* (based on *Alibertia edulis*). — *Amaioua edulis* (Rich.) Baill., *Hist. Pl. [Baillon]* 7: 387 [Feb. 1880] (Baillon 1880). — *Cordiera edulis* (Rich.) Kuntze, *Revis. Gen. Pl.* 1: 279 [5 Nov. 1891] (Kuntze 1891). — *Sabicea edulis* (Rich.) Seem., *Hooker's J. Bot. Kew Gard. Misc.* 3: 266 (Seemann 1851).

NOTES. — Incipiently domesticated by pre-Columbian Amerindians (Clement 1999; Levis *et al.* 2017). Persson & Delprete (2017) recognized 4 varieties in *Alibertia edulis* (Rich.) A.Rich. Only var. *edulis* is present in French Guiana.

VERNACULAR NAMES. — Pa: tukuyuy • Wp: yanipa lali • Cr: gouyavkaka-poule, gouyav-nwé • Br: apuruizinho, marmelada, puruhy, purui-pequeno.

HERBARIUM DATA (FG). — 29 collections at CAY. Sel. exs.: *J.B. Leblond s.n.* (lecto-, P[P00870033], designated by Delprete & Persson in Taylor *et al.* [2011: 136]).

SIZE. — Up to 10 m tall (Taylor *et al.* 2004).

[1435] *Alibertia latifolia* (Benth.) K.Schum.

Fl. Bras. [Martius] 6 (6): 386 [15 June 1889] (Schumann 1889). — *Cordiera latifolia* Benth., *J. Bot. [Hooker]* 3: 221 (Bentham 1841).

*Alibertia oligantha* K.Schum., *Fl. Bras. [Martius]* 6 (6): 385 [15 June 1889] (Schumann 1889). — *Cordiera oligantha* (K.Schum.) Kuntze, *Revis. Gen. Pl.* 1: 279 [5 Nov. 1891] (Kuntze 1891).

*Alibertia latifolia* var. *parvifolia* K.Schum., *Fl. Bras. [Martius]* 6 (6): 386 [15 June 1889] (Schumann 1889).

*Alibertia granulosa* Rusby, *Descr. S. Amer. Pl.* 133 [20 Dec. 1920] (Rusby 1920).

*Alibertia latifolia* var. *pargueniana* Steyerm., *Mem. New York Bot. Gard.* 12 (3): 225 (Steyermark 1965).

VERNACULAR NAMES. — Pa: tukuyuy.

HERBARIUM DATA (FG). — A single collection, *P. Grenand 1808*, 12 m.

## Genus *Alseis* Schott

[1436] *Alseis longifolia* Ducke

Arch. Jard. Bot. Rio de Janeiro 3: 256 (Ducke 1922).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *J.-J. de Granville B-4741*.

SIZE. — Up to 45 cm dbh (Boom & Delprete 2002).

## Genus *Amaioua* Aubl.

[1437] *Amaioua glomerulata* (Lam. ex Poir.) Delprete & C.H.Perss.

Adansonia, sér. 3, 34 (2): 356 [Dec. 2012] (Delprete & Persson 2012). — *Mussaenda glomerulata* Lam. ex Poir., *Encycl. [J. Lamarck et al.]* 4 (1): 393 [9 Feb. 1797] (Poiret 1797). — *Rondeletia glomerulata* (Lam. ex Poir.) Spreng., *Syst. Veg. [Sprengel]* 1: 707 ["1825" publ. late 1824] (Sprengel 1824).

*Amaioua fagifolia* Desf., *Mém. Mus. Hist. Nat.* 6: 14 (Desfontaines 1820).

*Amaioua corymbosa* Kunth, *Nova genera et species plantarum [H.B.K.]* 3: 419 [13 Mar. 1820] (Kunth 1820). — *Hexactina corymbosa* (Kunth) Willd. ex Schult.f., *Syst. Veg. [Roemer & Schultes]* 7 (1): 91 (Schultes 1829), *nom. nud. pro syn.*

VERNACULAR NAMES. — Pa: wakavu-kamwi • Ka: wayu, wayun • Wp: kuluwi, puluwi • Cr: bwa-négrès, bwa-pièr • Br: goiabeira-preta, purui-grande.

HERBARIUM DATA (FG). — 12 collections at CAY. Sel. exs.: *H. Puig 12055*.

INVENTORY DATA (FG). — 3 trees in 2 plots; Fmax < 1%; dbhinv = 14.6 cm.

[1438] *Amaioua guianensis* Aubl.
(Fig. 49A)

Hist. Pl. Guiane 2 (Suppl.): 13 [Jun.-Dec. 1775] (Aublet 1775). — *Hamelia glabra* Lam., *Encycl. [J. Lamarck et al.]* 3 (1): 65 [19 Oct. 1789] (Lamarck 1789), *nom. illeg. superfl.* (based on *Amaioua guianensis*). — *Hamelia sessiliflora* Willd., *Sp. Pl.*, ed. 4 1 (2): 981 [July 1798] (Willdenow 1798), *nom. illeg. superfl.* (based on *Amaioua guianensis*). — *Duhamelia glabra* Pers., *Syn. Pl. [Persoon]* 1: 203 [1 Apr.-15 June 1805] (Persoon 1805), *nom. illeg. superfl.* (based on *Amaioua guianensis*).
<figure><img src="image_70.png" /><figcaption>FIG. 47. — Proteaceae: *A, Panopsis sessilifolia* (Rich.) Sandwith (M.-F. Prévost & D. Sabatier 4000); *B, Roupala montana* Aubl. (D. Sabatier & M.-F. Prévost 4925). Putranjivaceae: *C, Drypetes fanshawei* Sandwith (D. Sabatier & J.-F. Molino 5001); *D, Drypetes variabilis* Uittien. © D. Sabatier/IRD.</figcaption></figure>

*Amaioua guianensis* var. *macrantha* Steyerm., *Mem. New York Bot. Gard.* 12 (3): 216 (Steyermark 1965).

NOTE. — Hyperdominant in Amazonia (ter Steege *et al.* 2020).

VERNACULAR NAMES. — Pa: maras-gamana • Ka: kapasi tuno • Wp: kuluwi, puluwi • Wn: akuli epa • Cr: bwa-négrès, bwa-pièr • Br: goabeira-preta, purui-grande.

HERBARIUM DATA (FG). — 113 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, BM[BM001008914] designated by Steyermark [1965: 216]).

INVENTORY DATA (FG). — 149 trees in 80 plots; Fmax = 1.6 %; dbhinv = 26.8 cm.

## Genus *Botryarrhena* Ducke

### [1439] *Botryarrhena pendula* Ducke

Notizbl. Bot. Gart. Berlin-Dahlem 11: 477 [11 July 1932] (Ducke 1932).

VERNACULAR NAMES. — Pa: iwan-etni, tukuyuy-kamwi.

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: *C. Feuillet et al.* 10067.

SIZE. — Tree to 20 m tall, dbh to 22.5(-45) cm.

## Genus *Capirona* Spruce

### [1440] *Capirona macrophylla* (Poepp.) Delprete (Fig. 49B)

*Phytotaxa* 443 (1): 102 [18 May 2020] (Delprete 2020). — *Condaminea macrophylla* Poepp., *Nova genera ac species plantarum* [*Poeppig & Endlicher*] 3: 30 [15-21 Aug. 1841] (Poeppig 1841).

*Capirona decorticans* Spruce, *J. Proc. Linn. Soc., Bot.* 3: 200 [Apr. 1859] (Spruce 1859).

*Monadelphanthus floridus* H.Karst., *Fl. Columb. [H. Karsten]* 1 (2): 67 [13 Feb. 1860] (Karsten 1860).
*Capirona duckei* Huber, *Bol. Mus. Goeldi Hist. Nat. Ethnogr.* 7: 185 (Huber 1913), *nom. nud*.

*Capirona leiophloea* Benoist, *Bull. Mus. Natl. Hist. Nat.* 27: 367 (Benoist 1921).

*Capirona huberiana* Ducke, *Arch. Jard. Bot. Rio de Janeiro* 3: 257 (Ducke 1922).

*Capirona surinamensis* Bremek., *Recueil Trav. Bot. Néerl.* 31: 261 (Bremekamp 1934).

*Loretoa peruviana* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 11 (5): 222 (Standley 1936).

*Capirona wurdackii* Steyerm., *Mem. New York Bot. Gard.* 10 (5): 190 (Steyermark 1964).

**VERNACULAR NAMES.** — Pa: wadidga, wadidga-priye, wadidga-wahuyo • Ka: akepiimyo, talipi, tapili • Wp: ayà pili • Wn: ëkepiimë • Nt: lisa pao, mutende • Cr: bwa-palika • Br: escorrega-macaco, mulatorana, pau-mulato, perna-de-moça.

**HERBARIUM DATA (FG).** — 20 collections at CAY. Sel. exs.: *O. Poncy 1048* (P[P00208816]).

**INVENTORY DATA (FG).** — 31 trees in 16 plots; $F_{max} < 1\%$; $dbh_{inv} = 35.5$ cm.

## Genus *Chimarrhis* Jacq.

### [1441] *Chimarrhis microcarpa* Standl.

*Bull. Torrey Bot. Club* 53 (7): 471 [Oct. 1926] (Standley 1926).

*Chimarrhis longistipulata* Bremek., *Recueil Trav. Bot. Néerl.* 31: 260 (Bremekamp 1934).

**VERNACULAR NAMES.** — Wp: walalu'i • Cr: bwa-chapèl • Fr: bois cathédrale.

**HERBARIUM DATA (FG).** — 34 collections at CAY. Sel. exs.: *S.A. Mori & J.J. Pipoly 15562*.

**INVENTORY DATA (FG).** — 84 trees in 16 plots; $F_{max} < 1\%$; $dbh_{inv} = 80$ cm.

### [1442] *Chimarrhis turbinata* DC.

*Prodr. [A. P. de Candolle* 4: 404 [late Sep. 1830] (Candolle 1830). — *Pseudochimarrhis turbinata* (DC.) Ducke, *Arch. Jard. Bot. Rio de Janeiro* 3: 255 (Ducke 1922). — *Chimarrhis duckei* Rizzini, *Revista Brasil. Biol.* 7: 277 (Rizzini 1947), *nom. illeg. superfl.* (based on *Chimarrhis turbinata*).

*Bathysa difformis* Benoist, *Bull. Mus. Natl. Hist. Nat.* 26: 185 (Benoist 1920). — *Pseudochimarrhis difformis* (Benoist) Benoist, *Arch. Bot.* 5 (Mém. 1): 265 [27 Mar. 1933] (Benoist 1933).

*Elaeagia brasiliensis* Standl., *Bull. Torrey Bot. Club* 60 (6): 395 [1 June 1933] (Standley 1933).

**VERNACULAR NAMES.** — Pa: kinuwup, warerey • Ka: abalaba, atakali, mapalaba, me:ku ubupo • Te: pukutsa'i • Wp: walalu'i • Wn: epalai, ëpalai • Nt: pali udu • Cr: bwa-chapèl, bwa-pagay, sitronèl-rouj • Fr: bois cathédrale • Br: carapanaúba-cinzero, pau-de-remo.

**HERBARIUM DATA (FG).** — 50 collections at CAY. Sel. exs.: *D. Sabatier & M.-F. Prévost 2116*.

**INVENTORY DATA (FG).** — 207 trees in 93 plots; $F_{max} = 2.3\%$; $dbh_{inv} = 114.6$ cm.

## Genus *Cordiera* A.Rich.

### [1443] *Cordiera myrciifolia* (K.Schum.) C.H.Perss. & Delprete

*Fl. Venez. Guayana* 8: 559 (Persson & Delprete 2004). — *Alibertia myrciifolia* K.Schum., *Fl. Bras. [Martius]* 6 (6): 393 [15 June 1889] (Schumann 1889). — *Cordiera myrciifolia* Spruce ex K.Schum., *Fl. Bras. [Martius]* 6 (6): 393 [15 June 1889] (Schumann 1889), *nom. nud. pro syn.*

*Alibertia uniflora* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 11 (5): 179 (Standley 1936).

*Alibertia myrciifolia* var. tepuiensis Steyerm., *Mem. New York Bot. Gard.* 12 (3): 226 (Steyermark 1965).

*Alibertia triloba* Steyerm., *Mem. New York Bot. Gard.* 12 (3): 227 (Steyermark 1965).

**VERNACULAR NAMES.** — Wn: malipaimë.

**HERBARIUM DATA (FG).** — 6 collections at CAY. Sel. exs.: *J.-F. Molino & D. Sabatier 1993*.

**INVENTORY DATA (FG).** — 4 trees in 4 plots; $F_{max} < 1\%$; $dbh_{inv} = 12.8$ cm.

### [1444] *Cordiera triflora* A.Rich.

*Prodr. [A. P. de Candolle* 4: 445 [late Sep. 1830] (Richard 1830). — *Alibertia triflora* (A.Rich.) K.Schum., *Fl. Bras. [Martius]* 6 (6): 392 [15 June 1889] (Schumann 1889).

*Octavia sessiliflora* DC., *Prodr. [A. P. de Candolle* 4: 464 [late Sep. 1830] (Candolle 1830). — *Nonatelia sessiliflora* (DC.) Kuntze, *Revis. Gen. Pl.* 1: 290 [5 Nov. 1891] (Kuntze 1891).

*Cordiera trifolia* Steud., *Nomencl. Bot. [Steuvel]*, ed. 2, 1: 419 (Steuvel 1840), orth. var.

*Alibertia tenuifolia* K.Krause, *Verh. Bot. Vereins Prov. Brandenburg* 50: 106 [30 Sep. 1908] (Krause 1908).

*Alibertia steinbachii* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 4 (8): 287 [24 Oct. 1929] (Standley 1929).

*Alibertia benensis* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 7 (3): 291 (Standley 1931).

**HERBARIUM DATA (FG).** — 4 collections at CAY. Sel. exs.: *L.C. Richard s.n.* (lecto-, P[P03821428] selected by Persson & Delprete [2010: 263]).

**SIZE.** — Up to 20 cm $dbh$ (Taylor *et al.* 2004).
## Genus *Cosmibuena* Ruiz & Pav.

[1445] *Cosmibuena grandiflora* (Ruiz & Pav.) Rusby

Bull. New York Bot. Gard. 4 (14): 368 [7 Dec. 1907] (Rusby 1907). — *Cinchona grandiflora* Ruiz & Pav., Fl. Peruv. [Ruiz & Pavón] 2: 54 (Ruiz & Pavón 1799). — *Cinchona obtusifolia* D.Dietr., Syn. Pl. [D. Dietrich] 1: 721 [July 1839] (Dietrich 1839), nom. illeg. superfl. (based on *Cinchona grandiflora*).

*Cosmibuena obtusifolia* Ruiz & Pav., Flora Peruviana 3: 3 (Ruiz & Pavón 1802). — *Buena obtusifolia* (Ruiz & Pav.) DC., Prodr. [A. P. de Candolle] 4: 356 [late Sep. 1830] (Candolle 1830).

*Cinchona quina* Ruiz & Pav., Flora Peruviana 3: 4 (Ruiz & Pavón 1802).

*Cinchona longiflora* Mutis ex Steud., Nomencl. Bot. [Steudel] 1: 196 (Steudel 1821), nom. nud. pro syn.

*Cinchona china* Ruiz ex DC., Prodr. [A. P. de Candolle] 4: 356 [late Sep. 1830] (Candolle 1830).

*Buena triflora* Benth., J. Bot. [Hooker] 3: 216 (Bentham 1841). — *Cosmibuena triflora* (Benth.) Klotzsch, Getreue Darstell. Gew. 14 (2): t. 15 (Klotzsch 1846).

*Buena latifolia* Benth., Pl. Hartw. [Bentham]: 191 [? 16 Sep. 1845] (Bentham 1845). — *Cosmibuena latifolia* (Benth.) Klotzsch ex Walp., Repert. Bot. Syst. [Walpers] 6 (1): 69 [3-5 Sep. 1846] (Walpers 1846). — *Cosmibuena obtusifolia* var. latifolia (Benth.) Hook.f., Bot. Mag. 102: t. 6239 (Hooker 1876). — *Cosmibuena grandiflora* var. latifolia (Benth.) Steyerm., Mem. New York Bot. Gard. 23: 296 (Steyermark 1972).

*Cosmibuena quinqueflora* Klotzsch, Getreue Darstell. Gew. 14 (2): t. 15 (Klotzsch 1846).

*Buena skinneri* Oerst., Vidensk. Meddel. Dansk Naturhist. Forn. Kjøbenhavn 1852: 48 (Oersted 1852). — *Cosmibuena skinneri* (Oerst.) Hemsl., Biol. Cent.-Amer., Bot. 2 (7): 12 [Apr. 1881] (Hemsley 1881).

*Cosmibuena gardenioides* Wernham, J. Bot. 51: 321 (Wernham 1913).

*Cosmibuena arborea* Standl., Contr. U.S. Natl. Herb. 17: 447 (Standley 1914).

*Cosmibuena ovalis* Standl., Contr. U.S. Natl. Herb. 18 (3): 137 [11 Feb. 1916] (Standley 1916).

NOTES. — Epiphytic or rarely terrestrial shrub or tree (rarely hemi-epiphytic strangling tree). *Cinchona china* is a misspell of *C. quina*.

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *G. Cremers & J.-J. de Granville 13176*.

SIZE. — Up to 14 m tall (Taylor *et al.* 2004).

## Genus *Coussarea* Aubl.

[1446] *Coussarea albescens* (DC.) Müll.Arg.

Fl. Bras. [Martius] 6 (5): 87 [1 July 1881] (Müller 1881), in obs. — *Faramea albescens* DC., Prodr. [A. P. de Candolle] 4: 498 [late Sep. 1830] (Candolle 1830).

*Coussarea racemosa* A.Rich., Prodr. [A. P. de Candolle] 4: 494 [late Sep. 1830] (Richard 1830).

NOTES. — The description and name of *C. racemosa* came from Richard (Dec. 1830: 177), the text of which Candolle had seen a preprint version. Thereafter Candolle (Sept. 1830: 498) cited Richard as the author of this name, so that in this case, the authority is “A.Rich.”, not “A.Rich. ex DC.” The names *F. albescens* and *C. racemosa* were published in the same volume and have thus equal priority. When transferring *F. albescens* to *Coussarea* Aubl., Müller (1881: 87) stated that it was synonymous with *C. racemosa*, and he chose *C. albescens* over *C. racemosa*, as indicated in the index of vol. 6 (5) of *Flora Brasiliensis* (Müller 1888) where the former appears in normal type and the latter in italics (i.e. as a synonym). Under Art. 11.5 of the ICN (Turland *et al.* 2018), we thus have to use *C. albescens*, relegating *C. racemosa* as its junior synonym.

VERNACULAR NAMES. — Pa: mavinbi, mavinbi-kamwi, mavinvi, mavinvi-kamwi.

HERBARIUM DATA (FG). — 52 collections at CAY. Sel. exs.: *L. C. Richard s.n.* (original material of *Coussarea racemosa*: P[P03914186]).

INVENTORY DATA (FG). — 75 trees in 24 plots; $F_{max}$ = 2.2 %; $dbh_{inv}$ = 18.5 cm.

[1447] *Coussarea grandis* Müll.Arg.

Fl. Bras. [Martius] 6 (5): 100 [1 July 1881] (Müller 1881).

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *G. Cremers 8397*.

INVENTORY DATA (FG). — 11 trees in 2 plots; $F_{max}$ = 1.5 %; $dbh_{inv}$ = 16.1 cm.

[1448] *Coussarea granvillei* Delprete & B.M.Boom

Brittonia 51 (4): 403 (Delprete & Boom 1999).

NOTE. — Known only from French Guiana.

HERBARIUM DATA (FG). — 14 collections at CAY. Sel. exs.: *J.-J. de Granville B-4744* (holo-, CAY[CAY007384]); *J.-J. de Granville et al. 8601*, 10 m × 10 cm.

[1449] *Coussarea hallei* Steyerm.

Mem. New York Bot. Gard. 23: 387 (Steyermark 1972).

NOTE. — Known only from French Guiana.

VERNACULAR NAMES. — Nt: ayee wato.

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *F. Hallé 1018* (holo-, P[P00836890]; iso-, CAY[CAY028108, CAY028109], P[P00836891]); *P. Petronelli 300*, dbh 10 cm.

[1450] *Coussarea lutea* (Aubl.) Delprete

Taxon 64: 609 (Delprete 2015). — *Nonatelia lutea* Aubl., Hist. Pl. Guiane 1: 190, t. 74 [only inflorescence and flowers] [Jun.-Dec. 1775] (Aublet 1775). — *Psychotria lutea* (Aubl.) Willd., Sp. Pl., ed. 4 1 (2): 971 [July 1798] (Willdenow 1798).

*Coussarea mapourioides* Bremek., Recueil Trav. Bot. Néerl. 31: 282 (Bremekamp 1934).
<figure><img src="image_71.png" /><figcaption>FIG. 48. — Rhabdodendraceae: **A**, *Rhabdodendron amazonicum* (Spruce ex Benth.) Huber (D. Sabatier & M.-F. Prévost 3022). Rhamnaceae: **B**, *Sarcomphalus cinnamomum* (Triana & Planch.) Hauenschild (D. Sabatier & M.-F. Prévost 2538). Rhizophoraceae **C**, *Cassipourea guianensis* Aubl. (J.-F. Molino & D. Sabatier 2844). Rosaceae: **D**, *Prunus myrtifolia* (L.) Urb. (D. Sabatier & J.-F. Molino 5527). A, B, D, © D. Sabatier/IRD; C, © J.-F. Molino/IRD.</figcaption></figure>

NOTE. — Several specimens of *C. lutea* collected in the Guianas have been identified as *Coussarea hirticalyx* Standl. (1930: 175), which is the species most similar to *C. lutea*. According to Taylor (1999), *C. hirticalyx* occurs from Amazonian Colombia to Peru. Taylor & Steyermark (2004: 564) cited a specimen from the Venezuelan Guayana (A. Fernández 7544) having a corolla tube up to 20 mm long; they expressed doubts regarding the delimitation of the two species. However, pending a more detailed comparison between the two taxa, *C. lutea* can be distinguished from *C. hirticalyx* by its persistent stipules (vs caducous in *C. hirticalyx*), corolla tube (8.5-)10.3-13(-20) mm long (vs 33-34 mm long), and fruit 9-10 × 8-9 mm (vs c. 18 × 14 mm).

HERBARIUM DATA (FG). — 7 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (Lectotype BM[BM000058142] designated by Delprete [2015: 609]).

SIZE. — Up to 10 m tall.

[1451] *Coussarea machadoana* Standl.
(Fig. 49C)

Bull. Torrey Bot. Club 60 (6): 395 [1 June 1933] (Standley 1933).

VERNACULAR NAMES. — Pa: wakāu, uwakaa.

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: S.A. Mori & J.J. Pipoly 15459.

INVENTORY DATA (FG). — 40 trees in 26 plots; Fmax < 1%; dbhinv = 20.8 cm.

[1452] *Coussarea paniculata* (Willd.) Standl.
(Fig. 49D)

J. Wash. Acad. Sci. 18: 282 (Standley 1928). — *Froelichia paniculata* Willd., Sp. Pl., ed. 4 1 (2): 607 [July 1798] (Willdenow 1798). — *Billardiera paniculata* Vahl, *Eclog. Amer.* 1: 13 (Vahl 1797), “*Billardieria*”, nom. inval. (combination not definitely accepted by the author). — *Froelichia paniculata* Vahl, *Eclog. Amer.* 3: 56 (Vahl 1807), isonym. — *Coussarea froelichia* A.Rich., *Prodr. [A. P. de Candolle]* 4: 494 [late Sep. 1830] (Richard 1830), nom. illeg. superfl. (based on *Billardiera paniculata*).

*Coussarea macrocarpa* A.Rich., *Prodr. [A. P. de Candolle]* 4: 494 [late Sep. 1830] (Richard 1830).

*Faramea martinii* DC., *Prodr. [A. P. de Candolle]* 4: 498 [late Sep. 1830] (Candolle 1830), “Martini”. — *Coussarea martinii* (DC.) Benth. & Hook.f. ex B.D.Jacks., *Index Kew.* 1 (1): 631 [6 Sep. 1893] (Jackson 1893), “Martini”.

*Coussarea darienensis* Steyerm., *Ceiba* 3 (1): 20 (Steyermark 1952).

*Coussarea morii* Dwyer, *Ann. Missouri Bot. Gard.* 67 (1): 130 (Dwyer 1980).
NOTES. — Vahl realised after the printing, but before publication of the first volume of his book, that the name *Billardiera* was already in use for another genus (*Billardiera* Sm. 1793). In his foreword, he therefore proposed the new name *Froelichia* Vahl, wherewith he invalidated all the species' names described under *Billardiera*. Furthermore, none of the new combinations in *Froelichia* were validly published in 1797, because no association was made between genus name and specific epithets. In fact, validation of Vahl's binomials occured when the index in the last volume was published (1807), but this was after Willdenow's publication of *F. paniculata*. *Coussarea paniculata* is based on the invalid *F. paniculata* Vahl, but this is an indirect reference to the valid *F. paniculata* Willd. The descriptions and names of *C. froelichia* and *C. macrocarpa* in Candolle (Sept. 1830: 494) came from Richard (Dec. 1830: 177), the text of which Candolle had seen a preprint version. Both names are thus to be ascribed to Richard alone (not "A.Rich. ex DC.").

VERNACULAR NAMES. — Pa: mavinbi, mavinvi • Ka: kofilan, maipuli elepali • Wp: tapi'i ka'alulu.

HERBARIUM DATA (FG). — 4 collections at CAY. Sel. exs.: *R. C. Ek* 1745.

INVENTORY DATA (FG). — 15 trees in 6 plots; Fmax = 1%; dbhinv = 20.9 cm.

## Genus *Coutarea* Aubl.

[1453] *Coutarea hexandra* (Jacq.) K.Schum. (Fig. 49E)

Fl. Bras. [Martius] 6 (6): 185 [15 June 1889] (Schumann 1889). — *Portlandia hexandra* Jacq., *Enumer. Syst. Pl.*: 16 [Aug.-Sep. 1760] (Jacquin 1760). — *Coutarea portlandia* Dum.Cours., *Bot. Cult.* 2: 561 [1-4 July 1802] (Dumont 1802), *nom. illeg. superfl.* (based on *Portlandia hexandra*).

*Coutarea speciosa* Aubl., *Hist. Pl. Guiane* 1: 314 [Jun.-Dec. 1775] (Aublet 1775). — *Portlandia speciosa* (Aubl.) Baill., *Hist. Pl. [Bail-lon]* 7: 381 [Feb. 1880] (Baillon 1880). — *Coutarea hexandra* var. *speciosa* (Aubl.) K.Schum., *Fl. Bras. [Martius]* 6 (6): 197 [15 June 1889] (Schumann 1889).

*Portlandia acuminata* Willd., *Syst. Veg. [Roemer & Schultes]* 5: 23 [Dec. 1819] (Willdenow 1819).

*Exostema souzanum* Mart., *Reise Bras. [Spix & Mart.]* 2: 789 (Mar-tius 1828). — *Cinchona souzana* (Mart.) Brign., *Mem. Mat. Fis. Soc. Ital. Sci. Modena, Pt. Mem. Fis.* 2 (1): 63 (Brignoli 1862).

*Coutarea campanilla* DC., *Prodr. [A. P. de Candolle]* 4: 350 [late Sep. 1830] (Candolle 1830). — *Coutarea hexandra* var. *campanilla* (DC.) Steyerm., *Mem. New York Bot. Gard.* 23: 298 (Steyermark 1972).

*Coutarea flavescens* Sessé & Moc. ex DC., *Prodr. [A. P. de Candolle]* 4: 350 [late Sep. 1830] (Candolle 1830).

*Coutarea pubescens* Pohl, *Pl. Bras. Icon. Descr.* 2 (4): 148 [Jan.-Sep. 1833] (Pohl 1833). — *Coutarea hexandra* var. *pubescens* (Pohl) K.Schum., *Fl. Bras. [Martius]* 6 (6): 198 [15 June 1889] (Schumann 1889). — *Coutarea hexandra* f. *pubescens* (Pohl) Steyerm., *Mem. New York Bot. Gard.* 23: 298 (Steyermark 1972).

*Bignonia triflora* Pav. ex DC., *Prodr. [A. P. de Candolle]* 9: 148 [1 Jan. 1845] (Candolle 1845), *nom. nud*.

*Gardenia hexagona* Lem., *Ill. Hort.* 16: 92 (Lemaire 1869).

*Coutarea scherffiana* André, *Ill. Hort.* 25: 120 (André 1878).

*Coutarea lindeniana* Baill., *Adansonia* [Baillon] 12: 300 (Baillon 1879).

*Coutarea hexandra* f. *tarapotensis* K.Schum., *Fl. Bras. [Martius]* 6 (6): 198 [15 June 1889] (Schumann 1889).

*Coutarea hexandra* var. *amazonica* K.Schum., *Fl. Bras. [Martius]* 6 (6): 197 [15 June 1889] (Schumann 1889).

*Coutarea hexandra* var. *fluminensis* K.Schum., *Fl. Bras. [Martius]* 6 (6): 197 [15 June 1889] (Schumann 1889).

*Coutarea hexandra* f. *albiflora* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 92 (Chodat & Hassler 1904).

*Coutarea hexandra* f. *grandiflora* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 92 (Chodat & Hassler 1904).

*Coutarea hexandra* f. *roseiflora* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 92 (Chodat & Hassler 1904).

*Coutarea hexandra* var. *calycina* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 92 (Chodat & Hassler 1904).

NOTE. — The name *Portlandia acuminata* Willd. is to be ascribed to Willdenow alone (Turland *et al.* 2018: Art. 46.3, Ex. 15).

VERNACULAR NAMES. — Pa: kwik-duwě • Ka: pakeli • Cr: quinaquina • Br: capança-vermelha, falsa-quina, quinarana.

HERBARIUM DATA (FG). — 48 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (Neotype of *Coutarea speciosa*, P-JU No. 9927a [P00680269], designated by Delprete [2015: 603], isotype P-JU No. 9927b [P00680270]).

INVENTORY DATA (FG). — 1 tree, dbh = 13 cm.

## Genus *Duroia* L.f.

[1454] *Duroia aquatica* (Aubl.) Bremek.

*Recueil Trav. Bot. Néerl.* 31: 270 (Bremekamp 1934). — *Coupoui aquatica* Aubl., *Hist. Pl. Guiane* 2 (Suppl.): 16 [Jun.-Dec. 1775] (Aublet 1775). — *Cupirana aubletiana* Miers, *Apocyn. S. Am.*: 16 [May-June 1878] (Miers 1878), *nom. illeg. superfl.* (based on *Coupoui aquatica*).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Pa: kinuwup • Ka: atagali, atakali, yalakalu pupo • Wp: yanipa u • Wn: mutupupot • Nt: gaan maamadosu, maamadosu • Cr: konfitir-makak, mari-pwèl • Br: purui-grande.

HERBARIUM DATA (FG). — 63 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (original material BM[BM000793078]).

INVENTORY DATA (FG). — 95 trees in 58 plots; Fmax < 1%; dbhinv = 19.9 cm.

[1455] *Duroia eriopila* L.f.

*Suppl. Pl.*: 209 [“1781” publ. Apr. 1782] (Linnaeus 1782). — *Genipa eriopila* (L.f.) Oken, *Allg. Naturgesch.* 3 (2): 886 (Oken 1841). — *Amaioua eriopila* (L.f.) Baill., *Hist. Pl. [Baillon]* 7: 387 [Feb. 1880] (Baillon 1880).

*Genipa merianae* Rich., *Actes Soc. Hist. Nat. Paris* 1: 107 [Oct. 1792] (Richard 1792). — *Gardenia merianae* (Rich.) Poir., *Encycl. [J.]*
Lamarck *et al.*) Suppl. 2: 708 [3 July 1812) (Poiret 1812), nom. illeg. (Duroia eriopila in synonymy).

*Amaioua surinamensis* Steud., *Flora* 26 (45): 763 [17 Dec. 1843] (SteuDEL 1843), “*Amajoua*”. — *Duroia surinamensis* (Steucl.) Benth. & Hook.f. ex B.D.Jacks., *Index Kew.* 1 (2): 804 [14 Dec. 1893] (Jackson 1893).

*Amaioua grandifolia* Miq., *Linnaea* 18: 291 [“1844” publ. Feb. (-May?) 1845] (Miquel 1845).

*Amaioua affinis* Miq., *Linnaea* 18: 613 [“1844” publ. prob. Aug. 1845] (Miquel 1845).

*Amaioua ursina* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 8 (3): 168 (Standley 1930).

*Duroia eriopila* f. glabra Steyerm., *Mem. New York Bot. Gard.* 12 (3): 203 (Steyermark 1965).

**VERNACULAR NAMES.** — Pa: kinuwup • Ka: atagali, atakali, me:ku ubupo, wayamu wolekotopo • Wp: yanipa u • Cr: konfitir-makak, mari-pwèl • Br: purui-grande.

**HERBARIUM DATA (FG).** — 22 collections at CAY. Sel. exs.: *P. Gre-nand* et al. 3442.

**INVENTORY DATA (FG).** — 36 trees in 28 plots; Fmax < 1%; dbhinv = 20 cm.

[1456] *Duroia longiflora* Ducke

Arch. Jard. Bot. Rio de Janeiro 4: 181 (Ducke 1925).

**VERNACULAR NAMES.** — Pa: inam-etni-seine, kuyau-kamwi • Nt: maamadosu.

**HERBARIUM DATA (FG).** — 36 collections at CAY. Sel. exs.: *S.A. Mori* et al. 21520.

**INVENTORY DATA (FG).** — 305 trees in 96 plots; Fmax = 4.2%; dbhinv = 52.5 cm.

[1457] *Duroia martiniana* (Miers) Bremek.

*Recueil Trav. Bot. Néerl.* 31: 270 (Bremekamp 1934). — *Cupirana martiniana* Miers, *Apocyn. S. Am.*: 17 [May-June 1878] (Miers 1878). — *Coupoui martiniana* (Miers) Kuntze, *Revis. Gen. Pl.* 2: 414 [5 Nov. 1891] (Kuntze 1891), “Cupuia”.

**NOTE.** — Known only from the Guiana Shield.

**VERNACULAR NAMES.** — Pa: kinuwup • Ka: atakali • Wp: yanipa u • Wn: mutupupot • Nt: maamadosu, pikin maamadosu.

**HERBARIUM DATA (FG).** — 49 collections at CAY. Sel. exs.: *C. Persson* et al. 2069.

**INVENTORY DATA (FG).** — 2 trees in 2 plots; Fmax < 1%; dbhinv = 11.6 cm.

[1458] *Duroia micrantha* (Ladbr.) Zarucchi & J.H.Kirkbr.

*Ann. Missouri Bot. Gard.* 77 (4): 851 (Zarucchi & Kirkbride 1990). — *Coupoui micrantha* Ladbr., *J. Bot.* 58: 176 (Ladbrook 1920).

*Duroia sprucei* Rusby, *Descr. S. Amer. Pl.* 133 [20 Dec. 1920] (Rusby 1920).

**HERBARIUM DATA (FG).** — 3 collections at CAY. Sel. exs.: *O. Poncy et al.* 1746.

**INVENTORY DATA (FG).** — 2 trees in 2 plots; Fmax < 1%; dbhinv = 29 cm.

Genus *Eumachia* DC.

[1459] *Eumachia boliviana* (Standl.) Delprete & J.H.Kirkbr.

*J. Bot. Res. Inst. Texas* 9: 76 (Delprete & Kirkbride 2015). — *Psychotria boliviana* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 7 (3): 302 (Standley 1931). — *Margaritopsis boliviana* (Standl.) C.M.Taylor, *Syst. Geogr. Pl.* 75: 170 (Taylor 2005).

*Psychotria lawrancei* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 17 (3): 281 (Standley 1937).

*Psychotria foetidiflora* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 22 (3): 202 (Standley 1940).

*Psychotria kukenanensis* Steyerm., *Fieldiana, Bot.* 28 (3): 601 (Steyermark 1953).

*Psychotria turboensis* Standl. ex Steyerm., *Acta Bot. Venez.* 4: 101. (Steyermark 1964)

*Psychotria puberulenta* Steyerm., *Mem. New York Bot. Gard.* 23: 488 (Steyermark 1972).

*Psychotria plowmanii* Steyerm., *Ann. Missouri Bot. Gard.* 71 (4): 1178 (Steyermark 1984).

*Psychotria ronaldii* Steyerm., *Ann. Missouri Bot. Gard.* 74 (2): 111 (Steyermark 1987).

**VERNACULAR NAMES.** — Pa: mavinvi-kamwi • Wp: tapi'i wapa'a.

**HERBARIUM DATA (FG).** — 2 collections at CAY. Sel. exs.: *C. Feuillet* et al. 10354.

**SIZE.** — Up to 12 m tall.

Genus *Faramea* Aubl.

[1460] *Faramea corymbosa* Aubl.

*Hist. Pl. Guiane* 1: 102 [Jun.-Dec. 1775] (Aublet 1775).

*Faramea pendula* Poepp., *Nova genera ac species plantarum* [*Poeppig & Endlicher*] 3: 29 [15-21 Aug. 1841] (Poeppig 1841).

*Faramea truncata* DC., *Prodr. [A. P. de Candolle]* 4: 496 [late Sep. 1830] (Candolle 1830).

**VERNACULAR NAMES.** — Wp: ka'a pelele.

**HERBARIUM DATA (FG).** — 24 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JJR[P00777662] designated by Lanjouw & Uittien [1940: 151]).

**INVENTORY DATA (FG).** — 1 tree, dbh = 10.2 cm.
[1461] *Faramea multiflora* A.Rich.

Prodr. [A. P. de Candolle] 4: 497 [late Sep. 1830] (Richard 1830). —
*Coussarea multiflora* (A.Rich.) Lemée, *Fl. Guyane Franç.* 3: 542
(Lemée 1954).

Coffea umbellata Vell., *Fl. Flumin.* 63 [“1825” publ. 7 Sep.-28 Nov. 1829] (Vellozo 1829), *nom. illeg. hom.*, *non* Ruiz & Pav. (Ruiz & Pavón 1799).

Psychotria cyanea Vell., *Fl. Flumin.* 68 [“1825” publ. 7 Sep.-28 Nov. 1829] (Vellozo 1829).

*Faramea salicifolia* C.Presl, *Symb. Bot. [C. Presl]* 2: 24 [July 1834] (Presl 1834). — *Faramea multiflora* var. salicifolia (C.Presl) Steyerm., *Mem. New York Bot. Gard.* 17 (1): 394 (Steyermark 1967).

*Faramea amazonica* Müll.Arg., *Flora* 58: 470 (Müller 1875). — *Faramea multiflora* var. *amazonica* (Müll.Arg.) Steyerm., *Mem. New York Bot. Gard.* 17 (1): 392 (Steyermark 1967).

*Faramea egensis* Müll.Arg., *Flora* 58: 470 (Müller 1875).

*Faramea parviflora* Müll.Arg., *Flora* 58: 470 (Müller 1875).

*Faramea salicifolia* f. *paniculata* Kuntze, *Revis. Gen. Pl.* 1: 282 [5 Nov. 1891] (Kuntze 1891).

*Faramea salicifolia* f. *subumbellata* Kuntze, *Revis. Gen. Pl.* 1: 282 [5 Nov. 1891] (Kuntze 1891).

*Faramea maynensis* Spruce ex Rusby, *Bull. New York Bot. Gard.* 4 (13): 300 [25 June 1906] (Rusby 1906). — *Faramea multiflora* var. *maynensis* (Spruce ex Rusby) Steyerm., *Mem. New York Bot. Gard.* 17 (1): 393 (Steyermark 1967).

Rudgea scandens K.Krause, *Bot. Jahrb. Syst.* 40 (3): 342 [24 Jan. 1908] (Krause 1908).

*Faramea benensis* Rusby, *Mem. New York Bot. Gard.* 7: 379 (Rusby 1927). — *Faramea multiflora* var. *benensis* (Rusby) Steyerm., *Mem. New York Bot. Gard.* 17 (1): 394 (Steyermark 1967).

*Faramea hondurae* Standl., *J. Wash. Acad. Sci.* 18: 169 (Standley 1928).

*Faramea stenura* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 4 (8): 331 [24 Oct. 1929] (Standley 1929).

*Faramea talamancarum* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 4 (8): 332 [24 Oct. 1929] (Standley 1929).

*Faramea cuencana* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 7 (2): 219 (Standley 1931).

*Faramea laxula* K.Krause, *Notizbl. Bot. Gart. Berlin-Dahlem* 15: 392 [30 Mar. 1941] (Krause 1941).

*Faramea cuatrecasasii* Standl. ex Steyerm., *Acta Biol. Venez.* 4 (1): 42 (Steyermark 1964).

*Faramea multiflora* var. *epedunculata* Steyerm., *Mem. New York Bot. Gard.* 17 (1): 392 (Steyermark 1967).

*Faramea talamancarum* f. *acutifolia* Dwyer, *Ann. Missouri Bot. Gard.* 54 (2): 141 [27 Oct. 1967] (Dwyer 1967).

NOTE. — The description and name of *F. multiflora* came from Richard (published in December 1830: 176), the text of which Candolle had seen a preprint version. The name is thus to be ascribed to Richard alone (not "A.Rich. ex DC.").

VERNACULAR NAMES. — Wp: kusili sī ka'a, tapi'i wapa'a sili, uwa kaya • Wn: iliwaimë • Br: chacrona.

HERBARIUM DATA (FG). — 69 collections at CAY. Sel. exs.: *L. C. Richard s.n.* (lecto-, P[P00836846] designated by Jardim [Taylor & Jardim 2020: 121]).

SIZE. — Up to 10 m tall (Taylor *et al.* 2004).

[1462] *Faramea occidentalis* (L.) A.Rich.

Mém. Rubiac. 96 [Dec. 1830] (Richard 1830). — *Ixora occidentalis* L., *Syst. Nat.*, ed. 10, 2: 893 [7 June 1759] (Linnaeus 1759). — *Coffea occidentalis* (L.) Jacq., * Enum. Syst. Pl. *: 16 [Aug.-Sep. 1760] (Jacquin 1760). — *Tetramerium odoratissimum* C.F.Gaertn., *Suppl. Carp.*: 90 [May 1806] (Gaertner 1806), *nom. illeg. superfl.* (based on several previously published plates and citing several published names in synonymy, notably *Coffea occidentalis*). — *Antoniana laurifolia* Tussac, *Fl. Antill.* 2: 62 (Tussac 1818), *nom. illeg. superfl.* (refers to the plates of Plumier and Brown on which is based *Ixora occidentalis*). — *Tetramerium occidentale* (L.) Nees & Mart., *Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur.* 12: 13 (Nees & Martius 1824). — *Faramea odoratissima* DC., *Prodr. [A. P. de Candolle]* 4: 496 [late Sep. 1830] (Candolle 1830), *nom. illeg. superfl.* (based on *Ixora occidentalis*). — *Coussarea odoratissima* M.Gómez, *Dicc. Bot. Nombres Vulg. Cub. Puerto-Riq.*: 28 (Gómez 1889), *nom. illeg. superfl.* (based on *Tetramerium odoratissimum*, i.e. on *Coffea occidentalis*).

*Faramea sertulifera* DC., *Prodr. [A. P. de Candolle]* 4: 496 [late Sep. 1830] (Candolle 1830). — *Coussarea sertulifera* (DC.) M.Gómez, *Anales Soc. Esp. Hist. Nat.* 23: 292 (Gómez 1894).

Darluca prostrata Raf., *New Fl. [Rafinesque]* 4: 56 (Rafinesque 1838), *nom. inval.* (simultaneously published with *Neleixa prostrata* Raf.)

*Neleixa prostrata* Raf., *New Fl. [Rafinesque]* 4: 56 (Rafinesque 1838), *nom. inval.* (simultaneously published with *Darluca prostrata* Raf.)

*Faramea tenuifolia* Rusby, *Mem. New York Bot. Gard.* 7: 380 (Rusby 1927).

*Faramea zetekii* Standl., *Contr. Arnold Arbor.* 5: 146 (Standley 1933).

*Faramea belizensis* Standl., *Publ. Carnegie Inst. Wash.* 461: 90 [26 Nov. 1935] (Standley 1935).

*Faramea occidentalis* subsp. *lonchocalyx* Steyerm., *Mem. New York Bot. Gard.* 17 (1): 385 (Steyermark 1967).

*Faramea occidentalis* var. *brachycalyx* Steyerm., *Mem. New York Bot. Gard.* 17 (1): 385 (Steyermark 1967).

*Faramea occidentalis* var. *meridionalis* Steyerm., *Mem. New York Bot. Gard.* 17 (1): 384 (Steyermark 1967).

*Faramea standleyana* L.O.Williams, *Phytologia* 26 (6): 490 [11 Oct. 1973] (Williams 1973).

NOTE. — Jacquin did not explicitly cite *Ixora occidentalis* L. as the basionym of his *Coffea occidentalis*, but the use of the same epithet and iconotype (“Plum. ic. 156, f. 2”) is an indirect reference to it.

VERNACULAR NAMES. — Pa: waaduk-purubumna.

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *M.-F. Prévost & P. Grenand* 3778, dbh 10 cm.
<figure><img src="image_72.png" /><figcaption>FIG. 49. — Rubiaceae: **A**, *Amaioua guianensis* Aubl. (D. Sabatier et al. 4792); **B**, *Capirona macrophylla* (Poepp.) Delprete (D. Sabatier 3554); **C**, *Coussarea machadoana* Standl. (D. Sabatier & J.-F. Molino 4839); **D**, *Coussarea paniculata* (Willd.) Standl. (D. Sabatier & J.-F. Molino 5307); **E**, *Coutarea hexandra* (Jacq.) K.Schum. (J.-F. Molino et al. 3382). A, © M.-F. Prévost/IRD; B-D, © D. Sabatier/IRD; E, © J.-F. Molino/IRD.</figcaption></figure>

[1463] *Faramea pedunculata* (Bremek.) Delprete

*Blumea* 51 (2): 361 (Delprete 2006). — *Faramea sessiliflora* Aubl. var. *pedunculata* Bremek., *Brittonia* 8 (4): 241 [Jan. 1957] (Bremekamp 1957).

NOTE. — Known only from the Guiana Shield.

VERNACULAR NAMES. — Wp: yanipa lali sili.

HERBARIUM DATA (FG). — 8 collections at CAY. Sel. exs.: R.S. Cowan 38808 (original material US [00129992]).

INVENTORY DATA (FG). — 2 trees in 2 plots; Fmax < 1%; dbhinv = 11.5 cm.

[1464] *Faramea polytriadophora* Bremek.

*Brittonia* 8 (4): 241 [Jan. 1957] (Bremekamp 1957).

HERBARIUM DATA (FG). — A single collection, D. Sabatier & E. Fonty 5590.

INVENTORY DATA (FG). — 3 trees in 2 plots; Fmax < 1%; dbhinv = 30.7 cm.

[1465] *Faramea sessilifolia* (Kunth) DC.

Prodr. [A. P. de Candolle] 4: 497 [late Sep. 1830] (Candolle 1830). — *Tetramerium sessilifolium* Kunth, *Nova genera et species plantarum* [H.B.K.] 3: 374 [21 Nov. 1819] (Kunth 1819). — *Ixora sessilifolia* (Kunth) Spreng., *Syst. Veg. [Sprengel]* 1: 409 ["1825" publ. late 1824] (Sprengel 1824).

*Faramea longifolia* Benth., *J. Bot. [Hooker]* 3: 234 (Bentham 1841).

*Faramea coarinensis* Müll.Arg., *Fl. Bras. [Martius]* 6 (5): 133 [1 July 1881] (Müller 1881).

*Faramea longifolia* var. *petiolaris* Müll.Arg., *Fl. Bras. [Martius]* 6 (5): 133 [1 July 1881] (Müller 1881).

*Faramea planitiarum* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 22 (3): 187 (Standley 1940).

*Faramea costata* Steyerm., *Mem. New York Bot. Gard.* 17 (1): 389 (Steyermark 1967).

HERBARIUM DATA (FG). — A single collection, *J.-J. de Granville 638*.

SIZE. — Suriname. *S. Koemar & K. McDonnell 46* (MO), 15 m.
[1466] *Faramea* sp. A

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *J.-F. Molino* 2144.

INVENTORY DATA (FG). — 2 trees in 1 plot; Fmax < 1%; dbhinv = 10.2 cm.

## Genus *Ferdinandusa* Pohl

[1467] *Ferdinandusa goudotiana* K.Schum.

Fl. Bras. [Martius] 6 (6): 211 [15 June 1889] (Schumann 1889).

*Ferdinandusa goudotiana* var. *eciliata* Steyerm., *Mem. New York Bot. Gard.* 23: 281 (Steyermark 1972).

*Ferdinandusa goudotiana* var. *psilocarpa* Steyerm., *Mem. New York Bot. Gard.* 23: 281 (Steyermark 1972).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *D. Loubry* 1892.

SIZE. — Up to 18 m tall (Taylor *et al.* 2007).

[1468] *Ferdinandusa paraensis* Ducke

Arch. Jard. Bot. Rio de Janeiro 3: 259 (Ducke 1922).

*Ferdinandusa paraensis* var. *palustris* Ducke, *Arch. Jard. Bot. Rio de Janeiro* 3: 259 (Ducke 1922).

VERNACULAR NAMES. — Pa: á-wakaha • Wp: wila to tawa • Br: acauá, bacabinha-quina.

HERBARIUM DATA (FG). — 35 collections at CAY. Sel. exs.: *S.A. Mori* et al. 21018.

INVENTORY DATA (FG). — 75 trees in 37 plots; Fmax = 1.6%; dbhinv = 40.9 cm.

[1469] *Ferdinandusa* sp. A
(Fig. 50A)

HERBARIUM DATA (FG). — 3 collections at CAY. Sel. exs.: *D. Sabatier* & *M.-F. Prévost* 4928.

INVENTORY DATA (FG). — 12 trees in 3 plots; Fmax = 1.4%; dbhinv = 16.2 cm.

## Genus *Genipa* L.

[1470] *Genipa americana* L. var. *americana*

Syst. Nat., ed. 10, 2: 931 [7 June 1759] (Linnaeus 1759), "americ". — *Gardenia genipa* Sw., *Prodr. [Swartz]*: 51 [20 Jun.-29 July 1788] (Swartz 1788), *nom. illeg. superfl.* (based on *Genipa americana*). — *Genipa grandifolia* Pers., *Syn. Pl. [Persoon]* 1: 198 [1 Apr.-15 June 1805] (Persoon 1805), *nom. illeg. superfl.* (based on *Genipa americana*).

*Genipa oblongifolia* Ruiz & Pav., *Flora Peruviana* 2: 67 (Ruiz & Pavón 1799). — *Gardenia oblongifolia* (Ruiz & Pav.) Poir., *Encycl. [J. Lamarck et al.]* Suppl. 2: 708 [3 July 1812] (Poiret 1812).

*Genipa humilis* Vell., *Fl. Flumin.* 94 [“1825” publ. 7 Sep.-28 Nov. 1829] (Vellozo 1829).

*Genipa barbata* C.Presl, *Symb. Bot. [C. Presl]* 2: 13 [July 1834] (Presl 1834).

*Genipa excelsa* K.Krause, *Bot. Jahrb. Syst.* 40 (3): 327 [24 Jan. 1908] (Krause 1908).

*Genipa codonocalyx* Standl., *Contr. U.S. Natl. Herb.* 17: 446 (Stand-ley 1914).

NOTE. — Domesticated by pre-Columbian Amerindians (Clement 1999; Levis *et al.* 2017).

VERNACULAR NAMES. — Pa: arasgu • Ka: tabulupo, tapulupo • Te: dzádupa uhu • Wp: yanipa, yanipa e'e • Wn: kupë • Nt: uman tapupa • Cr: jénipa, jinipa • Fr: génipa • Br: genipapo, jenipapo.

HERBARIUM DATA (FG). — 6 collections at CAY. Sel. exs.: *D. Sabatier* & *M.-F. Prévost* 4201.

SIZE. — Up to 60 cm dbh (Taylor *et al.* 2004).

[1471] *Genipa americana* var. *caruto* (Kunth) K.Schum.

Fl. Bras. [Martius] 6 (6): 352 [15 June 1889] (Schumann 1889). — *Genipa caruto* Kunth, *Nova genera et species plantarum* [H.B.K.] 3: 407 [13 Mar. 1820] (Kunth 1820).

*Genipa pubescens* DC., *Prodr. [A. P. de Candolle]* 4: 378 [late Sep. 1830] (Candolle 1830).

*Genipa americana* f. *grandifolia* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 171 (Chodat & Hassler 1904).

*Genipa americana* f. *parvifolia* Chodat & Hassl., *Bull. Herb. Boissier*, sér. 2, 4: 171 (Chodat & Hassler 1904).

*Genipa americana* f. *jorgensenii* Steyerm., *Mem. New York Bot. Gard.* 23: 353 (Steyermark 1972).

VERNACULAR NAMES. — Br: genipapo.

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *V. Hequet* 159.

SIZE. — Guyana. *M.J. Jansen-Jacobs et al.* 78 (MO), 8 m × 30 cm.

[1472] *Genipa spruceana* Steyerm.

*Mem. New York Bot. Gard.* 23: 353 (Steyermark 1972).

VERNACULAR NAMES. — Pa: amuwan, arasgu • Te: dzádupa'i • Wp: yanipa i • Wn: pisusuk • Nt: uman tapupa • Cr: jinipa-dilo, jiniparivié, ti-jinipa • Fr: petit génipa.

HERBARIUM DATA (FG). — 60 collections at CAY. Sel. exs.: *M.-F. Prévost* & *P. Grenand* 2032.

SIZE. — Brazil, Pará. *C.A. Cid Ferreira 8010* (MO), 10 m × 18 cm.
## Genus Guettarda L.

[1473] Guettarda argentea Lam.

Encycl. [J. Lamarck et al.] 3 (1): 54 [19 Oct. 1789] (Lamarck 1789). — Matthiola argentea (Lam.) Kuntze, Revis. Gen. Pl. 1: 288 [5 Nov. 1891] (Kuntze 1891).

Guettarda argentea var. glabrata Urb., Symb. Antill. [Urban] 6 (1): 47 [15 July 1909] (Urban 1909).

VERNACULAR NAMES. — Pa: seivan.

HERBARIUM DATA (FG). — 25 collections at CAY. Sel. exs.: D. Sabatier & J.-F. Molino 5027, dbh = 46.5 cm.

## Genus Iertia Schreb.

[1474] Iertia coccinea (Aubl.) J.F.Gmel.

Syst. Nat., ed. 13[bis], 2 (1): 567 [late Sep.-Nov. 1791] (Gmelin 1791). — Guettarda coccinea Aubl., Hist. Pl. Guiane 1: 317 [Jun.-Dec. 1775] (Aublet 1775). — Duhamelia coccinea (Aubl.) Dum. Cours., Bot. Cult., ed. 2, 4: 328 (Dumont 1811). — Phosanthus coccineus (Aubl.) Raf., Ann. Gen. Sci. Phys. 6: 82 (Rafinesque 1820), “coccinea”.

Iertia flava Miq., Linnaea 18: 613 [“1844” publ. prob. Aug. 1845] (Miquel 1845).

Iertia coccinea var. pentamera Bremek., Recueil Trav. Bot. Néerl. 31: 263 (Bremekamp 1934).

VERNACULAR NAMES. — Pa: audik-amana, audika-retni, tahuma • Wp: wainimi susu • Wn: palalipanuimë • Nt: kotika uman gonbe • Cr: lanmou-rouj • Br: coralleira.

HERBARIUM DATA (FG). — 96 collections at CAY. Sel. exs.: J.B. Aublet s.n. (lecto-, P[P02273439] designated by Boom [1984: 439]).

INVENTORY DATA (FG). — 14 trees in 12 plots; Fmax < 1%; dbhinv = 27.6 cm.

[1475] Iertia hypoleuca Benth.

J. Bot. [Hooker] 3: 220 (Bentham 1841). — Iertia coccinea var. hypoleuca (Benth.) K.Schum., Fl. Bras. [Martius] 6 (6): 286 [15 June 1889] (Schumann 1889).

Iertia breviflora Mart. ex Steud., Nomencl. Bot. [Steudel], ed. 2, 1: 825 (Steudel 1840), nom. nud.

Cassupa scarlatina K.Schum. & K.Krause, Verh. Bot. Vereins Prov. Brandenburg 50: 98 [30 Sep. 1908] (Schumann & Krause 1908). — Iertia scarlatina (K.Schum. & K.Krause) Standl., Publ. Field Mus. Nat. Hist., Bot. Ser. 8 (5): 346 (Standley 1931).

Iertia hoehnei K.Krause, Arch. Bot. São Paulo 1: 115 (Krause 1925).

VERNACULAR NAMES. — Ka: melamelulan • Wp: wainimi susu.

HERBARIUM DATA (FG). — 19 collections at CAY. Sel. exs.: L.E. Skog & C. Feuillet 7440.

SIZE. — Up to 15 m tall, to 35 cm dbh.

[1476] Iertia parviflora Vahl

Eclog. Amer. 2: 28 (Vahl 1798).

Brignolia acuminata DC., Prodr. [A. P. de Candolle] 4: 444 [late Sep. 1830] (Candolle 1830).

Brignolia pubigera Benth., J. Bot. [Hooker] 3: 219 (Bentham 1841).

Bruinsmania isertioides Miq., Linnaea 17: 73 (Miquel 1843).

Iertia glabra Ducke, Arch. Jard. Bot. Rio de Janeiro 4: 179 (Ducke 1925).

Iertia parviflora var. hirta Steyerm., Mem. New York Bot. Gard. 17 (1): 295 (Steyermark 1967).

Yutajea liesneri Steyerm., Ann. Missouri Bot. Gard. 74 (3): 676 (Steyermark 1987).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: L. Andersson 1993.

SIZE. — Up to 12 m tall, to 15 cm dbh.

[1477] Iertia spiciformis DC.

Prodr. [A. P. de Candolle] 4: 437 [late Sep. 1830] (Candolle 1830), “spicaeformis”.

Iertia commutata Miq., Stirp. Surinam. Select.: 172 [“1850” publ. Mar. 1851] (Miquel 1851).

Iertia bullata K.Schum., Fl. Bras. [Martius] 6 (6): 286 [15 June 1889] (Schumann 1889).

Iertia viscosa Ducke, Arch. Jard. Bot. Rio de Janeiro 3: 260 (Ducke 1922).

Iertia pterantha Bremek., Brittonia 8 (4): 242 [Jan. 1957] (Bremekamp 1957).

VERNACULAR NAMES. — Ka: melamelulan • Wp: wainimi susu.

HERBARIUM DATA (FG). — 93 collections at CAY. Sel. exs.: S.A. Mori et al. 26555.

INVENTORY DATA (FG). — 2 trees in 2 plots; Fmax < 1%; dbhinv = 11.1 cm.

## Genus Ixora L.

[1478] Ixora ferrea (Jacq.) Benth.

Linnaea 23: 447 (Bentham 1850). — Sideroxyloides ferreum Jacq., Select. Stirp. Amer. Hist.: 19 [5 Jan. 1763] (Jacquin 1763). — Siderodendrum ferreum (Jacq.) J.F.Gmel., Syst. Nat., ed. 13[bis], 2 (1): 242 [late Sep.-Nov. 1791] (Gmelin 1791). — Siderodendrum ferreum (Jacq.) Lam., Tabl. Encycl. 1[1 (2)]: 282 [13 Feb. 1792] (Lamarck 1792), isonym. — Siderodendrum triflorum Vahl, Eclog. Amer. 1: 10 (Vahl 1797), nom. illeg. superfl. (based on Sideroxyloides ferreum).

VERNACULAR NAMES. — Wp: walaku ki'īy.

HERBARIUM DATA (FG). — 48 collections at CAY. Sel. exs.: J.-J. de Granville et al. 6268.

SIZE. — Up to 12 m tall, to 15 cm dbh.
[1479] *Ixora panurensis* Müll.Arg.

*Flora* 58: 454 (Müller 1875).

*Ixora ulei* K.Krause, *Notizbl. Königl. Bot. Gart.* Berlin 6: 205 [15 May 1914] (Krause 1914).

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *D. Larpin* 882.

INVENTORY DATA (FG). — 5 trees in 2 plots; $F_{max} < 1\%$; $dbh_{inv} = 17.7$ cm.

[1480] *Ixora schomburgkiana* Benth.

*Linnaea* 23: 448 (Bentham 1850).

*Siderodendrum macrophyllum* Benth., *J. Bot. [Hooker]* 3: 220 (Bentham 1841), “*Siderodendron*”.

*Ixora gleasonii* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser. 4* (8): 344 [24 Oct. 1929] (Standley 1929).

NOTE. — Known only from the Guiana Shield.

HERBARIUM DATA (FG). — A single collection, *D. Sabatier* 1433.

SIZE. — Up to 10 m tall, to 15 cm dbh.

Genus *Kutchubaea* Fisch. ex DC.

[1481] *Kutchubaea insignis* Fisch. ex DC.
(Fig. 50D)

Prodr. [*A. P. de Candolle*] 4: 373 [late Sep. 1830] (Candolle 1830).

*Gardenia integra* A.Rich., *Mém. Rubiac.* 241 [Dec. 1830] (Richard 1830), *nom. illeg. superfl.* (includes original material of *Kutchubaea insignis*).

*Kutchubaea laxiflora* Benth. & Hook.f., *Gen. Pl. [Bentham & Hooker f.]* 2 (1): 98 [7-9 Apr. 1873] (Bentham & Hooker 1873), “*Kutchubaea*”, *pro parte quoad specim. in flore, nom. inval.* (genus name and epithet not associated).

NOTES. — The original description of *Gardenia integra* (Richard 1830: 241) is followed by: “*Crescit in Guianâ* (Richard). *Kutchubaea insignis* Fischer ex De Cand. *annot. in herb. Mus. Paris.*” The first sentence refers to *L.C. Richard s.n.* (P00836504), on whose left label Achille Richard wrote “*Gardenia integra* nob.” The second sentence is an indirect reference to another specimen at P, *J. Martin s.n.* [P00836503], which bears a label in Candolle’s hand that reads “*Kutchubaea insignis* Fisch.” The original material of *G. integra* thus comprises two collections, making lectotypification of this name necessary. *L.C. Richard s.n.*, barcode P00836504, is here designated as lectotype. Also, the publication of *K. insignis* Fisch. ex DC. predates that of *G. integra*; hence the latter name is superfluous and illegitimate.

VERNACULAR NAMES. — Pa: tukuyuy-kamwi.

HERBARIUM DATA (FG). — 9 collections at CAY. Sel. exs.: *J. Martin s.n.* (original material P[P00836503]).

INVENTORY DATA (FG). — 10 trees in 8 plots; $F_{max} < 1\%$; $dbh_{inv} = 44.2$ cm.

Genus *Ladenbergia* Klotzsch

[1482] *Ladenbergia lambertiana*
(A.Braun ex Mart.) Klotzsch

*Getreue Darstell. Gew.* 14 (2): t. 15 (Klotzsch 1846). — *Cinchona lambertiana* A.Braun ex Mart., *Reise Bras. [Spix & Mart.]* 3: 1286 (Martius 1831). — *Remijia lambertiana* (A.Braun ex Mart.) Wedd., *Ann. Sci. Nat., Bot. sér.* 3, 10: 13 [Dec. 1848] (Weddell 1848). — *Cascarilla lambertiana* (A.Braun ex Mart.) Wedd., *Hist. Nat. Quinquinas*: 85 (Weddell 1849). — *Buena lambertiana* (A.Braun ex Mart.) Wedd., *J. Linn. Soc., Bot.* 11: 187 [“1871” publ. Dec. 1869] (Weddell 1869).

*Ladenbergia schomburgkii* Klotzsch, *Getreue Darstell. Gew.* 14 (2): t. 15 (Klotzsch 1846). — *Cascarilla schomburgkii* (Klotzsch) Triana, *Revista Acad. Colomb. Ci. Exact.* 2: 410 (Triana 1938).

*Ladenbergia lucens* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 7 (4): 362 (Standley 1931).

*Ladenbergia puberula* Steyerm., *Mem. New York Bot. Gard.* 23: 270 (Steyermark 1972).

*Ladenbergia venamoensis* Steyerm., *Mem. New York Bot. Gard.* 23: 271 (Steyermark 1972).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *J.-J. de Granville & F. Crozier* 17010.

SIZE. — Up to 20 m tall (Steyermark 1974).

Genus *Morinda* L.

[1483] *Morinda citrifolia* L.

*Sp. Pl.* 1: 176 [1 May 1753] (Linnaeus 1753), nom. cons. — *Samama citrifolia* (L.) Kuntze, *Revis. Gen. Pl.* 1: 297 [5 Nov. 1891] (Kuntze 1891).

*Platanocephalus orientalis* Crantz, *Inst. Rei Herb.* 1: 349 [Jan.-July 1766] (Crantz 1766).

*Morinda tinctoria* Noronha, *Verh. Batav. Genootsch. Kunst.* 5 (Art. 4): 20 (Noronha 1790), *nom. nud.*

*Morinda tomentosa* B.Heyne ex Roth, *Syst. Veg. [Roemer & Schultes]* 5: 216 [Dec. 1819] (Roth 1819).

*Morinda angustifolia* Roth, *Nov. Pl. Sp.* 147 [Apr. 1821] (Roth 1821).

*Morinda chachuca* Buch.-Ham., *Trans. Linn. Soc. London* 13 (2): 536 (Buchanan-Hamilton 1822).

*Morinda nodosa* Buch.-Ham., *Trans. Linn. Soc. London* 13 (2): 537 (Buchanan-Hamilton 1822).

*Morinda mudia* Buch.-Ham., *Trans. Linn. Soc. London* 13 (2): 536 (Buchanan-Hamilton 1822).

*Morinda multiflora* Roxb., *Fl. Ind.* 2: 200 (Roxburgh 1824).

*Morinda stenophylla* Spreng., *Syst. Veg. [Sprengel]* 1: 749 [1825 publ. late 1824] (Sprengel 1824). — *Morinda coreia* Buch.-Ham. var. *stenophylla* (Spreng.) Chandrab., *Fl. Coimbatore* [*M. Chandrabose & N.C. Nair*]: xxviii, 142 (Chandrabose 1988).

*Morinda macrophylla* Desf., *Tabl. École Bot.*, ed. 3 [*Cat. Pl. Horti Paris.*], 404 (Desfontaines 1829).
*Morinda aspera* Wight & Arn., *Prodr. Fl. Ind. Orient.* 1: 420 [10 Oct. 1834] (Wight & Arnott 1834). — *Morinda pubescens* Sm. var. aspera (Wight & Arn.) M.Gangop., *Nelumbo* 59 (1): 24 (Gangopadhyay 2017).

*Morinda quadrangularis* G.Don, *Gen. Hist.* 3: 545 [8-15 Nov. 1834] (Don 1834).

*Morinda ligulata* Blanco, *Fl. Filip. [F.M. Blanco]*, ed. 2: 105 (Blanco 1845).

*Morinda littoralis* Blanco, *Fl. Filip. [F.M. Blanco]*, ed. 2: 109 (Blanco 1845), “litoralis”.

*Morinda teysmanniana* Miq., *Fl. Ned. Ind.* 2: 244 (Miquel 1857).

*Morinda zollingeriana* Miq., *Fl. Ned. Ind.* 2: 243 (Miquel 1857).

*Sarcocephalus leichhardtii* F.Muell., *Essay Pl. Burdekin:* 12 (Muel-ler 1860).

*Morinda citrifolia* L. f. potteri H.St.John, *Bull. Torrey Bot. Club* 111 (4): 482 (St.John 1984).

NOTE. — A species native to India, West Indies and Australia, cultivated and naturalised throughout the tropics. Cultivated and naturalised in Central America, the Greater Antilles, and in the Guianas; several naturalised populations have recently been observed in French Guiana (Delprete, pers. obs.)

VERNACULAR NAMES. — Fr: noni.

HERBARIUM DATA (FG). — 5 collections. Sel. exs.: *M.-F. Prévost 4686*.

SIZE. — Up to 12 m tall, to 15 cm dbh.

Genus *Pagamea* Aubl.

[1484] *Pagamea guianensis* Aubl.

*Hist. Pl. Guiane* 1: 113 [Jun.-Dec. 1775] (Aublet 1775), “*Guy-annensis*” on plate.

*Pagamea guianensis* var. angustifolia Progel, *Fl. Bras. [Martius]* 6 (1): 285 [1 Aug. 1868] (Progel 1868).

*Pagamea guianensis* var. parviflora Spruce ex Progel, *Fl. Bras. [Martius]* 6 (1): 285 [1 Aug. 1868] (Progel 1868).

HERBARIUM DATA (FG). — 41 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JJR[P00777682] designated by Lanjouw & Uittien [1940: 152]); *P.G. Delprete & F. Crozier 7150*.

INVENTORY DATA (FG). — 11 trees in 3 plots; Fmax = 1.4%; dbhinv = 13.8 cm.

[1485] *Pagamea thyrtsiflora* Spruce ex Benth.

*J. Proc. Linn. Soc., Bot.* 1: 110 [“1857” publ. 1856] (Bentham 1856).

HERBARIUM DATA (FG). — A single collection at CAY, *J.-J. de Granville 13833*.

SIZE. — Up to 10 m tall.

Genus *Palicourea* Aubl.

[1486] *Palicourea brachyloba* (Müll.Arg.) B.M.Boom

*Brittonia* 37 (2): 209 (Boom 1985). — *Psychotria brachyloba* Müll. Arg., *Fl. Bras. [Martius]* 6 (5): 243 [1 July 1881] (Müller 1881). — *Uragoga brachyloba* (Müll.Arg.) Kuntze, *Revis. Gen. Pl.* 2: 959 [5 Nov. 1891] (Kuntze 1891).

HERBARIUM DATA (FG). — 2 collections at CAY. Sel. exs.: *J.-J. de Granville B-5356*.

SIZE. — Up to 12 m tall (Boom & Delprete 2002).

[1487] *Palicourea guianensis* Aubl.

*Hist. Pl. Guiane* 1: 173 [Jun.-Dec. 1775] (Aublet 1775), “*Guy-annensis*” on plate. — *Stephanium guianense* (Aubl.) J.F.Gmel., *Syst. Nat.*, ed. 13[bis], 2 (1): 368 [late Sep.-Nov. 1791] (Gmelin 1791). — *Psychotria palicourea* Sw., *Fl. Ind. Occid.* 1: 433 [Nov. 1797] (Swartz 1797), *nom. illeg. superfl.* (based on *Palicourea guianensis*). — *Simira palicourea* Poir., *Encycl. [J. Lamarck et al.]* 7: 196 [6 July 1806] (Poiret 1806), *nom. illeg.* (based on the illegitimate *Psychotria palicourea*). — *Uragoga palicourea* Kuntze, *Revis. Gen. Pl.* 1: 300 [5 Nov. 1891] (Kuntze 1891), *nom. illeg. superfl.* (based on *Palicourea guianensis*).

*Palicourea barbinervia* DC., *Prodr. [A. P. de Candolle]* 4: 530 [late Sep. 1830] (Candolle 1830). — *Uragoga barbinervia* (DC.) Kuntze, *Revis. Gen. Pl.* 2: 959 [5 Nov. 1891] (Kuntze 1891). — *Palicourea guianensis* subsp. *barbinervia* (DC.) Steyerm., *Mem. New York Bot. Gard.* 23: 730 (Steyermark 1972).

*Psychotria lutea* Spreng. ex DC., *Prodr. [A. P. de Candolle]* 4: 530 [late Sep. 1830] (Candolle 1830), *nom. nud. pro syn.*

*Psychotria cataractarum* Müll.Arg., *Fl. Bras. [Martius]* 6 (5): 226 [1 July 1881] (Müller 1881). — *Nonatelia cataractarum* (Müll.Arg.) Kuntze, *Revis. Gen. Pl.* 1: 291 [5 Nov. 1891] (Kuntze 1891).

*Palicourea guianensis* var. tetramera Bremek., *Recueil Trav. Bot. Néerl.* 31: 283 (Bremekamp 1934).

*Palicourea guianensis* var. trimera Bremek., *Recueil Trav. Bot. Néerl.* 31: 283 (Bremekamp 1934).

*Palicourea guianensis* f. glabra Steyerm., *Mem. New York Bot. Gard.* 23: 730 (Steyermark 1972).

*Palicourea guianensis* subsp. *occidentalis* Steyerm., *Mem. New York Bot. Gard.* 23: 729 (Steyermark 1972).

*Palicourea guianensis* var. glabrescens Steyerm., *Mem. New York Bot. Gard.* 23: 730 (Steyermark 1972).

VERNACULAR NAMES. — Pa: audik-amana, audika-retni • Ka: maipuli elepali, maipyuli kiyelapole, matula, padula, patula • Wp: tewitulū u, tewitulū wate'e, tewitutū wate'e • Nt: ayee wato, ayuwato • Cr: bwa-fizi • Br: erva-de-rato.

HERBARIUM DATA (FG). — 63 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JJR[P00778113] designated by Lanjouw & Uittien [1940: 155]).

INVENTORY DATA (FG). — 29 trees in 17 plots; Fmax = 2.7%; dbhinv = 31.5 cm.
## Genus *Posoqueria* Aubl.

[1488] *Posoqueria gracilis* (Rudge) Roem. & Schult.

*Syst. Veg.* [Roemer & Schultes] 5: 227 [Dec. 1819] (Roemer & Schultes 1819). — *Solena gracilis* Rudge, *Pl. Guian.* [*Rudge*] 1 (3): 27 [June 1805] (Rudge 1805). — *Tocoyena gracilis* (Rudge) A.Rich., *Mém. Rubiac.*: 167 [Dec. 1830] (Richard 1830). — *Posoqueria latifolia* subsp. *gracilis* (Rudge) Steyerm., *Mem. New York Bot. Gard.* 17 (1): 327 (Steyermark 1967).

NOTE. — Even though volume 5 of the *Systema vegetabilium* was published shortly after Roemer's death, Schultes' foreword clearly confirms Roemer's prominent role. Therefore, contrary to Stafleu & Cowan's recommendation (1983: TL-2-9408), most names should be ascribed to Roemer and Schultes, not Schultes alone.

VERNACULAR NAMES. — Wp: aliwa.

HERBARIUM DATA (FG). — 14 collections at CAY. Sel. exs.: *E. Rudge s.n.* (original material BM[BM001008906]).

SIZE. — Guyana. *P. Mutchnick* 836 (MO), 20 m.

[1489] *Posoqueria latifolia* (Rudge) Roem. & Schult.

*Syst. Veg.* [Roemer & Schultes] 5: 227 [Dec. 1819] (Roemer & Schultes 1819). — *Solena latifolia* Rudge, *Pl. Guian.* [*Rudge*] 1 (3): 26 [June 1805] (Rudge 1805). — *Ucriana insignis* Willd. ex Spreng., *Syst. Veg.* [*Sprengel*] 1: 761 ["1825" publ. late 1824] (Sprengel 1824), *nom. illeg. superfl.* (based on *Tocoyena latifolia*).

*Tocoyena speciosa* Rich., *Actes Soc. Hist. Nat. Paris* 1: 107 [Oct. 1792] (Richard 1792).

*Tocoyena latifolia* Lam., *Tabl. Encycl.* 2[5 (2)]: 259 [31 Oct. 1819] (Lamarck 1819).

*Tocoyena insignis* Roem. & Schult., *Syst. Veg.* [Roemer & Schultes] 5: 229 [Dec. 1819] (Roemer & Schultes 1819). — *Posoqueria insignis* (Roem. & Schult.) Nees, *Flora* 4 (1): 301 [21 May 1821] (Nees 1821).

*Tocoyena longifolia* Kunth, *Nova genera et species plantarum [H.B.K.]* 3: 411 [13 Mar. 1820] (Kunth 1820). — *Ucriana humboldtii* Spreng., *Syst. Veg.* [*Sprengel*] 1: 761 ["1825" publ. late 1824] (Sprengel 1824), *nom. illeg. superfl.* (based on *Tocoyena longifolia*).

*Tocoyena mutisii* Kunth, *Nova genera et species plantarum [H.B.K.]* 3: 411 [13 Mar. 1820] (Kunth 1820). — *Ucriana mutisii* (Kunth) Spreng., *Syst. Veg.* [*Sprengel*] 1: 761 ["1825" publ. late 1824] (Sprengel 1824).

*Tocoyena macrophylla* Kunth, *Nova genera et species plantarum [H.B.K.]* 3: 412 [13 Mar. 1820] (Kunth 1820). — *Ucriana macrophylla* (Kunth) D.Dietr., *Syn. Pl. [D. Dietrich]* 1: 798 [July 1839] (Dietrich 1839). — *Posoqueria macrophylla* (Kunth) Hemsl., *Biol. Cent.-Amer., Bot.* 2 (7): 39 [Apr. 1881] (Hemsley 1881).

*Posoqueria revoluta* Schrad., *Gött. Gel. Anz.* 2: 714 [5 May 1821] (Schrader 1821). — *Solena revoluta* (Schrad.) D.Dietr., *Syn. Pl. [D. Dietrich]* 1: 799 [July 1839] (Dietrich 1839).

*Gardenia suaveolens* Vell., *Fl. Flumin.*: 102 ["1825" publ. 7 Sep.-28 Nov. 1829] (Vellozo 1829), "suave-olens".

*Posoqueria decora* DC., *Prodr. [A. P. de Candolle]* 4: 375 [late Sep. 1830] (Candolle 1830). — *Solena decora* (DC.) D.Dietr., *Syn. Pl. [D. Dietrich]* 1: 799 [July 1839] (Dietrich 1839).

*Posoqueria lucida* Mart., *Flora* 24 (2, Beibl.): 77 (Martius 1841).

*Posoqueria macropus* Mart., *Flora* 24 (2, Beibl.): 79 (Martius 1841).

*Stannia panamensis* Walp. & Duchass., *Linnaea* 23: 755 ["1850" publ. Jan. 1851] (Walpers & Duchassaing 1851). — *Posoqueria panamensis* (Walp. & Duchass.) Walp., *Ann. Bot. Syst.* [*Walpers*] 2 (5): 797 [21-24 Apr. 1852] (Walpers 1852).

*Stannia metensis* H.Karst., *Linnaea* 28: 441 (Karsten 1857). — *Posoqueria metensis* (H.Karst.) R.Knuth, *Repert. Spec. Nov. Regni Veg.* *Beih.* 43: 667 (Knuth 1928).

*Stannia grandiflora* H.Karst., *Fl. Columb. [H. Karsten]* 1 (1): 31 [6 Apr. 1859] (Karsten 1859). — *Posoqueria panamensis* subsp. *grandiflora* (H.Karst.) Steyerm., *Mem. New York Bot. Gard.* 17 (1): 325 (Steyermark 1967).

*Posoqueria multiflora* Lem., *Ill. Hort.* 16: t. 597 (Lemaire 1869), *nom. illeg. hom., non* (Willd.) Blume (1826).

*Posoqueria spraguei* Wernham, *Bull. Misc. Inform. Kew* 1914 (2): 66 [30 Mar. 1914] (Wernham 1914).

*Oxyanthus isthmia* W.Mill. & L.H.Bailey, *Stand. Cycl. Hortic.* 4: 2419 [23 Feb. 1916] (Miller & Bailey 1916).

*Posoqueria mutisii* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 11 (5): 232 (Standley 1936).

NOTE. — See note under *Posoqueria gracilis* as to Roemer and Schultes authority.

VERNACULAR NAMES. — Pa: mavinvi-kamwi, mbinbi-wašiune, pune-etni-kamwi • Ka: kuyaken elepali, pipya tamili • Wp: aliwa so ătă, aliwa so u, aliwa u • Wn: meku susu • Br: açucena-do-mato, lirio-do-mato.

HERBARIUM DATA (FG). — 96 collections at CAY. Sel. exs.: *J. Martin s.n.* (original material BM[BM001008911]).

INVENTORY DATA (FG). — 329 trees in 102 plots; $F_{max}$ = 2.8 %; $dbh_{inv}$ = 31.8 cm.

[1490] *Posoqueria longiflora* Aubl. (Fig. 50B)

*Hist. Pl. Guiane* 1: 134 [Jun.-Dec. 1775] (Aublet 1775). — *Kyrtanthus longiflorus* (Aubl.) J.F.Gmel., *Syst. Nat.*, ed. 13[bis], 2 (1): 362 [late Sep.-Nov. 1791] (Gmelin 1791). — *Solena longiflora* (Aubl.) Willd., *Sp. Pl.*, ed. 4 1 (2): 961 [July 1798] (Willdenow 1798).

*Willdenovia schreberi* J.F.Gmel., *Syst. Nat.*, ed. 13[bis], 2 (1): 362 [late Sep.-Nov. 1791] (Gmelin 1791).

*Tocoyena undulatifolia* A.Rich., *Mém. Rubiac.* 167 [Dec. 1830] (Richard 1830).

*Posoqueria acuminata* Mart., *Flora* 24 (2, Beibl.): 79 (Martius 1841).

VERNACULAR NAMES. — Pa: inuva-kamwi • Ka: aimyala posowepo, kapaya wati • Te: aliwa • Wp: aliwa • Wn: asisi, meku susu, mupapatutpë • Nt: agu siton • Cr: grènn-koumarou • Br: açucena-do-mato, papa-terra.

HERBARIUM DATA (FG). — 78 collections at CAY. Sel. exs.: *J.B. Aublet s.n.* (lecto-, P-JJR[P00777902] designated by Lanjouw & Uittien [1940: 156]).

SIZE. — Up to 10 m tall (Taylor *et al.* 2004).
Genus *Psychotria* L.

[1491] *Psychotria anceps* Kunth

*Nova genera et species plantarum [H.B.K.]* 3: 360 [21 Nov. 1819]
(Kunth 1819). — *Uragoga anceps* (Kunth) Kuntze, *Revis. Gen. Pl.*
2: 959 [5 Nov. 1891] (Kuntze 1891).

*Psychotria salicifolia* Kunth, *Nova genera et species plantarum [H.B.K.]* 3: 356 [21 Nov. 1819] (Kunth 1819). — *Uragoga salicifolia* (Kunth) Kuntze, *Revis. Gen. Pl.* 2: 962 [5 Nov. 1891] (Kuntze 1891).

*Psychotria lucida* Kunth, *Nova genera et species plantarum [H.B.K.]* 3: 361 [21 Nov. 1819] (Kunth 1819).

*Psychotria viburnoides* Kunth, *Nova genera et species plantarum [H.B.K.]* 3: 361 [21 Nov. 1819] (Kunth 1819). — *Uragoga viburnoides* (Kunth) Kuntze, *Revis. Gen. Pl.* 2: 963 [5 Nov. 1891] (Kuntze 1891), “*viburnodes*”.

*Psychotria patrisii* DC., *Prodr. [A. P. de Candolle]* 4: 510 [late Sep. 1830] (Candolle 1830). — *Uragoga patrisii* (DC.) Kuntze, *Revis. Gen. Pl.* 2: 962 [5 Nov. 1891] (Kuntze 1891). — *Mapouria patrisii* (DC.) Lemée, *Fl. Guyane Franç.* 3: 549 (Lemée 1954).

*Psychotria chlorantha* Benth., *J. Bot. [Hooker]* 3: 225 (Bentham 1841). — *Uragoga chlorantha* (Benth.) Kuntze, *Revis. Gen. Pl.* 2: 960 [5 Nov. 1891] (Kuntze 1891).

*Mapouria corymbifera* Müll.Arg., *Flora* 59: 464 (Müller 1876). — *Uragoga corymbifera* (Müll.Arg.) Kuntze, *Revis. Gen. Pl.* 2: 960 [5 Nov. 1891] (Kuntze 1891).

*Mapouria corumbensis* S.Moore, *Trans. Linn. Soc. London, Bot.* 4 (3): 380 [“1894-96” publ. Dec. 1895] (Moore 1895). — *Psychotria corumbensis* (S.Moore) Hoehne, *Ind. Bibl. Pl. Col. Com. Rondon:* 387 (Hoehne 1951).

*Psychotria familiarifolia* Wernham, *J. Bot.* 55: 253 (Wernham 1917).

*Mapouria biacuminata* Rusby, *Descr. S. Amer. Pl.* 136 [20 Dec. 1920] (Rusby 1920).

*Mapouria ornithophila* Bremek., *Recueil Trav. Bot. Néerl.* 31: 288 (Bremekamp 1934).

*Psychotria paracatuensis* Standl., *Publ. Field Mus. Nat. Hist., Bot. Ser.* 11 (5): 245 (Standley 1936).

*Psychotria anceps* var. *robustior* Steyerm., *Mem. New York Bot. Gard.* 23: 466 (Steyermark 1972).

HERBARIUM DATA (FG). — 20 collections at CAY. Sel. exs.: *D. Loubry 2413*.

INVENTORY DATA (FG). — 2 trees in 2 plots; Fmax < 1%; dbhinv = 13.4 cm.

[1492] *Psychotria carthagenensis* Jacq.

* Enum. Syst. Pl.*: 16 [Aug.-Sep. 1760] (Jacquin 1760). — *Uragoga carthagenensis* (Jacq.) Kuntze, *Revis. Gen. Pl.* 2: 959 [5 Nov. 1891] (Kuntze 1891).

*Psychotria ardisiifolia* Kunth, *Nova genera et species plantarum [H.B.K.]* 3: 359 [21 Nov. 1891] (Kunth 1891), “*ardisiaefolia*”.

*Psychotria floribunda* Kunth, *Nova genera et species plantarum [H.B.K.]* 3: 359 [21 Nov. 1891] (Kunth 1891). — *Uragoga floribunda* (Kunth) Kuntze, *Revis. Gen. Pl.* 2: 960 [5 Nov. 1891] (Kuntze 1891).

VERNACULAR NAMES. — Pa: pune-etni-seine • Ka: akami ekunali • Wp: tângala yiki, tângala yiki sili, tapi'i ka'alulu sili, uwa ki'iy • Br: rainha.

HERBARIUM DATA (FG). — 52 collections at CAY. Sel. exs.: *M.-F. Prévost 1106*.

INVENTORY DATA (FG). — 4 trees in 2 plots; Fmax < 1%; dbhinv = 14.4 cm.

[1493] *Psychotria ficigemma* DC.

*Prodr. [A. P. de Candolle]* 4: 510 [late Sep. 1830] (Candolle 1830). — *Uragoga ficigemma* (DC.) Kuntze, *Revis. Gen. Pl.* 2: 960 [5 Nov. 1891] (Kuntze 1891). — *Mapouria ficigemma* (DC.) Lemée, *Fl. Guyane Franç.* 3: 549 (Lemée 1954).

HERBARIUM DATA (FG). — 33 collections at CAY. Sel. exs.: *J.-B. Patris s.n.* (original material G-DC, not seen); *P.G. Delprete & F. Crozier 7127*.

INVENTORY DATA (FG). — 44 trees in 22 plots; Fmax = 2.1%; dbhinv = 49.6 cm.

[1494] *Psychotria guianensis* (Aubl.) Clos

*Mém. Acad. Sci. Toulouse*, sér. 7, 10: 217 (Clos 1878). — *Mapouria guianensis* Aubl., *Hist. Pl. Guiane* 1: 175 [Jun.-Dec. 1775] (Aublet 1775), “*Guyannensis*” on plate. — *Psychotria nitida* Willd., *Sp. Pl.*, ed. 4 1 (2): 963 [July 1798] (Willdenow 1798), *nom. illeg. superfl.* (based on *Mapouria guianensis*). — *Simira nitida* Poir., *Encycl. [J. Lamarck et al.]* 7: 196 [6 July 1806] (Poiret 1806), *nom. illeg.* (based on the illegitimate *Psychotria nitida*). — *Psychotria mapouria* Roem. & Schult., *Syst. Veg. [Roemer & Schultes]* 5: 187 [Dec. 1819] (Roemer & Schultes 1819), “*Mapuria*”, *nom. illeg.* (based on the illegitimate *Psychotria nitida*). — *Uragoga mapouria* Kuntze, *Revis. Gen. Pl.* 1: 300 [5 Nov. 1891] (Kuntze 1891), *nom. illeg. superfl.* (based on *Mapouria guianensis*).

*Psychotria magnoliifolia* Kunth, *Nova genera et species plantarum [H.B.K.]* 3: 360 [21 Nov. 1891] (Kunth 1891), “*magnoliaefolia*”. — *Uragoga magnoliifolia* (Kunth) Kuntze, *Revis. Gen. Pl.* 2: 961 [5 Nov. 1891] (Kuntze 1891), “*magnoliaefolia*”.

*Psychotria magnoliifolia* Willd. ex Roem. & Schult., *Syst. Veg. [Roemer & Schultes]* 5: 190 [Dec. 1819] (Roemer & Schultes 1819), “*magnoliaefolia*”. — *Psychotria willdenowii* DC., *Prodr. [A. P. de Candolle]* 4: 505 [late Sep. 1830] (Candolle 1830).

*Palicourea elliptica* Kunth, *Nova genera et species plantarum [H.B.K.]* 3: 369 [21 Nov. 1891] (Kunth 1891).

*Psychotria sambucina* Link ex Roem. & Schult., *Syst. Veg. [Roemer & Schultes]* 5: 188 [Dec. 1819] (Roemer & Schultes 1819). — *Mapouria sambucina* (Link ex Roem. & Schult.) Müll.Arg., *Flora* 59: 458 (Müller 1876). — *Uragoga sambucina* (Link ex Roem. & Schult.) Kuntze, *Revis. Gen. Pl.* 2: 962 [5 Nov. 1891] (Kuntze 1891).

*Psychotria elliptica* Humb. & Bonpl. ex Roem. & Schult., *Syst. Veg. [Roemer & Schultes]* 5: 189 [Dec. 1819] (Roemer & Schultes 1819). — *Uragoga elliptica* (Humb. & Bonpl. ex Roem. & Schult.) Kuntze, *Revis. Gen. Pl.* 1: 300 [5 Nov. 1891] (Kuntze 1891).

*Mapouria myriantha* Müll.Arg., *Fl. Bras. [Martius]* 6 (5): 400 [1 July 1881] (Müller 1881). — *Uragoga myriantha* (Müll.Arg.) Kuntze, *Revis. Gen. Pl.* 2: 961 [5 Nov. 1891] (Kuntze 1891).