<table><caption>**Table 2.** Ancestral state transitions estimated in MrBayes under the Mkv model. The degree of support for estimated synapomorphies is indicated via asterisks (*), three asterisks being the highest degree of support. “State fixed” indicates high support for daughter node coupled with uncertainty at the parent node due to state heterogeneity in other members of the clade. “Megalyra–Carminator clade” = †Cretodinapsis + Megalyra + †Prodinapsis + (Carminator + Cryptalyra).</caption><thead><tr><th>Char</th><th>State</th><th>BPP</th><th>State</th><th>BPP</th><th>Interpretation</th></tr></thead><tbody><tr><td>(A)</td><td colspan="2">Megalyridae node</td><td colspan="2">Megazarinae node</td><td>—</td></tr><tr><td>31</td><td>0</td><td>0.64</td><td>1</td><td>0.96</td><td>State fixed at daughter node</td></tr><tr><td>42</td><td>0</td><td>0.62</td><td>1</td><td>0.98</td><td>State fixed at daughter node</td></tr><tr><td>(B)</td><td colspan="2">Megalyridae node</td><td colspan="2">Megalyrinae node</td><td>—</td></tr><tr><td>18</td><td>1</td><td>0.51</td><td>2</td><td>0.96</td><td>State fixed at daughter node</td></tr><tr><td>39</td><td>1</td><td>0.77</td><td>2</td><td>0.97</td><td>State fixed at daughter node</td></tr><tr><td>43</td><td>0</td><td>0.78</td><td>1</td><td>0.97</td><td>State fixed at daughter node</td></tr><tr><td>44</td><td>0</td><td>0.79</td><td>1</td><td>0.91</td><td>State fixed at daughter node</td></tr><tr><td>(C)</td><td colspan="2">Megalyrinae node</td><td colspan="2">Rigel + Megalyridia node</td><td>—</td></tr><tr><td>6</td><td>1</td><td>0.84</td><td>0</td><td>0.99</td><td>State fixed at daughter node</td></tr><tr><td>13**</td><td>1</td><td>1.00</td><td>0</td><td>0.95</td><td>Well-supported synapomorphy</td></tr><tr><td>23</td><td>0</td><td>0.54</td><td>1</td><td>0.99</td><td>State fixed at daughter node</td></tr><tr><td>24</td><td>1</td><td>0.68</td><td>0</td><td>0.99</td><td>State fixed at daughter node</td></tr><tr><td>30**</td><td>0</td><td>0.93</td><td>1</td><td>1.00</td><td>Well-supported synapomorphy</td></tr><tr><td>47</td><td>2</td><td>0.75</td><td>1</td><td>1.00</td><td>State fixed at daughter node</td></tr><tr><td>48</td><td>2</td><td>0.81</td><td>1</td><td>1.00</td><td>State fixed at daughter node</td></tr><tr><td>49</td><td>2</td><td>0.78</td><td>1</td><td>0.97</td><td>State fixed at daughter node</td></tr><tr><td>(D)</td><td colspan="2">Megalyrinae node</td><td colspan="2">Megalyra–Carminator node</td><td>—</td></tr><tr><td>10**</td><td>0</td><td>0.97</td><td>1</td><td>0.85</td><td>Well-supported synapomorphy (note 1)</td></tr><tr><td>11**</td><td>0</td><td>0.99</td><td>1</td><td>0.93</td><td>Well-supported synapomorphy</td></tr><tr><td>30*</td><td>0</td><td>0.93</td><td>2</td><td>0.62</td><td>Weakly supported synapomorphy</td></tr><tr><td>38***</td><td>0</td><td>1.00</td><td>2</td><td>0.98</td><td>Highly supported synapomorphy (note 2)</td></tr><tr><td>40**</td><td>0</td><td>0.96</td><td>2</td><td>0.77</td><td>Well-supported synapomorphy (note 3)</td></tr><tr><td>44**</td><td>1</td><td>0.91</td><td>0</td><td>0.87</td><td>Well-supported synapomorphy (note 4)</td></tr><tr><td>(47*)</td><td>2</td><td>0.75</td><td>1</td><td>0.97</td><td>Very weakly supported synapomorphy (note 5)</td></tr><tr><td>48**</td><td>2</td><td>0.81</td><td>1</td><td>1.00</td><td>Well-supported synapomorphy (note 6)</td></tr></tbody></table>

Note 1. Implies synapomorphic reversal to 0 for *Cryptalyra + Carminator*.

Note 2. Variable in *Megalyra*; derivation of state 2 in *Megalyridia* implied to be homoplastic.

Note 3. Variable in †*Prodinapsis*; derivation of state 2 in *Megalyridia* implied to be homoplastic.

Note 4. Derivation of state 1 for *Cryptalyra + Carminator* implied to be synapomorphic.

Note 5. Pattern driven by †*Prodinapsis*; highly variable across tree; should be treated with skepticism.

Note 6. Implies derivation of state 1 for *Rigel + Megalyridia* is a homoplastic synapomorphy.

garding †Cretodinapsis, when this taxon is excluded from the parsimony analyses, †Prodinapsis is still retrieved as sister of *Megalyra* (Fig. 11), supported by the absence of parapsides (23:0) and the absence of 1A distally of cu-a (44:0). This sistergroup relationship is furthermore corroborated by the paleodistribution of the two genera, *Megalyra* being represented in Baltic amber, where †*Prodinapsis* is also widely found (Poinar and Shaw 2007; Perrichot 2009). On the other hand, under the conditions of the Bayesian analyses, the clade of †*Prodinapsis*, †Cretodinapsis, *Megalyra*, and *Cryptalyrin* is supported by seven characters (Table 2): presence of the median sulcus on the vertex and eye setae (10:1, 11:1), rugose metacoxal sculpture (30:2), absence of the fore wing Rs (38:2, 40:2), fore wing distal 1a darkened (44:0), and areolate-rugose propodeal sculpture (48:1). Therefore, the †Cretodinapsini is synonymized with *Megalyrin*, which is now composed of three genera.

Under Bayesian analyses, we also found support for the clade [*Rigel + Megalyridia*], which conflicts with the results of the parsimony analyses. With the equal rates

model, this grouping considerably strengthens the relationship of [*Megalyra + †Cretodinapsis + †Prodinapsis + (Carminator + Cryptalyra)*] (Fig. 12). [*Rigel + Megalyridia*] is supported by the flagellum longer than head + mesosoma (6:0, also in †*Cretolyra gen. nov.*, †*Kamyristi gen. nov.*, †*Megalava*, *Ettchellsia*, and the outgroup), the occipital carina basally curving towards the back of the head (13:0), the parapsides present (23:1, also in most Cretaceous taxa and all extant genera except *Megalyra*), the axillae continuous medially (24:0, also in *Dinapsis*, *Carminator*, *Ettchellsia*, *Megalyra*, *Neodinapsis* and some Cretaceous taxa), the metacoxa shagreened (30:1, also in *Carminator*, *Dinapsis*, *Neodinapsis* and †*Kamyristi exfrigore gen. et sp. nov.*), the Rs vein of the hind wing medium-sized (47:1, also in *Neodinapsis* and †*Prodinapsis*), the propodeum areolate-rugose (48:1, also in *Cryptalyra*, *Megalyra*, *Neodinapsis*, †*Cretodinapsis*, †*Kamyristi gen. nov.*, †*Megallica* and †*Prodinapsis*) and the ovipositor about as long as the body (49:1, also in *Dinapsis*, †*Megazar* and †*Prodinapsis*). Most of these characters are very homoplastic, except for the configuration
of the occipital carina, which is unique within the family. This finding highlights the instability of the relationships within the Megalyrinae, mainly because of the wide morphological diversity between and even within genera and the incomplete coding of some fossils.

The addition of new fossil taxa to the Megalyridae is not without consequences for the classification of the family. Only two of the non-monotypic tribes previously defined are consistently retrieved and can be diagnosed precisely (†Megalavini **trib. nov.**, Cryptalyrini); two others, Dinapsini s.l. and †Cretodinapsini, are not retrieved as monophyletic. To properly align classification with phylogeny, following the current tribal system, we propose a number of changes in the Megalyridae classification. First, we recognize the two main megaljid clades as subfamilies, the †Megazarinae **stat. nov.** and Megalyrinae. These two subfamilies are likely to represent distinct lineages within the Megalyridae, the †Megazarinae **stat. nov.** comprising exclusively Cretaceous taxa and the Megalyrinae constituting the crown group including all the extant taxa as well as a mix of extinct taxa from the Cretaceous onwards. Secondly, under parsimony, four additional non-conflicting groupings above the genus level are retrieved and can be consistently diagnosed and recognized as tribes: Cryptalyrini, Dinapsini s.str., Megalyrini (expanded with the inclusion of †Cretodinapsini), and †Megalavini **trib. nov.** Finally, three new monotypic tribes are created for Cretaceous genera, †Kamyristini **trib. nov.**, †Megallicini **trib. nov.**, and †Valaaini **trib. nov.** Such a large number of tribes (ten, of which six are monotypic) may seem exaggerated for a group as relictual as the Megalyridae but is necessary to maintain congruence with the phylogenetic results. Furthermore, the Megalyridae are an ancient family, whose past diversity is only at the dawn of its discovery and those tribes could prove of use for future taxonomic works. A summary of the revised classification is presented in Appendix S1.

Vilhelmsen et al. (2010a) stated that †*Cretodinap*
sis should be considered as an incertae sedis member
of Evaniomorpha despite the presence of several char-
acters reminiscent of the Megalyridae; these characters
are probably plesiomorphic and not unique to the family
(e.g., the vertex with a sulcus, the mesoscutum with a me-
dian mesoscutal sulcus, the large axillae, the similar fore
wing venation, the long ovipositor). However, it has so
far not been possible to establish the presence of a sub-
antennal groove and the configuration of the anterior thor-
acic spiracle in †*Cretodinap*
sis. While †*Cretodinap*
sis
usually jumped around depending on the parameters in
the analyses of Vilhelmsen et al. (2010a), they retrieved it
nested with extant genera when the †Maimetshidae were
excluded. Its position within the Megalyrinae is more sta-
ble in our analyses but still poorly supported. This would
strengthen the placement of †*Cretodinap*
sis among the
Megalyridae. However, it is unique in the subfamily in
having the vein M+Cu pigmented on the forewing while
it is spectral to absent in other megalyrines. This resem-
bles more the †Megazarinae **stat. nov.**, but the venation
is much reduced compared to other species and without

clarifying the diagnostic characters, it is not possible to
decide further on the position of †*Cretodinapsis*. Howev-
er, the material is unavailable for study at this time.

Hopefully, further improvement of the understanding
of the phylogeny of Megalyridae will be achieved through
the study of the fossil record. However, we do not current-
ly know of megalyrid morphotypes in Burmese amber
in addition to those we describe here; Perrichot (2009)
widely surveyed fossils from European Cretaceous and
Paleogene ambers. New data could be provided by the
descriptions of taxa in geographically distinct deposits
such as the Aptian Congolese amber, Turonian Raritan
amber, Campanian Canadian amber, lower Eocene Fush-
un (China) and Cambay (India) amber, and lower-middle
Miocene Ethiopian and Zhangpu (China) amber.

6. Conclusions

The description of new fossil Megalyridae has enhanced
our understanding of the evolutionary history of this fam-
ily. It is evident that the Megalyridae experienced exten-
sive radiation by the mid-Cretaceous, apparently includ-
ing possible close relatives of at least some of the extant
genera; however, a substantial portion of the Cretaceous
diversity, i.e., the †Megazarinae stat. nov., did not leave
any extant descendants. The general decline in diversi-
ty and contraction of distribution of the family, correlat-
ed with environmental changes (i.e., a general cooling
during the Cenozoic), is highlighted by the documenta-
tion of species from Siberia, located higher than 65° N
during Upper Cretaceous (Rasnitsyn et al. 2016; Nadein
and Perkovsky 2018), and now under a polar climate.
Whereas fossil megalyrids were only known from regions
where they are presently absent, the Burmese amber taxa
that are described herein are the first fossils found in a
paratropical habitat in which they are currently present
(i.e., *Ettchellsia* and *Carminator*). Thus, during the warm
climate of the mid-Cretaceous, the Megalyridae appear
to be at least as—and likely more—diverse compared to
the present, the Cretaceous genera surpassing the modern
genera in number, and possibly more widely distributed,
although confirmed megalyrid fossils from Gondwana
deposits are still missing.

7. Author contributions

**Manuel Brazidec:** Conceptualization; data curation; formal analysis; investigation; methodology; project administration; writing – original draft; writing – review and editing; visualization; validation. **Lars Vilhelmsen:** Conceptualization; formal analysis; investigation; methodology; resources; writing – review and editing; validation. **Brendon E. Boudinot:** Formal analysis; funding acquisition; investigation; methodology; resources; writing – review and editing; visualization; validation. **Adrian K. F. Richter:** Methodology; resources; writing – review and editing; visualization; validation. **Jörg U. Hammel:** Methodology; resources; writing – review and editing; visualization; validation. **Evgeny**
of the occipital carina, which is unique within the family. This finding highlights the instability of the relationships within the Megalyrinae, mainly because of the wide morphological diversity between and even within genera and the incomplete coding of some fossils.

The addition of new fossil taxa to the Megalyridae is not without consequences for the classification of the family. Only two of the non-monotypic tribes previously defined are consistently retrieved and can be diagnosed precisely (†Megalavini **trib. nov.**, Cryptalyrini); two others, Dinapsini s.l. and †Cretodinapsini, are not retrieved as monophyletic. To properly align classification with phylogeny, following the current tribal system, we propose a number of changes in the Megalyridae classification. First, we recognize the two main megaljid clades as subfamilies, the †Megazarinae **stat. nov.** and Megalyrinae. These two subfamilies are likely to represent distinct lineages within the Megalyridae, the †Megazarinae **stat. nov.** comprising exclusively Cretaceous taxa and the Megalyrinae constituting the crown group including all the extant taxa as well as a mix of extinct taxa from the Cretaceous onwards. Secondly, under parsimony, four additional non-conflicting groupings above the genus level are retrieved and can be consistently diagnosed and recognized as tribes: Cryptalyrini, Dinapsini s.str., Megalyrini (expanded with the inclusion of †Cretodinapsini), and †Megalavini **trib. nov.** Finally, three new monotypic tribes are created for Cretaceous genera, †Kamyristini **trib. nov.**, †Megallicini **trib. nov.**, and †Valaaini **trib. nov.** Such a large number of tribes (ten, of which six are monotypic) may seem exaggerated for a group as relictual as the Megalyridae but is necessary to maintain congruence with the phylogenetic results. Furthermore, the Megalyridae are an ancient family, whose past diversity is only at the dawn of its discovery and those tribes could prove of use for future taxonomic works. A summary of the revised classification is presented in Appendix S1.

Vilhelmsen et al. (2010a) stated that †*Cretodinap*
sis should be considered as an incertae sedis member
of Evaniomorpha despite the presence of several char-
acters reminiscent of the Megalyridae; these characters
are probably plesiomorphic and not unique to the family
(e.g., the vertex with a sulcus, the mesoscutum with a me-
dian mesoscutal sulcus, the large axillae, the similar fore
wing venation, the long ovipositor). However, it has so
far not been possible to establish the presence of a sub-
antennal groove and the configuration of the anterior thor-
acic spiracle in †*Cretodinap*
sis. While †*Cretodinap*
sis
usually jumped around depending on the parameters in
the analyses of Vilhelmsen et al. (2010a), they retrieved it
nested with extant genera when the †Maimetshidae were
excluded. Its position within the Megalyrinae is more sta-
ble in our analyses but still poorly supported. This would
strengthen the placement of †*Cretodinap*
sis among the
Megalyridae. However, it is unique in the subfamily in
having the vein M+Cu pigmented on the forewing while
it is spectral to absent in other megalyrines. This resem-
bles more the †Megazarinae **stat. nov.**, but the venation
is much reduced compared to other species and without

clarifying the diagnostic characters, it is not possible to
decide further on the position of †*Cretodinapsis*. Howev-
er, the material is unavailable for study at this time.

Hopefully, further improvement of the understanding
of the phylogeny of Megalyridae will be achieved through
the study of the fossil record. However, we do not current-
ly know of megalyrid morphotypes in Burmese amber
in addition to those we describe here; Perrichot (2009)
widely surveyed fossils from European Cretaceous and
Paleogene ambers. New data could be provided by the
descriptions of taxa in geographically distinct deposits
such as the Aptian Congolese amber, Turonian Raritan
amber, Campanian Canadian amber, lower Eocene Fush-
un (China) and Cambay (India) amber, and lower-middle
Miocene Ethiopian and Zhangpu (China) amber.

6. Conclusions

The description of new fossil Megalyridae has enhanced
our understanding of the evolutionary history of this fam-
ily. It is evident that the Megalyridae experienced exten-
sive radiation by the mid-Cretaceous, apparently includ-
ing possible close relatives of at least some of the extant
genera; however, a substantial portion of the Cretaceous
diversity, i.e., the †Megazarinae stat. nov., did not leave
any extant descendants. The general decline in diversi-
ty and contraction of distribution of the family, correlat-
ed with environmental changes (i.e., a general cooling
during the Cenozoic), is highlighted by the documenta-
tion of species from Siberia, located higher than 65° N
during Upper Cretaceous (Rasnitsyn et al. 2016; Nadein
and Perkovsky 2018), and now under a polar climate.
Whereas fossil megalyrids were only known from regions
where they are presently absent, the Burmese amber taxa
that are described herein are the first fossils found in a
paratropical habitat in which they are currently present
(i.e., *Ettchellsia* and *Carminator*). Thus, during the warm
climate of the mid-Cretaceous, the Megalyridae appear
to be at least as—and likely more—diverse compared to
the present, the Cretaceous genera surpassing the modern
genera in number, and possibly more widely distributed,
although confirmed megalyrid fossils from Gondwana
deposits are still missing.

7. Author contributions

**Manuel Brazidec:** Conceptualization; data curation; formal analysis; investigation; methodology; project administration; writing – original draft; writing – review and editing; visualization; validation. **Lars Vilhelmsen:** Conceptualization; formal analysis; investigation; methodology; resources; writing – review and editing; validation. **Brendon E. Boudinot:** Formal analysis; funding acquisition; investigation; methodology; resources; writing – review and editing; visualization; validation. **Adrian K. F. Richter:** Methodology; resources; writing – review and editing; visualization; validation. **Jörg U. Hammel:** Methodology; resources; writing – review and editing; visualization; validation. **Evgeny**
**E. Perkovsky**: Funding acquisition; resources; writing – review and editing; validation. **Yong Fan**: Resources; writing – review and editing; validation. **Zhen Wang**: Resources; writing – review and editing; visualization; validation. **Qiong Wu**: Resources; writing – review and editing; visualization; validation. **Bo Wang**: Funding acquisition; resources; writing – review and editing; visualization; validation. **Vincent Perrichot**: Conceptualization; data curation; funding acquisition; investigation; methodology; project administration; resources; writing – review and editing; visualization; supervision; validation.

## 8. Competing interests

The authors have declared that no competing interests exist.

## 9. Acknowledgments

We are very grateful to all persons and colleagues who kindly provided or facilitated access to the material studied here: the late Jens-Wilhelm Janzen and his daughter Eva (Seevetal, Germany); and Prof. Alexandr Rasnitsyn (PIN, Russia). We warmly thank the editorial team of Arthropod Systematics & Phylogeny and two anonymous reviewers for careful consideration and suggestions on our manuscript. We thank the Willi Hennig Society for making available the TNT software for free. This research was supported by the National Natural Science Foundation of China (42125201) and Second Tibetan Plateau Scientific Expendition and Research (2019QZKK0706). This work is a contribution to the Deep-time Digital Earth (DDE) Big Science Program. B.E.B. was supported by an Alexander von Humboldt research fellowship (2020–2022) and a Peter S. Buck research fellowship from the Smithsonian Institution (2023). We acknowledge provision of beamtime, related to the proposal BAG-20190010 and I-20200136- at beamline P05 at PETRA III at DESY, a member of the Helmholtz Association (HGF), and the support during the beam times by Hereon team members Fabian Wilde, Julian Moosmann, and Felix Beckmann. This research was supported in part through the Maxwell computational resources operated at Deutsches Elektronen-Synchrotron DESY, Hamburg, Germany. E.E.P. acknowledge the support of Scholars at Risk Ukraine (SARU) jointly funded by the Villum Foundation, Carlsberg Foundation and Novo Nordisk Foundation.

## 10. References

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Chen H, Liuhe B, Zhang X (2021) Two new species of the family Megalyridae (Hymenoptera) from China. ZooKeys 1043: 21–31. https://doi.org/10.3897/zookeys.1043.65223

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**E. Perkovsky**: Funding acquisition; resources; writing – review and editing; validation. **Yong Fan**: Resources; writing – review and editing; validation. **Zhen Wang**: Resources; writing – review and editing; visualization; validation. **Qiong Wu**: Resources; writing – review and editing; visualization; validation. **Bo Wang**: Funding acquisition; resources; writing – review and editing; visualization; validation. **Vincent Perrichot**: Conceptualization; data curation; funding acquisition; investigation; methodology; project administration; resources; writing – review and editing; visualization; supervision; validation.

## 8. Competing interests

The authors have declared that no competing interests exist.

## 9. Acknowledgments

We are very grateful to all persons and colleagues who kindly provided or facilitated access to the material studied here: the late Jens-Wilhelm Janzen and his daughter Eva (Seevetal, Germany); and Prof. Alexandr Rasnitsyn (PIN, Russia). We warmly thank the editorial team of Arthropod Systematics & Phylogeny and two anonymous reviewers for careful consideration and suggestions on our manuscript. We thank the Willi Hennig Society for making available the TNT software for free. This research was supported by the National Natural Science Foundation of China (42125201) and Second Tibetan Plateau Scientific Expendition and Research (2019QZKK0706). This work is a contribution to the Deep-time Digital Earth (DDE) Big Science Program. B.E.B. was supported by an Alexander von Humboldt research fellowship (2020–2022) and a Peter S. Buck research fellowship from the Smithsonian Institution (2023). We acknowledge provision of beamtime, related to the proposal BAG-20190010 and I-20200136- at beamline P05 at PETRA III at DESY, a member of the Helmholtz Association (HGF), and the support during the beam times by Hereon team members Fabian Wilde, Julian Moosmann, and Felix Beckmann. This research was supported in part through the Maxwell computational resources operated at Deutsches Elektronen-Synchrotron DESY, Hamburg, Germany. E.E.P. acknowledge the support of Scholars at Risk Ukraine (SARU) jointly funded by the Villum Foundation, Carlsberg Foundation and Novo Nordisk Foundation.

## 10. References

Baltazar CR (1962) *Etchellsia philippinensis* sp. nov. (Dinapsinae, Megalyridae, Hymenoptera). The Philippine Journal of Science 90: 219–220.

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Perrichot V (2009) Long-tailed wasps (Hymenoptera: Megalyridae) from Cretaceous and Paleogene European amber. Paleontological Contributions 1: 1–19. https://doi.org/10.17161/PC.1808.5468

Perrichot V, Nel A, Néraudeau D (2004) A new, enigmatic, evaniomorphan in the Albian amber of France (Insecta, Hymenoptera). Journal of Systematic Palaeontology 2(2): 159–162. https://doi.org/10.1017/S1477201904001245

Poinar GO, Shaw SR (2007) *Megalyra baltica* Poinar and Shaw n. sp. (Hymenoptera: Megalyridae), a long-tailed wasp from Baltic amber. Zootaxa 1478(1): 65–68. https://doi.org/10.11646/zootaxa.1478.1.7

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# Supplementary Material 1

## Appendices S1, S2

**Authors:** Brazidec M, Vilhelmsen L, Boudinot BE, Richter AKF, Hammel JU, Perkovksy EE, Fan Y, Whang Z, Wu Q, Wang B, Perrichot V (2024)

**Data type:** .zip

**Explanation notes:** Appendix S1. Revised classification of the Megalyridae. — Appendix S2. List of characters used in the phylogenetic analyses.

**Copyright notice:** This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbL/1.0). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited.

**Link:** https://doi.org/asp.82.e111148.suppl1
# Supplementary Material 1

## Appendices S1, S2

**Authors:** Brazidec M, Vilhelmsen L, Boudinot BE, Richter AKF, Hammel JU, Perkovksy EE, Fan Y, Whang Z, Wu Q, Wang B, Perrichot V (2024)

**Data type:** .zip

**Explanation notes:** Appendix S1. Revised classification of the Megalyridae. — Appendix S2. List of characters used in the phylogenetic analyses.

**Copyright notice:** This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbL/1.0). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited.

**Link:** https://doi.org/asp.82.e111148.suppl1