Schoenoplectiella erecta (Poir.) Lye ssp. raynalii
(Schuyler) Beentje (Cyperaceae) – a new record to India
from Ossudu Bird Sanctuary, Villupuram District, Tamil
Nadu

Pradeep Chandrasegrane, P Umamaheswari, Natesan Balachandran, Raphael Mathevet

<figure><img src="image_1.png" /></figure>

To cite this version:

Pradeep Chandrasegrane, P Umamaheswari, Natesan Balachandran, Raphael Mathevet. Schoenoplectiella erecta (Poir.) Lye ssp. raynalii (Schuyler) Beentje (Cyperaceae) – a new record to India from Ossudu Bird Sanctuary, Villupuram District, Tamil Nadu. Journal of Threatened Taxa, 2023, 15 (2), pp.22741-22745. 10.11609/jott.7988.15.2.22741-22745. hal-04080960

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Building evidence for conservation globally
# Journal of Threatened Taxa

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# Schoenoplectiella erecta (Poir.) Lye ssp. raynalii (Schuyler) Beentje (Cyperaceae) – a new record to India from Ossudu Bird Sanctuary, Villupuram District, Tamil Nadu

Chandrasegrane Pradeep<fnref n="1" /> ID, Paneerselvam Umamaheswari<fnref n="2" /> ID, Natesan Balachandran³ ID &
Raphael Mathevet<fnref n="4" /> ID

**Abstract:** *Schoenoplectiella erecta* subsp. *raynalii* (Cyperaceae) is recorded for the first time from India and Asia. This taxon was collected in Ossudu Bird Sanctuary, Villupuram district of Tamil Nadu, southern India. Detailed description including microscopic study of the glume and nutlets with digital images, morphological characters, habitat, and key characters between the two subspecies are provided.

**Keywords:** Coromandel coast, India, migratory birds, new record, Pondicherry, wetland.

A new genus *Schoenoplectiella* was established by Lye (2003) with 26 species and it was separated from the genus *Schoenoplectus* (Rchb.) Palla. based on the rbcL suprageneric phylogeny (Muasya et al. 1998) both the genera were differentiated from the heterogeneous and un-natural genus, *Scirpus* L. Characteristically, *Schoenoplectiella* has members that are annuals, rarely perennial, rhizome very short hidden among the culm-bases, prostrate or elongate, glumes entire at apex, and nutlets transversely rugulose to sharply ridged whereas *Schoenoplectus* has perennial, rhizome elongate,

creeping or ascending, glumes notched or emargiante or bifid, and nutlets generally smooth (Hayasaka 2012). Currently, the genus has 65 accepted species (POWO 2022) and these are distributed from warm temperate to tropical regions of Africa, America, Asia, and Madagascar (Verloove et al. 2016). In India, 10 species were recorded (Mao & Dash 2020) and only five species are known from Tamil Nadu (Narasimhan & Sheeba 2021).

During recurrent botanical surveys for the last three year (from September 2018 to August 2021) an interesting plant belonging to the genus *Schoenoplectiella* was collected from the shores of the freshwater Lake Ossudu, in Ossudu Bird Sanctuary, a protected area, in Tamil Nadu and Pondicherry regions, southeastern India. On critical microscopic examination and referring to available literatures (Smith 2003; Xanthos & Browning 2015) the collected voucher specimen was identified as *Schoenoplectiella erecta* ssp. *raynalii*. Our identification was confirmed further by comparing the isotype image deposited at K (A.M.Yalala 425-K000416875). Further

**Editor:** Pankaj Kumar, Department of Plant and Soil Science, Texas Tech University, Lubbock, Texas, USA. **Date of publication:** 26 February 2023 (online & print)

**Citation:** Pradeep, C., P. Umamaheswari, N. Balachandran & R. Mathevet (2023). *Schoenoplectiella erecta* (Poir.) Lye ssp. *raynalii* (Schuyler) Beentje (Cyperaceae) – a new record to India from Ossudu Bird Sanctuary, Villupuram District, Tamil Nadu. *Journal of Threatened Taxa* 15(2): 22741–22745. https://doi.org/10.11609/jott.7988.15.2.22741-22745

**Copyright:** © Pradeep et al. 2023. Creative Commons Attribution 4.0 International License. JOTT allows unrestricted use, reproduction, and distribution of this article in any medium by providing adequate credit to the author(s) and the source of publication.

**Funding:** AFD [Project: RUSE].

**Competing interests:** The authors declare no competing interests.

**Acknowledgements:** We thank Dr. D. Barboni for a careful review of the manuscript; RUSE AFD, France for the financial support for the survey; the head of forest force and wildlife, Chennai, and district forest officer, Villupuram district, Tamil Nadu granted permission to do botanical survey at Ossudu and Kazuveli wetlands; to the board of trustees, RBG, KEW for the digital image; University of South Florida Herbarium for their digital herbarium image and AURO herbarium for referring the *S. lateriflora* specimens. We also thanks to the anonymous reviewer(s) that shaped this manuscript in better form.

<figure><img src="image_7.png" /></figure>

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<footnote n="1">¹⁻⁴ Ecology Department, French Institute of Pondicherry, Pondicherry 605001, India.</footnote>

<footnote n="2">² Department of Botany, Kanchi Mamunivar Government Institute for Postgraduate Studies and Research, Puducherry 605008, India.</footnote>

<footnote n="4">⁴ UMR 5175 Centre d'Ecologie Fonctionnelle et Evolutive (CEFE), Campus CNRS, 1919 route de Mende, 34293 Montpellier, France.</footnote>

<footnote n="1">¹ pradeep.c@ifpindia.org, ² umamaheswari@ifpindia.org, ³ balachandran.n@ifpindia.org (corresponding author),</footnote>

<footnote n="4">⁴ raphael.mathevet@ifpindia.org</footnote>
<figure><img src="image_10.png" /><figcaption>Figure 1. Study area – Ossudu Bird Sanctuary.</figcaption></figure>

perusal of literatures (Cook 1998; Panda et al. 2002; Prasad & Singh 2002; Ansari et al. 2016; Kar et al. 2016; Mao & Dash 2020; Narasimhan & Sheeba 2021) we come to know that this taxon was not yet recorded in Asia, India, and regional floras; however, *Schoenoplectiella erecta* ssp. *erecta* was recorded from northern India. In the Lake Ossudu Bird Sanctuary, three *Schoenoplectiella* species also occur: *S. atriculata* (L.) Lye, *S. lateriflora* (J.F.Gmel.) Lye, and *S. juncoides* (Roxb.) Lye.

## MATERIALS AND METHODS

Botanical exploration and ecological studies were conducted from September 2018 to August 2021 in Ossudu (Figure 1) along the Coromandel Coast, Villupuram district of Tamil Nadu. Ossudu Bird Sanctuary falls in two political boundaries, viz., Union Territory of Pondicherry and Villupuram district of Tamil Nadu. During the survey an unfamiliar sedge was found and collected from the eastern shores of Ossudu Lake and the collected specimen was processed and deposited at Herbarium, French Institute of Pondicherry (HIFP) for further studies. Camera attached light microscope was used to examine the morphological features of glume and nutlets to understand the morphological differences. Recent publications (Mao & Dash 2020; Narasimhan & Sheeba 2021) and consultation of herbaria (Madras Herbarium (MH), Rapinath Herbarium Tiruchirapalli (RHT), Deccan Reginal Centre (DRC), Auroville Herbarium (AURO)) for the occurrence and international datasets (GBIF- the Global Biodiversity Information Facility— https://www gbif org, COL- Catalogue of Life— https:// www catalogueoflife org, POWO- Plants of the World Online— https://powo science kew org/ and USDA- United States Department of Agriculture— https:// plants usda gov) were referred for the global distribution

range of this species.

## RESULTS

The inflorescence of *Schoenoplectiella* is capitate or anthellate. The anthella of spikelets are simple to decompound due to the presence of branched or unbranched peduncles with few to numerous sessile spikelets and they are densely crowded. This characteristic feature was recorded in both the inflorescence of *S. lateriflora* and *S. erecta*. On critical examination of the herbarium specimens deposited at AURO (4751, 10317, 11940), *S. lateriflora* was 10–20 cm high with decompound inflorescence, peduncles 3–15 mm high, style 3-branched, achene trigonous whereas *S. erecta* is more than 30 cm high, inflorescence decompound, peduncles 15–65 mm long, secondary peduncles to 8–12 mm long, style 2-branched and nutlet plano-convex.

The two known subspecies, *Schoenoplectiella erecta* ssp. *raynalii* is similar to *S. erecta* ssp. *erecta* but differs by its style and nutlets. Following key can be used to differentiate them:

1. Nutlet biconvex, style 2-fid, slightly wrinkled ....
.................................................. *S. erecta* ssp. *erecta*

1. Nutlet plano-convex, style 3-fid, distinctly rugose ............................ *S. erecta* ssp. *raynalii*

## Taxonomic treatment

*Schoenoplectiella erecta* (Poiret) Lye ssp. *raynalii* (Schuyler) Beentje Fl. Trop. E. Africa, Cyperaceae, 34, 2010. (Image 1)

*Schoenoplectus erectus* (Poiret) Palla *ex* J. Raynal ssp. *raynalii* (Schuyler) Lye, Nordic J. Bot. 3(2): 243. 1983.

*Scirpus raynalii* Schuyler, Notul. Nat. Acad. Nat. Sci.
<figure><img src="image_11.png" /><figcaption>A</figcaption></figure>

<figure><img src="image_12.png" /><figcaption>B</figcaption></figure>

<figure><img src="image_13.png" /><figcaption>C</figcaption></figure>

<figure><img src="image_14.png" /><figcaption>D</figcaption></figure>

<figure><img src="image_15.png" /><figcaption>E</figcaption></figure>

<figure><img src="image_16.png" /><figcaption>F</figcaption></figure>

<figure><img src="image_17.png" /><figcaption>G</figcaption></figure>

Image 1. *Schoenoplectiella erecta* subsp. *raynali*: A—The present collection | B—Isotype image from KEW | C—in fluorescence in close view | D—plant in natural condition | E—dorsal, ventral and lateral view of glumes | F—gynoecium with 3-fid stigma | G—young and matured nutlet. © Balachandran & Uma Maheswari.
<figure><img src="image_18.png" /></figure>

Philadelphia 438: 1, figs. 1, 3, 6. 1971.

Type: BOTSWANA, Maun district, 6.2.1964, Yalala 425 (K-K000416875!-Isotype-digital image seen)

## Description

Annual herbs, culms densely tufted, 30-43 cm (including inflorescence bract), cylindrical, ridged when dry, 1.6-2 mm thick. Leaves 1-3, 2.5-13 cm long, rarely equalling the culm; sheaths brown, 5-9 cm long, ribbed, disintegrating to fibres. Inflorescence, pseudolateral, anthella-decompound in 2-5 pedunculate corymb, primary peduncle 0.5-6 cm long, secondary rachis 8-12 mm long; overtopped by lower bract, involucral bract stem like, erect, 4.5-13 cm long, secondary bract 0.5-4 cm long. Spikelets, in clusters, 1-5, 5-7 × 2-3 mm; green-cream, reddish brown when mature; ovoid-oblong, 3(5)-13(18) × 2-3.5 mm; glume straw-coloured, with brown mosaic, central region often greenish when fresh, ovate-obovate, 2.53.4(5) × 1.5 mm, smooth, mucronate, margins scarious. Perianth absent, stamens 3, style 3-fid. Achenes almost blackish brown when ripe, planoconvex when matured, obovoid, 1.2-1.6 × 1.1-1.5 mm, with sharp ridges along the margins, distinctly and transversely rugose.

Flowering and fruiting: from January-April.

Habitat: Along the shores of the fresh water lake at the elevational range between 30 and 40 m. It was found growing along with *S. lateriflorus* and *S. juncoides* of Cyperaceae, *Persicaria glabra* (Willd.) M.Gomez of Polygonaceae, *Ludwigia perennis* L. of Onagraceae, *Dinebra polystachyos* (R.Br.) E.A.Kellogg of Poaceae, *Aponogeton natans* (L.) Engl. & K.Krause of Aponogetonaceae.

Distribution: Africa, Asia, Australia, Mexico, and South America.

Specimen examined: India, Tamil Nadu, Villupuram district, Ossudu Bird Sanctuary, 11.9577° N, 79.7456° E, 18 m, 8 February 2020, Pradeep & Balachandran 27514 (HIFP!, two sheets).

## Conservation status

Considered as 'Least Concern' on the IUCN Red List of Threatened Species (Mesterházy 2020), and in recent days the range of distribution has been extended from Africa, America to Asia, and Australia. Probably the migratory granivorous ducks (Mallards/the whistling ducks) might have played a major role in extending the distribution of this species. This study also proves the report of Kleyheeg et al. (2019) that the granivorous water birds disperse viable seeds of wetland plants over long distance during their migration. The censuses

of large flocks of such migratory granivorous wintering or breeding ducks were recorded during this study and previous ones (Perennou 1990; Davidar 2011; Mathevet et al. 2020).

Notes: As per the revision of Hayasaka (2012) and Xanthos & Browning (2015) the subspecies 'raynalii' could be easily distinguished by having 3-fid stigmas, planoconvex nature of nutlet with distinct rugose surface, while its typical subspecies 'erecta', has 2-fid stigma, biconvex nutlet with moderate wrinkled surface. The surface of *S. lateriflora* nutlet is otherwise same that of *S. erecta* ssp. *raynalii* but the former species has trigonous nutlet with 2-fid stigmas.

In *Schoenoplectiella*, interspecific natural hybridization exhibits range of variations in plant height, culm width below the inflorescence, shape and length of overtopping inflorescence, glume dimensions at apex, anther crest length, style branch and length, achene dimensions and surface sculpturing were very well studied (Browning 1992; Hayasaka 2012). Meanwhile the variation in the shape of inflorescence was overlooked by previous taxonomic accounts (Ohwi 1944; Koyama 1958). Though, in India we observed that the length of primary and secondary peduncles of the inflorescence is much longer than the (iso)type specimen from KEW image and specimen from the University of South Florida Herbarium (20709).

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POWO (2022). https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:77107810-1. Accessed on 7 March 2022.

Smith, S.G. (2003). Cyperaceae. In: Ball, P.W., A.A. Reznick & D.F. Murray (ed.). Flora of North America Vol. 23; http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=10246. Accessed on 9 March 92022.

Verloove, F., A. Mesterhazy & J. Browning (2016). Studies in *Schoenoplectiella* (Cyperaceae) in tropical West Africa. *Phytotaxa* 283(1): 096–100.

Xanthos, M. & J. Browning (2015). Taxonomic re-evaluation of *Schoenoplectiella lateriflora* subsp. *laevinux* (Cyperacee) and a new record for *Schoenoplectiella erecta* subsp. *erecta*. *Kew Bulletin* 70: 36–41.

https://www.gbif.org/occurrence/1503282087. Accessed on 24 February 2022.

https://www.catalogueoflife.org/data/taxon/5L3DV. Accessed on 24 February 2022.

https://plants.usda.gov/home/plantProfile?symbol=SCERR2. Accessed on 9 March 2022.

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Dr. George Mathew, Kerala Forest Research Institute, Peechi, India

Dr. John Noyes, Natural History Museum, London, UK

Dr. Albert G. Orr, Griffith University, Nathan, Australia

Dr. Sameer Padhye, Katholieke Universiteit Leuven, Belgium

Dr. Nancy van der Poorten, Toronto, Canada

Dr. Kareen Schnabel, NIWA, Wellington, New Zealand

Dr. R.M. Sharma, (Retd.) Scientist, Zoological Survey of India, Pune, India

Dr. Manuj Siliwal, WILD, Coimbatore, Tamil Nadu, India

Dr. G.P. Sinha, Botanical Survey of India, Allahabad, India

Dr. K.A. Subramanian, Zoological Survey of India, New Alipore, Kolkata, India

Dr. P.M. Sureshan, Zoological Survey of India, Kozhikode, Kerala, India

Dr. R. Varatharajan, Manipur University, Imphal, Manipur, India

Dr. Eduard Vives, Museu de Ciències Naturals de Barcelona, Terrassa, Spain

Dr. James Young, Hong Kong Lepidopterists' Society, Hong Kong

Dr. R. Sundararaj, Institute of Wood Science & Technology, Bengaluru, India

Dr. M. Nithyanandan, Environmental Department, La Ala Al Kuwait Real Estate. Co. K.S.C., Kuwait

Dr. Himender Bharti, Punjabi University, Punjab, India

Mr. Purnendu Roy, London, UK

Dr. Saito Motoki, The Butterfly Society of Japan, Tokyo, Japan

Dr. Sanjay Sondhi, TITLI TRUST, Kalpavriksh, Dehradun, India

Dr. Nguyen Thi Phuong Lien, Vietnam Academy of Science and Technology, Hanoi, Vietnam

Mr. Nitin Kulkarni, Tropical Research Institute, Jabalpur, India

Dr. Robin Wen Jiang Ngiam, National Parks Board, Singapore

Dr. Lional Monod, Natural History Museum of Geneva, Genève, Switzerland.

Dr. Asheesh Shivam, Nehru Gram Bharti University, Allahabad, India

Dr. Rosana Moreira da Rocha, Universidade Federal do Paraná, Curitiba, Brasil

Dr. Kurt R. Arnold, North Dakota State University, Saxony, Germany

Dr. James M. Carpenter, American Museum of Natural History, New York, USA

Dr. David M. Claborn, Missouri State University, Springfield, USA

Dr. Kareen Schnabel, Marine Biologist, Wellington, New Zealand

Dr. Amazonas Chagas Júnior, Universidade Federal de Mato Grosso, Cuiabá, Brasil

Mr. Sonoon Jyoti Gogoi, Assam University, Silchar, Assam, India

Dr. Heo Chong Chin, Universiti Teknologi MARA (UiTM), Selangor, Malaysia

Dr. R.J. Shiel, University of Adelaide, SA 5005, Australia

Dr. Siddharth Kulkarni, The George Washington University, Washington, USA

Dr. Priyadarsanan Dharma Rajan, ATREE, Bengaluru, India

Dr. Phil Alderslade, CSIRO Marine and Atmospheric Research, Hobart, Australia

Dr. John E.N. Veron, Coral Reef Research, Townsville, Australia

Dr. Daniel Whitmore, State Museum of Natural History Stuttgart, Rosenstein, Germany.

Dr. Yu-Feng Hsu, National Taiwan Normal University, Taipei City, Taiwan

Dr. Keith V. Wolfe, Antioch, California, USA

Dr. Siddharth Kulkarni, The Hormiga Lab, The George Washington University, Washington, D.C., USA

Dr. Tomas Ditrich, Faculty of Education, University of South Bohemia in Ceske Budejovice, Czech Republic

Dr. Mihaly Foldvari, Natural History Museum, University of Oslo, Norway

Dr. V.P. Uniyal, Wildlife Institute of India, Dehradun, Uttarakhand 248001, India

Dr. John T.D. Caleb, Zoological Survey of India, Kolkata, West Bengal, India

Dr. Priyadarsanan Dharma Rajan, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Bangalore, Karnataka, India

## Fishes

Dr. Neelesh Dahanukar, IISER, Pune, Maharashtra, India

Dr. Topiltzin Contreras MacBeath, Universidad Autónoma del estado de Morelos, México

Dr. Heok Hee Ng, National University of Singapore, Science Drive, Singapore

Dr. Rajeev Raghavan, St. Albert's College, Kochi, Kerala, India

Dr. Robert D. Sluka, Chiltern Gateway Project, A Rocha UK, Southall, Middlesex, UK

Dr. E. Vivekanandan, Central Marine Fisheries Research Institute, Chennai, India

Dr. Davor Zanella, University of Zagreb, Zagreb, Croatia

Dr. A. Biju Kumar, University of Kerala, Thiruvananthapuram, Kerala, India

Dr. Akhilesh K.V., ICAR-Central Marine Fisheries Research Institute, Mumbai Research

Centre, Mumbai, Maharashtra, India

Dr. J.A. Johnson, Wildlife Institute of India, Dehradun, Uttarakhand, India

Dr. R. Ravinesh, Gujarat Institute of Desert Ecology, Gujarat, India

## Amphibians

Dr. Sushil K. Dutta, Indian Institute of Science, Bengaluru, Karnataka, India

Dr. Annemarie Ohler, Muséum national d'Histoire naturelle, Paris, France

## Reptiles

Dr. Gernot Vogel, Heidelberg, Germany

Dr. Raju Vyas, Vadodara, Gujarat, India

Dr. Pritpal S. Soorae, Environment Agency, Abu Dubai, UAE.

Prof. Dr. Wayne J. Fuller, Near East University, Mersin, Turkey

Prof. Chandrashekher U. Rivonker, Goa University, Taleigao Plateau, Goa, India

Dr. S.R. Ganesh, Chennai Snake Park, Chennai, Tamil Nadu, India

Dr. Himansu Sekhar Das, Terrestrial & Marine Biodiversity, Abu Dhabi, UAE

## Birds

Dr. Hem Sagar Baral, Charles Sturt University, NSW Australia

Mr. H. Byju, Coimbatore, Tamil Nadu, India

Dr. Chris Bowden, Royal Society for the Protection of Birds, Sand UK

Dr. Priya Davidar, Pondicherry University, Kalapet, Puducherry, India

Dr. J.W. Duckworth, IUCN SSC, Bath, UK

Dr. Rajah Jayapal, SACON, Coimbatore, Tamil Nadu, India

Dr. Rajiv S. Kalsi, M.L.N. College, Yamuna Nagar, Haryana, India

Dr. V. Santharam, Rishi Valley Education Centre, Chittoor Dt., Andhra Pradesh, India

Dr. S. Balachandran, Bombay Natural History Society, Mumbai, India

Mr. J. Praveen, Bengaluru, India

Dr. C. Srinivasulu, Osmania University, Hyderabad, India

Dr. K.S. Gopi Sundar, International Crane Foundation, Baraboo, USA

Dr. Gombobaatar Sundev, Professor of Ornithology, Ulaanbaatar, Mongolia

Prof. Reuven Yosef, International Birding & Research Centre, Eilat, Israel

Dr. Taej Mundkur, Wetlands International, Wageningen, The Netherlands

Dr. Carol Inskipp, Bishop Auckland Co., Durham, UK

Dr. Tim Inskipp, Bishop Auckland Co., Durham, UK

Dr. V. Gokula, National College, Tiruchirappalli, Tamil Nadu, India

Dr. Arkady Lelej, Russian Academy of Sciences, Vladivostok, Russia

Dr. Simon Dowell, Science Director, Chester Zoo, UK

Dr. Mário Gabriel Santiago dos Santos, Universidade de Trás-os-Montes e Alto Douro,

Quinta de Prados, Vila Real, Portugal

Dr. Grant Connette, Smithsonian Institution, Royal, VA, USA

Dr. M. Zafar-ul Islam, Prince Saud Al Faisal Wildlife Research Center, Taif, Saudi Arabia

## Mammals

Dr. Giovanni Amori, CNR - Institute of Ecosystem Studies, Rome, Italy

Dr. Anwaruddin Chowdhury, Guwahati, India

Dr. David Mallon, Zoological Society of London, UK

Dr. Shomita Mukherjee, SACON, Coimbatore, Tamil Nadu, India

Dr. Angie Appel, Wild Cat Network, Germany

Dr. P.O. Nameer, Kerala Agricultural University, Thrissur, Kerala, India

Dr. Ian Redmond, UNEP Convention on Migratory Species, Lansdown, UK

Dr. Heidi S. Riddle, Riddle's Elephant and Wildlife Sanctuary, Arkansas, USA

Dr. Karin Schwartz, George Mason University, Fairfax, Virginia

Dr. Lala A.K. Singh, Bhubaneswar, Orissa, India

Dr. Mewa Singh, Mysore University, Mysore, India

Dr. Paul Racey, University of Exeter, Devon, UK

Dr. Honnavalli N. Kumara, SACON, Anaikatty P.O., Coimbatore, Tamil Nadu, India

Dr. Nishith Dhariaya, HNG University, Patan, Gujarat, India

Dr. Spartaco Gippoliti, Socio Onorario Società Italiana per la Storia della Fauna "Giuseppe Altobello", Rome, Italy

Dr. Justus Joshua, Green Future Foundation, Tiruchirapalli, Tamil Nadu, India

Dr. H. Raghuram, The American College, Madurai, Tamil Nadu, India

Dr. Paul Bates, Harison Institute, Kent, UK

Dr. Jim Sanderson, Small Wild Cat Conservation Foundation, Hartford, USA

Dr. Dan Challender, University of Kent, Canterbury, UK

Dr. David Mallon, Manchester Metropolitan University, Derbyshire, UK

Dr. Brian L. Cypher, California State University-Stanislaus, Bakersfield, CA

Dr. S.S. Talmale, Zoological Survey of India, Pune, Maharashtra, India

Prof. Karan Bahadur Shah, Budhanilakantha Municipality, Kathmandu, Nepal

Dr. Susan Cheyne, Borneo Nature Foundation International, Palangkaraja, Indonesia

Dr. Hemanta Kafley, Wildlife Sciences, Tarleton State University, Texas, USA

## Other Disciplines

Dr. Aniruddha Belsare, Columbia MO 65203, USA (Veterinary)

Dr. Mandar S. Paingankar, University of Pune, Pune, Maharashtra, India (Molecular)

Dr. Jack Tordoff, Critical Ecosystem Partnership Fund, Arlington, USA (Communities)

Dr. Ulrike Streicher, University of Oregon, Eugene, USA (Veterinary)

Dr. Hari Balasubramanian, EcoAdvisors, Nova Scotia, Canada (Communities)

Dr. Rayanna Hellem Santos Bezerra, Universidade Federal de Sergipe, São Cristóvão, Brazil

Dr. Jamie R. Wood, Landcare Research, Canterbury, New Zealand

Dr. Wendy Collinson-Jonker, Endangered Wildlife Trust, Gauteng, South Africa

Dr. Rajeshkumar G. Jani, Anand Agricultural University, Anand, Gujarat, India

Dr. O.N. Tiwari, Senior Scientist, ICAR-Indian Agricultural Research Institute (IARI), New Delhi, India

Dr. L.D. Singla, Guru Angad Dev Veterinary and Animal Sciences University, Ludhiana, India

Dr. Rupika S. Rajakaruna, University of Peradeniya, Peradeniya, Sri Lanka

Dr. Bahar Baviskar, Wild-CER, Nagpur, Maharashtra 440013, India

## Reviewers 2019–2021

Due to paucity of space, the list of reviewers for 2018–2020 is available online.

The opinions expressed by the authors do not reflect the views of the Journal of Threatened Taxa, Wildlife Information Liaison Development Society, Zoo Outreach Organization, or any of the partners. The journal, the publisher, the host, and the partners are not responsible for the accuracy of the political boundaries shown in the maps by the authors.

**Journal of Threatened Taxa** is indexed/abstracted in Bibliography of Systematic Mycology, Biological Abstracts, BIOSIS Previews, CAB Abstracts, EBSCO, Google Scholar, Index Copernicus, Index Fungorum, JournalSeek, National Academy of Agricultural Sciences, NewJour, OCLC WorldCat, SCOPUS, Stanford University Libraries, Virtual Library of Biology, Zoological Records.

NAAS rating (India) 5.64

Print copies of the Journal are available at cost. Write to:

The Managing Editor, JoTT,

c/o Wildlife Information Liaison Development Society,

43/2 Varadarajulu Nagar, 5th Street West, Ganapathy, Coimbatore,

Tamil Nadu 641006, India

ravi@threatenedtaxa.org
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www.threatenedtaxa.org

OPEN ACCESS

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The Journal of Threatened Taxa (JoTT) is dedicated to building evidence for conservation globally by publishing peer-reviewed articles online every month at a reasonably rapid rate at www.threatenedtaxa.org. All articles published in JoTT are registered under Creative Commons Attribution 4.0 International License unless otherwise mentioned. JoTT allows allows unrestricted use, reproduction, and distribution of articles in any medium by providing adequate credit to the author(s) and the source of publication.

ISSN 0974-7907 (Online) | ISSN 0974-7893 (Print)

February 2023 | Vol. 15 | No. 2 | Pages: 22559–22770

Date of Publication: 26 February 2023 (Online & Print)

DOI: 10.11609/jott.2023.15.2.22559-22770

## Communications

**Sunda Clouded Leopard Neofelis diardi** (Cuvier, 1823) (Mammalia: Carnivora: Felidae) occupancy in Borneo: results of a pilot vehicle spotlight transect survey
– Jephte Sompud, Sze Lue Kee, Kurtis Jai-Chyi Pei, Paul Liau, Collin Goh & Anthony J. Giordano, Pp. 22559–22566

**On the occurrence of Eurasian Otter Lutra lutra** (Carnivora: Mustelidae) in Neeru stream of Chenab catchment, Jammu & Kashmir, India
– Dinesh Singh, Anil Thakar & Neeraj Sharma, Pp. 22567–22573

**Distribution of avifauna on twenty-one islands of the Gulf of Mannar Biosphere Reserve, India**
– H. Byju, N. Raveendran & S. Ravichandran, Pp. 22574–22585

**Habitats of House Sparrow Passer domesticus** (Linnaeus, 1758) in Rameswaram Island, Tamil Nadu, India
– M. Pandian, Pp. 22586–22596

**Seasonal diversity and dietary guild structure of birds in two Vindhyan gorge forests of Rajasthan, India**
– Ashvini Kumar Joshi, Pp. 22597–22605

**Differential kleptoparasitic interactions of Himalayan Vulture Gyps himalayensis with conspecifics and heterospecifics during various stages of breeding**
– Hameem Mushtaq Wani, Pp. 22606–22610

**Range extension of Isthmoheros tuyrensis**, a threatened species of fish (Cichlidae) in Panama: including new ecological and morphological data
– Arturo Dominici-Arosemena, Arturo Angulo, Haydee Osorio-Ugarte, Quiriatjaryn Ortega-Samaniego, Andrés Fraiz, Arminda Guerrel, Edgar Araúz, Jennyfer Montiel, Beatriz Medina, Yehudi Rodríguez-Arriatti, Yessenia González, Javier Pardo, Karly Urriola & Adrián Ramos-Merchante, Pp. 22611–22622

**Tadpole morphology of Jerdon's Narrow-mouthed Frog Uperodon montanus** (Jerdon, 1853) with a range and elevation extension report from Western Ghats, India
– Amit Hegde, Girish Kadadevaru & K.P. Dinesh, Pp. 22623–22631

**An annotated checklist of the economically important family of moths (Lepidoptera: Heterocera: Noctuidae) of the northern Western Ghats, India, with notes on their type species, diversity, distribution, host plants, and an unusual new faunistic record**
– Aparna Sureshchandra Kalawate, Prachee Surwade & S.N. Pawara, Pp. 22632–22653

**Report of a tussock moth genus Maeproctis** (Lepidoptera: Erebidae: Lymantriinae: Nygmiini) from India
– Gagan Preet Kour Bali & Amritpal Singh Kaleka, Pp. 22654–22660

**Butterflies of Silent Valley National Park and its environs, Western Ghats of Kerala, India**
– Kalesh Sadasivan, P.C. Sujitha, Toms Augustine, Edayillam Kunhikrishnan, Vinayan P. Nair, M. Divin Murukesh & Baiju Kochunarayanan, Pp. 22661–22676

**Notes on morphology and bionomics of Urolabida histrionica** (Westwood) (Heteroptera: Uro stylididae) from Assam, India
– Sachin Ranade & Hemant V. Ghate, Pp. 22677–22685

**Andromonoecy functional through heterostyly and large carpenter bees as principal pollinators in Solanum carolinense** L. (Solanaceae)
– Suvarna Raju Palathoti & Aluri Jacob Solomon Raju, Pp. 22686–22694

**An inventory of endemic and near endemic angiosperm flora of Biligiri Rangaswamy Temple Tiger Reserve, peninsular India**
– J. Jayanthi, Pp. 22695–22717

**Multidimensional time-lapse of a relict species Canarium strictum** Roxb. from a sacred landscape in Pune District, India
– Mukul Mahabaleshwarkar, Nivedita Ghayal, Supriya Mahabaleshwarkar & Vinaya Ghate, Pp. 22718–22725

**Rediscovery of Sewardiella tuberifera** Kash., a long-lost monotypic endemic Indian liverwort
– Sapana Pant, S.D. Tewari, Prachi Joshi, Manisha Bhandari & Richa Arya, Pp. 22726–22730

**Physcomitrium eurystomum** Sendtn. (Funariaceae: Bryophyta) and **Splashnobryum obtusum** (Brid.) Müll. Hal. (Splashnobryaceae: Bryophyta), two rare moss species from the Western Ghats of Kerala
– C. Nair Manju, P.M. Vineesha, B. Mufeed & K.P. Rajesh, Pp. 22731–22736

## Short Communications

**First record of the Great Seahorse Hippocampus kelloggi** Jordan & Snyder, 1901 (Actinopterygii: Syngnathiformes: Syngnathidae) from the northwestern coast of Bay of Bengal
– Anil Kumar Behera, Biswajit Mahari & Amrit Kumar Mishra, Pp. 22737–22740

**Schoenoplectiella erecta** (Poir.) Lye ssp. *raynallii* (Schuyler) Beentje (Cyperaceae) – a new record to India from Ossudu Bird Sanctuary, Villupuram District, Tamil Nadu
– Chandrasegrane Pradeep, Paneerselvam Umamaheswari, Natesan Balachandran & Raphael Mathevet, Pp. 22741–22745

## Notes

**Status of the Sumatran Striped Rabbit Nesolagus netscheri** in Isau-Isau Wildlife Reserve, South Sumatra Province, Indonesia
– Arum Setiawan, Muhammad Iqbal, Octavia Susilowati, Doni Setiawan, Martialis Puspito Khristy Maharsi & Indra Yustian, Pp. 22746–22748

**Photographic record of the butterfly ray Gymnura cf. poecilura** (Myliobatiformes: Gymnuridae) from the Bhagirathi-Hooghly River in West Bengal, eastern India
– Priyankar Chakraborty, Pp. 22749–22751

**First report of the fairyfly Schizophragma mitai** Triapitsyn (Hymenoptera: Mymaridae) from India with notes on *S. indica* Rehmat & Anis
– Anandhan Rameshkumar, Nazurius Anand, Sayan Sardar & Sarfrazul Islam Kazmi, Pp. 22752–22756

**Occurrence of Ranunculus sceleratus** L. (Ranunculaceae) from the Nilgiri District, Tamil Nadu, India
– J. Shashikanth, S.Mugendhiran & Digvijay Verma, Pp. 22757–22760

**First report of Meliola panici** on *Ottochloa nodosa* (Kunth) Dandy (Poaceae)
– Gopinathan Nair Gokul & Jacob Thomas, Pp. 22761–22763

**New record of an usneoied lichen Usnea hirta** (L.) Weber ex F.H.Wigg. from India
– K.S. Vinayaka, Archana R. Mesta & N. Rajeshwari, Pp. 22764–22766

**On the occurrence of two species of rare cyanobacterial genus Petalonema** M.J.Berkeley ex Wolle, 1887 (Cyanophyceae: Nostocales: Scytonemataceae) from eastern Himalaya, India
– Jai Prakash Keshri, Narendra Nath Koley & Jay Mal, Pp. 22767–22770

## Publisher & Host

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Threatened Taxa