Type: COLOMBIA: Río Magdalena, s.d., *J. Weir 26* (holotype BM [barcode BM001008877]).

**Notes:** In the protologue of *Schoenleinia thyrsoidea* Miers, Miers (1864: 186) cited the material studied as "Rio Magdalena. - Weir 26." and stated that "this plant approaches nearest to the genus *Schoenlinia* of Klotzsch, the type of which is *Exostemma cuspidata* St. Hil.; I find no species among the *Cinchona* group hitherto described, to which it can be referred." At BM there is a specimen, with barcode BM001008877, and the label "HERB. JOHN MIERS. Bequeathed 1879." On the same sheet are affixed three additional labels, with blue paper. One small label has the annotation "Schoenlinia thyrsoidea nob." handwritten by Miers. On the second label with blue paper is handwritten "Rio Magdalena, N. Grenada, Weir 26." On a third label with blue paper is present a long species description, handwritten by Miers, corresponding to that of his publication. This specimen is the holotype of this name.

29. **IXORA** L., Sp. Pl. 110. 1753.

FGT, vol. 40(2), p. 584:
"29-1. *Ixora araguaiensis* Delprete, J. Bot. Res. Inst. Texas 2(1): 456, fig. 2. 2008. Tipo: Brasil, Tocantins, Mun. Pium, Ilha do Bananal, Parque Nacional do Araguaia, Posto de fiscalização do Projeto Quelônios da Amazônia, 09°50'57"S, 50°11'31"W, 190 m, 26/III/1999 (fl), *M.A. da Silva, R.C. Mendonça, E. Cardoso, A.D. dos Santos, N.G. Sousa, N.R. Oliveira & J.T. dos Santos* 4164 (holótipo, IBGE; isótipo, NY)."

**Accepted name:** *Ixora araguaiensis* Delprete

Type: BRAZIL. Tocantins: Mun. Pium, Ilha do Bananal, Parque Nacional do Araguaia, Posto de fiscalização do Projeto Quelônios da Amazônia, 9°50'57"S, 50°11'31"W, 190 m, 26 Mar. 1999 (fl), *M.A. da Silva, R.C. Mendonça, E. Cardoso, A.D. dos Santos, N.G. Sousa, N.R. Oliveira & J.T. dos Santos* 4164 (holotype, IBGE [Acc. No. 046569]; isotype, NY [barcode 01085900]).

FGT, vol. 40(2), p. 589:
"29-2. *Ixora brevifolia* Benth., Linnaea 23: 448. 1850. Tipo: Brasil, Minas Gerais, sem localidade, 1845-1846 (fl), *Widgren 1119* (lectótipo, UPS V-134607, selecionado por Delprete, 2003; isolectótipo UPS V-102640)."

**Accepted name:** *Ixora brevifolia* Benth.

Type: BRAZIL. Minas Gerais: Without locality, "skogstrakter" [forest tracts], 1845-1846, *J.F. Widgren 1119* (UPS V-134607, lectotype designated by Delprete (2003: 1472); isolectotype UPS V-102640; isolectotype fragment NY [barcode 00688209; ex UPS V-134607]).

**Notes:** For synonyms and types see Delprete (2003).

FGT, vol. 40(2), p. 593:
"29-3. *Ixora casei* Hance in Walp., Ann. Bot. Syst. 2: 754. 1852. - Tipo: Ilhas Carolinas ("Strong Island"), s.d., *Q.W. Case s.n.* (holótipo, K)."

**Accepted name:** *Ixora casei* Hance in Walp.

Type: CAROLINE ISLANDS: Strong Island, s.d., *Q.W. Case s.n.* (holotype, K [barcode K000763383]).

For synonyms see Delprete (2010b: 593).

**Notes:** Hance (in Walpers 1852: 754) in the protologue of *Ixora casei* Hance cited the collection locality of the material studied as "in insulis Carolis ad "Strong's Island." In K, there are two specimens reporting that locality associated with this name.

The K specimen with barcode K000763379 has the annotations "Type specimen of Bot. Mag. t. 685" and "Ixora macrothyrsa J. & B.) I. duffii Veitch! See Fl. & Pom. 1878, p. 76 with woodcut) Ualan on Strong Island, Caroline Group, Kew Gardens Sept. 18, 1884" handwritten directly on the sheet. A third annotation says "Not I. macrothyrsa of Malaysia! See Bremekamp in Bull. Jard. Bot. Buit. sér. 3, 14, 278 (1937)" handwritten by Bremekamp directly on the sheet. The specimen consists of three loose leaves and two inflorescences. This specimen is not original material, because it was gathered from a plant cultivated in the Kew Gardens in 1884, reproduced from material collected in Strong's Island.

The K specimen with barcode K000763383 has the annotation "Ixora casei Hance in Walpers Ann. Vol. II, p. 754. Strong Island (Caroline Islands) An S. N. Case" handwritten in pencil directly on the sheet. This specimen is the holotype of *I. casei*.

FGT, vol. 40(2), p. 595:
"29-4. *Ixora chinensis* Lam., Encycl. Meth. 3: 344. 1789. - Tipo: China, "communicated by Sonnerat" (holótipo, P-Lam.)."
Type: COLOMBIA: Río Magdalena, s.d., *J. Weir 26* (holotype BM [barcode BM001008877]).

**Notes:** In the protologue of *Schoenleinia thyrsoidea* Miers, Miers (1864: 186) cited the material studied as "Rio Magdalena. - Weir 26." and stated that "this plant approaches nearest to the genus *Schoenlinia* of Klotzsch, the type of which is *Exostemma cuspidata* St. Hil.; I find no species among the *Cinchona* group hitherto described, to which it can be referred." At BM there is a specimen, with barcode BM001008877, and the label "HERB. JOHN MIERS. Bequeathed 1879." On the same sheet are affixed three additional labels, with blue paper. One small label has the annotation "Schoenlinia thyrsoidea nob." handwritten by Miers. On the second label with blue paper is handwritten "Rio Magdalena, N. Grenada, Weir 26." On a third label with blue paper is present a long species description, handwritten by Miers, corresponding to that of his publication. This specimen is the holotype of this name.

29. **IXORA** L., Sp. Pl. 110. 1753.

FGT, vol. 40(2), p. 584:
"29-1. *Ixora araguaiensis* Delprete, J. Bot. Res. Inst. Texas 2(1): 456, fig. 2. 2008. Tipo: Brasil, Tocantins, Mun. Pium, Ilha do Bananal, Parque Nacional do Araguaia, Posto de fiscalização do Projeto Quelônios da Amazônia, 09°50'57"S, 50°11'31"W, 190 m, 26/III/1999 (fl), *M.A. da Silva, R.C. Mendonça, E. Cardoso, A.D. dos Santos, N.G. Sousa, N.R. Oliveira & J.T. dos Santos* 4164 (holótipo, IBGE; isótipo, NY)."

**Accepted name:** *Ixora araguaiensis* Delprete

Type: BRAZIL. Tocantins: Mun. Pium, Ilha do Bananal, Parque Nacional do Araguaia, Posto de fiscalização do Projeto Quelônios da Amazônia, 9°50'57"S, 50°11'31"W, 190 m, 26 Mar. 1999 (fl), *M.A. da Silva, R.C. Mendonça, E. Cardoso, A.D. dos Santos, N.G. Sousa, N.R. Oliveira & J.T. dos Santos* 4164 (holotype, IBGE [Acc. No. 046569]; isotype, NY [barcode 01085900]).

FGT, vol. 40(2), p. 589:
"29-2. *Ixora brevifolia* Benth., Linnaea 23: 448. 1850. Tipo: Brasil, Minas Gerais, sem localidade, 1845-1846 (fl), *Widgren 1119* (lectótipo, UPS V-134607, selecionado por Delprete, 2003; isolectótipo UPS V-102640)."

**Accepted name:** *Ixora brevifolia* Benth.

Type: BRAZIL. Minas Gerais: Without locality, "skogstrakter" [forest tracts], 1845-1846, *J.F. Widgren 1119* (UPS V-134607, lectotype designated by Delprete (2003: 1472); isolectotype UPS V-102640; isolectotype fragment NY [barcode 00688209; ex UPS V-134607]).

**Notes:** For synonyms and types see Delprete (2003).

FGT, vol. 40(2), p. 593:
"29-3. *Ixora casei* Hance in Walp., Ann. Bot. Syst. 2: 754. 1852. - Tipo: Ilhas Carolinas ("Strong Island"), s.d., *Q.W. Case s.n.* (holótipo, K)."

**Accepted name:** *Ixora casei* Hance in Walp.

Type: CAROLINE ISLANDS: Strong Island, s.d., *Q.W. Case s.n.* (holotype, K [barcode K000763383]).

For synonyms see Delprete (2010b: 593).

**Notes:** Hance (in Walpers 1852: 754) in the protologue of *Ixora casei* Hance cited the collection locality of the material studied as "in insulis Carolis ad "Strong's Island." In K, there are two specimens reporting that locality associated with this name.

The K specimen with barcode K000763379 has the annotations "Type specimen of Bot. Mag. t. 685" and "Ixora macrothyrsa J. & B.) I. duffii Veitch! See Fl. & Pom. 1878, p. 76 with woodcut) Ualan on Strong Island, Caroline Group, Kew Gardens Sept. 18, 1884" handwritten directly on the sheet. A third annotation says "Not I. macrothyrsa of Malaysia! See Bremekamp in Bull. Jard. Bot. Buit. sér. 3, 14, 278 (1937)" handwritten by Bremekamp directly on the sheet. The specimen consists of three loose leaves and two inflorescences. This specimen is not original material, because it was gathered from a plant cultivated in the Kew Gardens in 1884, reproduced from material collected in Strong's Island.

The K specimen with barcode K000763383 has the annotation "Ixora casei Hance in Walpers Ann. Vol. II, p. 754. Strong Island (Caroline Islands) An S. N. Case" handwritten in pencil directly on the sheet. This specimen is the holotype of *I. casei*.

FGT, vol. 40(2), p. 595:
"29-4. *Ixora chinensis* Lam., Encycl. Meth. 3: 344. 1789.
- Tipo: China, "communicated by Sonnerat" (holótipo, P-Lam.)."
**Accepted name:** *Ixora chinensis* Lam.

Type: CHINA. Without locality, s.d., P. Sonnerat s.n. (first-step lectotype designated by Fosberg & Sachet (1989a: 77); P-LA [barcode P00308523], **second-step lectotype here designated**).

For synonyms see Delprete (2010b: 595).

**Notes:** In the protologue of *Ixora chinensis* Lam., Lamarck (1789: 344) cited "*Flamma sylvarum peregrina*", Rumph. Amb. 4. p. 107, t. 47." and stated "Je ne vois pas pourquoi Linné a associé le *Flamma sylvarum* de Rumphe avec le *Pavetta* de Rhéede, dont je traiterai après cette espèce. A la verité, ces deux plantes sont du même genre; mais elles sont très-distinctes l'une de l'autre, comme l'examen m'en a convaincu, les possedant toutes deux dans mon herbier." ("I don't see why Linnaeus associated *Flamma sylvarum* of Rumphius with *Pavetta* of Rhéede, which I will treat after this species. To the truth, these two plants belong to the same genus; but they are very distinct from each other, as the examination convinced me, both of them being present in my herbarium). He described the material studied of *I. chinensis* as "Cette espèce nous a été communiqué par M. Sonnerat, comme provenant de la Chine; elle croit aussi dans l'Isle de Java. (v.s.)." ("This plant was communicated by Mr. Sonnerat, as originated from China; it grows also in the Island of Java"). The abbreviation "v.s." (vidi siccum) means that he saw a specimen in his own herbarium. Therefore, original material of this name is a specimen collected by Sonnerat present in the Lamarck herbarium (P-Lam.) in Paris and plate 47 "*Flamma sylvarum peregrina*" of Rumphius's *Herbarium amboinense*, vol. 4 (1743). The geographic region studied in this publication is "Amboina" or "Amboyna", nowadays called Ambon Island, which is part of the Maluku Islands of Indonesia.

On Rumphius's plate 47 is depicted a branch with numerous leaves and an inflorescence with flower buds and flowers in anthesis. The leaves are depicted as short-petiolated, with blades acute at base and obtuse-subacute at apex, and with 6-8 secondary veins on each side of the midrib. The flower buds are depicted as acute at the apex. The corollas are drawn has having a long-narrow tube and short lobes round at the apex. On page 107, Rumphius described the provenance of the plant as "Elegans *Flammae silvarum* species circa annum 1675 in Amboinam fuit delata sub nomine floris Japanensis mirabilis, proquo & per decem annos istum habui, tandem vero a diversis Europaeis, qui diu in ista regione versati fuere, intellexi, hanc arborem sese non in Japana, sed in Java aliisque Malayensibus regionibus vidisse,

unde & hanc tanquam peregrinam species *Flammae silvarum* describam." The description that follows that statement is quite detailed and corresponds to all features depicted on plate 47.

Fosberg and Sachet (1989a: 77) cited the type of *Ixora chinensis* Lam. as "Type: China. "communicated by Sonnerat" (P-Herb. Lam. Not seen by us)." As their type citation was published before 2001, it should be treated as an inadvertent first-step lectotypification, because in P-LA there is a sheet with two different gatherings. On the right side of the sheet are affixed a small branch with one leaf and a terminal inflorescence, and two loose leaves, with barcode number P00308523. On the stem is affixed a label with the annotation "ixora chinensis lam. dict." The corollas have broadly ovate to almost round lobes, obtuse at the apex. This specimen, with barcode number P00308523, is here designated as the second-step lectotype of *Ixora chinensis*.

FGT, vol. 40(2), p. 598:
"29-5. *Ixora coccinea* L., Sp. Pl. 110. 1753; L., Gen. Pl., ed. 5, 48. 1754. - Tipo: "Habitat in Índia" (BM-LINN)."

**Accepted name:** *Ixora coccinea* L.

Type: [icon] "*Schetti*" in Rheede, Hort. Malab. 2: 17, tab. 13. 1679, lectotype designated by Fosberg and Sachet in Taxon 38: 488. 1989.

For synonyms see Delprete (2010b: 598–599).

**Notes:** Fosberg and Sachet (1989b: 486–489) showed that the various potential typifications of *Ixora coccinea* L. were either inadequate or invalid (Bremekamp, 1937: 198; Corner, 1941: 185; respectively). They concluded that, to maintain the traditional usage of the name for the species they designated Table 13 of Rheede's (1679) second volume of *Hortus Indicus Malabaricus* as the lectotype of this name. Jarvis (2007: 598) cited Fosberg and Sachet's discussion and typification, and accepted Rheede's plate as lectotype.

FGT, vol. 40(2), p. 601:
"29-6. *Ixora congestiflora* Delprete, J. Bot. Res. Inst. Texas 2(1): 456, fig. 1. 2008. Tipo: Brasil, Tocantins, Mun. Caseara, Parque Estadual do Cantão, transecto 11, floresta estacional semidecídua, 09°18'00"S, 50°01'57"W, 15/I/2000 (fl), *P.E. Nogueira & M. Richter* 692 (holótipo, IBGE; isótipo, NY)."
**Accepted name:** *Ixora chinensis* Lam.

Type: CHINA. Without locality, s.d., P. Sonnerat s.n. (first-step lectotype designated by Fosberg & Sachet (1989a: 77); P-LA [barcode P00308523], **second-step lectotype here designated**).

For synonyms see Delprete (2010b: 595).

**Notes:** In the protologue of *Ixora chinensis* Lam., Lamarck (1789: 344) cited "*Flamma sylvarum peregrina*", Rumph. Amb. 4. p. 107, t. 47." and stated "Je ne vois pas pourquoi Linné a associé le *Flamma sylvarum* de Rumphe avec le *Pavetta* de Rhéede, dont je traiterai après cette espèce. A la verité, ces deux plantes sont du même genre; mais elles sont très-distinctes l'une de l'autre, comme l'examen m'en a convaincu, les possedant toutes deux dans mon herbier." ("I don't see why Linnaeus associated *Flamma sylvarum* of Rumphius with *Pavetta* of Rhéede, which I will treat after this species. To the truth, these two plants belong to the same genus; but they are very distinct from each other, as the examination convinced me, both of them being present in my herbarium). He described the material studied of *I. chinensis* as "Cette espèce nous a été communiqué par M. Sonnerat, comme provenant de la Chine; elle croit aussi dans l'Isle de Java. (v.s.)." ("This plant was communicated by Mr. Sonnerat, as originated from China; it grows also in the Island of Java"). The abbreviation "v.s." (vidi siccum) means that he saw a specimen in his own herbarium. Therefore, original material of this name is a specimen collected by Sonnerat present in the Lamarck herbarium (P-Lam.) in Paris and plate 47 "*Flamma sylvarum peregrina*" of Rumphius's *Herbarium amboinense*, vol. 4 (1743). The geographic region studied in this publication is "Amboina" or "Amboyna", nowadays called Ambon Island, which is part of the Maluku Islands of Indonesia.

On Rumphius's plate 47 is depicted a branch with numerous leaves and an inflorescence with flower buds and flowers in anthesis. The leaves are depicted as short-petiolated, with blades acute at base and obtuse-subacute at apex, and with 6-8 secondary veins on each side of the midrib. The flower buds are depicted as acute at the apex. The corollas are drawn has having a long-narrow tube and short lobes round at the apex. On page 107, Rumphius described the provenance of the plant as "Elegans *Flammae silvarum* species circa annum 1675 in Amboinam fuit delata sub nomine floris Japanensis mirabilis, proquo & per decem annos istum habui, tandem vero a diversis Europaeis, qui diu in ista regione versati fuere, intellexi, hanc arborem sese non in Japana, sed in Java aliisque Malayensibus regionibus vidisse,

unde & hanc tanquam peregrinam species *Flammae silvarum* describam." The description that follows that statement is quite detailed and corresponds to all features depicted on plate 47.

Fosberg and Sachet (1989a: 77) cited the type of *Ixora chinensis* Lam. as "Type: China. "communicated by Sonnerat" (P-Herb. Lam. Not seen by us)." As their type citation was published before 2001, it should be treated as an inadvertent first-step lectotypification, because in P-LA there is a sheet with two different gatherings. On the right side of the sheet are affixed a small branch with one leaf and a terminal inflorescence, and two loose leaves, with barcode number P00308523. On the stem is affixed a label with the annotation "ixora chinensis lam. dict." The corollas have broadly ovate to almost round lobes, obtuse at the apex. This specimen, with barcode number P00308523, is here designated as the second-step lectotype of *Ixora chinensis*.

FGT, vol. 40(2), p. 598:
"29-5. *Ixora coccinea* L., Sp. Pl. 110. 1753; L., Gen. Pl., ed. 5, 48. 1754. - Tipo: "Habitat in Índia" (BM-LINN)."

**Accepted name:** *Ixora coccinea* L.

Type: [icon] "*Schetti*" in Rheede, Hort. Malab. 2: 17, tab. 13. 1679, lectotype designated by Fosberg and Sachet in Taxon 38: 488. 1989.

For synonyms see Delprete (2010b: 598–599).

**Notes:** Fosberg and Sachet (1989b: 486–489) showed that the various potential typifications of *Ixora coccinea* L. were either inadequate or invalid (Bremekamp, 1937: 198; Corner, 1941: 185; respectively). They concluded that, to maintain the traditional usage of the name for the species they designated Table 13 of Rheede's (1679) second volume of *Hortus Indicus Malabaricus* as the lectotype of this name. Jarvis (2007: 598) cited Fosberg and Sachet's discussion and typification, and accepted Rheede's plate as lectotype.

FGT, vol. 40(2), p. 601:
"29-6. *Ixora congestiflora* Delprete, J. Bot. Res. Inst. Texas 2(1): 456, fig. 1. 2008. Tipo: Brasil, Tocantins, Mun. Caseara, Parque Estadual do Cantão, transecto 11, floresta estacional semidecídua, 09°18'00"S, 50°01'57"W, 15/I/2000 (fl), *P.E. Nogueira & M. Richter* 692 (holótipo, IBGE; isótipo, NY)."
**Accepted name:** *Ixora congestiflora* Delprete

Type: BRAZIL. Tocantins: Mun. Caseara, Parque Estadual do Cantão, transecto 11, floresta estacional semidecídua, 9°18'0"S, 50°1'57"W, 15 Jan. 2000 (fl), P.E. Nogueira & M. Richter 692 (holotype, IBGE [Acc. No. 047624]; isotype, NY [barcode 01085899]).

FGT, vol. 40(2), p. 604:
"29-7. *Ixora finlaysoniana* Wall. ex G. Don, Gen. Hist. 3: 572. 1834. Tipo: Indias Orientais (holótipo, BM)."

Type: "EAST INDIES". Without locality, s.d. [1821-1823], G. Finlayson s.n. (holotype, K [barcode K001123205, Wallich Cat. No. 6166]; isotype, LE [barcode LE00017478]).

**Notes:** George Don (1834: 572) published *Ixora finlaysoniana* Wall. ex G. Don and cited "Wall. cat. no. 6166" and "Native of the East Indies." Delprete (2010b: 604) cited the holotype at BM, but apparently there is no original material associated with this name in that institution. At K there is a specimen, with barcode K001123205, with the handwritten label "6166 Ixora Finlaysoniana Wall. M. Finlayson." This specimen is the holotype of this name.

FGT, vol. 40(2), p. 605:
"29-8. *Ixora irwinii* Delprete, J. Bot. Res. Inst. Texas 2(1): 459, fig. 3. 2008. Tipo: Brasil, Tocantins, 1 km S de Araguaina, Rio das Lontras, 300 m, 15/III/1968 (fr), H.S. Irwin, H. Maxwell & D.C. Wasshausen 21221 (holótipo, UB; isótipo, NY)."

**Accepted name:** *Ixora irwinii* Delprete

Type: BRAZIL. Tocantins: 1 km S de Araguaina, Rio das Lontras, 300 m, 15 Mar. 1968 (fr), H.S. Irwin, H. Maxwell & D.C. Wasshausen 21221 (holotype, UB [barcode UB0040430]; isotypes, NY [barcode 01085901], US [barcode US02508907]).

FGT, vol. 40(2), p. 609:
"29-9. *Ixora pubescens* Willd. in Schult. & Schult. f., Mant. 3: 126. 1818 [sic! 1827]. - *Faramea vaginata* Benth., nom. nud., Linnaea 23: 454. 1850; non Griseb. (1866, nom. superfl.). Tipo: Brasil, Pará [provavelmente perto de Belém], s.d., *F.W. Sieber s.n.* (holótipo B-W 2807 [*dedit Hoffmannsegg*])."

**Accepted name:** *Ixora pubescens* Willd. in Schult. & Schult. f., Mant. 3: 126. 1827.

Type: BRAZIL. Pará: [probably Belém], s.d., F.W. Sieber in J.C. Hoffmannsegg s.n. (holotype B-W [barcode B-W 02807 -01 0]; isotype frag. G [barcode G00634186]).

**Notes:** Joseph A. Schultes and Julius H. Schultes (1827: 126) published *Ixora pubescens* Willd. in Schult. & Schult. f. (Art. 46.3, Ex. 15) and cited the material studied as "*In Para Brasiliae*, com. a *Hoffmannsegg*." That material was collected by Sieber, and Hoffmannsegg never set foot in Brazil. In B-W there is a specimen, with barcode B-W 02807 -01 0, with the handwritten label "arbuste rbr. gull. (Sieber)" affixed on the bottom right corner of the sheet. The annotation "Ix. pubescens" is handwritten directly on the upper right corner of that sheet. This is the specimen of *Ixora pubescens* studied by Willdenow and is the holotype.

In G there is a sheet, barcode G00634186, with an envelope containing one leaf and the fragment of an inflorescence. On the envelope is handwritten "Ixora pubescens Willd., Pará: Hoffmannsegg, fol. numis & flor." This is an isotype fragment of this name.

FGT, vol. 40(2), p. 609:

**Synonym:**

"*Ixora pubescens* var. *glabrifolia* Muell. Arg. in Mart., Fl. Bras. 6(5): 62. 1881. Tipo: Brasil, Pará, "in vicinibus Pará" [agora Belém], *Spruce 328* (lectótipo G, aqui seleccionado, isolectótipos G, K, foto-G [lectótipo] em US, foto-G [isolectótipo] em NY, foto-K em NY)."

Type: BRAZIL. Pará: "in vicinibus Pará" [near Belém], s.d. [Jul.-Aug. 1849], R. *Spruce 328* (first-step lectotype designated by Delprete (2010b: 609); G [barcode G00634184], **second-step lectotype here designated**; isolectotypes, FI [barcode FI004824], G [barcode G00634185], K [barcode K000424499], M [barcode M-0187733]; photo-G [F neg. No. 25677] in NY; photo-K in NY).

**Notes:** Müller Argoviensis (1881: 62) cited the gatherings *Spruce 328*, *Burchell 9456*, and *Burchell 9270* for *Ixora pubescens* var. *glabrifolia* Müll. Arg., without citing the herbarium of deposit. In G, there are two specimens of *Spruce 328*. Therefore, Delprete's (2010b: 609) statement "lectótipo G, aqui seleccionado" should be treated as first-step lectotypification. The specimen with the annotation "I. pubescens β glabrifolia, scripsit Müll.-
**Accepted name:** *Ixora congestiflora* Delprete

Type: BRAZIL. Tocantins: Mun. Caseara, Parque Estadual do Cantão, transecto 11, floresta estacional semidecídua, 9°18'0"S, 50°1'57"W, 15 Jan. 2000 (fl), P.E. Nogueira & M. Richter 692 (holotype, IBGE [Acc. No. 047624]; isotype, NY [barcode 01085899]).

FGT, vol. 40(2), p. 604:
"29-7. *Ixora finlaysoniana* Wall. ex G. Don, Gen. Hist. 3: 572. 1834. Tipo: Indias Orientais (holótipo, BM)."

Type: "EAST INDIES". Without locality, s.d. [1821-1823], G. Finlayson s.n. (holotype, K [barcode K001123205, Wallich Cat. No. 6166]; isotype, LE [barcode LE00017478]).

**Notes:** George Don (1834: 572) published *Ixora finlaysoniana* Wall. ex G. Don and cited "Wall. cat. no. 6166" and "Native of the East Indies." Delprete (2010b: 604) cited the holotype at BM, but apparently there is no original material associated with this name in that institution. At K there is a specimen, with barcode K001123205, with the handwritten label "6166 Ixora Finlaysoniana Wall. M. Finlayson." This specimen is the holotype of this name.

FGT, vol. 40(2), p. 605:
"29-8. *Ixora irwinii* Delprete, J. Bot. Res. Inst. Texas 2(1): 459, fig. 3. 2008. Tipo: Brasil, Tocantins, 1 km S de Araguaina, Rio das Lontras, 300 m, 15/III/1968 (fr), H.S. Irwin, H. Maxwell & D.C. Wasshausen 21221 (holótipo, UB; isótipo, NY)."

**Accepted name:** *Ixora irwinii* Delprete

Type: BRAZIL. Tocantins: 1 km S de Araguaina, Rio das Lontras, 300 m, 15 Mar. 1968 (fr), H.S. Irwin, H. Maxwell & D.C. Wasshausen 21221 (holotype, UB [barcode UB0040430]; isotypes, NY [barcode 01085901], US [barcode US02508907]).

FGT, vol. 40(2), p. 609:
"29-9. *Ixora pubescens* Willd. in Schult. & Schult. f., Mant. 3: 126. 1818 [sic! 1827]. - *Faramea vaginata* Benth., nom. nud., Linnaea 23: 454. 1850; non Griseb. (1866, nom. superfl.). Tipo: Brasil, Pará [provavelmente perto de Belém], s.d., *F.W. Sieber s.n.* (holótipo B-W 2807 [*dedit Hoffmannsegg*])."

**Accepted name:** *Ixora pubescens* Willd. in Schult. & Schult. f., Mant. 3: 126. 1827.

Type: BRAZIL. Pará: [probably Belém], s.d., F.W. Sieber in J.C. Hoffmannsegg s.n. (holotype B-W [barcode B-W 02807 -01 0]; isotype frag. G [barcode G00634186]).

**Notes:** Joseph A. Schultes and Julius H. Schultes (1827: 126) published *Ixora pubescens* Willd. in Schult. & Schult. f. (Art. 46.3, Ex. 15) and cited the material studied as "*In Para Brasiliae*, com. a *Hoffmannsegg*." That material was collected by Sieber, and Hoffmannsegg never set foot in Brazil. In B-W there is a specimen, with barcode B-W 02807 -01 0, with the handwritten label "arbuste rbr. gull. (Sieber)" affixed on the bottom right corner of the sheet. The annotation "Ix. pubescens" is handwritten directly on the upper right corner of that sheet. This is the specimen of *Ixora pubescens* studied by Willdenow and is the holotype.

In G there is a sheet, barcode G00634186, with an envelope containing one leaf and the fragment of an inflorescence. On the envelope is handwritten "Ixora pubescens Willd., Pará: Hoffmannsegg, fol. numis & flor." This is an isotype fragment of this name.

FGT, vol. 40(2), p. 609:

**Synonym:**

"*Ixora pubescens* var. *glabrifolia* Muell. Arg. in Mart., Fl. Bras. 6(5): 62. 1881. Tipo: Brasil, Pará, "in vicinibus Pará" [agora Belém], *Spruce 328* (lectótipo G, aqui seleccionado, isolectótipos G, K, foto-G [lectótipo] em US, foto-G [isolectótipo] em NY, foto-K em NY)."

Type: BRAZIL. Pará: "in vicinibus Pará" [near Belém], s.d. [Jul.-Aug. 1849], R. *Spruce 328* (first-step lectotype designated by Delprete (2010b: 609); G [barcode G00634184], **second-step lectotype here designated**; isolectotypes, FI [barcode FI004824], G [barcode G00634185], K [barcode K000424499], M [barcode M-0187733]; photo-G [F neg. No. 25677] in NY; photo-K in NY).

**Notes:** Müller Argoviensis (1881: 62) cited the gatherings *Spruce 328*, *Burchell 9456*, and *Burchell 9270* for *Ixora pubescens* var. *glabrifolia* Müll. Arg., without citing the herbarium of deposit. In G, there are two specimens of *Spruce 328*. Therefore, Delprete's (2010b: 609) statement "lectótipo G, aqui seleccionado" should be treated as first-step lectotypification. The specimen with the annotation "I. pubescens β glabrifolia, scripsit Müll.-
Arg.!”, with barcode G00634184, is here designated the
second-step lectotype of this name.

FGT, vol. 40(2), p. 611:
“29-10. **Ixora undulata** Roxb. in Carey & Wall. (Eds.),
Fl. Indica ou escr. Indian Pl. 1: 395. 1820. Tipo: Bang-
ladesh/India, “nativo das florestas de Bengala”, s.d.,
*Roxburg s.n.* (não visto).”

**Accepted name:** *Ixora undulata* Roxb. ex Sm.

**Correct author and bibliographic citation:** *Ixora undulata* Roxb. ex Sm. in Rees, Cycl. 19: Ixora No. 7. 1814.

Type: “EAST INDIES”: Without locality, 1808, *Lord Valentia s.n.*, Herb. Smith No. 189.9 (LINN [bar-code unknown], lectotype designated by I.M. Turner (2021: 397)).

**Synonym:**

*Ixora undulata* Roth in Roemer & Schultes, Syst. Veg. 3: 178. 1818, *nom. illeg.*, non Roxb. ex Sm. 1814 ["*Ixora undulata* B.Heyne ex Roth", Nov. Pl. Sp.: 91. 1821, isonym].

Type: [INDIA]: Without locality, s.d., *W. Roxburgh s.n.* (B-W [barcode B -W 02809 -01 0], neotype designated by Turner (2021: 397)).

*Ixora undulata* Roxb., Fl. Ind. 1: 385. 1820, *nom. illeg.*, non Roxb. ex Sm. 1814, nec Roth 1818.

Type: [INDIA]: Without locality, s.d., *W. Roxburgh s.n.* (K n.v. [barcode unknown], lectotype designated by Husain and Paul in J. Econ. Taxon. Bot., Addit. Ser. 6: 178. 1989).

**Notes:** Ian M. Turner (2021: 397) explained the lectotypification of *Ixora undulata* Roxb. ex Sm. and its synonyms as follows: “This is another case where the same name was published by different authors over a relatively short period. In this case they all concern the same species. No specimens likely to have been seen by Roth have been found. The species is confined to North India, so it is likely that Heyne was sent a specimen by Roxburgh, or received one when he visited Roxburgh in Calcutta in late 1811, before his departure from India on leave (Heyne, 1814). Therefore, selecting a Roxburgh specimen as neotype for Roth’s name seems acceptable. The earliest validation of *Ixora undulata* was by Smith.

There is a specimen from Roxburgh in Smith's herbarium in LINN, which is here designated as lectotype for Smith's name. There is more than one Roxburgh specimen of this species in K, so I am not currently able to state which specimen is the lectotype of *Ixora undulata* Roxb."

30. **LADENBERGIA** Klotzsch in Hayne, Getr. Darst. Gew. 14: tab. 15. 1846.

FGT, vol. 40(2), p. 614:
“30-1. **Ladenbergia cujabensis** Klotzsch in Hayne, Getr. Darst. Gew. 14: tab. 15. 1846. - *Remijia cujabensis* (Klotzsch) Wedd., Ann. Sci. Nat., Sér. 3, 10: 13. 1848. - *Cinchona cujabensis* Manso ex Klotzsch in Hayne, *pro syn.*, Getr. Darst. Gew. 14: tab. 15. 1846. - Tipo: Brasil, Mato Grosso, perto de Cuiabá, s.d., *Manso & Lhotsky 20* (holótipo, B, destruído, foto em NY; lectótipo, F, aqui escolhido, isolectótipo, G).”

**Accepted name:** *Ladenbergia cujabensis* Klotzsch ex Walp.

Type: BRAZIL. Mato Grosso: Near Cuiabá, 1832, *P.A.L. da Silva Manso & J. Lhotsky 20* (F [Acc. No. 768356], **lectotype here designated**; isolectotypes, B†, BR [barcode 00000552387, without collection number], F [Acc. No. 686819], G [(2 sheets) barcode G00405218]).

**Notes:** There are three citations of *Cinchona cujabensis* Manso prior or contemporary to the publication of *Ladenbergia cujabensis* Klotzsch ex Walp. (Walpers, 1846): Fürnrohr (1832: 26), Martius (1843: 57), and Mérat (1846: 614). However, those three publications cite *C. cujabensis* as a *nomen nudum*. Hence, the combinations based on *C. cujabensis* are invalid, because the basionym is a *nomen nudum*.

*Ladenbergia cujabensis* Klotzsch ex Walp. was first validly published by Walpers (1846: 67), who was the first author to provide a description of this species and attributed the name to Klotzsch.

Weddell (1848: 13) published the combination *Remijia cujabensis* (Klotzsch ex Walp.) Wedd. His citation was “8. R. Cujabensis. — Ladenbergia Cujabensis Kltzsch. l. c.” This is an indirect reference to *Ladenbergia cujabensis*, the basionym, because he cited Klotzsch.

Andersson (1997: 276) cited the authority of *Ladenbergia cujabensis* as "*Ladenbergia cujabensis* Klotzsch in Hayne, Getreue Darstell. Gew.] 14 t. 15. 1846." Hayne's (1846) publication consists of a series of
Arg.!”, with barcode G00634184, is here designated the
second-step lectotype of this name.

FGT, vol. 40(2), p. 611:
"29-10. **Ixora undulata** Roxb. in Carey & Wall. (Eds.),
Fl. Indica ou escr. Indian Pl. 1: 395. 1820. Tipo: Bang-
ladesh/India, "nativo das florestas de Bengala", s.d.,
*Roxburg s.n.* (não visto)."

**Accepted name:** *Ixora undulata* Roxb. ex Sm.

**Correct author and bibliographic citation:** *Ixora undulata* Roxb. ex Sm. in Rees, Cycl. 19: Ixora No. 7. 1814.

Type: "EAST INDIES": Without locality, 1808, *Lord Valentia s.n.*, Herb. Smith No. 189.9 (LINN [bar-code unknown], lectotype designated by I.M. Turner (2021: 397)).

**Synonym:**

*Ixora undulata* Roth in Roemer & Schultes, Syst. Veg. 3: 178. 1818, *nom. illeg.*, non Roxb. ex Sm. 1814 ["*Ixora undulata* B.Heyne ex Roth", Nov. Pl. Sp.: 91. 1821, isonym].

Type: [INDIA]: Without locality, s.d., *W. Roxburgh s.n.* (B-W [barcode B -W 02809 -01 0], neotype designated by Turner (2021: 397)).

*Ixora undulata* Roxb., Fl. Ind. 1: 385. 1820, *nom. illeg.*, non Roxb. ex Sm. 1814, nec Roth 1818.

Type: [INDIA]: Without locality, s.d., *W. Roxburgh s.n.* (K n.v. [barcode unknown], lectotype designated by Husain and Paul in J. Econ. Taxon. Bot., Addit. Ser. 6: 178. 1989).

**Notes:** Ian M. Turner (2021: 397) explained the lectotypification of *Ixora undulata* Roxb. ex Sm. and its synonyms as follows: "This is another case where the same name was published by different authors over a relatively short period. In this case they all concern the same species. No specimens likely to have been seen by Roth have been found. The species is confined to North India, so it is likely that Heyne was sent a specimen by Roxburgh, or received one when he visited Roxburgh in Calcutta in late 1811, before his departure from India on leave (Heyne, 1814). Therefore, selecting a Roxburgh specimen as neotype for Roth's name seems acceptable. The earliest validation of *Ixora undulata* was by Smith.

There is a specimen from Roxburgh in Smith's herbarium in LINN, which is here designated as lectotype for Smith's name. There is more than one Roxburgh specimen of this species in K, so I am not currently able to state which specimen is the lectotype of *Ixora undulata* Roxb."

30. **LADENBERGIA** Klotzsch in Hayne, Getr. Darst. Gew. 14: tab. 15. 1846.

FGT, vol. 40(2), p. 614:
"30-1. **Ladenbergia cujabensis** Klotzsch in Hayne, Getr. Darst. Gew. 14: tab. 15. 1846. - *Remijia cujabensis* (Klotzsch) Wedd., Ann. Sci. Nat., Sér. 3, 10: 13. 1848. - *Cinchona cujabensis* Manso ex Klotzsch in Hayne, *pro syn.*, Getr. Darst. Gew. 14: tab. 15. 1846. - Tipo: Brasil, Mato Grosso, perto de Cuiabá, s.d., *Manso & Lhotsky 20* (holótipo, B, destruído, foto em NY; lectótipo, F, aqui escolhido, isolectótipo, G)."

**Accepted name:** *Ladenbergia cujabensis* Klotzsch ex Walp.

Type: BRAZIL. Mato Grosso: Near Cuiabá, 1832, P.A.L. da Silva Manso & J. Lhotsky 20 (F [Acc. No. 768356], **lectotype here designated**; isolectotypes, B†, BR [barcode 00000552387, without collection number], F [Acc. No. 686819], G [(2 sheets) barcode G00405218]).

**Notes:** There are three citations of *Cinchona cujabensis* Manso prior or contemporary to the publication of *Ladenbergia cujabensis* Klotzsch ex Walp. (Walpers, 1846): Fürnrohr (1832: 26), Martius (1843: 57), and Mérat (1846: 614). However, those three publications cite *C. cujabensis* as a *nomen nudum*. Hence, the combinations based on *C. cujabensis* are invalid, because the basionym is a *nomen nudum*.

*Ladenbergia cujabensis* Klotzsch ex Walp. was first validly published by Walpers (1846: 67), who was the first author to provide a description of this species and attributed the name to Klotzsch.

Weddell (1848: 13) published the combination *Remijia cujabensis* (Klotzsch ex Walp.) Wedd. His citation was "8. R. Cujabensis. — Ladenbergia Cujabensis Kltzsch. l. c." This is an indirect reference to *Ladenbergia cujabensis*, the basionym, because he cited Klotzsch.

Andersson (1997: 276) cited the authority of *Ladenbergia cujabensis* as "*Ladenbergia cujabensis* Klotzsch in Hayne, Getreue Darstell. Gew.] 14 t. 15. 1846." Hayne's (1846) publication consists of a series of
plates accompanied by an unpaginated text. The text of plate 14, on line 22, says “8) L. (Casc.) cujabensis Kl. (Cinchona cujabensis Manso) Brasilia.” On line 22, there is no description, but a basionym is cited, *C. cujabensis*. Andersson (1997: 276) cited the type of *L. cujabensis* as “Type: *Manso & Lhotsky 20*; Brazil, Mato Grosso, vicinity of Cuiabá (B holotype, destroyed, photo F-155; F, G isotypes),” Andersson’s “holotype” cannot be corrected to “lectotype” because the B specimens was destroyed during WWII.

Delprete (2010b: 614) followed Andersson's citation of the authority of this name, by citing *Ladenbergia cujabensis* Klotzsch in Hayne. Delprete (2010b: 614) designated the lectotype of this name as a specimen at F. However, at F there are two sheets of *Manso & Lhotsky 20*, both consisting of fragments removed from material in G. Hence, a lectotype needs to be designated.

The F specimen with Acc. No. 686819 is a fragment with one loose leaf and an infructescence with open capsules. The other specimen at F, Acc. No. 768356, has a label with the heading “Herbier de Candolle” on which is handwritten “No. 20” and “M. da Silva Manso” and another label with the handwritten annotation “20. Ladenbergia cujabensis Kl. Cujabae. fl. nov. - leg. M. da Silva Manso 1832.” The F specimen with Acc. No. 768356 consists of a small branch with three leaves and an infructescence with open capsules and is here designated the lectotype of this name.

In G there are two sheets, with barcode G00405218, that are original material associated with this name. On the first sheet is affixed a label with the annotation “M. da Silva Manso 1832, 20, Cinchona Cuyabensis Manso, [...] Cuyaba, fl. Nov.” On both sheets are affixed branches with several leaves and inflorescences with dehisced capsules. This specimen, mounted on two sheets, is an isolectotype of *Ladenbergia cujabensis*.

31. **LIMNOSIPANEA** Hook. f., Hooker's Icon. Pl. 11: 38. 1868.

FGT, vol. 40(2), p. 621:
“31-1. *Limnosipanea erythreoides* (Cham.) K. Schum. in Mart., Fl. Bras. 6(6): 253, tab. 123. 1889. - *Sipanea erythraeoides* Cham., Linnaea 9: 242. 1834. Tipo: "Brasilia inter tropicos", s.d., *Sellow s.n.* (holótipo B, destruído).”

**Accepted name:** *Limnosipanea erythraeoides*
(Cham.) K. Schum.

Type: BRAZIL. Goiás: Near Caiapônia, "in arenosis humidiusque, Aldeia dos Cayapos" [sandy

humid savanna, near the indigenous village of the Cayapos (now the town of Caiapônia, ca. 16°57'S, 51°48'W)], Aug. 1826 (fl), *L. Riedel 409* (BR [barcode 00000824532], phenotype designated by Delprete (2022: 54); isotype BM [barcode BM000614341], FI [barcode FI018882], P [barcode P00729267], US [Acc. No. 254488]).

FGT, vol. 40(2), p. 624:
“31-2. *Limnosipanea palustris* (Seem.) Hook. f. in Hook., Ic. Pl. 11: 38, pl. 1050. 1868. - *Sipania palustris* Seem., Bot. Voy. Herald 136. 1854; non *Sipanea palustris* (A. Rich.) J.H. Kirkbr., nom. superfl. [= *Sipanea wilson-brownei* R.S. Cowan]. Tipo: Panamá, perto da cidade de Panamá, *Seeman 347* (holótipo, K; isótipo, K, fotos em NY).”

**Accepted name:** *Limnosipanea palustris* (Seem.) Hook. f.

**Correct citation:** *Limnosipanea palustris* (Seem.) Hook. f., Ic. pl. 11: 38. 1868. - *Sipanea* [as "*Sipania*"] *palustris* Seem., Bot. Voy. Herald 136. 1854; non *Sipanea palustris* (A. Rich.) J.H. Kirkbr., Brittonia 49: 360. 1997, *comb. illeg. superfl.* (= *Sipanea wilson-brownei* R.S. Cowan); non *Bertiera palustris* A. Rich. ex DC. (Sep. 1830: 392).

Type: PANAMA: Near Panama City, swamps, s.d. [1846-1849], *B.C. Seemann 347* (BM [barcode BM 000614301]), lectotype designated by Delprete (2022: 57); isotype K [without barcode]; photo-BM at BM, MO, NY, US).

FGT, vol. 40(2), p. 624:
**Synonym:**
“*Limnosipanea schomburgkii* Hook. f. in Hook., Icon. Pl. 1050. 1868.”

**Correct citation:** *Limnosipanea palustris* (Seem.) Hook. f., Ic. pl. 11: 38. 1868.

Type: GUYANA. [“Roraima”, sic!], Rovuma River, s.d. [1842-1843], *R.H. Schomburgk ser. II, 464* (= *M.R. Schomburgk 744*), (BM) [BM000614338] lectotype designated by Delprete (2022: 57); isotype types F [Acc. No. 766892], G [2 sheets, barcodes G00379139, G00379140], K [without barcode], P [barcode P00729461], US [barcode 00588519]; photo-P at NY; photo-G at F [Neg. No. 25681]).
plates accompanied by an unpaginated text. The text of plate 14, on line 22, says “8) L. (Casc.) cujabensis Kl. (Cinchona cujabensis Manso) Brasilia.” On line 22, there is no description, but a basionym is cited, *C. cujabensis*. Andersson (1997: 276) cited the type of *L. cujabensis* as “Type: *Manso & Lhotsky 20*; Brazil, Mato Grosso, vicinity of Cuiabá (B holotype, destroyed, photo F-155; F, G isotypes),” Andersson’s “holotype” cannot be corrected to “lectotype” because the B specimens was destroyed during WWII.

Delprete (2010b: 614) followed Andersson's citation of the authority of this name, by citing *Ladenbergia cujabensis* Klotzsch in Hayne. Delprete (2010b: 614) designated the lectotype of this name as a specimen at F. However, at F there are two sheets of *Manso & Lhotsky 20*, both consisting of fragments removed from material in G. Hence, a lectotype needs to be designated.

The F specimen with Acc. No. 686819 is a fragment with one loose leaf and an infructescence with open capsules. The other specimen at F, Acc. No. 768356, has a label with the heading “Herbier de Candolle” on which is handwritten “No. 20” and “M. da Silva Manso” and another label with the handwritten annotation “20. Ladenbergia cujabensis Kl. Cujabae. fl. nov. - leg. M. da Silva Manso 1832.” The F specimen with Acc. No. 768356 consists of a small branch with three leaves and an infructescence with open capsules and is here designated the lectotype of this name.

In G there are two sheets, with barcode G00405218, that are original material associated with this name. On the first sheet is affixed a label with the annotation “M. da Silva Manso 1832, 20, Cinchona Cuyabensis Manso, [...] Cuyaba, fl. Nov.” On both sheets are affixed branches with several leaves and inflorescences with dehisced capsules. This specimen, mounted on two sheets, is an isolectotype of *Ladenbergia cujabensis*.

31. **LIMNOSIPANEA** Hook. f., Hooker's Icon. Pl. 11: 38. 1868.

FGT, vol. 40(2), p. 621:
“31-1. *Limnosipanea erythreoides* (Cham.) K. Schum. in Mart., Fl. Bras. 6(6): 253, tab. 123. 1889. - *Sipanea erythraeoides* Cham., Linnaea 9: 242. 1834. Tipo: "Brasilia inter tropicos", s.d., *Sellow s.n.* (holótipo B, destruído).”

**Accepted name:** *Limnosipanea erythraeoides*
(Cham.) K. Schum.

Type: BRAZIL. Goiás: Near Caiapônia, "in arenosis humidiusque, Aldeia dos Cayapos" [sandy

humid savanna, near the indigenous village of the Cayapos (now the town of Caiapônia, ca. 16°57'S, 51°48'W)], Aug. 1826 (fl), *L. Riedel 409* (BR [barcode 00000824532], phenotype designated by Delprete (2022: 54); isotype BM [barcode BM000614341], FI [barcode FI018882], P [barcode P00729267], US [Acc. No. 254488]).

FGT, vol. 40(2), p. 624:
“31-2. *Limnosipanea palustris* (Seem.) Hook. f. in Hook., Ic. Pl. 11: 38, pl. 1050. 1868. - *Sipania palustris* Seem., Bot. Voy. Herald 136. 1854; non *Sipanea palustris* (A. Rich.) J.H. Kirkbr., nom. superfl. [= *Sipanea wilson-brownei* R.S. Cowan]. Tipo: Panamá, perto da cidade de Panamá, *Seeman 347* (holótipo, K; isótipo, K, fotos em NY).”

**Accepted name:** *Limnosipanea palustris* (Seem.) Hook. f.

**Correct citation:** *Limnosipanea palustris* (Seem.) Hook. f., Ic. pl. 11: 38. 1868. - *Sipanea* [as "*Sipania*"] *palustris* Seem., Bot. Voy. Herald 136. 1854; non *Sipanea palustris* (A. Rich.) J.H. Kirkbr., Brittonia 49: 360. 1997, *comb. illeg. superfl.* (= *Sipanea wilson-brownei* R.S. Cowan); non *Bertiera palustris* A. Rich. ex DC. (Sep. 1830: 392).

Type: PANAMA: Near Panama City, swamps, s.d. [1846-1849], *B.C. Seemann 347* (BM [barcode BM 000614301]), lectotype designated by Delprete (2022: 57); isotype K [without barcode]; photo-BM at BM, MO, NY, US).

FGT, vol. 40(2), p. 624:
**Synonym:**
“*Limnosipanea schomburgkii* Hook. f. in Hook., Icon. Pl. 1050. 1868.”

**Correct citation:** *Limnosipanea palustris* (Seem.) Hook. f., Ic. pl. 11: 38. 1868.

Type: GUYANA. [“Roraima”, sic!], Rovuma River, s.d. [1842-1843], *R.H. Schomburgk ser. II, 464* (= *M.R. Schomburgk 744*), (BM) [BM000614338] lectotype designated by Delprete (2022: 57); isotype types F [Acc. No. 766892], G [2 sheets, barcodes G00379139, G00379140], K [without barcode], P [barcode P00729461], US [barcode 00588519]; photo-P at NY; photo-G at F [Neg. No. 25681]).
34. **MANETTIA** Mutis ex L., Mant. 2: 553, 558. 1771, *nom. cons.*

FGT, vol. 40(2), p. 648:

"34-1. *Manettia cordifolia* Mart., Königl. Akad. Wiss. Königl. Gen.-Conserv. Wiss. Saaml. München 9: 95, pl. 7. 1824. - *Manettia ignita* var. *cordifolia* (Mart.) K. Schum. in Mart., Fl. Bras. 6(6): 170. 1889. - Tipo: Brasil, Minas Gerais, Villa Rica [agora Ouro Preto], s.d., *Martius s.n.* (holótipo M)."

Accepted name: *Manettia cordifolia* Mart.

Type: BRAZIL. Minas Gerais: "Villa Rica" [now the town of Ouro Preto], "habitat in sepibus, locis camporibus ad Villam Ricam etc. Provinciae Min. Geraes, Apr.", s.d. [Apr. 1818], C.F.P. *Martius s.n.* (M [barcode M-0198125], lectotype designated by Gauto et al. (2022: 430)).

**Notes:** Martius (1824: 96) cited the locality of *Manettia cordifolia* Mart., from material collected by himself, as "Habitat in sepibus et inter virgulta nec non in sylvarum prope Villam Ricam et alibi in Provincia Minarum [...]." At M there are seven specimens with this name, collected by Martius in Minas Gerais. On the stem of each specimen is attached a small tag with a number handwritten by Töpfer in the 1900s (for reference see Fleischmann & Gonella, 2020). Those specimens are described below.

Specimen with barcode M-0198125: label without heading, with the handwritten annotation "Nacibea cordifolia Mart., habitat in sepibus, locis camporibus ad Villam Ricam etc. Provinciae Min. Geraes, Apr." and the printed text "Dr. Martius Iter Brasil." A second label, handwritten by K. Schumann says "Manettia ignita M. var. cordifolia m" and the stamp "det Schumann in Fl. Bras." The specimen has a small tag attached to the stem with the number "3015" handwritten by Töpfer. This specimen consists of a significant plant portion with numerous leaves, numerous flower buds, and numerous flowers in anthesis. In addition, it has a label saying that it was collected in an open field near Villa Rica, now the town of Ouro Preto, Minas Gerais. This specimen was designated as the lectotype of *Manettia cordifolia* by Gauto et al. (2022: 430).

Specimen with barcode M-0198128: label without heading, with the handwritten annotation "Manettia, habitat in sylvis ad Mariana et V.R.ª [Villa Rica], Provinciae M. G., Apr." and the printed text "Dr. Martius Iter Brasil." A second label, handwritten by K. Schumann says "Manettia ignita m. var. cordifolia m" and the stamp "det. Schumann in Fl. Bras." On the specimen stem is attached a small tag

with the number "3034" handwritten by Töpfer.

Specimen with barcode M-0198129: label without heading, with the handwritten annotation "Nacibea punicea Mart., Habitat in Campis ad Grão Magor Montem [Grão Mogol Mountain], Provinciae Min. Geraes, Aug." and the printed text "Dr. Martius Iter Brasil." A second label, handwritten by K. Schumann says "Manettia ignita m. var. cordifolia m" and the stamp "det Schumann in Fl. Bras." On the specimen stem is attached a small tag with the number "3026" handwritten by Töpfer.

Specimen with barcode M-0198130: label with heading "Herbarium Regium Monacense" and the handwritten annotation "Manettia, Martius Iter Brasiliense." A second label, printed, says "Manettia [ignita K. Schum. var. cordifolia K. Schum]." On the specimen stem is attached a small tag with the number "3012" handwritten by Töpfer.

Specimen with barcode M-0198131: label with heading "Herbarium Regium Monacense" and the handwritten annotation "Manettia, Martius Iter Brasiliense." A second label has the printed text "Manettia [ignita K. Schum. var. cordifolia K. Schum]." On the specimen stem is attached a small tag with the number "3011" handwritten by Töpfer.

Specimen with barcode M-0198132: label with heading "Herbarium Regium Monacense" and the handwritten annotation "Manettia, Martius Iter Brasiliense." A second label has the printed text "Manettia [ignita K. Schum. var. cordifolia K. Schum]." On the specimen stem is attached a small tag with the number "3010" handwritten by Töpfer.

Specimen with barcode M-0198134: label without heading, and the handwritten annotation "Sabicea acuminata Mart., Mart. Obs. 638., Habitat in campestribus, prope Villam Ricam, Provinciae Min. Geraes, Apr." and the printed text ""Dr. Martius Iter Brasil." A second label, handwritten by K. Schumann says "Manettia ignita M. an M. gracilis Ch. et Schl. exempl. minimi imperfectum" and the stamp "det Schumann in Fl. Bras." On the specimen stem is attached a small tag with the number "3007" handwritten by Töpfer.

FGT, vol. 40(2), p. 656:

"34-2. *Manettia irwinii* Steyerm., Brittonia 30: 36. 1978. - Tipo: Brasil. Goiás: Serra dos Pireneus, Valley of Rio Corumbá, steep rocky slope 15 km N of Corumbá de Goiás, on road to Niquelândia, 1150 m, 17/I/1968 (fl), H.S. Irwin, H. Maxwell & D. Wasshausen 18688 (holótipo, UB; isótipos, K n.v., MBM, NY, VEN)."

Accepted name: *Manettia irwinii* Steyerm.
34. **MANETTIA** Mutis ex L., Mant. 2: 553, 558. 1771, *nom. cons.*

FGT, vol. 40(2), p. 648:

"34-1. *Manettia cordifolia* Mart., Königl. Akad. Wiss. Königl. Gen.-Conserv. Wiss. Saaml. München 9: 95, pl. 7. 1824. - *Manettia ignita* var. *cordifolia* (Mart.) K. Schum. in Mart., Fl. Bras. 6(6): 170. 1889. - Tipo: Brasil, Minas Gerais, Villa Rica [agora Ouro Preto], s.d., *Martius s.n.* (holótipo M)."

Accepted name: *Manettia cordifolia* Mart.

Type: BRAZIL. Minas Gerais: "Villa Rica" [now the town of Ouro Preto], "habitat in sepibus, locis camporibus ad Villam Ricam etc. Provinciae Min. Geraes, Apr.", s.d. [Apr. 1818], C.F.P. *Martius s.n.* (M [barcode M-0198125], lectotype designated by Gauto et al. (2022: 430)).

**Notes:** Martius (1824: 96) cited the locality of *Manettia cordifolia* Mart., from material collected by himself, as "Habitat in sepibus et inter virgulta nec non in sylvarum prope Villam Ricam et alibi in Provincia Minarum [...]." At M there are seven specimens with this name, collected by Martius in Minas Gerais. On the stem of each specimen is attached a small tag with a number handwritten by Töpfer in the 1900s (for reference see Fleischmann & Gonella, 2020). Those specimens are described below.

Specimen with barcode M-0198125: label without heading, with the handwritten annotation "Nacibea cordifolia Mart., habitat in sepibus, locis camporibus ad Villam Ricam etc. Provinciae Min. Geraes, Apr." and the printed text "Dr. Martius Iter Brasil." A second label, handwritten by K. Schumann says "Manettia ignita M. var. cordifolia m" and the stamp "det Schumann in Fl. Bras." The specimen has a small tag attached to the stem with the number "3015" handwritten by Töpfer. This specimen consists of a significant plant portion with numerous leaves, numerous flower buds, and numerous flowers in anthesis. In addition, it has a label saying that it was collected in an open field near Villa Rica, now the town of Ouro Preto, Minas Gerais. This specimen was designated as the lectotype of *Manettia cordifolia* by Gauto et al. (2022: 430).

Specimen with barcode M-0198128: label without heading, with the handwritten annotation "Manettia, habitat in sylvis ad Mariana et V.R.ª [Villa Rica], Provinciae M. G., Apr." and the printed text "Dr. Martius Iter Brasil." A second label, handwritten by K. Schumann says "Manettia ignita m. var. cordifolia m" and the stamp "det. Schumann in Fl. Bras." On the specimen stem is attached a small tag

with the number "3034" handwritten by Töpfer.

Specimen with barcode M-0198129: label without heading, with the handwritten annotation "Nacibea punicea Mart., Habitat in Campis ad Grão Magor Montem [Grão Mogol Mountain], Provinciae Min. Geraes, Aug." and the printed text "Dr. Martius Iter Brasil." A second label, handwritten by K. Schumann says "Manettia ignita m. var. cordifolia m" and the stamp "det Schumann in Fl. Bras." On the specimen stem is attached a small tag with the number "3026" handwritten by Töpfer.

Specimen with barcode M-0198130: label with heading "Herbarium Regium Monacense" and the handwritten annotation "Manettia, Martius Iter Brasiliense." A second label, printed, says "Manettia [ignita K. Schum. var. cordifolia K. Schum]." On the specimen stem is attached a small tag with the number "3012" handwritten by Töpfer.

Specimen with barcode M-0198131: label with heading "Herbarium Regium Monacense" and the handwritten annotation "Manettia, Martius Iter Brasiliense." A second label has the printed text "Manettia [ignita K. Schum. var. cordifolia K. Schum]." On the specimen stem is attached a small tag with the number "3011" handwritten by Töpfer.

Specimen with barcode M-0198132: label with heading "Herbarium Regium Monacense" and the handwritten annotation "Manettia, Martius Iter Brasiliense." A second label has the printed text "Manettia [ignita K. Schum. var. cordifolia K. Schum]." On the specimen stem is attached a small tag with the number "3010" handwritten by Töpfer.

Specimen with barcode M-0198134: label without heading, and the handwritten annotation "Sabicea acuminata Mart., Mart. Obs. 638., Habitat in campestribus, prope Villam Ricam, Provinciae Min. Geraes, Apr." and the printed text ""Dr. Martius Iter Brasil." A second label, handwritten by K. Schumann says "Manettia ignita M. an M. gracilis Ch. et Schl. exempl. minimi imperfectum" and the stamp "det Schumann in Fl. Bras." On the specimen stem is attached a small tag with the number "3007" handwritten by Töpfer.

FGT, vol. 40(2), p. 656:

"34-2. *Manettia irwinii* Steyerm., Brittonia 30: 36. 1978. - Tipo: Brasil. Goiás: Serra dos Pireneus, Valley of Rio Corumbá, steep rocky slope 15 km N of Corumbá de Goiás, on road to Niquelândia, 1150 m, 17/I/1968 (fl), H.S. Irwin, H. Maxwell & D. Wasshausen 18688 (holótipo, UB; isótipos, K n.v., MBM, NY, VEN)."

Accepted name: *Manettia irwinii* Steyerm.
Type: BRAZIL. Goiás: Serra dos Pireneus, Valley of Rio Corumbá, steep rocky slope 15 km N of Corumbá de Goiás, on road to Niquelândia, 1150 m, 17 Jan. 1968 (fl), H.S. Irwin, H. Maxwell & D. Wasshausen 18688 (holotype, UB [barcode UB0040326]; isotypes, K [barcode K000173773], MBM [barcode MBM068350], MO [Acc. No. 2817602], NY [barcode 00132193], US [barcode 00130538], VEN [Acc. No. 103020]).

FGT, vol. 40(2), p. 659:

"34-3. *Manettia luteo-rubra* (Vell.) Benth., Linnaea 23: 445. 1850. - *Guagnebina luteo-rubra* Vell., Fl. Flum. 46. 1825; Icon. 1: 121. 1831. Tipo: Brasil, coletor desconhecido, s.d., in Vellozo, Fl. Flum. Icon. 1: 121. 1831 (lectótipo)."

**Accepted name:** *Manettia luteorubra* (Vell.) Benth.

Type: BRAZIL. [State of Rio de Janeiro or São Paulo] [illustration]: Original parchment plate of *Florae Fluminensis* in the Manuscript Section of the Biblioteca Nacional of Rio de Janeiro [Catalogue No. mss1095062_125]], lectotype here designated.

**Notes:** Frei José Mariano da Conceição Vellozo (1742-1811) worked on the flora of the state of Rio de Janeiro and contiguous areas in the state of São Paulo and commended the drawings of the species to be published in his publication. According to Borgmeier (1937) and Carauta (1973), the text of *Florae Fluminensis* was printed in 1825 and distributed in 1829; and the illustrations in *Florae Fluminensis Icones* were edited in 1827 and published 1831. As Plate 121 of Vellozo's *Icones* was published two years after the publication of *Guagnebina luteorubra*, it cannot be treated as original material. An original drawing on parchment of *G. luteorubra* is kept in the Manuscript Section of the National Library in Rio de Janeiro. The original drawing prepared for Vellozo has the heading "Tetrand. Monog. GUAGNEBINA luteo rubra" printed at the top-center of the drawing, and the number "125" handwritten on the upper right corner. On the drawing is depicted a voluble branch with several solitary flowers. The original plate of *G. luteorubra*, with Catalogue No. mss1095062_125, is here designated the lectotype of this name.

35. **MITRACARPUS** Zucc. ex Schult. & Schult. f., Syst. Veg., Mant. 3: 210, 399. 1827.

FGT, vol. 40(2), p. 668:

"35-1. *Mitracarpus baturitensis* Sucre, Rodriguésia 26(38): 255. 1971. Tipo: Brasil, Ceará [Serra de Baturité], s.d., *A. Loefgren* 898 (holótipo, RB)."

**Accepted name:** *Mitracarpus baturitensis* Sucre

Type: BRAZIL. Ceará: [Serra de Baturité], s.d., *A. Loefgren* 898 (holotype, RB [Acc. No. 3899, barcode 00285709]).

FGT, vol. 40(2), p. 672:

"35-2. *Mitracarpus eritrichoides* Standl., Publ. Field Mus. Nat. Hist., Bot. 11: 223. 1936. Tipo: Brasil, Mato Grosso, Diamantino, nascente do Rio Paraguai, XII/1844 (fl, fr), *H.A. Weddell* 3090 (holótipo, P)."

**Accepted name:** *Mitracarpus eritrichoides* Standl.

Type: BRAZIL. Mato Grosso: Diamantino, nascente do Rio Paraguai, Dec. 1844 (fl, fr), *H.A. Weddell* 3090 (holotype, P [barcode P01090381], isotype fragment, F [ex P; Acc. No. 654376 barcode V0069777F]).

**Notes:** Standley (1936: 223-224) cited the type of *Mitracarpus eritrichoides* Standl. the specimen *Weddell* 3090 at P, which is the holotype (Souza and Cabral, 2010; Souza et al., 2010).

FGT, vol. 40(2), p. 676:

"35-3. *Mitracarpus hirtus* (L.) DC., Prodr. 4: 572. 1830. - *Spermacoce hirta* L., Sp. Pl. ed. 2. 148. 1762. Tipo: Jamaica, s.l., s.d., *Coletor Desconhecido s.n.* (holótipo, LINN 125.4, foto!)."

**Accepted name:** *Mitracarpus hirtus* (L.) DC.

Type: JAMAICA. Without locality, s.d. [1746-1755], *P. Browne s.n.* (holotype, LINN-HL 125.8).

**Notes:** Linnaeus (1762: 148) described *Spermacoce hirta* L. as "erecta subirsuta, folia elliptica, lineata, subtus pubescentia [...] verticilli multiflori" and cited "Brown. jam. 141. Habitat in Jamaica." Five years later, Linnaeus (1767: 115) re-described S. *hirta* as "scabra, fol. oblongis [...] Folia elliptica, lineata, scabra utrique (subtus vero imprimis nervis), non villosa, subpetiolata petiolis connexis membrana multiseta. Flores albi, tubulosi, laterales. Stamina fauce longiora. Antherae
Type: BRAZIL. Goiás: Serra dos Pireneus, Valley of Rio Corumbá, steep rocky slope 15 km N of Corumbá de Goiás, on road to Niquelândia, 1150 m, 17 Jan. 1968 (fl), H.S. Irwin, H. Maxwell & D. Wasshausen 18688 (holotype, UB [barcode UB0040326]; isotypes, K [barcode K000173773], MBM [barcode MBM068350], MO [Acc. No. 2817602], NY [barcode 00132193], US [barcode 00130538], VEN [Acc. No. 103020]).

FGT, vol. 40(2), p. 659:

"34-3. *Manettia luteo-rubra* (Vell.) Benth., Linnaea 23: 445. 1850. - *Guagnebina luteo-rubra* Vell., Fl. Flum. 46. 1825; Icon. 1: 121. 1831. Tipo: Brasil, coletor desconhecido, s.d., in Vellozo, Fl. Flum. Icon. 1: 121. 1831 (lectótipo)."

**Accepted name:** *Manettia luteorubra* (Vell.) Benth.

Type: BRAZIL. [State of Rio de Janeiro or São Paulo] [illustration]: Original parchment plate of *Florae Fluminensis* in the Manuscript Section of the Biblioteca Nacional of Rio de Janeiro [Catalogue No. mss1095062_125]], lectotype here designated.

**Notes:** Frei José Mariano da Conceição Vellozo (1742-1811) worked on the flora of the state of Rio de Janeiro and contiguous areas in the state of São Paulo and commended the drawings of the species to be published in his publication. According to Borgmeier (1937) and Carauta (1973), the text of *Florae Fluminensis* was printed in 1825 and distributed in 1829; and the illustrations in *Florae Fluminensis Icones* were edited in 1827 and published 1831. As Plate 121 of Vellozo's *Icones* was published two years after the publication of *Guagnebina luteorubra*, it cannot be treated as original material. An original drawing on parchment of *G. luteorubra* is kept in the Manuscript Section of the National Library in Rio de Janeiro. The original drawing prepared for Vellozo has the heading "Tetrand. Monog. GUAGNEBINA luteo rubra" printed at the top-center of the drawing, and the number "125" handwritten on the upper right corner. On the drawing is depicted a voluble branch with several solitary flowers. The original plate of *G. luteorubra*, with Catalogue No. mss1095062_125, is here designated the lectotype of this name.

35. **MITRACARPUS** Zucc. ex Schult. & Schult. f., Syst. Veg., Mant. 3: 210, 399. 1827.

FGT, vol. 40(2), p. 668:

"35-1. *Mitracarpus baturitensis* Sucre, Rodriguésia 26(38): 255. 1971. Tipo: Brasil, Ceará [Serra de Baturité], s.d., *A. Loefgren* 898 (holótipo, RB)."

**Accepted name:** *Mitracarpus baturitensis* Sucre

Type: BRAZIL. Ceará: [Serra de Baturité], s.d., *A. Loefgren* 898 (holotype, RB [Acc. No. 3899, barcode 00285709]).

FGT, vol. 40(2), p. 672:

"35-2. *Mitracarpus eritrichoides* Standl., Publ. Field Mus. Nat. Hist., Bot. 11: 223. 1936. Tipo: Brasil, Mato Grosso, Diamantino, nascente do Rio Paraguai, XII/1844 (fl, fr), *H.A. Weddell* 3090 (holótipo, P)."

**Accepted name:** *Mitracarpus eritrichoides* Standl.

Type: BRAZIL. Mato Grosso: Diamantino, nascente do Rio Paraguai, Dec. 1844 (fl, fr), *H.A. Weddell* 3090 (holotype, P [barcode P01090381], isotype fragment, F [ex P; Acc. No. 654376 barcode V0069777F]).

**Notes:** Standley (1936: 223-224) cited the type of *Mitracarpus eritrichoides* Standl. the specimen *Weddell* 3090 at P, which is the holotype (Souza and Cabral, 2010; Souza et al., 2010).

FGT, vol. 40(2), p. 676:

"35-3. *Mitracarpus hirtus* (L.) DC., Prodr. 4: 572. 1830. - *Spermacoce hirta* L., Sp. Pl. ed. 2. 148. 1762. Tipo: Jamaica, s.l., s.d., *Coletor Desconhecido s.n.* (holótipo, LINN 125.4, foto!)."

**Accepted name:** *Mitracarpus hirtus* (L.) DC.

Type: JAMAICA. Without locality, s.d. [1746-1755], *P. Browne s.n.* (holotype, LINN-HL 125.8).

**Notes:** Linnaeus (1762: 148) described *Spermacoce hirta* L. as "erecta subirsuta, folia elliptica, lineata, subtus pubescentia [...] verticilli multiflori" and cited "Brown. jam. 141. Habitat in Jamaica." Five years later, Linnaeus (1767: 115) re-described S. *hirta* as "scabra, fol. oblongis [...] Folia elliptica, lineata, scabra utrique (subtus vero imprimis nervis), non villosa, subpetiolata petiolis connexis membrana multiseta. Flores albi, tubulosi, laterales. Stamina fauce longiora. Antherae
violaceae” and did not cite any specimen or collection
locality.

The specimen LINN-HL 125.8 is annotated as “Sper-
macoce 3. Brown” by Solander, and “Spermacoce 3 his-
pida” by Linnaeus. Two other annotations are present:
“villosa? Swartz” and “villosa? Swartz, non hispida”, both
handwritten in pencil by J.E. Smith. On the sheet are two
branches with flowers and fruits. At the bottom of the
lower branch Linnaeus wrote “Br”, which confirms that
this specimen was collected by Patrick Brown in Jamaica.

The specimen LINN-HL 125.4 is annotated as “hir-
ta” by Linnaeus. No annotation by Linnaeus regarding
the collector is present on the sheet. This is the specimen
that he used to describe *Spermacoce hirta* in 1767. This
specimen consists of a sterile branch. Nicolson (1977:
572) wrote that “The description of flower and anther
color strongly suggests that Linnaeus prepared his
description from living material and pressed the flower-
ing specimen for his herbarium (now 125.4).” This speci-
men is not a species of *Mitracarpus*.

Verdcourt (1975: 317–322) provided a long discussion about the complex nomenclatural history and identity of this species, and stated that “the earliest name bestowed on the plant is *Spermacoce hirta* L., Sp. Pl. ed. 2, 1: 148 (1762) [...] The Brown reference (Civil and Nat. Hist. Jamaica: 141 (1756) is to SPERMACOCE 3. Erecta subirsuta, foliis oblongis venis arcuatis refertis, superioribus majoribus appropinquatus, floribus constipatis ad alas [...]” and “The actual Browne specimen referred to is still in existence and is preserved in the Linnean Herbarium as number 125.8. This specimen is undoubtedly to be taken as the holotype of *Spermacoce hirta* L. and is undoubtedly the same plant as *Mitracarpus villosus* (Sw.) DC.” (Verdcourt, 1975: 318–319).

Nicolson (1977) reconsidered the application of the
names *Mitracarpus hirtus* (L.) DC. vs. *M. villosus* (Sw.)
DC, and agreed with Verdcourt (1975) that LINN-HL
125.8 is the holotype of *Spermacoce hirta* L. (Linnaeus
1762: 148). Also, regarding *S. hirta* he stated that “I grant
that there is implicit exclusion of the type of *S. hir-
ta* Linnaeus (1762) from *S. hirta* Linnaeus (1767), since
the taxa circumscribed are clearly different, by modern
standards. However, there seems to be sufficient evi-
dence that Linnaeus (1767) explicitly excluded the type
of his 1762 name. By modern standards it appears that
Linnaeus simply misidentified his new material, 125.4,
with his earlier material, 125.8. Hence, *S. hirta* Lin-
naeus (1767) is not a later homonym of *S. hirta* Linnaeus
(1762), although the taxon to which he applied the name
is different.” (Nicolson 1977: 572).

Dwyer (1980: 284), Souza and Cabral (in Delprete
2010b: 676) and Souza et al. (2010: 334) erroneously cit-

ed the specimen LINN 125.4 as the holotype of *Sperma-
coce hirta*.

Jarvis (2007: 868) regarding *Spermacoce hirta* L. wrote “Lectotype (Verdcourt in Kew Bull. 30: 318. 1975): Herb. Linn. No. 125.8 (LINN). [...] *Note*: See discussion by Verdcourt, and Nicholson (in *Taxon* 26: 572. 1977) and Howard (*Fl. Less. Antilles* 6: 434. 1989).” It is clear that the sole original specimen of *S. hirta* L. (Linnaeus 1762: 148) is LINN 125.8, which is the holotype.

FGT, vol. 40(2), p. 683:
“35-4. *Mitracarpus microspermus* K. Schum. in Mart., Fl. Bras. 6(6): 83. 1888. Tipo: Brasil: [Roraima?], “ad fluvium Rio Branco”, 1840 (fl, fr), *Schomburgk 856 pro parte* (holótipo B, destruido; lectótipo BM, selecionado por E.B. Souza et al., 2010).”

**Accepted name:** *Mitracarpus microspermus* K. Schum.

Type: BRAZIL. [Roraima?], “ad fluvium Rio Brancos”, Jul. 1840 (fl, fr), *Rob. H. Schomburgk ser. I, 856 pro parte* (B†, photo F [F0BN000914]; BM [barcode BM014124066], lectotype designated by Souza et al. (Apr.–Jun. 2010: 338)).

**Notes:** Schumann (1888: 83–84) for *Mitracarpus microspermus* K. Schum. cited the gatherings *“Habitat in Guiana Anglica ad fluvium Rio Branco: Rob. Schomburgk n. 856 ex p.; in Guiana Batava: hb. Petropol.; in republica Guatemalensi ad Mniogalpa: Friedrichstal”* and cited *“M. scabrellum* Benth. in J. Bot. 3: 238 ex p. (Bentham, 1841)*” as a synonym of *M. microspermus*.

Souza et al. (Apr.–Jun. 2010: 338) designated as lectotype of *Mitracarpus microspermus* the specimen *Rob. Schomburgk ser. I, 856 pro parte* at BM. That specimen has barcode BM014124066, and has a label with the handwritten annotation “On the Rio Branco. In Guiana Anglica, legit Schomburgk No. 856!, VII 1840.” On the same label is handwritten “Mitracarpus scabrellum Benth! in Hook. Journ. III. p. 238.” However, there is no evidence on that sheet that it was studied by Schumann.

FGT, vol. 40(2), p. 683:

**Synonym:**

*Mitracarpus minutiflorus* K. Schum. in Mart., Fl. Bras. 6(6): 80. 1888. Tipo: Brasil, Goiás, “*inter urbem Goiaz et Cavalcante*”, 1828-1830 (fl, fr), *Burchell 7820* (holótipo B, destruído; lectótipo BR, selecionado por E.B. Souza et al. 2010; isolectótipo K).”
violaceae” and did not cite any specimen or collection
locality.

The specimen LINN-HL 125.8 is annotated as “Sper-
macoce 3. Brown” by Solander, and “Spermacoce 3 his-
pida” by Linnaeus. Two other annotations are present:
“villosa? Swartz” and “villosa? Swartz, non hispida”, both
handwritten in pencil by J.E. Smith. On the sheet are two
branches with flowers and fruits. At the bottom of the
lower branch Linnaeus wrote “Br”, which confirms that
this specimen was collected by Patrick Brown in Jamaica.

The specimen LINN-HL 125.4 is annotated as “hir-
ta” by Linnaeus. No annotation by Linnaeus regarding
the collector is present on the sheet. This is the specimen
that he used to describe *Spermacoce hirta* in 1767. This
specimen consists of a sterile branch. Nicolson (1977:
572) wrote that “The description of flower and anther
color strongly suggests that Linnaeus prepared his
description from living material and pressed the flower-
ing specimen for his herbarium (now 125.4).” This speci-
men is not a species of *Mitracarpus*.

Verdcourt (1975: 317–322) provided a long discussion about the complex nomenclatural history and identity of this species, and stated that “the earliest name bestowed on the plant is *Spermacoce hirta* L., Sp. Pl. ed. 2, 1: 148 (1762) [...] The Brown reference (Civil and Nat. Hist. Jamaica: 141 (1756) is to SPERMACOCE 3. Erecta subirsuta, foliis oblongis venis arcuatis refertis, superioribus majoribus appropinquatus, floribus constipatis ad alas [...]” and “The actual Browne specimen referred to is still in existence and is preserved in the Linnean Herbarium as number 125.8. This specimen is undoubtedly to be taken as the holotype of *Spermacoce hirta* L. and is undoubtedly the same plant as *Mitracarpus villosus* (Sw.) DC.” (Verdcourt, 1975: 318–319).

Nicolson (1977) reconsidered the application of the
names *Mitracarpus hirtus* (L.) DC. vs. *M. villosus* (Sw.)
DC, and agreed with Verdcourt (1975) that LINN-HL
125.8 is the holotype of *Spermacoce hirta* L. (Linnaeus
1762: 148). Also, regarding *S. hirta* he stated that “I grant
that there is implicit exclusion of the type of *S. hir-
ta* Linnaeus (1762) from *S. hirta* Linnaeus (1767), since
the taxa circumscribed are clearly different, by modern
standards. However, there seems to be sufficient evi-
dence that Linnaeus (1767) explicitly excluded the type
of his 1762 name. By modern standards it appears that
Linnaeus simply misidentified his new material, 125.4,
with his earlier material, 125.8. Hence, *S. hirta* Lin-
naeus (1767) is not a later homonym of *S. hirta* Linnaeus
(1762), although the taxon to which he applied the name
is different.” (Nicolson 1977: 572).

Dwyer (1980: 284), Souza and Cabral (in Delprete
2010b: 676) and Souza et al. (2010: 334) erroneously cit-

ed the specimen LINN 125.4 as the holotype of *Sperma-
coce hirta*.

Jarvis (2007: 868) regarding *Spermacoce hirta* L. wrote “Lectotype (Verdcourt in Kew Bull. 30: 318. 1975): Herb. Linn. No. 125.8 (LINN). [...] *Note*: See discussion by Verdcourt, and Nicholson (in *Taxon* 26: 572. 1977) and Howard (*Fl. Less. Antilles* 6: 434. 1989).” It is clear that the sole original specimen of *S. hirta* L. (Linnaeus 1762: 148) is LINN 125.8, which is the holotype.

FGT, vol. 40(2), p. 683:
“35-4. *Mitracarpus microspermus* K. Schum. in Mart., Fl. Bras. 6(6): 83. 1888. Tipo: Brasil: [Roraima?], “ad fluvium Rio Branco”, 1840 (fl, fr), *Schomburgk 856 pro parte* (holótipo B, destruido; lectótipo BM, selecionado por E.B. Souza et al., 2010).”

**Accepted name:** *Mitracarpus microspermus* K. Schum.

Type: BRAZIL. [Roraima?], “ad fluvium Rio Brancos”, Jul. 1840 (fl, fr), *Rob. H. Schomburgk ser. I, 856 pro parte* (B†, photo F [F0BN000914]; BM [barcode BM014124066], lectotype designated by Souza et al. (Apr.–Jun. 2010: 338)).

**Notes:** Schumann (1888: 83–84) for *Mitracarpus microspermus* K. Schum. cited the gatherings *Habitat in Guiana Anglica ad fluvium Rio Branco: Rob. Schomburgk n. 856 ex p.*; in *Guiana Batava: hb. Petropol.*; in *republica Guatemalensi ad Mniogalpa: Friedrichstal*” and cited “*M. scabrellum* Benth. in J. Bot. 3: 238 ex p. (Bentham, 1841)” as a synonym of *M. microspermus*.

Souza et al. (Apr.–Jun. 2010: 338) designated as lectotype of *Mitracarpus microspermus* the specimen *Rob. Schomburgk ser. I, 856 pro parte* at BM. That specimen has barcode BM014124066, and has a label with the handwritten annotation “On the Rio Branco. In Guiana Anglica, legit Schomburgk No. 856!, VII 1840.” On the same label is handwritten “Mitracarpus scabrellum Benth! in Hook. Journ. III. p. 238.” However, there is no evidence on that sheet that it was studied by Schumann.

FGT, vol. 40(2), p. 683:

**Synonym:**

*Mitracarpus minutiflorus* K. Schum. in Mart., Fl. Bras. 6(6): 80. 1888. Tipo: Brasil, Goiás, “*inter urbem Goiaz et Cavalcante*”, 1828-1830 (fl, fr), *Burchell 7820* (holótipo B, destruído; lectótipo BR, selecionado por E.B. Souza et al. 2010; isolectótipo K).”
Type: BRAZIL. Goiás: "*inter urbem Goiaz et Cavalcante*", 1828-1830 (fl, fr), W.J. Burchell 7820 (B†; BR [barcode 000000574153], lectotype designated by Souza et al. (Apr.-Jun. 2010: 338); isolectotypes GH [barcode 01154967], K [barcode K000447056]).

FGT, vol. 40(2), p. 688:

"35-5. *Mitracarpus parvulus* K. Schum. in Mart., Fl. Bras. 6(6): 84. 1888. Tipo: Tocantins, "ad Porto Real" [agora Porto Nacional], 1828-1830 (fl), *Burchell 8674* (holótipo B, destruído; lectótipo BR, selecionado por E.B. Souza et al. 2010; isolectótipos K, P). Paratipos (sin-tipos citados por K. Schumann): Brasil, Mato Grosso, "in siccis graminosis provinciae Goyas prope Cuyabá", III/1827 (fl, fr), *Riedel 870* (B, destruído, BR); Tocantins, "ad Porto Real" [agora Porto Nacional], 1828-1830 (fl), *Burchell 8658* (B, BR, K, P), "in Brasilia occidentali", *Tamberlik s.n.* (não localizado)."

**Accepted name:** *Mitracarpus parvulus* K.Schum.

Type: BRAZIL. Tocantins: "ad Porto Real" [now Porto Nacional], 1828-1830 (fl), W.J. Burchell 8674 (B†; BR [barcode 000000558758], lectotype designated by Souza et al. (Apr.-Jun. 2010: 340); isolectotypes K [barcode K000174166], P [barcode P03911661]).

FGT, vol. 40(2), p. 690:

"35-6. *Mitracarpus recurvatus* Standl., Publ. Field Mus., Bot. 8: 384. 1931. Tipo: Brasil, Minas Gerais ["Goyaz"], Paracatu, "prés de porto" [?], 1894 (fl, fr), A. Glaziou 21511 (holótipo, K; isótipos, G, P)."

**Accepted name:** *Mitracarpus recurvatus* Standl.

Type: BRAZIL. Minas Gerais ["Brazil. Chiefly Province of Goyaz"): Paracatu, "prés de Porto", 1894 (fl, fr), A.F.M. Glaziou 21511 (holotype, K [barcode K000174497]; isotypes, F [fragment from K, Acc. No. 630505], G [barcode G00436966], P [not traced]).

FGT, vol. 40(2), p. 694:

"35-7. *Mitracarpus robustus* E.B. Souza & E.L. Cabral, Rodriguésia 61: 345, fig. 8 A-G. 2010. Tipo: Brasil, Ceará, Mun Porteiras, Chapada do Araripe, 7°28'S, 39°08'W, 930 m, 30/III/2000 (fl), P.G. Delprete, E.B. Souza, F.S. Cavalcanti & L.W. Lima-Verde 7316 (holótipo, EAC; isótipos, HUEFS, NY)."

**Accepted name:** *Mitracarpus polygonifolius* (A.St. Hil.) R.M.Salas & E.B.Souza, Rodriguésia 66(3): 921. 2015.

Type: BRAZIL. Ceará: Mun Porteiras, Chapada do Araripe, 7°28'S, 39°08'W, 930 m, 30 Mar. 2000 (fl), P.G. Delprete, E.B. Souza, F.S. Cavalcanti & L.W. Lima-Verde 7316 (holotype, EAC [2 sheets, Acc. No. 28985]; isotypes, HUEFS [barcode HUEFS0080844], NY [barcode unknown]).

*Mitracarpus polygonifolius* (A.St.Hil.) R.M.Salas & E.B.Souza, Rodriguésia 66(3): 921. 2015. – *Spermacoce polygonifolia* A. St. Hil., Voy. Distr. Diam. 1: 381. 1833.

Type: BRAZIL. Minas Gerais: "Villa Rica" [now Ouro Preto], s.d., A. de Saint-Hilaire s.n. (P [barcode P03818586], **lectotype here designated**; isolectotype, MPU [barcode MPU022474]).

**Notes:** Souza and Cabral in Souza et al. (Apr.-Jun. 2010: 345) proposed *Mitracarpus robustus* E.B. Souza & E.L. Cabral as a new name for *Mitracarpus frigidus* var. *humboldtianus*, which is not a valid name. They cited the holotype of *M. robustus* as the specimen *Delprete et al. 7316* at EAC. The EAC specimen is mounted on two sheets, both with Accession No. 28985. According to Art. 8.3 of the *Code* (Turland et al. 2018), the two sheets are treated as a single specimen with multiple preparations.

Salas et al. (2015) concluded that *Spermacoce polygonifolia* A.St.Hil. is synonymous with *Mitracarpus robustus* E.B.Souza & E.L.Cabral. Because Saint Hilaire's name has nomenclatural priority, they published the new combination *Mitracarpus polygonifolius* (A.St.Hil.) R.M.Salas & E.B.Souza, which is the name to be used for this species.

Saint-Hilaire (1833: 381-383) published *Spermacoce polygonifolia* with a detailed description and comparison with several species of *Borreria* and *Spermacoce*, but did not cite any material examined or the herbarium of deposit. Two original specimens annotated as *S. polygonifolia* by Saint-Hilaire are found at P and MPU. Salas et al. (2015) cited the specimen at P as the holotype. However, because Saint-Hilaire did not cite any specimen or herbarium of deposit, the P specimen cannot be treated as the holotype. Also, Salas et al.'s citation cannot be treated as an inadvertent lectotypification, because, according to the Code, it should have been accompanied by "here designated" or a similar expression. Hence the specimen at P, with barcode P03818586, is here designated the lectotype of this name.
Type: BRAZIL. Goiás: "*inter urbem Goiaz et Cavalcante*", 1828-1830 (fl, fr), W.J. Burchell 7820 (B†; BR [barcode 000000574153], lectotype designated by Souza et al. (Apr.-Jun. 2010: 338); isolectotypes GH [barcode 01154967], K [barcode K000447056]).

FGT, vol. 40(2), p. 688:

"35-5. *Mitracarpus parvulus* K. Schum. in Mart., Fl. Bras. 6(6): 84. 1888. Tipo: Tocantins, "ad Porto Real" [agora Porto Nacional], 1828-1830 (fl), *Burchell 8674* (holótipo B, destruído; lectótipo BR, selecionado por E.B. Souza et al. 2010; isolectótipos K, P). Paratipos (sin-tipos citados por K. Schumann): Brasil, Mato Grosso, "in siccis graminosis provinciae Goyas prope Cuyabá", III/1827 (fl, fr), *Riedel 870* (B, destruído, BR); Tocantins, "ad Porto Real" [agora Porto Nacional], 1828-1830 (fl), *Burchell 8658* (B, BR, K, P), "in Brasilia occidentali", *Tamberlik s.n.* (não localizado)."

**Accepted name:** *Mitracarpus parvulus* K.Schum.

Type: BRAZIL. Tocantins: "ad Porto Real" [now Porto Nacional], 1828-1830 (fl), W.J. Burchell 8674 (B†; BR [barcode 000000558758], lectotype designated by Souza et al. (Apr.-Jun. 2010: 340); isolectotypes K [barcode K000174166], P [barcode P03911661]).

FGT, vol. 40(2), p. 690:

"35-6. *Mitracarpus recurvatus* Standl., Publ. Field Mus., Bot. 8: 384. 1931. Tipo: Brasil, Minas Gerais ["Goyaz"], Paracatu, "prés de porto" [?], 1894 (fl, fr), A. Glaziou 21511 (holótipo, K; isótipos, G, P)."

**Accepted name:** *Mitracarpus recurvatus* Standl.

Type: BRAZIL. Minas Gerais ["Brazil. Chiefly Province of Goyaz"): Paracatu, "prés de Porto", 1894 (fl, fr), A.F.M. Glaziou 21511 (holotype, K [barcode K000174497]; isotypes, F [fragment from K, Acc. No. 630505], G [barcode G00436966], P [not traced]).

FGT, vol. 40(2), p. 694:

"35-7. *Mitracarpus robustus* E.B. Souza & E.L. Cabral, Rodriguésia 61: 345, fig. 8 A-G. 2010. Tipo: Brasil, Ceará, Mun Porteiras, Chapada do Araripe, 7°28'S, 39°08'W, 930 m, 30/III/2000 (fl), P.G. Delprete, E.B. Souza, F.S. Cavalcanti & L.W. Lima-Verde 7316 (holótipo, EAC; isótipos, HUEFS, NY)."

**Accepted name:** *Mitracarpus polygonifolius* (A.St. Hil.) R.M.Salas & E.B.Souza, Rodriguésia 66(3): 921. 2015.

Type: BRAZIL. Ceará: Mun Porteiras, Chapada do Araripe, 7°28'S, 39°08'W, 930 m, 30 Mar. 2000 (fl), P.G. Delprete, E.B. Souza, F.S. Cavalcanti & L.W. Lima-Verde 7316 (holotype, EAC [2 sheets, Acc. No. 28985]; isotypes, HUEFS [barcode HUEFS0080844], NY [barcode unknown]).

*Mitracarpus polygonifolius* (A.St.Hil.) R.M.Salas & E.B.Souza, Rodriguésia 66(3): 921. 2015. – *Spermacoce polygonifolia* A. St. Hil., Voy. Distr. Diam. 1: 381. 1833.

Type: BRAZIL. Minas Gerais: "Villa Rica" [now Ouro Preto], s.d., A. de Saint-Hilaire s.n. (P [barcode P03818586], **lectotype here designated**; isolectotype, MPU [barcode MPU022474]).

**Notes:** Souza and Cabral in Souza et al. (Apr.-Jun. 2010: 345) proposed *Mitracarpus robustus* E.B. Souza & E.L. Cabral as a new name for *Mitracarpus frigidus* var. *humboldtianus*, which is not a valid name. They cited the holotype of *M. robustus* as the specimen *Delprete et al. 7316* at EAC. The EAC specimen is mounted on two sheets, both with Accession No. 28985. According to Art. 8.3 of the *Code* (Turland et al. 2018), the two sheets are treated as a single specimen with multiple preparations.

Salas et al. (2015) concluded that *Spermacoce polygonifolia* A.St.Hil. is synonymous with *Mitracarpus robustus* E.B.Souza & E.L.Cabral. Because Saint Hilaire's name has nomenclatural priority, they published the new combination *Mitracarpus polygonifolius* (A.St.Hil.) R.M.Salas & E.B.Souza, which is the name to be used for this species.

Saint-Hilaire (1833: 381-383) published *Spermacoce polygonifolia* with a detailed description and comparison with several species of *Borreria* and *Spermacoce*, but did not cite any material examined or the herbarium of deposit. Two original specimens annotated as *S. polygonifolia* by Saint-Hilaire are found at P and MPU. Salas et al. (2015) cited the specimen at P as the holotype. However, because Saint-Hilaire did not cite any specimen or herbarium of deposit, the P specimen cannot be treated as the holotype. Also, Salas et al.'s citation cannot be treated as an inadvertent lectotypification, because, according to the Code, it should have been accompanied by "here designated" or a similar expression. Hence the specimen at P, with barcode P03818586, is here designated the lectotype of this name.
FGT, vol. 40(2), p. 697:
“35-8. *Mitracarpus schininianus* E.L. Cabral, W.A. Medina & E.B. Souza, Candollea 64: 154. 2009 – *Mitracarpus frigidus* var. *glaberrimus* Chodat & Hassl., Bull. Herb. Boissier ser. 2, 4: 191. 1904. Tipo: Paraguai, Canindeyú, “Iter ad Yerbales, montium Sierra de Maracayú”, s.d. (fl, fr), *E. Hassler 5027* (lectótipo, G, selecionado por Cabral et al., 2009).”

**Accepted name:** *Mitracarpus schininianus*
E.L.Cabral, W.A.Medina & E.B.Souza

Type: PARAGUAY. Canindeyú: “In sylva Ipé hú Sierra Maracayú”, Nov. 1898-1899 (fl, fr), *E. Hassler 5027* (G [G001166094], lectotype designated by Cabral et al. (2009: 154); isolectotypes G [2 sheets, barcodes G001166095, G001166096]).

FGT, vol. 40(2), p. 699:
“35-9. *Mitracarpus steyermarkii* E.L. Cabral & Bacigalupo, Acta Bot. Bras. 11(1): 50. 1997. Tipo: Brasil, Bahia, Barreiras, 7 km S of Rio Piau, ca. 150 km SW of Barreiras, 850 m, 13/IV/1966 (fl), *H.S. Irwin, J.W. Grear Jr., R. Souza & R. Reis dos Santos 14690* (holótipo K; isótipo NY).”

**Accepted name:** *Mitracarpus steyermarkii*
E.L.Cabral & Bacigalupo

Type: BRAZIL. Bahia: Barreiras, 7 km S of Rio Piau, ca. 150 km SW of Barreiras, 850 m, 13 Apr. 1966 (fl), *H.S. Irwin, J.W. Grear Jr., R. Souza & R. Reis dos Santos 14690* (holotype K [barcode K000016261]; isotypes CTES [barcode CTES0013507], F [Acc. No. 1784171], NY [barcode 00132260]).

37. **OLDENLANDIA** L., Sp. Pl. 119. 1753.

FGT, vol. 40(2), p. 712:
“37-1. *Oldenlandia corymbosa* L., Sp. Pl. 119. 1753. - *Hedyotis corymbosa* (L.) Lam., Tabl. Encycl. 1: 272. 1792. - Tipo: Não citado (holótipo, LINN).”

**Accepted name:** *Oldenlandia corymbosa* L.

Type: [icon] Plumier, Nov. Pl. Amer. tab. 36. 1703 (lower portion of Tab. 36, fruits and flowers of “Oldenlandia”), lectotype designated by Verdcourt (1976: 308).

**Notes:** Linnaeus (1753: 119) published *Oldenlandia corymbosa* L. and added the annotation “Oldenlandia pedunculis multifloris, foliis lineari-lanceolatis. + Oldenlandia humilis hyssopifolia. *Plum. gen. 42. Erhet. pict. t. 4. f. I.* - *Habitat in America meridionali.*”

Verdcourt (1976: 308) designated as lectotype the fruits and flowers of “Oldenlandia” depicted on the lower portion of Table 36 of Plumier (1703). According to The Linnaean Plant Name Typification Project (https://data.nhm.ac.uk/dataset/the-linnaean-plant-name-typification-project) “Verdcourt’s choice of type appears to be the earliest.”

FGT, vol. 40(2), p. 716:
“37-2. *Oldenlandia lancifolia* (Schumach.) DC., Prodr. 4: 425. 1830. - *Hedyotis lancifolia* Schumach. in Schumach. & Thonn, Beskr. Guin. Pl. 92. 1828. - *Oldenlandia herbacea* sensu Bremek. in Pulle, Fl. Surinam 4: 126. 1934. Tipo: Africa, Gana, Valley of Aquapim, *Thonning 210* (lectótipo C, selecionado por Verdcourt, 1976, isolectótipo, S n.v.).”

**Accepted name:** *Oldenlandia lancifolia* (Schumach.) DC.

Type: GHANA: Valley of Aquapim, *P. Thonning 210* (C [barcode C10003933], lectotype designated by Verdcourt (1976: 292); isolectotype, S [Acc. No. S-G-3012]; possible isolectotype C [barcode C10003937, “legit Thonning in Guinea”, without collection number]).

FGT, vol. 40(2), p. 717:
“37-3. *Oldenlandia salzmannii* (DC.) Hook. f. in Benth. & Hook. f., Gen. Pl. 2: 58. 1873. - *Anotis salzmannii* DC., Prodr. 4: 433. 1830. - *Hedyotis salzmannii* (DC.) Steud., Nom. Bot., ed. 2, 1: 728. 1841. - Tipo: Brasil, Bahia, s.d., *Salzmann s.n.* (holótipo G-DC).”

**Accepted name:** *Oldenlandia salzmannii* (DC.) Hook. f.

Type: BRAZIL. Bahia: “circà Bahiam frequentes” [near Salvador], s.d. [1827-1831], *P. Salzmann s.n.* (holotype G-DC [barcode G00666349]; isotypes E [barcode E00500000], HAL [2 sheets, barcodes HAL0114263, HAL0114264], K [barcode K000015898], MO [Acc. No. 4213062], MPU [barcode MPU021483]).
FGT, vol. 40(2), p. 697:
“35-8. *Mitracarpus schininianus* E.L. Cabral, W.A. Medina & E.B. Souza, Candollea 64: 154. 2009 – *Mitracarpus frigidus* var. *glaberrimus* Chodat & Hassl., Bull. Herb. Boissier ser. 2, 4: 191. 1904. Tipo: Paraguai, Canindeyú, “Iter ad Yerbales, montium Sierra de Maracayú”, s.d. (fl, fr), *E. Hassler 5027* (lectótipo, G, selecionado por Cabral et al., 2009).”

**Accepted name:** *Mitracarpus schininianus*
E.L.Cabral, W.A.Medina & E.B.Souza

Type: PARAGUAY. Canindeyú: “In sylva Ipé hú Sierra Maracayú”, Nov. 1898-1899 (fl, fr), *E. Hassler 5027* (G [G001166094], lectotype designated by Cabral et al. (2009: 154); isolectotypes G [2 sheets, barcodes G001166095, G001166096]).

FGT, vol. 40(2), p. 699:
“35-9. *Mitracarpus steyermarkii* E.L. Cabral & Bacigalupo, Acta Bot. Bras. 11(1): 50. 1997. Tipo: Brasil, Bahia, Barreiras, 7 km S of Rio Piau, ca. 150 km SW of Barreiras, 850 m, 13/IV/1966 (fl), *H.S. Irwin, J.W. Grear Jr., R. Souza & R. Reis dos Santos 14690* (holótipo K; isótipo NY).”

**Accepted name:** *Mitracarpus steyermarkii*
E.L.Cabral & Bacigalupo

Type: BRAZIL. Bahia: Barreiras, 7 km S of Rio Piau, ca. 150 km SW of Barreiras, 850 m, 13 Apr. 1966 (fl), *H.S. Irwin, J.W. Grear Jr., R. Souza & R. Reis dos Santos 14690* (holotype K [barcode K000016261]; isotypes CTES [barcode CTES0013507], F [Acc. No. 1784171], NY [barcode 00132260]).

37. **OLDENLANDIA** L., Sp. Pl. 119. 1753.

FGT, vol. 40(2), p. 712:
“37-1. *Oldenlandia corymbosa* L., Sp. Pl. 119. 1753. - *Hedyotis corymbosa* (L.) Lam., Tabl. Encycl. 1: 272. 1792. - Tipo: Não citado (holótipo, LINN).”

**Accepted name:** *Oldenlandia corymbosa* L.

Type: [icon] Plumier, Nov. Pl. Amer. tab. 36. 1703 (lower portion of Tab. 36, fruits and flowers of “Oldenlandia”), lectotype designated by Verdcourt (1976: 308).

**Notes:** Linnaeus (1753: 119) published *Oldenlandia corymbosa* L. and added the annotation “Oldenlandia pedunculis multifloris, foliis lineari-lanceolatis. + Oldenlandia humilis hyssopifolia. *Plum. gen. 42. Erhet. pict. t. 4. f. I.* - *Habitat in America meridionali.*”

Verdcourt (1976: 308) designated as lectotype the fruits and flowers of “Oldenlandia” depicted on the lower portion of Table 36 of Plumier (1703). According to The Linnaean Plant Name Typification Project (https://data.nhm.ac.uk/dataset/the-linnaean-plant-name-typification-project) “Verdcourt’s choice of type appears to be the earliest.”

FGT, vol. 40(2), p. 716:
“37-2. *Oldenlandia lancifolia* (Schumach.) DC., Prodr. 4: 425. 1830. - *Hedyotis lancifolia* Schumach. in Schumach. & Thonn, Beskr. Guin. Pl. 92. 1828. - *Oldenlandia herbacea* sensu Bremek. in Pulle, Fl. Surinam 4: 126. 1934. Tipo: Africa, Gana, Valley of Aquapim, *Thonning 210* (lectótipo C, selecionado por Verdcourt, 1976, isolectótipo, S n.v.).”

**Accepted name:** *Oldenlandia lancifolia* (Schumach.) DC.

Type: GHANA: Valley of Aquapim, *P. Thonning 210* (C [barcode C10003933], lectotype designated by Verdcourt (1976: 292); isolectotype, S [Acc. No. S-G-3012]; possible isolectotype C [barcode C10003937, “legit Thonning in Guinea”, without collection number]).

FGT, vol. 40(2), p. 717:
“37-3. *Oldenlandia salzmannii* (DC.) Hook. f. in Benth. & Hook. f., Gen. Pl. 2: 58. 1873. - *Anotis salzmannii* DC., Prodr. 4: 433. 1830. - *Hedyotis salzmannii* (DC.) Steud., Nom. Bot., ed. 2, 1: 728. 1841. - Tipo: Brasil, Bahia, s.d., *Salzmann s.n.* (holótipo G-DC).”

**Accepted name:** *Oldenlandia salzmannii* (DC.) Hook. f.

Type: BRAZIL. Bahia: “circà Bahiam frequentes” [near Salvador], s.d. [1827-1831], *P. Salzmann s.n.* (holotype G-DC [barcode G00666349]; isotypes E [barcode E00500000], HAL [2 sheets, barcodes HAL0114263, HAL0114264], K [barcode K000015898], MO [Acc. No. 4213062], MPU [barcode MPU021483]).
**Note:** Bentham (1857: 109) cited the material studied of *Pagamea plicata* Spruce ex Benth. a gathering collected by Richard Spruce near São Gabriel da Cachoeira, Rio Negro, Amazonas, Brazil, without indicating the collection number or the herbarium of deposit. Bentham regularly studied specimens in both K and BM herbaria. The original gathering corresponds to *Spruce 2342*, which has duplicates in numerous herbaria. Delprete (2010b: 729) cited the BM specimen as the holotype. As Delprete's citation was published after 1 January 2001, according to the *Code* it cannot be interpreted as an inadvertent lectotypification. The K specimen with the stamp "Herbarium Benthamianum" and barcode K000265553, is here designated the lectotype of this name.

FGT, vol. 40(2), p. 732:

"39-2. *Pagamea guianensis* Aubl., Hist. Pl. Guiane 1: 113, fig. 44. 1775. - Tipo: Guiana Francesa, "prope e supra montem Serpent et a l'abitacion appellée Gallion", s.d., *J.B.C.F. Aublet s.n.* (holótipo BM, foto em NY)."

**Accepted name:** *Pagamea guianensis* Aubl.

**Type:** FRENCH GUIANA: Montagne Serpent [a series of hills parallel to the road between the town of Gallion and the bridge on the Comté River], "habitat propè & supra montem *Serpent* dictum" and "cet arbrisseau étoit en fleur et en fruit dans le mois d'Aout. [...] au sommet de la montagne Serpent & à l'abitation appellée Gallion", s.d. [1762-1764], *J.B.C.F. Aublet s.n.* (P-JJR 4: 143), lectotype designated by Lanjouw and Uittien (1940: 155); isolectotypes BM [barcode BM000624228], LINN-SM [No. 340.8]).

**Notes:** For additional information regarding the typification of *Pagamea guianensis* Aubl. see Delprete (2015).

40. **PALICOUREA Aubl.**, Hist. Pl. Guiane 172. 1775.

FGT, vol. 40(2), p. 739:

"40-1. *Palicourea coriacea* (Cham.) K. Schum. in Engl. & Prantl, Nat. Pflanzenfam. 4(4): 115. 1891. - *Patabea coriacea* Cham., Linnaea 9: 234. 1834. - Tipo: Brasil, sem localidade, s.d., *Sellow s.n.* (holótipo, B, destruído; foto em NY)."

**Accepted name:** *Palicourea coriacea* (Cham.) K.Schum.

**Type:** BRAZIL. Goiás: Mun. Cocalzinho, Parque Estadual dos Pireneus, trilha que segue do portal da Fazenda Capitão do Mato em direção do Morro do Cabeludo, vegetação de cerrado rupestre com áreas de campos encharcados, arbustinho multicaule, com geopódio basal lenhoso, caule de 40-60 cm, ráquis e brácteas amarelo vivo, 15°48'23-24"S, 48°49'20-29"W, alt. 1150 m, 19 Nov. 2006 (fl), *Delprete et al.* 9947 (RB [Acc. No. 494074, barcode 00584026], **neotype here designated**; isoneotypes NY [barcode 01146630], SPF [barcode SPF 189601], UFG [Acc. No. 43349]).

**Notes:** Chamisso (1834: 234-235) cited the material studied of *Patabea coriacea* Cham. as "E Brasilia misit Sellowius." The original material at B was destroyed during WWII. No other specimens associated with this name, collected by Sellow, could be found in any herbarium. Therefore, a neotype needs to be designated. The specimen *Delprete et al.* 9947 at RB is here designated the neotype of *Patabea coriacea* Cham. [= *Palicourea coriacea* (Cham.) K.Schum.], and the duplicates of this gathering, present in other herbaria, are isoneotypes.

FGT, vol. 40(2), p. 751:

"40-2. *Palicourea crocea* (Sw.) Roem. & Schult., Syst. Veg. 5: 193. 1819. - *Psychotria crocea* Sw., Prodr. Veg. Ind. Occ. 44. 1788. - Tipo: "Jamaica et Hispaniola", s.d., *Swartz s.n.* (holótipo, S n.v.)."

**Accepted name:** *Palicourea crocea* (Sw.) Roem. & Schult.

**Type:** JAMAICA. Without locality, s.d., O. *Swartz s.n.* (S [Acc. No. S-R-5316], **lectotype here designated**; isolectotypes LD [Acc. No. 1255937], M [barcode M-0198314], S [Acc. No. S-R-5317], SBT [barcode SBT13385]).

**Notes:** Olof Swartz (1788: 44) described *Psychotria crocea* Sw. as "[...] panicula erecta terminali crocea, pedunculis coloratis, Psychotrophium 2. Brown. jam. 160. t. 13. f. 2, I. *Jamaica, Hispaniola*" without citing any specimen. He wrongly cited the figure in Browne's publication. The plate associated with this name is Table 13, Figure 1, and not Figure 2 as cited by Swartz.

Under *Psychotria crocea*, Swartz cited "Psychotrophum 2", which was described and illustrated in Patrick Browne's (1756) *Civil and Natural History of Jamaica*. Browne (1756: 169) published the polynomial "Psychotrophum 2. Fructiculosum, foliis amplioribus ovatis,
**Note:** Bentham (1857: 109) cited the material studied of *Pagamea plicata* Spruce ex Benth. a gathering collected by Richard Spruce near São Gabriel da Cachoeira, Rio Negro, Amazonas, Brazil, without indicating the collection number or the herbarium of deposit. Bentham regularly studied specimens in both K and BM herbaria. The original gathering corresponds to *Spruce 2342*, which has duplicates in numerous herbaria. Delprete (2010b: 729) cited the BM specimen as the holotype. As Delprete's citation was published after 1 January 2001, according to the *Code* it cannot be interpreted as an inadvertent lectotypification. The K specimen with the stamp "Herbarium Benthamianum" and barcode K000265553, is here designated the lectotype of this name.

FGT, vol. 40(2), p. 732:

"39-2. *Pagamea guianensis* Aubl., Hist. Pl. Guiane 1: 113, fig. 44. 1775. - Tipo: Guiana Francesa, "prope e supra montem Serpent et a l'abitacion appellée Gallion", s.d., *J.B.C.F. Aublet s.n.* (holótipo BM, foto em NY)."

**Accepted name:** *Pagamea guianensis* Aubl.

**Type:** FRENCH GUIANA: Montagne Serpent [a series of hills parallel to the road between the town of Gallion and the bridge on the Comté River], "habitat propè & supra montem *Serpent* dictum" and "cet arbrisseau étoit en fleur et en fruit dans le mois d'Aout. [...] au sommet de la montagne Serpent & à l'abitation appellée Gallion", s.d. [1762-1764], *J.B.C.F. Aublet s.n.* (P-JJR 4: 143), lectotype designated by Lanjouw and Uittien (1940: 155); isolectotypes BM [barcode BM000624228], LINN-SM [No. 340.8]).

**Notes:** For additional information regarding the typification of *Pagamea guianensis* Aubl. see Delprete (2015).

40. **PALICOUREA Aubl.**, Hist. Pl. Guiane 172. 1775.

FGT, vol. 40(2), p. 739:

"40-1. *Palicourea coriacea* (Cham.) K. Schum. in Engl. & Prantl, Nat. Pflanzenfam. 4(4): 115. 1891. - *Patabea coriacea* Cham., Linnaea 9: 234. 1834. - Tipo: Brasil, sem localidade, s.d., *Sellow s.n.* (holótipo, B, destruído; foto em NY)."

**Accepted name:** *Palicourea coriacea* (Cham.) K.Schum.

**Type:** BRAZIL. Goiás: Mun. Cocalzinho, Parque Estadual dos Pireneus, trilha que segue do portal da Fazenda Capitão do Mato em direção do Morro do Cabeludo, vegetação de cerrado rupestre com áreas de campos encharcados, arbustinho multicaule, com geopódio basal lenhoso, caule de 40-60 cm, ráquis e brácteas amarelo vivo, 15°48'23-24"S, 48°49'20-29"W, alt. 1150 m, 19 Nov. 2006 (fl), *Delprete et al.* 9947 (RB [Acc. No. 494074, barcode 00584026], **neotype here designated**; isoneotypes NY [barcode 01146630], SPF [barcode SPF 189601], UFG [Acc. No. 43349]).

**Notes:** Chamisso (1834: 234-235) cited the material studied of *Patabea coriacea* Cham. as "E Brasilia misit Sellowius." The original material at B was destroyed during WWII. No other specimens associated with this name, collected by Sellow, could be found in any herbarium. Therefore, a neotype needs to be designated. The specimen *Delprete et al.* 9947 at RB is here designated the neotype of *Patabea coriacea* Cham. [= *Palicourea coriacea* (Cham.) K.Schum.], and the duplicates of this gathering, present in other herbaria, are isoneotypes.

FGT, vol. 40(2), p. 751:

"40-2. *Palicourea crocea* (Sw.) Roem. & Schult., Syst. Veg. 5: 193. 1819. - *Psychotria crocea* Sw., Prodr. Veg. Ind. Occ. 44. 1788. - Tipo: "Jamaica et Hispaniola", s.d., *Swartz s.n.* (holótipo, S n.v.)."

**Accepted name:** *Palicourea crocea* (Sw.) Roem. & Schult.

**Type:** JAMAICA. Without locality, s.d., O. *Swartz s.n.* (S [Acc. No. S-R-5316], **lectotype here designated**; isolectotypes LD [Acc. No. 1255937], M [barcode M-0198314], S [Acc. No. S-R-5317], SBT [barcode SBT13385]).

**Notes:** Olof Swartz (1788: 44) described *Psychotria crocea* Sw. as "[...] panicula erecta terminali crocea, pedunculis coloratis, Psychotrophium 2. Brown. jam. 160. t. 13. f. 2, I. *Jamaica, Hispaniola*" without citing any specimen. He wrongly cited the figure in Browne's publication. The plate associated with this name is Table 13, Figure 1, and not Figure 2 as cited by Swartz.

Under *Psychotria crocea*, Swartz cited "Psychotrophum 2", which was described and illustrated in Patrick Browne's (1756) *Civil and Natural History of Jamaica*. Browne (1756: 169) published the polynomial "Psychotrophum 2. Fructiculosum, foliis amplioribus ovatis,
stipulosis rigidis interpostis, ramulis crassioribus, racemis umbellatis, sustentaculis ternato-ternatis. Tab. 17. f. 2" However, his Psychotrophum 2 is on Table 13, figure 1 (on Table 17, Figure 2 is depicted a species of *Chione*). To this species he assigned the English vernacular name "The smaller succulent Psychotrophum", which was followed by a detailed description of the species. On Table 13, figure 1 of Browne's publication is depicted an inflorescence with flower buds, flowers in anthesis, and fruits. At the bottom of the branch are drawn a flower bud, a flower in anthesis, a dissected corolla, several representations of the hypanthium and style, and a mature fruit crowned by the persistent calyx. This drawing is original material of *Psychotria crocea*.

According to Stafleu and Cowan (1986: 116), the main set of Swartz's herbarium is at S, and the "West-Indian collections (1748-1786) at S, are not complete". They also indicated that additional Swartz specimens can be found in numerous herbaria.

Taylor (1989: 31) and Andersson (1992: 157) wrote that the holotype of *Psychotria crocea* [= *Palicourea crocea* (Sw.) Roem. & Schult.] is at BM. Taylor (1999a: 154) cited the type of this name as "Type: Jamaica, without locality, without date, O. Swartz s.n. (S holotype, photo JBSD)." Then, Taylor (2012b: 170) reverted her citation, and wrote that holotype of this name is at BM. After exhaustive search, no specimen collected by Swartz associated with this name was found in BM.

Delprete (2010b: 751) cited the holotype of this name as "holótipo, S n.v." This citation can not be interpreted as inadvertent lectotypification, because it is not accompanied by "here designated" or a similar expression. Searching for possible original material associated with this name, there are two specimens at S and specimens at SBT, M and LD collected by Swartz that should be considered as possible original material and are discussed below.

Two specimens associated with this name and collected by Swartz are present in S. The specimen with Accession No. S-R-5316 has the annotation "Ind. Occ. Swartz" handwritten by an unknown author on the bottom left corner of the sheet. The sheet also has the annotations "Palicourea crocea De Cand., Psychotria crocea Sw." handwritten by an unknown author on the bottom right corner. This specimen consists of two branches with leaves and terminal inflorescences with flower buds, a small branch with a terminal infructescence with mature fruits, and the distal portion of an inflorescence with flower buds. Considering the ample material present on this sheet, this specimen, with Accession No. S-R-5316, collected by Swartz in Jamaica, is here designated the lectotype of *Psychotria crocea* Sw.

The other S specimen, with Accession No. S-R-5317, is mounted on a smaller sheet glued onto a larger supporting sheet. On the smaller sheet is affixed a single branch with a fastigate inflorescence with the corollas fallen off, and a loose fruit. On the smaller sheet, there is the handwritten annotation "Psychotria crocea Sw! Palicourea crocea DC. - Jamaica: Ol. Swartz!" The annotation "Ol. Swartz!" means that this specimen was collected and studied by Swartz. On the bottom of the larger sheet are affixed two labels with the annotations "Psychotria crocea Swartz, Prodr. 44. 1788. [...] det. C.M. Taylor XI 1988" and "= Palicourea crocea (Sw.) R. & S. - det. C.M. Taylor XI 1988." This specimen is an isolectotype.

The SBT specimen, with barcode SBT13385, consists of a branch with numerous leaves and a terminal inflorescence with flower buds and flowers in anthesis. On the back of the sheet is handwritten in ink "Jamaica, Swartz. Palicourea crocea De Cand., 6 Psychotria Swartz crocea." This specimen is an isolectotype.

The M specimen with barcode M-0198314 has a label with the upper heading "Herbarium Regium Monacense" and the lower heading "Herbar. Schreberianum." The label has the handwritten annotation "Psychotria crocea Sw., Palicourea crocea R. & Schult., (nec De Cand., qui cum aliis confundere), Jamaica, Ol. Swartz." The specimen, collected by Swartz in Jamaica, is composed of a single branch with numerous leaves and a terminal inflorescence with flower buds. This specimen is an isolectotype.

The LD specimen, with Acc. No. 1255937, has the stamp "Museum Acharianum". It has two annotations handwritten directly on the sheet "Jamaica. Swartz" and "P. crocea", both by different unknown authors, not Swartz. The specimen, collected by Swartz in Jamaica, consists of a single branch with several leaves, and a terminal infructescence with three fruits. This specimen is an isolectotype.

FGT, vol. 40(2), p. 760:

"40-3. *Palicourea guianensis* Aubl., Hist. Pl. Guiane 1: 173. 1775. - *Psychotria guianensis* (Aubl.) Rusby, Mem. Torrey Bot. Club 3: 48. 1893. - Tipo: Guiana Francesa, Forêts d'Orapu, s.d., *J.B.C.F. Aublet s. n.* (holótipo, P)."

**Accepted name:** *Palicourea guianensis* Aubl.

Type: FRENCH GUIANA: Kaw Mountains, "in sylvis territorii Caux" and "arbrisseau en fleur dans les forêts de Caux, au mois de Février", s.d. [Feb-Mar 1764], *J.B.C.F. Aublet s.n.* (P-JJR 8: 268, lecto-
stipulosis rigidis interpostis, ramulis crassioribus, racemis umbellatis, sustentaculis ternato-ternatis. Tab. 17. f. 2" However, his Psychotrophum 2 is on Table 13, figure 1 (on Table 17, Figure 2 is depicted a species of *Chione*). To this species he assigned the English vernacular name "The smaller succulent Psychotrophum", which was followed by a detailed description of the species. On Table 13, figure 1 of Browne's publication is depicted an inflorescence with flower buds, flowers in anthesis, and fruits. At the bottom of the branch are drawn a flower bud, a flower in anthesis, a dissected corolla, several representations of the hypanthium and style, and a mature fruit crowned by the persistent calyx. This drawing is original material of *Psychotria crocea*.

According to Stafleu and Cowan (1986: 116), the main set of Swartz's herbarium is at S, and the "West-Indian collections (1748-1786) at S, are not complete". They also indicated that additional Swartz specimens can be found in numerous herbaria.

Taylor (1989: 31) and Andersson (1992: 157) wrote that the holotype of *Psychotria crocea* [= *Palicourea crocea* (Sw.) Roem. & Schult.] is at BM. Taylor (1999a: 154) cited the type of this name as "Type: Jamaica, without locality, without date, O. Swartz s.n. (S holotype, photo JBSD)." Then, Taylor (2012b: 170) reverted her citation, and wrote that holotype of this name is at BM. After exhaustive search, no specimen collected by Swartz associated with this name was found in BM.

Delprete (2010b: 751) cited the holotype of this name as "holótipo, S n.v." This citation can not be interpreted as inadvertent lectotypification, because it is not accompanied by "here designated" or a similar expression. Searching for possible original material associated with this name, there are two specimens at S and specimens at SBT, M and LD collected by Swartz that should be considered as possible original material and are discussed below.

Two specimens associated with this name and collected by Swartz are present in S. The specimen with Accession No. S-R-5316 has the annotation "Ind. Occ. Swartz" handwritten by an unknown author on the bottom left corner of the sheet. The sheet also has the annotations "Palicourea crocea De Cand., Psychotria crocea Sw." handwritten by an unknown author on the bottom right corner. This specimen consists of two branches with leaves and terminal inflorescences with flower buds, a small branch with a terminal infructescence with mature fruits, and the distal portion of an inflorescence with flower buds. Considering the ample material present on this sheet, this specimen, with Accession No. S-R-5316, collected by Swartz in Jamaica, is here designated the lectotype of *Psychotria crocea* Sw.

The other S specimen, with Accession No. S-R-5317, is mounted on a smaller sheet glued onto a larger supporting sheet. On the smaller sheet is affixed a single branch with a fastigate inflorescence with the corollas fallen off, and a loose fruit. On the smaller sheet, there is the handwritten annotation "Psychotria crocea Sw! Palicourea crocea DC. - Jamaica: Ol. Swartz!" The annotation "Ol. Swartz!" means that this specimen was collected and studied by Swartz. On the bottom of the larger sheet are affixed two labels with the annotations "Psychotria crocea Swartz, Prodr. 44. 1788. [...] det. C.M. Taylor XI 1988" and "= Palicourea crocea (Sw.) R. & S. - det. C.M. Taylor XI 1988." This specimen is an isolectotype.

The SBT specimen, with barcode SBT13385, consists of a branch with numerous leaves and a terminal inflorescence with flower buds and flowers in anthesis. On the back of the sheet is handwritten in ink "Jamaica, Swartz. Palicourea crocea De Cand., 6 Psychotria Swartz crocea." This specimen is an isolectotype.

The M specimen with barcode M-0198314 has a label with the upper heading "Herbarium Regium Monacense" and the lower heading "Herbar. Schreberianum." The label has the handwritten annotation "Psychotria crocea Sw., Palicourea crocea R. & Schult., (nec De Cand., qui cum aliis confundere), Jamaica, Ol. Swartz." The specimen, collected by Swartz in Jamaica, is composed of a single branch with numerous leaves and a terminal inflorescence with flower buds. This specimen is an isolectotype.

The LD specimen, with Acc. No. 1255937, has the stamp "Museum Acharianum". It has two annotations handwritten directly on the sheet "Jamaica. Swartz" and "P. crocea", both by different unknown authors, not Swartz. The specimen, collected by Swartz in Jamaica, consists of a single branch with several leaves, and a terminal infructescence with three fruits. This specimen is an isolectotype.

FGT, vol. 40(2), p. 760:

"40-3. *Palicourea guianensis* Aubl., Hist. Pl. Guiane 1: 173. 1775. - *Psychotria guianensis* (Aubl.) Rusby, Mem. Torrey Bot. Club 3: 48. 1893. - Tipo: Guiana Francesa, Forêts d'Orapu, s.d., *J.B.C.F. Aublet s. n.* (holótipo, P)."

**Accepted name:** *Palicourea guianensis* Aubl.

Type: FRENCH GUIANA: Kaw Mountains, "in sylvis territorii Caux" and "arbrisseau en fleur dans les forêts de Caux, au mois de Février", s.d. [Feb-Mar 1764], *J.B.C.F. Aublet s.n.* (P-JJR 8: 268, lecto-
type designated by Lanjouw and Uittien (1940: 155); isolectotype BM [barcode BM000058138]).

**Notes:** For additional information regarding the typification of *Palicourea guianensis* Aubl., see Delprete (2015).

FGT, vol. 40(2), p. 765: "40-4. *Palicourea macrobotrys* (Ruiz & Pav.) Roem. & Schult., Syst. Veg. 5: 194. 1819. - *Psychotria macrobotrys* Ruiz & Pav., Fl. Peruv. 2: 57, pl. 203, fig. a. 1799. Tipo: Perú, Huánuco, "in Andium nemoribus ad Cuchero et Chinchao", s.d., *Tafalla & Manzanilla* [in Herb. Ruiz & Pavón] s.n. (holótipo, MA; isótipo, MPU)."

**Accepted name:** *Palicourea macrobotrys* (Ruiz & Pav.) DC.

**Correction of authority:** *Palicourea macrobotrys* (Ruiz & Pav.) DC., Prodr. 4: 527. 1830. - "*Palicourea macrobotrys* (Ruiz & Pav.) Roem. & Schult.", Syst. Veg. 5: 194. 1819, name cited as a *Psychotria* species.

**Type:** PERU. Huánuco. [Protologue: "Habitat in *Andium* nemoribus imis ad *Cuchero* et *Chinchao* tractus"], "Cuchero", s.d. *J.J. Tafalla & J.A. Manzanilla* [in *Herb. H. Ruiz López & J.A. Pavón y Jiménez*] s.n. (MA [barcode MA 815923], *lectotype here designated*; isolectotypes K [barcode K000471538], MA [2 sheets, barcodes MA 815921, MA 815922], MPU [barcode MPU022082], drawing by Isidro Gálvez in Herbarium Archives of the Royal Botanical Exposition to the Viceroyalty of Peru [Acc. No. AJB04-D-0415_001]; isolectotype fragment F [Accession No. 591760]).

**Notes:** In the protologue of *Psychotria macrobotrys* Ruiz & Pav., Ruiz and Pavón (1799: 57) cited the collection locality as "Habitat in *Andium* nemoribus imis ad *Cuchero* et *Chinchao* tractus." Taylor (1999a: 232) cited the type of *P. macrobotrys* as "Habitat in Andium nemoribus ad Cuchero et Chinchao, Jul.-Aug., Ruiz and Pavón s.n. (MA holotype, n.v.)." Because there are several original specimens at MA, Taylor's type citation cannot be treated as a lectotypification. According to the *Code* (Turland et al., 2018), original material is "The set of specimens and illustrations from which a lectotype may be chosen (Art. 9.4, Notes 2 and 3, Art. F. 3.9, and Note 2 for details), or the holotype (Art. 9.1)." At MA, there are one original drawing and several original specimens, which are described and discussed below.

The original drawing of *Psychotria macrobotrys* made by Isidro Gálvez is preserved in the Herbarium Archives of the Royal Botanical Expedition to the Vice-royalty of Peru made by Hipólito Ruiz & José Pavón. On the colored drawing, two species of *Psychotria* are represented. On the left side is depicted a branch with an inflorescence with flowers and fruits of *Psychotria macrobotrys*, on the lower portion are drawn a dissected corolla, a stamen, hypanthium and style, a fruit, and two pyrenes. On the right side of the drawing is depicted a branch with an inflorescence with flowers and fruits of *Psychotria amethystina*, and just below are drawn a dissected corolla, a stamen, a hypanthium and style, a fruit, and two pyrenes. On the upper margin of the drawing is the handwritten number "203". Above the drawing is a label with the heading "REAL JARDÍN BOTÁNICO, CSIC - ARCHIVO, Real Expedición Botánica al Virreinado del Perú, AJB04-D-0415_001."

The MA specimen with barcode MA815921 has two labels. One label has the annotation "Palicourea macrobotrys D.C., Psychotria macrobotrys Fl. Per." handwritten by an unknown author. Just above that label is affixed another label, with the annotation "Psychotria macrobotrys Fl. P." handwritten by Hipólito Ruiz. On the sheet, are mounted two branches with several leaves and terminal inflorescences with flowers and young fruits.

The MA specimen with barcode MA815922 has a label with the annotation "Psychotria lutea, Cochero insi locis [?] et Julio, Psychotria macrobotrys Fl. P. vol. 2. p. 57. t. 203.f.a." handwritten by Hipólito Ruiz. On the sheet are mounted two branches with several leaves and terminal inflorescences with flowers and young fruits.

On the MA specimen with barcode MA815923, is glued a small sheet of paper with a detailed description of *Psychotria macrobotrys*, handwritten by Joseph Dombey, which is very similar to the description published by Ruiz and Pavón (1799: 57). Above that label is another label with the annotation "Psychotria macrobotrys Sp. Pl. Fl. Per. de Cuchero" handwritten by Hipólito Ruiz. On the sheet, are mounted two branches with several leaves and terminal inflorescences with flowers and young fruits. This specimen is here designated the lectotype of this name.

At K there is a sheet with barcode K000471538, and a small handwritten label (author unknown) with the annotation "Psychotria, Ex Herb de R et P., Lima." On the lower margin is handwritten "Palicourea macrobotrys (R. & P.) R. & S., PCS [Paul Carpenter Standley]". On the sheet, is mounted a branch with several leaves and a terminal inflorescence with numerous flower buds.

At MPU there is a sheet with barcode MPU022082 and with a handwritten label (author unknown) saying
"Psychotria macrobotrys Fl. Per., Pérou. dedit Pavon." On the lower right corner is a second label with the heading "Institut de Botanique de Montpellier" and the handwritten annotation "Palicourea macrobotrys Roem & Sch., Psychotria ______ Ruiz & Pav." On the sheet is mounted a branch with several leaves and a terminal infructescence.

At F there is a sheet, Accession No. 591760, on which is mounted an envelope containing a few flower buds and a leaf fragment. On the envelope is typewritten "Palicourea macrobotrys (R. & P.) R. & S., Peru, Ruiz & Pavón."

FGT, vol. 40(2), p. 769:
"40-5. **Palicourea marcgravii** A. St. Hil., Pl. Rem. Brés. 231, tab. 22, fig. A. 1824. - *Psychotria marcgravii* (A. St. Hil.) Spreng., Syst. Cur. Post. 79. 1827. - Tipo: Brasil, "cucilli entre Rossinha da Negra e Varge, route de Rio de Janeiro à Minas Gerais", s.d. (fl), *A. Saint-Hilaire 2209* (holótipo, P)."

**Accepted name:** *Palicourea marcgravii* A.St.Hil.

Type: [icon] Saint-Hilaire, Hist. Pl. Remarq. Bresil, Tab. 22, fig. A. 1824, **lectotype here designated.**

**Notes:** Augustin de Saint-Hilaire (1824: 231-232) provided a detailed description of *Palicourea marcgravii* A. St. Hil. and, among many other characters, stated that is a shrub 5-6 feet tall, with opposite leaves that are puberulous below when young or glabrous below when mature, terminal multiflorous cymes with puberulous rachis, corollas 5-7 lines [10.5-15 mm] long, gibbous at base, basally yellow and distally purple, 2-locular ovary, and ascending ovules. Above the description he cited "Galvania sp. 2da (Erva do rato) *Vell. Mss.* - Erva do rato *Marcg. Bras.* 60, fig. 2 (1). - *N. Vulg.* Erva do rato", and represented this species in Table 22, fig. A.

Delprete (2010b: 765) cited the type of *Palicourea marcgravii* as "A. Saint-Hilaire 2209* (holótipo, P).*" The specimen cited by Delprete, which now has barcode P00852566, cannot be treated as a holotype because Saint-Hilaire in the protologue did not cite any of his own collections.

Saint-Hilaire's (1824) Table 22, Figure A, of *Histoire des plantes les plus remarquables du Bresil et du Paraguay* well represents this species and is consistent with Saint-Hilaire's accurate description. Aside from a branch with numerous leaf pairs and two terminal inflorescences, at the base of the plate are depicted a longitudinally dissected corolla, an anther, a style with

two short lobes, and a hypanthium topped by the calyx and exceeding nectariferous disk. Hence, Saint-Hilaire's Table 22, fig. A is here designated the lectotype of *Palicourea marcgravii*.

FGT, vol. 40(2), p. 769:

**Synonym:**

*Palicourea marcgravii* A. St. Hil. var. *β, pubescens*
A.St.Hil., Pl. Rem. Brés. 232. 1824.

Type: [icon] Saint-Hilaire, Hist. Pl. Remarq. Bresil, Tab. 22, fig. A. 1824, **lectotype here designated.**

**Notes:** Saint-Hilaire (1824: 232) described *Palicourea marcgravii* var. *pubescens* A. St. Hil. as "V. *β, pubescens*; foliis subtùs pubescentibus aut puberulis." and cited the areas of occurrence as "Frequens in provinciis Minas Geraes et Pernambuco (Marcg.), ad marigines sylvarum primaevarum et in sylvis caeduis. Floret Decembre-Martio." Because in the original publication he described the leaves of *P. marcgravii* as puberulous below when young or glabrous below when mature, the morphological characters of var. *pubescens* fit within the variation of the species, and Saint-Hilaire's Table 22, fig. A is here designated the lectotype of *P. marcgravii* var. *pubescens*.

FGT, vol. 40(2), p. 783:

"40-6. **Palicourea officinalis** Mart. in Spix & Mart., Reise Bras. 2: 544. 1828. - *Psychotria medica* Muell. Arg. in Mart., Fl. Bras. 6(5): 232. 1881, *nom. nov. superfl.* Tipo: Brasil, Minas Gerais, "in arenosis siccis campestribus deserti ad flumen S. Francisco", 1818 (fl), *C.F.P. Martius s.n.* (holótipo, M)."

**Accepted name:** *Palicourea officinalis* Mart.

Type: BRAZIL. Minas Gerais: "campis editis Serro Frio", s.d. (fl), *C.F.P. Martius s.n.* (holotype, M [barcode M-0198485]).

**Notes:** Martius in Spix and Martius (1828: 544) did not cite the collection locality of *Palicourea officinalis* Mart. or the herbarium of deposit. At M there is a specimen with barcode M-0198485 and a label specifying "Dr. Martius Iter Brasil." On the label is annotated the name *Palicourea officinalis*, a short description, and the locality "campis editis Serro Frio" handwritten by Martius. This specimen is the holotype of this name.
"Psychotria macrobotrys Fl. Per., Pérou. dedit Pavon." On the lower right corner is a second label with the heading "Institut de Botanique de Montpellier" and the handwritten annotation "Palicourea macrobotrys Roem & Sch., Psychotria ______ Ruiz & Pav." On the sheet is mounted a branch with several leaves and a terminal infructescence.

At F there is a sheet, Accession No. 591760, on which is mounted an envelope containing a few flower buds and a leaf fragment. On the envelope is typewritten "Palicourea macrobotrys (R. & P.) R. & S., Peru, Ruiz & Pavón."

FGT, vol. 40(2), p. 769:
"40-5. **Palicourea marcgravii** A. St. Hil., Pl. Rem. Brés. 231, tab. 22, fig. A. 1824. - *Psychotria marcgravii* (A. St. Hil.) Spreng., Syst. Cur. Post. 79. 1827. - Tipo: Brasil, "cucilli entre Rossinha da Negra e Varge, route de Rio de Janeiro à Minas Gerais", s.d. (fl), *A. Saint-Hilaire 2209* (holótipo, P)."

**Accepted name:** *Palicourea marcgravii* A.St.Hil.

Type: [icon] Saint-Hilaire, Hist. Pl. Remarq. Bresil, Tab. 22, fig. A. 1824, **lectotype here designated.**

**Notes:** Augustin de Saint-Hilaire (1824: 231-232) provided a detailed description of *Palicourea marcgravii* A. St. Hil. and, among many other characters, stated that is a shrub 5-6 feet tall, with opposite leaves that are puberulous below when young or glabrous below when mature, terminal multiflorous cymes with puberulous rachis, corollas 5-7 lines [10.5-15 mm] long, gibbous at base, basally yellow and distally purple, 2-locular ovary, and ascending ovules. Above the description he cited "Galvania sp. 2da (Erva do rato) Vell. Mss. - Erva do rato Marcg. Bras. 60, fig. 2 (1). - N. Vulg. Erva do rato", and represented this species in Table 22, fig. A.

Delprete (2010b: 765) cited the type of *Palicourea marcgravii* as "A. Saint-Hilaire 2209* (holótipo, P).*" The specimen cited by Delprete, which now has barcode P00852566, cannot be treated as a holotype because Saint-Hilaire in the protologue did not cite any of his own collections.

Saint-Hilaire's (1824) Table 22, Figure A, of *Histoire des plantes les plus remarquables du Bresil et du Paraguay* well represents this species and is consistent with Saint-Hilaire's accurate description. Aside from a branch with numerous leaf pairs and two terminal inflorescences, at the base of the plate are depicted a longitudinally dissected corolla, an anther, a style with

two short lobes, and a hypanthium topped by the calyx and exceeding nectariferous disk. Hence, Saint-Hilaire's Table 22, fig. A is here designated the lectotype of *Palicourea marcgravii*.

FGT, vol. 40(2), p. 769:

**Synonym:**

*Palicourea marcgravii* A. St. Hil. var. *β, pubescens*
A.St.Hil., Pl. Rem. Brés. 232. 1824.

Type: [icon] Saint-Hilaire, Hist. Pl. Remarq. Bresil, Tab. 22, fig. A. 1824, **lectotype here designated.**

**Notes:** Saint-Hilaire (1824: 232) described *Palicourea marcgravii* var. *pubescens* A. St. Hil. as "V. *β, pubescens*; foliis subtùs pubescentibus aut puberulis." and cited the areas of occurrence as "Frequens in provinciis Minas Geraes et Pernambuco (Marcg.), ad marigines sylvarum primaevarum et in sylvis caeduis. Floret Decembre-Martio." Because in the original publication he described the leaves of *P. marcgravii* as puberulous below when young or glabrous below when mature, the morphological characters of var. *pubescens* fit within the variation of the species, and Saint-Hilaire's Table 22, fig. A is here designated the lectotype of *P. marcgravii* var. *pubescens*.

FGT, vol. 40(2), p. 783:

"40-6. **Palicourea officinalis** Mart. in Spix & Mart., Reise Bras. 2: 544. 1828. - *Psychotria medica* Muell. Arg. in Mart., Fl. Bras. 6(5): 232. 1881, *nom. nov. superfl.* Tipo: Brasil, Minas Gerais, "in arenosis siccis campestribus deserti ad flumen S. Francisco", 1818 (fl), *C.F.P. Martius s.n.* (holótipo, M)."

**Accepted name:** *Palicourea officinalis* Mart.

Type: BRAZIL. Minas Gerais: "campis editis Serro Frio", s.d. (fl), *C.F.P. Martius s.n.* (holotype, M [barcode M-0198485]).

**Notes:** Martius in Spix and Martius (1828: 544) did not cite the collection locality of *Palicourea officinalis* Mart. or the herbarium of deposit. At M there is a specimen with barcode M-0198485 and a label specifying "Dr. Martius Iter Brasil." On the label is annotated the name *Palicourea officinalis*, a short description, and the locality "campis editis Serro Frio" handwritten by Martius. This specimen is the holotype of this name.
FGT, vol. 40(2), p. 794:
“40-7. *Palicourea rigida* Kunth in Humb. & Bonpl., Nov. Gen. Sp. 3: 370 (quarto ed.). 1819. - *Psychotria rigida* var. *genuina* Muell. Arg. in Mart., Fl. Bras. 6(5): 230. 1881. Tipo: Venezuela, Sucre, “prope Quetepe et in Monte Cocollar”, s.d., *Humboldt & Bonpland s.n.* (holótipo, P-Bonpl; isótipo, B-Willd).”

**Accepted name:** *Palicourea rigida* Kunth

Type: VENEZUELA. Sucre: “prope Quetepe et in Monte Cocollar”, s.d., *J.A. Bonpland & F.W.H.A. Humboldt* 246 (holotype, P-Bonpl [barcode P00671115]; isotype B-W [barcode B-W 04106-02 0]).

FGT, vol. 40(2), p. 809:
“40-8. *Palicourea triphylla* DC., Prodr. 4: 526. 1830. - *Psychotria triphylla* (DC.) Muell. Arg. in Mart., Fl. Bras. 6(5): 233. 1881. Tipo: Guiana Francesa, sem localidade, s.d., *Patris s.n.* (holótipo, G-DC).”

**Accepted name:** *Palicourea triphylla* DC.

Type: FRENCH GUIANA: Without locality, s.d., *J.B. Patris s.n.* (holotype, G-DC [2 sheets, barcodes G00667830, G00667831]).

**Notes:** In G-DC there are two sheets associated with this name, which are kept together in the same folder and numbered as 1 and 2. Sheet number 1, with barcode G00667830, has one labed with the annotation “Cayenne” handwritten by Candolle, and one infructescence. Sheet number 2, barcode G00667831, has the labels with “Cayenne” and “Palicourea triphylla DC.” handwritten by Candolle, and branch with a few leaves and an inflorescence, and a separate node with a leaf pair. Because the two sheets are numbered consecutively and kept together, they are treated as one specimen, which is the holotype of *Palicourea triphylla* DC.

FGT, vol. 40(2), p. 813:
“40-9. *Palicourea urbaniana* Standl., Publ. Field Columbian Mus., Bot. Ser. 8: 219. 1930. - Tipo: Brasil, Goiás, sem localidade, 1894-1895 (fl, fr), *A. Glaziou* 21533 (holótipo, B, destruído).”

**Accepted name:** *Palicourea urbaniana* Standl.

Type: BRAZIL. Goiás: Capelinha de Santo Antonio, 1894-1895 [“25 Dec. 1894”] (fl, fr), *A.F.M. Glaziou* 21533 (K [barcode K000471481], **lectotype here designated**; isoctotypes P [barcode P00837027] fragment F [Acc. No. 611912 barcode V0070103F]).

**Notes:** The original material of *Palicourea urbaniana* Standl. at B was destroyed during WWII. Three specimens associated with this name are at F, K and P, which are discussed below. On the F sheet with Acc. No. 611912 is affixed an envelope in which are included fragments from B, consisting of two leaves and a distal portion of a branchlet with a few flower buds. On the envelope is typewritten “Palicourea urbaniana Standl. - Brazil: Capelinha de Santo Antonio, Goyaz, A. Glaziou 21533 - 1894-95. From type in hb. Berol.” On the same sheet is mounted a photograph of the destroyed B specimen. Above the photograph is annotated “C.N.H.M. negative number 49964.”

On the K sheet with barcode K000471481 are affixed two flowering branches and a label with the heading “Brazil: Chiefly Province of Goyaz” and printed at the bottom “Comm. Dr. A. Glaziou, 1896.” On the label is handwritten (author unknown) “21533. Palicourea sp.” Below the two branches is the annotation “Palicourea Urbaniana Standl! det. P.C.S.” handwritten directly on the sheet by Paul Standley. This specimen is here designated the lectotype of *Palicourea urbaniana*.

The P specimen, with barcode P00837027, has a label with the heading “HERB. MUS. PARIS”, the printed text “BRÉSIL, ..., Herbier de A. GLAZIOU, donné par Mme SIMART, sa fille, en 1907”, and the handwritten annotation “Psychotria Crulsiana Glaz. n. sp., (Goyaz) Capelinha de Sto. Antonio, 25 décembre 1894, Arbuste, n. 21533.” At the lower left corner of the sheet there is a label with the annotation “Capelinha de Sto. Antonio, le 25 décembre 1894, Crulsiana Glaz. n. sp. Arbuste” handwritten in pencil by Glaziou. *Psychotria crulsiana* Glaziou, published by Glaziou (1909b: 350) is a nomen nudum because it was listed with the sole description “Arbuste, fl. jaunes”, which is insufficient for distinguishing this taxon from numerous other yellow-flowered *Palicourea* species. The specimen P00837027 consists of a branch with numerous leaves and inflorescences and is an isotype of *Palicourea urbaniana*.

41. **PENTAS** Benth. in Hook., Bot. Mag. 70: pl. 4086. 1844.

FGT, vol. 40(2), p. 817:
“41-1. *Pentas lanceolata* (Forssk.) Deflers, Voy. Yemen, 142. 1889. - *Ophiorrhiza lanceolata* Forssk., Fl. Aegypt. Arab. 42. 1775. - *Manettia lanceolata* (Forssk.) Vahl.
FGT, vol. 40(2), p. 794:
“40-7. *Palicourea rigida* Kunth in Humb. & Bonpl., Nov. Gen. Sp. 3: 370 (quarto ed.). 1819. - *Psychotria rigida* var. *genuina* Muell. Arg. in Mart., Fl. Bras. 6(5): 230. 1881. Tipo: Venezuela, Sucre, “prope Quetepe et in Monte Cocollar”, s.d., *Humboldt & Bonpland s.n.* (holótipo, P-Bonpl; isótipo, B-Willd).”

**Accepted name:** *Palicourea rigida* Kunth

Type: VENEZUELA. Sucre: “prope Quetepe et in Monte Cocollar”, s.d., *J.A. Bonpland & F.W.H.A. Humboldt* 246 (holotype, P-Bonpl [barcode P00671115]; isotype B-W [barcode B-W 04106-02 0]).

FGT, vol. 40(2), p. 809:
“40-8. *Palicourea triphylla* DC., Prodr. 4: 526. 1830. - *Psychotria triphylla* (DC.) Muell. Arg. in Mart., Fl. Bras. 6(5): 233. 1881. Tipo: Guiana Francesa, sem localidade, s.d., *Patris s.n.* (holótipo, G-DC).”

**Accepted name:** *Palicourea triphylla* DC.

Type: FRENCH GUIANA: Without locality, s.d., *J.B. Patris s.n.* (holotype, G-DC [2 sheets, barcodes G00667830, G00667831]).

**Notes:** In G-DC there are two sheets associated with this name, which are kept together in the same folder and numbered as 1 and 2. Sheet number 1, with barcode G00667830, has one labed with the annotation “Cayenne” handwritten by Candolle, and one infructescence. Sheet number 2, barcode G00667831, has the labels with “Cayenne” and “Palicourea triphylla DC.” handwritten by Candolle, and branch with a few leaves and an inflorescence, and a separate node with a leaf pair. Because the two sheets are numbered consecutively and kept together, they are treated as one specimen, which is the holotype of *Palicourea triphylla* DC.

FGT, vol. 40(2), p. 813:
“40-9. *Palicourea urbaniana* Standl., Publ. Field Columbian Mus., Bot. Ser. 8: 219. 1930. - Tipo: Brasil, Goiás, sem localidade, 1894-1895 (fl, fr), *A. Glaziou* 21533 (holótipo, B, destruído).”

**Accepted name:** *Palicourea urbaniana* Standl.

Type: BRAZIL. Goiás: Capelinha de Santo Antonio, 1894-1895 [“25 Dec. 1894”] (fl, fr), *A.F.M. Glaziou* 21533 (K [barcode K000471481], **lectotype here designated**; isoctotypes P [barcode P00837027] fragment F [Acc. No. 611912 barcode V0070103F]).

**Notes:** The original material of *Palicourea urbaniana* Standl. at B was destroyed during WWII. Three specimens associated with this name are at F, K and P, which are discussed below. On the F sheet with Acc. No. 611912 is affixed an envelope in which are included fragments from B, consisting of two leaves and a distal portion of a branchlet with a few flower buds. On the envelope is typewritten “Palicourea urbaniana Standl. - Brazil: Capelinha de Santo Antonio, Goyaz, A. Glaziou 21533 - 1894-95. From type in hb. Berol.” On the same sheet is mounted a photograph of the destroyed B specimen. Above the photograph is annotated “C.N.H.M. negative number 49964.”

On the K sheet with barcode K000471481 are affixed two flowering branches and a label with the heading “Brazil: Chiefly Province of Goyaz” and printed at the bottom “Comm. Dr. A. Glaziou, 1896.” On the label is handwritten (author unknown) “21533. Palicourea sp.” Below the two branches is the annotation “Palicourea Urbaniana Standl! det. P.C.S.” handwritten directly on the sheet by Paul Standley. This specimen is here designated the lectotype of *Palicourea urbaniana*.

The P specimen, with barcode P00837027, has a label with the heading “HERB. MUS. PARIS”, the printed text “BRÉSIL, ..., Herbier de A. GLAZIOU, donné par Mme SIMART, sa fille, en 1907”, and the handwritten annotation “Psychotria Crulsiana Glaz. n. sp., (Goyaz) Capelinha de Sto. Antonio, 25 décembre 1894, Arbuste, n. 21533.” At the lower left corner of the sheet there is a label with the annotation “Capelinha de Sto. Antonio, le 25 décembre 1894, Crulsiana Glaz. n. sp. Arbuste” handwritten in pencil by Glaziou. *Psychotria crulsiana* Glaziou, published by Glaziou (1909b: 350) is a nomen nudum because it was listed with the sole description “Arbuste, fl. jaunes”, which is insufficient for distinguishing this taxon from numerous other yellow-flowered *Palicourea* species. The specimen P00837027 consists of a branch with numerous leaves and inflorescences and is an isotype of *Palicourea urbaniana*.

41. **PENTAS** Benth. in Hook., Bot. Mag. 70: pl. 4086. 1844.

FGT, vol. 40(2), p. 817:
“41-1. *Pentas lanceolata* (Forssk.) Deflers, Voy. Yemen, 142. 1889. - *Ophiorrhiza lanceolata* Forssk., Fl. Aegypt. Arab. 42. 1775. - *Manettia lanceolata* (Forssk.) Vahl. - *Palicourea urbaniana* Standl.”
Symb. Bot. 1: 12. 1790. *Neurocarpaea lanceolata* (Forssk.) R. Br. in Salt., Voy. Abyss. App. 4: 64. 1814. *Pseudomussaenda lanceolata* (Forssk.) Wernham, J. Bot. 54: 298. 1916. Tipo: Africa, Yemen, Hadie Mts., *Forsskål s.n.* (holótipo, C; isótipo, BM)."

**Accepted name:** *Pentas lanceolata* (Forssk.) Deflers

Type: YEMEN. Hadie Mountains, "In montibus altioribus *Hadie*, alibique", s.d. [1763], *Forsskål s.n.* (first-step lectotype designated by Verdcourt (1976: 208); C [barcode C10002685, *Addit no. 1526*], *second-step lectotype here designated*; dubious isolectotypes, BM [barcode BM000945087], C [barcodes C10002684, C10002686], S [Acc. Nos. S05-9977, S05-9978]].

**Notes:** Forsskål (1775: 42-43) along with the description of *Ophiorrhiza lanceolata* Forssk. cited material collected by himself in the locality "montibus altioribus *Hadie*, alibique" but did not cite any collection number. Verdcourt (1976: 208) cited the type of *O. lanceolata* as "Type: Yemen, Hadie Mts., *Forsskål* (C, holo.!, BM, iso.!). Verdcourt's citation is an inadvertent first-step lectotypification, because at C there are three specimens with this name collected by Forsskål, which are below discussed.

The specimen *Forsskål s.n.* [Addit no. 1525] at C, with barcode C10002684, has the stamp "Reinserted in the Herb. Forskalii from the General Herbarium at C, and recorded as Addit. No. 1525 [number handwritten]." On the upper corner of the sheet is the label "IDC microfiche foto Forsskål nr. 74I, 5-6 [numbers handwritten]." This specimen is a distal portion of a branch, with three leaf pairs and two inflorescences with flowers in anthesis and flower buds, and the base of the inflorescences there is an infructescence with mature capsules. On the reverse side of the sheet, on the upper right corner is handwritten (author unknown) "Coll. Forsskål." On the same side of the sheet is glued a piece of paper with the handwritten text "This plant has had a very chequered nomenclatural history & I am writing it of in the Kew Bulletin. The exact details will appear in my thesis. D.C. & Wernham both treated it as a synonym of *Mussaenda luteola* Dec & Wernham erected the genus & name *Pseudomussaenda lanceolata* (Forssk.) Wernham. He was quite wrong since this plant on the sheet is clearly a *Pentas* & the correct combination is *Pentas lanceolata* (Forssk.) Deflers. The other plant must now be called *Pseudomussaenda flava* Verdcourt ined. nom. nov. *Pentas lanceolata* is not at all equal to the cultivated *P. carnea* Benth. as has always been supposed.

Also *Pentas lanceolata* (Forssk.) K. Schum. is a misidentification since K. Schumann had *P. bussei* Krause (= P. coccinea Stapf) in mind. B.V." (Names underlined by Verdcourt). On that piece of paper is glued a smaller piece of paper with the handwritten date "28/12/50" and the printed annotation "Det. by Bernard Verdcourt".

The C specimen *Forsskål s.n.* [Addit no. 1526], with barcode C10002685, has the stamp "Reinserted in the Herb. Forskalii from the General Herbarium at C and recorded as Addit. No. 1527 [number handwritten]. At one time in the private "Herb. Leibmann [last word handwritten]." On the upper margin of the sheet is the label "IDC microfiche foto Forsskål nr. 74I, 7-8 [numbers handwritten]." On this sheet are present the distal portion of a branch with a few leaf pairs and one flower in anthesis, an inflorescence subtended by one leaf, with one flower in anthesis, and a branch with seven leaf pairs and a terminal infructescence with immature fruits. On the bottom of the sheet, it is handwritten by Vahl "Manettia lanceolata. Ophiorrhyza" [lanceolata] Forsk!"

The C specimen *Forsskål s.n.* [Addit no. 1527], with barcode C10002686, has the stamp "Reinserted in the Herb. Forskalii from the General Herbarium at C and recorded as Addit. No. 1527 [number handwritten]. At one time in the private "Herb. Leibmann [last word handwritten]." This is a small specimen with a few leaf pairs and two flowers in anthesis.

As the three above described specimens were collected by Forsskål, they represent original material of *Ophiorrhiza lanceolata*. However, there is no indication on those sheets that they belong to the same gathering. Specimen with barcode C10002685 is the most complete and is here designated the second-step lectotype of this name.

At BM there is a sheet with two different gatherings of *Pentas lanceolata* affixed on it. The specimen with number "1" and barcode BM000945087, has the annotation "Yemen: Mts. of Hadie, Forsskål, march 1763" penciled directly on the sheet by an unknown author (not Forsskål). This annotation was certainly added after the specimen was mounted on the sheet. A label affixed on the sheet has the annotation "1. Ex Oriente Forsskål." This is the specimen cited by Verdcourt (1976: 208) as "isotype" as it has been annotated by him as "Type of Pentas lanceolata (Forssk.) K. Schum. = Ophiorrhiza lanceolata Forssk., Det. BV, 1951, duplic. of Forsskål sp. in Copenhagen." Because this specimen has not been annotated by Forsskål, it is a dubious isolectotype.

Two specimens collected by Forsskål are at S. Specimen with Accession No. S05-9977 does not report the collection locality, is not annotated by him, and has
Symb. Bot. 1: 12. 1790. *Neurocarpaea lanceolata* (Forssk.) R. Br. in Salt., Voy. Abyss. App. 4: 64. 1814. *Pseudomussaenda lanceolata* (Forssk.) Wernham, J. Bot. 54: 298. 1916. Tipo: Africa, Yemen, Hadie Mts., *Forsskål s.n.* (holótipo, C; isótipo, BM)."

**Accepted name:** *Pentas lanceolata* (Forssk.) Deflers

Type: YEMEN. Hadie Mountains, "In montibus altioribus *Hadie*, alibique", s.d. [1763], *Forsskål s.n.* (first-step lectotype designated by Verdcourt (1976: 208); C [barcode C10002685, *Addit no. 1526*], **second-step lectotype here designated**; dubious isolectotypes, BM [barcode BM000945087], C [barcodes C10002684, C10002686], S [Acc. Nos. S05-9977, S05-9978]].

**Notes:** Forsskål (1775: 42-43) along with the description of *Ophiorrhiza lanceolata* Forssk. cited material collected by himself in the locality "montibus altioribus *Hadie*, alibique" but did not cite any collection number. Verdcourt (1976: 208) cited the type of *O. lanceolata* as "Type: Yemen, Hadie Mts., *Forsskål* (C, holo.!, BM, iso.!). Verdcourt's citation is an inadvertent first-step lectotypification, because at C there are three specimens with this name collected by Forsskål, which are below discussed.

The specimen *Forsskål s.n.* [Addit no. 1525] at C, with barcode C10002684, has the stamp "Reinserted in the Herb. Forskalii from the General Herbarium at C, and recorded as Addit. No. 1525 [number handwritten]." On the upper corner of the sheet is the label "IDC microfiche foto Forsskål nr. 74I, 5-6 [numbers handwritten]." This specimen is a distal portion of a branch, with three leaf pairs and two inflorescences with flowers in anthesis and flower buds, and the base of the inflorescences there is an infructescence with mature capsules. On the reverse side of the sheet, on the upper right corner is handwritten (author unknown) "Coll. Forsskål." On the same side of the sheet is glued a piece of paper with the handwritten text "This plant has had a very chequered nomenclatural history & I am writing it of in the Kew Bulletin. The exact details will appear in my thesis. D.C. & Wernham both treated it as a synonym of *Mussaenda luteola* Dec & Wernham erected the genus & name *Pseudomussaenda lanceolata* (Forssk.) Wernham. He was quite wrong since this plant on the sheet is clearly a *Pentas* & the correct combination is *Pentas lanceolata* (Forssk.) Deflers. The other plant must now be called *Pseudomussaenda flava* Verdcourt ined. nom. nov. *Pentas lanceolata* is not at all equal to the cultivated *P. carnea* Benth. as has always been supposed.

Also *Pentas lanceolata* (Forssk.) K. Schum. is a misidentification since K. Schumann had *P. bussei* Krause (= P. coccinea Stapf) in mind. B.V." (Names underlined by Verdcourt). On that piece of paper is glued a smaller piece of paper with the handwritten date "28/12/50" and the printed annotation "Det. by Bernard Verdcourt".

The C specimen *Forsskål s.n.* [Addit no. 1526], with barcode C10002685, has the stamp "Reinserted in the Herb. Forskalii from the General Herbarium at C and recorded as Addit. No. 1527 [number handwritten]. At one time in the private "Herb. Leibmann [last word handwritten]." On the upper margin of the sheet is the label "IDC microfiche foto Forsskål nr. 74I, 7-8 [numbers handwritten]." On this sheet are present the distal portion of a branch with a few leaf pairs and one flower in anthesis, an inflorescence subtended by one leaf, with one flower in anthesis, and a branch with seven leaf pairs and a terminal infructescence with immature fruits. On the bottom of the sheet, it is handwritten by Vahl "Manettia lanceolata. Ophiorrhyza" [lanceolata] Forsk!"

The C specimen *Forsskål s.n.* [Addit no. 1527], with barcode C10002686, has the stamp "Reinserted in the Herb. Forskalii from the General Herbarium at C and recorded as Addit. No. 1527 [number handwritten]. At one time in the private "Herb. Leibmann [last word handwritten]." This is a small specimen with a few leaf pairs and two flowers in anthesis.

As the three above described specimens were collected by Forsskål, they represent original material of *Ophiorrhiza lanceolata*. However, there is no indication on those sheets that they belong to the same gathering. Specimen with barcode C10002685 is the most complete and is here designated the second-step lectotype of this name.

At BM there is a sheet with two different gatherings of *Pentas lanceolata* affixed on it. The specimen with number "1" and barcode BM000945087, has the annotation "Yemen: Mts. of Hadie, Forsskål, march 1763" penciled directly on the sheet by an unknown author (not Forsskål). This annotation was certainly added after the specimen was mounted on the sheet. A label affixed on the sheet has the annotation "1. Ex Oriente Forsskål." This is the specimen cited by Verdcourt (1976: 208) as "isotype" as it has been annotated by him as "Type of Pentas lanceolata (Forssk.) K. Schum. = Ophiorrhiza lanceolata Forssk., Det. BV, 1951, duplic. of Forsskål sp. in Copenhagen." Because this specimen has not been annotated by Forsskål, it is a dubious isolectotype.

Two specimens collected by Forsskål are at S. Specimen with Accession No. S05-9977 does not report the collection locality, is not annotated by him, and has
the annotation “Possible type of Ophiorrhiza lanceolata Forssk.” handwritten in pencil by an unknown author directly on the sheet. The specimen consists of a branch with numerous leaves and a terminal inflorescence. Specimen with No. S05-9978 does not report the collection locality, is not annotated by him, and has the annotation “Possible type of Ophiorrhiza lanceolata Forssk.” handwritten by the same unknown author in pencil directly on the sheet. The specimen consists of a small branch with a few leaves and a terminal inflorescence, mounted on a smaller sheet glued on a larger sheet. On the smaller sheet, below the small branch, is handwritten “Mussaenda luteola Delile” in black ink, by an unknown author. The two S specimens are here treated as dubious isolectotypes of *Ophiorrhiza lanceolata*.

42. **PERAMA** Aubl., Pl. Guian. Franc. 1: 54, pl. 18. 1775.

FGT, vol. 40(2), p. 822:
“42-1. **Perama hirsuta** Aubl., Hist. Pl. Guiane 54, pl. 18. 1775. - *Mattuschkea hirsuta* (Aubl.) Vahl, Symb. 3: 11. 1794. - Tipo: Guiana Francesa: “In locis humidis & arenosis Aurora & Orapu”, VII/[1762-1764] (fl), J.B.C.F. Aublet s.n. (holótipo, BM, foto em NY).”

**Accepted name:** *Perama hirsuta* Aubl.

Type: FRENCH GUIANA: Near Roura and near the Orapu River, moist open fields with sandy soil, “in locis humidis & arenosis Aroura & Orapu” and “dans les lieux humides & sablonneux des quartiers d’Aroura & d’Orapu”, s.d. [Apr, Sep 1763, Mar 1764], J.B.C.F. Aublet s.n. (P-JJR 3: 117), lectotype designated by Lanjouw and Uittien (1940: 155); isolectotype, BM [barcode BM001009065]).

**Notes:** For further information about the typification of *Perama hirsuta* Aubl., see Delprete (2015).

43. **POSOQUERIA** Aubl., Hist. Pl. Guiane 133. 1775.

FGT, vol. 40(2), p. 831:
“43-1. **Posoqueria latifolia** (Rudge) Roem. & Schult., Syst. Pl. 5: 227. 1819. - *Solena latifolia* Rudge, Pl. Guian. 1: 26, tab. 40. 1806. - Tipo: Guiana Francesa, Rudge, Pl. Guian., pl. 40. 1806 (lectótipo).”

**Accepted name:** *Posoqueria latifolia* (Rudge) Roem. & Schult.

**Correct bibliographic citation:** *Posoqueria latifolia* (Rudge) Roem. & Schult., Syst. Pl. 5: 227. 1819.
- *Solena latifolia* Rudge, Pl. Guian. 1: 26. 1805; tab. 40. 1806 (the plate is not original material because it was published after the publication of the name).
- *Tocoyena latifolia* (Rudge) Poir., Tabl. Encycl. 2(5): 259. 1819. [on page 259 for *Tocoyena latifolia* (Rudge) Poir. is cited “Tab. 163, fig. 2”; Plate 163 has the title “Tocoyena” and in the plate are present two different taxa; on fig. 2 is depicted a species of *Posoqueria*, while on fig. 1 is depicted a species of *Tocoyena*].

Type: FRENCH GUIANA: Without locality, s.d., J. Martin s.n. (BM [barcode BM001008912], lectotype here designated; isolectotypes BM [barcode BM001008911], FI-W [2 sheets, barcodes FI067338, FI067339]).

**Notes:** According to Stafleu and Cowan (1983: 972), the description of *Solena latifolia* is in part 3 of Rudge’s (1805) *Plantarum Guianae*, which was published on June 1805, while Plate 40 is in part 4, which was published on April-May 1806. Therefore Plate 40 is not original material. The material used by Rudge to describe the taxa in his *Plantarum Guianae Rariorum Icones et Descriptio-nes* was collected by Joseph Martin in French Guiana. The fate of Martin’s specimens was described by Stearn and Williams (1957) and summarized by Stafleu and Cowan (1983: 971-972). Succinctly, in 1803, France and England were at war, and the specimens collected by Joseph Martin in French Guiana, originally intended for the Museum of Natural History of Paris, were confiscated by two British privateers and brought to London. About 400 of these specimens were bought by the British Museum (BM), and 772 were bought by Edward Rudge (1763-1846) and included in his own herbarium (Stafleu & Cowan, 1983: 971-972). Also, Rudge gave a partial set of those specimens to Banks, whose herbarium became the founding collection of BM. After Rudge’s death, his widow donated his herbarium to BM in 1847. Therefore, most of Martin’s collections converged to BM, although some of his specimens are also reported to be at FI or FI-Webb (Stafleu & Cowan, 1983: 971-972). Those specimens have the handwritten information “Guiana. Martin”, which may give the false impression that they were collected in modern day Guyana, but they actually came from French Guiana.

Rudge (1805: 27) below the description of *Solena latifolia* added the following observation “OBS. In figureà Prosoquerieae [sic! Posoqueriae] apud Aublet, limbus regularis depingitur, nihilominus valde suspicor irregularem esse: in ejus specimine, nunc in Herbario
the annotation “Possible type of Ophiorrhiza lanceolata Forssk.” handwritten in pencil by an unknown author directly on the sheet. The specimen consists of a branch with numerous leaves and a terminal inflorescence. Specimen with No. S05-9978 does not report the collection locality, is not annotated by him, and has the annotation “Possible type of Ophiorrhiza lanceolata Forssk.” handwritten by the same unknown author in pencil directly on the sheet. The specimen consists of a small branch with a few leaves and a terminal inflorescence, mounted on a smaller sheet glued on a larger sheet. On the smaller sheet, below the small branch, is handwritten “Mussaenda luteola Delile” in black ink, by an unknown author. The two S specimens are here treated as dubious isolectotypes of *Ophiorrhiza lanceolata*.

42. **PERAMA** Aubl., Pl. Guian. Franc. 1: 54, pl. 18. 1775.

FGT, vol. 40(2), p. 822:
“42-1. **Perama hirsuta** Aubl., Hist. Pl. Guiane 54, pl. 18. 1775. - *Mattuschkea hirsuta* (Aubl.) Vahl, Symb. 3: 11. 1794. - Tipo: Guiana Francesa: “In locis humidis & arenosis Aurora & Orapu”, VII/[1762-1764] (fl), J.B.C.F. Aublet s.n. (holótipo, BM, foto em NY).”

**Accepted name:** *Perama hirsuta* Aubl.

Type: FRENCH GUIANA: Near Roura and near the Orapu River, moist open fields with sandy soil, “in locis humidis & arenosis Aroura & Orapu” and “dans les lieux humides & sablonneux des quartiers d’Aroura & d’Orapu”, s.d. [Apr, Sep 1763, Mar 1764], J.B.C.F. Aublet s.n. (P-JJR 3: 117), lectotype designated by Lanjouw and Uittien (1940: 155); isolectotype, BM [barcode BM001009065]).

**Notes:** For further information about the typification of *Perama hirsuta* Aubl., see Delprete (2015).

43. **POSOQUERIA** Aubl., Hist. Pl. Guiane 133. 1775.

FGT, vol. 40(2), p. 831:
“43-1. **Posoqueria latifolia** (Rudge) Roem. & Schult., Syst. Pl. 5: 227. 1819. - *Solena latifolia* Rudge, Pl. Guian. 1: 26, tab. 40. 1806. - Tipo: Guiana Francesa, Rudge, Pl. Guian., pl. 40. 1806 (lectótipo).”

**Accepted name:** *Posoqueria latifolia* (Rudge) Roem. & Schult.

**Correct bibliographic citation:** *Posoqueria latifolia* (Rudge) Roem. & Schult., Syst. Pl. 5: 227. 1819.
- *Solena latifolia* Rudge, Pl. Guian. 1: 26. 1805; tab. 40. 1806 (the plate is not original material because it was published after the publication of the name).
- *Tocoyena latifolia* (Rudge) Poir., Tabl. Encycl. 2(5): 259. 1819. [on page 259 for *Tocoyena latifolia* (Rudge) Poir. is cited “Tab. 163, fig. 2”; Plate 163 has the title “Tocoyena” and in the plate are present two different taxa; on fig. 2 is depicted a species of *Posoqueria*, while on fig. 1 is depicted a species of *Tocoyena*].

Type: FRENCH GUIANA: Without locality, s.d., J. Martin s.n. (BM [barcode BM001008912], lectotype here designated; isolectotypes BM [barcode BM001008911], FI-W [2 sheets, barcodes FI067338, FI067339]).

**Notes:** According to Stafleu and Cowan (1983: 972), the description of *Solena latifolia* is in part 3 of Rudge’s (1805) *Plantarum Guianae*, which was published on June 1805, while Plate 40 is in part 4, which was published on April-May 1806. Therefore Plate 40 is not original material. The material used by Rudge to describe the taxa in his *Plantarum Guianae Rariorum Icones et Descriptio-nes* was collected by Joseph Martin in French Guiana. The fate of Martin’s specimens was described by Stearn and Williams (1957) and summarized by Stafleu and Cowan (1983: 971-972). Succinctly, in 1803, France and England were at war, and the specimens collected by Joseph Martin in French Guiana, originally intended for the Museum of Natural History of Paris, were confiscated by two British privateers and brought to London. About 400 of these specimens were bought by the British Museum (BM), and 772 were bought by Edward Rudge (1763-1846) and included in his own herbarium (Stafleu & Cowan, 1983: 971-972). Also, Rudge gave a partial set of those specimens to Banks, whose herbarium became the founding collection of BM. After Rudge’s death, his widow donated his herbarium to BM in 1847. Therefore, most of Martin’s collections converged to BM, although some of his specimens are also reported to be at FI or FI-Webb (Stafleu & Cowan, 1983: 971-972). Those specimens have the handwritten information “Guiana. Martin”, which may give the false impression that they were collected in modern day Guyana, but they actually came from French Guiana.

Rudge (1805: 27) below the description of *Solena latifolia* added the following observation “OBS. In figureà Prosoquerieae [sic! Posoqueriae] apud Aublet, limbus regularis depingitur, nihilominus valde suspicor irregularem esse: in ejus specimine, nunc in Herbario
Banksiano conservato, Antherae, quae persingulares, certe similes sunt: itaque hanc stirpem congenerem esse judicavi.” (Translation: “In the figure of *Posoqueria* by Aublet, the limb margin is depicted as regular, it does not seem irregular at all: in his [Aublet’s] specimen, preserved in Banks’ herbarium, the Anthers are peculiar, certainly similar: therefore, I judge this species as congeneric”). In Rudge’s observation, he compared his new taxon with an Aublet specimen of *Posoqueria longiflora* Aubl. [see Delprete (2015: 613) for discussion of that specimen] present in Bank’s herbarium (now integrated in BM), but he did not cite where the material of *S. latifolia* that he studied is kept. Taylor (2012c: 191) cited the type of *Solena latifolia* as “Holotipo: Guayana Francesa, *Martin s.n.* (BM)” without citing the barcode number(s).

In BM there are two sheets collected by Martin in French Guiana annotated with this name. The BM sheet with barcode BM001008912, on the upper left corner has a small handwritten label with the annotation “Guiana, Martin”. On the bottom of the sheet is penciled “Posoqueria latifolia Rom. & Sch., Solena latifolia Rudge Ic.” At the lower right corner is penciled “2”. This specimen consists of a branch with several leaves and an inflorescence with numerous flower buds at different stages of development and three flowers in anthesis. No annotation by Rudge is also present on this sheet. This specimen is here designated the lectotype of *Solena latifolia*.

The BM sheet with barcode BM001008911, on the upper left corner has a small label with the handwritten annotation “Guiana, Martin”. On the bottom of the sheet is penciled “Posoqueria latifolia Rom. & Sch., Solena latifolia Rudge Ic.” At the lower right corner is penciled “2”. The specimen is a branch with several leaves and an inflorescence with four flower buds and two flowers in anthesis. This sheet also does not have any annotation by Rudge and is an isotype of *Solena latifolia*.

In FI-W there are three specimens annotated as “*Tocoyena latifolia*” that were integrated in the Webb Herbarium from the Desfontaine herbarium, as indicated by the labels present on those sheets. Those specimens need to be evaluated as possible original material of *Solena latifolia*, and are discussed below.

The FI-W specimen with barcode FI067338 has a label with the annotation “Cayenne, Martin”. A second label has the printed text “Herb. Webbianum” and “Ex Herb. Desfontaines”, and “Tocoyena latifolia Poir.” handwritten in pencil by an unknown author. The specimen consists of a branch with several leaves and two terminal inflorescences. It is part of the original gathering by Martin in French Guiana. Although it does not have any annotation by Rudge, it is here treated as original material, and is an isotype of *Solena latifolia*.

The FI-W specimen with barcode FI067339 has a handwritten label with “Posoqueria” stroke through and the annotation “Tocoyena latifolia Poiret, decrit dans les actes de la société d’histoire nat. par M. Richard. Cayenne. Martin.” A second label has the headings “Herb. Webbianum” and “Ex Herb. Desfontaines”. This specimen, consisting of a branch with several leaves and a terminal inflorescence, is part of the original gathering by Martin in French Guiana. Although it does not have any annotation by Rudge, it is here treated as original material, and is an isotype of *Solena latifolia*.

The FI-W specimen with barcode FI067340 has the handwritten annotation “Tocoyena species nova, Confer. Posoqueria Aublet, Cayenne, Martin.” On the sheet is affixed a second label with the printed heading “Herb. Webbianum” and “Ex Herb. Desfontaines”. The specimen consists of several loose leaves and one flower in anthesis. Because of the actinomorphic corolla with acute lobes, this specimen belongs to the genus *Tocoyena*, and is not original material of *Solena latifolia*.

FGT, vol. 40(2), p. 837:

“43-2. *Posoqueria longiflora* Aubl., Hist. Pl. Guiane 1: 134, tab. 51. 1775; non Roxb. (1824). - *Solena longiflora* Willd., Sp. Pl. 1: 961. 1798. - *Kyrtanthus longiflorus* (Aubl.) Gmel., Syst. Nat. 2(1): 362. 1791. Tipo: Guiane Francesa, “sur le bords de grands riviers de la Guiane”, *Aublet s.n.* (holótipo, BM).”

**Accepted name:** *Posoqueria longiflora* Aubl.

Type: FRENCH GUIANA: Margins of great riv- ers, “Floret Novembri; fructum sert Januario [...] ad ripas fluviorum” and “sur les bords des grandes rivieres de la Guiane [...] en fleur dans le mois de Novembre & en fruit au mois de Janvier”, s.d. [1762– 1764], *J.B.C.F. Aublet s.n.* (P-JJR (P-JJR 6: 199), lecto- type designated by Lanjouw and Uittien (1940: 156); isolectotype, BM [barcode BM001008904]).

**Notes:** For further information about the typifica-
tion of *Posoqueria longiflora* Aubl., see Delprete (2015).

44. **PSYCHOTRIA** L., Syst. Nat., ed. 10, 929, 1364. 1759,
*nom. cons.*

FGT, vol. 40(2), p. 851:

“44-1. *Psychotria anceps* Kunth in Humb. & Bonpl., Nov. Gen. Sp. 3: 360. 1819. Tipo: Colômbia, Rio Tequen-
Banksiano conservato, Antherae, quae persingulares, certe similes sunt: itaque hanc stirpem congenerem esse judicavi.” (Translation: “In the figure of *Posoqueria* by Aublet, the limb margin is depicted as regular, it does not seem irregular at all: in his [Aublet’s] specimen, preserved in Banks’ herbarium, the Anthers are peculiar, certainly similar: therefore, I judge this species as congeneric”). In Rudge’s observation, he compared his new taxon with an Aublet specimen of *Posoqueria longiflora* Aubl. [see Delprete (2015: 613) for discussion of that specimen] present in Bank’s herbarium (now integrated in BM), but he did not cite where the material of *S. latifolia* that he studied is kept. Taylor (2012c: 191) cited the type of *Solena latifolia* as “Holotipo: Guayana Francesa, *Martin s.n.* (BM)” without citing the barcode number(s).

In BM there are two sheets collected by Martin in French Guiana annotated with this name. The BM sheet with barcode BM001008912, on the upper left corner has a small handwritten label with the annotation “Guiana, Martin”. On the bottom of the sheet is penciled “Posoqueria latifolia Rom. & Sch., Solena latifolia Rudge Ic.” At the lower right corner is penciled “2”. This specimen consists of a branch with several leaves and an inflorescence with numerous flower buds at different stages of development and three flowers in anthesis. No annotation by Rudge is also present on this sheet. This specimen is here designated the lectotype of *Solena latifolia*.

The BM sheet with barcode BM001008911, on the upper left corner has a small label with the handwritten annotation “Guiana, Martin”. On the bottom of the sheet is penciled “Posoqueria latifolia Rom. & Sch., Solena latifolia Rudge Ic.” At the lower right corner is penciled “2”. The specimen is a branch with several leaves and an inflorescence with four flower buds and two flowers in anthesis. This sheet also does not have any annotation by Rudge and is an isotype of *Solena latifolia*.

In FI-W there are three specimens annotated as “*Tocoyena latifolia*” that were integrated in the Webb Herbarium from the Desfontaine herbarium, as indicated by the labels present on those sheets. Those specimens need to be evaluated as possible original material of *Solena latifolia*, and are discussed below.

The FI-W specimen with barcode FI067338 has a label with the annotation “Cayenne, Martin”. A second label has the printed text “Herb. Webbianum” and “Ex Herb. Desfontaines”, and “Tocoyena latifolia Poir.” handwritten in pencil by an unknown author. The specimen consists of a branch with several leaves and two terminal inflorescences. It is part of the original gathering by Martin in French Guiana. Although it does not have any annotation by Rudge, it is here treated as original material, and is an isotype of *Solena latifolia*.

The FI-W specimen with barcode FI067339 has a handwritten label with “Posoqueria” stroke through and the annotation “Tocoyena latifolia Poiret, decrit dans les actes de la société d’histoire nat. par M. Richard. Cayenne. Martin.” A second label has the headings “Herb. Webbianum” and “Ex Herb. Desfontaines”. This specimen, consisting of a branch with several leaves and a terminal inflorescence, is part of the original gathering by Martin in French Guiana. Although it does not have any annotation by Rudge, it is here treated as original material, and is an isotype of *Solena latifolia*.

The FI-W specimen with barcode FI067340 has the handwritten annotation “Tocoyena species nova, Confer. Posoqueria Aublet, Cayenne, Martin.” On the sheet is affixed a second label with the printed heading “Herb. Webbianum” and “Ex Herb. Desfontaines”. The specimen consists of several loose leaves and one flower in anthesis. Because of the actinomorphic corolla with acute lobes, this specimen belongs to the genus *Tocoyena*, and is not original material of *Solena latifolia*.

FGT, vol. 40(2), p. 837:

“43-2. *Posoqueria longiflora* Aubl., Hist. Pl. Guiane 1: 134, tab. 51. 1775; non Roxb. (1824). - *Solena longiflora* Willd., Sp. Pl. 1: 961. 1798. - *Kyrtanthus longiflorus* (Aubl.) Gmel., Syst. Nat. 2(1): 362. 1791. Tipo: Guiane Francesa, “sur le bords de grands riviers de la Guiane”, *Aublet s.n.* (holótipo, BM).”

Accepted name: *Posoqueria longiflora* Aubl.

Type: FRENCH GUIANA: Margins of great riv- ers, “Floret Novembri; fructum sert Januario [...] ad ripas fluviorum” and “sur les bords des grandes rivieres de la Guiane [...] en fleur dans le mois de Novembre & en fruit au mois de Janvier”, s.d. [1762– 1764], *J.B.C.F. Aublet s.n.* (P-JJR (P-JJR 6: 199), lecto- type designated by Lanjouw and Uittien (1940: 156); isolectotype, BM [barcode BM001008904]).

**Notes:** For further information about the typifica-
tion of *Posoqueria longiflora* Aubl., see Delprete (2015).

44. **PSYCHOTRIA** L., Syst. Nat., ed. 10, 929, 1364. 1759,
*nom. cons.*

FGT, vol. 40(2), p. 851:

“44-1. *Psychotria anceps* Kunth in Humb. & Bonpl., Nov. Gen. Sp. 3: 360. 1819. Tipo: Colômbia, Rio Tequen-
dama, s.d., *Humboldt & Bonpland s.n.* (holótipo P-Bonpl., isótipo, B destruído). [Subg. Psychotria].”

**Accepted name:** *Psychotria anceps* Kunth

**Original material:** Colombia. Cundinamarca: Bogotá River, Tequendama Falls, “Crescit in temperatis subfrigidis Regni Novo-Granatensis, juxta cataractam Tequendamae, alt. 1300 hex. Floret Augusto”, s.d., *A. Bonpland & F.W.H.A. Humbold s.n.* (lost).

Type: COLOMBIA. Boyacá: Canada Castero, 20 km de Sta. Elena, piste Mani-Sta. Elena, 25 Feb. 1971 (fl). *C. Sastre* 862 (P [barcode P04579979], **neotype here designated**; isoneotype COL [Acc. No. 140074, barcode COL000157756]).

**Notes:** The fate of the botanical specimens gathered by Bonpland and Humboldt has been discussed by several authors (Delprete 2001; Hiepko 2006; Kirkbride and Wiersema 2020; Lack 2004a, 2004b, 2009, 2018; Stauffer et al. 2012). After his return to Paris, Humboldt divided those collections in several sets, to be studied by different specialists. The main set, although incomplete, is currently kept in the Bonpland Herbarium (P-Bonpl.), in the Museum of Natural History in Paris. Bonpland was initially in charge of writing the descriptions for the *Nova genera et species plantarum*, but after several months he renounced the task. Then, Humboldt invited Carl Ludwig Willdenow (1765–1812) to take up this huge task, to whom he sent a set in Berlin, but he died shortly after. Then, Humboldt invited Carl Sigismund Kunth (1788–1850), who was in Berlin, to take up this enormous work. Kunth arrived in Paris in 1813 and started immediately to write the descriptions for the *Nova genera*. In 1816, Bonpland emigrated from France to Argentina, with his personal herbarium, which was a third, incomplete set of the collections that he made with Humboldt. Therefore, Kunth and Bonpland where both in Paris from 1813 to 1816, and it is possible that Kunth was able to study the specimens in Bonpland’s personal herbarium during that period. Kunth went to La Rochelle, just before Bonpland’s departure, to convince him to leave his personal set and his field books in France, but he was only able to obtain Bonpland’s field books (“*Journal Botanique*”). Bonpland eventually returned his personal herbarium from Argentina in 1832, which arrived in Paris in 1833, and was then integrated in the P general herbarium in 1837. Those specimens have the label “Herbier de l’Amérique équatoriale, donné par M. A. Bonpland” and Kunth’s Rubiaceae names were published before 1833, therefore they cannot be treated as original material.

Kunth studied the specimens collected by Bonpland and Humboldt in Paris. For many names published by Kunth, no specimens are found in the herbaria where Bonpland and Humboldt’s collections are deposited. This is not surprising, as the absence of original specimens in P-Bonpl. has puzzled many botanists (e.g., Stauffer et al. 2012). For example, out of the about 350 monocotyledon names described by Kunth, Stauffer et al. (2012) reported that 235 original specimens are not present in P-Bonpl. After an exhaustive search in P and P-Bonpl. no original specimen of *Psychotria anceps* was found. Jurai Paule (B herbarium curator) confirmed the absence of original material *P. anceps* in B-W. Because no original specimen associated with this name could be found, it is therefore necessary to designate a neotype. The specimen *Sastre* 862 at P with barcode P04579979 is here designated the neotype of this name, and the duplicate at COL is an isoneotype.

FGT, vol. 40(2), p. 856:

“44-2. *Psychotria bracteocardia* (DC.) Muell. Arg., in Mart., Fl. Bras. 6(5): 362. 1881. - *Cephaelis bracteocardia* DC., Prodr. 4: 534. 1830. - *Uragoga bracteocardia* (DC.) Kuntze, Rev. Gen. 2: 959. 1891. Tipo: Brasil, Bahia, *Salzmann s.n.* (holótipo, G-DC; isótipos, MPU [2]). [Subg. Heteropsychotria].”

**Accepted name:** *Palicourea bracteocardia* (DC.) Delprete & J.H.Kirkbr., J. Bot. Res. Inst. Texas 10(2): 414. 2016.

Type: BRAZIL. Bahia: [“in collibus umbrosis”], s.d. [1827–1830], *P. Salzmann s.n.* (first-step lectotype designated by Steyermark (1972: 689); G-DC [barcode G00300922], second-step lectotype designated by Delprete and J.H. Kirkbride (2016: 414); isolectotypes, MPU [4 sheets, barcodes MPU022236, MPU022237, MPU022238, MPU022239]).

FGT, vol. 40(2), p. 860:

“44-3. *Psychotria capitata* Ruiz & Pav., Fl. Peruv. 2: 59, pl. 206, fig. a. 1709. Tipo: Peru, Dept. Huánuco, Chinchao, s.d., *Ruiz & Pavón s.n.* (holótipo, MA). [Subg. Heteropsychotria].”

**Accepted name:** *Palicourea violacea* (Aubl.) A.Rich., Mem. Fam. Rubiac. 95. 1830.

Type: PERU. Huánuco: Chinchao, “in Andium montibus nemorosis per Chinchao runcationes,” s.d., *H.*
dama, s.d., *Humboldt & Bonpland s.n.* (holótipo P-Bonpl., isótipo, B destruído). [Subg. Psychotria].”

**Accepted name:** *Psychotria anceps* Kunth

**Original material:** Colombia. Cundinamarca: Bogotá River, Tequendama Falls, “Crescit in temperatis subfrigidis Regni Novo-Granatensis, juxta cataractam Tequendamae, alt. 1300 hex. Floret Augusto”, s.d., *A. Bonpland & F.W.H.A. Humbold s.n.* (lost).

Type: COLOMBIA. Boyacá: Canada Castero, 20 km de Sta. Elena, piste Mani-Sta. Elena, 25 Feb. 1971 (fl). *C. Sastre* 862 (P [barcode P04579979], **neotype here designated**; isoneotype COL [Acc. No. 140074, barcode COL000157756]).

**Notes:** The fate of the botanical specimens gathered by Bonpland and Humboldt has been discussed by several authors (Delprete 2001; Hiepko 2006; Kirkbride and Wiersema 2020; Lack 2004a, 2004b, 2009, 2018; Stauffer et al. 2012). After his return to Paris, Humboldt divided those collections in several sets, to be studied by different specialists. The main set, although incomplete, is currently kept in the Bonpland Herbarium (P-Bonpl.), in the Museum of Natural History in Paris. Bonpland was initially in charge of writing the descriptions for the *Nova genera et species plantarum*, but after several months he renounced the task. Then, Humboldt invited Carl Ludwig Willdenow (1765–1812) to take up this huge task, to whom he sent a set in Berlin, but he died shortly after. Then, Humboldt invited Carl Sigismund Kunth (1788–1850), who was in Berlin, to take up this enormous work. Kunth arrived in Paris in 1813 and started immediately to write the descriptions for the *Nova genera*. In 1816, Bonpland emigrated from France to Argentina, with his personal herbarium, which was a third, incomplete set of the collections that he made with Humboldt. Therefore, Kunth and Bonpland where both in Paris from 1813 to 1816, and it is possible that Kunth was able to study the specimens in Bonpland’s personal herbarium during that period. Kunth went to La Rochelle, just before Bonpland’s departure, to convince him to leave his personal set and his field books in France, but he was only able to obtain Bonpland’s field books (“*Journal Botanique*”). Bonpland eventually returned his personal herbarium from Argentina in 1832, which arrived in Paris in 1833, and was then integrated in the P general herbarium in 1837. Those specimens have the label “Herbier de l’Amérique équatoriale, donné par M. A. Bonpland” and Kunth’s Rubiaceae names were published before 1833, therefore they cannot be treated as original material.

Kunth studied the specimens collected by Bonpland and Humboldt in Paris. For many names published by Kunth, no specimens are found in the herbaria where Bonpland and Humboldt’s collections are deposited. This is not surprising, as the absence of original specimens in P-Bonpl. has puzzled many botanists (e.g., Stauffer et al. 2012). For example, out of the about 350 monocotyledon names described by Kunth, Stauffer et al. (2012) reported that 235 original specimens are not present in P-Bonpl. After an exhaustive search in P and P-Bonpl. no original specimen of *Psychotria anceps* was found. Jurai Paule (B herbarium curator) confirmed the absence of original material *P. anceps* in B-W. Because no original specimen associated with this name could be found, it is therefore necessary to designate a neotype. The specimen *Sastre* 862 at P with barcode P04579979 is here designated the neotype of this name, and the duplicate at COL is an isoneotype.

FGT, vol. 40(2), p. 856:

“44-2. *Psychotria bracteocardia* (DC.) Muell. Arg., in Mart., Fl. Bras. 6(5): 362. 1881. - *Cephaelis bracteocardia* DC., Prodr. 4: 534. 1830. - *Uragoga bracteocardia* (DC.) Kuntze, Rev. Gen. 2: 959. 1891. Tipo: Brasil, Bahia, *Salzmann s.n.* (holótipo, G-DC; isótipos, MPU [2]). [Subg. Heteropsychotria].”

**Accepted name:** *Palicourea bracteocardia* (DC.) Delprete & J.H.Kirkbr., J. Bot. Res. Inst. Texas 10(2): 414. 2016.

Type: BRAZIL. Bahia: [“in collibus umbrosis”], s.d. [1827–1830], *P. Salzmann s.n.* (first-step lectotype designated by Steyermark (1972: 689); G-DC [barcode G00300922], second-step lectotype designated by Delprete and J.H. Kirkbride (2016: 414); isolectotypes, MPU [4 sheets, barcodes MPU022236, MPU022237, MPU022238, MPU022239]).

FGT, vol. 40(2), p. 860:

“44-3. *Psychotria capitata* Ruiz & Pav., Fl. Peruv. 2: 59, pl. 206, fig. a. 1709. Tipo: Peru, Dept. Huánuco, Chinchao, s.d., *Ruiz & Pavón s.n.* (holótipo, MA). [Subg. Heteropsychotria].”

**Accepted name:** *Palicourea violacea* (Aubl.) A.Rich., Mem. Fam. Rubiac. 95. 1830.

Type: PERU. Huánuco: Chinchao, “in Andium montibus nemorosis per Chinchao runcationes,” s.d., *H.*
Ruiz López & J.A. Pavón y Jiménez s.n. (MA [bar-
code MA815948]), lectotype designated by Delprete
and Kirkbride (2016: 435). – PERU. Junín: Schunke
Hacienda, above San Ramon, 1300–1700 m, dense
forest, Aug.–Sep. 1923, C. Schunke A19 (US [Acc.
No. 1470372], epitype designated by Delprete and
Kirkbride (2016: 435)).

FGT, vol. 40(2), p. 866:
“44-4. *Psychotria carthagenensis* Jacq., Enum. Pl. Carib.
16. 1760. Tipo: Colômbia, Bolívar, Cartagena, s.d., *Jac-
quin s.n.* (LE?, W?). [Subg. Psychotria].”

**Accepted name:** *Psychotria carthagenensis* Jacq.

**Original material:** Colombia. Bolívar: “habitat Carthagenae”, Aug., *N.J. Jacquin s.n.* (not traced).

Type: [Icon] “Psychotria carthagenensis” in Jac-
quin, Select. Stirp. Amer. Hist. tab. 174, fig. 22.
1763, neotype designated by Taylor in Taylor et al.
(2020: 6. e-publication). – COLOMBIA. Magdale-
na: Santa Marta, 200 ft. [ca. 60 m], Mar. 1898–1901,
H.H. *Smith 1802* (MO [Acc. No. 124263, barcode
MO-1765070], epitype designated by Taylor in Tay-
lor et al. (2020: 6, e-publication); isoepitype US [Acc.
No. 533747, barcode 02518872)].

**Notes:** In the protologue of *Psychotria carthagenensis* Jacq., Jacquin (1760: 16) did not cite any locality or herbarium specimen. No original specimen associated with this name is present at LE, P or W. Because of absence of original specimens, Taylor et al. (2020) designated as lectotype table 174, figure 22 of Jacquin's (1763) *Selecta-rum Stirpium Americanarum Historia*. In that figure, *P. carthagenensis* is represented only by one leaf, one flower, and one fruit, with features that could apply to numerous species of *Psychotria* and are insufficient to fix the appli-cation of the name. Also, as table 174, figure 22 was pub-lished three years after the publication of the name, it is not original material. Hence, Taylor's lectotype citation, according to Art. 9.10 of the *Code*, can be corrected to a neotype designation. The epitype specimen designated by Taylor, *H.H. Smith 1802* at MO, is valid and is here fol-lowed, and an isoepitype specimen is present in US.

FGT, vol. 40(2), p. 877:
“44-5. *Psychotria colorata* (Willd. ex Roem. & Schult.)
Muell. Arg. in Mart., Fl. Bras. 6(5): 372. 1881. - *Cepha-
elis colorata* Willd. ex Roem. & Schult., Syst. Veg. 5: 214.

1819. Tipo: Brasil, s.d., *Sieber s.n.* (B-Willd 4149). [Subg. Heteropsychotria].”

**Accepted name:** *Palicourea colorata* (Hoffmanns. ex Willd.) Delprete & J.H.Kirkbr., J. Bot. Res. Inst. Texas 10(2): 416. 2016.

Type: BRAZIL: [Amazon Basin], without locality, s.d., *F.W. Sieber s.n.* [dedit J.C. Hoffmansegg] (holotype: B-W 04149-01(a) B-W 04149-02(b)).

**Notes:** For additional synonyms and types of *Palicourea colorata* (Hoffmanns. ex Roem. & Schult.) Delprete & Kirkbr., see Delprete and Kirkbride (2016).

FGT, vol. 40(2), p. 877:

**Synonym:**

*“Psychotria megapontica* Muell. Arg. in Mart., Fl. Bras. 6(5): 362. 1881. *Uragoga megapontica* (Muell. Arg.) Kuntze, Revis. Gen. Pl. 2: 961. 1891. - *Psychotria colorata* (Willd. ex Roem. & Schult.) Muell. Arg. ssp. *megapontica* (Muell. Arg.) Steyerm., Mem. New York Bot. Gard. 23: 685. 1972. Tipo: Brasil, Goiás, “Megaponte” [sic! “Meia Ponte” agora Pirenópolis], s.d., *Pohl 976* (holótipo G; sin-tipo B, destruído; foto-B em NY).”

Type: BRAZIL. Goiás: “Megaponte” [sic! “Meia Ponte,” now town of Pirenopolis], s.d., *J.E. Pohl 976* (G [barcode G 00301215], lectotype designated by Delprete and Kirkbride (2016: 416)).

**Notes:** Müller Argoviensis (1881: 362) for *Psychotria megapontica* Müll. Arg. cited the only gathering *Pohl 976* without citing the herbarium of deposit. Delprete (2010b: 877) cited the specimen at G as the holotype. According to the *Code*, Delprete’s citation cannot be corrected as a lectotype designation because it lacks “here designated” or a similar expression. Delprete & Kirkbride (2016: 416) designated the same specimen as lectotype.

FGT, vol. 40(2), p. 886:
“44-6. *Psychotria deflexa* DC., Prodr. 4: 510. 1830. Tipo: Guiana Francesa, s.d., *Patris s.n.* (holótipo, G-DC). [Subg. Heteropsychotria].”

**Accepted name:** *Palicourea deflexa* (DC.) Borhidi, Acta Bot. Hung. 53: 243. 2012 [“2011”].

Type: FRENCH GUIANA: Without locality, s.d., *J.B. Patris s.n.* (first-step lectotype designated by
Ruiz López & J.A. Pavón y Jiménez s.n. (MA [bar-code MA815948]), lectotype designated by Delprete and Kirkbride (2016: 435). – PERU. Junín: Schunke Hacienda, above San Ramon, 1300–1700 m, dense forest, Aug.–Sep. 1923, C. Schunke A19 (US [Acc. No. 1470372], epitype designated by Delprete and Kirkbride (2016: 435)).

FGT, vol. 40(2), p. 866:
“44-4. *Psychotria carthagenensis* Jacq., Enum. Pl. Carib. 16. 1760. Tipo: Colômbia, Bolívar, Cartagena, s.d., *Jacquin s.n.* (LE?, W?). [Subg. Psychotria].”

**Accepted name:** *Psychotria carthagenensis* Jacq.

**Original material:** Colombia. Bolívar: “habitat Carthagenae”, Aug., *N.J. Jacquin s.n.* (not traced).

Type: [Icon] “Psychotria carthagenensis” in Jacquin, Select. Stirp. Amer. Hist. tab. 174, fig. 22. 1763, neotype designated by Taylor in Taylor et al. (2020: 6. e-publication). – COLOMBIA. Magdalena: Santa Marta, 200 ft. [ca. 60 m], Mar. 1898–1901, *H.H. Smith 1802* (MO [Acc. No. 124263, barcode MO-1765070], epitype designated by Taylor in Taylor et al. (2020: 6, e-publication); isoepitype US [Acc. No. 533747, barcode 02518872]).

**Notes:** In the protologue of *Psychotria carthagenensis* Jacq., Jacquin (1760: 16) did not cite any locality or herbarium specimen. No original specimen associated with this name is present at LE, P or W. Because of absence of original specimens, Taylor et al. (2020) designated as lectotype table 174, figure 22 of Jacquin's (1763) *Selecta-rum Stirpium Americanarum Historia*. In that figure, *P. carthagenensis* is represented only by one leaf, one flower, and one fruit, with features that could apply to numerous species of *Psychotria* and are insufficient to fix the application of the name. Also, as table 174, figure 22 was published three years after the publication of the name, it is not original material. Hence, Taylor's lectotype citation, according to Art. 9.10 of the *Code*, can be corrected to a neotype designation. The epitype specimen designated by Taylor, *H.H. Smith 1802* at MO, is valid and is here followed, and an isoepitype specimen is present in US.

FGT, vol. 40(2), p. 877:
“44-5. *Psychotria colorata* (Willd. ex Roem. & Schult.) Muell. Arg. in Mart., Fl. Bras. 6(5): 372. 1881. - *Cepha-elis colorata* Willd. ex Roem. & Schult., Syst. Veg. 5: 214.

1819. Tipo: Brasil, s.d., *Sieber s.n.* (B-Willd 4149). [Subg. Heteropsychotria].”

**Accepted name:** *Palicourea colorata* (Hoffmanns. ex Willd.) Delprete & J.H.Kirkbr., J. Bot. Res. Inst. Texas 10(2): 416. 2016.

Type: BRAZIL: [Amazon Basin], without locality, s.d., *F.W. Sieber s.n.* [dedit J.C. Hoffmansegg] (holotype: B-W 04149-01(a) B-W 04149-02(b)).

**Notes:** For additional synonyms and types of *Palicourea colorata* (Hoffmanns. ex Roem. & Schult.) Delprete & Kirkbr., see Delprete and Kirkbride (2016).

FGT, vol. 40(2), p. 877:

**Synonym:**

“*Psychotria megapontica* Muell. Arg. in Mart., Fl. Bras. 6(5): 362. 1881. *Uragoga megapontica* (Muell. Arg.) Kuntze, Revis. Gen. Pl. 2: 961. 1891. - *Psychotria colorata* (Willd. ex Roem. & Schult.) Muell. Arg. ssp. *megapontica* (Muell. Arg.) Steyerm., Mem. New York Bot. Gard. 23: 685. 1972. Tipo: Brasil, Goiás, “Megaponte” [sic! “Meia Ponte” agora Pirenópolis], s.d., *Pohl 976* (holótipo G; sin-tipo B, destruído; foto-B em NY).”

Type: BRAZIL. Goiás: “Megaponte” [sic! “Meia Ponte,” now town of Pirenopolis], s.d., *J.E. Pohl 976* (G [barcode G 00301215], lectotype designated by Delprete and Kirkbride (2016: 416)).

**Notes:** Müller Argoviensis (1881: 362) for *Psychotria megapontica* Müll. Arg. cited the only gathering *Pohl 976* without citing the herbarium of deposit. Delprete (2010b: 877) cited the specimen at G as the holotype. According to the *Code*, Delprete’s citation cannot be corrected as a lectotype designation because it lacks “here designated” or a similar expression. Delprete & Kirkbride (2016: 416) designated the same specimen as lectotype.

FGT, vol. 40(2), p. 886:
“44-6. *Psychotria deflexa* DC., Prodr. 4: 510. 1830. Tipo: Guiana Francesa, s.d., *Patris s.n.* (holótipo, G-DC). [Subg. Heteropsychotria].”

**Accepted name:** *Palicourea deflexa* (DC.) Borhidi, Acta Bot. Hung. 53: 243. 2012 [“2011”].

Type: FRENCH GUIANA: Without locality, s.d., *J.B. Patris s.n.* (first-step lectotype designated by
Steyermark (1972: 502); G-DC [barcode G00478847 barcode, sheet No. 4, with two flowering branches, annotated as "Psychotria deflea DC." by Candolle], second-step lectotype designated by Delprete and Kirkbride (2016: 418); isolectotypes, G-DC [barcode G00478847, sheet No. 3, with flower buds)]).

**Notes.** In G-DC, inside the folder annotated as "Psychotria deflexa DC., De Cand. prodr. 4 p. 510 n. 57", aside from the two preparations designated by Delprete and Kirkbride (2016: 418) as lectotype and isolectotype, there are two additional specimens, both with barcode G00478849, on sheet No. 1 (sterile) and sheet No. 2 (with flower buds). Those two additional specimens were annotated by Delprete in 2013 as "dubious isolectotypes G-DC, probably not the same species." The two specimens with barcode G00478849 were identified by A. Berger in 2022 as *Palicourea octocuspis* Müll. Arg.

FGT, vol. 40(2), p. 890:

"44-7. *Psychotria goyazensis* Muell. Arg., Flora 59: 548, 551. 1876. Tipo: Brasil, Goiás, "prope urbem Goyaz" [cidade de Goiás], s.d., *Burchell* 6723 (holótipo, G). [Subg. Heteropsychotria]."

**Accepted name:** *Palicourea brevicollis* (Müll. Arg.) C.M. Taylor in Taylor and Hollowel, Novon 25(1): 85-86. 2016.

**Notes:** For a clarification of the species delimitation, full synonymy and typification of *Palicourea brevicollis* (Müll. Arg.) C.M.Taylor is presented below. Additional information is provided by Taylor in Taylor and Hollowel (2016: 85-86).

**Synonymy:**

*Palicourea brevicollis* (Müll. Arg.) C.M.Taylor in C.M.Taylor and Hollowel, Novon 25(1): 85. 2016.
- *Psychotria brevicollis* Müll. Arg., Flora 59: 548, 552. 1876c. - *Uragoga brachytere* Kuntze, Revis. Gen. Pl. 2: 955. 1891, replacement name, not *Uragoga brevicollis* (Müll. Arg.) Kuntze (based on *Mapouria brevicollis* Müll. Arg., 1876b). Type: Brazil. Minas Gerais: Caldas, s.d., *A.F. Regnell III.116 p.p.* (lectotype BR [barcode BR000000531615], designated by C.M. Taylor in Taylor and Hollowell (2016: 85).

*Psychotria regnellii* Müll. Arg., Flora 548, 552. 1876. - *Uragoga regnellii* (Müll. Arg.) Kuntze, Revis. Gen. Pl. 2: 952. 1891.

Type: BRAZIL. Minas Gerais: Caldas, s.d., *A.F. Regnell III.116 p.p.* (BR [barcode BR000000531569], lectotype designated by C.M.Taylor in Taylor and Hollowell (2016: 85)).

*Psychotria goyazensis* Müll. Arg., Flora 59: 548, 551. 1876. - *Palicourea goyazensis* (Müll. Arg.) Borhidi, Acta Bot. Hung. 59(1-2): 34. 2017.

Type: BRAZIL. Goiás: "prope urbem Goyaz" [town of Goiás], s.d., *W.J. Burchell* 6723 (BR [BR000000531632], **lectotype here designated**; isolectotype K [barcode K000174296]; isolectotype fragment G [barcode G00096064]).

**Notes:** In the protologue of *Psychotria goyazensis* Müll.Arg., Müller Argoviensis (1876c: 548, 551) cited the gathering *Burchell* 6723 collected in the town of Goiás, the first capital of the homonymous state. In BR there is a specimen from the Martius Herbarium, with barcode BR000000531632, annotated by Müller Argoviensis. The specimen label has the annotation "Brasilia: Prope urbem Goyaz, 1861, Burchell no. 6723" and is here designated as the lectotype of this name. Fragments consisting of a few fruits and a leaf portion of *Burchell* 6723 are in an envelope in G annotated by Müller Argoviensis, and represent an isolectotype. A duplicate of *Burchell* 6723 in K, not annotated by him, is another isolectotype.

FGT, vol. 40(2), p. 895:

"44-8. *Psychotria gracilenta* Muell. Arg., Flora 59: 542, 545. 1876. Tipo: Brasil, Bahia, sem localidade, s.d., *Blanchet* 1590 (holótipo, G). [Subg. Heteropsychotria]."

**Accepted name:** *Palicourea gracilenta* (Müll.Arg.) Delprete & J.H.Kirkbr., J. Bot. Res. Inst. Texas 10(2): 421. 2016.

**Notes:** Clarification of the species delimitation, full synonymy and typification of *Palicourea gracilenta* (Müll. Arg.) Delprete & J.H.Kirkbr. is presented below.

*Palicourea gracilenta* (Müll.Arg.) Delprete & J.H.Kirkbr., J. Bot. Res. Inst. Texas 10(2): 421. 2016. - *Psychotria gracilenta* Müll. Arg., Flora 59:542, 545. 1876.

Type: BRAZIL. Bahia: without locality, s.d. [1834], *J.S. Blanchet* 1590 (G [barcode G00300770], lectotype designated by Delprete and Kirkbride (2016: 421); isolectotype BM [barcode BM000624171]).
Steyermark (1972: 502); G-DC [barcode G00478847 barcode, sheet No. 4, with two flowering branches, annotated as "Psychotria deflea DC." by Candolle], second-step lectotype designated by Delprete and Kirkbride (2016: 418); isolectotypes, G-DC [barcode G00478847, sheet No. 3, with flower buds)]).

**Notes.** In G-DC, inside the folder annotated as "Psychotria deflexa DC., De Cand. prodr. 4 p. 510 n. 57", aside from the two preparations designated by Delprete and Kirkbride (2016: 418) as lectotype and isolectotype, there are two additional specimens, both with barcode G00478849, on sheet No. 1 (sterile) and sheet No. 2 (with flower buds). Those two additional specimens were annotated by Delprete in 2013 as "dubious isolectotypes G-DC, probably not the same species." The two specimens with barcode G00478849 were identified by A. Berger in 2022 as *Palicourea octocuspis* Müll. Arg.

FGT, vol. 40(2), p. 890:

"44-7. *Psychotria goyazensis* Muell. Arg., Flora 59: 548, 551. 1876. Tipo: Brasil, Goiás, "prope urbem Goyaz" [cidade de Goiás], s.d., *Burchell* 6723 (holótipo, G). [Subg. Heteropsychotria]."

**Accepted name:** *Palicourea brevicollis* (Müll. Arg.) C.M. Taylor in Taylor and Hollowel, Novon 25(1): 85-86. 2016.

**Notes:** For a clarification of the species delimitation, full synonymy and typification of *Palicourea brevicollis* (Müll. Arg.) C.M.Taylor is presented below. Additional information is provided by Taylor in Taylor and Hollowel (2016: 85-86).

**Synonymy:**

*Palicourea brevicollis* (Müll. Arg.) C.M.Taylor in C.M.Taylor and Hollowel, Novon 25(1): 85. 2016.
- *Psychotria brevicollis* Müll. Arg., Flora 59: 548, 552. 1876c. - *Uragoga brachytere* Kuntze, Revis. Gen. Pl. 2: 955. 1891, replacement name, not *Uragoga brevicollis* (Müll. Arg.) Kuntze (based on *Mapouria brevicollis* Müll. Arg., 1876b). Type: Brazil. Minas Gerais: Caldas, s.d., *A.F. Regnell III.116 p.p.* (lectotype BR [barcode BR000000531615], designated by C.M. Taylor in Taylor and Hollowell (2016: 85).

*Psychotria regnellii* Müll. Arg., Flora 548, 552. 1876. - *Uragoga regnellii* (Müll. Arg.) Kuntze, Revis. Gen. Pl. 2: 952. 1891.

Type: BRAZIL. Minas Gerais: Caldas, s.d., *A.F. Regnell III.116 p.p.* (BR [barcode BR000000531569], lectotype designated by C.M.Taylor in Taylor and Hollowell (2016: 85)).

*Psychotria goyazensis* Müll. Arg., Flora 59: 548, 551. 1876. - *Palicourea goyazensis* (Müll. Arg.) Borhidi, Acta Bot. Hung. 59(1-2): 34. 2017.

Type: BRAZIL. Goiás: "prope urbem Goyaz" [town of Goiás], s.d., *W.J. Burchell* 6723 (BR [BR000000531632], **lectotype here designated**; isolectotype K [barcode K000174296]; isolectotype fragment G [barcode G00096064]).

**Notes:** In the protologue of *Psychotria goyazensis* Müll.Arg., Müller Argoviensis (1876c: 548, 551) cited the gathering *Burchell* 6723 collected in the town of Goiás, the first capital of the homonymous state. In BR there is a specimen from the Martius Herbarium, with barcode BR000000531632, annotated by Müller Argoviensis. The specimen label has the annotation "Brasilia: Prope urbem Goyaz, 1861, Burchell no. 6723" and is here designated as the lectotype of this name. Fragments consisting of a few fruits and a leaf portion of *Burchell* 6723 are in an envelope in G annotated by Müller Argoviensis, and represent an isolectotype. A duplicate of *Burchell* 6723 in K, not annotated by him, is another isolectotype.

FGT, vol. 40(2), p. 895:

"44-8. *Psychotria gracilenta* Muell. Arg., Flora 59: 542, 545. 1876. Tipo: Brasil, Bahia, sem localidade, s.d., *Blanchet* 1590 (holótipo, G). [Subg. Heteropsychotria]."

**Accepted name:** *Palicourea gracilenta* (Müll.Arg.) Delprete & J.H.Kirkbr., J. Bot. Res. Inst. Texas 10(2): 421. 2016.

**Notes:** Clarification of the species delimitation, full synonymy and typification of *Palicourea gracilenta* (Müll. Arg.) Delprete & J.H.Kirkbr. is presented below.

*Palicourea gracilenta* (Müll.Arg.) Delprete & J.H.Kirkbr., J. Bot. Res. Inst. Texas 10(2): 421. 2016. - *Psychotria gracilenta* Müll. Arg., Flora 59:542, 545. 1876.

Type: BRAZIL. Bahia: without locality, s.d. [1834], *J.S. Blanchet* 1590 (G [barcode G00300770], lectotype designated by Delprete and Kirkbride (2016: 421); isolectotype BM [barcode BM000624171]).
*Psychotria brachybotrya* Müll.Arg. in Martius, Fl. Bras. 6(5):327. 1881.

Type: BRAZIL. Amazonas: Rio Negro, Sao Gabriel da Cachoeira, Mar. 1852, *R. Spruce 2190* (first-step lectotype designated by Steyermark (1972: 609); P [barcode P00836988], second-step lectotype designated by Delprete and Kirkbride (2016: 421); isolectotypes, K [2 sheets, barcodes K000173545, K00173546], P [barcode P00836988], W [barcode 1889-0014256]).

*Psychotria iquitosensis* Standl., Publ. Columbian Mus., Bot. Ser. 8:195. 1930.

Type: PERU. Loreto: Near Iquitos, 14 Jul. 1929, *Ll. Williams 1391* (holotype (F [Acc. No. 604636 barcode V0041058F]; isotypes fragm. G [barcode G00300771], S [Acc. No. S05-1086]).

FGT, vol. 40(2), p. 898:
“44-9. **Psychotria hoffmannseggiana** (Willd. ex Roem. & Schult.) Muell.Arg. in Mart., Fl. Bras 6(5): 336. 1881.
- *Cephaelis hoffmannseggiana* Willd. ex Roem. & Schult., Syst. Veg. 5: 214. 1819. Tipo: Brasil, Pará, s.d., *Sieber s.n.* (holótipo B-W 4155 [dedit Hoffmannsegg], foto em NY) [Subg. Heteropsychotria].”

**Accepted name:** *Palicourea hoffmannseggiana*
(Roem. & Schult.) Borhidi

FGT vol. 40(2), fig. 86 (page 926): Figure 86 is labeled as *Psychotria officinalis* (Aubl.) Sandwith, but is *Palicourea hoffmannseggiana* (Roem. & Schult.) Borhidi.

**Notes:** The species delimitation of *Psychotria hoffmannseggiana* (Roem. & Schult.) Müll. Arg. presented by Delprete (2010b: 898-904) was incorrect. To clarify the delimitation of this species, as here recognized, full synonymy and typification is presented below.

*Palicourea hoffmannseggiana* (Roem. & Schult.) Borhidi, Acta Bot. Hung. 53: 245. 2012 [“2011”]. - *Cephaelis hoffmannseggiana* Roem. & Schult., Syst. Veg. 5: 214. 1819. - *Psychotria hoffmannseggiana* (Roem. & Schult.) Müll. Arg. in Martius, Fl. Bras. 6(5):336. 1881.

Type: BRAZIL. Pará: without locality, s.d., *F.W. Sieber s.n.* [dedit *J.C. Hoffmannsegg*] (holotype B-W [barcode B-W 04155 01 0], photo at NY).

*Cephaelis rubra* Hoffmanns. ex Roem. & Schult., Syst. Veg. 5: 214. 1819. - *Psychotria rubra* (Hoffmanns. ex Roem. & Schult.) Müll.Arg. in Martius, Fl. Bras. 6(5): 336. 1881.

Type: BRAZIL. [Amazon Basin]: without locality, s.d., *F.W. Sieber s.n.* [dedit *J.C. Hoffmannsegg*] (holotype B-W [barcode B-W 04150 -01 0]).

*Psychotria barbiflora* DC., Prodr. 4: 509. 1830.

Type: BRAZIL. Bahia: without locality, 1831, *P. Salzmann s.n.* (first-step lectotype designated by Steyermark (1972: 602); G [barcode G00300780] second-step lectotype designated by Delprete and Kirkbride (2016: 422); possible isolectotypes, MPU [2 sheets, MPU022106, MPU022107], P [barcode P00836992]).

*Carapichea patrisii* DC., Prodr. 4: 536. 1830. *Cephaelis patrisii* (DC.) D. Dietr., Syn. Pl. 1: 773. 1839. - *Uragoga carapichea* (DC.) Kuntze, Revis. Gen. Pl. 2:955. 1891; non *Uragoga patrisii* (DC.) Kuntze (1891).

Type: FRENCH GUIANA: Without locality, s.d., *J.B. Patris s.n.* (G-DC) [without barcode, sheet N. 4], lectotype designated by Delprete and Kirkbride (2016: 422); possible isolectotypes, G-DC [without barcode, sheets N. 1, 2 and 3)].

*Psychotria furcata* DC., Prodr. 4: 512. 1830.

Type: PANAMA: Without locality, s.d., *T. Haenke s.n.* (holotype: G-DC [barcode G00478835]).

*Psychotria heterocephala* Müll.Arg. in Martius, Fl. Bras. 6(5): 333. 1881.

Type: BRAZIL. [State unknown]: Rio São João, s.d., *A.C.V. Schott 5301* (G [barcode G00300784], lectotype designated by Delprete and Kirkbride (2016: 422)).

*Psychotria bahiensis* Müll.Arg. in Martius, Fl. Bras. 6(5): 338. 1881, nom. illeg. [non *Psychotria bahiensis* DC. 1830].

Type: BRAZIL. Bahia: without locality, s.d., *J.S. Blanchet 1815* (G [barcode G00301031], lectotype designated by Delprete and Kirkbride (2016: 422); isolectotypes, F [Acc. No. 734074], P [barcode P00836959]).

*Psychotria flavicans* Müll.Arg. in Martius, Fl. Bras. 6(5): 339. 1881.
*Psychotria brachybotrya* Müll.Arg. in Martius, Fl. Bras. 6(5):327. 1881.

Type: BRAZIL. Amazonas: Rio Negro, Sao Gabriel da Cachoeira, Mar. 1852, *R. Spruce 2190* (first-step lectotype designated by Steyermark (1972: 609); P [barcode P00836988], second-step lectotype designated by Delprete and Kirkbride (2016: 421); isolectotypes, K [2 sheets, barcodes K000173545, K00173546], P [barcode P00836988], W [barcode 1889-0014256]).

*Psychotria iquitosensis* Standl., Publ. Columbian Mus., Bot. Ser. 8:195. 1930.

Type: PERU. Loreto: Near Iquitos, 14 Jul. 1929, *Ll. Williams 1391* (holotype (F [Acc. No. 604636 barcode V0041058F]; isotypes fragm. G [barcode G00300771], S [Acc. No. S05-1086]).

FGT, vol. 40(2), p. 898:
“44-9. **Psychotria hoffmannseggiana** (Willd. ex Roem. & Schult.) Muell.Arg. in Mart., Fl. Bras 6(5): 336. 1881.
- *Cephaelis hoffmannseggiana* Willd. ex Roem. & Schult., Syst. Veg. 5: 214. 1819. Tipo: Brasil, Pará, s.d., *Sieber s.n.* (holótipo B-W 4155 [dedit Hoffmannsegg], foto em NY) [Subg. Heteropsychotria].”

**Accepted name:** *Palicourea hoffmannseggiana*
(Roem. & Schult.) Borhidi

FGT vol. 40(2), fig. 86 (page 926): Figure 86 is labeled as *Psychotria officinalis* (Aubl.) Sandwith, but is *Palicourea hoffmannseggiana* (Roem. & Schult.) Borhidi.

**Notes:** The species delimitation of *Psychotria hoffmannseggiana* (Roem. & Schult.) Müll. Arg. presented by Delprete (2010b: 898-904) was incorrect. To clarify the delimitation of this species, as here recognized, full synonymy and typification is presented below.

*Palicourea hoffmannseggiana* (Roem. & Schult.) Borhidi, Acta Bot. Hung. 53: 245. 2012 [“2011”]. - *Cephaelis hoffmannseggiana* Roem. & Schult., Syst. Veg. 5: 214. 1819. - *Psychotria hoffmannseggiana* (Roem. & Schult.) Müll. Arg. in Martius, Fl. Bras. 6(5):336. 1881.

Type: BRAZIL. Pará: without locality, s.d., *F.W. Sieber s.n.* [dedit *J.C. Hoffmannsegg*] (holotype B-W [barcode B-W 04155 01 0], photo at NY).

*Cephaelis rubra* Hoffmanns. ex Roem. & Schult., Syst. Veg. 5: 214. 1819. - *Psychotria rubra* (Hoffmanns. ex Roem. & Schult.) Müll.Arg. in Martius, Fl. Bras. 6(5): 336. 1881.

Type: BRAZIL. [Amazon Basin]: without locality, s.d., *F.W. Sieber s.n.* [dedit *J.C. Hoffmannsegg*] (holotype B-W [barcode B-W 04150 -01 0]).

*Psychotria barbiflora* DC., Prodr. 4: 509. 1830.

Type: BRAZIL. Bahia: without locality, 1831, *P. Salzmann s.n.* (first-step lectotype designated by Steyermark (1972: 602); G [barcode G00300780] second-step lectotype designated by Delprete and Kirkbride (2016: 422); possible isolectotypes, MPU [2 sheets, MPU022106, MPU022107], P [barcode P00836992]).

*Carapichea patrisii* DC., Prodr. 4: 536. 1830. *Cephaelis patrisii* (DC.) D. Dietr., Syn. Pl. 1: 773. 1839. - *Uragoga carapichea* (DC.) Kuntze, Revis. Gen. Pl. 2:955. 1891; non *Uragoga patrisii* (DC.) Kuntze (1891).

Type: FRENCH GUIANA: Without locality, s.d., *J.B. Patris s.n.* (G-DC) [without barcode, sheet N. 4], lectotype designated by Delprete and Kirkbride (2016: 422); possible isolectotypes, G-DC [without barcode, sheets N. 1, 2 and 3)].

*Psychotria furcata* DC., Prodr. 4: 512. 1830.

Type: PANAMA: Without locality, s.d., *T. Haenke s.n.* (holotype: G-DC [barcode G00478835]).

*Psychotria heterocephala* Müll.Arg. in Martius, Fl. Bras. 6(5): 333. 1881.

Type: BRAZIL. [State unknown]: Rio São João, s.d., *A.C.V. Schott 5301* (G [barcode G00300784], lectotype designated by Delprete and Kirkbride (2016: 422)).

*Psychotria bahiensis* Müll.Arg. in Martius, Fl. Bras. 6(5): 338. 1881, *nom. illeg.* [non *Psychotria bahiensis* DC. 1830].

Type: BRAZIL. Bahia: without locality, s.d., *J.S. Blanchet 1815* (G [barcode G00301031], lectotype designated by Delprete and Kirkbride (2016: 422); isolectotypes, F [Acc. No. 734074], P [barcode P00836959]).

*Psychotria flavicans* Müll.Arg. in Martius, Fl. Bras. 6(5): 339. 1881.
Type: BRAZIL. Bahia: Igreja Velha, 1841, *J.S. Blan-
chet* 3246 (G [barcode G00300761, sheet No. 1], lecto-
type designated by Delprete and Kirkbride (2016: 422);
isolectotypes, BM [barcode BM000624170], F [2 sheets,
Acc. Nos. 520907, 734075], G [barcode G00300761, sheet
No. 2], NY [barcode 00132671], P [2 sheets, barcodes
P00837029, P00837030], U [barcode U0006199]).

*Psychotria hoffmannseggiana* f. *pubescens* Steyerm.,
Mem. New York Bot. Gard. 23: 607. 1972.

Type: GUYANA: Kaieteur Plateau, vicinity of Kaieteur Falls, along W rim of Potaro River, ca. 1400 ft [= 426 m], 18 Feb. 1962, R.S. Cowan & T.R. Soderstrom 1897 (holotype VEN [Acc. No. 78483]; isotypes MICH [barcode 1108255], US [barcode 00138792]).

FGT, vol. 40(2), p. 905:
“44-10. **Psychotria leiocarpa** Cham. & Schltdl., Linnaea 4: 22. 1829. Tipo: Brasil, “E. Brasilia tropica, varietaremque vicinus meridionalibus”, s.d., Sellow s.n. (holótipo B, destruído). [Subg. Heteropsychotria].”

**Accepted name:** *Palicourea leiocarpa* (Cham. & Schltdl.) Delprete, **comb. nov.**

Type: BRAZIL: Without locality, s.d., *F. Sellow s.n.* (K [barcode K000174314], **neotype here designated**).

**Notes:** In the protologue of *Psychotria leiocarpa* Cham. & Schltdl., Chamisso and Schlechtendal (1829a: 22) cited material collected by Sellow in Brazil. The material at B studied by those authors was destroyed during WWII. The sole specimen associated with this name that could be found is at K, and has the handwritten annotation “Brazil, Sello.” That specimen, with barcode K000174314, is a branch with numerous inflorescences with flower buds and flowers in anthesis and has the penciled annotation (author unknown) “Psychotria leiocarpa Cham. et Schl.” Because there is no evidence on the specimen that it was studied by Chamisso and Schlechtendal, it is here designated the neotype of this name.

Chamisso and Schlechtendal (1829a: 22-23) described two varieties under *Psychotria leiocarpa*: var. *tropica* Cham. & Schltdl. and var. *extratropica* Cham. & Schltdl. Müller Argoviensis (1881: 281) described two additional varieties under *Psy. leiocarpa*, var. *constricta* Müll.Arg. and var. *intermedia* Müll.Arg. These four varieties have been treated as synonymous, and no infraspecific rank is here recognized for this species. The typifi-

cation of these varietal names is beyond the scope of the
present study.

FGT, vol. 40(2), p. 909:
“44-11. *Psychotria lupulina* Benth., Hook. J. Bot. 3: 230. 1841. Tipo: Guyana, s.d., *Rob. Schomburgk 26* (holótipo, K, isótipo, BM; foto-K em NY) [Subg. Heteropsychotria].”

**Accepted name:** *Palicourea justicifolia* (Rugde) Delprete & J.H.Kirkbr., J. Bot. Res. Inst. Texas 10(2): 424. 2016.

Type: GUYANA: Without locality, s.d. [1836], *Rob. H. Schomburgk, Ser. I, 26 pro parte* “in the later sets” (BM [barcode BM001008948], lectotype designated by Delprete and Kirkbride (2016: 424); isolectotypes, BR [barcode 00000531573], E [2 sheets, barcodes E00285066, E00505324], F [Acc. No. 734131], G [2 sheets, barcodes G00300487, G00300593], GH [barcode 00095189], TCD [barcode TCD0005769], U [barcode U0006214]).

Notes: Full synonymy and typification of this spe-
cies is presented in Delprete and Kirkbride (2016).

FGT, vol. 40(2), p. 914:
“44-12. **Psychotria mapourioides** DC., Prodr. 4: 509. 1830. Tipo: Guiana Francesa, Rio Sinnamary, s.d., *J.B.C.F. Aublet s.n.* (holótipo, BM). [Subg. Psychotria].”

**Accepted name:** *Psychotria pedunculosa* Rich.,
Actes Soc. Hist. Nat. Paris 1: 107. 1792.

**Notes:** In the protologue of *Psychotria mapourioides* DC., Candolle (1830: 509) cited the material studied as "In Guianâ Gallicâ legit cl. Patris." Delprete's (2010b: 914) citation of the type specimen as collected by Aublet and present in BM is a typographical error, as the original material of this name was collected by Patris and is in G-DC. This species has traditionally been known as *P. mapourioides* and is the name that was used in FGT. A recent article by Taylor et al. (2020) established that the corrected name for this species is *P. pedunculosa* Rich., which predates the name traditionally used. For further discussion about the delimitation this species and details on the typification of synonyms, see Taylor et al. (2020). For a clarification of the species as presently recognized, full synonymy and typification is presented below.