**Notes:** In the protologue of *Tocoyena brasiliensis* Mart., Martius (1841: 82-83) cited the locality and phenology of the material studied as "Crescit in Monte Telegraphi prope Sebastianopolin; Junio floret" without citing the collector and herbarium of deposit. Schumann (1889: 348), among other collections studied of *T. brasiliensis*, cited the specimen "in monte Telegraphenberg: Luchnath, floret Junio". Most likely, Schumann's citation refers to the original specimen cited by Martius. The BR specimen, barcode 000000578046, has a label with heading "Herbarium Martii" and the handwritten information "Tocoyena brasiliensis Mart., frutex arborescens; - flores viridis-lutei. Thelegraphenberg, Brasil: Leg. Luschnath, Juny 1834." The BR specimen is the holotype of *Tocoyena brasiliensis*. A second label on this specimen has the annotation "Tocoyena brasiliensis Mart." by K. Schumann, and a third label has the annotation "Tocoyena sellowiana (Cham. & Schltdl.) Schum., det. A.L. Prado, 25.04.1984." Prado (1987), in an unpublished Master Dissertation, treated *T. brasiliensis* and *T. sellowiana* as synonymous. However, *T. sellowiana* is recognized as a distinct species by Siberbauer-Gottsberger et al. (1992) and Delprete (2010c). Siberbauer-Gottsberger et al. (1992: 160) differentiated *T. brasiliensis* in having "Corolla tube entrance villous; leaves obovate, 15-17 x 6-7.5 cm (NE. to E. Brazil)", whereas *T. sellowiana* has "Corolla tube entrance glabrous; leaves oblong-lanceolate, smaller, mostly 7-9 x 2.5-5.5 cm (E. Brazil)." This distinction between the two species is followed here. *Tocoyena brasiliensis* occurs throughout the Cerrado Biome and not just "NE to E Brazil", as stated by Siberbauer-Gottsberger et al. (1992: 160). As Delprete (2010c: 1352-1354) wrote, *T. brasiliensis* occurs from the state of Amapá to the state of São Paulo, and the holotype specimen is from the state of Rio de Janeiro.

FGT, vol. 40(3), p. 1354:

"59-3. *Tocoyena formosa* (Cham. & Schltdl.) K. Schum. in Mart., Fl. Bras. 6(6): 347. 1889. - *Gardenia formosa* Cham. & Schltdl., Linnaea 4: 200. 1829. Tipo: Brasil meridional, s.d., *Sellow s.n.* (holótipo B, destruído; frag-B at F)."

**Accepted name:** *Tocoyena formosa* (Cham. & Schltdl.) K.Schum.

Type: BRAZIL. Goiás: Mun. Monte Alegre, rod. Monte Alegre - Campos Belos (BR-010), km 137,

arvoreta de 4 m com tronco principal contorto e copa umbeliforme, botões florais verdes, corola branca e muito perfumada durante a antese, tornando-se amarela após a antese, 13°9'5"S, 46°45'57"W, 580 m, 1 Nov. 2005 (fl), P.G. Delprete, H. Lorenzi, J. Lombardi & E. Gonçalves 9228 (GB [barcode GB-0210435], **neotype here designated**; isoneotypes, CAY [barcode CAY223351], HPL [Acc. No. 10984], HTO [Acc. No. 9991], UFG [Acc. No. 40686]).

**Notes:** Chamisso and Schlechtendal (1829b: 200-201) cited the material studied of *Gardenia formosa* Cham. & Schltdl. as "E Brasilia aequinocialis misit Sellowius unicum, prohdor, specimen floriferum, fructu deficiente." (Equatorial Brazil, unfortunately a sole flowering specimen collected by Sellow, without fruit). According to Stafleu and Cowan (1976: 482; 1985: 190, respectively) "Chamisso's own herbarium was also acquired by LE." and "Diederich Franz Leonard von Schlechtendal - Herbarium and types: HAL. Several of the types of his Berlin period were at B (mainly destroyed). - Plants for the botanical gardens of Berlin and Halle, and material from his herbarium (but collected by others) are e.g. at CAS, E, FI, G, L, M, MW, WAG." The original material at B was destroyed during WWII.

After exhaustive search, no original specimen of *Gardenia formosa* was found at HAL (Marcus Lehnert, HAL Curator, pers. comm., 5 Sep. 2022). A fragment of the original specimen at B was reported in previous literature (e.g., Delprete 2010c) to be present at F; however, after exhaustive search, there is no original fragment of *G. formosa* at F (Lucia Kawasaki, pers. comm., 2 Aug. 2022).

A specimen at K, barcode K000424433, has a label with the heading "Ex Museo botanico Berolinensi", the printed text "Brasilia, leg. Sellow", the handwritten annotation "Tocoyena formosa K. Schum.", and the stamp "determ. C. Schumann". A smaller label has the printed information "Brasilia. Reliquiae d. Sello. No.", the stamp "15 nov. 1907" and the annotation "25 III". It has been annotated as "Tocoyena formosa var. pseudobrasiliensis Gottsb. & Ehrend." by Siberbauer-Gottsberger in 1998. This specimen consists of a small branch with several leaves and an immature fruit. Because it has a fruit, it cannot be original material, as Chamisso and Schlechtendal wrote that they only studied a flowering specimen.

In absence of original material, a phenotype needs to be selected, and possibly with duplicates distributed in several herbaria. The gathering *Delprete et al. 9228* has duplicates distributed in five herbaria, and the specimen at GB is here designated as the neotype of *Gardenia formosa*.
**Notes:** In the protologue of *Tocoyena brasiliensis* Mart., Martius (1841: 82-83) cited the locality and phenology of the material studied as "Crescit in Monte Telegraphi prope Sebastianopolin; Junio floret" without citing the collector and herbarium of deposit. Schumann (1889: 348), among other collections studied of *T. brasiliensis*, cited the specimen "in monte Telegraphenberg: Luchnath, floret Junio". Most likely, Schumann's citation refers to the original specimen cited by Martius. The BR specimen, barcode 000000578046, has a label with heading "Herbarium Martii" and the handwritten information "Tocoyena brasiliensis Mart., frutex arborescens; - flores viridis-lutei. Thelegraphenberg, Brasil: Leg. Luschnath, Juny 1834." The BR specimen is the holotype of *Tocoyena brasiliensis*. A second label on this specimen has the annotation "Tocoyena brasiliensis Mart." by K. Schumann, and a third label has the annotation "Tocoyena sellowiana (Cham. & Schltdl.) Schum., det. A.L. Prado, 25.04.1984." Prado (1987), in an unpublished Master Dissertation, treated *T. brasiliensis* and *T. sellowiana* as synonymous. However, *T. sellowiana* is recognized as a distinct species by Siberbauer-Gottsberger et al. (1992) and Delprete (2010c). Siberbauer-Gottsberger et al. (1992: 160) differentiated *T. brasiliensis* in having "Corolla tube entrance villous; leaves obovate, 15-17 x 6-7.5 cm (NE. to E. Brazil)", whereas *T. sellowiana* has "Corolla tube entrance glabrous; leaves oblong-lanceolate, smaller, mostly 7-9 x 2.5-5.5 cm (E. Brazil)." This distinction between the two species is followed here. *Tocoyena brasiliensis* occurs throughout the Cerrado Biome and not just "NE to E Brazil", as stated by Siberbauer-Gottsberger et al. (1992: 160). As Delprete (2010c: 1352-1354) wrote, *T. brasiliensis* occurs from the state of Amapá to the state of São Paulo, and the holotype specimen is from the state of Rio de Janeiro.

FGT, vol. 40(3), p. 1354:

"59-3. *Tocoyena formosa* (Cham. & Schltdl.) K. Schum. in Mart., Fl. Bras. 6(6): 347. 1889. - *Gardenia formosa* Cham. & Schltdl., Linnaea 4: 200. 1829. Tipo: Brasil meridional, s.d., *Sellow s.n.* (holótipo B, destruído; frag-B at F)."

**Accepted name:** *Tocoyena formosa* (Cham. & Schltdl.) K.Schum.

Type: BRAZIL. Goiás: Mun. Monte Alegre, rod. Monte Alegre - Campos Belos (BR-010), km 137,

arvoreta de 4 m com tronco principal contorto e copa umbeliforme, botões florais verdes, corola branca e muito perfumada durante a antese, tornando-se amarela após a antese, 13°9'5"S, 46°45'57"W, 580 m, 1 Nov. 2005 (fl), P.G. Delprete, H. Lorenzi, J. Lombardi & E. Gonçalves 9228 (GB [barcode GB-0210435], **neotype here designated**; isoneotypes, CAY [barcode CAY223351], HPL [Acc. No. 10984], HTO [Acc. No. 9991], UFG [Acc. No. 40686]).

**Notes:** Chamisso and Schlechtendal (1829b: 200-201) cited the material studied of *Gardenia formosa* Cham. & Schltdl. as "E Brasilia aequinocialis misit Sellowius unicum, prohdor, specimen floriferum, fructu deficiente." (Equatorial Brazil, unfortunately a sole flowering specimen collected by Sellow, without fruit). According to Stafleu and Cowan (1976: 482; 1985: 190, respectively) "Chamisso's own herbarium was also acquired by LE." and "Diederich Franz Leonard von Schlechtendal - Herbarium and types: HAL. Several of the types of his Berlin period were at B (mainly destroyed). - Plants for the botanical gardens of Berlin and Halle, and material from his herbarium (but collected by others) are e.g. at CAS, E, FI, G, L, M, MW, WAG." The original material at B was destroyed during WWII.

After exhaustive search, no original specimen of *Gardenia formosa* was found at HAL (Marcus Lehnert, HAL Curator, pers. comm., 5 Sep. 2022). A fragment of the original specimen at B was reported in previous literature (e.g., Delprete 2010c) to be present at F; however, after exhaustive search, there is no original fragment of *G. formosa* at F (Lucia Kawasaki, pers. comm., 2 Aug. 2022).

A specimen at K, barcode K000424433, has a label with the heading "Ex Museo botanico Berolinensi", the printed text "Brasilia, leg. Sellow", the handwritten annotation "Tocoyena formosa K. Schum.", and the stamp "determ. C. Schumann". A smaller label has the printed information "Brasilia. Reliquiae d. Sello. No.", the stamp "15 nov. 1907" and the annotation "25 III". It has been annotated as "Tocoyena formosa var. pseudobrasiliensis Gottsb. & Ehrend." by Siberbauer-Gottsberger in 1998. This specimen consists of a small branch with several leaves and an immature fruit. Because it has a fruit, it cannot be original material, as Chamisso and Schlechtendal wrote that they only studied a flowering specimen.

In absence of original material, a phenotype needs to be selected, and possibly with duplicates distributed in several herbaria. The gathering *Delprete et al. 9228* has duplicates distributed in five herbaria, and the specimen at GB is here designated as the neotype of *Gardenia formosa*.
FGT, vol. 40(3), p. 1355:

**Synonym:**

*Tocoyena formosa* var. *pseudobrasiliensis* Gottsb. & Ehrend., Pl. Syst. Evol. 181: 162. 1992.

Type: BRAZIL. São Paulo: Mun. Botucatu, 18 km N, 14 km E of São Manuel, along São Manuel-Piracicaba Hwy, near ex-railway station Treze de Maio, 22º45'S, 48º25'W, 550 m, 1 Nov. 1971 (fl), I. Gottsberger & G. Gottsberger 11-11171 (holotype, UB [barcode UB0040411, Acc.No. 12034]; isotypes, K [barcode K000172525], MBM [barcode MBM267187], NY [barcode 00581278], SP [barcode SP001575] UB [UB1063124, Acc. No. 11696 (the only specimen in SpeciesLink))].

**Notes:** Silberbauer-Gottsberger et al. (1992: 162) for *Tocoyena formosa* var. *pseudobrasiliensis* Gottsb. & Ehrend. cited the holotype at UB and one of the isotypes at UB, but they did not specify which of the two specimens is the holotype. In UB there is only one specimen, with barcode UB0040411 and Acc. No. 12034, with the authors' original label specifying that it is the holotype, which is followed here.

Silberbauer-Gottsberger et al. (1992: 162) cited an isotype specimen at GI, but Martin de Jong (GI Curator, pers. comm., 28 August 2022) reported that there is no isotype specimen there, nor the paratype. The same authors also cited an isotype specimen at BOTU, but Henrique Borgatto (BOTU Curator, pers. comm., 12 September 2022) reported that there is no isotype specimen there, nor the paratype.

This variety has not been recognized by Deprete (2010c: 1354-1371), who treated *Tocoyena formosa* as a variable species, without any infraspecific taxa, and is here maintained as such.

FGT, vol. 40(3), p. 1372:

"59-4. *Tocoyena viscidula* Mart., Flora 24, suppl. 2: 80-81. 1841. Tipo: Brasil, Bahia, in silvis Caatinga mediterraneis, s.d., *Martius s.n.* (holótipo, M n.v., foto-M em UEC n.v., UFMT)."

**Accepted name:** *Tocoyena viscidula* Mart.

Type: BRAZIL. Bahia: "in silvis Caatinga mediterraneis", s.d., *C.F.P. Martius* 3301 (holotype, M [barcode M-0189486], photo in UEC, UFMT).

**Notes:** In the protologue of *Tocoyena viscidula* Mart., Martius (1841: 80-81) cited the material studied

as "In silvis Caatinga mediterraneis prov. Bahiensis." At M there is a specimen, barcode M-0189486, with the label with the printed text "Dr. Martius Iter Brasil" and the handwritten annotation "[Habitat in] sylv. Catinga indes., [Provinciae] Bahiensis, Nov." This specimen is the holotype of *T. viscidula*.

60. **UNCARIA** Schreber, *nom. cons.*, Gen. 125. 1789.

FGT, vol. 40(3), p. 1377:

"60-1. *Uncaria guianensis* (Aubl.) Gmel., Syst. Nat. 2: 370. 1791. - *Ourouparia guianensis* Aubl., Hist. Pl. Guiana 1: 177, t. 68. 1775. - *Uncaria aculeata* Willd. in Usteri (baseado em *Ourouparia guianensis*), Delect. Opusc. Bot. 2: 200. 1793. - *Nauclea guianensis* (Aubl.) Poir. in Lam., Encycl. Mét. 4: 436. 1797. - *Uruparia versicolor* Raf. (baseado em *Ourouparia guianensis*), Sylv. Tell. 148. 1830. - *Uruparia guianensis* (Aubl.) Kuntze, Rev. Gen. Pl. 1: 301. 1891. Tipo: Guiana Francesa, *Aublet s.n.* (holótipo, BM)."

**Accepted name:** *Uncaria guianensis* (Aubl.) Gmel.

Type: FRENCH GUIANA: Galibi Creek, at 40 miles from the coast, "in sylvis ad ripas amnis Galibiensis, quadraginta milliaribus, & ampliùs à maris littore" and "bord de la crique des Galibis [...] en plein fleur dans le mois de Mai", s.d. [Apr-May 1763], *J.B.C.F. Aublet s.n.* (P-JJR 8: 270, lectotype designated by Lanjouw and Uittien (1940: 154); isolectotype, BM [barcode BM001008725]).

**Notes:** For further information about the typification of *Ourouparia guianensis* Aubl., see Delprete (2015).

61. **WARSZEWICZIA** Klotzsch., Flora 36: 716. 1853.

FGT, vol. 40(3), p. 1383:

"61-1. *Warszewiczia longistaminea* K. Schum. in Mart., Fl. Bras. 6(6): 218. 1889. Tipo: Brasil, Tocantins, Porto Nacional ["Goyaz, prope Porto Real"], s.d., *Burchell 8607* (holótipo, B, destruído; foto-B em NY e US; lectótipo, NY, aqui escolhido; isolectótipo F)."

**Accepted name:** *Warszewiczia longistaminea* K.Schum.

Type: BRAZIL. Tocantins: Porto Nacional ["Goyaz, prope Porto Real"], s.d., *W.J. Burchell* 8607 (B†; NY [barcode 00259228], lectotype designated by
FGT, vol. 40(3), p. 1355:

**Synonym:**

*Tocoyena formosa* var. *pseudobrasiliensis* Gottsb. & Ehrend., Pl. Syst. Evol. 181: 162. 1992.

Type: BRAZIL. São Paulo: Mun. Botucatu, 18 km N, 14 km E of São Manuel, along São Manuel-Piracicaba Hwy, near ex-railway station Treze de Maio, 22º45'S, 48º25'W, 550 m, 1 Nov. 1971 (fl), I. Gottsberger & G. Gottsberger 11-11171 (holotype, UB [barcode UB0040411, Acc.No. 12034]; isotypes, K [barcode K000172525], MBM [barcode MBM267187], NY [barcode 00581278], SP [barcode SP001575] UB [UB1063124, Acc. No. 11696 (the only specimen in SpeciesLink))].

**Notes:** Silberbauer-Gottsberger et al. (1992: 162) for *Tocoyena formosa* var. *pseudobrasiliensis* Gottsb. & Ehrend. cited the holotype at UB and one of the isotypes at UB, but they did not specify which of the two specimens is the holotype. In UB there is only one specimen, with barcode UB0040411 and Acc. No. 12034, with the authors' original label specifying that it is the holotype, which is followed here.

Silberbauer-Gottsberger et al. (1992: 162) cited an isotype specimen at GI, but Martin de Jong (GI Curator, pers. comm., 28 August 2022) reported that there is no isotype specimen there, nor the paratype. The same authors also cited an isotype specimen at BOTU, but Henrique Borgatto (BOTU Curator, pers. comm., 12 September 2022) reported that there is no isotype specimen there, nor the paratype.

This variety has not been recognized by Deprete (2010c: 1354-1371), who treated *Tocoyena formosa* as a variable species, without any infraspecific taxa, and is here maintained as such.

FGT, vol. 40(3), p. 1372:

"59-4. *Tocoyena viscidula* Mart., Flora 24, suppl. 2: 80-81. 1841. Tipo: Brasil, Bahia, in silvis Caatinga mediterraneis, s.d., *Martius s.n.* (holótipo, M n.v., foto-M em UEC n.v., UFMT)."

**Accepted name:** *Tocoyena viscidula* Mart.

Type: BRAZIL. Bahia: "in silvis Caatinga mediterraneis", s.d., *C.F.P. Martius* 3301 (holotype, M [barcode M-0189486], photo in UEC, UFMT).

**Notes:** In the protologue of *Tocoyena viscidula* Mart., Martius (1841: 80-81) cited the material studied

as "In silvis Caatinga mediterraneis prov. Bahiensis." At M there is a specimen, barcode M-0189486, with the label with the printed text "Dr. Martius Iter Brasil" and the handwritten annotation "[Habitat in] sylv. Catinga indes., [Provinciae] Bahiensis, Nov." This specimen is the holotype of *T. viscidula*.

60. **UNCARIA** Schreber, *nom. cons.*, Gen. 125. 1789.

FGT, vol. 40(3), p. 1377:

"60-1. *Uncaria guianensis* (Aubl.) Gmel., Syst. Nat. 2: 370. 1791. - *Ourouparia guianensis* Aubl., Hist. Pl. Guiane 1: 177, t. 68. 1775. - *Uncaria aculeata* Willd. in Usteri (baseado em *Ourouparia guianensis*), Delect. Opusc. Bot. 2: 200. 1793. - *Nauclea guianensis* (Aubl.) Poir. in Lam., Encycl. Mét. 4: 436. 1797. - *Uruparia versicolor* Raf. (baseado em *Ourouparia guianensis*), Sylv. Tell. 148. 1830. - *Uruparia guianensis* (Aubl.) Kuntze, Rev. Gen. Pl. 1: 301. 1891. Tipo: Guiana Francesa, *Aublet s.n.* (holótipo, BM)."

**Accepted name:** *Uncaria guianensis* (Aubl.) Gmel.

Type: FRENCH GUIANA: Galibi Creek, at 40 miles from the coast, "in sylvis ad ripas amnis Galibiensis, quadraginta milliaribus, & ampliùs à maris littore" and "bord de la crique des Galibis [...] en plein fleur dans le mois de Mai", s.d. [Apr-May 1763], *J.B.C.F. Aublet s.n.* (P-JJR 8: 270, lectotype designated by Lanjouw and Uittien (1940: 154); isolectotype, BM [barcode BM001008725]).

**Notes:** For further information about the typification of *Ourouparia guianensis* Aubl., see Deprete (2015).

61. **WARSZEWICZIA** Klotzsch., Flora 36: 716. 1853.

FGT, vol. 40(3), p. 1383:

"61-1. *Warszewiczia longistaminea* K. Schum. in Mart., Fl. Bras. 6(6): 218. 1889. Tipo: Brasil, Tocantins, Porto Nacional ["Goyaz, prope Porto Real"], s.d., *Burchell 8607* (holótipo, B, destruído; foto-B em NY e US; lectótipo, NY, aqui escolhido; isolectótipo F)."

**Accepted name:** *Warszewiczia longistaminea* K.Schum.

Type: BRAZIL. Tocantins: Porto Nacional ["Goyaz, prope Porto Real"], s.d., *W.J. Burchell* 8607 (B†; NY [barcode 00259228], lectotype designated by
Delprete (2010c: 1383); isolectotypes, BR [barcode 000000532717], GH [barcode 00094780], K [3 sheets, barcodes K0001173867, K0001173868, K000173900], US [barcode 00137681]; isolectotype fragment F [ex P, Acc. No. 971120]; photo-B in F [F0BN000069] and NY).

**Notes:** In the protologue of *Warszewiczia longistaminea* K.Schum., Schumann (1889: 218–219) cited the material studied as “*Habitat in provincia Goyaz prope Porto Real: Burchell n. 8607*.” The material examined by Schumann, at B, was destroyed during WWII. The specimen of *Burchell 8607* at NY was designated as the lectotype of this name by Delprete (2010c: 1383).

FGT, vol. 40(3), p. 1386:

“61-2. *Warszewiczia schwackei* K. Schum. in Mart., Fl. Bras. 6(6): 219. 1889. Tipo: Brasil, Amazonas, Manaus, II/1855 (fl), *Spruce 3840* (holótipo, B, destruído; lectótipo, NY, aqui escolhido; isolectótipo K; foto-K em NY).”

**Accepted name:** *Warszewiczia schwackei* K.Schum.

Type: BRAZIL. Amazonas: [Rio Negro, Manaus] "Habitat in provincia Alto Amazonas prope Manáos", Feb. 1855 (fl), *R. Spruce 3840* (B†; NY [barcode 00259229], lectotype designated by Delprete (2010c: 1386); isolectotypes, BR [barcode 000000532706], E [barcode E00285370], F [Acc. No. 767832], G [2 sheets, barcodes G00436823, G00436824], GH [barcode 00057422], K [2 sheets, barcodes K000173865, K000173866], LD [Acc. No. 1816690]; isolectotype fragment F [ex G, Acc. No. 635234]; photo-K in NY).

## ACKNOWLEDGMENTS

I wish to thank the following Herbarium Curators and Researchers for searching for original material and for sending valuable digital images: Juraj Paule (B, B-W), Ranee Prakash (BM), Olivier Lachenaud (BR), Gerardo Aymard (COL), Lucia Kawasaki (F), Nicolas Fumeaux (G, G-DC), Martin de Jong (GI), Marcus Lehnert (HAL), Hajo Esser and Andreas Fleischmann (M), Eva García Ibáñez and Javier Fuertes Aguilar (MA), Esther Garcia Guillen (MA-RJB Archives), Amy Weiss and Matthew Pace (NY), Odile Poncy, Cécile Aupic, and Sovanmoly Hul (P, P-Bonp., P-J, P-JJR, P-Lam), Maria Candida Mamede (SP), Vera Lucia Gomes Klein, Danielle Diniz and Carmen Monteiro (UFG), Joseph

H. Kirkbride Jr. (US), and Christian Bräuchler (W). I wish to express my gratitude to the staff members of the projects Jstor Global Plants (https://plants.jstor.org/), Reflora - Brazilian Plants: Historic Rescue and Virtual Herbarium for knowledge and conservation of the Brazilian Flora (http://reflora.jbrj.gov.br/reflora/PrincipalUC/PrincipalUC.do), and INCT - Herbário Virtual da Flora e Fungos (http://inct.splink.org.br/) for making the images of millions of herbarium specimens available to the general public. Important help regarding nomenclatural issues was supplied by Joseph H. Kirkbride Jr., John H. Wiersema (both at the Smithsonian Institution, Washington, D.C., U.S.A.), Kanchi Gandhi (Harvard University, Cambridge, Massachusetts, U.S.A.), and John McNeill (Royal Botanic Garden, Edinburgh, Scotland), who made considerable contributions in solving numerous nomenclatural problems, and interpreting the *Code*. Last, but no less important, I wish to express my gratitude to Joseph H. Kirkbride Jr. and two anonymous reviewers for the careful revision of the manuscript and their significant comments and corrections. Finally, I am very grateful to the Centro Studi Erbario Tropicale (FT herbarium), University of Florence (Italy) for financially supporting this publication.

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Adams CD, Taylor CM. 2012. *Spermacoce*. In: Davidse G, Sousa MS, Knapp S, Chiang F. and Collaborators (eds.), *Flora Mesoamericana*, vol. 4, part 2, Rubiaceae a Verbenaceae. Universidad Nacional Autónoma de México, Missouri Botanical Garden & The Natural History Museum (London); St. Louis, Missouri Botanical Garden Press, pp. 275–282.

Andersson L. 1992. A provisional checklist of Neotropical Rubiaceae. *Scripta Botanica Belgica*. 1: 1–230.

Andersson L. 1997. Synopsis of the genus *Ladenbergia* (Rubiaceae). *Nordic Journal of Botany*. 17: 255–299. https://doi.org/10.1111/j.1756-1051.1997.tb00316.x

Applequist W L. 2013. Report on the Nomenclature Committee of Vascular plants: 65. Taxon. 62: 1315–1326. https://doi.org/10.12705/626.49

Applequist WL. 2023. Report on the Nomenclature Committee of Vascular plants: 73. Taxon. 72: 179–204. https://doi.org/10.1002/tax.12871

Aublet JBCF. 1775. Histoire des plantes de la Guiane française. 4 vols. Paris. P. F. Didot Jeune.

Bacigalupo NM, Cabral EL. 1999. Revisión de las especies del género *Diodia* (Rubiaceae, Spermacoceae). *Darwiniana*. 37: 153–165. https://doi.org/10.14522/darwiniana.2014.371-2.348
Delprete (2010c: 1383); isolectotypes, BR [barcode 000000532717], GH [barcode 00094780], K [3 sheets, barcodes K0001173867, K0001173868, K000173900], US [barcode 00137681]; isolectotype fragment F [ex P, Acc. No. 971120]; photo-B in F [F0BN000069] and NY).

**Notes:** In the protologue of *Warszewiczia longistaminea* K.Schum., Schumann (1889: 218–219) cited the material studied as “*Habitat in provincia Goyaz prope Porto Real: Burchell n. 8607*.” The material examined by Schumann, at B, was destroyed during WWII. The specimen of *Burchell 8607* at NY was designated as the lectotype of this name by Delprete (2010c: 1383).

FGT, vol. 40(3), p. 1386:

“61-2. *Warszewiczia schwackei* K. Schum. in Mart., Fl. Bras. 6(6): 219. 1889. Tipo: Brasil, Amazonas, Manaus, II/1855 (fl), *Spruce 3840* (holótipo, B, destruído; lectótipo, NY, aqui escolhido; isolectótipo K; foto-K em NY).”

**Accepted name:** *Warszewiczia schwackei* K.Schum.

Type: BRAZIL. Amazonas: [Rio Negro, Manaus] "Habitat in provincia Alto Amazonas prope Manáos", Feb. 1855 (fl), *R. Spruce 3840* (B†; NY [barcode 00259229], lectotype designated by Delprete (2010c: 1386); isolectotypes, BR [barcode 000000532706], E [barcode E00285370], F [Acc. No. 767832], G [2 sheets, barcodes G00436823, G00436824], GH [barcode 00057422], K [2 sheets, barcodes K000173865, K000173866], LD [Acc. No. 1816690]; isolectotype fragment F [ex G, Acc. No. 635234]; photo-K in NY).

## ACKNOWLEDGMENTS

I wish to thank the following Herbarium Curators and Researchers for searching for original material and for sending valuable digital images: Juraj Paule (B, B-W), Ranee Prakash (BM), Olivier Lachenaud (BR), Gerardo Aymard (COL), Lucia Kawasaki (F), Nicolas Fumeaux (G, G-DC), Martin de Jong (GI), Marcus Lehnert (HAL), Hajo Esser and Andreas Fleischmann (M), Eva García Ibáñez and Javier Fuertes Aguilar (MA), Esther Garcia Guillen (MA-RJB Archives), Amy Weiss and Matthew Pace (NY), Odile Poncy, Cécile Aupic, and Sovanmoly Hul (P, P-Bonp., P-J, P-JJR, P-Lam), Maria Candida Mamede (SP), Vera Lucia Gomes Klein, Danielle Diniz and Carmen Monteiro (UFG), Joseph

H. Kirkbride Jr. (US), and Christian Bräuchler (W). I wish to express my gratitude to the staff members of the projects Jstor Global Plants (https://plants.jstor.org/), Reflora - Brazilian Plants: Historic Rescue and Virtual Herbarium for knowledge and conservation of the Brazilian Flora (http://reflora.jbrj.gov.br/reflora/PrincipalUC/PrincipalUC.do), and INCT - Herbário Virtual da Flora e Fungos (http://inct.splink.org.br/) for making the images of millions of herbarium specimens available to the general public. Important help regarding nomenclatural issues was supplied by Joseph H. Kirkbride Jr., John H. Wiersema (both at the Smithsonian Institution, Washington, D.C., U.S.A.), Kanchi Gandhi (Harvard University, Cambridge, Massachusetts, U.S.A.), and John McNeill (Royal Botanic Garden, Edinburgh, Scotland), who made considerable contributions in solving numerous nomenclatural problems, and interpreting the Code. Last, but no less important, I wish to express my gratitude to Joseph H. Kirkbride Jr. and two anonymous reviewers for the careful revision of the manuscript and their significant comments and corrections. Finally, I am very grateful to the Centro Studi Erbario Tropicale (FT herbarium), University of Florence (Italy) for financially supporting this publication.

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Bentham G. 1841. XVIII. Contribution towards a Flora of South America. – Enumeration of plants collected by Mr. Schomburgk in British Guiana. Journal of Botany (Hooker). 3: 212-221.

Bentham G. 1850. Plantae regnellianae - Rubiaceae. Linnaea. 23: 443-466.

Bentham G. 1853. On some new genera and species of Brazilian Rubiaceae. Hooker's Journal of Botany Kew Garden Miscellaneous. 5: 229-236.

Bentham G. 1857. Notes on Loganiaceae. Journal of the Proceedings of the Linnean Society, Botany. 1: 52-114.

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https://books.google.com.br/books?id=UNpCAAAAAcAAJ&printsec=frontcover&hl=fr&source=gbs_ge_summary_r&cad=0#v=onepage&q&f=false

Borgmeier T. 1961. A historia da "Flora Fluminensis" de Frei Vellozo. Vellozia. 1: 36-44.

Borhidi, A. 2017. La circuscripción de *Palicourea* subgen. *Heteropsychotria* (Rubiaceae Palicoureeae). Acta Botanica Hungarica. 59: 25-61.

Bremekamp, CEB. 1934. Notes on the Rubiaceae of Surinam. Recueil des Travaux Botaniques Néerlandais. 31: 248-308.

Bremekamp CEB. 1937. The Malaysian species of *Ixora* (Rubiaceae). Bulletin du Jardin Botanique de Buitenzorg, ser. 3. 14: 197-367.

Bridson D., Verdcourt B. 1988. *Coffea*. In: Polhill RM, Flora of Tropical East Africa, Rubiaceae (Part 2). Rotterdam, Published in the behalf of the East African Governments by A.A. Balkema, pp. 703-723.

Browne P. 1756. Civil and Natural History of Jamaica in three parts. London, Published by the Author, 503 pp., 49 pls.

Cabaña Fader AA, Salas RM, Dessein S, Cabral EL. 2016. Synopsis of *Hexasepalum* (Rubiaceae), the priority name for *Diodella* and a new species from Brazil. Systematic Botany. 41: 408-422. https:doi.org/10.1600/036364416X691803

Cabral EL. 1992 ["1991"). Rehabilitación del género *Galianthe* (Rubiaceae). Boletin de la Sociedad Argentina de Botánica. 27: 235-249.

Cabral EL. 2004. Dos especies nuevas de *Galianthe* (Rubiaceae) para Brasil. Bonplandia. 13: 15-19.

Cabral EL. 2009. Revisión sinóptica de *Galianthe* subgen. *Galianthe* (Rubiaceae: Spermacoceae), con una sección nueva. Annals of the Missouri Botanical Garden. 96: 27-50. https://doi.org/10.3417/2006193

Cabral EL, Bacigalupo NM. 2000. Novedades taxonómicas en *Galianthe* y *Borreria* (Rubiaceae-Spermacoceae). Bonplandia. 10: 119-128.

Cabral EL, Bacigalupo NM. 2005. Novelties in Spermacoceae (Rubiaceae) from Bolivia and Paraguay. Brittonia. 57: 129-140. https://www.jstor.org/stable/4099007

Cabral EL, Cabaña Fader AA. 2022. *Diodia*. In: Zuloaga FO, Zanotti CA (Eds.), Flora Argentina - Flora Vascular de la República de Argentina, vol. 19(3). Pasaje Garro, Talleres Trama S.A., pp. 254-255.

Cabral EL, Salas RM. 2022a. *Chomelia*. In: Zuloaga FO, Zanotti CA (Eds.), Flora Argentina - Flora Vascular de la República de Argentina, vol. 19(3). Pasaje Garro, Talleres Trama S.A., pp. 239-240.

Cabral EL, Salas RM. 2022b. *Coccocypselum*. In: Zuloaga FO, Zanotti CA (Eds.), Flora Argentina - Flora Vascular de la República de Argentina, vol. 19(3). Pasaje Garro, Talleres Trama S.A., pp. 240-242.

Cabral EL, Salas RM. 2022c. *Coussarea*. In: Zuloaga FO, Zanotti CA (Eds.), Flora Argentina - Flora Vascular de la República de Argentina, vol. 19(3). Pasaje Garro, Talleres Trama S.A., pp. 246-247.

Cabral EL, Salas RM. 2022d. *Hamelia*. In: Zuloaga FO, Zanotti CA (Eds.), Flora Argentina - Flora Vascular de la República de Argentina, vol. 19(3). Pasaje Garro, Talleres Trama S.A., pp. 300-301.

Cabral EL, Medina WA, Souza E B. 2009. Novedades en el género *Mitracarpus* (Spermacoceae-Rubiaceae) para la flora del Paraguay. Candollea. 64: 152-156. https://ri.conicet.gov.ar/handle/11336/36634

Cabral EL, Miguel LM, Salas RM. 2011. Dos especies nuevas de *Borreria* (Rubiaceae), sinopsis y clave de las especies para Bahia, Brasil. Acta Botanica Brasileira. 25: 255-276. https://doi.org/10.1590/S0102-33062011000200002

Candolle AP de [Sep.] 1830. Rubiaceae. Prodromus ..., vol. 4, pp. 341-622. Paris, Treuttel & Würtz.

Carauta JPP. 1969. A data efetiva da "Flora Fluminensis". Vellozia. 7: 26-33.

Carauta JPP. 1973. The text of Vellozo's Flora Fluminensis and its effective date of publication. Taxon. 22: 281-284. https://onlinelibrary.wiley.com/doi/abs/10.2307/1218138
Barbosa MR. 2007. *Guettarda*. In: MGL Wanderley (Coord.), Flora Fanerogâmica do Estado de São Paulo, vol. 5, pp. 346-348. São Paulo, Instituto de Botânica, Imprensa Oficial do Estado de São Paulo.

Bentham G. 1841. XVIII. Contribution towards a Flora of South America. – Enumeration of plants collected by Mr. Schomburgk in British Guiana. Journal of Botany (Hooker). 3: 212-221.

Bentham G. 1850. Plantae regnellianae - Rubiaceae. Linnaea. 23: 443-466.

Bentham G. 1853. On some new genera and species of Brazilian Rubiaceae. Hooker's Journal of Botany Kew Garden Miscellaneous. 5: 229-236.

Bentham G. 1857. Notes on Loganiaceae. Journal of the Proceedings of the Linnean Society, Botany. 1: 52-114.

Bolingbroke H. 1807. Voyage to the Demerary, containing a statistical account of the settlements there and those on the Essequebo, the Berbice, and other contiguous rivers of Guyana. London, R. Phillips.

https://www.biodiversitylibrary.org/item/164274#page/13/mode/1up

Bolingbroke H. 1809. Voyage to the Demerary, containing a statistical account of the settlements there and those on the Essequebo, the Berbice, and other contiguous rivers of Guyana. London, R. Phillips.

https://books.google.com.br/books?id=UNpCAAAAAcAAJ&printsec=frontcover&hl=fr&source=gbs_ge_summary_r&cad=0#v=onepage&q&f=false

Borgmeier T. 1961. A historia da "Flora Fluminensis" de Frei Vellozo. Vellozia. 1: 36-44.

Borhidi, A. 2017. La circuscripción de *Palicourea* subgen. *Heteropsychotria* (Rubiaceae Palicoureeae). Acta Botanica Hungarica. 59: 25-61.

Bremekamp, CEB. 1934. Notes on the Rubiaceae of Surinam. Recueil des Travaux Botaniques Néerlandais. 31: 248-308.

Bremekamp CEB. 1937. The Malaysian species of *Ixora* (Rubiaceae). Bulletin du Jardin Botanique de Buitenzorg, ser. 3. 14: 197-367.

Bridson D., Verdcourt B. 1988. *Coffea*. In: Polhill RM, Flora of Tropical East Africa, Rubiaceae (Part 2). Rotterdam, Published in the behalf of the East African Governments by A.A. Balkema, pp. 703-723.

Browne P. 1756. Civil and Natural History of Jamaica in three parts. London, Published by the Author, 503 pp., 49 pls.

Cabaña Fader AA, Salas RM, Dessein S, Cabral EL. 2016. Synopsis of *Hexasepalum* (Rubiaceae), the priority name for *Diodella* and a new species from Brazil. Systematic Botany. 41: 408-422. https:doi.org/10.1600/036364416X691803

Cabral EL. 1992 ["1991"). Rehabilitación del género *Galianthe* (Rubiaceae). Boletín de la Sociedad Argentina de Botánica. 27: 235-249.

Cabral EL. 2004. Dos especies nuevas de *Galianthe* (Rubiaceae) para Brasil. Bonplandia. 13: 15-19.

Cabral EL. 2009. Revisión sinóptica de *Galianthe* subgen. *Galianthe* (Rubiaceae: Spermacoceae), con una sección nueva. Annals of the Missouri Botanical Garden. 96: 27-50. https://doi.org/10.3417/2006193

Cabral EL, Bacigalupo NM. 2000. Novedades taxonómicas en *Galianthe* y *Borreria* (Rubiaceae-Spermacoceae). Bonplandia. 10: 119-128.

Cabral EL, Bacigalupo NM. 2005. Novelties in Spermacoceae (Rubiaceae) from Bolivia and Paraguay. Brittonia. 57: 129-140. https://www.jstor.org/stable/4099007

Cabral EL, Cabaña Fader AA. 2022. *Diodia*. In: Zuloaga FO, Zanotti CA (Eds.), Flora Argentina - Flora Vascular de la República de Argentina, vol. 19(3). Pasaje Garro, Talleres Trama S.A., pp. 254-255.

Cabral EL, Salas RM. 2022a. *Chomelia*. In: Zuloaga FO, Zanotti CA (Eds.), Flora Argentina - Flora Vascular de la República de Argentina, vol. 19(3). Pasaje Garro, Talleres Trama S.A., pp. 239-240.

Cabral EL, Salas RM. 2022b. *Coccocypselum*. In: Zuloaga FO, Zanotti CA (Eds.), Flora Argentina - Flora Vascular de la República de Argentina, vol. 19(3). Pasaje Garro, Talleres Trama S.A., pp. 240-242.

Cabral EL, Salas RM. 2022c. *Coussarea*. In: Zuloaga FO, Zanotti CA (Eds.), Flora Argentina - Flora Vascular de la República de Argentina, vol. 19(3). Pasaje Garro, Talleres Trama S.A., pp. 246-247.

Cabral EL, Salas RM. 2022d. *Hamelia*. In: Zuloaga FO, Zanotti CA (Eds.), Flora Argentina - Flora Vascular de la República de Argentina, vol. 19(3). Pasaje Garro, Talleres Trama S.A., pp. 300-301.

Cabral EL, Medina WA, Souza E B. 2009. Novedades en el género *Mitracarpus* (Spermacoceae-Rubiaceae) para la flora del Paraguay. Candollea. 64: 152-156. https://ri.conicet.gov.ar/handle/11336/36634

Cabral EL, Miguel LM, Salas RM. 2011. Dos especies nuevas de *Borreria* (Rubiaceae), sinopsis y clave de las especies para Bahia, Brasil. Acta Botanica Brasileira. 25: 255-276. https://doi.org/10.1590/S0102-33062011000200002

Candolle AP de [Sep.] 1830. Rubiaceae. Prodromus ..., vol. 4, pp. 341-622. Paris, Treuttel & Würtz.

Carauta JPP. 1969. A data efetiva da "Flora Fluminensis". Vellozia. 7: 26-33.

Carauta JPP. 1973. The text of Vellozo's Flora Fluminensis and its effective date of publication. Taxon. 22: 281-284. https://onlinelibrary.wiley.com/doi/abs/10.2307/1218138
Carmo JAM, Sala RM, Sobrado SV, Simões AO. 2019. (2707) Proposal to conserve the name *Psyllocarpus* against *Tapanhuacanga* (Rubiaceae: Spermacoceae). Taxon. 68: 869-871. https://doi.org/10.1002/tax.12109

Chamisso A. 1834. Spicilegium plantarum e familis jam prius recesitis, praesertim Brasiliensium. Rubiaceae. Linnaea. 9: 214-261.

Chamisso A, Schlechtendal D. 1828. De plantis in expedizione speculatoria Romanzoffiana observatis disserere pergut. Rubiaceae. Sectio I. Stellatae & Sectio III. Spermacoceae. Linnaea. 3: 309-366.

Chamisso A, Schlechtendal D. 1829a. De plantis in expedizione speculatoria Romanzoffiana observatis disserere pergut. Rubiaceae: Machaonia. - Sectio IV. Psychotriaceae. Linnaea. 4: 1-36.

Chamisso A, Schlechtendal D. 1829b. De plantis in expedizione speculatoria Romanzoffiana observatis disserere pergut. Rubiaceae. Quae supersunt. Linnaea. 4: 129-434.

Chodat R, Hassler É. 1904. Plantae Hasslerianae soit Enumeration des plantes récoltées au Paraguay par le Dr Émile Hassler, d'Aarau (Suisse) de 1885 à 1902. Bulletin de l'Herbier Boissier. 2: 61-92.

Coelho GF. 2008. Expedição histórica nos sertões de Goiás: São José do Duro. Goiânia, ICBC, Reedição ampliada, 168 pp.

Corner EJH. 1941. Notes on the systematy and distribution of Malayan Phanerogams, IV. *Ixora*. The Gardens' Bulletin, Straits Settlements. 21: 177-236.

Costa CB. 2004. Revisão taxonômica de *Coccocypselum* P. Br. (Rubiaceae). Unpublished Doctoral Thesis. São Paulo, Universidade de São Paulo, 154 pp.

Costa CB, Delprete PG. 2010. *Coccocypselum*. In: Delprete PG, Rubiaceae, Parte 1: Introdução, Gêneros A-H. In: Rizzo JA (Coord.), Flora dos Estados de Goiás e Tocantins, vol. 40. Goiânia, Universidade Federal de Goiás, IRD and UFG, Editora e Grafica Vieira, pp. 174-211.

Cruls L. 1894. Relatório da Comissão exploradora do Planalto Central do Brazil. Rio de Janeiro, H. Lombaerts & C., Impressores do Observatorio.

D'Arcy WG. 1970. Jacquin names, some notes on their typification. Taxon. 19: 554-560. https://doi.org/10.2307/1218948

Delprete PG. 1997. Revision and typification of Brazilian *Augusta* (Rubiaceae, Rondeletieae), with ecological observation on riverine vegetation of Cerrado and Atlantic Forest. Brittonia. 49: 487-497. https://doi.org/10.2307/2807739

Delprete PG. 1999a. Rondeletieae (Rubiaceae), Part I (*Rustia*, *Tresanthera*, *Condaminea*, *Picardaea*, *Pogonopus*, *Chimarrhis*, *Dioicodendron*, *Molopanthera*, *Doli-

*chodelphys*, and *Parachimarrhis*). Flora Neotropica Monographs vol. 77. Bronx, New York Botanical Garden Press, 226 pp.

Delprete PG. 1999b. Rubiaceae-Condamineae. In: G. Harling & L. Andersson (Eds.), Flora of Ecuador vol. 62. Copenhagen, Council for Nordic Publications in Botany: 3-52.

Delprete PG. 2001. The Neotropical Rubiaceae collected by Humboldt and Bonpland and described by Bonpland, Kunth, and Roemer and Schultes. In: Aymard GA (Ed.), Alexander Von Humboldt: Homenaje al Bicentenario de su Llegada a Tierras Venezolanas. BioLlania, Edición Especial N° 7: 22-51.

Delprete PG. 2003. Revision and typification of some species of *Ixora* (Rubiaceae) from central and southern Brazil. Sida. 20: 1471-1480. https://www.jstor.org/stable/41961005

Delprete PG. 2007. New combinations and new synonyms in the genus *Spermacoce* (Rubiaceae) for the Flora of Goiás and Tocantins (Brazil) and the Flora of the Guianas. Journal of the Botanical Research Institute of Texas. 1: 1023-1030. https://www.jstor.org/stable/41971535

Delprete PG. 2008a. A new species of *Chomeilia* (Rubiaceae, Guettardeae) from the Brazilian Planalto. Blumea. 53: 393-398. https://doi.org/10.3767/000651908X608034

Delprete PG. 2008b. Three new species of *Ixora* (Rubiaceae) from the state of Tocantins, Brazil. Journal of the Botanical Research Institute of Texas. 2: 455-462. https://www.jstor.org/stable/41971659

Delprete PG. 2008c. Revision of *Tocoyena* (Rubiaceae: Gardenieae) from the states of Goiás and Tocantins and a new species endemic to the white-sand areas in the Brazilian Cerrado. Journal of the Botanical Research Institute of Texas. 2: 983-993. https://www.jstor.org/stable/41971744

Delprete PG. 2010a. Rubiaceae, Parte 1: Introdução, Gêneros A-H. In: Rizzo JA (Coord.), Flora dos Estados de Goiás e Tocantins, vol. 40. Goiânia, Universidade Federal de Goiás, IRD and UFG, Editora e Grafica Vieira, pp. 1-580.

Delprete PG. 2010b. Rubiaceae, Parte 2: Gêneros I-R. In: Rizzo JA (Coord.), Flora dos Estados de Goiás e Tocantins, vol. 40. Goiânia, Universidade Federal de Goiás, IRD and UFG, Editora e Grafica Vieira, pp. 581-1098.

Delprete PG. 2010c. Rubiaceae, Parte 3: Gêneros S-W, Índices. In: Rizzo JA (Coord.), Flora dos Estados de Goiás e Tocantins, vol. 40. Goiânia, Universidade Federal de Goiás, IRD and UFG, Editora e Grafica Vieira, pp. 1099-1610.
Carmo JAM, Sala RM, Sobrado SV, Simões AO. 2019. (2707) Proposal to conserve the name *Psyllocarpus* against *Tapanhuacanga* (Rubiaceae: Spermacoceae). Taxon. 68: 869-871. https://doi.org/10.1002/tax.12109

Chamisso A. 1834. Spicilegium plantarum e familis jam prius recesitis, praesertim Brasiliensium. Rubiaceae. Linnaea. 9: 214-261.

Chamisso A, Schlechtendal D. 1828. De plantis in expe-
dizione speculatoria Romanzoffiana observatis dis-
serere pergut. Rubiaceae. Sectio I. Stellatae & Sectio
III. Spermacoceae. Linnaea. 3: 309-366.

Chamisso A, Schlechtendal D. 1829a. De plantis in expe-
dizione speculatoria Romanzoffiana observatis dis-
serere pergut. Rubiaceae: Machaonia. – Sectio IV.
Psychotriaceae. Linnaea. 4: 1-36.

Chamisso A, Schlechtendal D. 1829b. De plantis in expe-
dizione speculatoria Romanzoffiana observatis dis-
serere pergut. Rubiaceae. Quae supersunt. Linnaea. 4:
129-434.

Chodat R, Hassler É. 1904. Plantae Hasslerianae soit Enu-
meration des plantes récoltées au Paraguay par le Dr
Émile Hassler, d'Aarau (Suisse) de 1885 à 1902. Bul-
letin de l'Herbier Boissier. 2: 61-92.

Coelho GF. 2008. Expedição histórica nos sertões de Goiás: São José do Duro. Goiânia, ICBC, Reedição ampliada, 168 pp.

Corner EJH. 1941. Notes on the systematy and distribu-
tion of Malayan Phanerogams, IV. *Ixora*. The Gar-
dens' Bulletin, Straits Settlements. 21: 177-236.

Costa CB. 2004. Revisão taxonômica de *Coccocypselum* P. Br. (Rubiaceae). Unpublished Doctoral Thesis. São Paulo, Universidade de São Paulo, 154 pp.

Costa CB, Delprete PG. 2010. *Coccocypselum*. In: Delprete PG, Rubiaceae, Parte 1: Introdução, Gêneros A-H. In: Rizzo JA (Coord.), Flora dos Estados de Goiás e Tocantins, vol. 40. Goiânia, Universidade Federal de Goiás, IRD and UFG, Editora e Grafica Vieira, pp. 174-211.

Cruls L. 1894. Relatório da Comissão exploradora do Pla-
nalto Central do Brazil. Rio de Janeiro, H. Lombaerts
& C., Impressores do Observatorio.

D'Arcy WG. 1970. Jacquin names, some notes on their typification. Taxon. 19: 554-560. https://doi.org/10.2307/1218948

Delprete PG. 1997. Revision and typification of Brazilian *Augusta* (Rubiaceae, Rondeletieae), with ecological observation on riverine vegetation of Cerrado and Atlantic Forest. Brittonia. 49: 487-497. https://doi.org/10.2307/2807739

Delprete PG. 1999a. Rondeletieae (Rubiaceae), Part I (Rustia, Tresanthera, Condaminea, Picardaea, Pogonopus, Chimarrhis, Dioicodendron, Molopanthera, Doli-

*chodelphys*, and *Parachimarrhis*). Flora Neotropica
Monographs vol. 77. Bronx, New York Botanical Gar-
den Press, 226 pp.

Delprete PG. 1999b. Rubiaceae-Condamineae. In: G. Harling & L. Andersson (Eds.), Flora of Ecuador vol. 62. Copenhagen, Council for Nordic Publications in Botany: 3-52.

Delprete PG. 2001. The Neotropical Rubiaceae col-
lected by Humboldt and Bonpland and described
by Bonpland, Kunth, and Roemer and Schultes. In:
Aymard GA (Ed.), Alexander Von Humboldt: Hom-
enaje al Bicentenario de su Llegada a Tierras Vene-
zolanas. BioLlania, Edición Especial N° 7: 22-51.

Delprete PG. 2003. Revision and typification of some spe-
cies of *Ixora* (Rubiaceae) from central and southern
Brazil. Sida. 20: 1471-1480. https://www.jstor.org/sta-
ble/41961005

Delprete PG. 2007. New combinations and new synony-
mies in the genus *Spermacoce* (Rubiaceae) for the
Flora of Goiás and Tocantins (Brazil) and the Flora
of the Guianas. Journal of the Botanical Research
Institute of Texas. 1: 1023-1030. https://www.jstor.
org/stable/41971535

Delprete PG. 2008a. A new species of *Chome-
lia* (Rubiaceae, Guettardeae) from the Brazil-
ian Planalto. Blumea. 53: 393-398. https://doi.
org/10.3767/000651908X608034

Delprete PG. 2008b. Three new species of *Ixora* (Rubiaceae) from the state of Tocantins, Brazil. Journal of the Botanical Research Institute of Texas. 2: 455-462. https://www.jstor.org/stable/41971659

Delprete PG. 2008c. Revision of *Tocoyena* (Rubiaceae: Gardenieae) from the states of Goiás and Tocantins and a new species endemic to the white-sand areas in the Brazilian Cerrado. Journal of the Botanical Research Institute of Texas. 2: 983-993. https://www.jstor.org/stable/41971744

Delprete PG. 2010a. Rubiaceae, Parte 1: Introdução, Gêneros A-H. In: Rizzo JA (Coord.), Flora dos Estados de Goiás e Tocantins, vol. 40. Goiânia, Universidade Federal de Goiás, IRD and UFG, Editora e Grafica Vieira, pp. 1-580.

Delprete PG. 2010b. Rubiaceae, Parte 2: Gêneros I-R. In: Rizzo JA (Coord.), Flora dos Estados de Goiás e Tocantins, vol. 40. Goiânia, Universidade Federal de Goiás, IRD and UFG, Editora e Grafica Vieira, pp. 581-1098.

Delprete PG. 2010c. Rubiaceae, Parte 3: Gêneros S-W, Índices. In: Rizzo JA (Coord.), Flora dos Estados de Goiás e Tocantins, vol. 40. Goiânia, Universidade Federal de Goiás, IRD and UFG, Editora e Grafica Vieira, pp. 1099-1610.
Delprete PG. 2015. Typification and etymology of Aublet's Rubiaceae names. Taxon. 64: 595-624. https://www.jstor.org/stable/taxon.64.3.595

Delprete PG. 2022. Monograph of tribe Sipaneeae (Rubiaceae, Ixoroideae) - a Neotropical group with its center of diversity on the Guiana Shield. Webbia. 77(1), suppl. 1: 1-284. https://doi.org/10.36253/jopt-13963

Delprete PG, Cortés-B R. 2007 ["2006"). A synopsis of the Rubiaceae of the states of Mato Grosso and Mato Grosso do Sul, central-western Brazil, with a key to genera, and a preliminary species list. Revista de Biologia Neotropical. 3: 13-96. https://doi.org/10.5216/rbn.v3i1.2813

Delprete PG, Kirkbride JH Jr. 2016. New combinations and new names in *Palicourea* (Rubiaceae) for species of *Psychotria* subgenus *Heteropsychotria* occurring in the Guianas. Journal of the Botanical Research Institute of Texas. 10: 409-442. https://www.jstor.org/stable/44858580

Delprete PG, Persson C. 2010. *Alibertia*. In: Delprete PG, Rubiaceae, Parte 1: Introdução, Gêneros A-H. In: Rizzo JA (Coord.), Flora dos Estados de Goiás e Tocantins, vol. 40. Goiânia, Universidade Federal de Goiás, IRD and UFG, Editora e Grafica Vieira, pp. 69-88.

Delprete PG, Persson C. 2011. Alibertia Group. In: Taylor CM, Sanchez-Gonzalez J, Hammel B, Lorence DH, Persson C, Delprete PG, Gereau RE, Rubiacearum americanarum magna hama pars XXVIII: new taxa, new combinations, new names, and lectotypification for several species found in Mexico and Central America. Novon. 21: 133-148. https://www.jstor.org/stable/23018508

Delprete PG, Persson C. 2017. *Alibertia*. Pp. 57-118. In: Persson CH, Delprete PG. The Alibertia group (Gardenieae - Rubiaceae) - Part 1. Flora Neotropica Monographs vol. 119. Bronx, New York Botanical Garden Press.

Delprete PG, Smith LB, Klein RB. 2004. Rubiáceas, Volume 1 - Gêneros de A-G: 1. *Alseis* até 19. *Galium*. In: Reis A (Ed.), Flora Ilustrada Catarinense. Itajaí, Herbário Barbosa Rodrigues, pp. 1-344.

Delprete PG, Smith LB, Klein RB. 2005. Rubiáceas, Volume 2 - Gêneros de G-Z: 20. *Gardenia* até 46. *Tocoyena*. In: Reis A (ed.), Flora Ilustrada Catarinense. Itajaí, Herbário Barbosa Rodrigues, pp. 345-843.

Dempster LT. 1990. The genus *Galium* in South America. IV. Allertonia. 5: 283-345. https://www.jstor.org/stable/23188164

Devecchi, M. F. & J. R. Pirani. 2021. Simaroubaceae. In: Rizzo JA, Gomes Klein VL (Coord.), Flora dos Estados de Goiás e Tocantins, vol. 50. Goiânia, Cegraf UFG (Universidade Federal de Goiás), 45 pp.

Dillenius JJ. 1732. Hortus Elthamensis, 2 vols. London, published by the Author, 437 pp. + 324 pls.

Don G. 1834. A general history of the dichlamydeous plants, vol. 3. London, J.G. & F. Rivington, 867 pp.

Dwyer JD. 1980. In: Woodson E et al. (eds.), Flora of Panama, part IX. Family 179. Rubiaceae. Annals of the Missouri Botanical Garden. 67: 1-522. https://doi.org/10.2307/2398954

Ehrendorfer F. 1955. Revision of the genus *Relbunium* (Endl.) Benth. & Hook. (Rubiaceae-Galieae). Botanische Jahrbücher fur Systematik. 76: 516-553.

Ek RC. 1990. Index of Guyana plant collectors. In: ARA Görts-van Rijn (ed.), Flora of the Guianas, Supplementary Series, Fascicle 1. Koenigstein, Koetz Scientific Books, 85 pp.

Elias TS. 1976. A monograph of the genus *Hamelia* (Rubiaceae). Memoirs of the New York Botanical Garden. 26: 81-144.

Ellis J. 1761. LXXXII. An account on the plants *Halesia* and *Gardenia*: In a letter from John Ellis, Esq; F.R.S. to Philip Carteret Webb, Esq; R.R.S. Philosophical Transactions of the Royal Society of London. 51: 929-935, pls. 22-23.

Estrella E. 1991. La Naturaleza Americana y las Expediciones Botánicas Españolas del Siglo XVIII. In: Cevallos IC (ed.), Flora Huayaquilensis, La Espedición Botánicas de Juan Tafalla a la Real Audiencia de Quito, 1799-1808. Quito, Abya-Yala Editing.

Eschwege WL. 1822. Geognostisches Gemälde von Brasilien und wahrscheinliches Muttergestein der Diamanten. Weimar, Landes-Industrie-Comptoir, 44 pp.

Fleischmann A, Gonella PM. 2020. Typification and authorship of *Drosera intermedia* (Droseraceae). Taxon. 69: 153-160. https://doi.org/10.1002/tax.12158

Florentín JE, Cabral EL, Salas RM. 2022. Lectotypificaciones de nombres ligados a *Galianthe* (Rubiaceae) para la flora de Argentina. Darwiniana, n.s. 10: 29-34. https://doi.org/10.14522/darwiniana.2022.101.1007

Florentín MN, Florentín JE, Salas RM. 2020. Integrative taxonomic analyses sheds light on three historically disputed American *Spermacoce* species, and a key to the American species of *Spermacoce* (Spermacoceae, Rubiaceae). Systematic Botany. 45: 585-606. https://doi.org/10.1600/036364420X15935294613464

Forsskål P. 1775. Flora Aegyptiaco-Arabica: sive descriptions plantarum. Hauniae, Ex Officina Mölleri.

Fosberg FR, Sachet H.-H. 1989a. Three cultivated Ixoras (Rubiaceae). Baileya. 23: 74-85.
Delprete PG. 2015. Typification and etymology of Aublet's Rubiaceae names. Taxon. 64: 595-624. https://www.jstor.org/stable/taxon.64.3.595

Delprete PG. 2022. Monograph of tribe Sipaneeae (Rubiaceae, Ixoroideae) - a Neotropical group with its center of diversity on the Guiana Shield. Webbia. 77(1), suppl. 1: 1-284. https://doi.org/10.36253/jopt-13963

Delprete PG, Cortés-B R. 2007 ["2006"). A synopsis of the Rubiaceae of the states of Mato Grosso and Mato Grosso do Sul, central-western Brazil, with a key to genera, and a preliminary species list. Revista de Biologia Neotropical. 3: 13-96. https://doi.org/10.5216/rbn.v3i1.2813

Delprete PG, Kirkbride JH Jr. 2016. New combinations and new names in *Palicourea* (Rubiaceae) for species of *Psychotria* subgenus *Heteropsychotria* occurring in the Guianas. Journal of the Botanical Research Institute of Texas. 10: 409-442. https://www.jstor.org/stable/44858580

Delprete PG, Persson C. 2010. *Alibertia*. In: Delprete PG, Rubiaceae, Parte 1: Introdução, Gêneros A-H. In: Rizzo JA (Coord.), Flora dos Estados de Goiás e Tocantins, vol. 40. Goiânia, Universidade Federal de Goiás, IRD and UFG, Editora e Grafica Vieira, pp. 69-88.

Delprete PG, Persson C. 2011. Alibertia Group. In: Taylor CM, Sanchez-Gonzalez J, Hammel B, Lorence DH, Persson C, Delprete PG, Gereau RE, Rubiacearum americanarum magna hama pars XXVIII: new taxa, new combinations, new names, and lectotypification for several species found in Mexico and Central America. Novon. 21: 133-148. https://www.jstor.org/stable/23018508

Delprete PG, Persson C. 2017. *Alibertia*. Pp. 57-118. In: Persson CH, Delprete PG. The Alibertia group (Gardenieae - Rubiaceae) - Part 1. Flora Neotropica Monographs vol. 119. Bronx, New York Botanical Garden Press.

Delprete PG, Smith LB, Klein RB. 2004. Rubiáceas, Volume 1 - Gêneros de A-G: 1. *Alseis* até 19. *Galium*. In: Reis A (Ed.), Flora Ilustrada Catarinense. Itajaí, Herbário Barbosa Rodrigues, pp. 1-344.

Delprete PG, Smith LB, Klein RB. 2005. Rubiáceas, Volume 2 - Gêneros de G-Z: 20. *Gardenia* até 46. *Tocoyena*. In: Reis A (ed.), Flora Ilustrada Catarinense. Itajaí, Herbário Barbosa Rodrigues, pp. 345-843.

Dempster LT. 1990. The genus *Galium* in South America. IV. Allertonia. 5: 283-345. https://www.jstor.org/stable/23188164

Devecchi, M. F. & J. R. Pirani. 2021. Simaroubaceae. In: Rizzo JA, Gomes Klein VL (Coord.), Flora dos Estados de Goiás e Tocantins, vol. 50. Goiânia, Cegraf UFG (Universidade Federal de Goiás), 45 pp.

Dillenius JJ. 1732. Hortus Elthamensis, 2 vols. London, published by the Author, 437 pp. + 324 pls.

Don G. 1834. A general history of the dichlamydeous plants, vol. 3. London, J.G. & F. Rivington, 867 pp.

Dwyer JD. 1980. In: Woodson E et al. (eds.), Flora of Panama, part IX. Family 179. Rubiaceae. Annals of the Missouri Botanical Garden. 67: 1-522. https://doi.org/10.2307/2398954

Ehrendorfer F. 1955. Revision of the genus *Relbunium* (Endl.) Benth. & Hook. (Rubiaceae-Galieae). Botanische Jahrbücher fur Systematik. 76: 516-553.

Ek RC. 1990. Index of Guyana plant collectors. In: ARA Görts-van Rijn (ed.), Flora of the Guianas, Supplementary Series, Fascicle 1. Koenigstein, Koetz Scientific Books, 85 pp.

Elias TS. 1976. A monograph of the genus *Hamelia* (Rubiaceae). Memoirs of the New York Botanical Garden. 26: 81-144.

Ellis J. 1761. LXXXII. An account on the plants *Halesia* and *Gardenia*: In a letter from John Ellis, Esq; F.R.S. to Philip Carteret Webb, Esq; R.R.S. Philosophical Transactions of the Royal Society of London. 51: 929-935, pls. 22-23.

Estrella E. 1991. La Naturaleza Americana y las Expediciones Botánicas Españolas del Siglo XVIII. In: Cevallos IC (ed.), Flora Huayaquilensis, La Espedición Botánicas de Juan Tafalla a la Real Audiencia de Quito, 1799-1808. Quito, Abya-Yala Editing.

Eschwege WL. 1822. Geognostisches Gemälde von Brasilien und wahrscheinliches Muttergestein der Diamanten. Weimar, Landes-Industrie-Comptoir, 44 pp.

Fleischmann A, Gonella PM. 2020. Typification and authorship of *Drosera intermedia* (Droseraceae). Taxon. 69: 153-160. https://doi.org/10.1002/tax.12158

Florentín JE, Cabral EL, Salas RM. 2022. Lectotypificaciones de nombres ligados a *Galianthe* (Rubiaceae) para la flora de Argentina. Darwiniana, n.s. 10: 29-34. https://doi.org/10.14522/darwiniana.2022.101.1007

Florentín MN, Florentín JE, Salas RM. 2020. Integrative taxonomic analyses sheds light on three historically disputed American *Spermacoce* species, and a key to the American species of *Spermacoce* (Spermacoceae, Rubiaceae). Systematic Botany. 45: 585-606. https://doi.org/10.1600/036364420X15935294613464

Forsskål P. 1775. Flora Aegyptiaco-Arabica: sive descriptions plantarum. Hauniae, Ex Officina Mölleri.

Fosberg FR, Sachet H.-H. 1989a. Three cultivated Ixoras (Rubiaceae). Baileya. 23: 74-85.
Fosberg FR, Sachet H.-H. 1989b. Lectotypification of *Ixora coccinea* L. (Rubiaceae). Taxon. 38: 486-489. https://doi.org/10.1002/j.1996-8175.1989.tb00989.x

Franck AR, Gann GD, Salle J, Farid A. 2021. Sharpening plant taxonomy in South Florida: *Baccharis* and *Melanthera* (Asteraceae), *Borreria* and *Chiococca* (Rubiaceae), and *Lantana* (Verbenaceae). Phytologia. 103: 29-68.

Fürnrohr, A. E. 1832. [*Cinchona cujabensis* Manso, nom. nud.] Nachricht die Sammlungen des Hrn. Dr. Lhotsky betreffend. Flora. 15(2), Beibl. 3: 25-[26]-40.

Galli U. 2017. Professor José Ângelo Rizzo: Mistura de árvore e gente. Goiânia, Editora Kelps, 190 pp.

Gardner G. 1975. Viagem ao interior do Brasil, principalmente nas províncias do Norte e nos distritos do ouro e do diamante durante os anos de 1836-1841; tradução de Milton Amado, apresentação de Mário Ferri. Belo Horizonte, Ed. Itatiaia; São Paulo, Ed. de Universidade de São Paulo. Reconquista do Brasil, vol. 13, 260 pp.

Gauto SY, Florentin MN, Gonzalez AM, Salas RM. 2022. Tipificación de nombres en *Manettia* (Rubiaceae, Spermacoceae) para la Flora de Argentina y Paraguay. Darwiniana. 10: 429-434. https://doi.org/10.14522/darwiniana.2022.102.1078

Glaziou AFM. 1905. Liste des plantes du Brésil Central recuillies en 1861-1865. Bulletin de la Société Botanique de France 56, Mémoires 3a: 1-112 (Introduction, Fam. 1: Renonculacées to Fam. 42: Ampélidées). https://www.biodiversitylibrary.org/page/3461395#page/120/mode/1up

Glaziou AFM. 1906. Liste des plantes du Brésil Central recuillies en 1861-1865. Bulletin de la Société Botanique de France 56, Mémoires 3b: 113-200 (Fam. 42: Ampélidées to Fam. 50: Crassulacées).

Glaziou AFM. 1907. Liste des plantes du Brésil Central recuillies en 1861-1865. Bulletin de la Société Botanique de France 56, Mémoires 3c: 201-296 (Fam. 51: Droséracées to Fam. 56: Mélastomacées).

Glaziou AFM. 1909a. Liste des plantes du Brésil Central recuillies en 1861-1865. Bulletin de la Société Botanique de France 56, Mémoires 3d: 297-328 (Fam. 56: Mélastomacées to Fam. 67: Ombellifères).

Glaziou AFM. 1909b. Liste des plantes du Brésil Central recuillies en 1861-1865. Bulletin de la Société Botanique de France 56, Mémoires 3d: 329-392 (Fam. 67: Ombellifères to Fam. 74: Composées [Fam. 71: Rubiaceées pp. 334-365]).

Glaziou AFM. 1910. Liste des plantes du Brésil Central recuillies en 1861-1865. Bulletin de la Société Botanique de France 56, Mémoires 3e: 393-488 (Fam. 74: Composées to Fam. 93. Convolvulacées).

Glaziou AFM. 1911. Liste des plantes du Brésil Central recuillies en 1861-1865. Bulletin de la Société Botanique de France 56, Mémoires 3f: 489-661 (93. Convolvulacées to Fam. 129. Cératophyllacées, Errata, Table Alphabetique des familles).

Gomes BA. 1801. Memoria sobre a ipecacuanha fusa do Brasil, ou cipó das nossas boticas. Typografia Chalcographica, Typoplastica e Litteraria do Arco do Cego, Lisboa. https://www.abebooks.com/Plantas-Medicinais-Brasil-Bernardino-Antonio-Gomes/30592978589/bd#&gid=1&pid=4

Gomes M. 2003. Reavaliação taxonômica de algumas espécies dos gêneros *Coussarea* Aubl. e *Faramea* Aubl. (Rubiaceae). Acta Botanica Brasilica. 17: 449-466. https://doi.org/10.1590/S0102-33062003000300011

Gray A. 1860. Notes upon some Rubiaceae collected in the United States South-Sea Exploring Expedition under Captain Wilkes, with Characters of New Species, &c. Proceedings of the American Academy of Arts and Sciences. 4: 33-50.

Grisebach AHR. 1861. Rubiaceae. Flora of the British West Indian Islands, Part. 4. London, Loell Reeve & Co., pp. 315-410.

Grisebach AHR. 1879. Symbolae ad Floram Argentinam. Abhandlungen der Königlichen Gesellschaft der Wissenschaften zu Göttingen. 24: [3]-345.

Gustafsson CGR. 1998. The neotropical *Rosenbergiodendron* (Rubiaceae, Gardenieae). Brittonia. 50: 452-466.

Heyne B. 1814. Tracts, historical and statistical, on India; with journals of several tours through various parts of the peninsula: also, An account of Sumatra, in a series of letters. London, printed for Robert Baldwin. https://archive.org/details/in.ernet.dli.2015.41907

Hiepko P. 2006. Humboldt, his botanical mentor Willdenow, and the fate of the collections of Humboldt & Bonpland. Botanische Jahrbücher fur Systematik. 126: 509-516. https://doi.org/10.1127/0006-8152/2006/0126-0509

Howard RA. 1989. Dicotyledoneae – Part 3. Flora of the Lesser Antilles, Leeward and Windward Islands, vol. 6. Arnold Arboretum, Harvard University, Jamaica Plain, 658 pp.

Husain T, Paul SR. 1989. Taxonomic studies on Indian species of genus *Ixora* L. (Rubiaceae). Journal of Economic and Taxonomic Botany, Additional Series. 6: 1-205.

Jacquin NJ. 1760. Enumeratio Systematica Plantarum, quas in insulis Caribaeis... Lugduni Bataviorum, Apud Theodorum Haak, 41 pp.

Jacquin NJ. 1763. Selectarum stirpium americanarum historia. Wien, ex Officina Krausiana, viii + 284 pp.

Jarvis C. 2007. Order out of Chaos: Linnaean plant names and their types. London, Linnean Society of London & Natural History Museum, 1016 pp.
Fosberg FR, Sachet H.-H. 1989b. Lectotypification of *Ixora coccinea* L. (Rubiaceae). Taxon. 38: 486-489. https://doi.org/10.1002/j.1996-8175.1989.tb00989.x

Franck AR, Gann GD, Salle J, Farid A. 2021. Sharpening plant taxonomy in South Florida: *Baccharis* and *Melanthera* (Asteraceae), *Borreria* and *Chiococca* (Rubiaceae), and *Lantana* (Verbenaceae). Phytologia. 103: 29-68.

Fürnrohr, A. E. 1832. [*Cinchona cujabensis* Manso, nom. nud.] Nachricht die Sammlungen des Hrn. Dr. Lhotsky betreffend. Flora. 15(2), Beibl. 3: 25-[26]-40.

Galli U. 2017. Professor José Ângelo Rizzo: Mistura de árvore e gente. Goiânia, Editora Kelps, 190 pp.

Gardner G. 1975. Viagem ao interior do Brasil, principalmente nas províncias do Norte e nos distritos do ouro e do diamante durante os anos de 1836-1841; tradução de Milton Amado, apresentação de Mário Ferri. Belo Horizonte, Ed. Itatiaia; São Paulo, Ed. de Universidade de São Paulo. Reconquista do Brasil, vol. 13, 260 pp.

Gauto SY, Florentin MN, Gonzalez AM, Salas RM. 2022. Tipificación de nombres en *Manettia* (Rubiaceae, Spermacoceae) para la Flora de Argentina y Paraguay. Darwiniana. 10: 429-434. https://doi.org/10.14522/darwiniana.2022.102.1078

Glaziou AFM. 1905. Liste des plantes du Brésil Central recuillies en 1861-1865. Bulletin de la Société Botanique de France 56, Mémoires 3a: 1-112 (Introduction, Fam. 1: Renonculacées to Fam. 42: Ampélidées). https://www.biodiversitylibrary.org/page/3461395#page/120/mode/1up

Glaziou AFM. 1906. Liste des plantes du Brésil Central recuillies en 1861-1865. Bulletin de la Société Botanique de France 56, Mémoires 3b: 113-200 (Fam. 42: Ampélidées to Fam. 50: Crassulacées).

Glaziou AFM. 1907. Liste des plantes du Brésil Central recuillies en 1861-1865. Bulletin de la Société Botanique de France 56, Mémoires 3c: 201-296 (Fam. 51: Droséracées to Fam. 56: Mélastomacées).

Glaziou AFM. 1909a. Liste des plantes du Brésil Central recuillies en 1861-1865. Bulletin de la Société Botanique de France 56, Mémoires 3d: 297-328 (Fam. 56: Mélastomacées to Fam. 67: Ombellifères).

Glaziou AFM. 1909b. Liste des plantes du Brésil Central recuillies en 1861-1865. Bulletin de la Société Botanique de France 56, Mémoires 3d: 329-392 (Fam. 67: Ombellifères to Fam. 74: Composées [Fam. 71: Rubiaceées pp. 334-365]).

Glaziou AFM. 1910. Liste des plantes du Brésil Central recuillies en 1861-1865. Bulletin de la Société Botanique de France 56, Mémoires 3e: 393-488 (Fam. 74: Composées to Fam. 93. Convolvulacées).

Glaziou AFM. 1911. Liste des plantes du Brésil Central recuillies en 1861-1865. Bulletin de la Société Botanique de France 56, Mémoires 3f: 489-661 (93. Convolvulacées to Fam. 129. Cératophyllacées, Errata, Table Alphabetique des familles).

Gomes BA. 1801. Memoria sobre a ipecacuanha fusa do Brasil, ou cipó das nossas boticas. Typografia Chalcographica, Typoplastica e Litteraria do Arco do Cego, Lisboa. https://www.abebooks.com/Plantas-Medicinais-Brasil-Bernardino-Antonio-Gomes/30592978589/bd#&gid=1&pid=4

Gomes M. 2003. Reavaliação taxonômica de algumas espécies dos gêneros *Coussarea* Aubl. e *Faramea* Aubl. (Rubiaceae). Acta Botanica Brasilica. 17: 449-466. https://doi.org/10.1590/S0102-33062003000300011

Gray A. 1860. Notes upon some Rubiaceae collected in the United States South-Sea Exploring Expedition under Captain Wilkes, with Characters of New Species, &c. Proceedings of the American Academy of Arts and Sciences. 4: 33-50.

Grisebach AHR. 1861. Rubiaceae. Flora of the British West Indian Islands, Part. 4. London, Loell Reeve & Co., pp. 315-410.

Grisebach AHR. 1879. Symbolae ad Floram Argentinam. Abhandlungen der Königlichen Gesellschaft der Wissenschaften zu Göttingen. 24: [3]-345.

Gustafsson CGR. 1998. The neotropical *Rosenbergiodendron* (Rubiaceae, Gardenieae). Brittonia. 50: 452-466.

Heyne B. 1814. Tracts, historical and statistical, on India; with journals of several tours through various parts of the peninsula: also, An account of Sumatra, in a series of letters. London, printed for Robert Baldwin. https://archive.org/details/in.ernet.dli.2015.41907

Hiepko P. 2006. Humboldt, his botanical mentor Willdenow, and the fate of the collections of Humboldt & Bonpland. Botanische Jahrbücher fur Systematik. 126: 509-516. https://doi.org/10.1127/0006-8152/2006/0126-0509

Howard RA. 1989. Dicotyledoneae – Part 3. Flora of the Lesser Antilles, Leeward and Windward Islands, vol. 6. Arnold Arboretum, Harvard University, Jamaica Plain, 658 pp.

Husain T, Paul SR. 1989. Taxonomic studies on Indian species of genus *Ixora* L. (Rubiaceae). Journal of Economic and Taxonomic Botany, Additional Series. 6: 1-205.

Jacquin NJ. 1760. Enumeratio Systematica Plantarum, quas in insulis Caribaeis... Lugduni Bataviorum, Apud Theodorum Haak, 41 pp.

Jacquin NJ. 1763. Selectarum stirpium americanarum historia. Wien, ex Officina Krausiana, viii + 284 pp.

Jarvis C. 2007. Order out of Chaos: Linnaean plant names and their types. London, Linnean Society of London & Natural History Museum, 1016 pp.
Judkevich MD, Salas RM, González AM. 2015. Revisión de *Randia* (Rubiaceae) en Argentina, taxonomía y morfoanatomía. Boletín de la Sociedad Argentina de Botánica. 50: 607-625. https://doi.org/10.31055/1851.2372.v50.n4.12920

Kainulainen K, Persson C, Eriksson T, Bremer B. 2010. Molecular systematics and morphological character evolution of the Condamineae. American Journal of Botany. 97: 1961-1981. https://doi.org/10.3732/ajb.1000090

Kirkbride JH Jr. 1976. A revision of the genus *Declieu-xia* (Rubiaceae). Memoirs of the New York Botanical Garden. 28: 1-87.

Kirkbride JH Jr. 1979. Revision of the genus *Psyllocarpus* (Rubiaceae). Smithsonian Contributions to Botany. 41: 1-52.

Kirkbride JH Jr. 1982. Rubiaceae types in the Triana collections at the Instituto de Ciencias Naturales, Universidad Nacional, Bogota, Colombia. Taxon. 31: 303-307. https://doi.org/10.2307/1219996

Kirkbride JH Jr., 1997. Manipulus rubiacearum-VI. Britain. 49: 354-379. https://doi.org/10.2307/2807836

Kirkbride JH Jr. 2014. *Hexasepalum teres* (Rubiaceae), a new combination. Journal of the Botanical Research Institute of Texas. 8: 17-18.

Kirkbride JH Jr., Delprete PG. 2012. A new variety of *Declieuxia cacuminis* (Rubiaceae) from the state of Tocantins, Brazil, and a review of the varieties recognized in the species. Journal of the Botanical Research Institue of Texas. 6: 391-396. https://www.jstor.org/stable/41972427

Kirkbride JH Jr., Delprete PG. 2015. New combinations in *Hexasepalum* (Rubiaceae, Spermacoceae). Journal of the Botanical Research Institue of Texas. 9: 103-106. https://www.jstor.org/stable/24621250

Kirkbride JH Jr., Wiersema JH. 2020. Nomenclature anomalies among Kunth's Rubiaceae. Taxon. 69: 605-612. https://doi.org/10.1002/tax.12183

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Lack HW. 2009. Alexander von Humboldt and the Botanical Exploration of the Americas, 1st ed. Munich, Prestl.

Lack HW. 2018. Alexander von Humboldt and the Botanical Exploration of the Americas, 2nd ed. Munich, Prestl.

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Stauffer FW, Stauffer J, Dorr LJ. 2012. Bonpland and Humboldt specimens, field notes, and herbaria; new insights from a study of the monocotyledons collected in Venezuela. Candollea. 67: 75-130.

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Taylor CM. 1989. Revision of *Palicourea* (Rubiaceae) in Mexico and Central America. Systematic Botany Monographs (St. Louis). 26: 1–102.

Taylor CM. 1999a. 162(18). Rubiaceae-Psychotrieae (1). In: Harling G, Andersson L (Eds.), Flora of Ecuador 62, 162 (part 3), Göteborg, University of Göteborg, Stockholm, Section of Botany, Riksmuseum, & Quito, Pontificia Universidad Católica del Ecuador; Trelleborg, Berlings, Skogs AB, pp. [133]–241.

Taylor CM. 1999b. 162(20). Rubiaceae-Coussareae. In: Harling G, Andersson L (Eds.), Flora of Ecuador 62, 162 (part 3). Göteborg, University of Göteborg, Stockholm, Section of Botany, Riksmuseum, & Quito, Pontificia Universidad Católica del Ecuador; Trelleborg, Berlings, Skogs AB, pp. [245]–314.

Taylor CM. 2005. *Margaritopsis* (Rubiaceae, Psychotrieae) in the Neotropics. Systematics and Geography of Plants. 75: 161–177. https://www.jstor.org/stable/3668574

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Taylor CM. 2012a. *Coccocypselum*. In: Davidse G, Sousa Sánchez M, Knapp S, Chiang Cabrera F (ed.), Flora Mesoamericana, vol. 4, part 2, Rubiaceae a Verbenaceae. Universidad Nacional Autónoma de México, Instituto de Biología, Missouri Botanical Garden, The Natural History Museum (London), Saint Louis, Missouri Botanical Garden Press, pp. 57–60.

Taylor CM. 2012b. *Palicourea*. In: Davidse G, Sousa Sánchez M, Knapp S, Chiang Cabrera F (ed.), Flora Mesoamericana, vol. 4, part 2, Rubiaceae a Verbenaceae. Universidad Nacional Autónoma de México, Instituto de Biología, Missouri Botanical Garden, The
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Taylor CM. 2012c. *Posoqueria*. In: Davidse G, Sousa Sánchez M, Knapp S, Chiang Cabrera F (ed.), Flora Mesoamericana, vol. 4, part 2, Rubiaceae a Verbenaceae. Universidad Nacional Autónoma de México, Instituto de Biología, Missouri Botanical Garden, The Natural History Museum (London), Saint Louis, Missouri Botanical Garden Press, pp. 188–191.

Taylor CM. 2012d. *Psychotria*. In: Davidse G, Sousa Sánchez M, Knapp S, Chiang Cabrera F (ed.), Flora Mesoamericana, vol. 4, part 2, Rubiaceae a Verbenaceae. Universidad Nacional Autónoma de México, Instituto de Biología, Missouri Botanical Garden, The Natural History Museum (London), Saint Louis, Missouri Botanical Garden Press, pp. 192–241.

Taylor CM. 2015. Rubiacearum Americanarum Magna Hama Pars XXXIV: The new group *Palcourea* sect. *Tricephalium* with eight new species and a new subspecies (Palcoureeae). Novon. 24: 55–95. https://doi.org/10.3417/2015001

Taylor CM. 2017. Rubiacearum Americanarum Magna Hama Pars XXXVI: New species and taxonomic changes in *Palcourea* (Palcoureeae). Novon. 25: 239–258. https://doi.org/10.3417/2015045

Taylor CM, Hollowell VC. 2016. Rubiacearum americanarum magna hama pars XXXV: The new group *Palcourea* sect. *Nonatelia*, with five new species (Palcoureeae). Novon. 25: 69–110. https://doi.org/10.3417/2015012

Taylor CM, Jardim JG. 2020. Rubiacearum americanarum magna hama pars XLVI: New species and taxonomic changes in *Faramea* of Central and South America (Rubiaceae, Coussareeae). Novon. 28: 108–142. https://doi.org/10.3417/2019552

Taylor CM, Neill DA, Gereau RE. 2011. Rubiacearum americanarum magna hama pars XXIX: Overview of the neotropical genus *Schizocalyx* (Condamineae) and description of two new species. Novon. 21: 496–507. https://doi.org/10.3417/2008095

Taylor CM, Gereau RE, Callmander M. 2020. The identity of *Mapouria* (Rubiaceae, Psychotrieae). Taxon. 69: 1072–1084. https://doi.org/10.1002/tax.12278

Taylor CM, Razafimandimbison SG, Barrabé L, Jardim JG., Barbosa MRV. 2017. *Eumachia* expanded, a pantropical genus distinct from *Psychotria* (Rubiaceae, Palcoureeae). Candollea. 72: 290–318. https://doi.org/10.15553/c2017v722a6

Tepe EJ. 2018. A series of unfortunate events: the forgotten botanist and the misattribution of a type collection. Phytokkeys. 109: 33–39. http://doi.org/10.3897/phytok Keys.109.28144

Turland NJ, Wiersema JH, Barrie FR, Greuter W, Hawsworth DL, Herendeen PS, Knapp S, Kusber W-H, Li D-Z, Marhold K, May TW, McNeill J, Monro AM, Prado J, Price MJ, Smith GF (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile, vol. 159. Glashütten: Koeltz Botanical Books, 254 pp. https://doi.org/10.12705/Code.2018.

Turner IM. 2021. Heyne, Roth, Roemer and Schultes, and the plant names published in *Novae plantarum species praesertim Indiae orientalis*. Taxon. 70: 365–428. https://doi.org/10.1002/tax.12449

Ule E. 1894. Relatorio do Dr. Ernesto Ule. Botânico da Comissão. Annexo VI. In L. Cruls (Ed.) Comissão exploradora do Planalto central do Brazil. Governo do Brasil, Rio de Janeiro: 339–365.

Ule E. 2003. Relatorio do Dr. Ernesto Ule. Botânico da Comissão. Annexo VI. Comissão Exploradora do Planalto Central do Brasil. Edição fac-similar. Edições do Senado Federal, vol. 22. Conselho Editorial, Brasília, DF, 380 pp.

Vandelli D. 1788. Florae Lusitanicae et Brasiliensis Specimen. Coimbra, Ex Typografia Academico-Regia, 96 pp.

Vegter IH. 1976. Index Herbariorum Part II Collectors M (4). Utrecht, M. Bohn, Scheltema & Holkema.

Vellozo JMC. 1829 [“1825”] Florae Fluminensis, seu descriptionum plantarum praefectura fluminensi sponte nascendum... Rio de Janeiro: Ex Typographia Nationali.

Vellozo JMC. 1831 [“1827”] Florae Fluminensis Icones nunc primo eduntur. Paris, Oficina Litográfica Senefelder, 11 vols.

Verdcourt B. 1975. Studies in the Rubiaceae–Rubioideae for the ‘Flora of Tropical East Africa’: I. Kew Bulletin. 30: 247–326. https://doi.org/10.2307/4102911

Verdcourt B. 1976. Rubiaceae (Part 1). In: Pohlill RM (Ed.), Flora of Tropical East Africa. Published on behalf of the East African Community by the Crown Agents for Oversea Governments, London, Whitefriars Press LTD., 414 pp.

Walpers WG. 1843. Rubiaceae. Novorum Actorum Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum 19, Suppl. 1: 349–357.

Walpers WG. 1846. [Cinchona cujabensis Manso, nom. nud.] Repertorium Botanices Rystematicae 6: 67. Fridesici Hofmeister, Leipzig.

Ward DB. 2008. Thomas Walter typification project, V: Neotypes and epotypes for 63 Walter names of genera D through Z. Journal of the Botanical Research Institute of Texas. 2: 475–486. https://www.jstor.org/stable/41971664
Natural History Museum (London), Saint Louis, Missouri Botanical Garden Press, pp. 163–181.

Taylor CM. 2012c. *Posoqueria*. In: Davidse G, Sousa Sánchez M, Knapp S, Chiang Cabrera F (ed.), Flora Mesoamericana, vol. 4, part 2, Rubiaceae a Verbenaceae. Universidad Nacional Autónoma de México, Instituto de Biología, Missouri Botanical Garden, The Natural History Museum (London), Saint Louis, Missouri Botanical Garden Press, pp. 188–191.

Taylor CM. 2012d. *Psychotria*. In: Davidse G, Sousa Sánchez M, Knapp S, Chiang Cabrera F (ed.), Flora Mesoamericana, vol. 4, part 2, Rubiaceae a Verbenaceae. Universidad Nacional Autónoma de México, Instituto de Biología, Missouri Botanical Garden, The Natural History Museum (London), Saint Louis, Missouri Botanical Garden Press, pp. 192–241.

Taylor CM. 2015. Rubiacearum Americanarum Magna Hama Pars XXXIV: The new group *Palcourea* sect. *Tricephalium* with eight new species and a new subspecies (Palcoureeae). Novon. 24: 55–95. https://doi.org/10.3417/2015001

Taylor CM. 2017. Rubiacearum Americanarum Magna Hama Pars XXXVI: New species and taxonomic changes in *Palcourea* (Palcoureeae). Novon. 25: 239–258. https://doi.org/10.3417/2015045

Taylor CM, Hollowell VC. 2016. Rubiacearum americanarum magna hama pars XXXV: The new group *Palcourea* sect. *Nonatelia*, with five new species (Palcoureeae). Novon. 25: 69–110. https://doi.org/10.3417/2015012

Taylor CM, Jardim JG. 2020. Rubiacearum americanarum magna hama pars XLVI: New species and taxonomic changes in *Faramea* of Central and South America (Rubiaceae, Coussareeae). Novon. 28: 108–142. https://doi.org/10.3417/2019552

Taylor CM, Neill DA, Gereau RE. 2011. Rubiacearum americanarum magna hama pars XXIX: Overview of the neotropical genus *Schizocalyx* (Condamineae) and description of two new species. Novon. 21: 496–507. https://doi.org/10.3417/2008095

Taylor CM, Gereau RE, Callmander M. 2020. The identity of *Mapouria* (Rubiaceae, Psychotrieae). Taxon. 69: 1072–1084. https://doi.org/10.1002/tax.12278

Taylor CM, Razafimandimbison SG, Barrabé L, Jardim JG., Barbosa MRV. 2017. *Eumachia* expanded, a pantropical genus distinct from *Psychotria* (Rubiaceae, Palcoureeae). Candollea. 72: 290–318. https://doi.org/10.15553/c2017v722a6

Tepe EJ. 2018. A series of unfortunate events: the forgotten botanist and the misattribution of a type collection. Phytokkeys. 109: 33–39. http://doi.org/10.3897/phytok Keys.109.28144

Turland NJ, Wiersema JH, Barrie FR, Greuter W, Hawsworth DL, Herendeen PS, Knapp S, Kusber W-H, Li D-Z, Marhold K, May TW, McNeill J, Monro AM, Prado J, Price MJ, Smith GF (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile, vol. 159. Glashütten: Koeltz Botanical Books, 254 pp. https://doi.org/10.12705/Code.2018.

Turner IM. 2021. Heyne, Roth, Roemer and Schultes, and the plant names published in *Novae plantarum species praesertim Indiae orientalis*. Taxon. 70: 365–428. https://doi.org/10.1002/tax.12449

Ule E. 1894. Relatorio do Dr. Ernesto Ule. Botânico da Comissão. Annexo VI. In L. Cruls (Ed.) Comissão exploradora do Planalto central do Brazil. Governo do Brasil, Rio de Janeiro: 339–365.

Ule E. 2003. Relatorio do Dr. Ernesto Ule. Botânico da Comissão. Annexo VI. Comissão Exploradora do Planalto Central do Brasil. Edição fac-similar. Edições do Senado Federal, vol. 22. Conselho Editorial, Brasília, DF, 380 pp.

Vandelli D. 1788. Florae Lusitanicae et Brasiliensis Specimen. Coimbra, Ex Typografia Academico-Regia, 96 pp.

Vegter IH. 1976. Index Herbariorum Part II Collectors M (4). Utrecht, M. Bohn, Scheltema & Holkema.

Vellozo JMC. 1829 [“1825”] Florae Fluminensis, seu descriptionum plantarum praefectura fluminensi sponte nascendum... Rio de Janeiro: Ex Typographia Nationali.

Vellozo JMC. 1831 [“1827”] Florae Fluminensis Icones nunc primo eduntur. Paris, Oficina Litográfica Senefelder, 11 vols.

Verdcourt B. 1975. Studies in the Rubiaceae–Rubioideae for the ‘Flora of Tropical East Africa’: I. Kew Bulletin. 30: 247–326. https://doi.org/10.2307/4102911

Verdcourt B. 1976. Rubiaceae (Part 1). In: Pohlill RM (Ed.), Flora of Tropical East Africa. Published on behalf of the East African Community by the Crown Agents for Oversea Governments, London, Whitefriars Press LTD., 414 pp.

Walpers WG. 1843. Rubiaceae. Novorum Actorum Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum 19, Suppl. 1: 349–357.

Walpers WG. 1846. [Cinchona cujabensis Manso, nom. nud.] Repertorium Botanices Rystematicae 6: 67. Fridesici Hofmeister, Leipzig.

Ward DB. 2008. Thomas Walter typification project, V: Neotypes and epotypes for 63 Walter names of genera D through Z. Journal of the Botanical Research Institute of Texas. 2: 475–486. https://www.jstor.org/stable/41971664
Weddell HA. 1854. Notice sur quelques Rubiacées de l'Amérique tropicale. Annales des Sciences Naturelles, Botanique, sér. 4. 1: 65-80, 10 pls.

Wernham HF. 1914. A monograph of the genus *Sabicea*. London, Longmans & Co., B. Quatrich, and Dulau & Co., and British Museum (Natural History), 82 pp.

Wiersema JH. 2015. Application of the name *Lemna punctata* G. Mey., the type of *Landoltia* Les & D. J. Crawford. Plant Biology (Stuttgart) 1: 5-9 (Epubl. 18 Jun. 2014). https://doi.org/10.1111/plb.12209

Wiersema JH, Delprete PG, Kirkbride JH Jr, Franck AR. 2017. A new weed in Florida, *Spermacoce latifolia*, and the distinction between *S. alata* and *S. latifolia* (Spermacoceae, Rubiaceae). Castanea. 82: 114-131. https://doi.org/10.2179/17-124

Willdenow CL. 1798. Caroli a Linné Species Plantarum, Editio Quarta, vol. 1, part 2. Berlin, Impensis G.C. Nauk, pp. [497]-1568.

Wurdack JJ. 1970. Erroneous data in Glaziou collections of Melastomataceae. Taxon. 19: 911-913. https://doi.org/10.2307/1218309

Zappi D. 2003. Revision of *Rudgea* (Rubiaceae) in southeastern and southern Brazil. Kew Bulletin. 58: 513-596. https://doi.org/10.2307/4111145

## APPENDIX 1. NEW TYPIFICATIONS IN THE PRESENT ARTICLE. ACCEPTED NAMES ARE IN **BOLD**

### Lectotypifications (81):

*Amaioua intermedia* Mart.

*Borreria angustifolia* Cham. & Schltdl. [≡ *Galianthe angustifolia* (Cham. & Schltdl.) E.L.Cabral].

*Borreria ovalifolia* M.Martens & Galeotti [≡ *Spermacoce ovalifolia* (M.Martens & Galeotti) Hemsl.].

*Borreria suaveolens* G.Mey. [≡ *Spermacoce suaveolens* (G. Mey.) Kuntze].

*Borreria umbellata* Spreng. [≡ *Emmeorhiza umbellata* (Spreng.) K.Schum.].

*Borreria vulpina* Standl. [≡ *Spermacoce vulpina* (Standl.) Govaerts].

*Borreria warmingii* K.Schum. [≡ *Spermacoce warmingii* (K.Schum.) Kuntze].

*Chomelia pohliana* Muell.Arg.

*Chomelia ribesioides* Benth. ex A.Gray

*Coccocypselum condalia* Pers.

*Coccocypselum erythrocephalum* Cham. & Schltdl.

*Coccocypselum hasslerianum* Chodat

*Condalia lanceolata* Ruiz & Pav. [≡ *Coccocypselum lanceolatum* (Ruiz & Pav.) Pers.].

*Coussarea platyphylla* Muell.Arg.

*Dialypetalanthus fuscescens* Kuhlmann

*Exostema formosum* Cham. & Schltdl. ex DC. [≡ *Rustia formosa* (Cham. & Schltdl. ex DC.) Klotzsch].

*Exostema formosum* var. β *laeve* DC. [≡ *Rustia formosa*].

*Faramea bracteata* Benth.

*Faramea contracta* Walp. [≡ *Coussarea contracta* (Walp.) Benth. & Hook. f. ex Müll. Arg.].

*Faramea hydrangeaefolia* Benth. [≡ *Coussarea hydrangeifolia* (Benth.) Benth. & Hook. f. ex Müll. Arg.].

*Faramea multiflora* A.Rich. ex DC.

*Faramea nitida* Benth.

*Ferdinandusa elliptica* Pohl

*Ferdinandusa ovalis* Pohl [= *Ferdinandusa elliptica* Pohl].

*Ferdinandusa pubescens* Wedd. [= *Ferdinandusa speciosa* Pohl].

*Ferdinandusa speciosa* Pohl

*Galium megapotamicum* Spreng.

*Gardenia longiflora* Ruiz & Pav. [≡ *Rosenbergiodendron longiflorum* (Ruiz. & Pav.) Fagerl.].

*Gardenia sessilis* Vell. [≡ *Cordiera sessilis* (Vell.) Kuntze].

*Guagnebina luteorubra* Vell. [≡ *Manettia luteorubra* (Vell.) Benth.].

*Guettarda burchelliana* Muell.Arg. [= *Guettarda pohliana*].

*Guettarda burchelliana* var. *opaca* Muell.Arg. [= *Guettarda pohliana*].

*Guettarda pohliana* Muell.Arg.

*Guettarda viburnoides* var. *genuina* Muell.Arg. [= *Guettarda viburnoides* Cham. & Schltdl.].

*Guettarda viburnoides* var. *pannosa* Muell.Arg. [= *Guettarda viburnoides* Cham. & Schltdl.].

*Guettarda viburnoides* var. *viburnoides* Muell.Arg. [= *Guettarda viburnoides* Cham. & Schltdl.]

*Ixora chinensis* Lam.

*Ixora pubescens* var. *glabrifolia* Muell.Arg. [= *Ixora pubescens* Willd. in Schult. & Schult. f.].

*Ladenbergia cujabensis* Klotzsch

*Malanea macrophylla* Bartl. ex Griseb.

*Malanea parviflora* Muell.Arg. [≡ *Chomelia parviflora* (Muell.Arg.) Muell. Arg.].

*Malanea ribesioides* var. *tomentosa* Muell. Arg. [= *Chomelia ribesioides* Benth. ex A.Gray].

*Malanea ribesioides* var. *villosa* Muell. Arg. [= *Chomelia ribesioides* Benth. ex A.Gray].

*Ophiorrhiza lanceolata* Forssk. [≡ *Pentas lanceolata* (Forssk.) Deflers].

*Pagamea plicata* Spruce ex Benth.

*Palicourea marcgravii* A.St.Hil.

*Palicourea marcgravii* var. *pubescens* A.St.Hil. [= *Palicourea marcgravii* A.St.Hil.].

*Palicourea urbaniana* Standl.

*Psychotria crocea* Sw. [≡ *Palicourea crocea* (Sw.) Roem. & Schult.].
Weddell HA. 1854. Notice sur quelques Rubiacées de l'Amérique tropicale. Annales des Sciences Naturelles, Botanique, sér. 4. 1: 65-80, 10 pls.

Wernham HF. 1914. A monograph of the genus *Sabicea*. London, Longmans & Co., B. Quatrich, and Dulau & Co., and British Museum (Natural History), 82 pp.

Wiersema JH. 2015. Application of the name *Lemna punctata* G. Mey., the type of *Landoltia* Les & D. J. Crawford. Plant Biology (Stuttgart) 1: 5-9 (Epubl. 18 Jun. 2014). https://doi.org/10.1111/plb.12209

Wiersema JH, Delprete PG, Kirkbride JH Jr, Franck AR. 2017. A new weed in Florida, *Spermacoce latifolia*, and the distinction between *S. alata* and *S. latifolia* (Spermacoceae, Rubiaceae). Castanea. 82: 114-131. https://doi.org/10.2179/17-124

Willdenow CL. 1798. Caroli a Linné Species Plantarum, Editio Quarta, vol. 1, part 2. Berlin, Impensis G.C. Nauk, pp. [497]-1568.

Wurdack JJ. 1970. Erroneous data in Glaziou collections of Melastomataceae. Taxon. 19: 911-913. https://doi.org/10.2307/1218309

Zappi D. 2003. Revision of *Rudgea* (Rubiaceae) in southeastern and southern Brazil. Kew Bulletin. 58: 513-596. https://doi.org/10.2307/4111145

## APPENDIX 1. NEW TYPIFICATIONS IN THE PRESENT ARTICLE. ACCEPTED NAMES ARE IN **BOLD**

### Lectotypifications (81):

*Amaioua intermedia* Mart.

*Borreria angustifolia* Cham. & Schltdl. [≡ *Galianthe angustifolia* (Cham. & Schltdl.) E.L.Cabral].

*Borreria ovalifolia* M.Martens & Galeotti [≡ *Spermacoce ovalifolia* (M.Martens & Galeotti) Hemsl.].

*Borreria suaveolens* G.Mey. [≡ *Spermacoce suaveolens* (G. Mey.) Kuntze].

*Borreria umbellata* Spreng. [≡ *Emmeorhiza umbellata* (Spreng.) K.Schum.].

*Borreria vulpina* Standl. [≡ *Spermacoce vulpina* (Standl.) Govaerts].

*Borreria warmingii* K.Schum. [≡ *Spermacoce warmingii* (K.Schum.) Kuntze].

*Chomelia pohliana* Muell.Arg.

*Chomelia ribesioides* Benth. ex A.Gray

*Coccocystalum condalia* Pers.

*Coccocystalum erythrocephalum* Cham. & Schltdl.

*Coccocystalum hasslerianum* Chodat

*Condalia lanceolata* Ruiz & Pav. [≡ *Coccocystalum lanceolatum* (Ruiz & Pav.) Pers.].

*Coussarea platyphylla* Muell.Arg.

*Dialypetalanthus fuscescens* Kuhlmann

*Exostema formosum* Cham. & Schltdl. ex DC. [≡ *Rustia formosa* (Cham. & Schltdl. ex DC.) Klotzsch].

*Exostema formosum* var. β *laeve* DC. [≡ *Rustia formosa*].

*Faramea bracteata* Benth.

*Faramea contracta* Walp. [≡ *Coussarea contracta* (Walp.) Benth. & Hook. f. ex Müll. Arg.].

*Faramea hydrangeaefolia* Benth. [≡ *Coussarea hydrangeifolia* (Benth.) Benth. & Hook. f. ex Müll. Arg.].

*Faramea multiflora* A.Rich. ex DC.

*Faramea nitida* Benth.

*Ferdinandusa elliptica* Pohl

*Ferdinandusa ovalis* Pohl [= *Ferdinandusa elliptica* Pohl].

*Ferdinandusa pubescens* Wedd. [= *Ferdinandusa speciosa* Pohl].

*Ferdinandusa speciosa* Pohl

*Galium megapotamicum* Spreng.

*Gardenia longiflora* Ruiz & Pav. [≡ *Rosenbergiodendron longiflorum* (Ruiz. & Pav.) Fagerl.].

*Gardenia sessilis* Vell. [≡ *Cordiera sessilis* (Vell.) Kuntze].

*Guagnebina luteorubra* Vell. [≡ *Manettia luteorubra* (Vell.) Benth.].

*Guettarda burchelliana* Muell.Arg. [= *Guettarda pohliana*].

*Guettarda burchelliana* var. *opaca* Muell.Arg. [= *Guettarda pohliana*].

*Guettarda pohliana* Muell.Arg.

*Guettarda viburnoides* var. *genuina* Muell.Arg. [= *Guettarda viburnoides* Cham. & Schltdl.].

*Guettarda viburnoides* var. *pannosa* Muell.Arg. [= *Guettarda viburnoides* Cham. & Schltdl.].

*Guettarda viburnoides* var. *viburnoides* Muell.Arg. [= *Guettarda viburnoides* Cham. & Schltdl.]

*Ixora chinensis* Lam.

*Ixora pubescens* var. *glabrifolia* Muell.Arg. [= *Ixora pubescens* Willd. in Schult. & Schult. f.].

*Ladenbergia cujabensis* Klotzsch

*Malanea macrophylla* Bartl. ex Griseb.

*Malanea parviflora* Muell.Arg. [≡ *Chomelia parviflora* (Muell.Arg.) Muell. Arg.].

*Malanea ribesioides* var. *tomentosa* Muell. Arg. [= *Chomelia ribesioides* Benth. ex A.Gray].

*Malanea ribesioides* var. *villosa* Muell. Arg. [= *Chomelia ribesioides* Benth. ex A.Gray].

*Ophiorrhiza lanceolata* Forssk. [≡ *Pentas lanceolata* (Forssk.) Deflers].

*Pagamea plicata* Spruce ex Benth.

*Palicourea marcgravii* A.St.Hil.

*Palicourea marcgravii* var. *pubescens* A.St.Hil. [= *Palicourea marcgravii* A.St.Hil.].

*Palicourea urbaniana* Standl.

*Psychotria crocea* Sw. [≡ *Palicourea crocea* (Sw.) Roem. & Schult.].
*Psychotria macrobotrys* Ruiz & Pav. [= **Palicourea macrobotrys** (Ruiz & Pav.) DC.].

*Psychotria microcephala* (Willd. ex Roem. & Schult.)
Muell.Arg. var. *tripotamica* Muell. Arg. [= **Palicourea prunifolia** (Kunth) Borhidi].

*Psychotria racemosa* Rich. [= **Palicourea racemosa** (Aubl.) G.Nicholson].

*Psychotria rupestris* Muell.Arg. [= **Palicourea rupestris** (Müll.Arg.) Delprete].

*Psychotria schuechiana* Müll.Arg. [= **Palicourea subtriflora** (Muell.Arg.) Delprete].

*Psychotria stachyoides* Benth. [= **Palicourea rupestris** (Müll.Arg.) Delprete].

*Psychotria stipulosa* Muell. Arg. [= **Palicourea stipulosa** (Müll.Arg.) Borhidi].

*Psychotria subtriflora* Muell. Arg. [= **Palicourea subtriflora** (Muell.Arg.) Delprete].

*Psychotria vellosiana* Benth. [= **Palicourea sessilis** (Vell.) C.M.Taylor].

*Psychotria villosa* Vell., nom. illeg. [= **Palicourea tenerior** (Cham.) Delprete & J.H.Kirkbr.].

*Psychotria viridis* Ruiz & Pav.

*Psychotria xanthocephala* Muell.Arg. [= **Palicourea prunifolia** (Kunth) Borhidi].

*Retiniphyllum kuhlmannii* Standl.

*Richardia brasiliensis* Gomes

*Richardsonia grandiflora* Cham. & Schltdl. [= **Richardia grandiflora** (Cham. & Schltdl.) Steud.].

*Rubia noxia* A. St. Hil. [= **Galium noxium** (A.St.Hil.) Dempster].

*Rudgea burchelliana* Muell.Arg.

*Rudgea erioloba* Benth.

*Rudgea goyazensis* Muell.Arg.

*Rudgea jacobinensis* Muell.Arg. [= **Rudgea erioloba** Benth.].

*Rudgea longiflora* Benth.

*Rudgea myrsinifolia* Benth.

*Rudgea tomentosa* Rusby

*Sabicea brasiliensis* Wernham

*Sabicea grisea* Cham. & Schltdl.

*Sabicea humilis* var. *lanceolata* Wernham [= **Sabicea humilis**].

*Sabicea moorei* Wernham [= **Sabicea humilis** S.Moore].

*Spermacoce peruviana* Pers. [= **Galianthe peruviana** (Pers.) E.L.Cabral].

*Spermacoce polygonifolia* A.St.Hil. [= **Mitracarpus polygynolius** (A.St.Hil.) R.M.Salas & E.B.Souza].

*Sprucea rubescens* Benth. [= **Simira rubescens** (Benth.) Bremek. ex Steyerm.].

*Staelia aurea* K.Schum.

*Staelia capitata* K.Schum. [= **Planaltina capitata** (K.Schum.) R.M.Salas & E.L.Cabral].

**Neotypifications (11):**

*Borreria scabiosoides* Cham. & Schltdl. [= **Spermacoce scabiosoides** (Cham. & Schltdl.) Kuntze].

*Borreria tenuis* DC. [= **Spermacoce neotenuis** Govaerts].

*Gardenia formosa* Cham. & Schltdl. [= **Tocoyena formosa** (Cham. & Schltdl.) K.Schum.].

*Gonzalagunia dicocca* Cham. & Schltdl.

*Hamelia patens* Jacq.

*Patabea coriacea* Cham. [= **Palicourea coriacea** (Cham.) K.Schum.].

*Psychotria anceps* Kunth

*Psychotria leiocarpa* Cham. & Schltdl. [= **Palicourea leiocarpa** (Cham. & Schltdl.) Delprete].

*Psychotria racemosa* Rich. [= **Palicourea racemosa** (Aubl.) G.Nicholson].

*Psychotria stachyoides* Benth. [= **Palicourea stachyoides** (Benth.) Delprete].

*Schwenkfeldia aurea* Spreng. [= **Coccocypselum aureum** (Spreng.) Cham. & Schltdl.].
*Psychotria macrobotrys* Ruiz & Pav. [= **Palicourea macrobotrys** (Ruiz & Pav.) DC.].

*Psychotria microcephala* (Willd. ex Roem. & Schult.)
Muell.Arg. var. *tripotamica* Muell. Arg. [= **Palicourea prunifolia** (Kunth) Borhidi].

*Psychotria racemosa* Rich. [= **Palicourea racemosa** (Aubl.) G.Nicholson].

*Psychotria rupestris* Muell.Arg. [= **Palicourea rupestris** (Müll.Arg.) Delprete].

*Psychotria schuechiana* Müll.Arg. [= **Palicourea subtriflora** (Muell.Arg.) Delprete].

*Psychotria stachyoides* Benth. [= **Palicourea rupestris** (Müll.Arg.) Delprete].

*Psychotria stipulosa* Muell. Arg. [= **Palicourea stipulosa** (Müll.Arg.) Borhidi].

*Psychotria subtriflora* Muell. Arg. [= **Palicourea subtriflora** (Muell.Arg.) Delprete].

*Psychotria vellosiana* Benth. [= **Palicourea sessilis** (Vell.) C.M.Taylor].

*Psychotria villosa* Vell., nom. illeg. [= **Palicourea tenerior** (Cham.) Delprete & J.H.Kirkbr.].

*Psychotria viridis* Ruiz & Pav.

*Psychotria xanthocephala* Muell.Arg. [= **Palicourea prunifolia** (Kunth) Borhidi].

*Retiniphyllum kuhlmannii* Standl.

*Richardia brasiliensis* Gomes

*Richardsonia grandiflora* Cham. & Schltdl. [= **Richardia grandiflora** (Cham. & Schltdl.) Steud.].

*Rubia noxia* A. St. Hil. [= **Galium noxium** (A.St.Hil.) Dempster].

*Rudgea burchelliana* Muell.Arg.

*Rudgea erioloba* Benth.

*Rudgea goyazensis* Muell.Arg.

*Rudgea jacobinensis* Muell.Arg. [= **Rudgea erioloba** Benth.].

*Rudgea longiflora* Benth.

*Rudgea myrsinifolia* Benth.

*Rudgea tomentosa* Rusby

*Sabicea brasiliensis* Wernham

*Sabicea grisea* Cham. & Schltdl.

*Sabicea humilis* var. *lanceolata* Wernham [= **Sabicea humilis**].

*Sabicea moorei* Wernham [= **Sabicea humilis** S.Moore].

*Spermacoce peruviana* Pers. [= **Galianthe peruviana** (Pers.) E.L.Cabral].

*Spermacoce polygonifolia* A.St.Hil. [= **Mitracarpus polygynolius** (A.St.Hil.) R.M.Salas & E.B.Souza].

*Sprucea rubescens* Benth. [= **Simira rubescens** (Benth.) Bremek. ex Steyerm.].

*Staelia aurea* K.Schum.

*Staelia capitata* K.Schum. [= **Planaltina capitata** (K.Schum.) R.M.Salas & E.L.Cabral].

### Neotypifications (11):

*Borreria scabiosoides* Cham. & Schltdl. [= **Spermacoce scabiosoides** (Cham. & Schltdl.) Kuntze].

*Borreria tenuis* DC. [= **Spermacoce neotenuis** Govaerts].

*Gardenia formosa* Cham. & Schltdl. [= **Tocoyena formosa** (Cham. & Schltdl.) K.Schum.].

*Gonzalagunia dicocca* Cham. & Schltdl.

*Hamelia patens* Jacq.

*Patabea coriacea* Cham. [= **Palicourea coriacea** (Cham.) K.Schum.].

*Psychotria anceps* Kunth

*Psychotria leiocarpa* Cham. & Schltdl. [= **Palicourea leiocarpa** (Cham. & Schltdl.) Delprete].

*Psychotria racemosa* Rich. [= **Palicourea racemosa** (Aubl.) G.Nicholson].

*Psychotria stachyoides* Benth. [= **Palicourea stachyoides** (Benth.) Delprete].

*Schwenkfeldia aurea* Spreng. [= **Coccocypselum aureum** (Spreng.) Cham. & Schltdl.].