This species belongs to the large *rhenanus* species group as setae S4, JV3 and JV4 are present, the setae on dorsal shield are setiform, approximately equal in length except Z4/Z5, setae r3 and R1 are inserted on lateral integument and setae on dorsal shield in the z-Z and s-S series shorter than distances between their bases.

In the literature, Pritchard described this species and Baker (1962) from Congo and Rwanda, recorded than from Nigeria (Matthysse and Denmark 1981), from Indonesia (Oomen 1982), from Kenya (El-Banhawy *et al.* 2009) and our specimens are very similar to the original description based on drawings. This species has very original characters.

Surprisingly, no measurements were given in all papers mentioning this species, even the original description. Measurements are given in the table 38 for the first time for the three adult female specimens and for the single adult male specimen.

This species if mentioned for the first time from La Réunion, from primary forest of medium altitude. Its biology is totally unknown.

**Specimens examined:** 3 ♀♀ + 1 ♂ in total, all measured. Forêt de Bélouve - Gite (aasl 1500m, Long 55°33'36" E, Lat 21°6'0" S), 3 ♀♀ on *C. borbonica*, 28/1/2017; Forêt de Bélouve - Trou de fer (aasl 1300 m, Long 55°33'36" E, Lat 21°2'24" S), 1 ♂ on *A. heterophylla*, 28/1/2017.

**Remarks:** this species has short subequal setae except Z5 in females which are around the double length of all other dorsal setae. The ventrianal shield of the female quadrangular, almost square, and as large as wide. Setae Z5, JV5 and the four macrosetae of the leg IV are knobbed (Table 38).

## Typhlodromus (Anthoseius) transvaalensis (Nesbitt)

*Kampimodromus transvaalensis* Nesbitt 1951: 55.

*Neoseiulus transvaalensis*, Muma 1961: 295.

<table><caption>**Table 37** Character measurements of adult females of *Kuzinellus scytinus* collected in this study with those in previous studies (localities followed by the number of specimens measured).</caption><thead><tr><th colspan="2">La Réunion</th><th>Africa</th><th colspan="2">Holotype</th></tr><tr><th>Characters</th><th>2, this study</th><th>3</th><th>Madagascar 1</th><th>Madagascar 2</th></tr></thead><tbody><tr><td>Dsl</td><td><strong>323</strong> (310 – 335)</td><td>329 (317 – 341)</td><td>1</td><td>325</td></tr><tr><td>Dsw</td><td><strong>170</strong> (155 – 185)</td><td>147 (157 – 184)</td><td>163</td><td>180</td></tr><tr><td>j1</td><td><strong>18</strong></td><td>18</td><td>15</td><td>17</td></tr><tr><td>j3</td><td><strong>18</strong> (15 – 20)</td><td>17 (16 – 18)</td><td>17</td><td>18</td></tr><tr><td>j4</td><td><strong>16</strong> (15 – 18)</td><td>14 (13 – 16)</td><td>15</td><td>14</td></tr><tr><td>j5</td><td><strong>16</strong> (15 – 18)</td><td>13 (11 – 14)</td><td>14</td><td>13</td></tr><tr><td>j6</td><td><strong>18</strong></td><td>15 (14 – 16)</td><td>17</td><td>17</td></tr><tr><td>J2</td><td><strong>23</strong></td><td>19</td><td>22</td><td>20</td></tr><tr><td>J5</td><td><strong>13</strong></td><td>11 (10 – 11)</td><td>9</td><td>10</td></tr><tr><td>r3</td><td><strong>21</strong> (20 – 23)</td><td>20 (19 – 21)</td><td>23</td><td>20</td></tr><tr><td>R1</td><td><strong>16</strong> (15 – 18)</td><td>15 (14 – 16)</td><td>17</td><td>16</td></tr><tr><td>s4</td><td><strong>25</strong></td><td>21 (19 – 22)</td><td>22</td><td>21</td></tr><tr><td>s6</td><td><strong>25</strong></td><td>23 (22 – 24)</td><td>24</td><td>25</td></tr><tr><td>S2</td><td><strong>28</strong></td><td>26 (24 – 27)</td><td>28</td><td>25</td></tr><tr><td>S4</td><td><strong>30</strong></td><td>26 (26 – 27)</td><td>31</td><td>28</td></tr><tr><td>S5</td><td><strong>20</strong></td><td>18 (16 – 19)</td><td>21</td><td>21</td></tr><tr><td>z2</td><td><strong>16</strong> (15 – 18)</td><td>15 (13 – 16)</td><td>17</td><td>17</td></tr><tr><td>z3</td><td><strong>19</strong> (18 – 20)</td><td>18 (14 – 21)</td><td>20</td><td>20</td></tr><tr><td>z4</td><td><strong>21</strong> (20 – 23)</td><td>18 (16 – 21)</td><td>19</td><td>20</td></tr><tr><td>z5</td><td><strong>16</strong> (15 – 18)</td><td>14 (11 – 16)</td><td>16</td><td>15</td></tr><tr><td>z6</td><td><strong>18</strong></td><td>15 (14 – 16)</td><td>17</td><td>17</td></tr><tr><td>Z4</td><td><strong>35</strong></td><td>31 (30 – 32)</td><td>34</td><td>33</td></tr><tr><td>Z5</td><td><strong>56</strong> (55 – 58)</td><td>51 (50 – 53)</td><td>53</td><td>51</td></tr></tbody></table><table><thead><tr><th colspan="2">La Réunion</th><th>Africa</th><th colspan="2">Holotype</th></tr><tr><th>Characters</th><th>2, this study</th><th>3</th><th>Madagascar 1</th><th>Madagascar 2</th></tr></thead><tbody><tr><td>st1-st1</td><td><strong>48</strong></td><td>-</td><td>1</td><td>47</td></tr><tr><td>st2-st2</td><td><strong>54</strong> (53 – 55)</td><td>54 (53 – 56)</td><td>1</td><td>-</td></tr><tr><td>st3-st3</td><td><strong>60</strong></td><td>-</td><td>1</td><td>-</td></tr><tr><td>st1-st3</td><td><strong>63</strong> (60 – 65)</td><td>62 (59 – 64)</td><td>1</td><td>-</td></tr><tr><td>st4-st4</td><td><strong>95</strong></td><td>-</td><td>1</td><td>-</td></tr><tr><td>st5-st5</td><td><strong>63</strong></td><td>51 (50 – 53)</td><td>1</td><td>-</td></tr><tr><td>Lisl</td><td><strong>20</strong></td><td>-</td><td>1</td><td>-</td></tr><tr><td>Lisw</td><td><strong>4</strong> (3 – 5)</td><td>-</td><td>1</td><td>-</td></tr><tr><td>Sisl</td><td><strong>15</strong> (13 – 18)</td><td>-</td><td>1</td><td>-</td></tr><tr><td>Vsl</td><td><strong>113</strong></td><td>107 (106 – 109)</td><td>1</td><td>-</td></tr><tr><td>Vsw ZV2</td><td><strong>88 – 90</strong></td><td>89 (86 – 93)</td><td>1</td><td>-</td></tr><tr><td>Vsw anus</td><td><strong>80</strong></td><td>73 (72 – 74)</td><td>1</td><td>-</td></tr><tr><td>JV5</td><td><strong>38</strong> (35 – 40)</td><td>-</td><td>1</td><td>-</td></tr><tr><td>StIV</td><td><strong>33</strong> (30 – 35)</td><td>31 (27 – 35)</td><td>1</td><td>-</td></tr><tr><td>Scl</td><td><strong>32</strong> (33 – 35)</td><td>41 (35 – 46)</td><td>1</td><td>-</td></tr><tr><td>Sew</td><td><strong>7</strong> (5 – 8)</td><td>-</td><td>1</td><td>-</td></tr><tr><td>Fdl</td><td><strong>28</strong></td><td>25</td><td>1</td><td>-</td></tr><tr><td>No teeth Fd</td><td><strong>6</strong></td><td>6</td><td>1</td><td>-</td></tr><tr><td>Mdl</td><td><strong>26</strong> (25 – 28)</td><td>27</td><td>1</td><td>-</td></tr><tr><td>No teeth Md</td><td><strong>3</strong></td><td>3</td><td>1</td><td>-</td></tr></tbody></table>

Sources of measurements – Africa (Burundi 1♀, Rwanda 1♀): Moraes *et al.* (2008); Rwanda: Moraes *et al.* (2008); Holotype Madagascar 1: Chazeau (1970); Holotype Madagascar 2: Chant & Yoshida-Shaul (1986); -: not provided.
*Clavidromus transvaalensis*, Muma & Denmark 1968: 238; Muma & Denmark 1970: 128; Moraes *et al.* 1986: 182.

*Typhlodromus transvaalensis*, Chant & Baker 1965: 5; Schicha 1981a: 36; Moraes *et al.* 2004a: 355; Chant & McMurtry 1994: 252, 2007: 157.

*Typhlodromus jackmickleyi*, De Leon 1958: 75; van Der Merwe 1968: 23 (synonymy according to Muma & Denmark 1968).

*Typhlodromus pectinatus*, Athias-Henriot 1958a: 179 (synonymy according to Muma & Denmark 1968).

This species has elongate serrated dorsal setae, setae Z1 and JV3 absent, an elongate calyx of the spermatheca, leg IV with 3 macrosetae and few teeth on chelicerae. It belongs to the *transvaalensis* species group of the subgenus *Anthoseius* of the genus *Typhlodromus* (Chant and McMurtry 1994).

According to McMurtry *et al.* (2013), *T. transvaalensis* is a type III phytoseiid and a generalist predator that feeds on mites, insects and pollen. It completed its life cycle when fed on the eriophyid mites *Eriophyes dioscoridis* Soliman and Abou-Awad and *Eriophyes olive* Zaher and Abou-Awad, eggs of the scale insect *Parlatoria zizyphus* (Lucas) and pollen of *R. communis* in experimental conditions. The percentage of individuals attaining maturity was less than 20% when nymphs of the tetranychid mite, *T. urticae* Koch, were provided. The development was faster and reproduction was higher when *T. transvaalensis* fed on eriophyid mites. *T. urticae* was an unsuitable feeding and reproduction substrate. The daily reproduction was as low as 0.4 and 0.8 egg/ female/ day when females were maintained on pollen grains of *R. communis* and eggs of *P. zizyphus*. The adult female daily consumed 126, 97 and 6 individuals of *E. olivi*, *E. dioscoridis* and *T. urticae*, respectively (Momen and Hussein 1999). Adult female *T. transvaalensis* were more efficient at predating all stages of *P. latus* (Banks) than *Tetranychus bastosi* Tuttle, Baker and Sales. The *T. transvaalensis* life cycle was shorter with diets including *R. communis* pollen, but *Zea mays* pollen was also suitable for reproduction. The results indicate that *T. transvaalensis* is a generalist predator with high potential for controlling *P. latus* in *Jatropha curcas* plantations and that the presence of *R. communis* and *Z. mays* crops boosts its development and reproduction (Canarte *et al.* 2017). This species is widely distributed all over the world (Demite *et al.* 2019). It was already recorded from *La Réunion* in

<table><caption>**Table 38** Character measurements of adult females and one adult male of *Typhlodromus (Anthoseius) ndibu* collected in this study (localities followed by the number of specimens measured between brackets).</caption><thead><tr><th rowspan="2">Characters</th><th colspan="2">♀</th><th colspan="2">♂</th><th rowspan="2">Characters</th><th colspan="2">♀</th><th colspan="2">♂</th></tr><tr><th>Réunion<br>3, this study</th><th>La Réunion<br>1, this study</th><th>Réunion<br>3, this study</th><th>La Réunion<br>1, this study</th><th>Réunion<br>3, this study</th><th>La Réunion<br>1, this study</th><th>Réunion<br>3, this study</th><th>La Réunion<br>1, this study</th></tr></thead><tbody><tr><td>Dsl</td><td>312 (303 – 328)</td><td>263</td><td>z2</td><td>13</td><td>Vsl</td><td>111 (110 – 113)</td><td>100</td><td></td><td></td></tr><tr><td>Dsw</td><td>203 (200 – 205)</td><td>175</td><td>z3</td><td>13</td><td>Vsw ZV2</td><td>113 (110 – 115)</td><td>135</td><td></td><td></td></tr><tr><td>j1</td><td>15</td><td>15</td><td>z4</td><td>13</td><td>Vsw anus</td><td>96 (90 – 100)</td><td>88</td><td></td><td></td></tr><tr><td>j3</td><td>13 (10 – 15)</td><td>16</td><td>z5</td><td>10</td><td>Distance between sol.</td><td>20</td><td>15</td><td></td><td></td></tr><tr><td>j4</td><td>9 (8 – 10)</td><td>10</td><td>Z4</td><td>14 (13 – 15)</td><td>JV5</td><td>19 (15 – 23) knob.</td><td>15 knob.</td><td></td><td></td></tr><tr><td>j5</td><td>10</td><td>12</td><td>Z5</td><td>31 (30 – 33) knob.</td><td>SgeIV</td><td>13 (12 – 13) knob.</td><td>10 knob.</td><td></td><td></td></tr><tr><td>j6</td><td>11 (10 – 13)</td><td>13</td><td>st1-st1</td><td>41 (40 – 43)</td><td>StiIV</td><td>12 (12 – 13) knob.</td><td>13 knob.</td><td></td><td></td></tr><tr><td>J2</td><td>12 (10 – 13)</td><td>15</td><td>st2-st2</td><td>49 (48 – 50)</td><td>StIV</td><td>23 (23 – 24) knob.</td><td>16 knob.</td><td></td><td></td></tr><tr><td>J5</td><td>10</td><td>8</td><td>st3-st3</td><td>55</td><td>SttIV</td><td>21 (20 – 23) knob.</td><td>15 knob.</td><td></td><td></td></tr><tr><td>r3</td><td>15</td><td>15</td><td>st1-st3</td><td>61 (53 – 65)</td><td>Scl</td><td>8</td><td></td><td></td><td></td></tr><tr><td>R1</td><td>15</td><td>14</td><td>st4-st4</td><td>56 (55 – 58)</td><td>Scw</td><td>5</td><td></td><td></td><td></td></tr><tr><td>s4</td><td>14 (13 – 15)</td><td>18</td><td>Gensl</td><td>107 (103 – 110)</td><td>Fdl</td><td>23 (20 – 25)</td><td>18</td><td></td><td></td></tr><tr><td>s6</td><td>15</td><td>18</td><td>Gensw st5</td><td>64 (63 – 65)</td><td>No teeth Fd</td><td>8</td><td></td><td></td><td></td></tr><tr><td>S2</td><td>15</td><td>18</td><td>Gensw post. corn.</td><td>81 (80 – 83)</td><td>Mdl</td><td>24 (23 – 25)</td><td>23</td><td></td><td></td></tr><tr><td>S4</td><td>17 (15 – 18)</td><td>15</td><td>st5-st5</td><td>58</td><td>No teeth Md</td><td>3</td><td></td><td></td><td></td></tr><tr><td>S5</td><td>14 (13 – 15)</td><td>13</td><td>Lis1</td><td>20</td><td>Shaft</td><td>Not applicable</td><td>13</td><td></td><td></td></tr><tr><td></td><td></td><td></td><td>Lisw</td><td>7 (5 – 8)</td><td></td><td></td><td></td><td></td><td></td></tr></tbody></table>

Sources of measurements – No measurements are available in the whole literature; - : not provided.
previous surveys (Quilici *et al.* 2000). If all details of collections were provided in the previous paper, measurements of specimens collected and identified were not given. Measurements of specimens collected during this study are provided in table 39.

**Specimens examined:** 70 ♀♀ + 4 im. in total, 20 ♀♀ measured. Saint-Paul – Savannah (aasl 61 m, Long 55°29'43" E, Lat 21°20'41" S), 5 ♀♀ in *Frankliniothrips* sp. rearings, 16/6/2015; Petite Île – Les bas, Doris Morel farm (aasl 230 m, long 55°34'1" E, lat 21°21'22" S), 1 ♀ on *P. lanceolata*, 31/5/2016; Saint-Pierre – Bassin-Plat CIRAD Research Station (aasl 153 m, Long 55°29'18" E, Lat 21°19'25" S), 1 ♀ in plot CC, 16/8/2016; 1 ♀ in plot BM, 23/8/2016; Petite Île – Piton Bloc, Yébo Luguy farm (aasl 973 m, Long 55°34'64" E, Lat 21°18'64" S), 1 ♀ on *B. catharticus*, 18/10/2016; Saint-Pierre – Ligne Paradis, La Coccinelle Inc. (aasl 164 m, Long 55°28'59" E, Lat 21°18'55" S), 22 ♀♀ + 3 im. in rearings of beneficial insects, 22/2/2017; 4 ♀♀ in rearings of beneficial insects, 11/6/2017; 32 ♀♀ in rearings of beneficial insects, 1/8/2017; 3 ♀♀ + 1 im. in rearings of beneficial insects, 11/8/2017.

**Remarks:** all measurement values (Table 39) agree well with those already published on this species with only very slight variations. Measurement values of female specimens of La Réunion are very similar with values for specimens of Kenya and South Africa. Despite the large number of females collected, sometimes from a single location, we did not find any males in our study, suggesting that this predatory mite can reproduce through thelytokous parthenogenesis, as observed by Kishimoto (2015).

## Conclusion

Before this 10th paper, the fauna of Phytoseiidae of La Réunion Island was limited to 33 recorded species including 24 Amblyseiinae, 5 Phytoseiinae and 4 Typhlodrominae, among which 8 species that had been described as new.

This paper reports on results of surveys done recently (2015-2018) and add 19 newly recorded species among which 3 are new to Science and 21 already known species but with additional data. The number of species for La Réunion Island now reached to 52 among which 11 have been described only from this Island (*P. haroldi* was found in Mauritius recently).

Among these 19 species newly recorded, at least nine are already well known as biological control agents (BCA). This must be underlined as new regulations on importation of macro-organisms are proposed in a lot of countries and specifically for countries like France that have very far over-sea territories. Therefore, it is impossible to import and of course to sell and use exotic species if they are not indigenous in La Réunion Island that is considered as a territory. An importation permit must be requested, but it is expensive and chances to obtain are generally very low (Kreiter *et al.* 2016). The knowledge of the biodiversity, especially of efficient biological control agents from overseas territories, not only for fundamental biodiversity knowledge or conversation purposes but also for agricultural and economical ones, is so of a considerable importance.

## Acknowledgements

Acknowledgements are due to the institution that have granted travel and accommodations to the senior author in La Réunion during more than 3 months: Montpellier SupAgro, CIRAD (UMR PVBMT and UR Hortsys), Armeflhor and La Coccinelle.

Part of the present study was realised within the framework of the project Agrum'Aide (2014-2018) managed by CIRAD (the leader was Fabrice Le Bellec). This study was financially supported by the French Agency of Biodiversity (formerly ONEMA), in the framework of the tender 'Biodiversité-Ecophyto'.

Many thanks are also due to FDGDON La Réunion (23 Rue Jules Thirel – Cour de l'Usine de Savannah, 97460 Saint Paul), ANSES (Laboratoire de la Santé des Végétaux, Unité
<table><caption>**Table 39** Character measurements of adult females of *Typhlodromus (Anthoseius) transvaalensis* collected in this study with those in previous studies (localities followed by the number of specimens measured).</caption><thead><tr><th>Characters</th><th>La Réunion<br>20, this study</th><th>Africa<br>9</th><th>Egypt<br>5</th><th>Japan<br>10</th><th>New Caledonia<br>1</th><th>South Africa<br>52</th><th>Taiwan<br>2</th><th>Holotype<br>South Africa, 1</th></tr></thead><tbody><tr><td>Dsl</td><td><b>362</b> (343 – 395)</td><td>352 (325 – 373)</td><td>365 (335 – 407)</td><td>389 (383 – 395)</td><td>356</td><td>356 – 367</td><td>331</td><td>-</td></tr><tr><td>Dsw</td><td><b>211</b> (200 – 223)</td><td>210 (188 – 232)</td><td>192 (170 – 219)</td><td>253 (251 – 255)</td><td>179</td><td>215 – 230</td><td>173</td><td>-</td></tr><tr><td>j1</td><td><b>28</b> (25 – 33)</td><td>28 (25 – 30)</td><td>30 (27 – 33)</td><td>33 (33 – 34)</td><td>29</td><td>29 – 33</td><td>26</td><td>23</td></tr><tr><td>j3</td><td><b>39</b> (35 – 43)</td><td>38 (36 – 40)</td><td>41 (38 – 48)</td><td>43 (42 – 43)</td><td>39</td><td>37 – 42</td><td>35</td><td>36</td></tr><tr><td>j4</td><td><b>31</b> (28 – 36)</td><td>30 (26 – 33)</td><td>32 (30 – 37)</td><td>31 (31 – 32)</td><td>28</td><td>29 – 33</td><td>28</td><td>27</td></tr><tr><td>j5</td><td><b>32</b> (30 – 35)</td><td>30 (26 – 33)</td><td>31 (30 – 36)</td><td>32 (32 – 33)</td><td>30</td><td>29 – 33</td><td>26</td><td>28</td></tr><tr><td>j6</td><td><b>39</b> (35 – 45)</td><td>38 (35 – 41)</td><td>38 (36 – 42)</td><td>39 (38 – 39)</td><td>34</td><td>37 – 42</td><td>34</td><td>36</td></tr><tr><td>J2</td><td><b>46</b> (40 – 50)</td><td>44 (40 – 46)</td><td>44 (41 – 51)</td><td>46 (45 – 46)</td><td>43</td><td>45 – 48</td><td>34</td><td>45</td></tr><tr><td>J5</td><td><b>10</b> (8 – 13)</td><td>9 (9 – 10)</td><td>11 (7 – 14)</td><td>11</td><td>10</td><td>11 – 13</td><td>10</td><td>10</td></tr><tr><td>r3</td><td><b>35</b> (33 – 38)</td><td>32 (29 – 35)</td><td>34 (31 – 37)</td><td>38 (37 – 38)</td><td>36</td><td>34</td><td>26</td><td>34</td></tr><tr><td>R1</td><td><b>43</b> (40 – 48)</td><td>40 (38 – 41)</td><td>41 (39 – 44)</td><td>43 (42 – 43)</td><td>40</td><td>39 – 43</td><td>36</td><td>45</td></tr><tr><td>s4</td><td><b>46</b> (40 – 50)</td><td>45 (42 – 49)</td><td>49 (43 – 56)</td><td>50 (49 – 50)</td><td>45</td><td>45 – 48</td><td>41</td><td>45</td></tr><tr><td>s6</td><td><b>50</b> (45 – 53)</td><td>49 (46 – 54)</td><td>50 (47 – 54)</td><td>50 (49 – 51)</td><td>43</td><td>47 – 52</td><td>46</td><td>45</td></tr><tr><td>S2</td><td><b>56</b> (53 – 60)</td><td>55 (50 – 57)</td><td>50 (49 – 64)</td><td>56</td><td>55</td><td>52 – 56</td><td>50</td><td>71</td></tr><tr><td>S4</td><td><b>58</b> (55 – 63)</td><td>56 (53 – 56)</td><td>54 (50 – 60)</td><td>57 (57 – 58)</td><td>51</td><td>56 – 59</td><td>50</td><td>58</td></tr><tr><td>S5</td><td><b>7</b> (5 – 10)</td><td>9 (8 – 11)</td><td>11 (10 – 11)</td><td>8 (8 – 9)</td><td>9</td><td>11 – 13</td><td>6</td><td>5</td></tr><tr><td>z2</td><td><b>25</b> (23 – 28)</td><td>24 (21 – 28)</td><td>26 (24 – 33)</td><td>26 (25 – 26)</td><td>21</td><td>23 – 26</td><td>22</td><td>21</td></tr><tr><td>z3</td><td><b>40</b> (35 – 43)</td><td>38 (36 – 40)</td><td>41 (37 – 48)</td><td>40</td><td>36</td><td>35 – 40</td><td>36</td><td>36</td></tr><tr><td>z4</td><td><b>41</b> (38 – 45)</td><td>41 (39 – 41)</td><td>45 (40 – 55)</td><td>45 (45 – 46)</td><td>42</td><td>37 – 42</td><td>41</td><td>42</td></tr><tr><td>z5</td><td><b>28</b> (25 – 30)</td><td>22 (21 – 28)</td><td>28 (25 – 37)</td><td>29 (28 – 29)</td><td>24</td><td>25 – 29</td><td>24</td><td>27</td></tr><tr><td>Z4</td><td><b>58</b> (53 – 60)</td><td>52 (47 – 60)</td><td>56 (53 – 63)</td><td>56 (55 – 56)</td><td>51</td><td>52 – 57</td><td>53</td><td>56</td></tr><tr><td>Z5</td><td><b>65</b> (58 – 70)</td><td>62 (57 – 70)</td><td>64 (60 – 71)</td><td>67 (66 – 68)</td><td>64</td><td>62 – 68</td><td>56</td><td>63</td></tr><tr><td>st1-st1</td><td><b>61</b> (58 – 65)</td><td>-</td><td>62 (61 – 65)</td><td>-</td><td>-</td><td>-</td><td>-</td><td>-</td></tr><tr><td>st2-st2</td><td><b>66</b> (64 – 69)</td><td>60 (54 – 60)</td><td>65 (62 – 73)</td><td>-</td><td>-</td><td>63 – 68</td><td>-</td><td>-</td></tr><tr><td>st3-st3</td><td><b>75</b> (70 – 80)</td><td>-</td><td>75 (70 – 83)</td><td>-</td><td>75</td><td>-</td><td>-</td><td>-</td></tr><tr><td>st1-st3</td><td><b>71</b> (68 – 75)</td><td>68 (65 – 70)</td><td>-</td><td>-</td><td>69</td><td>92 – 100</td><td>-</td><td>-</td></tr><tr><td>st4-st4</td><td><b>95</b> (78 – 115)</td><td>-</td><td>85 (78 – 88)</td><td>-</td><td>-</td><td>-</td><td>-</td><td>-</td></tr><tr><td>st5-st5</td><td><b>78</b> (73 – 85)</td><td>74 (67 – 80)</td><td>82 (78 – 90)</td><td>-</td><td>-</td><td>78 – 84</td><td>-</td><td>-</td></tr><tr><td>Lisl</td><td><b>30</b> (23 – 35)</td><td>-</td><td>-</td><td>-</td><td>29</td><td>-</td><td>24</td><td>-</td></tr><tr><td>Lisw</td><td><b>3</b> (3 – 5)</td><td>-</td><td>-</td><td>-</td><td>-</td><td>-</td><td>3</td><td>-</td></tr><tr><td>Sisl</td><td><b>12</b> (8 – 18)</td><td>-</td><td>-</td><td>-</td><td>12</td><td>-</td><td>9</td><td>-</td></tr><tr><td>Vsl</td><td><b>124</b> (115 – 133)</td><td>118 (107 – 130)</td><td>112 (82 – 131)</td><td>136 (133 – 138)</td><td>111</td><td>112 – 120</td><td>106</td><td>111</td></tr><tr><td>Vsw ZV2</td><td><b>80</b> (73 – 88)</td><td>76 (73 – 78)</td><td>79 (73 – 85)</td><td>83 (83 – 84)</td><td>84</td><td>74 – 81</td><td>65</td><td>79</td></tr><tr><td>Vsw anus</td><td><b>82</b> (75 – 88)</td><td>78 (73 – 84)</td><td>87 (71 – 130)</td><td>-</td><td>-</td><td>-</td><td>-</td><td>-</td></tr><tr><td>JV5</td><td><b>58</b> (50 – 63)</td><td>-</td><td>63 (60 – 65)</td><td>60 (60 – 61)</td><td>57</td><td>-</td><td>-</td><td>60</td></tr><tr><td>Scl</td><td><b>19</b> (8 – 28)</td><td>19 (13 – 30)</td><td>10 (9 – 10)</td><td>-</td><td>27</td><td>-</td><td>-</td><td>-</td></tr><tr><td>Scw</td><td><b>8</b> (5 – 10)</td><td>-</td><td>-</td><td>-</td><td>3</td><td>-</td><td>-</td><td>-</td></tr><tr><td>SgeIV</td><td><b>26</b> (23 – 28)</td><td>26 (25 – 26)</td><td>26 (24 – 28)</td><td>27</td><td>25</td><td>24 – 27</td><td>22</td><td>20</td></tr><tr><td>StiIV</td><td><b>28</b> (25 – 30)</td><td>27 (24 – 29)</td><td>28 (27 – 34)</td><td>28</td><td>26</td><td>27 – 30</td><td>26</td><td>27</td></tr><tr><td>StIV</td><td><b>47</b> (43 – 50)</td><td>46 (40 – 50)</td><td>48 (45 – 51)</td><td>48 (48 – 49)</td><td>44</td><td>45 – 48</td><td>58</td><td>45</td></tr><tr><td>Fdl</td><td><b>29</b> (25 – 30)</td><td>28 (26 – 30)</td><td>31 (30 – 32)</td><td>-</td><td>28</td><td>30</td><td>-</td><td>-</td></tr><tr><td>No teeth Fd</td><td><b>2</b></td><td>2</td><td>2</td><td>3</td><td>2</td><td>-</td><td>-</td><td>-</td></tr><tr><td>Mdl</td><td><b>29</b> (25 – 33)</td><td>30 (30 – 31)</td><td>31 (30 – 31)</td><td>1</td><td>29</td><td>30</td><td>-</td><td>-</td></tr><tr><td>No teeth Md</td><td><b>1</b></td><td>1</td><td>1</td><td>1</td><td>1</td><td>-</td><td>-</td><td>-</td></tr></tbody></table>

Sources of measurements – Africa (Kenya 2♀♀, Cape Verde 1♀, South Africa 6♀♀): Ueckermann *et al.* (2008); Egypt: Abo-Shnaf & Moraes (2014); Japan: Ehara & Kishimoto (2007); South Africa (identified as *T. jackmickleyi* but synonymized by Denmark & Muma 1968): van der Merwe (1968); Taiwan: Tseng (1983); Holotype South Africa and New Caledonia: Schicha (1981a); -: not provided.
Ravageurs et Agents Pathogènes tropicaux, Site de La Réunion, 7 Chemin de l'Irat, 97410 Saint-Pierre), and UMR PVBMT (same address) which have provided with several samples.

We are grateful to the growers for permission to collect in their farms: Pierre and Ingrid Avril, EARL Montvert in Le Montvert-les-Hauts, Jean-Louis Robert, in Le 19e, Plaine des Caffres, Janick Bénard in Le Tampon, Grand Tampon, Yebo Lugy in Petite-Île, Piton-Bloc, Doris Morel in Petite-Île, Les Bas, Delaunay Jean-Max in Vincendo and Aldo Grace in Le Tampon, Bras de Pontho.

Huge thanks to Maëva Labouyrie, Inès Seka, András Kovács, and Hélder Silva, students of Master of Montpellier SupAgro, who have worked enthusiastically for a master project with the senior author on the material of La Réunion Island from March to July 2018.

We are very grateful to Professor Gilberto J. de Moraes (University of Sao Paulo, Escuala Superior de Agricultura Luiz de Queiroz, Departamento Entomologia and Acarologia, Piracicaba, Brasil) for organising a loan of the holotype, paratypes and some additional materials of *Neoseiulus recifensis* Gondim Jr and Moraes and Drs Danuta Knihinicki and Lauren Drysdale (NSW Department of Primary Industries, Orange Agricultural Institute, Biosecurity Collections, Orange NSW, Australia) for organising a loan of the holotype of *Neoseiulus houstoni* (Schicha).

I would like also to thank my colleague Professor Marie-Stéphane Tixier who have examined the synonymy of the three species of *Neoseiulus* with a polytomous key we are developing in UMR CBGP under her supervision and has reached the same conclusion than the senior author on the synonymy of the three species impossible to distinguish on the basis of morphological characters.

We are finally very grateful to Professor Francisco Ferragut who has given his opinion on specimens of *Amblyseius neoankaratrae* from la Réunion Island after the publication of his paper on Phytoseiidae of Mauritius (2019) in which this species is mentioned and thanks to whom this paper was greatly improved.

I finally would like to thank heartfully the two reviewers for very valuable constructive comments and great improvements of this paper.

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