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# Challenging the Wallacean shortfall: A total assessment of insect diversity on Guadeloupe (French West Indies), a checklist and bibliography

François Meurgey, Thibault Ramage

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François Meurgey, Thibault Ramage. Challenging the Wallacean shortfall: A total assessment of insect diversity on Guadeloupe (French West Indies), a checklist and bibliography. Insecta Mundi, 2020, 0786, pp.1-183. mnhn-04171729

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8-28-2020

# Challenging the Wallacean shortfall: A total assessment of insect diversity on Guadeloupe (French West Indies), a checklist and bibliography

François Meurgey

Muséum d'Histoire Naturelle, Nantes, francois.meurgey@nantesmetropole.fr

Thibault Ramage

Paris, France, thibault.ramage@hotmail.fr

Follow this and additional works at: https://digitalcommons.unl.edu/insectamundi

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Part of the Ecology and Evolutionary Biology Commons, and the Entomology Commons

Meurgey, François and Ramage, Thibault, "Challenging the Wallacean shortfall: A total assessment of insect diversity on Guadeloupe (French West Indies), a checklist and bibliography" (2020). *Insecta Mundi*. 1281.

https://digitalcommons.unl.edu/insectamundi/1281
Challenging the Wallacean shortfall:
A total assessment of insect diversity on Guadeloupe
(French West Indies), a checklist and bibliography

François Meurgey

Entomology Department, Muséum d'Histoire Naturelle
12 rue Voltaire
44000 Nantes, France

Thibault Ramage

UMS 2006 PatriNat, AFB-CNRS-MNHN
36 rue Geoffroy St Hilaire
75005 Paris, France

Date of issue: August 28, 2020
François Meurgey and Thibault Ramage

Challenging the Wallacean shortfall: A total assessment of insect diversity on
Guadeloupe (French West Indies), a checklist and bibliography

Insecta Mundi 0786: 1–183

ZooBank Registered: urn:lsid:zoobank.org:pub:FBA700C6-87CE-4969-8899-FDA057D6B8DA

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Table of Contents

<table><tr><td>Introduction</td><td>1</td></tr><tr><td>The Islands</td><td>2</td></tr><tr><td>Materials and Methods</td><td>3</td></tr><tr><td>Results</td><td>4</td></tr><tr><td>Checklist of Species</td><td>9</td></tr><tr><td>Order Zygentoma</td><td>9</td></tr><tr><td>Order Ephemeroptera</td><td>9</td></tr><tr><td>Order Odonata</td><td>10</td></tr><tr><td>Order Orthoptera</td><td>13</td></tr><tr><td>Order Phasmida</td><td>15</td></tr><tr><td>Order Dermaptera</td><td>16</td></tr><tr><td>Order Mantodea</td><td>17</td></tr><tr><td>Order Blattodea</td><td>17</td></tr><tr><td>Order Psocodea</td><td>20</td></tr><tr><td>Order Thysanoptera</td><td>21</td></tr><tr><td>Order Hemiptera</td><td>25</td></tr><tr><td>Order Hymenoptera</td><td>45</td></tr><tr><td>Order Strepsiptera</td><td>55</td></tr><tr><td>Order Coleoptera</td><td>55</td></tr><tr><td>Order Neuroptera</td><td>122</td></tr><tr><td>Order Trichoptera</td><td>123</td></tr><tr><td>Order Lepidoptera</td><td>126</td></tr><tr><td>Order Diptera</td><td>151</td></tr><tr><td>Order Siphonaptera</td><td>164</td></tr><tr><td>Conclusion: The Wallacean Shortfall</td><td>164</td></tr><tr><td>Acknowledgments</td><td>165</td></tr><tr><td>Literature Cited</td><td>165</td></tr></table>
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# Challenging the Wallacean shortfall: A total assessment of insect diversity on Guadeloupe (French West Indies), a checklist and bibliography

## François Meurgey

Entomology Department, Muséum d'Histoire Naturelle
12 rue Voltaire
44000 Nantes, France
francois.meurgey@nantesmetropole.fr

## Thibault Ramage

UMS 2006 PatriNat, AFB-CNRS-MNHN
36 rue Geoffroy St Hilaire
75005 Paris, France
thibault.ramage@hotmail.fr

**Abstract.** This work is the first attempt to integrate into one list and quantify all the known species of Guadeloupean insects. It includes all the species known to us to be reported for the island of Guadeloupe until the 31st of December 2019. A total of 3097 valid species are listed, of which the largest component is the beetles (1447 species). Preliminarily, 632 species (20%) are considered to be strictly endemic to Guadeloupe, and 250 species are recognized as adventive or introduced. The bibliography complements the taxonomic information and includes over 350 titles. Brief annotations are also given on the history of entomology in Guadeloupe. This list includes four *new island records* for Guadeloupe: *Azeta rhodogaster* Guenée, 1852 (Lepidoptera: Erebidae), *Eudocima apta* (Walker, 1857) (Lepidoptera: Erebidae), *Urania leilus* (Linnaeus, 1758) (Lepidoptera: Uraniidae) and *Micronotus quadriundulatus* (Redtenbacher, 1892) (Orthoptera: Tetrigidae), all recently collected in Guadeloupe.

**Key words.** Insecta, database, new island records.

**Résumé.** Ce travail constitue une première tentative d'intégrer dans une même liste et de quantifier l'ensemble des espèces d'insectes actuellement connues en Guadeloupe. Il inclut l'ensemble des espèces répertoriées en Guadeloupe jusqu'au 31 Décembre 2019. Une liste de 3097 espèces valides est donnée, dont la plus grande composante sont les Coléoptères (1447 espèces). Les espèces endémiques strictes de Guadeloupe, en l'état actuel de nos connaissances, sont au nombre de 632 (20%) et 250 espèces sont reconnues comme introduites. Une bibliographie complète de plus de 350 entrées est également donnée, ainsi que de brèves informations sur le statut actuel des connaissances. La liste présentée comprend quatre nouvelles mentions pour la Guadeloupe; *Azeta rhodogaster* Guenée, 1852 (Lepidoptera: Erebidae), *Eudocima apta* (Walker, 1857) (Lepidoptera, Erebidae), *Urania leilus* (Linnaeus, 1758) (Lepidoptera: Uraniidae) et *Micronotus quadriundulatus* (Redtenbacher, 1892) (Orthoptera: Tetrigidae), récemment capturés en Guadeloupe.

**Mots-clés.** Insectes, base de données, nouvelles mentions.

## Introduction

This work addresses the questions of “What and how much do we know about the diversity of insects in Guadeloupe?” Until now the answers to these questions were out of our reach, as the data on Guadeloupean species are mostly dispersed across hundreds of zoological publications. To our knowledge, the only previous publication dealing with the quantification of the known arthropod fauna of Guadeloupe was a short popular summary commenting on such efforts (Redaud 1996).

In spite of being one of the larger faunas in the Lesser Antilles, Guadeloupean insects have only recently received significant attention from taxonomists. This checklist integrates, for the first time, updated knowledge on the insect fauna of this island. The publication of several earlier checklists provided an important starting point to attempt the compilation of a comprehensive list of the known Guadeloupean insects.
The various groups treated were the Coleoptera (Chalumeau 1983; Chalumeau and Touroult 2005; Peck et al. 2014), Ephemeroptera (Hofmann et al. 1999), Odonata (Meurgey and Picard 2011), Orthoptera (Perez-Gelabert 2001), Phasmatodea (Lelong and Langlois 2001), Thysanoptera, and Diptera (Etienne 2005; Etienne et al. 2003, 2015). Some of these publications have already had the desired effect of eliciting more studies in those groups.

## The Islands

**Physical geography.** With an area of 1434 km² (554 mi²) bipartite Guadeloupe is the largest island in the Lesser Antilles. It is located between 15°50' N to 16°31' N and 61°00' W to 61°48' W. It lies between the islands of Montserrat (53 km to the northwest) and Dominica (35 km to the south-southeast) in the Leeward Islands group, in the northern half of the Lesser Antilles (Fig. 1).

Guadeloupe is composed of the main pair of islands of Grande-Terre and Basse-Terre, which are collectively called Guadeloupe, the smaller satellite islands of Marie-Galante and La Désirade, and five small islands collectively called Les Saintes. The maximum elevation of 1467 m is at the summit of la Soufrière volcano, Basse-Terre, the highest point in the Lesser Antilles. Guadeloupe is a tropical island which has been heavily altered by clearing for large- and small-scale agriculture (especially on Grande-Terre), but with a significant area of coastal mangrove and regenerating or little-disturbed lowland seasonal forest and submontane wet forest (especially on Basse-Terre).

**Habitats.** Much of the island of Basse-Terre is still forested, and these areas are mostly public forest lands administered by the Parc National de Guadeloupe. The rest is urban, suburban, small scale agriculture,

<figure><img src="image_5.png" /><figcaption>**Figure 1.** The islands of the Caribbean, with an arrow pointing to Guadeloupe in the Lesser Antilles.</figcaption></figure>
and large agricultural plantations, especially of sugarcane and banana.

There is a diversity of altered and natural habitats from coastal mangrove, dry coastal thorn scrub, seasonally xeric forest and seasonal tropical humid forests grading into submontane rain forest. Most of the forest exists in the steep uplands of Basse-Terre and in the National Park. These forests are not easy of access except along a well-developed system of forestry roads and over 150 km of hiking trails. The vegetation types have been described by Stehlé (1936, 1946, 1949) and Beard (1949), who stated that Guadeloupe contains the greatest and finest extent of mountain forests in the Lesser Antilles. A comprehensive ecological map based on potential vegetation, with an interpretative text, is available for the Guadeloupe archipelago (Rousteau et al. 1996).

## Materials and Methods

**Construction of the checklist of known species.** In 2011, we started a Guadeloupean list, as a part of our on-going studies of the insect fauna of the Lesser Antilles. A preliminary checklist of 2710 species was published on the behalf of the Parc National de Guadeloupe and integrated into the French national taxonomic database (TAXREF: Gargominy et al. 2017) on behalf of the natural heritage inventory of France (MNHN, online). Since then, the French taxonomic database was updated on a yearly basis with the version 13.0 available online (http://inpn.mnhn.fr/programme/referentiel-taxonomique-taxref). The checklist is also available online (http://inpn.mnhn.fr/programme/referentiel-taxonomique-taxref?lg=en) and will be updated on a yearly basis.

Between 2011 and December 2019 we returned to the list and started to correct it, to add new published records and rearrange the species list according to the taxonomic changes. This was done by checking carefully all the available publications on the insects from the Lesser Antilles the Caribbean and more generally all the published works on the New World Insecta, except data from technical reports or internet records. Publications were kindly provided by authors, colleagues or searched from the Zoological Record. It is likely that, for some groups, certain lists are incomplete, and some species names are obsolete. This work is based only on published data. No doubt some researchers are aware of or already described or collected more species than listed here. We know, for example, of several on-going and unpublished studies (Orthoptera, Blattodea, Hymenoptera...) containing newly recorded species and even new species.

**Format of the checklist.** This list is based on published records, specimens in collections, and our surveys in Guadeloupe, and was compiled in a spreadsheet with the taxonomic hierarchy of family, subfamily, tribe (occasionally), genus, species, subspecies, author's name and the first citation for each taxon, followed by more recent records from the literature. For this second effort, we focused only on valid taxa. These lists were coded using the TAXREF format (Michel et al. 2017) in order to integrate the species database. Data on endemism and introduction from the literature were coded as *** = single island endemic, ** = endemic to one or more Lesser Antillean islands, * = Caribbean endemic and I = Introduced or adventive.

Species are presented alphabetically within the class, orders and families. The list does not attempt to provide a strict and modern classification for all groups. The classificatory arrangements within the orders of insects are sometimes phylogenetic, following Zhang (2013), but most often the families are listed alphabetically. The classification of orders follows Zhang (2013), notably concerning orders Blattodea (Beccaloni and Eggleton 2013) and Mantodea (Otte et al. 2013).

Classification details vary among groups, mostly according to the level of available knowledge. As knowledge of phylogenetic relationships grows, some groups will experience significant taxonomic changes. An attempt was made to update and correct the species names through the assistance of the most recent taxonomic catalogs available.

These procedures have hopefully minimized the number of errors and out of date names in this checklist. In any case, this checklist should not be considered a finished catalog nor a taxonomic reference, but a starting point towards a more complete taxonomic inventory of the Guadeloupean insects. Throughout this work, an attempt is made to identify and quantify species that are unique or endemic to the island, from the literature. Data on endemicity should become more refined as the West Indian arthropod fauna becomes better known.
**Bibliographic references and species records.** The bibliography (i.e. Literature Cited) complements the species list and facilitates access to the necessary information for the study, rational use, and conservation of this fauna. It includes over 350 titles of mixed biological disciplines, containing useful information on Guadeloupean insects. For Coleoptera, we choose to list all the species included in the works of Peck et al. (2014, 2016) which are recent syntheses of the known species. For other orders, we tried to obtain most of the known references, even if they do not deal with the fauna of Guadeloupe. This work was done by checking over 3000 titles dealing with the fauna of Guadeloupe and the West Indies in general, with the help of experts, Google Scholar, and the Zoological Record.

**New records.** While it is not the intention of this paper to extensively list new island records for Guadeloupe from museum specimens or our field work, some have been included. These are indicated with a statement “new island record” along with the name of the person making the identification, the acronym of the collection and the details of the observation.

## Results

All taxa are listed under order, family, subfamily, sometimes tribe and genus. Species considered to be introduced or adventive are marked with a “I”, endemic species and genera are marked as follows: *: Caribbean endemic, ** Lesser Antillean Endemic, *** Single island endemic.

**Assessment of the total Guadeloupe island insect fauna.** There is a total of 29 extant insect orders worldwide (Zhang 2013), 19 of which are represented in Guadeloupe (Table 1). Two new additional orders, Embioptera and Raphidioptera, were recently found by respectively the senior author and

<table><caption>**Table 1.** Overall number of species known by order in Guadeloupe. Si End=single island endemic; WI End=West Indian endemic.</caption><thead><tr><td>Order</td><td>Species</td><td>Genera</td><td>Families</td><td>Si End</td><td>WI End</td><td>Introduced</td></tr></thead><tbody><tr><td>Blattodea</td><td>51</td><td>35</td><td>7</td><td>8</td><td>0</td><td>3</td></tr><tr><td>Coleoptera</td><td>1447</td><td>758</td><td>77</td><td>496</td><td>362</td><td>99</td></tr><tr><td>Dermaptera</td><td>14</td><td>10</td><td>5</td><td>0</td><td>0</td><td>1</td></tr><tr><td>Diptera</td><td>304</td><td>61</td><td>23</td><td>33</td><td>36</td><td>109</td></tr><tr><td>Ephemeroptera</td><td>11</td><td>9</td><td>4</td><td>9</td><td>0</td><td>0</td></tr><tr><td>Hemiptera</td><td>385</td><td>279</td><td>68</td><td>20</td><td>3</td><td>8</td></tr><tr><td>Hymenoptera</td><td>190</td><td>79</td><td>20</td><td>33</td><td>7</td><td>19</td></tr><tr><td>Lepidoptera</td><td>463</td><td>309</td><td>21</td><td>7</td><td>10</td><td>6</td></tr><tr><td>Mantodea</td><td>1</td><td>1</td><td>1</td><td>0</td><td>1</td><td>0</td></tr><tr><td>Neuroptera</td><td>11</td><td>9</td><td>3</td><td>1</td><td>0</td><td>0</td></tr><tr><td>Odonata</td><td>38</td><td>24</td><td>5</td><td>2</td><td>2</td><td>0</td></tr><tr><td>Orthoptera</td><td>30</td><td>24</td><td>7</td><td>8</td><td>2</td><td>3</td></tr><tr><td>Phasmida</td><td>9</td><td>7</td><td>3</td><td>5</td><td>0</td><td>0</td></tr><tr><td>Psocoptera</td><td>17</td><td>10</td><td>9</td><td>0</td><td>0</td><td>0</td></tr><tr><td>Siphonaptera</td><td>1</td><td>1</td><td>1</td><td>0</td><td>0</td><td>1</td></tr><tr><td>Strepsiptera</td><td>1</td><td>1</td><td>1</td><td>0</td><td>0</td><td>0</td></tr><tr><td>Thysanoptera</td><td>76</td><td>52</td><td>4</td><td>0</td><td>0</td><td>1</td></tr><tr><td>Trichoptera</td><td>42</td><td>20</td><td>9</td><td>10</td><td>21</td><td>0</td></tr><tr><td>Zygentoma</td><td>6</td><td>4</td><td>2</td><td>0</td><td>1</td><td>0</td></tr><tr><td>Total</td><td>3097</td><td>1693</td><td>270</td><td>632</td><td>445</td><td>250</td></tr></tbody></table>
Gérard Chovet (pers. comm.) in the wet forests of Basse-Terre, though these were collected too recently to include in this checklist. The absence of the remaining eight orders is noted: Plecoptera, Mecoptera, Megaloptera, Archaeognatha, Grylloblattodea, Mantophasmatodea, Zoraptera, and Phthiraptera. The lack of Grylloblattodea and Mantophasmatodea is easily explained by their distributions, Asia and North America for the former, and Africa for the latter. Some of these taxa marginally succeeded in colonizing the forest floor in some tropical regions but this colonization didn't happen in Guadeloupe. The other orders in the list could be either a genuine absence of these orders, a lack of publication on the subject or a lack of specific research in the literature on these groups. An obvious lack of study is the absence of records of Phthiraptera (bird lice). There must be many species on the birds of Guadeloupe. Overall, 3097 species are inventoried from Guadeloupe. Coleoptera accounts for almost the half of these taxa (47%), followed by Lepidoptera (15%) and Hemiptera (13%).

**Early mode of colonization.** The Lesser Antillean islands were colonized primarily from continental America and adjacent islands by overwater dispersal (Peck et al. 2014), islands playing the role of "stepping stones" and facilitating dispersal across the Caribbean. Some orders, such as Odonata, have reached islands by aerial colonization, and this colonization is still active with the presence of two African species regularly recorded in the Lesser Antilles; *Hemianax ephippiger* (Burmeister) and *Tramea basilaris* (Palisot de Beauvois) (Meurgey 2006b; Meurgey and Picard 2011).

**Comments on the list.** Zygentoma are represented by six species. This order is in need of further study. We know only one old publication dating from 1972. The Ephemeroptera consists of 11 species relatively recently studied (Hofmann et al. 1999) in which eight are endemic to Guadeloupe (73%).

Order Odonata is relatively rich in Guadeloupe with 38 species and consists of 29 Anisoptera and 9 Zygoptera. The paucity of zygopterans is explained by the fact that the species in this suborder possess less flight capabilities than anisopterans (see Corbet 1999). Endemism in odonate fauna is low with only five species (Meurgey and Picard 2011). Two are single island endemics, three others are Lesser Antilles endemics and two species are West Indian endemics. This order has been particularly well studied in Guadeloupe during the past 15 years (Meurgey and Picard 2011), and research on taxonomy has been carried out. These studies proved *Macrothemis meurgeyi* Daigle, 2007 and *Brechmorhoga archboldi* (Donnelly, 1970) to be junior synonyms of other known Antillean or continental species (Meurgey 2016a, b, c). As a result, the endemic species in Guadeloupe are zygopterans. This is realistic considering speciation in this suborder due to the limited dispersal of species.

Two of the 30 species of Orthoptera in Guadeloupe are endemic: *Karukerana aguilari* Bonfils, 1965 and *Rhumosa bolognei* Hugel and Desutter-Grandcolas, 2018. This fauna has been relatively well studied, but there are undoubtedly several species from the canopy that still need to be described. We know about several new island records and several new species to be described (S. Hugel pers.comm.).

Nine species of the order Phasmida are recorded from Guadeloupe, with five endemic to Guadeloupe. This order has been well-studied in the past ten years and is considered as well known in Guadeloupe, except for their ecology.

Dermaptera are represented by 14 species but none are considered endemic. This order needs more study in Guadeloupe.

Mantodea are poorly represented with only one species, *Oligonyx insularis* Bonfils, described from Guadeloupe in 1961. This small, uncommon species lives in semi-deciduous forests and nothing is known about its ecology. The species *Acontista multicolor* (Saussure, 1870) is considered by many authors to be present in Guadeloupe. Saussure (1870) described this species based on specimens from "Guadalupe," not "Guadeloupe". He also described two other species in this genus from French Guiana and Brazil, countries located on the mainland. We have encountered the label "Guadalupe" for many species of dragonflies considered as present in Guadeloupe and other islands that were in fact described from South America. Considering that (i) in 1870 the word Guadeloupe was widely known to refer to the Lesser Antillean island, (ii) the lack of specimens of this species in collections, (iii) the lack of recent records from the West Indies, and (iv):- the fact that Guadalupe is also an island and/or province of Mexico, Brazil, Costa-Rica and even Colombia; we consider *Acontista multicolor* absent from Guadeloupe and the mention from the island to be erroneous due to the misinterpretation of the type locality.
Blattodea are more diverse, with 51 species in which eight are endemic. Most of the endemic cockroaches are found in the drier parts of the islands, especially on Grande-Terre. Some of these are synanthropic species and commonly live in houses.

Thysanoptera are represented by 76 species, all widely distributed over the Caribbean and the mainland. Almost all of them are considered agricultural pests.

Hemiptera are represented in Guadeloupe by 385 species, belonging to 68 families and including 20 single-island endemic species. More studies are needed for this order in Guadeloupe, especially on the non-agricultural pest species. In this order, family Cicadidae is represented by two species, one being endemic to the Caribbean, *Tympanoterpes grisea* (Germar, 1830), and another introduced from South America *Fidicina mannifera* (Fabricius, 1803).

Hymenoptera are represented by 20 families and 190 species in Guadeloupe, with 40 endemic species, and no less than 19 introduced species, especially in the family Formicidae. Within this order, Anthophila were poorly known prior to Meurgey (2014) and Meurgey et al. (2015), which focused on the systematics and distribution of species. These works have increased the number of species from six to 24 and have contributed to a better understanding of both biology and ecology of bees. Additionally, there are at least seven new species of Halictidae from Guadeloupe that have not yet been described (A. Pauly pers. comm.). Compared to the Coleoptera (1447 species), Lepidoptera (464 species), Hemiptera (390 species) and Diptera (302 species), the order Hymenoptera is still poorly known in Guadeloupe. The number of species in the parasitoid families will undoubtedly increase in the future; recent surveys conducted by the authors in Guadeloupe have yielded six new records of Aculeata species, demonstrating the undersampling of this order (unpublished data).

Coleoptera are by far the most diverse insects in Guadeloupe, with 1447 species to date, roughly 41% of which are endemics (593 spp.; Peck et al. 2014). Families Curculionidae (445 species) and Staphylinidae (154 species) are the most diverse among all the Coleoptera, containing 41% of the species total, followed by Chrysomelidae (99 species), Scarabaeidae (74 species) and Cerambycidae (71 species). The number of beetles recorded from Guadeloupe is consistent with the fact that the order Coleoptera is by far the most species-rich worldwide (Zhang 2013). It also reflects the fact that Coleoptera have been studied more in Guadeloupe than on other Lesser Antillean islands (Peck 2009, 2011, 2016; Peck et al. 2012, 2014).

In spite of its agricultural interest as beneficial insects, the order Neuroptera has not been studied since the 1970s, with a list of species for the Caribbean (Penny 1977). Recent collecting in Guadeloupe by the senior author has increased the number of species in the family Myrmeleontidae from two to five (Giacomino 2015), all widely distributed in tropical America. More recently, new collection records from Guadeloupe revealed the presence of an additional species of Mantispidae (Hoffman et al. 2017).

Trichoptera are represented by 42 species, 21 being endemic to Guadeloupe (50%). This order has been studied by Botosaneanu (2000, 2002) and is considered well-known in Guadeloupe.

Lepidoptera are represented by 463 species in Guadeloupe, of which 407 are moths and 57 are butterflies (Papilionoidea). Only seven species are endemic to Guadeloupe, and five others restricted to the Lesser Antilles. This order has been well studied by Brévignon (2002) for the butterflies and by Zagatti et al. (2006) for the moths. Family Pterophoridae has been studied recently (Bigot and Etienne 2009) and is considered well-known.

Diptera are represented by 304 species, with 69 endemic species (23%). These species are distributed into 23 families. Family Agromyzidae is the richest among Diptera with 88 species recorded, followed by Ceratopogonidae (49 species) and Culicidae (24 species). This order is badly in need of study in the Lesser Antilles and several new species have yet to be described, especially in the families Asilidae and Bombyliidae.

Siphonaptera are represented by only one species in Guadeloupe, *Ctenocephalides felis felis* (Bouché, 1835), introduced by man.

**Level of knowledge.** The Guadeloupean insect fauna is better known than any other Lesser Antillean island due to significant advances made in the last 20 years. Our actual knowledge is mainly due to efforts of the large amateur and professional community involved in collecting material for study and in taxonomic work mainly from France and other countries. The most significant of these, have come from Guadeloupe, are the books by Chalumeau (1983) on scarabaeid beetles, by Chalumeau and Touroult (2005) on cerambycid beetles and by Meurgey and Picard on Odonata (2011).
Most significant taxonomic works are mainly due to researchers from France, such as Botosaneanu (2002) on caddisflies, Etienne (see references) on Hemiptera, Meurgey (see references) on Odonata, Lelong and Langlois (2001) on phasmids, and Brévignon and Brévignon (2011) on butterflies. Surveys in protected areas at the request of managers and in general on Guadeloupe has also allowed for obtaining of material from remote areas. However, this survey is far from complete, even for some groups with a relative high level of knowledge, such as Phasmida, Dermaptera and Coleoptera (in absolute numbers), which has long been the subject of intense collecting and taxonomic work by mostly French and US researchers.

Relative to its size, the insect fauna of Guadeloupe is better known than the fauna of the other islands of the Lesser Antilles and can be compared only to some of the better-known Greater Antillean islands (see Table 2) (Peck et al. 2012, 2014). In the Lesser Antilles, the best-known fauna is that of Grenada which as an outdated preliminary catalog of its insects (Woodruff et al. 1998), which nonetheless should help in the compilation of a complete catalog. Despite the fact that we know of many publications for the other Lesser Antillean islands, such as St. Vincent or Dominica no catalog or synthesis is actually available. Because of the very preliminary documentation of this fauna, and despite the dramatic environmental changes that have occurred in Guadeloupe in the last 200 years, we basically have no data about the conservation status of insect populations. It is probable that many local extinctions have occurred, especially on Grande-Terre, due to the deforestation of extensive areas of the island. Table 1 summarizes the number of species found in this checklist per major arthropod group, also including data on endemic species.

Table 2 summarizes knowledge on other island insect faunas. In the older Greater Antilles, the rich insect fauna is mainly due to the size of the islands, their proximity to the mainland and probably the older geological history of this archipelago. These have allowed the accumulation of a higher diversity in their insect faunas. In the Lesser Antilles, Grenada is the only well-documented island, regarding its insects as a whole. This island possesses a richness similar to that of Guadeloupe despite its smaller size, but this is correlated with its closer distance to the mainland. The relatively small number of insects in Guadeloupe compared to Grenada could be better explained by a lack of surveys in Guadeloupe along with a greater level of alteration of its environment.

**Endemicity.** Among the 3097 insect species recorded from Guadeloupe, 1077 (35%) are considered endemic to the Caribbean. Of those endemics, 632 (59%) are endemic to Guadeloupe, and 445 (41%) are found on other West Indian islands. The vast majority of known endemics are Coleoptera; they account for 78% of single-island endemic species and for 80% of the total number of endemic species. Diptera account for 5% of single island endemic taxa and 6% of the total endemic taxa. These analyses should be considered with caution, due to the fact that the knowledge of endemicity is more limited for other insect orders.

**Introduced species.** Many species of plants and animals have been introduced into Guadeloupe by industry, agriculture and commerce. In this checklist, a total of 250 species are identified as introduced or adventive (8%). These numbers are likely to be gross underestimates that hopefully will become more accurate with further studies on this fauna. Most of the introduced species are not considered to be invasive and have often been established for a long time in Guadeloupe. Diptera account for 45% of introduced species, followed by Coleoptera (3%) and Hymenoptera (1%). In the order Hymenoptera, almost all the introduced species are ants, which were probably introduced along with exotic plants. From a general point of view, almost all the listed introduced species are either agricultural pests, such as Hemiptera, Hymenoptera and some Coleoptera, or biological control agents used against other introduced species. However, there are also some ectoparasitic species introduced with vertebrates during European colonization, such as the cat flea *Ctenocephalides felis* (Siphonaptera).
<table><caption>Table 2. Comparison of the recorded number of insect species between Guadeloupe and other oceanic islands and island groups.</caption><thead><tr><th rowspan="2"></th><th colspan="3">Greater Antilles</th><th colspan="2">Lesser Antilles</th><th colspan="2">Macaronesia</th><th colspan="2">Pacific Ocean</th></tr><tr><th>Cuba</th><th>Hispaniola</th><th>Puerto Rico</th><th>Bermuda</th><th>Guadeloupe</th><th>Grenada</th><th>Canary Islands</th><th>Galapagos Islands</th><th>French Polynesia</th><th>Samoa</th></tr></thead><tbody><tr><th>Area (km²)</th><td>110,860</td><td>76,480</td><td>9,104</td><td>53,3</td><td>1,434</td><td>340</td><td>7,447</td><td>7,900</td><td>4,167</td><td>2,934</td></tr><tr><th>Distance from mainland (km)</th><td>781</td><td>927</td><td>829</td><td>1695</td><td>860</td><td>234</td><td>250</td><td>1356</td><td>6000</td><td>3923</td></tr><tr><th># of spp.</th><td>8,316</td><td>5,676</td><td>5,066</td><td>1,218</td><td>3,097</td><td>2,101</td><td>6,042</td><td>1,616</td><td>2,556</td><td>2,523</td></tr><tr><th>Reference</th><td>Genaro and Tejuca 2001</td><td>Perez-Gelbert 2008</td><td>Maldonado Capriles 1996</td><td>Sterrer 1998</td><td>This work</td><td>Woodruff et al. 1998</td><td>Izquierdo et al. 2004</td><td>Bensted-Smith 2002</td><td>Ramage 2017</td><td>Kami and Miller 1998</td></tr></tbody></table>
# Checklist of Species

## CLASS Insecta

SUBCLASS Dicondylia Hennig, 1953.

ORDER Zygentoma Börner, 1904. (6 SPECIES).

Key reference: Wygodzinsky (1972).

## FAMILY Lepismatidae Latreille, 1802.

Genus *Ctenolepisma* Escherich, 1905.

*Ctenolepisma dubitalis* Wygodzinsky, 1959. Wygodzinsky (1972); Meurgey (2011b). *

*Ctenolepisma targioniana* Sivestri, 1908. Wygodzinsky (1972); Meurgey (2011b).

*Ctenolepisma versluysi* Escherich, 1905. Wygodzinsky (1959); Meurgey (2011b).

Genus *Heterolepisma* Escherich, 1905.

*Heterolepisma horni* Stach, 1933. Wygodzinsky (1972); Meurgey (2011b).

Genus *Stylifera* Stach, 1932.

*Stylifera impudica* (Escherich, 1905). Wygodzinsky (1972); Meurgey (2011b).

## FAMILY Nicoletiidae Escherich, 1905.

Genus *Nicoletia* Gervais, 1843.

*Nicoletia phytophila* Gervais, 1844. Mendes (1986).

## INFRACLASS Paleoptera Martynov, 1923.

ORDER Ephemeroptera Hyatt and Arms, 1890. (11 SPECIES)

SUPERFAMILY Baetoidea Ulmer, 1920.

FAMILY Baetidae Leach, 1815.

Genus *Americabaetis* Kluge, 1992.

*Americabaetis humilis* Hofmann and Thomas, 1999. Hofmann et al. (1999).

*Americabaetis spinosus* Hofmann and Thomas, 1999. Hofmann, Sartori and Thomas (1999).***

Genus *Callibaetis* Eaton, 1881.

*Callibaetis floridanus* Banks, 1900. Hofmann et al. (1999).

Genus *Cloeodes* Traver, 1938.

*Cloeodes caraibensis* Hofmann and Thomas, 1999. Hofmann et al. (1999).***

Genus *Fallceon* Waltz and McCafferty, 1987.

*Fallceon ater* Hofmann and Thomas, 1999. Hofmann et al. (1999). Hofmann et al. (1999).***

## SUPERFAMILY Caenoidea Newman, 1853.

FAMILY Caenidae Newman, 1853.

Genus *Caenis* Stephens, 1835.

*Caenis catherinae* Hofmann and Thomas, 1999. Hofmann et al. (1999).***

*Caenis femina* Hofmann and Thomas, 1999. Hofmann et al. (1999).***

## SUPERFAMILY Ephemeroelloidea Klapalek, 1909.

FAMILY Leptohyphidae Edmunds and Traver, 1954.

Genus *Allenhyphes* Hofmann and Sartori, 1999. Hofmann et al. (1999).

*Allenhyphes flinti* (Allen, 1973). Hofmann, et al. (1999).***

Genus *Leptohyphes* Eaton, 1882.

*Leptohyphes guadeloupensis* Hofmann and Thomas, 1999. Hofmann et al. (1999).***
Genus *Tricorythodes* Ulmer, 1920.

*Tricorythodes griseus* Hofmann and Thomas, 1999. Hofmann et al. (1999).***

SUPERFAMILY Leptophlebioidea Banks, 1900.

FAMILY Leptophlebiidae Banks, 1900.

Genus *Hagenulopsis* Ulmer, 1919.

*Hagenulopsis guadeloupensis* Hofmann and Thomas, 1999. Hofmann et al. (1999).***

ORDER Odonata Fabricius, 1793. (38 species).

SUBORDER Anisoptera Selys, 1854.

SUPERFAMILY Aeshnoidea Leach, 1815.

FAMILY Aeshnidae Rambur, 1842.

Genus *Anax* Leach, 1815.

*Anax amazili* (Burmeister, 1839). Meurgey and Perron (2004); Meurgey (2005); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

*Anax concolor* Brauer, 1865. Meurgey and Perron (2004); Meurgey (2004d); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

*Anax junius* (Drury, 1773). Klots (1932); Goyaud (1994); Grand (1996); Hofmann (1999); Grand (2000); Pincebourde (2000); Grand (2002); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

Genus *Coryphaeschna* Williamson, 1903.

*Coryphaeschna adnexa* (Hagen, 1861). Goyaud (1994); Hofmann (1999); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

Genus *Hemianax* Selys, 1883.

*Hemianax ephippiger* (Burmeister, 1839). Meurgey (2006b); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

Genus *Rhionaeschna* Förster, 1909.

*Rhionaeschna psilus* (Calvert, 1947). Pincebourde (2000); Pont (2004); Meurgey (2004c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

Genus *Triacanthagyna* Selys, 1883.

*Triacanthagyna caribbea* Williamson, 1923. Goyaud (1994); Meurgey (2004b); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

*Triacanthagyna septima* (Selys in Sagra, 1857). Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

SUPERFAMILY Libelluloidea Rambur, 1842.

FAMILY Libellulidae Rambur, 1842.

Genus *Brachymesia* Kirby, 1889.

*Brachymesia furcata* (Hagen, 1861). Klots (1932); Goyaud (1994); Hofmann (1999); Pincebourde (2000); Grand (2002); Meurgey and Williamson (2002); Levasseur (2006); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

*Brachymesia herbida* (Gundlach, 1889). Goyaud (1994); Hofmann (1999); Dupont (2000); Grand (2000); Jacquemin (2000); Mashaal (2000); Pincebourde (2000); Grand (2002); Meurgey and Williamson (2002); Ilbert and Menegaux (2004); Pont (2004); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

Genus *Brechmorhoga* Kirby, 1894.

*Brechmorhoga praecox grenadensis* (Kirby, 1894). Donnelly (1970); Hofmann (1999); Grand (2002); Meurgey (2007); Meurgey (2008b); Meurgey and Picard (2011); Meurgey (2013) as Brechmorhoga archboldi Donnelly, 1970; Meurgey (2016b).
Genus *Dythemis* Hagen, 1861.

*Dythemis sterilis* Hagen, 1861. Klots (1932); Goyaud (1994); Hofmann (1999); Dupont (2000); Grand (2000); Jacquemin (2000); Mashaal (2000); Pincebourde (2000); Grand (2002); Meurgey and Williamson (2002); Pont (2003); Ilbert and Menegaux (2004); Pont (2004); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey and Poiron (2011); Meurgey (2013).

Genus *Erythemis* Hagen, 1861.

*Erythemis vesiculosa* (Fabricius, 1775). Goyaud (1994); Grand (1996); Hofmann (1999); Dupont (2000); Grand (2000); Jacquemin (2000); Mashaal (2000); Pincebourde (2000); Grand (2002); Meurgey and Williamson (2002); Pont (2003); Ilbert and Menegaux (2004); Pont (2004); Levasseur (2006); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

Genus *Erythrodiplax* Brauer, 1868.

*Erythrodiplax berenice* (Drury, 1773). Meurgey (2004a); Meurgey (2007).

*Erythrodiplax umbrata* (Linnaeus, 1758). Klots (1932); Goyaud (1994); Grand (1996); Hofmann (1999); Dupont (2000); Grand (2000); Mashaal (2000); Pincebourde (2000); Grand (2002); Meurgey and Williamson (2002); Pont (2003); Ilbert and Menegaux (2004); Pont (2004); Levasseur (2006); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

Genus *Macrothemis* Hagen, 1868.

*Macrothemis celeno* (Selys in Sagra, 1857). Goyaud (1994); Hofmann (1999); Jacquemin (2000); Mashaal (2000); Pincebourde (2000); Grand (2002); Pont (2003); Ilbert and Menegaux (2004); Pont (2004); Daigle (2007); Meurgey (2007); Meurgey (2009); Meurgey (2011a); Meurgey and Picard (2011); Meurgey (2013) (as *Macrothemis meurgeyi* Daigle, 2007); Meurgey (2016a). ***

Genus *Miathyria* Kirby, 1889.

*Miathyria marcella* (Selys in Sagra, 1857). Goyaud (1994); Dommanget (2000); Dupont (2000); Grand (2000); Levasseur (2006); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

Genus *Micrathyria* Kirby, 1889.

*Micrathyria aequalis* (Hagen, 1861). Goyaud (1994); Grand (1996); Hofmann (1999); Dupont (2000); Grand (2002); Pincebourde (2000); Mashaal (2000); Grand (2002); Meurgey and Williamson (2002); Pont (2004); Levasseur (2006); Meurgey (2006c); Meurgey and Picard (2011).

*Micrathyria didyma* (Selys in Sagra, 1857). Klots (1932); Dupont (2000); Pincebourde (2000); Pont (2004); Levasseur (2006); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

Genus *Orthemis* Hagen, 1861.

*Orthemis macrostigma* (Rambur, 1842). Goyaud (1994); Grand (1996); Hofmann (1999); Dupont (2000); Grand (2000); Jacquemin (2000); Mashaal (2000); Pincebourde (2000); Grand (2002); Meurgey and Williamson (2002); Pont (2003); Ilbert and Menegaux (2004); Pont (2004); Levasseur (2006); Meurgey and Daigle (2007); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

Genus *Pantala* Hagen, 1861.

*Pantala flavescens* (Fabricius, 1798). Goyaud (1994); Hofmann (1999); Grand (2000); Mashaal (2000); Grand (2002); Meurgey and Williamson (2002); Pont (2003); Ilbert and Menegaux (2004); Pont (2004); Meurgey (2005); Levasseur (2006); Meurgey (2006a); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

*Pantala hymenaea* (Say, 1839). Meurgey (2008a); Meurgey and Picard (2011); Meurgey (2013).

Genus *Tauriphila* Kirby, 1889.

*Tauriphila australis* (Hagen, 1867). Guezennec and Guezennec (2009); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

Genus *Tholymis* Hagen, 1867.

*Tholymis citrina* Hagen, 1867. Grand (2000); Meurgey and Williamson (2002); Meurgey and Weber (2005); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).
Genus *Tramea* Hagen, 1861.

*Tramea abdominalis* (Rambur, 1842). Klots (1932); Goyaud (1994); Grand (1996); Hofmann (1999); Dupont (2000); Grand (2000); Jacquemin (2000); Mashaal (2000); Pincebourde (2000); Grand (2002); Meurgey and Williamson (2002); Pont (2003); Ilbert and Menegaux (2004); Pont (2004); Meurgey (2005); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

*Tramea basilaris* (Palisot de Beauvois, 1805). Meurgey (2008a); Meurgey and Picard (2011); Meurgey (2013).

*Tramea binotata* (Rambur, 1842). Hofmann (1999); Pincebourde (2000); Meurgey (2004); Meurgey and Perron (2004); Meurgey (2004c); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

*Tramea calverti* Muttkowski, 1910. Meurgey (2004a); Meurgey (2004e); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

*Tramea insularis* Hagen, 1861; Meurgey and Picard (2011); Meurgey (2013).

SUBORDER Zygoptera Selys, 1854.

SUPERFAMILY Coenagrionoidea Kirby, 1890.

FAMILY Coenagrionidae Kirby, 1890.

Genus *Argia* Rambur, 1842.

*Argia concinna* (Rambur, 1842). Goyaud (1994); Hofmann (1999); Jacquemin (2000); Mashaal (2000); Pincebourde (2000); Grand (2002); Meurgey and Williamson (2002); Pont (2003); Ilbert and Menegaux (2004); Pont (2004); Meurgey (2007); Meurgey (2009); Meurgey (2011a); Meurgey and Picard (2011); Meurgey (2013). **

Genus *Enallagma* Charpentier, 1840.

*Enallagma coecum* (Hagen, 1861). Hofmann (1999); Pincebourde (2000); Levasseur (2006); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013). *

Genus *Ischnura* Charpentier, 1840.

*Ischnura capreolus* (Hagen, 1861). Hofmann (1999); Dommanget 2000; Grand (2000); Pincebourde (2000); Meurgey and Weber (2007); Meurgey (2007); Meurgey and Picard (2011).

*Ischnura hastata* (Say, 1839). Goyaud (1994); Grand (2002); Meurgey and Williamson (2002); Meurgey and Perron (2004); Pont (2004); Meurgey (2006c); Meurgey and Weber (2007); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

*Ischnura ramburii* (Selys in Sagra, 1857). Goyaud (1994); Grand (1996); Hofmann (1999); Dupont (2000); Grand (2000); Jacquemin (2000); Mashaal (2000); Pincebourde (2000); Grand (2002); Meurgey and Williamson (2002); Pont (2003); Ilbert and Menegaux (2004); Pont (2004); Levasseur (2006); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

Genus *Telebasis* Selys, 1865.

*Telebasis corallina* (Selys, 1876). Goyaud (1994); Hofmann (1999); Dommanget 2000; Grand (2002); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

FAMILY Protoneuridae Tillyard, 1917.

Genus *Protoneura* Selys in Sagra, 1857.

*Protoneura romanae* Meurgey, 2006. Meurgey (2006d); Meurgey (2007); Meurgey and Faucheux (2009); Meurgey (2010); Meurgey and Picard (2011); Meurgey (2013). ***

SUPERFAMILY Lestoidea Tillyard, 1913.

FAMILY Lestidae Calvert, 1901.

Genus *Lestes* Leach in Brewester, 1815.

*Lestes forficula* Rambur, 1842. Goyaud (1994); Grand (1996); Hofmann (1999); Jacquemin (2000); Mashaal (2000); Grand (2002); Meurgey and Williamson (2002); Ilbert and Menegaux (2004); Pont (2004); Levasseur (2006); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).
*Lestes tenuatus* Rambur, 1842. Rambur (1842); Goyaud (1994); Hofmann (1999); Meurgey and Williamson (2002); Meurgey (2006c); Meurgey (2007); Meurgey and Picard (2011); Meurgey (2013).

## ORDER Orthoptera Latreille, 1810. (30 SPECIES).

### SUBORDER Caelifera Ander, 1939.

### SUPERFAMILY Acridoidea MacLeay, 1821.

### FAMILY Acrididae MacLeay, 1821.

### SUBFAMILY Cyrtacanthacridinae Kirby, 1902.

### TRIBE Cyrtacanthacridini Kirby, 1910.

Genus *Schistocerca* Stål, 1873.

*Schistocerca nitens caribbeana* Dirsh, 1974. Bland and Desutter-Grandcolas (2003).

*Schistocerca pallens* (Thunberg, 1815). Gruner and Riom (1977); Bland and Desutter-Grandcolas (2003).

### SUBFAMILY Gomphocerinae Fieber, 1853.

### TRIBE Orphulellini Otte, 1979.

Genus *Orphulella* Giglio-Tos, 1894.

*Orphulella punctata* (DeGeer, 1773). Bland and Desutter-Grandcolas (2003).

### TRIBE Scyllinini Brunner von Wattenwyl, 1893.

Genus *Rhammatocerus* Saussure, 1861.

*Rhammatocerus cyanipes* (Fabricius, 1775). Bland and Desutter-Grandcolas (2003).

### FAMILY Tetrigidae Rambur, 1838.

### SUBFAMILY Tetriginae Rambur, 1838.

Genus *Micronotus* Hancock, 1902.

*Micronotus quadriundulatus* (Redtenbacher, 1892).

New island record (Meurgey, MHNN/SHNLH). Guadeloupe, Goyave, Bonfils, 06.VII.2017, 6499174 – 1781730, 86m, F. Meurgey leg (coll. SHNLH). Guadeloupe, Vieux-Habitants, La Grive-lière, 03.II.2018, 365977 – 1777514, 183m, F. Meurgey leg (coll. SHNLH), confirmed by S. Hugel (Pers. Com.).

### SUBORDER Ensifera Chopard, 1922.

### SUPERFAMILY Grylloidea Laicharting, 1781.

### FAMILY Gryllidae Laicharting, 1781.

### SUBFAMILY Eneopterinae Saussure, 1893.

Genus *Antillicharis* Otte and Perez-Gelabert, 2009.

*Antillicharis antillarum* (Saussure, 1874). Strohecker (1953); Bland and Desutter-Grandcolas (2003). ***

*Antillicharis unicolor* (Olivier, 1791). Kirby (1906); Bland and Desutter-Grandcolas (2003). ***

Genus *Carylla* Otte and Perez-Gelabert, 2009.

*Carylla fuscifrons* (Desutter-Grandcolas, 2003). Desutter-Grandcolas (2003); Bland and Desutter-Grandcolas (2003); Otte and Perez-Gelabert (2009).

### SUBFAMILY Gryllinae Laicharting, 1781.

### TRIBE Gryllini Laicharting, 1781.

Genus *Gryllus* Linnaeus, 1758.

*Gryllus assimilis* (Fabricius, 1775). Rehn (1905); Gruner and Riom (1977).
TRIBE Modicogryllini Otte and Alexander, 1983.

Genus *Gryllodes* Saussure, 1874.

*Gryllodes sigillatus* (Walker, 1869). Bland and Desutter-Grandcolas (2003).

SUBFAMILY Luzarinae Hebard, 1928.

SUBTRIBE Amphiacustina Hubbell, 1938.

Genus *Amphiacusta* Saussure, 1874.

*Amphiacusta annulipes* (Serville, 1831). Strohecker (1953).

*Amphiacusta caraiba* (Saussure, 1897). Kirby (1906); Desutter-Grandcolas and Otte (1997).

FAMILY Gryllotalpidae Leach, 1815.

SUBFAMILY Gryllotalpinae Leach, 1815.

Genus *Neocurtilla* Kirby, 1906.

*Neocurtilla hexadactyla* (Perty, 1832). Frank and McCoy (2014). I

SUBFAMILY Scapteriscinae Zeuner, 1939.

Genus *Scapteriscus* Scudder, 1868.

*Scapteriscus abbreviatus* Scudder, 1869. Frank and McCoy (2014). I

*Scapteriscus didactylus* (Latreille, 1804). Maros et al. (2005); Frank and McCoy (2014). I

FAMILY Phalangopsidae Blanchard, 1845.

SUBFAMILY Paragryllinae Desutter-Grandcolas, 1987.

TRIBE Paragryllini Desutter-Grandcolas, 1987.

Genus *Paragryllus* Guérin-Méneville, 1844.

*Paragryllus martini* Guérin-Méneville, 1844. ***

Genus *Unithema* Desutter-Grandcolas, 1991.

*Unithema guadelupensis* Desutter-Grandcolas, 1991. ***

SUPERFAMILY Stenopelmatoidea Burmeister, 1838.

FAMILY Anostostomatidae Saussure, 1859

SUBFAMILY Lutosinae Gorochov, 1988.

Genus *Rhumosa* Hugel and Desutter-Grandcolas, 2018.

*Rhumosa bolognei* Hugel and Desutter-Grandcolas, 2018. ***

FAMILY Gryllacrididae Blanchard, 1845.

SUBFAMILY Gryllacridinae Karny, 1929.

Genus *Abelona* Karny, 1937.

*Abelona guadeloupensis* (Karny, 1935).

SUPERFAMILY Tettigonioidea Krauss, 1902.

FAMILY Tettigoniidae Krauss, 1902.

SUBFAMILY Conocephalinae Burmeister, 1838.

TRIBE Conocephalini Burmeister, 1838.

Genus *Conocephalus* Thunberg, 1815.

*Conocephalus cinereus* Thunberg, 1815. Bland and Desutter-Grandcolas (2003).

TRIBE Copiphorini Karny, 1912.

Genus *Acantheremus* Karny, 1907.

*Acantheremus bonfilsi* Hugel and Morin, 2003. ***
*Acantheremus dominicanus* Naskrecki, 1997. Hugel (2009). **

Genus *Neoconocephalus* Karny, 1907.

*Neoconocephalus maxillosus* (Fabricius, 1775). Bland and Desutter-Grandcolas (2003).

*Neoconocephalus triops* (Linnaeus, 1758). Gruner and Riom (1977).

SUBFAMILY Phaneropterinae Burmeister, 1838.

Genus *Microcentrum* Scudder, 1862.

*Microcentrum triangulatum* Brunner von Wattenwyl, 1878. Kirby (1906); Gruner and Riom (1977); Bland and Desutter-Grandcolas (2003).

Genus *Orophus* Saussure, 1859.

*Orophus decoratus* (Walker, 1869). Bland and Desutter-Grandcolas (2003).

Genus *Philophyllia* Stål, 1873.

*Philophyllia guttulata* Stål, 1873. Gruner and Riom (1977).

SUBFAMILY Pseudophyllinae Burmeister, 1838.

TRIBE Cocconotini Brunner von Wattenwyl, 1895.

Genus *Nesonotus* Beier, 1960.

*Nesonotus reticulatus* (Fabricius, 1793). Kirby (1906); Bonfils (1966); Gruner and Riom (1977); Stumpner et al. (2013). ***

TRIBE Pleminiini Brunner von Wattenwyl, 1895.

Genus *Xerophyllopteryx* Rehn, 1905.

*Xerophyllopteryx fumosa* (Brunner von Wattenwyl, 1895. Kirby (1906); Bonfils (1966); Gruner and Riom (1977); Stumpner et al. (2013). **

TRIBE Pterophyllini Hebard, 1941.

Genus *Karukerana* Bonfils, 1965.

*Karukerana aguilari* Bonfils, 1965. Gruner and Riom (1977); Stumpner et al. (2013).***

ORDER Phasmida Leach, 1815. (9 SPECIES).

FAMILY Diipheromeridae Karny, 1923.

SUBFAMILY Diipheromerinae Kirby, 1904.

TRIBE Diipheromerini Kirby, 1904.

Genus *Bacteria* Latreille, Lepeletier de Saint Fargeau, Audinet-Serville and Guérin-Méneville, 1825.

*Bacteria donskoffi* (Langlois and Lelong, 1998). Lelong and Langlois. (2001). ***

*Bacteria ferula* (Fabricius, 1793). Langlois and Lelong (1997); Lelong and Langlois (1998); Lelong and Langlois (2001).

Genus *Clonistria* Stål, 1875.

*Clonistria guadeloupensis* Redtenbacher, 1908. Langlois and Lelong (1997); Lelong and Langlois (1998); Lelong and Langlois (2001). ***

Genus *Paraclonistria* Langlois and Lelong, 1998.

*Paraclonistria nigramala* Lelong and Langlois, 1998. ***
FAMILY Phasmatidae Gray, 1835.

SUBFAMILY Cladomorphinae Brunner von Wattenwyl, 1893.

TRIBE Cranidiini Günther, 1953.

Genus *Diapherodes* Gray, 1835.

*Diapherodes angulata* (Fabricius, 1793). Langlois and Lelong (1997); Lelong and Langlois (1998); Lelong and Langlois (2001); Hennemann et al. (2016).

FAMILY Prisopodidae Brunner von Wattenwyl, 1893.

SUBFAMILY Prisopodinae Brunner von Wattenwyl, 1893.

TRIBE Paraprisopodini Zompro, 2004.

Genus *Melopasma* Redtenbacher, 1906.

*Melopasma antillarum* (Caudell, 1914). Langlois and Lelong (1997); Langlois and Lelong (1998); Lelong and Langlois (2001).

FAMILY Pseudophasmatidae Bradley and Galil, 1977.

SUBFAMILY Xerosomatinae Zompro, 2004.

TRIBE Hesperophasmatini Bradley and Galil, 1977.

Genus *Hesperopasma* Rehn, 1901.

*Hesperopasma pavisae* Langlois and Lelong, 1998. Langlois and Lelong (1998). ***

Genus *Lamponius* Stål, 1875.

*Lamponius guerini* (Saussure, 1868). Langlois and Lelong (1997); Lelong and Langlois (1998); Lelong and Langlois (2001); Hennemann et al. (2016). ***

*Lamponius lethargicus* Lelong and Langlois, 1998. Lelong and Langlois (1998); Lelong P. and al. 2001. ***

ORDER Dermaptera de Geer, 1773 (14 SPECIES).

FAMILY Anisolabididae Verhoeff, 1902.

SUBFAMILY Anisolabidinae Verhoeff, 1902.

Genus *Anisolabis* Fieber, 1853.

*Anisolabis maritima* (Bonelli, 1832). Bonfils (1972).

Genus *Carcinophora* Scudder, 1876.

*Carcinophora percheroni* (Guérin and Percheron, 1838). Bonfils (1972); Meurgey (2011b).

Genus *Euborellia* Burr, 1909.

*Euborellia annulipes* (Lucas, 1847). Bonfils (1972); Meurgey (2011b). I

*Euborellia caraibea* Hebard, 1922. Bonfils (1972); Meurgey (2011b).

*Euborellia femoralis* (Dohrn, 1863). Bonfils (1972); Meurgey (2011b).

FAMILY Forficulidae Latreille, 1810.

SUBFAMILY Forficulinae Latreille, 1810.

Genus *Doru* Burr, 1907.

*Doru albipes* (Fabricius, 1787). Bonfils (1972); Meurgey (2011b).

FAMILY Labiduridae Zacher, 1911.

SUBFAMILY Labidurinae Zacher, 1911.

Genus *Labidura* Leach, 1815.

*Labidura riparia* (Pallas, 1773). Bonfils (1972); Meurgey (2011b).

*Labidura xanthopus* (Stål, 1855).
## FAMILY Spongiphoridae Verhoeff, 1902.

### SUBFAMILY Labiinae Burr, 1909.

Genus *Circolabia* Steinmann, 1987.

*Circolabia arcuata* (Scudder, 1876). Bonfils (1972); Meurgey (2011b).

Genus *Paralabella* Steinmann, 1990.

*Paralabella curvicauda* (Motschulsky, 1863). Bonfils (1972); Meurgey (2011b).

*Paralabella dorsalis* (Burmeister, 1838). Bonfils (1972); Meurgey (2011b).

### SUBFAMILY Spongiphorinae Verhoeff, 1902.

Genus *Marava* Burr, 1911.

*Marava arachidis* (Yersin, 1860). Bonfils (1972); Meurgey (2011b).

*Marava dominicae* (Rehn and Hebard, 1917). Bonfils (1972); Meurgey (2011b).

Genus *Spongiphora* Audinet-Serville, 1831.

*Spongiphora croceipennis* Audinet-Serville, 1831. Bonfils (1972); Meurgey (2011b).

Genus *Spongovostox* Burr, 1911.

*Spongovostox ghilianii* (Dohrn, 1864). Bonfils (1972); Meurgey (2011b).

## ORDER Mantodea Burmeister, 1838. (1 SPECIES).

### FAMILY Thespidae Giglio-Tos, 1919.

### SUBFAMILY Oligonicinae Giglio-Tos, 1919.

### TRIBE Oligonicini Giglio-Tos, 1919.

Genus *Oligonyx* Saussure, 1869.

*Oligonyx insularis* Bonfils, 1967. Bonfils (1967). **

## SUBCLASS Pterygota Lang, 1888.

## INFRACLASS Neoptera Martynov, 1923.

## ORDER Blattodea Brunner von Wattenwyl, 1882 (51 SPECIES).

### SUPERFAMILY Corydioidea Saussure, 1864.

### FAMILY Corydiidae Saussure, 1864.

### SUBFAMILY Holocompsinae Burmeister, 1838.

Genus *Holocompsa* Burmeister, 1838.

*Holocompsa nitidula* (Fabricius, 1781). Bonfils (1969).

### SUPERFAMILY Blaberoidea Saussure, 1864.

### FAMILY Ectobiidae Brunner von Wattenwyl, 1865.

### SUBFAMILY Blatellinae Karny, 1908.

Genus *Blattella* Caudell, 1903.

*Blattella germanica* (Linnaeus, 1767). Bonfils (1969).

Genus *Cahita* Hebard, 1923.

*Cahita palposa* Bonfils, 1969. ***

Genus *Ischnoptera* Burmeister, 1838.

*Ischnoptera rufa* (De Geer, 1773).

Genus *Symploce* Hebard, 1916.

*Symploce pallens* (Stephens, 1835). Bonfils (1969).

Genus *Euthlastoblatta* Hebard, 1917.

*Euthlastoblatta absimilis* (Gurney, 1937). Bonfils (1969).

*Euthlastoblatta absimilis montana* (Bonfils, 1969). ***
Genus *Nelipophygus* Rehn and Hebard, 1927.

*Nelipophygus cicatricosus* Bonfils, 1969. ***

SUBFAMILY Nyctiborinae Brunner von Wattenwyl, 1893.

Genus *Nyctibora* Burmeister, 1838.

*Nyctibora princisi* Bonfils, 1975. Bonfils (1975). ***

SUBFAMILY Pseudophyllodromiinae Hebard, 1929.

Genus *Cariblatta* Hebard, 1916.

*Cariblatta antiguensis* (Saussure and Zehntner, 1893). Bonfils (1969).

*Cariblatta guadeloupensis* Bonfils, 1969. Bonfils (1969). ***

*Cariblatta plagia* Rehn and Hebard, 1927. Bonfils (1969).

Genus *Chorisoneura* Brunner von Wattenwyl, 1865.

*Chorisoneura formosella* Rehn and Hebard, 1927. Bonfils (1969).

Genus *Neoblattella* Shelford, 1911.

*Neoblattella guadeloupensis* Bonfils, 1969. ***

*Neoblattella perdentata* Bonfils, 1969. ***

Genus *Plectoptera* Saussure, 1864.

*Plectoptera dominicae* Rehn and Hebard, 1927. Bonfils (1969).

*Plectoptera perscita* Rehn and Hebard, 1927. Bonfils (1969).

*Plectoptera rhabdota* Rehn and Hebard, 1927. Bonfils (1969).

Genus *Supella* Shelford, 1911.

*Supella longipalpa* (Fabricius, 1798). Bonfils (1969).

FAMILY Blaberidae Brunner von Wattenwyl, 1865.

SUBFAMILY Blaberinae Brunner von Wattenwyl, 1865.

Genus *Phoetalia* Stål, 1874.

*Phoetalia pallida* (Brunner von Wattenwyl, 1865). Bonfils (1969).

SUBFAMILY Epilamprinae Brunner von Wattenwyl, 1865.

Genus *Epilampra* Burmeister, 1838.

*Epilampra abdomennigrum* (De Geer, 1773). Bonfils (1969).

*Epilampra fugax* (Bonfils, 1969). ***

Genus *Hemiblabera* Saussure, 1893.

*Hemiblabera granulata* Saussure, 1893. Bonfils (1969); Gruner and Riom (1977).

SUBFAMILY Oxyhaloinae Roth, 1971.

Genus *Rhyparobia* Krauss, 1892.

*Rhyparobia maderae* (Fabricius, 1781). Bonfils (1969).

SUBFAMILY Panchlorinae Brunner von Wattenwyl, 1865.

Genus *Panchlora* Burmeister, 1838.

*Panchlora nivea* (Linnaeus, 1758). Bonfils (1969).

*Panchlora sagax* Rehn and Hebard, 1927. Bonfils (1969).

*Panchlora viridis* (Fabricius, 1775). Bonfils (1969).

Genus *Phoraspis* Audinet-Serville, 1831.

*Phoraspis atomaria* Blanchard, 1837. Bonfils (1969).
*Phoraspis pantherina* Blanchard, 1837. Bonfils (1969).

**SUBFAMILY Pycnoscelinae McKittrick, 1964.**

Genus *Pycnoscelus* Scudder, 1862.

*Pycnoscelus surinamensis* (Linnaeus, 1758). Rehn and Hebard (1927); Bonfils (1969); Gruner and Riom (1977).

**SUPERFAMILY Blattoidea Latreille, 1810.**

**EPIFAMILY Blattoidae Latreille, 1810.**

**FAMILY Blattidae Latreille, 1810.**

**SUBFAMILY Blattinae Latreille, 1810.**

Genus *Blatta* Linnaeus, 1758.

*Blatta orientalis* Linnaeus, 1758. Rehn and Hebard (1927); Bonfils (1969).

Genus *Henicotyle* Rehn and Hebard, 1927.

*Henicotyle antillarum* (Brunner von Wattenwyl, 1892). Bonfils (1969).

Genus *Pelmatosilpha* Dohrn, 1887.

*Pelmatosilpha purpurascens* Kirby, 1903. Bonfils (1969).

Genus *Periplaneta* Burmeister, 1838.

*Periplaneta americana* (Linnaeus, 1758). Bonfils (1969). I

*Periplaneta australasiae* (Fabricius, 1775). Bonfils (1969).

*Periplaneta brunnea* Burmeister, 1838. Bonfils (1969).

**EPIFAMILY Termitoidae Latreille, 1802.**

**FAMILY Kalotermitidae Froggatt, 1897.**

Genus *Comatermes* Krishna, 1961.

*Comatermes perfectus* (Hagen, 1858). Scheffrahn et al. (2017).

Genus *Cryptotermes* Banks, 1906.

*Cryptotermes brevis* (Walker, 1853). Darlington (1992); Scheffrahn et al. (1994); Aiguillon et al. (2017). I

*Cryptotermes darlingtonae* Scheffrahn and Krecek, 1999. Aiguillon et al. (2017).

*Cryptotermes havilandi* (Sjöstedt, 1900). Darlington (1992); Scheffrahn et al. (1994); Scheffrahn and Krecek (1999); Aiguillon et al. (2017).

Genus *Incisitermes* Krishna, 1961.

*Incisitermes incisus* (Silvestri, 1901). Darlington (1992); Scheffrahn et al. (1994); Aiguillon et al. (2017).

*Incisitermes tabogae* (Snyder, 1924). Darlington (1992); Scheffrahn et al. (1994); Contantino (1998); Aiguillon et al. (2017).

Genus *Procryptotermes* Holmgren, 1910.

*Procryptotermes corniceps* (Snyder, 1923). Darlington (1992); Scheffrahn et al. (1994); Contantino (1998).

*Procryptotermes leewardensis* Holmgren, 1910. Aiguillon et al. (2017).

**FAMILY Rhinotermitidae Froggatt, 1897.**

**SUBFAMILY Coptotermitinae Holmgren, 1910.**

Genus *Coptotermes* Wasmann, 1896.

*Coptotermes sjostedti* Holmgren, 1911. Scheffrahn et al. (2004); Aiguillon et al. (2017). I
SUBFAMILY Heterotermitinae Froggat, 1897.

Genus *Heterotermes* Froggatt, 1896.

*Heterotermes tenuis* (Hagen, 1858). Darlington (1992); Adamson (1948); Scheffrahn et al. (1994); Aiguillon et al. (2017).

SUBFAMILY Rhinotermitinae Froggat, 1897.

Genus *Rhinotermes* Hagen, 1858.

*Rhinotermes marginalis* (Linnaeus, 1758). Darlington (1992); Scheffrahn et al. (1994); Contantino (1998); Aiguillon et al. (2017).

FAMILY Termitidae Latreille, 1802.

SUBFAMILY Apicotermitinae Grassé and Noirot, 1954.

Genus *Anoplotermes* Müller, 1873.

*Anoplotermes meridianus* Emerson, 1925. Scheffrahn et al. (1994); Aiguillon et al. (2017).

Genus *Disjunctitermes* Scheffrahn, 2017.

*Disjunctitermes insularis* Scheffrahn, 2017. Scheffrahn *et al*. (2017).

SUBFAMILY Nasutitermitinae Hare, 1937.

Genus *Nasutitermes* Dudley, 1890.

*Nasutitermes corniger* (Motschulsky, 1855). Adamson (1948); Darlington (1992); Scheffrahn et al. (1994); Scheffrahn et al. (2005); Aiguillon et al. (2017).

*Nasutitermes ephratae* (Holmgren, 1910). Budesinsky et al. (2005); Darlington (1992); Aiguillon et al. (2017).

SUBFAMILY Termitinae Latreille, 1802.

Genus *Termes* Linnaeus, 1758.

*Termes hispaniolae* (Banks, 1918). Scheffrahn et al. (1994); Aiguillon et al. (2017).

ORDER Psocodea Hennig, 1966 = Psocoptera Shipley, 1904. (17 SPECIES).

SUBORDER Psocomorpha Roesler, 1944.

INFRAORDER Homilopsocidea Pearman, 1936.

FAMILY Ectopsocidae Roesler, 1944.

Genus *Ectopsocus* McLachlan, 1899.

*Ectopsocus maindroni* Badonnel, 1935.

*Ectopsocus thibaudi* Badonnel, 1979.

*Ectopsocus titschacki* Jentsch, 1939.

*Ectopsocus vilhenai* Badonnel, 1955.

FAMILY Lachesillidae Pearman, 1936.

Genus *Lachesilla* Westwood, 1840.

*Lachesilla caribe* García Aldrete, 1991. García Aldrete (1996).

FAMILY Peripsocidae Pearman, 1936.

Genus *Peripsocus* Hagen, 1866.

*Peripsocus potosi* Mockford, 1971.
INFRAORDER Psocetae Pearman, 1936.

FAMILY Hemipsocidae Pearman, 1936.

Genus *Hemipsocus* Selys-Longchamps, 1872.

*Hemipsocus roseus* (Hagen, 1859). Badonnel (1981).

FAMILY Psocidae Samouelle, 1819.

Genus *Indiopsocus* Mockford, 1974.

*Indiopsocus caraibensis acutus* Badonnel, 1989.

*Indiopsocus caraibensis caraibensis* Badonnel, 1989.

*Indiopsocus etiennei* Badonnel, 1989.

*Indiopsocus obrieni* Badonnel, 1989.

SUBORDER Troctomorpha Roesler, 1940.

INFRAORDER Nanopsocetae Pearman, 1936.

FAMILY Liposcelididae Broadhead, 1950.

Genus *Liposcelis* Motschulsky, 1852.

*Liposcelis entomophila* (Enderlein, 1907). Badonnel (1981).

FAMILY Pachytroctidae Enderlein, 1904.

Genus *Tapinella* Enderlein, 1908.

*Tapinella dichromoptera* Badonnel, 1988.

*Tapinella olmeca* Mockford, 1975. Badonnel (1981).

SUBORDER Trogiomorpha Roesler, 1940.

INFRAORDER Atropetae Pearman, 1936.

FAMILY Lepidopsocidae Pearman, 1936.

Genus *Echmepteryx* Aaron, 1886.

*Echmepteryx lunulata* Thornton, Lee and Chui, 1972. Badonnel (1981).

Genus *Neolepolepis* Mockford, 1993.

*Neolepolepis caribensis* (Turner, 1975). Badonnel (1981).

INFRAORDER Psocthropetae Pearman, 1936.

FAMILY Psyllipsocidae Lienhard and Smithers, 2002.

Genus *Psocathropos* Ribaga, 1899.

*Psocathropos lachlani* Ribaga, 1899. Badonnel (1981).

ORDER Thysanoptera Haliday, 1836. (76 SPECIES).

SUBORDER Terebrantia Haliday, 1836.

FAMILY Aeolothripidae Uzel, 1895.

Genus *Franklinothrips* Back, 1912.

*Franklinothrips vespiformis* (D. L. Crawford, 1909). Mound and Reynaud (2005), Etienne et al. (2015).

FAMILY Heterothripidae Bagnall, 1912.

Genus *Heterothrips* Hood, 1908.

*Heterothrips decacornis* Crawford, 1909. Etienne et al. (2015).
## FAMILY Thripidae Stephens, 1829.

Genus *Elixothrips* Stannard and Mitri, 1962

*Elixothrips brevisetis* (Bagnall, 1919). Etienne et al. (2015).

## SUBFAMILY Panchaetothripinae Bagnall, 1912.

Genus *Anisopilothrips* Stannard and Mitri 1962.

*Anisopilothrips venustulus* (Priesner, 1923). Etienne et al. (2015).

Genus *Arachisothrips* Stannard, 1952.

*Arachisothrips millsi* Stannard, 1952. Etienne et al. (2015).

Genus *Dinurothrips* Hood, 1913.

*Dinurothrips hookeri* Hood, 1913. Etienne et al. (2015).

Genus *Heliothrips* Haliday, 1836.

*Heliothrips haemorrhoidalis* (Bouché, 1833). Etienne et al. (2015).

Genus *Hercinothrips* Bagnall 1932.

*Hercinothrips femoralis* (Reuter, 1891). Etienne et al. (2015).

Genus *Parthenothrips* Uzel 1895.

*Parthenothrips dracaenae* (Heeger, 1854). Etienne et al. (2015).

Genus *Retithrips* Marchal, 1910.

*Retithrips syriacus* (Mayet, 1890). Etienne et al. (2015).

Genus *Selenothrips* Karny, 1911.

*Selenothrips rubrocinatus* (Giard, 1901). Denmark and Wolfenbarger (2010), Etienne et al. (2015).

Genus *Sericothrips* Haliday, 1836.

*Sericothrips tricinctus* Hood, 1927. Etienne et al. (2015).

## SUBFAMILY Thripinae Stephens, 1829.

Genus *Arorathrips* (Bhatti, 1990).

*Arorathrips spinipes* (Hood, 1915). Etienne et al. (2015).

Genus *Aurantothrips* Bhatti, 1978.

*Aurantothrips orchidaceus* (Bagnall, 1909). Etienne et al. (2015).

Genus *Ceratothripoides* Bagnall, 1918.

*Ceratothripoides brunneus* Bagnall, 1918. Etienne et al. (2015).

Genus *Chaetanaphothrips* Priesner, 1926.

*Chaetanaphothrips leeuweni* (Karny, 1914). Etienne et al. (2015).

*Chaetanaphothrips orchidii* (Moulton, 1907). Etienne et al. (2015).

*Chaetanaphothrips signipennis* (Bagnall, 1914). Etienne et al. (2015).

Genus *Coremothrips* Hood, 1925.

*Coremothrips pallidus* Hood, 1925. Etienne et al. (2015).

Genus *Corynothrips* Williams, 1913.

*Corynothrips stenopterus* Williams, 1913. Etienne et al. (2015).

Genus *Cricothrips* Trybom, 1913.

*Cricothrips trinidadensis* (Hood, 1956). Etienne et al. (2015).
Genus *Dendrothripoides* Bagnall, 1923.

*Dendrothripoides innoxius* (Karny, 1914). Etienne et al. (2015).

Genus *Echinothrips* Moulton, 1911.

*Echinothrips americanus* Morgan, 1913. Etienne et al. (2015).

*Echinopthrips caribbeanus* Hood, 1955. Etienne et al. (2015).

Genus *Frankliniella* Karny, 1910.

*Frankliniella borinquen* Hood, 1942. Etienne et al. (2015).

*Frankliniella breviseta* Moulton, 1948. Etienne et al. (2015).

*Frankliniella bruneri* Watson, 1925. Etienne et al. (2015).

*Frankliniella cephalica* (Crawford, 1910). Etienne et al. (2015).

*Frankliniella insularis* (Franklin, 1908). Etienne et al. (2015).

*Frankliniella kelliae* Sakimura, 1981. Etienne et al. (2015).

*Frankliniella occidentalis* (Pergande, 1895). Etienne et al. (2015).

*Frankliniella parvula* Hood, 1925. Etienne (2015). Etienne et al. (2015).

*Frankliniella rostrata* Priesner, 1932. Etienne (2015). Etienne et al. (2015).

*Frankliniella williamsi* Hood, 1915. Wang et al. (2010); Etienne et al. (2015).

Genus *Fulmekiola* Karny, 1925.

*Fulmekiola serrata* (Kobus, 1893). Etienne et al. (2015).

Genus *Pseudothrips* Hinds, 1902.

*Pseudothrips inequalis* (Beach, 1896). Etienne et al. (2015).

Genus *Rhamphothrips* Karny, 1913.

*Rhamphothrips pandens* Sakimura, 1983. Etienne et al. (2015).

Genus *Scirtothrips* Shull, 1909.

*Scirtothrips dorsalis* Hood, 1919. Etienne et al. (2015).

Genus *Trichromothrips* Priesner, 1930.

*Trichromothrips xanthius* (Williams, 1917). Etienne et al. (2015).

Genus *Thrips* Linnaeus, 1758.

*Thrips palmi* Karny, 1925. Etienne et al. (2015).

*Thrips simplex* (Morison, 1930). Etienne et al. (2015).

*Thrips tabaci* Lindeman, 1889. Etienne et al. (2015).

## SUBFAMILY Dendrothripinae Priesner, 1925

Genus *Leucothrips* Reuter, 1904.

*Leucothrips furcatus* Hood, 1831. Etienne et al. (2015).

## SUBFAMILY Sericothripinae Karny, 1921.

Genus *Hydatothrips* Karny, 1913.

*Hydatothrips tricinctus* (Hood, 1928). Etienne et al. (2015).

Genus *Neohydatothrips* John, 1929.

*Neohydatothrips portoricensis* (Morgan, 1925). Etienne et al. (2015).

*Neohydatothrips signifer* (Priesner, 1932). Etienne et al. (2015).
SUBORDER Tubulifera Haliday, 1836.

FAMILY Phlaeothripidae Uzel, 1895.

SUBFAMILY Idolothripinae Bagnall, 1908.

Genus *Bactrothrips* Karny, 1912.

*Bactrothrips* sp. nr. *hesperus* (Moulton, 1907). Etienne et al. (2015).

Genus *Diceratothrips* Bagnall, 1908.

*Diceratothrips bicornis* Bagnall, 1908. Etienne et al. (2015).

Genus *Elaphothrips* Girault, 1927.

*Elaphrothrips jacotguillarmodi* Johansen, 1979. Etienne et al. (2015).

Genus *Ethirothrips* Karny, 1925.

*Ethirothrips* sp. Etienne et al. (2015).

Genus *Gastrothrips* Hood, 1912.

*Gastrothrips abditus* Hood, 1935. Etienne et al. (2015).

*Gastrothrips anolis* Morgan, 1925. Etienne et al. (2015).

Genus *Nesothrips* Kirkaldy, 1907.

*Nesothrips lativentris* (Karny, 1913). Etienne et al. (2015).

*Nesothrips minor* (Bagnall, 1921). Etienne et al. (2015).

Genus *Ophthalmothrips* Hood, 1919.

*Ophthalmothrips longisetis* (Bournier, 1968). Etienne et al. (2015).

Genus *Adraneothrips* Hood, 1925.

*Adraneothrips alternatus* Hood, 1925. Etienne et al. (2015).

Genus *Aleurodothrips* Franklin, 1909.

*Aleurodothrips fasciapennis* (Franklin, 1908). Etienne et al. (2015).

Genus *Bamboosiella* Ananthakrishnan, 1957.

*Bamboosiella cingulata* (Hood, 1919). Etienne et al. (2015).

Genus *Docessissophothrips* Uzel, 1895.

*Docessissophothrips bonfilsi* Bournier, 1992. Etienne et al. (2015).

Genus *Dolichothrips* Karny, 1912.

*Dolichothrips indicus* (Hood, 1919). Etienne et al. (2015).

Genus *Gynaikothrips* Zimmermann, 1900.

*Gynaikothrips ficorum* (Marchal, 1908). Etienne et al. (2015).

*Gynaikothrips uzeli* (Zimmermann, 1900). Etienne et al. (2015).

Genus *Haplothrips* Amyot and Serville 1843.

*Haplothrips gowdeyi* (Franklin, 1908). Etienne et al. (2015).

Genus *Holopothrips* Hood, 1914.

*Holopothrips ananasi* Costa Lima, 1935. Etienne et al. (2015).

*Holopothrips inquilinus* (Bournier, 1993). Etienne et al. (2015).

*Holopothrips tabebuia* Cabrera and Segarra, 2008. Michel et al. (2008); Etienne et al. (2015).

*Holopothrips tenuis* Hood, 1914. Etienne et al. (2015).

Genus *Hoplothrips* Amyot and Audinet-Serville, 1843.

*Hoplothrips polypori* Bournier, 1992. Etienne et al. (2015).
Genus *Karnyothrips* Watson, 1923.

*Karnyothrips flavipes* (Jones, 1912). Etienne et al. (2015).

*Karnyothrips merrilli* (Watson, 1920). Etienne et al. (2015).

*Karnyothrips rhopalocerus* (Hood, 1925). Etienne et al. (2015).

Genus *Leptothrips* Hood, 1909.

*Leptothrips* sp. aff. *obesus* Johansen, 1987. Etienne et al. (2015).

Genus *Liothrips* Uzel, 1895.

*Liothrips brevicornis* Hood, 1913. Etienne et al. (2015).

Genus *Streptothrips* Priesner, 1932.

*Streptothrips rostratus* Bournier, 1992. Etienne et al. (2015).

Genus *Symphyothrips* Hood and Williams, 1915.

*Symphyothrips punctatus* Hood and Williams, 1915. Etienne et al. (2015).

Genus *Williamsiella* Hood, 1925.

*Williamsiella* sp. Etienne et al. (2015).

## ORDER Hemiptera Linnaeus, 1758 (385 SPECIES).

### SUBORDER Cicadomorpha Evans, 1946.

### SUPERFAMILY Cercopoidea Leach, 1815.

### FAMILY Aphrophoridae Amyot and Audinet-Serville, 1843.

Genus *Philaenus* Stål, 1864.

*Philaenus spumarius* (Linnaeus, 1758). Lethierry (1881).

### SUPERFAMILY Cicadoidea Westwood, 1840.

### FAMILY Cicadidae Latreille, 1802.

### SUBFAMILY Cicadinae Latreille, 1802.

### TRIBE FIDICININI Distant, 1905.

Genus *Fidicina* Amyot and Audinet-Serville, 1843.

*Fidicina mannifera* (Fabricius, 1803). I

Genus *Tympanoterpes* Stål, 1862.

*Tympanoterpes grisea* (Germar, 1830). Lethierry (1881); Meurgey (2011b).

### SUPERFAMILY Membracoidea Rafinesque, 1815.

### FAMILY Cicadellidae Latreille, 1802.

### SUBFAMILY Aphrodinae Haupt, 1927.

### TRIBE Xestocephalini Baker, 1915.

Genus *Xestocephalus* Van Duzee, 1892.

*Xestocephalus pulicarius* Van Duzee, 1894. Bonfils and Delplanque (1971); Meurgey (2011b).

### SUBFAMILY Cicadellinae Van Duzee, 1916.

### TRIBE Cicadellini Van Duzee, 1916.

Genus *Carneocephala* Ball, 1927.

*Carneocephala sagittifera* (Uhler, 1895). Bonfils and Delplanque (1971); Meurgey (2011b).

Genus *Cicadella* Latreille, 1817.

*Cicadella prolixa* (Lethierry, 1881). Bonfils and Delplanque (1971); Meurgey (2011b).

Genus *Draeculacephala* Ball 1902.

*Draeculacephala clypeata* Osborn, 1926. Dumbardon-Martial and Pierre (2016).
Genus *Hortensia* Metcalf and Bruner, 1936.

*Hortensia similis* (Walker, 1851). Bonfils and Delplanque (1971); Meurgey (2011b); Dumbardon-Martial and Pierre (2016).

Genus *Poeciloscarta* Stål, 1869.

*Poeciloscarta guadulpensis* (Lethierry, 1881). Lethierry (1881); Dumbardon-Martial and Pierre (2016).***

Genus *Sibovia* Metcalf, 1952.

*Sibovia nielsoni* Young, 1977. Dumbardon-Martial and Pierre (2016).

Genus *Stirellus* Osborn and Ball, 1902.

*Stirellus bicolor* (Van Duzee, 1892). Dumbardon-Martial and Pierre (2016).

Genus *Tylozygus* Fieber, 1866.

*Tylozygus fasciatus* (Walker, 1851). Bonfils and Delplanque (1971); Meurgey (2011b).

*Tylozygus geometricus* (Signoret, 1854). Dumbardon-Martial and Pierre (2016).

SUBFAMILY Coelidiinae Dohrn, 1859.

TRIBE Coelidiini Dohrn, 1859.

Genus *Coelidia* Germar, 1821.

*Coelidia semiflava* Lethierry, 1881. Lethierry (1881).

SUBFAMILY Deltcephalinae Dallas, 1870.

TRIBE Acinopterini Oman, 1943.

Genus *Acinopterus* van Duzee, 1892.

*Acinopterus angulatus* Lawson, 1922. Dumbardon-Martial and Pierre (2016).

TRIBE Chiasmini Distant, 1908.

Genus *Exitianus* Ball, 1929.

*Exitianus exitiosus* (Uhler, 1880). Bonfils and Delplanque (1971); Meurgey (2011b).

TRIBE Deltcephalini Dallas, 1870.

Genus *Deltcephalus* Burmeister, 1838.

*Deltcephalus flavicosta* (Stål, 1862). Bonfils and Delplanque (1971); Meurgey (2011b).

*Deltcephalus maculellus* (Osborn, 2010). Dumbardon-Martial and Pierre (2016).

Genus *Graminella* DeLong, 1936.

*Graminella cognita* Caldwell, 1952. Bonfils and Delplanque (1971); Meurgey (2011b).

*Graminella colona* (Uhler, 1895). Bonfils and Delplanque (1971); Meurgey (2011b).

*Graminella nigrifrons* (Forbes, 1885). Bonfils and Delplanque (1971); Meurgey (2011b).

*Graminella punctata* Caldwell, 1952. Bonfils and Delplanque (1971); Meurgey (2011b).

*Graminella sonora* (Ball, 1900). Dumbardon-Martial and Pierre (2016).

Genus *Sanctanus* Ball, 1932.

*Sanctanus* (*Cruciatanus) fasciatus (Osborn, 1900). Bonfils and Delplanque (1971); Meurgey (2011b).

TRIBE Hecalini Distant, 1908.

Genus *Spangbergiella* Signoret, 1879.

*Spangbergiella vulnerata* (Uhler, 1877). Bonfils and Delplanque (1971); Meurgey (2011b).
TRIBE Macrostelini Kirkaldy, 1906.

Genus *Balclutha* Kirkaldy, 1900.

*Balclutha abdominalis* (van Duzee, 1892). Dumbardon-Martial and Pierre (2016).

*Balclutha flavescens* (Baker, 1903). Bonfils and Delplanque (1971); Meurgey (2011b) [as junior synonym *Nesosteles virescens* (Osborn, 1926)].

*Balclutha frontalis* (Ferrari, 1882). Bonfils and Delplanque (1971); Meurgey (2011b) [as junior synonym *Nesosteles guajanae* (Delong, 1923)].

*Balclutha incisa* (Matsumura, 1902). Dumbardon-Martial and Pierre (2016).

Genus *Cicadulina* China, 1926.

*Cicadulina tortilla* Caldwell, 1952. Meurgey (2011b).

Genus *Macrosteles* Fieber. 1866.

*Macrosteles fascifrons* (Stål, 1858). Bonfils and Delplanque (1971); Meurgey (2011b).

TRIBE Paralimnini Distant, 1908.

Genus *Laevicephalus* DeLong, 1926.

*Laevicephalus nigripennis* (DeLong, 1923). Bonfils and Delplanque (1971); Meurgey (2011b).

TRIBE Pendarini Dmitriev, 2009.

Genus *Chlorotettix* Van Duzee 1892.

*Chlorotettix minimus* Baker, 1898. Bonfils and Delplanque (1971); Meurgey (2011b).

Genus *Ileopeltus* Cwikla, 1988.

*Ileopeltus tethys* (Van Duzee, 1907). Dumbardon-Martial and Pierre (2016).

TRIBE Stenometopiini Baker, 1923.

Genus *Stirellus* Osborn and Ball, 1902.

*Stirellus bicolor* (Van Duzee, 1892). Bonfils and Delplanque (1971); Meurgey (2011b).

SUBFAMILY Gyponinae Stål, 1870.

Genus *Gypona* Germar, 1821.

*Gypona (Marganalana) arunda* (DeLong and Freytag, 1964). Dumbardon-Martial and Pierre (2016).

Genus *Polana* Delong, 1942.

*Polana (Bulbusana) insulana* Freytag and Cwikla, 1982. Dumbardon-Martial and Pierre (2016). **

SUBFAMILY Ledrinae Kirschbaum, 1868.

Genus *Xerophloea* Germar, 1839.

*Xerophloea viridis* (Fabricius, 1794). Bonfils and Delplanque (1971); Meurgey (2011b); Dumbardon-Martial (2016).

SUBFAMILY Megophthalminae Kirkaldy, 1906.

TRIBE Agalliini Kirkaldy, 1901.

Genus *Agallia* Curtis, 1833.

*Agallia albidula* Uhler, 1895. Bonfils and Delplanque (1971); Meurgey (2011b).

SUBFAMILY Typhlocybinae Kirschbaum, 1868.

TRIBE Alebrini McAtee, 1926.

Genus *Protalebrella* Young, 1952.

*Protalebrella brasiliensis* (Baker, 1899). Bonfils and Delplanque (1971); Meurgey (2011b).
**TRIBE Dikraneurini McAtee, 1926.**

Genus *Typhlocybella* Baker, 1903.
*Typhlocybella minima* Baker, 1903. Bonfils and Delplanque (1971); Meurgey (2011b).

**TRIBE Empoascini Distant, 1908.**

Genus *Empoasca* Walsh, 1862.
*Empoasca fabae* (Harris, 1841). Bonfils and Delplanque (1971); Meurgey (2011b).

**FAMILY Membracidae Rafinesque, 1815.**

Genus *Antillotolania* Ramos, 1957.
*Antillotolania microcentroides* Cryan and Bartlett, 2002. *

Genus *Iria* Stål, 1867.
*Iria lethierryi* Funkhouser, 1927. ***

Genus *Monobelus* Stål, 1866.
*Monobelus nasutus* Stål, 1869. ***

Genus *Togotolania* Cryan and Deitz, 2002.
*Togotolania brachycorna* Cryan and Deitz, 2002.

**SUBORDER Fulgoromorpha Evans, 1946.**

**FAMILY Acanaloniidae Amyot and Audinet-Serville, 1843.**

Genus *Acanalonia* Spinola, 1839.
*Acanalonia viriditerminata* (Lethierry, 1881). Lethierry (1881). **

**FAMILY Cixiidae Spinola, 1839.**

Genus *Bothriocera* Burmeister, 1835.
*Bothriocera eborea* Fennah, 1943. Bonfils and Delplanque (1971); Meurgey (2011b).
*Bothriocera substigmatica* (Lethierry, 1881) ***. Lethierry (1881); Fennah (1945); Meurgey (2011b).

Genus *Pintalia* Stål, 1862.
*Pintalia pictipennis* Stål, 1862. Lethierry (1881); Meurgey (2011b).

**FAMILY Delphacidae Leach, 1815.**

Genus *Caenodelphax* Fennah, 1965.
*Caenodelphax teapae* (Fowler, 1905). Bonfils and Delplanque (1971); Meurgey (2011b); Dumbardon-Martial (2016).

Genus *Delphacodes* Fieber, 1866.
*Delphacodes nigrifacies* (Muir, 1918). Bonfils and Delplanque (1971); Meurgey (2011b).

Genus *Euides* Fieber, 1866.
*Euides weedi* (Van Duzee, 1897). Bonfils and Delplanque (1971); Meurgey (2011b).

Genus *Metadelphax* Wagner, 1963.
*Metadelphax propinqua* (Fieber, 1866). Bonfils and Delplanque (1971); Meurgey (2011b).

Genus *Nilaparvata* Distant, 1906.
*Nilaparvata muiri* China, 1925. Bonfils and Delplanque (1971); Meurgey (2011b).

Genus *Peregrinus* Kirkaldy, 1904.
*Peregrinus maidis* (Ashmead, 1890). Bonfils and Delplanque (1971).
Genus *Saccharosydne* Kirkaldy, 1907.

*Saccharosydne saccharivora* (Westwood, 1833). Bonfils and Delplanque (1971); Meurgey (2011b).

Genus *Sogatella* Fennah, 1956.

*Sogatella kolophon* (Kirkaldy, 1907). Bonfils et al. (1994); Vayssières et al. (2001); Meurgey (2011b).

## FAMILY Flatidae Spinola, 1839.

Genus *Anormenis* Melichar, 1923.

*Anormenis nivea* (Lethierry, 1881). Lethierry (1881).

Genus *Antillormenis* Fennah, 1942.

*Antillormenis albicostalis* (Fennah, 1941). Bonfils and Delplanque (1971); Meurgey (2011b).

Genus *Hansenia* Melichar, 1901.

*Hansenia nigra* (Lethierry, 1881). Lethierry (1881).

## FAMILY Tropiduchidae Stål, 1866.

Genus *Dioxyomus* Fennah, 1945.

*Dioxyomus longiceps* (Lethierry, 1881). Lethierry (1881); Meurgey (2011b).

Genus *Tangidia* Uhler, 1895.

*Tangidia triangulator* (Lethierry, 1881). Lethierry (1881); Meurgey (2011b).

## SUBORDER Heteroptera Latreille, 1810.

## INFRAORDER Cimicomorpha Leston, Pendergrast and Southwood, 1954.

## SUPERFAMILY Cimicoidea Latreille, 1802.

## FAMILY Anthocoridae Fieber, 1836.

## SUBFAMILY Lasiochilinae Crayon, 1972.

Genus *Lasiochilus* Reuter, 1871.

*Lasiochilus microps* Champion, 1900. Ford (1979); Carpintero (2002); Meurgey (2011b).

*Lasiochilus pallidulus* Reuter, 1871. Lethierry (1881); Ford (1979); Carpintero (2002); Meurgey (2011b).

*Lasiochilus varicolor* Uhler, 1894. Ford (1979); Carpintero (2002); Meurgey (2011b).

## SUBFAMILY Lyctocorinae Reuter, 1884.

## TRIBE Dufouriellini Van Duzee, 1916

Genus *Alofa* Herring, 1976.

*Alofa sodalis* (Buchanan-White, 1879). Carpintero (2002).

Genus *Buchananiella* Poppius, 1909.

*Buchananiella bicolor* Poppius, 1909. Ford (1979); Meurgey (2011b).

Genus *Cardiastethus* Fieber, 1860.

*Cardiastethus flavus* Poppius, 1909. Carpintero (2002); Meurgey (2011b).

*Cardiastethus assimilis* (Reuter, 1871). Ford (1979); Meurgey (2011b).

## TRIBE Scolopini Carayon, 1954.

Genus *Calliodis* Reuter, 1871.

*Calliodis maculipennis* (Reuter, 1884). Ford (1979); Carpintero (2002); Meurgey (2011b); Carpintero (2012).

*Calliodis signatus* (Poppius, 1909). Carpintero (2002); Meurgey (2011b); Carpintero (2012).
Genus *Montandoniola* Poppius, 1909.

*Montandoniola confusa* Streito and Matocq, 2009. Pluot-Sigwalt et al. (2009); Meurgey (2011b).

*Montandoniola moraguesi* (Puton, 1896). Pluot-Sigwalt et al. (2009); Meurgey (2011b).

TRIBE Xylocorini Carayon, 1972.

Genus *Xylocoris* Dufour, 1831.

*Xylocoris sordidus* (Reuter, 1871). Carpintero (2002).

FAMILY Nabidae Costa, 1853.

SUBFAMILY Nabinae Costa, 1853.

TRIBE Arachnocorini Scott, 1881.

Genus *Arachnocoris* Scott, 1881.

*Arachnocoris karukerae* Lopez-Moncet, 1990.

SUPERFAMILY Miroidea Hahn, 1833.

FAMILY Miridae Hahn, 1831.

SUBFAMILY Bryocorinae Bärensprung, 1860.

TRIBE Bryocorini Bärensprung, 1860.

Genus *Monalocoris* Dahlbom, 1851.

*Monalocoris nigrocollaris* Carvalho, 1989.

TRIBE Eccritotarsini Carvalho, 1957.

Genus *Neofurius* Distant, 1884.

*Neofurius albiceps* (Lethierry, 1881). Meurgey (2011b). ***

SUBFAMILY Cylapinae Kirkaldy, 1903.

TRIBE Fulviini Uhler, 1886.

Genus *Fulvius* Stål, 1862.

*Fulvius anthocoroides* (Reuter, 1875). Carvalho and Costa (1994); Meurgey (2011b).

*Fulvius submaculatus* Poppius, 1909. Pluot-Sigwalt and Chérot (2013).

SUBFAMILY Deraeocorinae Douglas and Scott, 1865.

TRIBE Clivinematini Reuter, 1871.

Genus *Hemicerocoris* Lethierry, 1881.

*Hemicerocoris nigritarsis* Lethierry, 1881. Lethierry (1881); Meurgey (2011b). ***

TRIBE Hyalodini Carvalho and Drake, 1943.

Genus *Hyaliodes* Reuter, 1876.

*Hyaliodes guadalupensis* Carvalho, 1985.

SUBFAMILY Mirinae Hahn, 1831.

TRIBE Mirini Hahn, 1831.

Genus *Derophthalma* Berg, 1883.

*Derophthalma irrorata* (Lethierry, 1881). Lethierry (1881); Meurgey (2011b).

Genus *Taylorilygus* Leston 1952.

*Taylorilygus apicalis* (Fieber, 1861). Lethierry (1881); Meurgey (2011b).
SUPERFAMILY Reduvioidea Latreille, 1807.

FAMILY Reduviidae Latreille, 1807.

SUBFAMILY Emesinae Amyot and Audinet-Serville, 1843.

TRIBE Leistarchini Stål, 1862.

Genus *Empicoris* Wolff, 1811.

*Empicoris armatus* (Champion, 1898). Meurgey (2011b).

Genus *Ghinallelia* Wygodzinsky 1966.

*Ghinallelia carayoni* Villiers, 1978. Villiers (1978); Meurgey (2011b).

*Ghinallelia insularis* Villiers, 1978. Villiers (1978); Meurgey (2011b).

Genus *Ploiaria* Scopoli, 1786.

*Ploiaria guadeloupensis* Villiers, 1978. Villiers (1978); Meurgey (2011b).

*Ploiaria matilei* Dispons and Villiers, 1967. Villiers (1978); Meurgey (2011b).

SUBFAMILY Harpactorinae Amyot and Audinet-Serville 1843.

TRIBE Harpactocorini Amyot and Audinet-Serville 1843.

Genus *Lophoscutus* Kormilev, 1951.

*Lophoscutus dudichi* (Kormilev, 1949). Kormilev and Doesburg (1991); Meurgey (2011b).

*Lophoscutus insularis* (Dudich, 1922). Kormilev and Van Doesburg (1989); Kormilev and Doesburg (1991); Meurgey (2011b).

Genus *Zelus* Fabricius, 1803.

*Zelus impar* Kuhlgatz, 1902. Hart (1987).

*Zelus longipes* (Linnaeus, 1767). Simmonds (1956); Gruner and Riom (1977); Villiers (1978); Meurgey (2011b).

*Zelus rubidus* (Lepeletier de Saint Fargeau and Audinet-Serville, 1825). Lethierry (1881); Meurgey (2011b).

SUBFAMILY Peiratinae Stål, 1859.

Genus *Sirthenea* Spinola, 1840.

*Sirthenea stria pedestris* Horvath, 1909. Villiers (1978); Meurgey (2011b).

*Sirthenea stria stria* (Fabricius, 1794). Willemse (1985); Meurgey (2011b).

SUBFAMILY Stenopodainae Amyot and Audinet-Serville, 1843.

Genus *Oncocephalus* Klug, 1830.

*Oncocephalus validispinis* Reuter, 1882. Villiers (1978); Meurgey (2011b).

SUPERFAMILY Tingoidea Laporte de Castelnau, 1833.

FAMILY Tingidae Laporte de Castelnau, 1833.

Genus *Corythaica* Stål, 1873.

*Corythaica planaris* Uhler, 1893. Gruner and Riom (1977); Meurgey (2011b).

Genus *Pseudacysta* Blatchley, 1926.

*Pseudacysta perseae* (Heidemann, 1908). Etienne and Streito (2008); Meurgey (2011b).

INFRAORDER Enicocephalomorpha Stichel, 1955.

SUPERFAMILY Enicocephaloidea Stål, 1860.

FAMILY Enicocephalidae Stål, 1860.

Genus *Henicocephalus* Westwood, 1837.

*Henicocephalus flavicollis* Westwood, 1837. Villiers (1978); Meurgey (2011b).

*Henicocephalus wygodzinskyi* Villiers, 1978. Villiers (1978); Meurgey (2011b).
Genus *Lomagostus* Villiers, 1958.

*Lomagostus aguilari* Villiers, 1978. Villiers (1978); Meurgey (2011b).

INFRAORDER Gerromorpha Popov, 1971.

SUPERFAMILY Gerroidea Leach, 1815.

FAMILY Gerridae Leach, 1815.

SUBFAMILY Gerrinae Leach, 1815.

Genus *Limnogonus* Stål, 1868.

*Limnogonus franciscanus* (Stål, 1859). de Kort-Gommers and Nieser (1969).

SUBFAMILY Rhagadotarsinae Lundblad, 1933.

Genus *Rheumatobates* Bergroth, 1892.

*Rheumatobates imitator* (Uhler, 1894). de Kort-Gommers and Nieser (1969).

FAMILY Veliidae Brullé, 1836.

SUBFAMILY Microveliinae China and Usinger, 1949.

Genus *Microvelia* Westwood, 1834.

*Microvelia pulchella* Westwood, 1834. Cordeiro and Moreira (2015).

SUPERFAMILY Mesovelioidea Douglas and Scott, 1867.

FAMILY Mesoveliidae Douglas and Scott, 1867.

SUBFAMILY Mesoveliinae Douglas and Scott, 1867.

Genus *Mesovelia* Mulsant and Rey, 1852.

*Mesovelia mulsanti* White, 1879. Horvath (1929); Moreira et al. (2008); Meurgey (2011b).

INFRAORDER Nepomorpha Popov, 1968.

SUPERFAMILY Nepoidea Latreille, 1802.

FAMILY Belostomatidae Leach, 1815.

SUBFAMILY Belostomatinae Leach, 1815.

Genus *Belostoma* Latreille, 1807.

*Belostoma boscii* Lepeletier de Saint Fargeau and Audinet-Serville, 1825. Gruner and Riom (1977); Meurgey (2011b).

*Belostoma subspinosus* Amyot and Audinet-Serville, 1843. Gruner and Riom (1977); Meurgey (2011b).

Genus *Zaitha* Amyot and Audinet-Serville, 1843.

*Zaitha margineguttata* Dufour, 1863. Lethierry (1881); Meurgey (2011b).

FAMILY Nepidae Latreille, 1802.

SUBFAMILY Ranatrinae Douglas and Scott, 1865.

TRIBE Ranatrini Douglas and Scott, 1865.

Genus *Ranatra* Fabricius, 1790.

*Ranatra galantae* Gruner and Riom, 1977. Meurgey (2011b).

SUPERFAMILY Notonectoidea Latreille, 1802.

FAMILY Notonectidae Latreille, 1802.

SUBFAMILY Anisopinae Hutchinson, 1929.

Genus *Buenoa* Kirkaldy, 1904.

*Buenoa antigone* (Kirkaldy, 1899). Gruner and Riom (1977); Meurgey (2011b).
SUBFAMILY Notonectinae Latreille, 1802.

Genus *Notonecta* Linnaeus, 1758.

*Notonecta indica* Linnaeus, 1771. Gruner and Riom (1977); Meurgey (2011b).

SUPERFAMILY Ochteroidea Kirkaldy, 1906.

FAMILY Gelastocoridae Kirkaldy, 1897.

SUBFAMILY Gelastocorinae Kirkaldy, 1897.

Genus *Lethocerus* Mayr, 1853.

*Lethocerus annulipes* (Herrich-Schaffer, 1845). Gruner and Riom (1977); Meurgey (2011b).

Genus *Pelocoris* Stål, 1876.

*Pelocoris convexus* Nieser, 1969. Gruner and Riom (1977); Meurgey (2011b).

INFRAORDER Pentatomorpha Leston, Pendergrast and Southwood, 1954.

SUPERFAMILY Aradoidea Brullé, 1836.

FAMILY Aradidae Spinola, 1837.

SUBFAMILY Aneurinae Douglas and Scott 1865.

Genus *Aneurosoma* Champion, 1898.

*Aneurosoma dissimile* (Bergroth, 1889). Kormilev and Froeschner (1987); Panizzi and Grazia (2015).

Genus *Aneurus* Curtis, 1825.

*Aneurus nasutus* Kormilev, 1966. Kormilev and Froeschner (1987); Panizzi and Grazia (2015).***.

*Aneurus vauriei* Kormilev, 1964. Kormilev and Froeschner (1987); Panizzi and Grazia (2015).

Genus *Iralunelus* Štys, 1974.

*Iralunelus politus* (Say, 1832). Lethierry (1881); Kormilev and Froeschner (1987); Meurgey (2011b); Panizzi and Grazia (2015).

SUBFAMILY Calisiinae Stål, 1873.

Genus *Calisius* Stål, 1860.

*Calisius contubernalis* Bergroth, 1913. Kormilev and Froeschner (1987); Panizzi and Grazia (2015). I.

*Calisius elegantulus* Bergroth, 1913. Kormilev and Froeschner (1987); Panizzi and Grazia (2015).***.

SUBFAMILY Carventinae Usinger, 1950.

Genus *Acaricoris* Harris and Drake, 1944

*Acaricoris austeris* Drake and Kormilev, 1958. Kormilev and Froeschner (1987); Panizzi and Grazia (2015).***.

Genus *Aglaocoris* Drake and Maldonado, 1955.

*Aglaocoris comes* Drake, 1956. Kormilev and Froeschner (1987); Panizzi and Grazia (2015).***.

*Aglaocoris invisus* Drake, 1957. Kormilev and Froeschner (1987); Panizzi and Grazia (2015).***.

*Aglaocoris vicinus* Drake, 1957. Kormilev and Froeschner (1987); Panizzi and Grazia (2015).***.

Genus *Antillaptera* Heiss, 2019.

*Antillaptera bassetterrana* Heiss, 2019. Heiss (2019).***.

Genus *Eretmocoris* Harris and Drake, 1944.

*Eretmocoris productus* Usinger and Matsuda, 1959. Kormilev and Froeschner (1987); Panizzi and Grazia (2015).***.

Genus *Glyptocoris* Harris and Drake, 1944.

*Glyptocoris insularis* (Drake, 1957). Kormilev and Froeschner (1987); Panizzi and Grazia (2015).***.

*Glyptocoris verus* Drake, 1956. Kormilev (1983); Kormilev and Froeschner (1987); Meurgey (2011b); Panizzi and Grazia (2015).***.
Genus *Kolpodaptera* Usinger and Matsuda, 1959.

*Kolpodaptera prominens* Usinger and Matsuda, 1959. Kormilev and Froeschner (1987); Panizzi and Grazia (2015).***

**SUPERFAMILY Coreoidea Leach, 1815.**

**FAMILY Alydidae Amyot and Audinet-Serville, 1843.**

Genus *Megalotomus* Fieber, 1860.

*Megalotomus pallescens* Stål 1860. Lethierry (1881).

Genus *Stenocoris* Burmeister, 1839.

*Stenocoris tipuloides* (De Geer, 1773). Lethierry (1881).

**FAMILY Coreidae Leach, 1815.**

**SUBFAMILY Coreinae Leach, 1815.**

**TRIBE Anisoscelini Laporte de Castelnau, 1840.**

Genus *Leptoglossus* Guérin-Méneville, 1831.

*Leptoglossus stigma* (Herbst, 1784). Lethierry (1881).

Genus *Phthia* Stål, 1862.

*Phthia lunata* (Fabricius, 1787). Lethierry (1881).

**TRIBE Hypselonotini Bergroth, 1913.**

Genus *Althos* Kirkaldy, 1904.

*Althos obscurator* (Fabricius, 1803). Lethierry (1881).

Genus *Anasa* Amyot and Audinet-Serville, 1843.

*Anasa bellator* (Fabricius, 1787). Lethierry (1881).

*Anasa scorbutica* (Fabricius, 1775). Lethierry (1881).

Genus *Dallacoris* Osuna, 1981.

*Dallacoris pictus* (Drury, 1770). Gruner and Riom (1977); Meurgey (2011b).

Genus *Phthiacnemia* Brailovsky, 2009.

*Phthiacnemia picta* (Drury, 1770). Lethierry (1881).

Genus *Spartocera* Laporte de Castelnau, 1832.

*Spartocera alternata* Dallas, 1852. Lethierry (1881).

*Spartocera batatas* (Fabricius, 1798). Lethierry (1881); Meurgey (2011b).

**FAMILY Rhopalidae Amyot and Audinet-Serville, 1843.**

Genus *Jadera* Stål, 1862.

*Jadera sanguinolenta* (Fabricius, 1775). Lethierry (1881).

**SUPERFAMILY Lygaeoidea Schilling, 1829.**

**FAMILY Geocoridae Baerensprung, 1860.**

Genus *Geocoris* Fallén, 1814.

*Geocoris deficiens* Lethierry, 1881. Lethierry (1881); Meurgey (2011b).

**FAMILY Lygaeidae Schilling, 1829.**

Genus *Kleidocerys* Stephens, 1829.

*Kleidocerys pictipes* Stål, 1874. Lethierry (1881).

Genus *Ochrimnus* Stål, 1874.

*Ochrimnus lineoloides* (Slater, 1964). Slater and Baranowski (1990); Meurgey (2011b).
Genus *Oncopeltus* Stål, 1868.

*Oncopeltus aulicus* (Fabricius, 1775). Slater and Baranowski (1990); Meurgey (2011b).

*Oncopeltus cingulifer* Stål, 1874. Slater and Baranowski (1979); Meurgey (2011b).

*Oncopeltus fasciatus* (Dallas, 1852). Lethierry (1881); Meurgey (2011b).

## FAMILY Pyrrhocoridae Fieber, 1860.

Genus *Dysdercus* Amyot and Audinet-Serville, 1835.

*Dysdercus andreae* (Linnaeus, 1758). Lethierry (1881); Gruner and Riom (1977); Klausner et al. (1981); Meurgey (2011b).

*Dysdercus delauneyi* Lethierry, 1881. Lethierry (1881); Meurgey (2011b).

## FAMILY Rhyparochromidae Amyot and Audinet-Serville, 1843.

Genus *Bathydema* Uhler, 1893.

*Bathydema sericea* (Lethierry, 1881). Lethierry (1881).

Genus *Cligenes* Distant, 1893.

*Cligenes distinctus* Distant, 1893. Brambila (2000).

Genus *Neopamera* Harrington, 1980.

139. *Neopamera bilobata* (Say, 1832). Melo et al. (2004); Meurgey (2011b).

Genus *Ozophora* Uhler, 1871.

*Ozophora burmeisteri* (Guérin-Méneville, 1857). Lethierry (1881); Meurgey (2011b).

*Ozophora hirsuta* Slater and Baranowski, 1979. Meurgey (2011b).

Genus *Pseudopachybrachius* Malipatil, 1978

*Pseudopachybrachius vinctus* (Say, 1831). Zheng and Slater (1984).

## SUPERFAMILY Pentatomoidea Reuter, 1910.

### FAMILY Cydnidae Billberg, 1820.

Genus *Cydnus* Fabricius, 1803.

*Cydnus insularis* Westwood, 1837. Lethierry (1881).

### FAMILY Pentatomidae Leach, 1815.

#### SUBFAMILY Asopinae Spinola, 1850.

Genus *Podisus* Herrich-Schäffer, 1851.

*Podisus distinctus* (Stål, 1860). Lethierry (1881).

#### SUBFAMILY Discocephalinae Fieber, 1860.

Genus *Macropygium* Spinola, 1837.

*Macropygium reticulare* (Fabricius, 1803). Lethierry (1881).

#### SUBFAMILY Edessinae Kirkaldy, 1909.

Genus *Edessa* Fabricius, 1803.

*Edessa bifida* (Say, 1831). Lethierry (1881).

*Edessa meditabunda* (Fabricius, 1794). Lethierry (1881).

#### SUBFAMILY Pentatominae Leach, 1815.

Genus *Acrosternum* Fieber, 1860.

*Acrosternum marginatum* (Palisot de Beauvois, 1817). Lethierry (1881); Rider (1987); Perez-Gelabert and Thomas (2005); Meurgey (2011b).
Genus *Loxa* Amyot and Audinet-Serville, 1843.

*Loxa flavicollis* (Drury, 1773). Lethierry (1881).

Genus *Mormidea* Amyot and Audinet-Serville, 1843.

*Mormidea ypsilon* (Linnaeus, 1758). Lethierry (1881).

Genus *Nezara* Amyot and Audinet-Serville, 1843.

*Nezara viridula* (Linnaeus, 1758). Meglič et al. (2001); Miklas et al. (2003); Kavar et al. (2006); Pavlovčič et al. (2008); Meurgey (2011b); Li et al. (2014).

Genus *Palomena* Mulsant and Rey, 1866.

*Palomena prasina* (Linnaeus, 1760). Lethierry (1881).

Genus *Piezodorus* Fieber, 1860.

*Piezodorus guildini* (Westwood, 1837). Lethierry (1881).

Genus *Thyanta* Stål, 1862.

*Thyanta testacea* (Dallas, 1851). Perez-Gelabert and Thomas (2005); Meurgey (2011b).

Genus *Tibraca* Stål, 1860.

*Tibraca limbativentris* Stål, 1860. Perez-Gelabert and Thomas (2005); Meurgey (2011b).

## FAMILY Scutelleridae Leach, 1815.

Genus *Tetyra* Fabricius, 1803.

*Tetyra pinguis* Germar, 1839. Lethierry (1881).

## FAMILY Tessaratomidae Stål, 1864.

Genus *Piezosternum* Amyot and Audinet-Serville, 1843.

*Piezosternum subulatum* (Thunberg, 1783). Lethierry (1881).

## SUBORDER Sternorrhyncha Amyot and Serville 1843.

## SUPERFAMILY Aleyrodoidea Westwood, 1840.

## FAMILY Aleyrodidae Westwood, 1840.

## SUBFAMILY Aleurodicinae Quaintance and Baker, 1913.

Genus *Aleurodicus* Douglas, 1892.

*Aleurodicus cocois* (Curtis, 1846). Streito et al. (2007); Meurgey (2011b).

*Aleurodicus dispersus* Russell, 1965. Streito et al. (2007); Meurgey (2011b).

*Aleurodicus maritimus* Hempel, 1923. Streito et al. (2007); Meurgey (2011b).

*Aleurodicus pulvinatus* (Maskell, 1895). Streito et al. (2007); Meurgey (2011b).

Genus *Aleuroglandulus* Bondar, 1923.

*Aleuroglandulus subtilis* Bondar, 1923. Streito et al. (2007); Meurgey (2011b).

Genus *Aleurotulus* Quaintance and Baker, 1914.

*Aleurotulus anthuricola* Nakahara, 1989. Streito et al. (2007); Meurgey (2011b).

*Aleurotulus pteridophytae* Martin, 1994. Streito et al. (2007); Meurgey (2011b).

Genus *Aleurovreus* Martin, 2005.

*Aleurovreus insignis* (Bondar, 1923). Streito et al. (2007); Meurgey (2011b).

## SUBFAMILY Aleyrodinae Westwood, 1840.

Genus *Aleurocanthus* Quaintance and Baker, 1914.

*Aleurocanthus woglumi* Ashby, 1915. Streito et al. (2007); Meurgey (2011b).
Genus *Aleuroclava* Singh, 1931.

*Aleuroclava jasmini* (Takahashi, 1932). Streito et al. (2007); Meurgey (2011b).

Genus *Aleurothrixus* Quaintance and Baker, 1914.

*Aleurothrixus antidesmae* Takahashi, 1933. Streito et al. (2007); Kumar-Dubey et al. (2010).

*Aleurothrixus floccosus* (Maskell, 1896). Streito et al. (2007); Ghahari et al. (2009); Meurgey (2011b).

*Aleurothrixus myrtacei* Bondar, 1923. Streito et al. (2007); Meurgey (2011b).

Genus *Aleurotrachelus* Quaintance and Baker, 1914.

*Aleurotrachelus atratus* Hempel, 1922. Streito et al. (2007); Meurgey (2011b).

*Aleurotrachelus trachoides* (Back, 1912). Streito et al. (2007); Meurgey (2011b).

Genus *Asiothrixus* Dubey, Ko and Martin, 2010.

*Asiothrixus antidesmae* Takahashi, 1933. Meurgey (2011b).

Genus *Bemisia* Quaintance and Baker, 1914.

*Bemisia tabaci* (Gennadius, 1889). Sauvion et al. (1999); Pavis et al. (2003); Sauvion et al. (2005); Streito et al. (2007); Meurgey (2011b). I

Genus *Crenidorsum* Russell, 1945.

*Crenidorsum aroidephagus* Martin and Aguiar, 2001. Streito et al. (2007); Meurgey (2011b).

Genus *Dialeurolonga* Dozier, 1928.

*Dialeurolonga simplex* Takahashi, 1955. Streito et al. (2007); Meurgey (2011b).

Genus *Metaleurodicus* Quaintance and Baker, 1913.

*Metaleurodicus griseus* (Dozier, 1936). Streito et al. (2007); Meurgey (2011b).

*Metaleurodicus minimus* (Quaintance, 1900). Meurgey (2011b).

Genus *Orchamoplatus* Russell, 1958.

*Orchamoplatus mammaeferus* (Quaintance and Baker, 1917). Streito et al. (2007); Meurgey (2011b). I

Genus *Parabemisia* Takahashi, 1952.

*Parabemisia myricae* (Kuwana, 1927). Streito et al. (2007); Meurgey (2011b). I

Genus *Paraleyrodes* Quaintance, 1909.

*Paraleyrodes bondari* Peracchi, 1971. Streito et al. (2007); Meurgey (2011b).

*Paraleyrodes pulverans* Bondar, 1923. Streito et al. (2007); Meurgey (2011b).

*Paraleyrodes urichii* Quaintance and Baker, 1913. Streito et al. (2007); Meurgey (2011b).

Genus *Singhiella* Sampson, 1943.

*Singhiella citrifolii* (Morgan, 1893). Streito et al. (2007); Meurgey (2011b).

*Singhiella simplex* (Singh, 1931). Germain et al. (2014). I

Genus *Tetraleurodes* Cockerell, 1902.

*Tetraleurodes acaciae* (Quaintance, 1900). Streito et al. (2007); Meurgey (2011b).

Genus *Trialeurodes* Cockerell, 1902.

*Trialeurodes vaporariorum* (Westwood, 1856). Streito et al. (2007); Meurgey (2011b).

*Trialeurodes variabilis* (Quaintance, 1900). Streito et al. (2007); Meurgey (2011b).

**SUPERFAMILY Aphidoidea Geoffroy, 1762.**

**FAMILY Aphididae Latreille, 1802.**

Genus *Aphis* Linnaeus, 1758.

*Aphis coreopsidis* (Thomas, 1878). Etienne (2005); Meurgey (2011b).

*Aphis craccivora* Koch, 1854. Stary et al. (1987); Etienne (2005); Meurgey (2011b).
*Aphis gossypii* Glover, 1877. Stary et al. (1987); Etienne (2005); Sauvion et al. (2005); Meurgey (2011b).

*Aphis illinoiensis* Shimer, 1866. Stary et al. (1987); Etienne (2005); Meurgey (2011b).

*Aphis nerii* Boyer de Fonscolombe, 1841. Etienne (2005); Meurgey (2011b).

*Aphis spiraecola* Patch, 1914. Etienne (2005); Meurgey (2011b).

*Genus Aulacorthum* Mordvilko, 1914.

*Aulacorthum solani* Kaltenbach, 1843. Etienne (2005); Meurgey (2011b).

*Genus Brachycaudus* van der Goot, 1913.

*Brachycaudus helichrysi* (Kaltenbach, 1843). Etienne (2005); Meurgey (2011b).

*Genus Capitophorus* van der Goot, 1913.

*Capitophorus hippophaes* Walker, 1852. Etienne (2005); Meurgey (2011b).

*Genus Carolinaia* Wilson, 1911.

*Carolinaia cyperi* Ainslie, 1915. Etienne (2005); Meurgey (2011b).

*Genus Cavariella* Del Guercio, 1911.

*Cavariella aegopodii* (Scopoli, 1763). Etienne (2005); Meurgey (2011b).

*Genus Cerataphis* Lichtenstein, 1882.

*Cerataphis brasiliensis* (Hempel, 1901). Etienne (2005); Meurgey (2011b).

*Cerataphis orchidearum* (Westwood, 1879). Etienne (2005); Meurgey (2011b).

*Genus Cinara* Curtis, 1835.

*Cinara tujafilina* (Del Guercio, 1909). Etienne and Champoiseau (2011).

*Genus Geopemphigus* Hille Ris Lambers, 1933.

*Geopemphigus floccosus* (Moreira, 1925). Etienne (2005); Meurgey (2011b).

*Genus Hyperomyzus* Börner, 1933.

*Hyperomyzus carduellinus* (Theobald, 1915). Etienne (2005); Meurgey (2011b).

*Hyperomyzus lactucae* (Linnaeus, 1758). Etienne (2005); Meurgey (2011b).

*Genus Hysteroneura* Davis, 1919.

*Hysteroneura setariae* (Thomas, 1878). Etienne (2005); Meurgey (2011b).

*Genus Lipaphis* Mordvilko, 1928.

*Lipaphis erysimi* (Kaltenbach, 1843). Etienne (2005); Meurgey (2011b).

*Genus Macrosiphoniella* Del Guercio, 1911.

*Macrosiphoniella sanborni* (Gillette, 1908). Etienne (2005); Meurgey (2011b).

*Genus Macrosiphum* Passerini, 1860.

*Macrosiphum euphorbiae* (Thomas, 1878). Etienne (2005); Meurgey (2011b).

*Genus Melanaphis* van der Goot, 1917.

*Melanaphis sacchari* (Zehntner, 1897). Etienne (2005); Meurgey (2011b).

*Genus Microparsus* Patch, 1909.

*Microparsus brasiliensis* (Moreira, 1925). Etienne (2005); Meurgey (2011b).

*Genus Myzus* Passerini, 1860.

*Myzus ornatus* Laing, 1932. Etienne (2005); Meurgey (2011b).

*Myzus persicae* Sulzer, 1776. Etienne (2005); Meurgey (2011b).

*Genus Neotoxoptera* Theobald, 1915.

*Neotoxoptera formosana* (Takahashi, 1921). Etienne and Champoiseau (2011).
Genus *Pentalonia* Coquerel, 1859.

*Pentalonia nigronervosa* Coquerel, 1859. Etienne (2005); Meurgey (2011b).

Genus *Pseudoregma* Doncaster, 1966.

*Pseudoregma panicola* (Takahashi, 1921). Etienne (2005); Meurgey (2011b).

Genus *Rhodobium* Hille Ris Lambers, 1947.

*Rhodobium porosum* (Sanderson, 1900). Etienne (2005); Meurgey (2011b).

Genus *Rhopalosiphum* Koch, 1854.

*Rhopalosiphum maidis* (Fitch, 1856). Stary et al. (1987); Etienne (2005); Meurgey (2011b).

*Rhopalosiphum nymphaeae* (Linnaeus, 1761). Etienne (2005); Meurgey (2011b).

*Rhopalosiphum padi* (Linnaeus, 1758). Etienne (2005); Meurgey (2011b).

*Rhopalosiphum rufiabdominale* (Sasaki, 1899). Etienne (2005); Meurgey (2011b).

Genus *Sarucallis* Shinji, 1922.

*Sarucallis kahawaluokalani* (Kirkaldy, 1907). Etienne et al. (2018). I.

Genus *Schizaphis* Börner, 1931.

*Schizaphis graminum* (Rondani, 1852). Etienne (2005); Meurgey (2011b).

Genus *Sipha* Passerini, 1860.

*Sipha flava* (Forbes, 1884). Etienne (2005); Meurgey (2011b).

Genus *Sitobion* Mordvilko, 1914.

*Sitobion avenae* (Fabricius, 1775). Etienne (2005); Meurgey (2011b).

*Sitobion mesosphaeri* (Tissot, 1934). Etienne (2005); Meurgey (2011b).

Genus *Tetraneura* Hartig, 1841.

*Tetraneura nigriabdominalis* (Sasaki, 1899). Etienne (2005); Meurgey (2011b).

Genus *Toxoptera* Koch, 1856.

*Toxoptera aurantii* (Boyer de Fonscolombe, 1841). Stary et al. (1987); Etienne (2005); Meurgey (2011b).

*Toxoptera citricidus* (Kirkaldy, 1907). Yokomi et al. (1994); Etienne (2005); Meurgey (2011b).

Genus *Uroleucon* Mordvilko, 1914.

*Uroleucon ambrosiae* (Thomas, 1878). Etienne (2005); Meurgey (2011b).

*Uroleucon erigeronense* (Thomas, 1878). Etienne (2005); Meurgey (2011b).

Genus *Utamphorophora* Knowlton, 1946.

*Utamphorophora commelinensis* (Smith, 1960). Etienne (2005); Meurgey (2011b).

## SUPERFAMILY Coccoidea Handlirsch, 1903.

## FAMILY Asterolecaniidae Berlese, 1898.

Genus *Asterolecanium* Targioni Tozzetti, 1868.

*Asterolecanium arabidis* (Signoret, 1876). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Bambusaspis* Cockerell, 1902.

*Bambusaspis bambusae* (Boisduval, 1869). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Bambusaspis miliaris* (Boisduval, 1869). Matile-Ferrero and Etienne (2006).

*Bambusaspis mimica* (Russell, 1941). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Palmaspis* Bodenheimer, 1951.

*Palmaspis palmae* (Cockerell, 1893). Matile-Ferrero and Etienne (2006).

Genus *Russellaspis* Bodenheimer, 1951.

*Russellaspis pustulans* (Cockerell, 1892). Matile-Ferrero and Etienne (2006); Meurgey (2011b).
## FAMILY Cerococcidae Balachowski, 1942.

Genus *Cerococcus* Comstock, 1882.

*Cerococcus deklei* Kosztarab and Vest, 1966. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

## FAMILY Coccidae Fallen, 1814.

Genus *Cardiococcus* Cockerell, 1903.

*Cardiococcus umbonatus* Cockerell, 1903. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Ceroplastes* Gray, 1828.

*Ceroplastes ceriferus* (Fabricius, 1798). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Ceroplastes cirripediformis* Comstock, 1881. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Ceroplastes dugesii* Lichtenstein, 1885. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Ceroplastes floridensis* Comstock, 1881. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Ceroplastes rubens* Maskell, 1893. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Ceroplastes rusci* (Linnaeus, 1758). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Ceroplastes stellifer* (Westwood, 1871). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Coccus* Linnaeus, 1758.

*Coccus hesperidum hesperidum* (Linnaeus, 1758). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Coccus longulus* (Douglas, 1887). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Coccus moestus* De Lotto, 1959. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Coccus viridis* (Green, 1889). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Eucalymnatus* Cockerell in Cockerell and Parrott, 1901.

*Eucalymnatus tessellatus* (Signoret, 1873). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Milviscutulus* Williams and Watson, 1990.

*Milviscutulus mangiferae* (Green, 1889). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Parasaissetia* Takahashi, 1955.

*Parasaissetia nigra* (Nietner, 1861). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Phalacrococcus* Hodges and Hodgson, 2010.

*Phalacrococcus howertoni* Hodges and Hodgson, 2010. Etienne et al. (2014).

Genus *Philephedra* Cockerell, 1898.

*Philephedra broadwayi* (Cockerell, 1896). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Philephedra tuberculosa* Nakahara and Gill, 1985. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Protopulvinaria* Cockerell, 1894.

*Protopulvinaria pyriformis* (Cockerell, 1894). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Pseudokermes* Cockerell, 1895.

*Pseudokermes vitreus* (Cockerell, 1894). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Pulvinaria* Targioni Tozzetti, 1866.

*Pulvinaria psidii* Maskell, 1893. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Pulvinaria urbicola* Cockerell, 1893. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Saissetia* Deplanche, 1859.

*Saissetia coffeae* (Walker, 1852). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Saissetia miranda* (Cockerell and Parrott in Cockerell, 1899). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Saissetia neglecta* De Lotto, 1969. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Saissetia oleae* (Olivier, 1791). Matile-Ferrero and Etienne (2006); Meurgey (2011b).
## FAMILY Diaspididae Targioni-Tozzetti, 1868.

Genus *Abgrallaspis* Balachowsky, 1948.

*Abgrallaspis cyanophylli* (Signoret, 1869). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Andaspis* MacGillivray, 1921.

*Andaspis punicae* (Laing, 1929). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Aonidiella* Berlese and Leonardi, 1896.

*Aonidiella comperei* McKenzie, 1937. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Aonidiella orientalis* (Newstead, 1894). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Aonidomytilus* Leonardi, 1903.

*Aonidomytilus albus* (Cockerell, 1893). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Aspidiella* Leonardi, 1898.

*Aspidiella hartii* (Cockerell, 1895). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Aspidiella sacchari* (Cockerell, 1893). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Aspidiotus* Bouché, 1833.

*Aspidiotus destructor* Signoret, 1869. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Aspidiotus excisus* Green, 1896. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Aspidiotus nerii* Bouché, 1833. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Aulacaspis* Cockerell, 1836.

*Aulacaspis tubercularis* Newstead, 1906. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Aulacaspis yasumatsui* Takagi, 1977. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Chortinaspis* Ferris, 1938.

*Chortinaspis subchortina* (Laing, 1929). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Chrysomphalus* Ashmead, 1880.

*Chrysomphalus aonidum* (Linnaeus, 1758). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Chrysomphalus dictyospermi* (Morgan, 1889). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Comstockiella* Cockerell, 1896.

*Comstockiella sabalis* (Comstock, 1883). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Dactylaspis* Ferris, 1937.

*Dactylaspis crotonis* (Cockerell, 1893). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Diaspis* Costa, 1828.

*Diaspis boisduvalii* Signoret, 1869. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Diaspis bromeliae* (Kerner, 1778).

Genus *Duplaspidiotus* MacGillivray, 1921.

*Duplaspidiotus tesseratus* (Grandpré and Charmoy, 1899). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Fiorinia* Targioni Tozzetti, 1868.

*Fiorinia proboscidaria* Green, 1900. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Furcaspis* Lindinger, 1908.

*Furcaspis biformis* (Cockerell, 1893).

Genus *Hemiberlesia* Cockerell, 1897.

*Hemiberlesia lataniae* (Signoret, 1869). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Hemiberlesia palmae* (Cockerell, 1892). Matile-Ferrero and Etienne (2006); Meurgey (2011b).
Genus *Howardia* Berlese and Leonardi, 1896.

*Howardia biclavis* (Comstock, 1883). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Ischnaspis* Douglas, 1887.

*Ischnaspis longirostris* (Signoret, 1882). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Lepidosaphes* Shimer, 1868.

*Lepidosaphes beckii* (Newman, 1869). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Lepidosaphes gloverii* (Packard, 1869). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Lepidosaphes rubrovittata* Cockerell, 1905. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Lepidosaphes tokionis* (Kuwana, 1902). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Lepidosaphes vermiculus* Mamet, 1937. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Melanaspis* Cockerell, 1897.

*Melanaspis coccolobae* Ferris, 1943. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Morganella* Cockerell, 1897.

*Morganella longispina* (Morgan, 1889). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Mycetaspis* Cockerell, 1897.

*Mycetaspis personata* (Comstock, 1883). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Odonaspis* Leonardi, 1897.

*Odonaspis greenii* Cockerell, 1902. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Odonaspis ruthae* Kotinsky, 1915. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Odonaspis secreta* (Cockerell, 1896). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Parlatoria* Targioni Tozzetti, 1868.

*Parlatoria cinerea* Hadden, 1909. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Parlatoria crotonis* Douglas, 1887. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Pinnaspis* Cockerell, 1892.

*Pinnaspis aspidistrae* (Signoret, 1869). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Pinnaspis buxi* (Bouché, 1851). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Pinnaspis strachani* (Cooley, 1899). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Pinnaspis theae* (Maskell, 1891). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Pseudaonidia* Cockerell, 1897.

*Pseudaonidia trilobitiformis* (Green, 1896). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Pseudaulacaspis* MacGillivray, 1921.

*Pseudaulacaspis pentagona* (Targioni Tozzetti, 1886). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Pseudoparlatoria* Cockerell, 1892.

*Pseudoparlatoria ostreata* Cockerell, 1892. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Selenaspidus* Cockerell, 1897.

*Selenaspidus articulatus* (Morgan, 1889). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Umbaspis* MacGillivray, 1921.

*Umbaspis regularis* (Newstead, 1911). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Unaspis* MacGillivray, 1921.

*Unaspis citri* (Comstock, 1883). Matile-Ferrero and Etienne (2006); Meurgey (2011b).
## FAMILY Eriococcidae Cockerell, 1903.

Genus *Eriococcus* Targioni Tozzetti, 1868.

*Eriococcus dubius* Cockerell, 1896. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

## FAMILY Margarodidae Guilging, 1829.

Genus *Icerya* Signoret, 1875.

*Icerya purchasi* Maskell, 1879. Matile-Ferrero and Etienne (2006); Etienne and Matile-Ferrero (2008); Meurgey (2011b).

*Icerya seychellarum* (Westwood, 1855). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

## FAMILY Monophlebidae Maskell, 1880.

Genus *Crypticerya* Cockerell, 1894.

*Crypticerya genistae* (Hempel, 1912). Matile-Ferrero and Etienne (2006); Meurgey (2011b). *Crypticerya rosae* (Riley and Howard, 1890). Matile-Ferrero and Etienne (2006); Etienne and Matile-Ferrero (2008); Meurgey (2011b).

## FAMILY Ortheziidae Amyot and Audinet-Serville, 1843.

Genus *Insignorthezia* Kozár, 2004.

*Insignorthezia insignis* (Browne, 1887). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Mixorthezia* Morrison, 1925.

*Mixorthezia fodiens* (Giard, 1897). ***

*Mixorthezia reynei* (Laing, 1925). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Nipponorthezinella* Kozár, 2004.

*Nipponorthezinella guadalcanalia* (Morrison, 1952). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Ortheziola* Šulc, 1895.

*Ortheziola fodiens* (Giard, 1897). Matile-Ferrero et al. (2006); Meurgey (2011b).

Genus *Praelongorthezia* Kozár, 2004.

*Praelongorthezia praelonga* (Douglas, 1891). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

## FAMILY Pseudococcidae Heymons, 1915.

Genus *Antonina* Signoret, 1875.

*Antonina crawl* Cockerell, 1900.

*Antonina graminis* (Maskell, 1897). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Chaetococcus* Maskell, 1898.

*Chaetococcus bambusae* (Maskell, 1893). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Chorizococcus* McKenzie, 1960.

*Chorizococcus rostellum* (Lobdell, 1930). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Dysmicoccus* Ferris, 1950.

*Dysmicoccus boninsis* (Kuwana, 1909). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Dysmicoccus brevipes* (Cockerell, 1893). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Dysmicoccus debregeasiae* (Green, 1922). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Dysmicoccus texensis* (Tinsley, 1900). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Ehrhornia* Ferris, 1918.

*Ehrhornia cupressi* (Ehrhorn, 1911). I
Genus *Ferrisia* Fullaway, 1923.

*Ferrisia virgata* (Cockerell, 1893). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Geococcus* Green, 1902.

*Geococcus coffeae* Green, 1933. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Hypogeococcus* Rau, 1938.

*Hypogeococcus pungens* Granara de Willink, 1981. Matile-Ferrero and Etienne (2006); Meurgey (2011b); German-Ramirez et al. (2014).

Genus *Maconelicoccus* Ezzat, 1958.

*Maconelicoccus hirsutus* (Green, 1908). Etienne et al. (1998); Matile-Ferrero et al. (2000); Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Nipaecoccus* Šulc, 1945.

*Nipaecoccus annonae* Williams and Granara de Willink, 1992. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Nipaecoccus guazumae* (Balachowsky, 1959). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Nipaecoccus nipae* (Maskell, 1893). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Palmicultor* Williams, 1963.

*Palmicultor palmarum* (Ehrhorn, 1916). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Paracoccus* Ezzat and McConnell, 1956.

*Paracoccus marginatus* Williams and Granara de Willink, 1992. Matile-Ferrero and Etienne (1998); Matile-Ferrero et al. (2000); Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Phenacoccus* Cockerell, 1893.

*Phenacoccus herreni* Cox and Williams, 1981. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Phenacoccus madeirensis* Green, 1923. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Phenacoccus parvus* Morrison, 1924. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Phenacoccus solani* Ferris, 1918. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Phenacoccus solenopsis* Tinsley, 1898. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Planococcus* Ferris, 1950.

*Planococcus citri* (Risso, 1813). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Planococcus halli* Ezzat and McConnell, 1956. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Planococcus minor* (Maskell, 1897). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Pseudococcus* Westwood, 1840.

*Pseudococcus cryptus* Hempel, 1918. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Pseudococcus elisae* Borchsenius, 1947. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Pseudococcus jackbeardsleyi* Gimpel and Miller, 1996. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Pseudococcus longispinus* (Targioni Tozzetti, 1867). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Pseudococcus maritimus* (Ehrhorn, 1900). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Pseudococcus viburni* (Signoret, 1875). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Puto* Signoret, 1876.

*Puto barberi* (Cockerell, 1895). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Rhizoecus* Kunckel d'Herculais, 1878.

*Rhizoecus americanus* (Hambleton, 1946). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Rhizoecus amorphophalli* Betrem, 1940. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Rhizoecus arabicus* Hambleton, 1976. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Rhizoecus eloti* Giard, 1897. Matile-Ferrero, Williams and Etienne (2006); Meurgey (2011b). ***
*Rhizoecus mayanus* (Hambleton, 1946). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

*Rhizoecus variabilis* Hambleton, 1978. Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Ripersiella* Tinsley, 1899.

*Ripersiella bituberculata* (McKenzie, 1960). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Saccharicoccus* Ferris, 1950.

*Saccharicoccus sacchari* (Cockerell, 1895). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Spilococcus* Ferris, 1950.

*Spilococcus mamillariae* (Bouché, 1844). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

Genus *Trionymus* Berg, 1899.

*Trionymus radicicola* (Morrison, 1926). Matile-Ferrero and Etienne (2006); Meurgey (2011b).

## SUPERFAMILY PSYLLOIDEA LATREILLE, 1807.

### FAMILY PSYLLIDAE LATREILLE, 1807.

Genus *Acizzia* Heslop-Harrison, 1961.

*Acizzia uncatoides* (Ferris and Klyver, 1932).

Genus *Diaphorina* Löw, 1880.

*Diaphorina citri* Kuwayama, 1908. Etienne et al. (1998); Etienne et al. (2001); Halbert and Núñez (2004); Pluke et al. (2005); Meurgey (2011b); Boykin et al. (2012).

Genus *Euceropsylla* Boselli, 1929.

*Euceropsylla cayeyensis* (Caldwell, 1942).

## ORDER HYMenOPTERA LINnaeus, 1758 (190 SPECIES).

### SUBORDER Apocrita Gerstaecker, 1867.

#### SUPERFAMILY Apoidea Latreille, 1802.

##### FAMILY Apidae Latreille, 1802.

###### SUBFAMILY Apinae Latreille, 1802.

##### TRIBE Anthophorini Dahlbom, 1835

Genus *Anthophora* Latreille, 1803

*Anthophora* (*Mystacanthophora*) *tricolor* (Fabricius, 1775). Meurgey (2014b); Meurgey (2016c).

##### TRIBE Apini Latreille, 1802.

Genus *Apis* Linnaeus, 1758.

*Apis mellifera* Linnaeus, 1758. Meurgey (2014b); Meurgey (2016c). I

*Apis mellifera adansonii* (Latreille, 1804). Chalumeau and Delplanque (1977); Meurgey (2011b); Meurgey (2014b); Meurgey (2016c).

*Apis mellifera ligustica* Spinola, 1806. Chalumeau and Delplanque (1977); Gruner and Riom (1977); Meurgey (2011b); Meurgey (2014b); Meurgey (2016c).

##### TRIBE Centridini Cockerell and Cockerell, 1901.

Genus *Centris* Fabricius, 1804

*Centris* (*Centris*) *decolorata* Lepeletier de Saint Fargeau, 1841. Meurgey (2014b); Meurgey (2016c).

*Centris* (*Heterocentris*) *lanipes* (Fabricius, 1775). Genaro and Franz (2008); Meurgey (2014b); Meurgey (2016c).

*Centris* (*Centris*) *smithii* Cresson, 1879. Meurgey (2015); Meurgey (2016c).

*Centris* (*Centris*) *versicolor* (Fabricius, 1775). Meurgey (2014b); Meurgey (2016c). **
TRIBE Ericrocidini Cockerell and Atkins, 1902.

Genus *Mesoplia* Lepeletier de Saint Fargeau, 1841.

*Mesoplia azurea* (Lepeletier de Saint Fargeau and Audinet-Serville, 1825). Meurgey (2014b); Meurgey (2016c).**

TRIBE Meliponini Lepeletier de Saint Fargeau, 1836.

Genus *Melipona* Illiger, 1806.

*Melipona variegatipes* Gribodo, 1893. Ashmead (1900); Meurgey (2014b); Meurgey (2016c).**

SUBFAMILY Eucerinae Latreille, 1802

TRIBE Eucerini Latreille, 1802

Genus *Melissodes* Latreille, 1829.

*Melissodes* (*Melissodes*) *martinicensis* Cockerell, 1917. Meurgey (2016c).

*Melissodes* (*Ecplectica*) *trifasciatus* Cresson, 1878. Meurgey (2015); Meurgey (2016c). *

TRIBE Exomalopsini Michener, 1944

Genus *Exomalopsis* Spinola, 1853.

*Exomalopsis* (*Exomalopsis*) *analis* Spinola 1853. Meurgey (2014b); Meurgey (2014c); Meurgey (2016c).

*Exomalopsis* (*Exomalopsis*) *bartschi* Timberlake, 1980. Meurgey (2014b); Meurgey (2014c); Meurgey (2016c).***

*Exomalopsis* (*Exomalopsis*) *similis* (Cresson, 1865). Genaro and Franz (2008); Meurgey (2014b); Meurgey (2014c); Meurgey (2016c).

SUBFAMILY Xylocopinae Latreille, 1802.

TRIBE Xylocopini Latreille, 1802.

Genus *Xylocopa* Latreille, 1802.

*Xylocopa* (*Neoxylocopa*) *caribea* (Lepeletier de Saint Fargeau, 1841). Meurgey (2014b); Meurgey (2016c). **

*Xylocopa* (*Neoxylocopa*) *fimbriata* (Fabricius, 1804). Meurgey (2014b); Meurgey (2016c).

*Xylocopa* (*Neoxylocopa*) *mordax* Smith, 1874. Genaro and Franz (2008); Meurgey (2014b); Meurgey (2016c). *

FAMILY Megachilidae Latreille, 1802.

SUBFAMILY Megachilinae Latreille, 1802.

TRIBE Megachilini Latreille, 1802.

Genus *Coelioxys* Latreille, 1809.

*Coelioxys abdominalis* Guérin-Méneville, 1844. Meurgey (2016); Meurgey (2016c).

*Coelioxys rufipes* Guérin-Méneville, 1844. Meurgey (2014b); Meurgey (2016c).

Genus *Megachile* Latreille, 1802.

*Megachile* (*Eutricharea*) *concinna* Smith, 1879. (Meurgey 2014b); Meurgey (2016c). I

*Megachile* (*Pseudomegachile*) *lanata* (Fabricius, 1775). Gruner and Riom (1977); Genaro and Franz (2008); Meurgey (2014b); Meurgey (2016c).

*Megachile* (*Callomegachile*) *rufipennis* (Fabricius, 1793). Genaro and Franz (2008); Meurgey (2014b); Meurgey (2016c).

*Megachile* (*Pseudocentron*) *vitraci* Pérez 1884. Meurgey (2014b); Meurgey (2016c). **
FAMILY Crabronidae Latreille, 1802.

SUBFAMILY Bembicinae Latreille, 1802.

TRIBE Bembicini Latreille, 1802.

Genus *Stictia* Illiger, 1807.

*Stictia signata* (Linnaeus, 1758). Genaro (2006).

SUBFAMILY Crabroninae Latreille, 1802.

TRIBE Crabronini Latreille, 1802.

Genus *Rhopalum* Stephens, 1829.

*Rhopalum duclosi* Leclercq, 2002. Leclercq (2002); Meurgey (2011b). ***

FAMILY Sphecidae Latreille, 1802.

SUBFAMILY Sceliphrinae Ashmead, 1899.

TRIBE Sceliphrini Ashmead, 1899.

Genus *Sceliphron* Klug, 1801.

*Sceliphron caementarium* (Drury, 1773). Van Der Vecht and van Breugel (1968).

*Sceliphron fasciatum* (Lepeletier de Saint Fargeau, 1845). Van Der Vecht and van Breugel (1968).

SUBFAMILY Sphecinae Latreille, 1802.

TRIBE Sphecini Latreille, 1802.

Genus *Sphex* Linnaeus, 1758.

*Sphex dorsalis* Lepeletier de Saint Fargeau, 1845. Genaro (2006).

*Sphex resinipes* (Fernald, 1906). Pulawski (2013).

SUPERFAMILY Chalcidoidea Latreille, 1817.

FAMILY Aphelinidae Thomson, 1876.

SUBFAMILY Aphelininae Thomson, 1876.

Genus *Aphelinus* Dalman, 1820.

*Aphelinus gossypii* Timberlake, 1924. Hayat and Fatima (1992).

Genus *Aphytis* Howard, 1900.

*Aphytis lepidosaphes* Compere, 1955.

SUBFAMILY Coccophaginae Förster, 1878.

Genus *Encarsia* Förster, 1878.

*Encarsia brasiliensis* (Hempel, 1904).

*Encarsia cubensis* Gahan, 1931.

*Encarsia guadeloupae* Viggiani, 1987.

*Encarsia luteola* Howard, 1895.

*Encarsia nigricephala* Dozier, 1937.

*Encarsia pergandiella* Howard, 1907.

*Encarsia quaintancei* Howard, 1907.

*Encarsia sophia* (Girault and Dodd, 1915).

SUBFAMILY Eretmocerinae Shafee and Khan, 1978.

Genus *Eretmocerus* Haldeman, 1850.

*Eretmocerus cocois* Delvare, 2008. Delvare et al. (2008).

*Eretmocerus desantisi* Rose, 2001. Delvare et al. (2008).

*Eretmocerus gracilis* Rose, 2001. Delvare et al. (2008).
## FAMILY Chalcididae Latreille, 1817.

### SUBFAMILY Chalcidinae Latreille, 1817.

Genus *Conura* Spinola, 1837.

*Conura fulvovariegata* (Cameron, 1884).

*Conura pseudofulvovariegata* (Becker, 1989).

*Conura quadrilineata* (Cameron, 1913).

## FAMILY Encyrtidae Walker, 1837.

### SUBFAMILY Encyrtinae Walker, 1837.

Genus *Ixodiphagus* Howard, 1907.

*Ixodiphagus hookeri* (Howard, 1908).

Genus *Ooencyrtus* Ashmead, 1900.

*Ooencyrtus submetallicus* (Howard, 1897).

### SUBFAMILY Tetracneminae Howard, 1892.

Genus *Anagyrus* Howard, 1896.

*Anagyrus diversicornis* (Howard, 1894).

*Anagyrus kamali* Moursi, 1948.

*Anagyrus punctulatus* Agarwal and Alam, 1959.

Genus *Leptomastix* Förster, 1856.

*Leptomastix dactylopii* Howard, 1885. Meurgey (2011b).

## FAMILY Eulophidae Westwood, 1829.

### SUBFAMILY Entedoninae Förster, 1856.

Genus *Goetheana* Girault, 1920.

*Goetheana parvipennis* (Gahan, 1927). Etienne et al. (2015).

*Goetheana shakesparei* Girault, 1920. Bennett et al. (1993).

Genus *Horismenus* Walker, 1843.

*Horismenus alnus* Hansson, 2009.

*Horismenus brachycaulis* Hansson, 2009.

*Horismenus lanius* Hansson, 2009.

Genus *Thripoctenus* Crawford, 1911.

*Thripoctenus javae* (Girault, 1917). Etienne et al. (2015). I

### SUBFAMILY Eulophinae Westwood, 1829.

Genus *Euplectrus* Westwood, 1832.

*Euplectrus platyhypenae* Howard, 1885. Zhou et al. (2003); Meurgey (2011b).

### SUBFAMILY Tetrastichinae Förster, 1856.

Genus *Aprostocetus* Westwood, 1833.

*Aprostocetus gala* (Walker, 1847). Etienne and Delvare (1991); Meurgey (2011b).

Genus *Quadrastichus* Girault, 1913.

*Quadrastichus erythrinae* Kim, 2004. Etienne and Dumbardon-Martial (2013). I

*Quadrastichus haitiensis* (Gahan, 1929). Etienne and Delvare (1991); Meurgey (2011b).

Genus *Tamarixia* Mercet, 1924.

*Tamarixia radiata* (Waterston, 1922). Etienne et al. (2001).
Genus *Thripastichus* Graham, 1987.

*Thripastichus gentilei* (Del Guercio, 1931). Etienne et al. (2015).

Genus *Trisecodes* Delvare and LaSalle, 2000.

*Trisecodes agromyzae* Delvare and LaSalle, 2000.

**FAMILY Eurytomidae Walker, 1832.**

**SUBFAMILY Eurytominae Walker, 1832.**

Genus *Eurytoma* Illiger, 1807.

*Eurytoma attiva* Burks, 1958. Gates and Delvare (2008); Meurgey (2011b).

*Eurytoma obtusiventris* Gahan, 1934.

*Eurytoma tricoloripes* Girault, 1915.

Genus *Phylloxeroxenus* Ashmead, 1888.

*Phylloxeroxenus cressoni* (Howard, 1897). Gates and Delvare (2008); Meurgey (2011b).

**SUBFAMILY Rileyinae Ashmead, 1904.**

Genus *Rileya* Ashmead, 1888.

*Rileya pulchra* (Ashmead, 1894).

**FAMILY Mymaridae Haliday, 1833.**

Genus *Anagrus* Haliday, 1833.

*Anagrus breviphragma* Soyka, 1956.

*Anagrus flaveolus* Waterhouse, 1913.

**FAMILY Trichogrammatidae Haliday, 1851.**

**SUBFAMILY Oligositinae Ashmead, 1904.**

Genus *Megaphragma* Timberlake, 1924.

*Megaphragma caribea* Delvare, 1993. Meurgey (2011b). ***

*Megaphragma mymaripenne* Timberlake, 1924. Meurgey (2011b).

**SUBFAMILY Trichogrammatinae Haliday, 1851.**

Genus *Ceratogramma* De Santis, 1957.

*Ceratogramma etiennei* Delvare, 1988.

Genus *Trichogramma* Westwood, 1833.

*Trichogramma minutum* Riley, 1871.

**SUPERFAMILY Chrysidoidea Latreille, 1802.**

**FAMILY Chrysididae Latreille, 1802.**

**SUBFAMILY Amiseginae Moscáry, 1890.**

Genus *Adelphe* Mocsáry, 1890.

*Adelphe nesos* Kimsey, 1986. Kimsey (1986); Meurgey (2011b).

**FAMILY Dryinidae Haliday, 1840.**

Genus *Anteon* Jurine, 1807.

*Anteon guadeloupense* Olmi, 1989. Olmi et al. (2000).

*Anteon micros* Olmi, 1984. Olmi et al. (2000).

Genus *Gonatopus* Ljungh, 1810.

*Gonatopus cubensis* (Richards, 1969). Genaro and Portuondo (1997); Olmi et al. (2000); Meurgey (2011b).
*Gonatopus flavipes* Olmi, 1984. Olmi et al. (2000).

*Gonatopus flavoides* Olmi, 1993. Olmi et al. (2000).

*Gonatopus variistriatus* (Fenton, 1927). Olmi et al. (2000).

**SUPERFAMILY Cynipoidea Latreille, 1802.**

**FAMILY Figitidae Thomson, 1862.**

**SUBFAMILY Eucoilinae Thomson, 1862.**

Genus *Cothonaspis* Hartig in Germar, 1840.

*Cothonaspis boulardii* Kelner-Pillault, 1979. Claret and Carton (1980); Allemand et al. (2002); Meurgey (2011b).

Genus *Zaeucoila* Ashmead, 1903.

*Zaeucoila incompleta* (Kieffer, 1907). Buffington et al. (2017).

*Zaeucoila infuscata* Gallardo and Buffington, 2017. Buffington et al. (2017).

*Zaeucoila lignys* Buffington and Gallardo, 2017. Buffington et al. (2017).

*Zaeucoila robusta* (Ashmead, 1894). Buffington et al. (2017).

**SUPERFAMILY Ichneumonoidea Latreille, 1802.**

**FAMILY Braconidae Nees, 1811.**

**SUBFAMILY Aphidiinae Haliday, 1833.**

**TRIBE Aphidiini Haliday, 1833.**

Genus *Lysiphlebus* Forster, 1862.

*Lysiphlebus testaceipes* (Cresson, 1880). Stary et al. (1987); Meurgey (2011b).

**SUBFAMILY Microgastrinae Förster, 1862.**

**TRIBE Cotesiini Mason, 1981.**

Genus *Cotesia* Cameron, 1891.

*Cotesia americana* (Lepeletier de Saint Fargeau, 1825). Ashmead (1900); Meurgey (2011b).

*Cotesia flavipes* Cameron, 1891. Delplanque (1976); Delattre (1978); Cochereau (1991); Meurgey (2011b).

**FAMILY Ichneumonidae Latreille, 1802.**

**SUBFAMILY Pimplinae Wesmael, 1845.**

**TRIBE Pimplini Wesmael, 1845.**

Genus *Pimpla* Fabricius, 1804.

*Pimpla turionellae* (Linnaeus, 1758). Ashmead (1900); Meurgey (2011b).

**SUPERFAMILY Platygastroidea Haliday, 1833.**

**FAMILY Platygastridae (Masner and Huggert 1989).**

**SUBFAMILY Sceliotrachelinae Brues, 1908.**

Genus *Fidiobia* Ashmead, 1894.

*Fidiobia dominica* Evans and Peña, 2005. Evans et al. (2005); Meurgey (2011b).

**SUPERFAMILY Vespoidea Latreille, 1802.**

**FAMILY Formicidae Latreille, 1809.**

**SUBFAMILY Amblyoponinae Forel, 1893.**

**TRIBE Amblyoponini Forel, 1893.**

Genus *Prionopelta* Mayr, 1866.

*Prionopelta antillana* Forel, 1909. Jaffe and Lattke (1994); Meurgey (2011b).
SUBFAMILY Dolichoderinae Forel, 1878.

TRIBE Leptomyrmecini Emery, 1913.

Genus *Azteca* Forel, 1878.

*Azteca delpini antillana* Forel, 1899. Jaffe et al. (1989); Jaffe and Lattke (1994); Meurgey (2011b).

TRIBE Tapinomini Emery, 1913.

Genus *Tapinoma* Foerster, 1850.

*Tapinoma litorale* Wheeler, 1905. Jaffe et al. (1989); Meurgey (2011b).

*Tapinoma melanocephalum* (Fabricius, 1793). Jaffe and Lattke (1994); Meurgey (2011b). I

SUBFAMILY Dorylinae Leach, 1815.

Genus *Ooceraea* Roger, 1862.

*Ooceraea biroi* (Forel, 1907). Galkowski (2016). I

SUBFAMILY Ectatomminae Emery, 1895.

TRIBE Ectatommini Emery, 1895.

Genus *Ectatomma* F. Smith, 1858.

*Ectatomma ruidum* (Roger, 1860). Jaffe et al. (1989); Jaffe and Lattke (1994); Passera et al. (1994); Meurgey (2011b).

Genus *Gnamptogenys* Roger, 1863.

*Gnamptogenys striatula* Mayr, 1884. Jaffe and Lattke (1994); Meurgey (2011b).

TRIBE Typhlomyrmecini Emery, 1911.

Genus *Typhlomyrmex* Mayr, 1862.

*Typhlomyrmex pusillus* Emery, 1894. Galkowski (2016).

SUBFAMILY Formicinae Latreille, 1809.

TRIBE Camponotini Forel, 1878.

Genus *Camponotus* Mayr, 1861.

*Camponotus conspicuus williamsi* Forel, 1916. Galkowski (2016).

*Camponotus pittieri* Forel, 1899. Meurgey (2011b).

*Camponotus planatus* Roger, 1863. Galkowski (2016).

*Camponotus sexguttatus* (Fabricius, 1793). Jaffe and Lattke (1994); Meurgey (2011b).

*Camponotus sexguttatus antiguanus* Wheeler, 1923.

TRIBE Lasiini Ashmead, 1905.

Genus *Nylanderia* Emery, 1906.

*Nylanderia pubens* (Forel, 1893). Meurgey (2011b).

*Nylanderia steinheili* (Forel, 1893). Meurgey (2011b).

Genus *Paratrechina* Motschoulsky, 1863.

*Paratrechina longicornis* (Latreille, 1802). Jaffe et al. (1989); Jaffe and Lattke (1994); Meurgey (2011b). I

TRIBE Myrmelachistini Forel, 1912.

Genus *Brachymyrmex* Mayr, 1868.

*Brachymyrmex cordemoyi* Forel, 1895. Galkowski (2016).
TRIBE Plagiolepidini Forel, 1886.

Genus *Plagiolepis* Mayr, 1861.

*Plagiolepis alluaudi* Emery, 1894. Wetterer (2013).

SUBFAMILY Myrmicinae Lepeletier de Saint Fargeau, 1835.

TRIBE Attini F. Smith, 1858.

Genus *Acromyrmex* Mayr, 1865.

*Acromyrmex octospinosus* (Reich, 1793). Kermarrec (1975); Papa and Papa (1982); Febvay et al. (1985); Febvay et al. (1986); Rahbé et al. (1988); Gomel et al. (1989); Jaffe and Lattke (1994); Patin (2007); Meurgey (2011b). I

Genus *Cyphomyrmex* Mayr, 1862.

*Cyphomyrmex minutus* Mayr, 1862. Célini et al. (2013).

*Cyphomyrmex rimosus* (Spinola, 1851). Jaffe and Lattke (1994).

Genus *Eurhopalothrix* Brown and Kempf, 1961.

*Eurhopalothrix guadeloupensis* Longino, 2013. Longino (2013).

Genus *Mycetomoellerius* Solomon, Rabeling, Sosa-Calvo and Schultz, 2019.

*Mycetomoellerius jamaicensis* (André, 1893). Jaffe and Lattke (1994); Meurgey (2011b).

Genus *Mycocepurus* Forel, 1893.

*Mycocepurus smithii* (Forel, 1893). Jaffe and Lattke (1994); Meurgey (2011b).

Genus *Octostruma* Forel, 1912.

*Octostruma lutzi* (Wheeler, 1913). Longino (2013).

Genus *Pheidole* Westwood, 1839.

*Pheidole antillana* Forel, 1893. Galkowski (2016).

*Pheidole exigua* Mayr, 1884. Galkowski (2016).

*Pheidole fallax* Mayr, 1870. Wheeler (1923); Jaffe et al. (1989); Jaffe and Lattke (1994); Meurgey (2011b).

*Pheidole flavens* Roger, 1863. Galkowski (2016).

*Pheidole kukrana* Wilson, 2003. Galkowski (2016).

*Pheidole laudatana* Wilson, 2003. Galkowski (2016).

*Pheidole megacephala* (Fabricius, 1793). Jaffe and Lattke (1994); Meurgey (2011b). I

*Pheidole mendicula* Wheeler, 1925. Galkowski (2016).

*Pheidole orbica* Forel, 1893. Galkowski (2016).

*Pheidole sculptior* (Forel, 1893). Kermarrec (1981); Meurgey (2011b).

*Pheidole subarmata* Mayr, 1884. Galkowski (2016).

*Pheidole teneriffana* Forel, 1893. Wetterer (2011).

Genus *Sericomyrmex* Mayr, 1865.

*Sericomyrmex mayri* Forel, 1912. Galkowski (2016); Ješovnik and Schultz (2017).

Genus *Strumigenys* Smith, 1860.

*Strumigenys alberti* (Forel, 1893). Jaffe and Lattke (1994); Meurgey (2011b).

*Strumigenys eggersi* Emery, 1890. Galkowski (2016).

*Strumigenys elongata* Roger, 1863. Galkowski (2016).

*Strumigenys emmae* (Emery, 1890). Jaffe and Lattke (1994); Meurgey (2011b). I

*Strumigenys enopla* (Bolton, 2000). Galkowski (2016).

*Strumigenys fridericimuelleri* Forel, 1886. Galkowski (2016).

*Strumigenys louisianae* Roger, 1863. Galkowski (2016).

*Strumigenys margaritae* Forel, 1893. Galkowski (2016).

*Strumigenys rogeri* Emery, 1890. Wetterer (2012). I
*Strumigenys smithii* Forel, 1886. Jaffe and Lattke (1994); Meurgey (2011b).

*Strumigenys subedentata* (Mayr, 1887). Galkowski (2016).

Genus *Wasmannia* Forel, 1893.

*Wasmannia europunctata* (Roger, 1863). Jaffe et al. (1989); Jaffe and Lattke (1994); Wetterer and Porter (2003); Meurgey (2011b). I

*Wasmannia sigmoidea* (Mayr, 1884). Galkowski (2016).

TRIBE Crematogastrini Forel, 1893.

Genus *Cardiocondyla* Emery, 1869.

*Cardiocondyla emeryi* Forel, 1881. Wetterer (2013). I

*Cardiocondyla mauritanica* Forel, 1890. Galkowski (2016). I

*Cardiocondyla minutior* Forel, 1899. Galkowski (2016). I

*Cardiocondyla obscurior* Wheeler, 1929. Galkowski (2016). I

*Cardiocondyla wroughtonii* (Forel, 1890). Meurgey (2011b). I

Genus *Crematogaster* Lund, 1831.

*Crematogaster curvispinosa* Mayr, 1862. Galkowski (2016).

Genus *Tetramorium* Mayr, 1855.

*Tetramorium bicarinatum* (Nylander, 1846). Jaffe and Lattke (1994); Meurgey (2011b). I

*Tetramorium caldarium* (Roger, 1857). Wetterer et al. (2015); Galkowski (2016). I

*Tetramorium simillimum* (Smith, 1851). Jaffe and Lattke (1994); Meurgey (2011b).

Genus *Trichomyrmex* Mayr, 1865.

*Trichomyrmex destructor* (Jerdon, 1851). Jaffe and Lattke (1994); Meurgey (2011b).

TRIBE Solenopsidini Forel, 1893.

Genus *Monomorium* Mayr, 1855.

*Monomorium ebeninum* Forel, 1891. Jaffe and Lattke (1994); Meurgey (2011b).

*Monomorium floricola* (Jerdon, 1851). Jaffe and Lattke (1994); Meurgey (2011b).

*Monomorium minimum* (Buckley, 1867). Jaffe and Lattke (1994); Meurgey (2011b).

*Monomorium salomonis* (Linnaeus, 1758). Galkowski (2016).

Genus *Rogeria* Emery, 1894.

*Rogeria curvipubens* Emery, 1894. Galkowski (2016).

*Rogeria foreli* Emery, 1894. Jaffe and Lattke (1994); Meurgey (2011b).

*Rogeria scobinata* Kugler, 1994. Galkowski (2016).

Genus *Solenopsis* Westwood, 1840.

*Solenopsis azteca* Forel, 1893. Galkowski (2016).

*Solenopsis brevicornis* Emery, 1888. Galkowski (2016).

*Solenopsis geminata* (Fabricius, 1804). Wheeler (1923); Kermarrec (1981); Jaffe et al. (1989); Jaffe and Lattke (1994); Meurgey (2011b). I

*Solenopsis globularia* (Smith, 1858). Galkowski (2016).

*Solenopsis invicta* Buren, 1972. Wetterer and Davis (2010); Meurgey (2011b); Wetterer (2013).

*Solenopsis pygmaea* Forel, 1901. Galkowski (2016).

*Solenopsis vinsoni* Pacheco and Mackay, 2013. Galkowski (2016).

Genus *Syllophopsis* Santschi, 1915.

*Syllophopsis subcoeca* (Emery, 1894). Galkowski (2016).
SUBFAMILY Ponerinae Lepeletier de Saint Fargeau, 1835.

TRIBE Platythyreini Emery, 1901.

Genus *Platythyrea* Roger, 1863.

*Platythyrea punctata* (Smith, 1858). Seal et al. (2011).

*Platythyrea sinuata* (Roger, 1860). Jaffe and Lattke (1994); Meurgey (2011b).

TRIBE Ponerini Lepeletier de Saint Fargeau, 1835.

Genus *Anochetus* Mayr, 1861.

*Anochetus mayri* Emery, 1884. Galkowski (2016).

Genus *Hypoponera* Santschi, 1938.

*Hypoponera gleadowi* (Forel, 1895). Jaffe and Lattke (1994); Meurgey (2011b).

*Hypoponera opacior* (Forel, 1893). Galkowski (2016).

*Hypoponera punctatissima* (Roger, 1859). Jaffe and Lattke (1994); Meurgey (2011b).

*Hypoponera ragusai* (Emery, 1894). Galkowski (2016).

Genus *Leptogenys* Roger, 1861.

*Leptogenys arcuata* Roger, 1861. Jaffe and Lattke (1994); Meurgey (2011b).

Genus *Odontomachus* Latreille, 1804.

*Odontomachus bauri* Emery, 1892. Jaffe and Lattke (1994); Meurgey (2011b).

*Odontomachus haematodus* (Linnaeus, 1758). Jaffe and Lattke (1994); Meurgey (2011b).

*Odontomachus insularis* Guérin-Méneville, 1844. Jaffe and Lattke (1994); Meurgey (2011b).

*Odontomachus ruginodis* Smith, 1937. Galkowski (2016).

Genus *Pachycondyla* F. Smith, 1858.

*Pachycondyla harpax* (Fabricius, 1804). Galkowski (2016).

Genus *Pseudoponera* Emery, 1900.

*Pseudoponera stigma* (Fabricius, 1804). Jaffe and Lattke (1994); Meurgey (2011b).

SUBFAMILY Pseudomyrmecinae M.R. Smith, 1952.

TRIBE Pseudomyrmecini M.R. Smith, 1952.

Genus *Pseudomyrmex* Lund, 1831.

*Pseudomyrmex curacaensis* (Forel, 1912). Jaffe and Lattke (1994); Meurgey (2011b).

*Pseudomyrmex flavidulus* (Smith, 1858). Jaffe and Lattke (1994); Meurgey (2011b).

*Pseudomyrmex gracilis* (Fabricius, 1804). Wetterer (2010); Meurgey (2011b).

FAMILY Mutillidae Latreille, 1803.

SUBFAMILY Mutillinae Latreille, 1802.

TRIBE Mutillini Latreille, 1802.

Genus *Timulla* Ashmead, 1899.

*Timulla guadeloupensis* Mickel, 1937. Mickel (1938); Evans (1972).

FAMILY Pompilidae Latreille, 1805.

SUBFAMILY Pepsinae Lepeletier de Saint Fargeau, 1845.

Genus *Pepsis* Fabricius, 1805.

*Pepsis ruficornis* (Fabricius, 1775). Vardy (2002); Meurgey (2011b).

SUBFAMILY Pompilinae Latreille, 1805.

Genus *Anoplius* Dufour, 1834.

*Anoplius* (*Notiochoares*) *amethystinus amethystinus* (Fabricius, 1793). Waichert (2012).
**FAMILY Scoliidae Latreille, 1802.**

**SUBFAMILY Scoliinae Latreille, 1802.**

**TRIBE Campsomerini Betrem, 1972.**

Genus *Dielis* Saussure and Sichel, 1864.

*Dielis dorsata* (Fabricius, 1787). Gruner and Riom (1977); Meurgey (2011b).

**FAMILY Tiphiidae Leach, 1815.**

**SUBFAMILY Myzininae Ashmead, 1899.**

Genus *Myzinum* Latreille, 1803.

*Myzinum haemorrhoidalis haemorrhoidalis* (Fabricius, 1775). Meurgey (2011b);

**FAMILY Vespidae Latreille, 1802.**

**SUBFAMILY Eumeninae Leach, 1815.**

Genus *Odynerus* Latreille, 1802.

*Odynerus figulus* (Saussure, 1855). Ashmead (1900); Meurgey (2011b).

Genus *Pachodynerus* Saussure, 1875.

*Pachodynerus guadulpensis* (Saussure, 1853). Bequaert (1948); Evans (1972); Menke (1986); Meurgey (2011b).

**SUBFAMILY Polistinae Lepeletier de Saint Fargeau, 1836.**

Genus *Polistes* Latreille, 1802.

*Polistes crinitus* (Felton, 1765). Meurgey (2011b).

**ORDER Strepsiptera Kirby, 1813. (1 SPECIES).**

**SUBORDER Stylopidia Kinzelbach, 1969.**

**FAMILY Halictophagidae Perkins, 1905.**

Genus *Stenocranophilus* Pierce, 1914.

*Stenocranophilus quadratus* Pierce, 1914. Simmonds (1956).

**ORDER Coleoptera Linnaeus, 1758. (1447 SPECIES).**

Family-group names of Coleoptera are presented roughly in the same order used by Peck et al. (2014), except when this conflicts with more recent systematic studies of Coleoptera (e.g. McKenna et al. 2019). For a complete list of references, see Peck et al. (2014).

**SUBORDER Adephaga Schellenberg, 1806.**

**FAMILY Rhysodidae Laporte de Castelnau, 1840.**

Genus *Clinidium* Kirby, 1830.

*Clinidium planum* (Chevrolat, 1844). ***

*Clinidium smithsonianum* Bell and Bell, 1985.

**FAMILY Carabidae Latreille, 1802.**

**SUBFAMILY Carabinae Latreille, 1802.**

Genus *Calosoma* Weber, 1801.

*Calosoma alternans* (Fabricius, 1792). Malausa (1977); Lemaire (2017).

**Recent record.** (Meurgey, MHNN/SHNLH): Guadeloupe, Deshaies, Morne Duranton, 11.IV.2018, 628731 – 1801514, 136m, F. Meurgey leg (coll. SHNLH).

**SUBFAMILY Paussinae Latreille, 1806.**

**TRIBE Ozaenini Hope, 1838.**

Genus *Pachyteles* Perty, 1830.

*Pachyteles delauneyi* Fleutiaux and Sallé, 1890. ***
SUBFAMILY Cicindelinae Latreille, 1802.

TRIBE Cicindelini Latreille, 1802.

Genus *Brasiella* Rivalier, 1954.

*Brasiella argentata* (Fabricius, 1801).

*Brasiella argentata pallipes* (Fleutiaux and Sallé, 1889). **.

Genus *Cicindelidia* Rivalier, 1954.

*Cicindelidia trifasciata* (Fabricius, 1781).

Genus *Cylindera* Westwood, 1831.

*Cylindera suturalis* (Fabricius, 1798).

*Cylindera (Plectographa) suturalis guadeloupensis* (Fleutiaux and Sallé, 1889). ***.

SUBFAMILY Scaritinae Bonelli, 1810.

TRIBE Clivinini Rafinesque, 1815.

Genus *Halocoryza* Alluaud, 1919.

*Halocoryza arenaria* (Darlington, 1939). I

Genus *Oxydrepanus* Putzeys, 1866.

*Oxydrepanus micans* Putzeys, 1866.

Genus *Semiardistomis* Kult, 1950.

*Semiardistomis laevistriatus* (Fleutiaux and Salle, 1889). ***

TRIBE Scaritini Bonelli, 1810.

Genus *Ardistomis* Putzeys, 1846.

*Ardistomis atripennis* Putzeys, 1866. ***

*Ardistomis guadeloupensis* Kult, 1950. ***

Genus *Aspidoglossa* Putzeys, 1846.

*Aspidoglossa cribrata* Putzeys, 1846.

*Aspidoglossa schach* (Fabricius, 1792).

Genus *Clivina* Latreille, 1802.

*Clivina elongata* Chaudoir, 1843.

*Clivina fasciata* Putzeys, 1846. I

*Clivina (Paraclivina) latiuscula* Putzeys, 1866. ***

*Clivina marginipennis* Putzeys, 1846.

*Clivina tuberculata* Putzeys, 1846.

Genus *Scarites* Fabricius, 1775.

*Scarites octocoelus* (Chaudoir, 1855). I

SUBFAMILY Trechinae Bonelli, 1810.

TRIBE Bembidiini Stephens, 1827.

Genus *Elaphropus* Motschulsky, 1839.

*Elaphropus fumigatus* (Motschulsky, 1851).

Genus *Mioptachys* Bates, 1882.

*Mioptachys autumnalis* (Bates, 1882).

Genus *Tachys* Dejean, 1821.

*Tachys abruptus* Darlington, 1934. **

*Tachys albipes* (Leconte, 1863). I

*Tachys ensenadae* Mutchler, 1934. Lemaire (2017).
*Tachys pallidus* Chaudoir, 1868. Lemaire (2017).

### TRIBE Trechini Bonelli, 1810.

Genus *Perileptus* Schaum, 1860.

*Perileptus dentifer* Darlington, 1935. Peck (2016).

### SUBFAMILY Harpalinae Bonelli, 1810.

#### TRIBE Harpalini Bonelli, 1810.

Genus *Amblygnathus* Dejean, 1829.

*Amblygnathus cephalotes* Dejean, 1829. Shpeley et al. (2017).

*Amblygnathus gilvipes* Ball and Maddison, 1987. Shpeley et al. (2017).

Genus *Athrostictus* Bates, 1878.

*Athrostictus paganus* (Dejean, 1831). Shpeley et al. (2017). *

Genus *Selenophorus* Dejean, 1831.

*Selenophorus alternans* Dejean, 1829. Shpeley et al. (2017).

*Selenophorus discopunctatus* Dejean, 1829. Lemaire (2017). Shpeley et al. (2017).

*Selenophorus integer* (Fabricius, 1801). Lemaire (2017). Lemaire (2017); Shpeley et al. (2017).

*Selenophorus irec* Shpeley, Hunting and Ball, 2017. Shpeley et al. (2017). ***

*Selenophorus latior* Darlington, 1934. Shpeley et al. (2017). *

*Selenophorus parumpunctatus* (Dejean, 1829). Lemaire (2017). Shpeley et al. (2017).

*Selenophorus propinquus* Putzeys, 1874. Lemaire (2017). Shpeley et al. (2017).

*Selenophorus striatopunctatus* Putzeys, 1878. Shpeley et al. (2017).

*Selenophorus sinuatus* Gyllenhal, 1806. ***

*Selenophorus subaeneus* Reiche, 1843.

*Selenophorus subquadratus* Putzeys, 1878. Shpeley et al. (2017). *

#### TRIBE Pentagonicini Bates, 1873.

Genus *Pentagonica* Schmidt-Gobel, 1846.

*Pentagonica flavipes* (LeConte, 1853).

#### TRIBE Platynini Bonelli, 1810.

Genus *Dyscolus* Dejean, 1831.

*Dyscolus lherminieri* (Chaudoir, 1842) Lemaire (2017). **

Genus *Glyptolenus* Bates, 1878.

*Glyptolenus chalybaeus* Dejean, 1831.

Genus *Platynus* Bonelli, 1810.

*Platynus alternans* (Chaudoir, 1878). ***

*Platynus dejeani* (Chaudoir, 1859). ***

*Platynus elongatus* (Chaudoir, 1878). ***

*Platynus memnonius* (Dejean, 1831). ***

#### TRIBE Perigonini Horn, 1881.

Genus *Perigona* Laporte de Castelnau, 1835.

*Perigona guadeloupensis* Fleutiaux and Sallé, 1889. ***

*Perigona nigriceps* (Dejean, 1831) Lemaire (2017). I

*Perigona picea* Darlington, 1934. *
TRIBE Lachnophorini LeConte, 1853.

Genus *Anchonoderus* Reiche, 1843.

*Anchonoderus subaeneus* Reiche, 1843. I

TRIBE Lebiini Bonelli, 1810.

Genus *Apenes* LeConte, 1851.

*Apenes chalumeaui* Ball and Shpeley, 1992.**

*Apenes marginalis* (Dejean, 1831). Lemaire (2017).

*Apenes purpurata* Fleutiaux and Sallé, 1890. **

*Apenes sculpticeps* Ball and Shpeley, 2009.

*Apenes variegata* Dejean, 1825. Lemaire (2017).

Genus *Cymindis* Latreille, 1806.

*Cymindis binotata* Fischer von Waldheim, 1820.

*Cymindis pallipes* Fabricius, 1792.

Genus *Lebia* Latreille, 1802.

*Lebia frenata* (Chaudoir, 1871).

*Lebia frenata chevrolati* Blackwelder, 1944.

Genus *Plochionus* Dejean, 1821.

*Plochionus amandus* Newman, 1840.

Genus *Somotrichus* Seidlitz 1887.

*Somotrichus unifasciatus* (Dejean, 1831). I

TRIBE Galeritini Jeannel, 1810.

Genus *Galerita* Fabricius, 1801.

*Galerita americana* (Linnaeus, 1758).

*Galerita microcostata* Darlington, 1934.

*Galerita tristis* Reiche, 1842.

*Galerita unicolor* Latreille and Dejean, 1823.

FAMILY Gyrinidae Latreille, 1810.

SUBFAMILY Gyrininae Latreille, 1810.

TRIBE Dineutini Desmarest, 1851.

Genus *Dineutus* Macleay, 1825.

*Dineutus metallicus* Aubé, 1838. **

TRIBE Gyrinini Latreille, 1810.

Genus *Gyrinus* Geoffroy, 1762.

*Gyrinus rugifer* Régimbart, 1883. **

TRIBE Orectochilini Régimbart, 1882.

Genus *Gyretes* Brullé, 1835.

*Gyretes morio* Aubé, 1838. **

FAMILY Haliplidae Aubé, 1836.

Genus *Haliplus* Latreille, 1802.

*Haliplus gravidoides* Vondel and Spangler, 2008.

*Haliplus gravidus* Aubé, 1838.
## FAMILY Noteridae Thomson, 1860.

## SUBFAMILY Noterinae Thomson, 1860.

### TRIBE Noterini Thomson, 1860.

Genus *Canthydrus* Sharp, 1882.

*Canthydrus subsignatus* Sharp, 1882.

Genus *Hydrocanthus* Say, 1823.

*Hydrocanthus advena* Sharpe, 1882.

Genus *Suphis* Aubé, 1836.

*Suphis cimicoides* Aubé, 1836.

Genus *Suphisellus* Crotch, 1873.

*Suphisellus binotatus* (Fleutiaux and Sallé, 1889). ***

*Suphisellus nigrinus* (Aubé, 1838).

### TRIBE Pronoterini Nilsson, 2005.

Genus *Pronoterus* Sharp, 1882.

*Pronoterus obscuripennis* Fleutiaux and Sallé, 1889. ***

## SUBFAMILY Notomicrinae Zimmermann, 1919.

### TRIBE Notomicrini Zimmermann, 1919.

Genus *Notomicrus* Sharp, 1882

*Notomicrus chailliei* Manuel, 2015. ***

*Notomicrus femineus* Manuel, 2015. ***

*Notomicrus sabrouxi* Manuel, 2015. ***

*Notomicrus sharpi* Balfour-Browne, 1939.

## FAMILY Dytiscidae Leach, 1815.

## SUBFAMILY Colymbetinae Erichson, 1837.

### TRIBE Colymbetini Erichson, 1837.

Genus *Meridiorhantus* Balke, Hajek and Hendrich, 2017.

*Meridiorhantus calidus* (Fabricius, 1792).

## SUBFAMILY Copelatinae Branden, 1885.

### TRIBE Copelatini Branden, 1885.

Genus *Copelatus* Erichson, 1832.

*Copelatus guadelupensis* Legros, 1948. ***

*Copelatus insulanus* Guignot, 1939. ***

*Copelatus posticatus* (Fabricius, 1801).

*Copelatus vitraci* Legros, 1948.

## SUBFAMILY Laccophilinae Gistel, 1848.

### TRIBE Laccophilini Gistel, 1848.

Genus *Laccomimus* Toledo and Michat, 2015.

*Laccomimus bordoni* Toledo and Michat, 2015. Manuel (2017).

Genus *Laccophilus* Leach, 1815.

*Laccophilus proximus* Say, 1823.

*Laccophilus subsignatus* Sharp, 1882.
SUBFAMILY Hydrodytinae Miller, 2001.

Genus *Hydrodytes* Miller, 2001.

*Hydrodytes opalinus* (Zimmermann, 1921). Scheers and Manuel (2019).

SUBFAMILY Hydroporinae Aubé, 1836.

TRIBE Bidessini Sharp, 1880.

Genus *Hydrovatus* Motschulsky, 1853.

*Hydrovatus caraibus* Sharp, 1882.

*Hydrovatus pustulatus* (Melsheimer, 1844).

TRIBE Hyphydrini Gistel, 1848.

Genus *Pachydrus* Sharp, 1882.

*Pachydrus globosus* (Aubé, 1838).

TRIBE Methlini Branden, 1885

Genus *Celina* Aubé, 1837.

*Celina dufaui* Legros, 1948. **

TRIBE Vatellini Sharp, 1880.

Genus *Derovatellus* Sharp, 1882.

*Derovatellus lentus* (Wehncke, 1876). Manuel (2015).

SUBFAMILY Dytiscinae Leach, 1815.

TRIBE Hydaticini Sharp, 1882.

Genus *Hydaticus* Leach, 1817.

*Hydaticus rimosus* Aubé, 1838.

TRIBE Aciliini Thomson, 1867.

Genus *Thermonectus* Dejean, 1833.

*Thermonectus basillaris* (Harris, 1829).

*Thermonectus circumscriptus* (Latreille, 1809).

*Thermonectus margineguttatus* (Aubé, 1838).

TRIBE Eretini Crotch, 1873.

Genus *Eretes* Laporte de Castelnau, 1833.

*Eretes griseus* (Fabricius, 1781).

*Eretes occidentalis* Erichson, 1847.

TRIBE Cybistrini Sharp, 1882.

Genus *Megadytes* Sharp, 1882.

*Megadytes* (*Bifurcitus)* *giganteus* (Laporte de Castelnau, 1835). I

SUBORDER POLYPHAGA Emery, 1886.

SUPERFAMILY Nosodendroidea Erichson, 1846.

FAMILY Nosodendridae Erichson, 1846.

Genus *Nosodendron* Latreille, 1804.

*Nosodendron punctatostriatum* Chevrolat, 1864. *
INFRAORDER Staphyliniformia.

SUPERFAMILY Hydrophiloidea Latreille, 1802.

FAMILY Hydrophilidae Latreille, 1802.

SUBFAMILY Hydrophilinae Latreille, 1802.

TRIBE Berosini Mulsant, 1844.

Genus *Berosus* Leach, 1817.

*Berosus interstitialis* Knisch, 1924.

*Berosus undatus* (Fabricius, 1792).

Genus *Derallus* Sharp, 1882.

*Derallus rudis* Sharp, 1887.

TRIBE Anacaenini Hansen, 1991

Genus *Paracymus* Thomson, 1867.

*Paracymus delatus* Wooldridge, 1971. ***

TRIBE Hydrophilini Latreille, 1802.

Genus *Enochrus* Thomson, 1859.

*Enochrus aequalis* (Sharp, 1882).

*Enochrus bartletti* Short, 2004. *

Genus *Helochares* Mulsant, 1844.

*Helochares abbreviatus* (Fabricius, 1801).

*Helochares guadelupensis* d'Orchymont, 1926. ***

Genus *Hydrobiomorpha* Blackburn, 1888.

*Hydrobiomorpha phallica* (d'Orchymont, 1928).

Genus *Hydrobius* Leach, 1815.

*Hydrobius armatus* Sharpe, 1882.

Genus *Hydrophilus* O. F. Müller, 1764.

*Hydrophilus insularis* Laporte de Castelnau, 1840.

Genus *Tropisternus* Solier, 1834.

Subgenus *Pristoternus* Orchymont, 1936.

*Tropisternus chalybeus* Castelnau, 1840.

*Tropisternus lateralis* (Fabricius, 1775). I

SUBFAMILY Sphaeridiinae Latreille, 1802.

TRIBE Coelostomatini Hayden, 1891.

Genus *Dactylosternum* Wollaston, 1854.

*Dactylosternum abdominale* (Fabricius, 1792). I

Genus *Phaenonotum* Sharp, 1882.

*Phaenonotum exstriatum* (Say, 1835).

TRIBE Omicrini Smetana, 1975.

Genus *Omicrus* Sharp, 1879.

*Omicrus subopacus* Smetana, 1975. ***
TRIBE Megasternini Mulsant, 1844.

Genus *Cercyon* Leach, 1817.

*Cercyon cribratum* Laporte de Castelnau, 1840. ***

*Cercyon nigriceps* (Marsham, 1802). I

Genus *Pelosoma* Mulsant, 1844.

*Pelosoma rufipes* (Fleutiaux and Sallé, 1889). ***

FAMILY Histeridae Gyllenhaal, 1808.

SUBFAMILY Abraeinae MacLeay, 1819.

TRIBE Abraeini MacLeay, 1819.

Genus *Abraeus* Leach, 1817.

*Abraeus dufaui* Desbordes, 1914. ***

TRIBE Acritini Wenzel, 1944.

Genus *Aeletes* Horn, 1873.

*Aeletes gulliver* (Marseul, 1856).

SUBFAMILY Trypanaeinae Marseul, 1857.

Genus *Trypanaeus* Eschscholtz, 1829.

*Trypanaeus flavipennis* Marseul, 1856.

*Trypanaeus luteivestris* Marseul, 1860.

SUBFAMILY Saprinaeae Blanchard, 1845.

Genus *Euspilotus* Lewis, 1907.

*Euspilotus azurescens* (Marseul, 1855).

Subgenus *Hesperosaprinus* Wenzel in Arnett, 1962.

*Euspilotus* (*Hesperosaprinus*) *azureus* (Sahlberg, 1823).

*Euspilotus* (*Hesperosaprinus*) *insularis* (Marseul, 1855). ***

SUBFAMILY Dendrophilinae Reitter, 1909.

TRIBE Bacaniini Kryzhanovskij, 1976.

Genus *Bacanius* LeConte, 1853.

*Bacanius ferrugineus* Bickhardt, 1918. ***

TRIBE Paromalini Reitter, 1909.

Genus *Carcinops* Marseul, 1855.

*Carcinops troglodytes* (Paykull, 1811). I

SUBFAMILY Histerinae Gyllenhal, 1808.

TRIBE Omalodini Kryzhanovskij, 1972

Genus *Omalodes* Dejean, 1833.

*Omalodes laevinotus* Marseul, 1853.

TRIBE Exosternini Bickhardt, 1914.

Genus *Atholus* Thomson, 1859.

*Atholus bimaculatus* (Linnaeus, 1758). I

*Atholus confinis* (Erichson, 1834). I
TRIBE Histerini Gyllenhal, 1808.

Genus *Hister* Linnaeus, 1758.
*Hister servus* Erichson, 1834.

SUPERFAMILY Staphylinoidea Latreille, 1802.

FAMILY Jacobsoniidae Heller, 1926.

Genus *Derolathrus* Sharp, 1908.
*Derolathrus sharpi* Grouvelle and Raffray, 1912.

FAMILY Hydraenidae Mulsant, 1844.

SUBFAMILY Hydraeninae Mulsant, 1844.

TRIBE Hydraenini Mulsant, 1844.

Genus *Hydraena* Kugelann, 1794.
*Hydraena guadelupensis* d'Orchymont, 1923.
*Hydraena insularis* d'Orchymont, 1945. **
*Hydraena phallica* d'Orchymont, 1930.

SUBFAMILY Ochthebiinae Thomson, 1859.

TRIBE Ochthebiini Thomson, 1859.

Genus *Ochthebius* Leach, 1815.
*Ochthebius attritus* LeConte, 1878.

FAMILY Ptiliidae Erichson, 1845.

SUBFAMILY Acrotrichinae Reitter, 1909.

TRIBE Acrotrichini Reitter, 1909.

Genus *Acrotrichis* Motschulsky, 1848.
*Acrotrichis rufescens* (Matthews, 1894).

TRIBE Nphanini Portevin, 1929.

Genus *Nphanes* Thomson, 1859.
*Nphanes meridionalis* Matthews, 1872.

FAMILY Leiodidae Fleming, 1821.

SUBFAMILY Leiodinae Fleming, 1821.

TRIBE Leiodini Fleming, 1821.

Genus *Zeadolopus* Broun, 1903.
*Zeadolopus parvantilliensis* Peck and Cook, 2014.

TRIBE Pseudoleiodini Portevin, 1926.

Genus *Aglyptinus* Cockerell, 1906.
*Aglyptinus guadelupensis* Portevin, 1942. ***
*Aglyptinus kaszabi* Hlisnikovský, 1964. ***

FAMILY Staphylinidae Latreille, 1802.

SUBFAMILY Aleocharinae Fleming, 1821.

TRIBE Hoplandrini Fenyes 1920.

Genus *Acantoxyura* Pace, 1987.
*Acantoxyura armata* Pace, 1987.
*Acantoxyura spinifera* Pace, 1987. ***
Genus *Hoplandria* Kraatz, 1857.
*Hoplandria guadeloupensis* Pace, 1987. ***

TRIBE Lomechusini Fleming, 1821.

Genus *Apalonia* Casey, 1906.
*Apalonia semiscapa* (Pace, 1987).

Genus *Macrogerodonia* Bernhauer, 1951.
*Macrogerodonia cursoria* Pace, 1987. **
*Macrogerodonia trichonota* Pace, 1987. **

Genus *Meronera* Sharp, 1887.
*Meronera albicincta* (Erichson, 1839).
*Meronera fulvicornis* Pace, 1991. Peck (2016).

TRIBE Autaliini Thomson 1859.

Genus *Eudera* Fauvel, 1865.
*Eudera didyma* (Erichson, 1839).

TRIBE Placusini Mulsant and Rey, 1871.

Genus *Euvira* Sharp, 1883.
*Euvira conifera* Pace, 1987. ***
*Euvira laeviuscula* Pace, 1987. ***
*Euvira maculata* Pace, 1987. ***

TRIBE Oxypodini Thomson, 1859.

Genus *Feluva* Blackwelder, 1952.
*Feluva guadalupensis* Pace, 1987. ***

TRIBE Athetini Casey, 1910.

Genus *Heterostiba* Pace, 1983.
*Heterostiba antillarum* Pace, 1987.

Genus *Leptonia* Sharp, 1883.
*Leptonia guadalupensis* Pace, 1987. **
*Leptonia megalomera* Pace, 1987. ***

Genus *Stethusa* Casey, 1910.
*Stethusa dichroa* (Gravenhorst, 1802).
*Stethusa lurida* (Erichson, 1839).

Genus *Plesiomalota* Pace, 1983.
*Plesiomalota tenella* Pace, 1987. ***

TRIBE Aleocharini Fleming, 1821.

Genus *Acrotona* Thomson, 1859.
*Acrotona flavoterminata* (Cameron, 1923). *
*Acrotona semilacera* (Pace, 1996). ***

Genus *Aleochara* Gravenhorst, 1802.
*Aleochara bilineata* Gyllenhal, 1810. I
*Aleochara notula* Erichsson, 1839.
*Aleochara puberula* Klug, 1832. I
*Aleochara taeniata* Erichson, 1839.

Genus *Atheta* Thomson, 1858.

*Atheta alternata* (Erichson, 1839).

*Atheta conformis* (Erichson, 1839).

*Atheta nigripennis* (Erichson, 1839). *

*Atheta nigripennis rufa* Pace, 1985.

TRIBE Diestotini Mulsant and Rey, 1871.

Genus *Diestota* Mulsant and Rey, 1870.

*Diestota cliens* Pace, 1987. ***

*Diestota didyma* (Erichson, 1840).

*Diestota fasciata* Pace, 1987. ***

*Diestota guadalupensis* Pace, 1987. ***

*Diestota laesicollis* (Erichson, 1840).

*Diestota laxiventris* Pace, 1987. ***

*Diestota melanura* (Erichson, 1839). *

*Diestota sperata* Sharp, 1876.

TRIBE Homalotini Heer, 1839.

Genus *Coenonica* Kraatz, 1857.

*Coenonica puncticollis* Kraatz, 1857.

Genus *Gyrophaena* Mannerheim, 1830.

*Gyrophaena guadelupensis* Pace, 1987. ***

Genus *Heterota* Mulsant and Rey, 1874.

*Heterota plumbea* (Waterhouse, 1858).

Genus *Thecturota* Casey, 1893.

*Thecturota antillarum* Pace, 1987. ***

TRIBE Hypocyptini Laporte de Castelnau, 1835.

Genus *Holobus* Solier, 1849.

*Holobus albidicornis* (Bernhauer, 1923). ***

*Holobus guadeloupeae* (Frank, 1972). **

*Holobus hypocyptina* (Bernhauer, 1923). ***

Genus *Oligota* Mannerheim, 1831.

*Oligota parva* Kraatz, 1862.

TRIBE Oxypodini Thomson, 1859.

Genus *Phloeopora* Erichson, 1837.

*Phloeopora religata* Erichson, 1839. ***

TRIBE Phytosini Thomson, 1867.

Genus *Phytosus* Curtis, 1838.

*Phytosus caribeanus* Haghebaert, 1993. ***

Genus *Placusa* Erichson, 1837.

*Placusa basiventris* Pace, 1987. ***

*Placusa lateralis* Pace, 1987. ***

*Placusa oblita* Pace, 1987. ***

*Placusa praepes* Pace, 1987. ***
*Placusa subtilis* Pace, 1987. ***

*Placusa trivialis* Pace, 1987. ***

### TRIBE Pronomaeini Mulsant and Rey, 1873.

Genus *Myllaena* Erichson, 1837.

*Myllaena guadalupensis* Pace, 1987. ***

Genus *Pronomaea* Erichson, 1837.

*Pronomaea debilis* Cameron, 1922. *

### SUBFAMILY Osoriinae Erichson, 1839.

### TRIBE OsoriinI Erichson, 1839.

Genus *Mimogonus* Fauvel, 1903.

*Mimogonus fumator* (Fauvel, 1889). I

### TRIBE Thoracophorini Reitter, 1909.

Genus *Clavilispinus* Bernhauer, 1926.

*Clavilispinus exiguus* (Erichson, 1840).

*Clavilispinus guadeloupensis* Irmler, 1991.

*Clavilispinus megacephalus* (Fauvel, 1865).

*Clavilispinus politus* (Sharp, 1887).

Genus *Espeson* Schaufuss, 1882.

*Espeson euplectoides* Fauvel, 1902.

*Espeson moratus* Schaufuss, 1882.

Genus *Glyptoma* Erichson, 1840.

*Glyptoma guadelupensis* (Wendeler, 1930). ***

Genus *Lispinodes* Sharp, 1880.

*Lispinodes nitens* (Fauvel, 1902).

Genus *Lispinus* Erichson, 1839.

*Lispinus attenuatus* Erichson, 1840.

*Lispinus cordobensis* Bernhauer, 1929.

*Lispinus insularis* Fauvel, 1863.

*Lispinus linearis* Erichson, 1840.

Genus *Nacaeus* Blackwelder, 1942.

*Nacaeus danforthi* (Blackwelder, 1943).

*Nacaeus flavipennis* (Fauve, 1865).

*Nacaeus guadeloupeae* (Blackwelder, 1943). ***

*Nacaeus impressicollis* (Motschulsky, 1857).

*Nacaeus nigrifrons* (Fauvel, 1863).

*Nacaeus planellus* (Sharp, 1887).

*Nacaeus sculpturatus* (Sharp, 1887).

*Nacaeus sulciger* Irmler, 2003.

Genus *Pseudespeson* Lecoq, 1994.

*Pseudespeson crassulus* (Fauvel, 1902). **

Genus *Tannea* Blackwelder, 1952.

*Tannea tenella* (Erichson, 1840).
Genus *Thoracophorus* Motschulsky, 1837.

*Thoracophorus brevicristatus* (Horn, 1871). I

*Thoracophorus exilis* (Erichson, 1840). *

*Thoracophorus filum* Sharp, 1887. Irmler (2015).

*Thoracophorus guadalupensis* Cameron, 1913.

*Thoracophorus sculptilis* (Erichson, 1840).

*Thoracophorus simplex* Wendeler, 1930. *

SUBFAMILY Oxytelinae Fleming, 1821.

TRIBE Oxytelini Fleming, 1821.

Genus *Parosus* Sharp, 1887.

*Parosus antillarum* Wendeler, 1928. ***

Genus *Trogactus* Sharp, 1887.

*Trogactus cornucopius* (Blackwelder, 1943). **

Genus *Anotylus* Thomson, 1859.

*Anotylus insignitus* (Gravenhorst, 1806). I

Genus *Apocellus* Erichson, 1840.

*Apocellus ustulatus* Erichson, 1840. *

Genus *Oxytelus* Gravenhorst, 1802.

*Oxytelus incisus* Motschulsky, 1857. I

Genus *Platystethus* Mannerheim, 1830.

*Platystethus spiculus* Erichson, 1840. I

TRIBE Thinobiini Sahlberg, 1876.

Genus *Bledius* Leach *in* Samouelle, 1819.

*Bledius caribbeanus* Blackwelder, 1943.

Genus *Carpelimus* Kirby, 1819.

*Carpelimus beattyi* Blackwelder, 1943. *

*Carpelimus correctus* Blackwelder, 1943.

*Carpelimus flavipes* (Erichson, 1840).

*Carpelimus fulvipes* (Erichson, 1840).

*Carpelimus imitator* (Bierig, 1935). *

Genus *Thinodromus* Kraatz, 1857.

*Thinodromus croceipes* (Fauvel, 1868). *

SUBFAMILY Paederinae Fleming, 1821.

TRIBE Paederini Fleming, 1821.

SUBTRIBE Cryptobiina Casey, 1905.

Genus *Biocrypta* Casey, 1905.

*Biocrypta centralis* (Sharp, 1885).

*Biocrypta fulvipes* (Erichson, 1840).

Genus *Ochthephilum* Stephens, 1829.

*Ochthephilum vitraci* Bernhauer, 1916. ***
TRIBE Lathrobiini Laporte de Castelnau, 1835.
SUBTRIBE Medonina Casey, 1905.

Genus *Hypomedon* Mulsant and Rey, 1878.
*Hypomedon debilicornis* (Wollaston, 1857). I

Genus *Lithocharis* Boisduval and Lacordaire, 1835.
*Lithocharis dorsalis* Erichson, 1840.
*Lithocharis infuscata* Erichson, 1840.
*Lithocharis limbata* Erichson, 1840.
*Lithocharis ochracea* (Gravenhorst, 1802). I
*Lithocharis secunda* Blackwelder, 1943. *
*Lithocharis sororcula* Kraatz, 1859. I
*Lithocharis vilis* Kraatz, 1859.

Genus *Stilomedon* Sharp, 1886.
*Stilomedon connexus* (Sharp, 1876). *

SUBTRIBE Scopaeina Mulsant and Rey, 1878.

Genus *Scopaeus* Erichson, 1840.
*Scopaeus antennalis* Cameron, 1913. *

SUBTRIBE Stilicopsina Casey, 1905.

Genus *Stamnoderus* Sharp, 1886.
*Stamnoderus labeo* (Erichson, 1840). *

TRIBE Pinophilini Normann, 1837.

Genus *Palaminus* Erichson, 1840.
*Palaminus apterus* Bernhauer, 1918. ***
*Palaminus dubius* Notman, 1929. ***
*Palaminus insularis* Cameron, 1913. *
*Palaminus variabilis* Erichson, 1840.

Genus *Pinophilus* Gravenhorst, 1802.
*Pinophilus insigniventris* Bernhauer, 1918. ***

SUBFAMILY Piestinae Erichson, 1839.

Genus *Piestus* Gravenhorst, 1806.
*Piestus capricornis* Laporte de Castelnau, 1835.
*Piestus fulvipes* Erichson, 1840.
*Piestus penicillatus* (Dalman, 1821).
*Piestus pygmaeus* Laporte de Castelnau, 1835.

SUBFAMILY Pselaphinae Latreille, 1802.
TRIBE Brachyglutini Raffray, 1904.

Genus *Eupsenius* LeConte, 1849.
*Eupsenius politus* Reitter, 1883. *

Genus *Reichenbachia* Leach, 1826.
*Reichenbachia guadalupensis* Raffray in Grouvelle and Raffray, 1908. ***

Genus *Scalenarthrus* LeConte, 1880.
*Scelenarthrus guadelupensis* Raffray in Grouvelle and Raffray, 1912. ***
*Scelenarthrus pectinicornis* Raffray, 1904. **
**TRIBE Jubini Raffray, 1904.**

Genus *Jubus* Schaufuss, 1872.
*Jubus insularis* Raffray in Grouvelle and Raffray, 1908. ***

**TRIBE Euplectini Streubel, 1839.**

Genus *Euplectus* Leach, 1817.
*Euplectus insularis* Raffray in Grouvelle and Raffray, 1908. ***

**TRIBE Iniocyphini Park, 1951.**

Genus *Bythinophysis* Raffray, 1908.
*Bythinophysis humilis* (Raffray in Grouvelle and Raffray, 1908). ***

**TRIBE Pselaphini Latreille, 1802.**

Genus *Neopselaphus* Jeannel, 1951.
*Neopselaphus chalumeaui* Besuchet, 1987. ***

**TRIBE Trichonychini Reitter, 1882.**

Genus *Biblomimus* Raffray, 1904.
*Biblomimus impressa* (Raffray in Grouvelle and Raffray, 1908). ***

Genus *Melba* Casey, 1897.
*Melba crassipes* Raffray in Grouvelle and Raffray, 1908. ***
*Melba fleutiauxi* Raffray, 1890. ***
*Melba frontalis* Raffray in Grouvelle and Raffray, 1908. ***

**TRIBE Trogastrini Jeannel, 1949.**

Genus *Rhexinia* Raffray, 1890.
*Rhexinia versicolor* Raffray in Grouvelle and Raffray, 1908. ***

**TRIBE Tyrini Reitter, 1882.**

Genus *Hamotus* Aubé, 1844.
*Hamotus hirtus* Raffray, 1905. **

**SUBFAMILY Scaphidiinae Latreille, 1806.**

**TRIBE Scaphidiini Latreille, 1806.**

Genus *Baeocera* Erichson, 1845.
*Baeocera dufaui* Pic, 1920. ***

**SUBFAMILY Scydmaeninae Leach, 1815.**

**TRIBE Cyrtoscydmini L.W. Schaufuss, 1889.**

Genus *Euconnus* Thomson, 1859.
*Euconnus gouadeloupensis* Franz, 1980. ***

**TRIBE Scydmaenini Leach, 1815.**

Genus *Phloeonomus* Heer, 1838.
*Phloeonomus pedicularius* (Erichson, 1840).

Genus *Scydmaenus* Latreille, 1802.
*Scydmaenus gouadeloupensis* Franz, 1984. *
SUBFAMILY Staphylininae Latreille, 1802.

TRIBE Diochini Casey, 1906.

Genus *Diochus* Erichson, 1839.

*Diochus nanus* Erichson, 1839.

TRIBE Staphylinini Latreille, 1802.

Genus *Belonuchus* Nordmann, 1837.

*Belonuchus bugnioni* Fauvel, 1901. **

Genus *Cafius* Stephens, 1829.

*Cafius bistriatus* (Erichson, 1840).

*Cafius caribeanus* Bierig, 1934.

*Cafius subtilis* Cameron, 1922.

Genus *Gabronthus* Tottenham, 1955.

*Gabronthus thermarum* (Aubé, 1850).

Genus *Philonthus* Stephens, 1829.

*Philonthus discoideus* (Gravenhorst, 1802).

*Philonthus hepaticus* Erichson, 1840.

*Philonthus longicornis* Stephens, 1832.

*Philonthus ventralis* (Gravenhorst, 1802).

TRIBE Xantholinini Erichson, 1839.

Genus *Neohypnus* Coiffait and Sáiz, 1964.

*Neohypnus attenuatus* (Erichson, 1839). I

*Neohypnus illucens* (Erichson, 1839).

Genus *Xantholinus* Dejean, 1821.

*Xantholinus solitarius* Blackwelder, 1943. ***

SUBFAMILY Steninae MacLeay, 1825.

Genus *Stenus* Latreille, 1797.

*Stenus antillensis* Benick, 1917.

SUBFAMILY Tachyporinae MacLeay, 1825.

TRIBE Tachyporini MacLeay, 1825.

Genus *Cilea* Jacquelin du Val, 1856.

*Cilea silphoides* (Linnaeus, 1767). I

Genus *Sepedophilus* Gistel, 1856.

*Sepedophilus interruptus* (Erichson, 1839).

INFRAORDER Scarabaeiformia Crowson, 1960.

SUPERFAMILY Scarabaeoidea Latreille, 1802.

FAMILY Passalidae Leach, 1815.

SUBFAMILY Passalinae Leach, 1815.

TRIBE Passalini Leach, 1815.

Genus *Passalus* Fabricius, 1792.

*Passalus unicornis* Lepeltier and Audinet-Serville, 1825. **
Genus *Spasalus* Kaup in Harold, 1869.

*Spasalus puncticollis* (Lepeletier de Saint Fargeau and Audinet-Serville, 1825). Boucher et al. (2015). **

## FAMILY Trogidae MacLeay, 1819.

Genus *Omorgus* Erichson, 1847.

*Omorgus suberosus* (Fabricius, 1775). Meurgey (2017d).

## FAMILY Hybosoridae Erichson, 1847.

### SUBFAMILY Ceratocanthinae Martínez, 1968.

### TRIBE Ceratocanthini Martinez, 1968.

Genus *Ceratocanthus* White, 1842.

*Ceratocanthus bonfilsi* Chalumeau, 1977. ***

Genus *Cloeotus* Germar, 1843.

*Cloeotus pyritosus* Erichson, 1843.

Genus *Germarostes* Paulian, 1982.

*Germarostes allorgei* (Paulian, 1947). **

*Germarostes pauliani* (Chalumeau and Cambefort, 1976). ***

## FAMILY Scarabaeidae Latreille, 1802.

### SUBFAMILY Aphodiinae Leach, 1815.

### TRIBE Aphodiini Leach, 1815.

Genus *Calamosternus* Motschulsky, 1859.

*Calamosternus granarius* (Linnaeus, 1767).

Genus *Labarrus* Mulsant and Rey, 1870.

*Labarrus lividus* (Olivier, 1789). I

*Labarrus pseudolividus* (Balthasar, 1941). I.

Genus *Nialaphodius* Kolbe, 1908.

*Nialaphodius nigrita* (Fabricius, 1801). I

## TRIBE Eupariini Schmitt, 1910.

Genus *Ataeniopsis* Petrovitz, 1973.

*Ataeniopsis armasi* Chalumeau, 1982. *

Genus *Ataenius* Harold, 1867.

*Ataenius aequalis* Chapin, 1940.

*Ataenius armasi* Chalumeau, 1982.

*Ataenius beattyi* Chapin, 1940. *

*Ataenius brevicollis* (Wollaston, 1854).

*Ataenius carinator* Harold, 1874.

*Ataenius cartwrighti* Chalumeau and Gruner, 1974. ***

*Ataenius elongatus* Beauvoir, 1805.

*Ataenius gracilis* (Melsheimer, 1844).

*Ataenius haroldi* Steinheil, 1872.

*Ataenius hygrophilus* Paulian, 1947. ***

*Ataenius imbricatus* Melsheimer, 1844.

*Ataenius insulae* Chalumeau and Gruner, 1974. ***

*Ataenius liogaster* Bates, 1887.

*Ataenius luteomargo* Chapin, 1940.

*Ataenius morator* Harold, 1869.
*Ataenius picinus* Harold, 1867.

*Ataenius scabrellus* Schmidt, 1909.

*Ataenius scutellaris* Harold, 1867.

*Ataenius stercorator* (Fabricius, 1775).

*Ataenius strigicaudus* Bates, 1887.

*Ataenius temperei* Chalumeau and Gruner, 1974. **

Genus *Euparia* Lepeletier de Saint Fargeau and Audinet-Serville, 1828.

*Euparia baraudi* Chalumeau and Gruner, 1974. **

Genus *Iguazua* Stebnicka, 1997.

*Iguazua blackwelderi* (Chapin, 1940).

Genus *Saprosites* Redtenbacher, 1858.

*Saprosites dufaui* Paulian, 1947. **

*Saprosites exaratus* (Fleutiaux and Sallé, 1889). **

TRIBE Psammodiini Mulsant, 1842.

Genus *Psammodius* Fallén, 1807.

*Psammodius cameneni* Chalumeau, 1976. *

*Psammodius viti* Chalumeau, 1983. ***

SUBTRIBE Rhysesemina Pittino and Mariani, 1986.

Genus *Platytomus* Mulsant, 1842.

*Platytomus parvulus* (Chevrolat, 1864).

TRIBE Rhyparini Schmidt, 1910

Genus *Rhyparus* Westwood, 1845

*Rhyparus spilmani* Cartwright and Chalumeau, 1977. **

**Recent record.** Guadeloupe, Vieux-Habitants, La Grivelière, 03.II.2018, 365977 – 1777514, 183m, F. Meurgey leg (coll. SHNLH).

SUBFAMILY Scarabaeinae Latreille, 1802.

TRIBE Canthonini Péringuey, 1901.

Genus *Pseudocanthon* Bates, 1887.

*Pseudocanthon caeranus* Matthews, 1966. **

*Pseudocanthon vitraci* (Fleutiaux and Sallé, 1889). ***

TRIBE Ateuchini Martinez, 1951.

Genus *Ateuchus* Weber, 1801.

*Ateuchus illaesum* (Harold, 1868).

*Ateuchus insulare* (Fleutiaux and Sallé, 1889). ***

Genus *Canthidium* Erichson, 1847.

*Canthidium melanocephalum* Olivier, 1789.

Genus *Uroxys* Westwood, 1842.

*Uroxys productus* (Fleutiaux and Sallé, 1889). ***

TRIBE Onthophagini Lacordaire, 1856.

Genus *Digitonthophagus* Balthasar, 1959.

*Digitonthophagus gazella* (Fabricius, 1787). I
Genus *Onthophagus* Latreille, 1802.
*Onthophagus antillarum* Arrow, 1903. **

SUBFAMILY Orphninae Erichson, 1847.
TRIBE Aegidiini Paulian, 1984.

Genus *Aegidium* Westwood, 1846.
*Aegidium chalumeaui* Rojkoff and Frolov, 2017.***
*Aegidium parvulum* Westwood, 1846. ***

SUBFAMILY Melolonthinae Leach, 1819.
TRIBE Melolonthini Leach, 1819.

Genus *Lachnosterna* Hope, 1837.
*Lachnosterna guadulpensis* Blanchard, 1837.

Genus *Phyllophaga* Harris, 1827.
*Phyllophaga denticulata* (Blanchard, 1850).
*Phyllophaga fuscipennis* (Moser, 1918). ***
*Phyllophaga mariaegalantae* Chalumeau and Gruner, 1976. ***
*Phyllophaga patrueloides* Paulian, 1947. ***
*Phyllophaga pauliani* Chalumeau and Gruner, 1976. ***
*Phyllophaga plaei* (Blanchard, 1850). *
*Phyllophaga sandersoniella* Chalumeau and Gruner, 1976. ***

SUBFAMILY Rutelinae MacLeay, 1819.
TRIBE Anomalini Blanchard, 1851.

Genus *Anomala* Samouelle, 1819.
*Anomala insularis* (Castelnau, 1840). **
*Anomala luciae* Blanchard, 1850. *

TRIBE Rutelini MacLeay, 1819.

Genus *Anticheira* Eschscholtz, 1818.
*Anticheira lucida* Olivier, 1789.

Genus *Cnemida* Kirby, 1827.
*Cnemida retusa* (Fabricius, 1801).

Genus *Macraspis* MacLeay, 1819.
*Macraspis tristis* Castelnau, 1840. **

Genus *Rutela* Latreille, 1802.
*Rutela striata striata* (Olivier, 1789). **

TRIBE Geniatini Burmeister, 1844.

Genus *Leucothyreus* MacLeay, 1819.
*Leucothyreus guadulpiensis* Burmeister, 1844. **

SUBFAMILY Dynastinae MacLeay, 1819.
TRIBE Cyclocephalini Laporte de Castelnau, 1840.

Genus *Cyclocephala* Dejean, 1821.
*Cyclocephala immaculata* (Olivier, 1789). Ratcliffe and Cave (2015).
*Cyclocephala insulicola* Arrow, 1937. Ratcliffe and Cave (2015).***
*Cyclocephala mafaffa* Burmeister, 1847. Ratcliffe and Cave (2015).
* *Cyclocephala melanocephala* (Fabricius, 1775). Ratcliffe and Cave (2015).

* *Cyclocephala tridentata* (Fabricius, 1801). Ratcliffe and Cave (2015).

Genus *Dyscinetus* Harold, 1869.

* *Dyscinetus questeli* Chalumeau, 1983. Ratcliffe and Cave (2015).***

**TRIBE Dynastini MacLeay, 1819.**

Genus *Dynastes* Kirby, 1825.

* *Dynastes hercules hercules* (Linnaeus, 1758). Huang (2014); Ratcliffe and Cave (2015). **

**TRIBE Oryctini Mulsant, 1842.**

Genus *Strategus* Kirby, 1828.

* *Strategus syphax* (Fabricius, 1775). Ratcliffe and Cave (2015). **

**TRIBE Pentodontini Mulsant, 1842.**

Genus *Tomarus* Erichson, 1847.

* *Tomarus cuniculus* (Fabricius, 1801). Ratcliffe and Cave (2015). *Tomarus ebenus* (Degeer, 1774). Ratcliffe and Cave (2015).

**TRIBE Phileurini Burmeister, 1847.**

Genus *Hemiphileurus* Kolbe, 1910.

* *Hemiphileurus laeviceps* (Arrow, 1947). Ratcliffe and Cave (2015). **

Genus *Phileurus* Latreille, 1807.

* *Phileurus didymus* Linnaeus, 1758.

* *Phileurus valgus* Olivier, 1789. Sautière (2014). Ratcliffe and Cave (2015).

SUBFAMILY Cetoniinae Leach, 1815.

**TRIBE Cetoniini Leach, 1815.**

Genus *Protaetia* Burmeister, 1842.

* *Protaetia fusca* (Herbst, 1790). Meurgey (2017a). I

**TRIBE Gymnetini Kirby, 1827.**

Genus *Gymnetis* MacLeay, 1819.

* *Gymnetis lanius* (Linnaeus, 1766). *

**INFRAORDER Elateriformia Crowson, 1960.**

**SUPERFAMILY Scirtoidea Fleming, 1821.**

**FAMILY Scirtidae Fleming, 1821.**

Genus *Cyphon* Paykull, 1799.

* *Cyphon degoutteti* Pic, 1918. ***

Genus *Microcara* Thomson, 1859.

* *Microcara dufaui* Legros, 1947. ***

Genus *Ora* Clark, 1865.

* *Ora dufaui* Legros, 1947. ***

Genus *Scirtes* Illiger, 1807.

* *Scirtes dufaui* Pic, 1916. ***
*Scirtes insularis* Champion, 1897. **

*Scirtes testaceus* Fabricius, 1801.

**SUPERFAMILY Buprestoidea Leach, 1815.**

**FAMILY Buprestidae Leach, 1815.**

**SUBFAMILY Polycestinae Lacordaire, 1857.**

**TRIBE Haplostethini LeConte, 1861.**

Genus *Micrasta* Kerremans, 1893.

*Micrasta creola* Obenberger, 1936. ***

*Micrasta gyleki* Obenberger, 1917. ***

*Micrasta pygmaeola* Obenberger, 1917. ***

*Micrasta strandi* Obenberger, 1936. ***

**TRIBE Acmaeoderini Kerremans, 1893.**

Genus *Acmaedera* Eschscholtz, 1829.

*Acmaedera flavomarginata* (Gray, 1832). I

*Acmaedera villiersi* Descarpentries, 1981. **

**TRIBE Polycestini Lacordaire, 1857.**

Genus *Polycesta* Dejean, 1833.

*Polycesta depressa* (Linnaeus, 1771).

**SUBFAMILY Chrysochroinae Laporte de Castelnau, 1835.**

**TRIBE Chrysochroini Laporte de Castelnau, 1835.**

Genus *Chalcophora* Dejean, 1833.

*Chalcophora humboldti* (Laporte and Gory, 1836). ***

**TRIBE Paraleptodemini Cobos, 1975.**

Genus *Euplectalecia* Obenberger, 1924.

*Euplectalecia erythropa* (Gory, 1840). **

**SUBFAMILY Buprestinae Leach, 1815.**

**TRIBE Chrysobothrini Gory and Laporte de Castelnau, 1836.**

Genus *Chrysobothris* Eschscholtz, 1829.

*Chrysobothris guadeloupensis* Descarpentries, 1981. ***

*Chrysobothris tranquebarica* Gmelin, 1788.

**SUBFAMILY Agrilinae Laporte de Castelnau, 1835.**

**TRIBE Agrilini Laporte de Castelnau, 1835.**

Genus *Aphanisticus* Latreille, 1829.

*Aphanisticus cochinchinae seminulum* Obenberger, 1929. I

**TRIBE Tracheini Laporte de Castelnau, 1835.**

Genus *Neotrachys* Obenberger, 1923.

*Neotrachys dominicanus* Théry, 1947. Meurgey (2017b). **

*Neotrachys guadeloupensis* (Fleutiaux and Sallé, 1889). ***
Genus *Taphrocerus* Solier, 1833.
*Taphrocerus chalumeaui* Hespenheide, 1998. ***

SUPERFAMILY Byrrhoidea Latreille, 1804.

FAMILY Elmidae Curtis, 1830.

SUBFAMILY Elminae Curtis, 1830.

Genus *Hexacylloepus* Hinton, 1940.
*Hexacylloepus smithi* (Grouvelle, 1898).

SUBFAMILY Larainae LeConte, 1861

Genus *Hexanchorus* Sharp, 1882.
*Hexanchorus caraibus* (Coquerel, 1851). **

FAMILY Dryopidae Billberg, 1820.

Genus *Pelonomus* Erichson, 1847.
*Pelonomus picipes* (Olivier, 1791). ***

FAMILY Limnichidae Erichson, 1846.

SUBFAMILY Cephalobyrrhinae Champion, 1925.

Genus *Throscinus* LeConte, 1874.
*Throscinus aethiops* Darlington, 1936. *

SUBFAMILY Limnichinae Erichson, 1846.

TRIBE Limnichini Erichson, 1846.

Genus *Limnichus* Latreille, 1829.
*Limnichus sulcatulus* Pic, 1922. ***

FAMILY Heteroceridae MacLeay, 1825.

Genus *Tropicus* Pacheco, 1964.
*Tropicus arawak* Bameul, 1995. ***

FAMILY Psephenidae Lacordaire, 1854.

Genus *Psephenops* Grouvelle, 1898.
*Psephenops smithi* Grouvelle, 1898. **
*Psephenops smithi guadeloupensis* Bameul, 2001. ***

FAMILY Ptilodactylidae Laporte de Castelnau, 1836.

SUBFAMILY Ptilodactylinae Laporte de Castelnau, 1836.

Genus *Ptilodactyla* Illiger, 1807.
*Ptilodactyla emarginata* Chevrolat, 1870. *
*Ptilodactyla guadelupensis* Legros, 1947. ***
*Ptilodactyla macrophthalma* Legros, 1947. **
*Ptilodactyla militaris* Chevrolat, 1870. *

Genus *Stirophora* Champion, 1897.
*Stirophora lyciformis* (Champion, 1897).

FAMILY Chelonariidae Blanchard, 1845.

Genus *Chelonarium* Fabricius, 1801.
*Chelonarium pilosellum* Chevrolat, 1880. **
SUPERFAMILY Dascilloidea Guérin-Méneville, 1843.

FAMILY Rhipiceridae Latreille, 1834.

Genus *Callirhipis* Latreille, 1829.

*Callirhipis lherminieri* Castelnau, 1834. **

SUPERFAMILY Elateroidea Leach, 1815.

FAMILY Eucnemidae Eschscholtz, 1829.

SUBFAMILY Melasinae Fleming, 1821.

TRIBE Melasini Fleming, 1821.

Genus *Isorhipis* Lacordaire, 1835.

*Isorhipis picteti* (Bonvouloir, 1871).

TRIBE Dirhagini Reitter, 1911.

Genus *Adelothyreus* Horn, 1890.

*Adelothyreus curtus* Fleutiaux, 1907. **

*Adelothyreus dufaui* Fleutiaux, 1907. **

*Adelothyreus mouffleti* Bonvouloir, 1875. ***

Genus *Entomophthalmus* Bonvouloir, 1871.

*Entomophthalmus americanus* Bonvouloir, 1872.

Genus *Rhagomicrus* Fleutiaux, 1902.

*Rhagomicrus solitarius* Fleutiaux, 1911. ***

SUBFAMILY Macraulacinae Fleutiaux, 1923.

TRIBE Macraulacini Fleutiaux, 1923.

Genus *Dromaeolus* Kiesenwetter, 1858.

*Dromaeolus fastidiosus* Bonvouloir, 1871.

*Dromaeolus palpalis* Fleutiaux, 1911. ***

*Dromaeolus subcylindricus* Fleutiaux, 1911. ***

Genus *Fornax* Laporte de Castelnau, 1835.

*Fornax adjectus* Horn, 1890.

*Fornax guadeloupensis* Fleutiaux and Sallé, 1889.

*Fornax insitus* Horn, 1890.

Genus *Plesiofornax* Coquerel, 1866.

*Plesiofornax colonus* (Fleutiaux, 1911). ***

*Plesiofornax dufaui* Fleutiaux, 1911. **

TRIBE Nematodini Leiler, 1976.

Genus *Nematodes* Berthold, 1827.

*Nematodes biimpressus* Fleutiaux, 1911. **

*Nematodes guadelupensis* Fleutiaux, 1911. ***

FAMILY Throscidae Laporte de Castelnau, 1840.

SUBFAMILY Throscinae Laporte de Castelnau, 1840.

TRIBE Throscini Laporte de Castelnau, 1840.

Genus *Aulonothroscus* Horn, 1890.

*Aulonothroscus bicarinatus* Fleutiaux, 1911. ***
**FAMILY Elateridae Leach, 1815.**

**SUBFAMILY Elaterinae Leach, 1815.**

**TRIBE Agriotini Champion, 1894.**

Genus *Agriotes* Eschscholtz, 1829.

*Agriotes guadelupensis* Candèze, 1865. **

**TRIBE Ampedini Fleutiaux, 1947.**

Genus *Anoplischius* Candèze, 1859.

*Anoplischius brunneus* (Fleutiaux and Sallé, 1889). ***

*Anoplischius sulcifrons* Candèze, 1859. ***

Genus *Crepidius* Candèze, 1859.

*Crepidius rhipiphorus* Candèze, 1859. ***

Genus *Dicrepidius* Eschscholtz, 1829.

*Dicrepidius cavifrons* Candèze, 1881.

*Dicrepidius distinctus* Fleutiaux, 1911. **

*Dicrepidius elegans* Fleutiaux and Sallé, 1889. ***

*Dicrepidius ignotus* Fleutiaux and Sallé, 1889. ***

*Dicrepidius palmatus* Candèze, 1859.

*Dicrepidius ramicornis* (Palisot de Beauvois, 1805).

Genus *Dipropus* Germar, 1839.

*Dipropus brunneus* (Candèze, 1859).

*Dipropus puberulus* (Boheman, 1858).

**TRIBE Physorhinini Candèze, 1895.**

Genus *Anchastomorphus* Champion, 1895.

*Anchastomorphus dufaui* Fleutiaux, 1911. ***

Genus *Anchastus* LeConte, 1853.

*Anchastus insularis* Candèze, 1889. **

*Anchastus sautierei* Chassain, 2008. ***

*Anchastus terminatus* Candèze, 1864. ***

Genus *Physorhinus* Germar, 1840.

*Physorhinus insularis* Candèze, 1881. **

**SUBFAMILY Cardiophorinae Candèze, 1859.**

Genus *Esthesopus* Eschscholtz, 1829.

*Esthesopus grenadensis* Champion, 1897. **

*Esthesopus poedicus* Candèze, 1860. *

Genus *Horistonotus* Candèze, 1860.

*Horistonotus asthenicus* Candèze, 1860. *

**SUBFAMILY Lissominae Laporte de Castelnau, 1835.**

**TRIBE Lissomini Laporte de Castelnau, 1835.**

Genus *Drapetes* Dejean, 1821.

*Drapetes mediorufus* Fleutiaux, 1911. ***

*Drapetes nigricans* Bonvouloir, 1859.

*Drapetes sellatus* Bonvouloir, 1859.
Genus *Lissomus* Dalman, 1824.

*Lissomus impressifrons* Bonvouloir, 1859.

*Lissomus insularis* Cobos, 1966. ***

*Lissomus punctulatus* Dalman, 1824.

SUBFAMILY Agrypninae Candèze, 1857.

TRIBE Agrypnini Candèze, 1857.

Genus *Lacon* Laporte de Castelnau, 1836.

*Lacon modestus* (Boisduval, 1835).

*Lacon modestus* var. *guadeloupensis* Candèze, 1857.

*Lacon subcostatus* (Candèze, 1857). *

TRIBE Conoderini Schoenherr, 1833.

Genus *Aeolus* Eschscholtz, 1829.

*Aeolus melliculus* Candèze, 1859.

*Aeolus rubricatus* Candèze, 1895.

Genus *Conoderus* Eschscholtz, 1829.

*Conoderus belti* (Champion, 1895).

*Conoderus bifoveatus* (Palisot de Beauvois, 1805). *

*Conoderus castaneus* (Fabricius, 1792). *

*Conoderus delauneyi* (Fleutiaux, 1911). **

*Conoderus lividus* (DeGeer, 1774).

*Conoderus posticus* (Eschscholtz, 1822).

*Conoderus pinguis* Candèze 1859.

*Conoderus posticus* var. *sticturus* Candèze, 1859.

*Conoderus rufidens* (Fabricius, 1801).

*Conoderus vitraci* (Fleutiaux, 1911). **

Genus *Heteroderes* Latreille, 1834.

*Heteroderes amplicollis* (Gyllenhal, 1817).

TRIBE Pyrophorini Candèze, 1863.

Genus *Lygelater* Costa, 1975.

*Lygelater ignitus* (Fabricius, 1787).

Genus *Pyrophorus* Illiger, 1809.

*Pyrophorus hesperus* Candèze, 1803.

*Pyrophorus pellucens* Eschscholtz, 1830.

*Pyrophorus phosphorescens* Laporte de Castelnau, 1840.

TRIBE Hemirhipini Candèze, 1857.

Genus *Chalcolepidius* Eschscholtz, 1829.

*Chalcolepidius obscurus* Laporte de Castelnau, 1836. *

*Chalcolepidius webbi* LeConte, 1854.

FAMILY Lycidae Laporte de Castelnau, 1836.

SUBFAMILY Lycinae Laporte de Castelnau, 1836.

TRIBE Calopterini Green, 1949.

Genus *Mesopteron* Bourgeois, 1905.

*Mesopteron pecticornis* (Chevrolat, 1870). ***
FAMILY Lampyridae Rafinesque, 1815.

SUBFAMILY Lampyrinae Rafinesque, 1815.

TRIBE Cratomorphini Green, 1948.

Genus *Aspisoma* Laporte de Castlenau, 1833.

*Aspisoma ignitum* (Linnaeus, 1767).

*Aspisoma sexpunctatum* (Motschulsky, 1854).

TRIBE Photinini LeConte, 1881.

Genus *Photinus* Laporte de Castelnau, 1833.

*Photinus discoideus* (Sahlberg, 1823). ***

*Photinus vitiosus* Gemminger, 1870. *

Genus *Robopus* Motschulsky, 1853.

*Robopus vittatus* (Olivier, 1790). *

*Robopus vittiger* (Gyllenhal, 1817).

FAMILY Cantharidae Imhoff, 1856.

SUBFAMILY Silinae Mulsant, 1862.

Genus *Tylocerus* Dalman, 1833.

*Tylocerus cinctipennis* (Fleutiaux and Sallé, 1889). ***

*Tylocerus maculicornis* (Fleutiaux and Sallé, 1889). ***

*Tylocerus unilineatus* Pic, 1938. ***

Genus *Tytthonyx* LeConte, 1851.

*Tytthonyx guadeloupensis* (Fleutiaux and Sallé, 1889). **

INFRAORDER Bostrichiformia Forbes, 1926.

SUPERFAMILY Bostrichoidea Latreille, 1802.

FAMILY Dermestidae Latreille, 1804.

SUBFAMILY Dermestinae Latreille, 1804.

TRIBE Dermestini Latreille, 1804.

Genus *Dermestes* Linnaeus, 1758.

*Dermestes ater* DeGeer, 1774.

*Dermestes carnivorus* Fabricius, 1775.

*Dermestes lardarius* Linnaeus, 1758.

*Dermestes maculatus* DeGeer, 1774.

SUBFAMILY Thorictinae Agassiz, 1846.

TRIBE Thaumaphrastini Anderson, 1949.

Genus *Thorictodes* Reitter, 1875.

*Thorictodes heydeni* Reitter, 1875.

SUBFAMILY Megatominae Leach, 1815.

TRIBE Megatomini Leach, 1815.

Genus *Orphinus* Motschulsky, 1858.

*Orphinus fulvipes* (Guérin-Méneville, 1838).

FAMILY Bostrichidae Latreille, 1802.

SUBFAMILY Polyaconinae Lesne, 1896.

Genus *Melalgus* Dejean, 1833.

*Melalgus caribbeanus* (Lesne, 1906).
*Melalgus gonagrus* (Fabricius, 1798).

SUBFAMILY Apatinae Jacquelin du Val, 1861.
TRIBE Apatini Jacquelin du Val 1861.

Genus *Apate* Fabricius, 1775.
*Apate cephalotes* (Olivier, 1790). I

SUBFAMILY Bostrichinae Latreille, 1802.
TRIBE Bostrichini Latreille, 1802.

Genus *Amphicerus* Guérin-Méneville, 1845.
*Amphicerus cornutus* Pallas, 1772.

TRIBE Xyloperthini Lesne, 1921.

Genus *Tetrapriocera* Horn, 1878.
*Tetrapriocera longicornis* (Olivier, 1795).

Genus *Xylomeira* Lesne, 1901.
*Xylomeira tridens* (Fabricius, 1792).

SUBFAMILY Dinoderinae Thomson, 1863.

Genus *Dinoderus* Stephens, 1830.
*Dinoderus bifoveolatus* (Wollaston, 1858).
*Dinoderus distinctus* Lesne, 1897. I
*Dinoderus minutus* (Fabricius, 1775). I

Genus *Rhyzopertha* Stephens, 1830.
*Rhyzopertha dominica* (Fabricius 1792).

SUBFAMILY Lyctinae Billberg, 1820.
TRIBE Lyctini Billberg, 1820.

Genus *Lyctus* Fabricius, 1792.
*Lyctus brunneus* (Stephens, 1830).
*Lyctus caribeanus* Lesne, 1931.

Genus *Minthea* Pascoe, 1866.
*Minthea rugicollis* (Walker, 1858).

FAMILY Ptinidae Latreille, 1802.
SUBFAMILY Ptininae Latreille, 1802.
TRIBE Ptinini Latreille, 1802.

Genus *Pitnus* Gorham, 1880.
*Pitnus antillanus* Bellés, 1992. *

Genus *Ptinus* Linnaeus, 1767.
*Ptinus dufaui* Pic, 1906. **
*Ptinus tessellatus* Gorham, 1898. *

SUBFAMILY Ernobiinae Pic, 1912.
TRIBE Ernobiini Pic, 1912.

Genus *Scymnuseutheca* Pic, 1909.
*Scymnuseutheca apicalis* Pic, 1909. ***
SUBFAMILY Anobiinae Fleming, 1821.

TRIBE Calymmaderini White, 1982.

Genus *Calymmaderus* Solier, 1849.

*Calymmaderus bibliothecarum* (Poey, 1851). *

*Calymmaderus brevissimus* (Pic, 1909). ***

*Calymmaderus dufaui* (Pic, 1906). **

TRIBE Stegobiini White, 1982.

Genus *Stegobium* Motschulsky, 1860.

*Stegobium paniceum* (Linnaeus, 1758). I

SUBFAMILY Xyletininae Gistel, 1848.

TRIBE Xyletini Gistel, 1856.

Genus *Xyletinus* Latreille, 1809.

*Xyletinus marmoratus* Pic, 1911. **

TRIBE Lasiodermini White, 1982.

Genus *Lasioderma* Stephens, 1835.

*Lasioderma serricorne* (Fabricius, 1792). I

SUBFAMILY Dorcatominae Thomson, 1859.

Genus *Petalium* LeConte, 1861.

*Petalium antillarum* Pic, 1903.

*Petalium fauveli* Pic, 1905. **

*Petalium fleutiauxi* Lepesme, 1947. ***

*Petalium pici* Lepesme, 1947. ***

*Petalium punctatum* Pic, 1911. ***

Genus *Protheca* LeConte, 1865.

*Protheca guadalupensis* (Pic, 1909). ***

Genus *Pseudodorcatoma* Pic, 1905.

*Pseudodorcatoma mariei* Lepesme, 1947. **

*Pseudodorcatoma ornata* Pic, 1905. ***

Genus *Stichtoptychus* Fall, 1905.

*Stichtoptychus dufaui* (Pic, 1911). **

SUBFAMILY Mesocoelopodinae Mulsant and Rey, 1864.

TRIBE Mesocoelopodini Mulsant and Rey, 1864.

Genus *Cryptorama* Fall, 1905.

*Cryptorama antillensis* White, 1984. *

*Cryptorama dufaui* (Pic, 1909). ***

*Cryptorama sericeum* (Pic, 1909). **

Genus *Tricorynus* Waterhouse, 1849.

*Tricorynus lepesmei* White, 1965. ***

*Tricorynus pierrei* (Lepesme, 1947). **

*Tricorynus sallei* (Guérin-Méneville, 1851). *
INFRAORDER Cucujiformia Lameere, 1938.

SUPERFAMILY Cleroidea Latreille, 1802.

FAMILY Trogossitidae Latreille, 1802.

SUBFAMILY Lophocaterinae Crowson, 1964.

TRIBE Lophocaterini Crowson, 1964.

Genus *Lophocateres* Olliff, 1883.

*Lophocateres pusillus* (Klug, 1832).

SUBFAMILY Trogossitinae Latreille, 1802.

TRIBE Trogossitini Latreille, 1802.

Genus *Corticotomus* Sharp, 1891.

*Corticotomus dufaui* (Léveillé, 1907). ***

Genus *Nemozoma* Latreille, 1804.

*Nemosoma fleutiauxi* Lepesme, 1947. ***

Genus *Temnoscheila* Westwood, 1835.

*Temnoscheila obscura* (Reitter, 1875). **

Genus *Tenebroides* Piller and Mitterpacher 1783.

*Tenebroides floridanus* Schaeffer, 1918.

*Tenebroides mauritanicus* (Linnaeus, 1758). I

*Tenebroides punctulata* Reitter, 1875. *

*Tenebroides transversicollis* Jacquelin du Val, 1857. *

FAMILY Thanerocleridae Chapin, 1924.

Genus *Thaneroclerus* Lefèbvre, 1838.

*Thaneroclerus buquet* (Lefèbvre, 1835). I

FAMILY Cleridae Latreille, 1802.

SUBFAMILY Epiphloeinae Kuwert, 1893.

Genus *Madoniella* Pic, 1935.

*Madoniella caporaali* Pic, 1935. ***

*Madoniella minor* Pic, 1935. ***

*Madoniella pici* Lepesme, 1947. **

SUBFAMILY Enopliinae Cistel, 1856.

Genus *Neorthopleura* Barr, 1976.

*Neorthopleura subfasciatum* (Chevrolat, 1874).

SUBFAMILY Korynetinae Laporte de Castelnau, 1836.

Genus *Korynetes* Herbst, 1792.

*Korynetes caeruleus* (De Geer, 1775).

Genus *Necrobia* Olivier, 1795.

*Necrobia rufipes* (De Geer, 1775).

SUBFAMILY Tarsosteninae Jacquelin du Val, 1861.

Genus *Tarsostenus* Spinola, 1844.

*Tarsostenus univittatus* (Rossi, 1792).
## FAMILY Malachiidae Fleming, 1821.

### SUBFAMILY Malachiinae Fleming, 1821.

Genus *Ablechrus* Waterhouse, 1877.

*Ablechrus nigrocaeruleus* (Gorham, 1898). Constantin (2017). **

## FAMILY Melyridae Leach, 1815.

Genus *Melyrodes* Gorham, 1882.

*Melyrodes aureohirta* Constantin and Schiller, 2014. Constantin and Schiller (2014). ***

## SUPERFAMILY Coccinelloidea Latreille, 1807.

### FAMILY Bothrideridae Erichson, 1845.

#### SUBFAMILY Bothriderinae Erichson, 1845.

Genus *Bothrideres* Dejean, 1835.

*Bothrideres chevrolati* (Grouvelle in Grouvelle and Raffray, 1908). **

*Bothrideres dufaui* (Grouvelle in Grouvelle and Raffray, 1908). **

*Bothrideres planus* Chevrolat, 1864. *

Genus *Sosylus* Erichson, 1845.

*Sosylus castaneus* Pascoe, 1863.

## FAMILY Cerylonidae Billberg, 1820.

### SUBFAMILY Ceryloninae Billberg, 1820.

Genus *Cautomus* Sharp, 1885.

*Cautomus infimus* Grouvelle in Grouvelle and Raffray, 1908. ***

Genus *Mychocerus* Erichson, 1845.

*Mychocerus discretus* (Casey, 1890).

Genus *Philothermus* Aubé, 1843.

*Philothermus exaratus* (Chevrolat, 1864).

*Philothermus guadeloupensis* Grouvelle, 1902. ***

*Philothermus puberulus* Schwarz, 1878.

### SUBFAMILY Euxestinae Grouvelle, 1908.

Genus *Euxestus* Wollaston, 1858.

*Euxestus erithacus* (Chevrolat, 1863).

Genus *Metacerylon* Grouvelle, 1906.

*Metacerylon dufaui* Grouvelle and Raffray, 1912. ***

## FAMILY Coccinellidae Latreille, 1807.

### SUBFAMILY Coccinellinae Latreille, 1807.

#### TRIBE Azyini Mulsant, 1850.

Genus *Azya* Mulsant, 1850.

*Azya scutata* Mulsant, 1850.

Genus *Pseudoazya* Gordon, 1980.

*Pseudoazya trinitatis* (Marshall, 1912).

#### TRIBE Chilocorini Mulsant, 1846.

Genus *Chilocorus* Leach, 1815.

*Chilocorus cacti* (Linnaeus, 1767). I
Genus *Cladis* Mulsant, 1850. *

*Cladis nitidula* (Fabricius, 1792). *

### TRIBE Coccidulini Mulsant, 1846.

Genus *Coleomegilla* Timberlake, 1920.

*Coleomegilla maculata* (DeGeer, 1775).

Genus *Cycloneda* Crotch, 1871.

*Cycloneda delauneyi* (Fleutiaux and Sallé, 1889). **

*Cycloneda sanguinea* (Linnaeus, 1763).

Genus *Psyllobora* Dejean, 1833.

*Psyllobora lineola* Fabricius, 1792. *

### TRIBE Coccinellini Latreille, 1807.

Genus *Coelophora* Mulsant, 1850.

*Coelophora inaequalis* (Fabricius, 1775). Nicolas (2012); Peck (2016). I.

### TRIBE Cryptognathini Mulsant, 1850.

Genus *Pentilia* Mulsant, 1850.

*Pentilia egena* Mulsant, 1850.

### TRIBE Diomini Gordon, 1999.

Genus *Decadiomus* Chapin, 1933.

*Decadiomus hubbardi* Chapin, 1933. Peck (2016). *

*Decadiomus hughesi* Gordon and Hilburn, 1990. Peck (2016). **

Genus *Diomus* Mulsant, 1850.

*Diomus austrinus* Gordon, 1976. Peck (2016). **

*Diomus ochroderus* (Mulsant, 1850). *

*Diomus roseicollis* (Mulsant, 1853).

*Diomus seminulus* Mulsant, 1850. Peck (2016).

*Diomus terminatus* Say, 1835. Peck (2016).

*Diomus thoracicus* (Fabricius, 1801).

### TRIBE Epilachnini Mulsant, 1846.

Genus *Epilachna* Chevrolat, 1836.

*Epilachna borealis* Fabricius, 1875. Peck (2016). I.

### TRIBE Hyperaspidini Mulsant, 1846.

Genus *Hyperaspis* Chevrolat in Dejean, 1837.

*Hyperaspis connectens* Thunberg, 1808. Peck (2016).

*Hyperaspis festiva* Mulsant, 1850. Peck (2016).

### TRIBE Noviini Mulsant, 1850.

Genus *Anovia* Casey, 1908.

*Anovia peruviana* Gordon, 1972. Peck (2016).

*Anovia punica* Gordon, 1972. Peck (2016).

Genus *Rodolia* Mulsant, 1850.

*Rodolia cardinalis* (Mulsant, 1850). Peck (2016). I.
TRIBE Scymnillini Casey, 1899

Genus *Zilus* Mulsant, 1850.
*Zilus eleutherae* (Casey, 1899). Peck (2016).

TRIBE Scymnini Mulsant, 1846.

Genus *Cryptolaemus* Mulsant, 1853.
*Cryptolaemus montrouzieri* Mulsant, 1853. I

Genus *Nephaspis* Casey, 1899.
*Nephaspis namolica* Gordon, 1982. Peck (2016).
*Nephaspis oculata* (Blatchley, 1917). Peck (2016).

Genus *Scymnus* Kugelann, 1794.
*Scymnus cyanipennis* Mulsant, 1850. Peck (2016). **
*Scymnus floralis* Fabricius, 1792. Peck (2016).
*Scymnus gilae* Casey, 1899. Peck (2016).
*Scymnus phloeus* Mulsant, 1850. Peck (2016). *

TRIBE Stethorini Dobzhansky, 1924.

Genus *Stethorus* Weise, 1885.
*Stethorus pinachi* Gordon and Chapin, 1983. Peck (2016).
*Stethorus pseudocaribus* Gordon and Chapin, 1983. Peck (2016).

TRIBE Sticholotidini Weise, 1901.

Genus *Neaptera* Gordon, 1991.
*Neaptera korschefskyi* (Duverger, 1986). ***
*Neaptera viola* Gordon, 1991. **

Genus *Sticholotis* Crotch, 1874.
*Sticholotis ruficeps* Weise, 1902. I

SUBFAMILY Microweiseinae Leng, 1920.

TRIBE Microweiseini Leng, 1920.

Genus *Coccidophilus* Brèthes, 1905.
*Coccidophilus cariba* Gordon, 1978. *
*Coccidophilus nigra* Duverger, 1986. ***

TRIBE Serangiini Pope, 1962.

Genus *Delphastus* Casey, 1899.
*Delphastus barti* Duverger, 1986. ***
*Delphastus nebulosus* Chapin, 1940. *

FAMILY Corylophidae LeConte, 1852.

SUBFAMILY Corylophinae LeConte, 1852.

TRIBE Peltinodini Paulian, 1950.

Genus *Holopsis* Broun, 1883.
*Holopsis pellucidus* (Matthews, 1899). ***

TRIBE Orthoperini Jacquelin du Val, 1857.

Genus *Orthoperus* Stephens, 1829.
*Orthoperus minutissimus* Matthews, 1899. **
TRIBE Rypobiini Paulian, 1950.

Genus *Hoplicnema* Matthews, 1899. *

*Hoplicnema sallei* Matthews, 1899. *

FAMILY Murmidiidae Jacquelin du Val, 1858

Genus *Botrodus* Casey, 1890.

*Botrodus dufaui* Grouvelle and Raffray, 1912. **

SUPERFAMILY Cucujoidea Latreille, 1802.

FAMILY Cryptophagidae Kirby, 1826.

SUBFAMILY Atomariinae LeConte, 1861

TRIBE Atomariini LeConte, 1861.

Genus *Curelius* Casey, 1900.

*Curelius japonicus* (Reitter, 1878). Peck (2016).

FAMILY Endomychidae Leach, 1815.

Genus *Eidoreus* Sharp, 1885.

*Eidoreus minutus* Sharp, 1885.

FAMILY Erotylidae Latreille, 1802.

SUBFAMILY Cryptophilinae Casey, 1900.

TRIBE Cryptophilini Casey, 1900.

Genus *Cryptophilus* Reitter, 1874.

*Cryptophilus integer* (Heer, 1841).

SUBFAMILY Erotylinae Latreille 1802.

TRIBE Erotylini Latreille, 1802.

Genus *Iphiclus* Chevrolat, 1837.

*Iphiclus guadeloupensis* (Fabricius, 1792). ***

SUBFAMILY Languriinae Hope, 1840.

TRIBE Hapalipini Leschen, 2003.

Genus Hapalips Reitter, 1877.

*Hapalips angulosus* Grouvelle in Grouvelle and Raffray, 1908. **

*Hapalips delauneyi* Grouvelle in Grouvelle and Raffray, 1908. ***

*Hapalips dufaui* Grouvelle in Grouvelle and Raffray, 1908. ***

*Hapalips guadeloupensis* Grouvelle, 1902. ***

*Hapalips sharpi* Grouvelle in Grouvelle and Raffray, 1908. ***

Genus *Platoberus* Sharp, 1900.

*Platoberus dufaui* Grouvelle, 1916. **

*Platoberus latus* Sharp, 1900.

SUBFAMILY Loberinae Bruce, 1951.

Genus *Loberus* LeConte, 1861.

*Loberus testaceus* Reitter, 1875. *

*Loberus vitraci* Grouvelle, 1902. ***

SUBFAMILY Xenoscelinae Ganglbauer, 1899.

Genus *Telmatoscius* Sharp, 1900.

*Telmatoscius dufaui* Grouvelle and Raffray, 1912. ***