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## Acknowledgements

The authors would like to thank the following people; for reagents and fly stocks: Emmanuel Derivery, Debbie Andrew, Jordan Raff, Paul Conduit, Daniel St. Johnston, Claudio Sunkel, Yu-Chiun Wang, Jens Januschke, Cayetano Gonzales, Magali Suzanne, Markus Affolter, Ron Vale; for use of the otracks software: Guy Blanchard; for image analysis: Jérôme Boulanger. The work is supported by the Medical Research Council (file reference number U105178780).

## Author contributions

Conceptualisation, K.R. and G.Gill.; Methodology, K.R., G.Gill. G.Gird.; Investigation, K.R., G.Gill., G.Gird.; Writing—Original Draft, K.R., G.Gill.; Funding Acquisition, K.R.

## Competing interests

The authors declare no competing interests.

## Additional information

**Supplementary information** The online version contains supplementary material available at https://doi.org/10.1038/s41467-021-24332-0.

**Correspondence** and requests for materials should be addressed to K.R.

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