<table><tr><td>Tipula serrulata</td><td>Loew, 1864</td></tr><tr><td>Tipula tacomicola</td><td>Alexander, 1949</td></tr></table>

## Subgenus Yamatotipula

Matsumura, 1916

<table><tr><td>Tipula aleutica</td><td>Alexander, 1923</td></tr><tr><td>Tipula albocaudata</td><td>Doane, 1901</td></tr><tr><td>Tipula caloptera</td><td>Loew, 1863</td></tr><tr><td>Tipula carsoni</td><td>Alexander, 1963</td></tr><tr><td>Tipula cayuga</td><td>Alexander, 1915</td></tr><tr><td>Tipula cognata</td><td>Doane, 1901</td></tr></table>

<table><tr><td>Tipula colteri</td><td>Alexander, 1943</td></tr><tr><td>Tipula concava</td><td>Alexander, 1926</td></tr><tr><td>Tipula continentalis</td><td>Alexander, 1941</td></tr><tr><td>Tipula dejecta</td><td>Walker, 1856</td></tr></table>

<table><tr><td>Tipula eluta</td><td>Loew, 1863</td></tr><tr><td>Tipula footeana</td><td>Alexander, 1961</td></tr><tr><td>Tipula furca</td><td>Walker, 1848</td></tr></table>

<table><tr><td>Tipula glendenningi</td><td>Alexander, 1943</td></tr><tr><td>Tipula grenfelli</td><td>Alexander, 1928</td></tr><tr><td>Tipula iroquois</td><td>Alexander, 1915</td></tr></table>

<table><tr><td>Tipula jacobus</td><td>Alexander, 1931</td></tr><tr><td>Tipula kennicotti</td><td>Alexander, 1915</td></tr></table>

<table><tr><td>Tipula noveboracensis</td><td>Alexander, 1919</td></tr><tr><td>Tipula sayi</td><td>Alexander, 1911</td></tr></table>

<table><tr><td>Tipula spernax</td><td>Osten Sacken, 1877</td></tr><tr><td>Tipula strepens</td><td>Loew, 1863</td></tr><tr><td>Tipula sulphurea</td><td>Doane, 1901</td></tr></table>

<table><tr><td>Tipula tephrocephala</td><td>Loew, 1864</td></tr><tr><td>Tipula tricolor</td><td>Fabricius, 1775</td></tr></table>

<table><tr><td>Tipula vicina</td><td>Dietz, 1917</td></tr></table>

<table><tr><td>- - NU - - MB ON QC - - - - -</td><td>Serrulate Large Crane Fly</td><td>Tipule serrulée</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Mount Rainier Large Crane Fly</td><td>Tipule du mont Rainier</td></tr></table>

Syn.: Tipula alexandriana Dietz, 1917; Tipula edmundsi Alexander, 1948

<table><tr><td>- - BC - - - - - - - - -</td><td>Colter's Large Crane Fly</td><td>Tipule de Colter</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Concave Large Crane Fly</td><td>Tipule concave</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Continental Large Crane Fly</td><td>Tipule continentale</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Dejected Large Crane Fly</td><td>Tipule déprimée</td></tr></table>

Syn.: Tipula brevicollis Alexander, 1915; Tipula fumosa Doane, 1901

<table><tr><td>- - BC - - - - - - - - -</td><td>Washed Large Crane Fly</td><td>Tipule lavée</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Foote's Large Crane Fly</td><td>Tipule de Foote</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Forked Large Crane Fly</td><td>Tipule fourchue</td></tr></table>

Syn.: Tipula bella Loew, 1863; Tipula fuscifer Harris, 1925

<table><tr><td>- - BC - - - - - - - - -</td><td>Glendenning's Large Crane Fly</td><td>Tipule de Glendenning</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Grenfell's Large Crane Fly</td><td>Tipule de Grenfell</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Iroquois Large Crane Fly</td><td>Tipule des Iroquois</td></tr></table>

Syn.: Tipula cincta Loew, 1863

<table><tr><td>- - BC - - - - - - - - -</td><td>James's Large Crane Fly</td><td>Tipule de James</td></tr><tr><td>- - AB - - - - - - - - -</td><td>Kennicott's Large Crane Fly</td><td>Tipule de Kennicott</td></tr></table>

Syn.: Tipula diluta Doane, 1901; Tipula parvemarginata Alexander, 1926; Tipula tetra Alexander, 1915

<table><tr><td>- - BC - - - - - - - - -</td><td>New York Large Crane Fly</td><td>Tipule de New York</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Say's Large Crane Fly</td><td>Tipule de Say</td></tr></table>

Syn.: Tipula costalis Say, 1823

<table><tr><td>- - BC - - - - - - - - -</td><td>Scornful Large Crane Fly</td><td>Tipule dédaigneuse</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Rustling Large Crane Fly</td><td>Tipule froissée</td></tr><tr><td>- - AB SK MB ON QC - - - - -</td><td>Sulphurous Large Crane Fly</td><td>Tipule sulfureuse</td></tr></table>

Syn.: Tipula sulphurea sulphurea Doane, 1901

<table><tr><td>- - BC - - - - - - - - -</td><td>Grey-headed Large Crane Fly</td><td>Tipule à tête grise</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Tricoloured Large Crane Fly</td><td>Tipule tricolore</td></tr></table>

Syn.: Tipula vitrea van der Wulp, 1881

<table><tr><td>- - BC - - - - - - - - -</td><td>Local Large Crane Fly</td><td>Tipule voisine</td></tr></table>



<table><tr><td>- - BC - - - - - - - - -</td><td>Aleutian Islands Large Crane Fly</td><td>Tipule des îles Aléoutiennes</td></tr><tr><td>- - BC - - - - - - - - -</td><td>White-tailed Large Crane Fly</td><td>Tipule à queue blanche</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Nicely Winged Large Crane Fly</td><td>Tipule à belles ailes</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Carson's Large Crane Fly</td><td>Tipule de Carson</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Simmons's Woods Large Crane Fly</td><td>Tipule de la forêt de Simmons</td></tr><tr><td>- - BC - - - - - - - - -</td><td>Olympia Large Crane Fly</td><td>Tipule d'Olympia</td></tr></table>
<figure><img src="image_6.png" /></figure>
Further Acknowledgments

Thanks to John Acorn (University of Alberta), Trond Andersen (University Museum of Bergen), Patrick
Ashe (Irish Biogeographical Society), David Beresford (Trent University), Bohdan Bilyj (Biotaxa), Annette
Borkent, Christopher J. Borkent, Stephanie Boucher (McGill University), Derek Bridgehouse (Halifax),
Scott E. Brooks (CNC, Ottawa), Fenja Brodo (Canadian Museum of Nature), Matthias Buck (Royal Al-
berta Museum), Trevor Burt (Ottawa), Syd Cannings (Whitehorse), Peter Cranston (Australia National
University), William Crins (Peterborough), Jeffrey M. Cumming (CNC, Ottawa), Deanna Dodgson (Win-
nipeg), Torbjørn Ekrem (Department of Natural History, NTNU University Museum), John Epler (Florida),
Eric Fisher (Sacramento), Steve Gaimari (California Department of Food and Agriculture), Wojciech
Gilka (Department of Invertebrate Zoology and Parasitology, University of Gdańsk), Deirdre E. Griffiths
(for providing unpublished material compiled by GD Griffiths on the Anthonyiidae), Martin Hauser (Cal-
ifornia Department of Food and Agriculture), Patrick Hudson (USGS), Michael Irwin (Illinois Natural His-
tory Survey), Gail Kampmeier (Illinois Natural History Survey), John Klymko (Sackville), Peter Langton
(Northern Ireland), Michelle Locke (CNC, Ottawa), Owen Lonsdale (CNC, Ottawa), Evgenyi Makarchenko
(Federal Scientific Centre of the East Asia Terrestrial Biodiversity), Stephen A. Marshall (University of
Guelph), Jon Martin (University of Melbourne), Tristan McKnight (Tucson), Gil Felipe Gonçalves Miranda
(Ottawa), Kevin Moran (CNC, Ottawa), Behnam Motamedinia (CNC, Ottawa), Karen Needham (Beaty
Biodiversity Museum, Vancouver), James O'Hara (CNC, Ottawa), Steve Paiero (University of Guelph),
Thomas Pape (Natural History Museum of Denmark), Adrian Pont (Oxford University), Bruno Rossaro
(Università degli Studi di Milano), Steffen Roth (University Museum of Bergen), Cory Sheffield (Roy-
al Saskatchewan Museum), Bradley J. Sinclair (CNC, Ottawa), Jeff Skevington (Ottawa), Martin Spies
(Zoologische Staatssammlung Munich - ZSM), Elisabeth Stur (Department of Natural History, NTNU
University Museum), John Swann (University of Calgary), Shaun Winterton (California Department of
Food and Agriculture), Richard Yank (Ottawa), and Andrew Young (University of Guelph).

We would like also to thank all the staff of the Diptera Collection of the Canadian National Collec-
tion of Insects, Arachnids and Nematodes (Agriculture and Agri-Food Canada, Ottawa) for providing
support in the generation of this checklist. Specimen data came from many collections; thanks go to
the staff members responsible for them. The main collections used are: Canadian National Collection
of Insects, Arachnids, and Nematodes (CNC), Ottawa; Beaty Biodiversity Museum, University of BC,
Vancouver; Canadian Forest Service, Great Lakes Forestry Centre, Sault Ste. Marie; Canadian Forest
Service, Northern Forestry Centre, Edmonton; Canadian Forest Service, Pacific Forestry Centre, Victoria;
E.H. Strickland Museum, University of Alberta, Edmonton; Illinois Natural History Survey, Champaign;
Insectarium de Montréal's scientific collection (IMQC); Lyman Entomological Museum, McGill Universi-
ty, Montreal; Manitoba Museum, Winnipeg; New Brunswick Museum, Saint John; Ohio State University,
Columbus; Ouellet-Robert entomological collection (QMOR), Université de Montréal, Montreal; Royal
Alberta Museum, Edmonton; Royal BC Museum, Victoria; Royal Ontario Museum, Toronto; Royal Sas-
katchewan Museum, Regina; University of Calgary Insect Collection, Calgary; University of Guelph Col-
lection, Guelph; US National Museum, Washington; Wallis Roughley Museum of Entomology, University
of Manitoba, Winnipeg.

We are grateful to those who posted photographs on iNaturalist and BugGuide and the experts
who identified them.
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The checklist of flies (Diptera) in Canada is a fundamental piece of work. It represents the first national list of all flies ever done for the country. Knowing which species occur in Canada will enable the assessment of their conservation status. Flies represent a large proportion of biodiversity, and thus their inclusion is important.