Question: What is the higher taxon of Gnathostomata, which was introduced at -525000000-0-0?
Paraphrase: What is the higher taxon of Gnathostomata, which was presented at -525000000-0-0?
SPARQL (Wikidata): SELECT ?answer WHERE { wd:Q26214 wdt:P171 ?answer . ?answer wdt:P580 ?x FILTER(contains(YEAR(?x),'0000'))}
Answer: Vertebrata, Olfactores