# Satyrini

De Satyrinii zijn een geslachtengroep van vlinders in de onderfamilie Satyrinae van de familie Nymphalidae. De hier gegeven onderverdeling in subtribus en geslachten volgt die van Peña et al. van 2006, aangevuld uit het overzicht van geslachten door de Nymphalidae Systematics Group.

## Subtribus en geslachten
### Parargina Tutt, 1896

* Pararge Hübner, 1819
* Chonala Moore, 1893*
* Kirinia Moore, 1893
** = Esperarge Nekrutenko, 1988
** = Esperia Nekrutenko, 1987 non Esperia Hübner, 1825 (Oecophoridae)
** = Esperella Nekrutenko, 1987 non Esperella Vosmaer, 1887 (Demospongiae)
* Lasiommata Westwood, 1841
** = Amecera Butler, 1867
* Lopinga Moore, 1893
** = Crebeta Moore, 1893
** = Polyargia Verity, 1957
* Nosea Koiwaya, 1993*
* Orinoma Gray, 1846*
* Rhaphicera Butler, 1867*
* Tatinga Moore, 1893*

### Lethina Reuter, 1896

* Lethe Hübner, 1819
** = Tanaoptera Billberg, 1820
** = Zophoessa Doubleday, 1849
** = Debis Doubleday, 1849
** = Charma Doherty, 1886
** = Tansima Moore, 1881
** = Rangbia Moore, 1892
** = Nemetis Moore, 1892
** = Pegada Moore, 1892
** = Kirrodesa Moore, 1892
** = Placilla Moore, 1892
** = Archondesa Moore, 1892
** = Choranesa Moore, 1892
** = Dionana Moore, 1892
** = Sinchula Moore, 1892
** = Kerrata Moore, 1892
** = Putlia Moore, 1892
** = Harima Moore, 1892
** = Magula Fruhstorfer, 1911
** = Hermias Fruhstorfer, 1911
* Aphysoneura Karsch, 1894*
** = Rhaphiceropsis Sharpe, 1894
* Enodia Hübner, 1819
* Hanipha Moore, 1880*
* Mandarinia Leech, 1892*
* Neope Moore, 1866
** = Enope Moore, 1857 non Enope Walker, 1854 (Erebidae)
** = Neope Butler, 1867
** = Blanaida Kirby, 1871
** = Patala Moore, 1892
* † Neorinella Martins Neto et al., 1993*
* † Neorinopsis Butler, 1873*
* Ninguta Moore, 1892*
** = Aranda Fruhstorfer, 1909
* † Pseudoneorina Nel & Descimon, 1986*
* Ptychandra C. & R. Felder, 1861*
* † Satyrites Scudder, 1872*
** = Lethites Scudder, 1875
* Satyrodes Scudder, 1875
** = Argus Scopoli, 1777

### Mycalesina Reuter, 1896

* Mycalesis Hübner, 1818
* Bicyclus Kirby, 1871
* Bletogona C. & R. Felder, 1867*
* Brakefieldia Aduse-Poku, Lees & Wahlberg, 2016
* Hallelesis Condamin, 1961
* Heteropsis Westwood, 1850*
* Nirvanopsis Vane-Wright, 2003*
* Orsotriaena Wallengren, 1858
* Pseudomycalesis Tsukada & Nishiyama, 1979*
** voor synoniemen, zie de subtribus

### Eritina Miller, 1968

* Erites Westwood, 1851*
* Coelites Westwood, 1850*

### Ragadiina Herrich-Schaeffer, 1864

* Ragadia Westwood, 1851*
* Acrophtalmia C. & R. Felder, 1861*
* Acropolis Hemming, 1934*
** = Pharia Fruhstorfer, 1911 non Pharia Gray, 1840 (Asteroidea)

### Coenonymphina Tutt, 1896

* Coenonympha Hübner, 1819
** = Chortobius Dunning & Pickard, 1858
** = Sicca Verity, 1953
* Lyela Swinhoe, 1908*
** = Dubierebia Muschamp, 1915
* Sinonympha Lee, 1974*
* Triphysa Zeller, 1850*
** = Phryne Herrich-Schäffer, 1844

### Hypocystina Miller, 1968

* Hypocysta Westwood, 1851
* Altiapa Parsons, 1986*
* Argynnina Butler, 1867*
* Argyronympha Mathew, 1886
* Argyrophenga Doubleday, 1845
* Dodonidia Butler, 1884
* Erebiola Fereday, 1879
* Erycinidia Rothschild & Jordan, 1905*
** = Pieridopsis Rothschild & Jordan, 1905
* Geitoneura Butler, 1867
* Harsiesis Fruhstorfer, 1911*
* Heteronympha Wallengren, 1858
** = Hipparchioides Butler, 1867
* Hyalodia Jordan, 1924*
* Lamprolenis Godman & Salvin, 1881
* Nesoxenica Waterhouse & Lyell, 1914
** = Xeniconympha Novicki, 1923
* Oreixenica Waterhouse & Lyell, 1914
* Paratisiphone Watkins, 1928*
* Percnodaimon Butler, 1876
* Platypthima Rothschild & Jordan, 1905*
* Tisiphone Hübner, 1819
** = Xenica Westwood, 1851
* Zipaetis Hewitson, 1863

### Euptychiina Miller, 1968

* Euptychia Hübner, 1818
* Amphidecta Butler, 1867
* Archeuptychia Forster, 1964*
* Caenoptychia Le Cerf, 1919*
** = Ristia Gagarin, 1936
* Caeruleuptychia Forster, 1964
** = Weymerana Forster, 1964
* Capronnieria Forster, 1964*
* Cepheuptychia Forster, 1964
* Cercyeuptychia Miller & Emmel, 1971*
* Chloreuptychia Forster, 1964
* Cissia Doubleday, 1848
** = Argyreuptychia Forster, 1964
** = Vareuptychia Forster, 1964
* Coeruleotaygetis Forster, 1964*
* Cyllopsis Felder, 1869
* Erichthodes Forster, 1964
* Euptychoides Forster, 1964
* Forsterinaria Gray, 1973
** = Haywardina Forster, 1964 non Haywardina Aczel, 1952 (Diptera)
** = Forsterina Lamas, 1977 non Forsterina Lehtinen, 1967 (Araneae)
* Godartiana Forster, 1964
* Guainanaza Freitas & Peña, 2006*
* Harjesia Forster, 1964
* Hermeuptychia Forster, 1964
* Magneuptychia Forster, 1964
* Megeuptychia Forster, 1964*
* Megisto Hübner, 1819*
* Moneuptychia Forster, 1964
** = Carminda Ebert & Dias, 1997
* Neonympha Hübner, 1818
* Oressinoma Doubleday, 1849
** = Ocalis Westwood, 1851
** = Ocalis Boisduval, 1870
* Palaeonympha Butler, 1871
* Paramacera Butler, 1868
* Parataygetis Forster, 1964
* Pareuptychia Forster, 1964
* Paryphthimoides Forster, 1964
* Pharneuptychia Forster, 1964
* Pindis Felder, 1869
* Posttaygetis Forster, 1964
* Praefaunula Forster, 1964*
* Pseudeuptychia Forster, 1964*
* Pseudodebis Forster, 1964*
* Rareuptychia Forster, 1964
* Satyrotaygetis Forster, 1964*
* Splendeuptychia Forster, 1964
* Taydebis Freitas, 2003*
* Taygetina Forster, 1964*
* Taygetis Hübner, 1819
* Taygetomorpha Miller, 2004*
* Yphthimoides Forster, 1964
* Zischkaia Forster, 1964

### Erebiina Tutt, 1896

* Erebia Dalman, 1816
** = Syngea Hübner, 1819
** = Epigea Hübner, 1819
** = Phorcis Hübner, 1819
** = Marica Hübner, 1819
** = Gorgo Hübner, 1819
** = Triariia Verity, 1953
** = Truncaefalcia Verity, 1953
** = Medusia Verity, 1953
** = Simplicia Verity, 1953
** = Atercoloratus Bang-Haas, 1938

### Maniolina Grote, 1897

* Maniola Schrank, 1801
** = Epinephele Hübner, 1819
* Aphantopus Wallengren, 1853
* Cercyonis Scudder, 1875
* Hyponephele Muschamp, 1915
* Proterebia Roos & Arnschied, 1980
* Pyronia Hübner, 1819
** = Idata de Lesse, 1952
** = Pasiphana de Lesse, 1952

### Pronophilina Reuter, 1896

* Pronophila Doubleday, 1849
** = Mygona Westwood, 1851
* Altopedaliodes Forster, 1964
* Antopedaliodes Forster, 1964
* Apexacuta Pyrcz, 2004
* Argyrophorus Blanchard, 1852
* Arhuaco Adams & Bernard, 1977
* Auca Hayward, 1953
* Calisto Hübner, 1823
* Cheimas Thieme, 1907
* Chillanella Herrera, 1966
* Corades Hewitson, 1849
** = Corades Doubleday, 1848 (nomen nudum)
** = Panarche Thieme, 1907
* Corderopedaliodes Forster, 1964
* Cosmosatyrus C. & R. Felder, 1867
* Daedalma Hewitson, 1858
* Dangond Adams & Bernard, 1979
* Diaphanos Adams & Bernard, 1981
* Drucina Butler, 1872
* Druphila Pyrcz, 2004
* Elina Blanchard, 1852
* Eretris Thieme, 1905
* Etcheverrius Herrera, 1965
* Eteona Doubleday, 1848
* Faunula C. & R. Felder, 1867
* Foetterleia Viloria, 2004
** = Foetterleia Viloria, 2003 (nom. nud.)
* Gyrocheilus Butler, 1867
* Haywardella Herrera, 1966
* Homoeonympha C. & R. Felder, 1867
** = Stygnus Felder & Felder, 1867 non Stygnus Perty, 1833 (Opiliones)
** = Stygnolepis Strand, 1942
** = Erebina Bryk, 1944
* Ianussiusa Pyrcz & Viloria, 2004
* Idioneurula Strand, 1932
** = Idioneura Felder & Felder, 1867 non Idioneura Selys, 1860 (Odonata)
* Junea Hemming, 1964
** = Polymastus Thieme, 1907
* Lasiophila C. & R. Felder, 1859
* Lymanopoda Westwood, 1851
** = Sarromia Westwood, 1851
** = Zabirnia Hewitson, 1877
** = Trophonina Röber, 1889
* Manerebia Staudinger, 1897
** = Penrosada Brown, 1944
** = Posteuptychia Forster, 1964
* Mygona Thieme, 1907
* Nelia Hayward, 1953
* Neomaenas Wallengren, 1858
** = Stibomorpha Butler, 1874
* Neomaniola Hayward, 1949
** = Pseudomaniola Weymer, 1890
* Neopedaliodes Viloria, Miller & Miller, 2004
** = Neopedaliodes Viloria, 2002 (nom. nud.)
* Neosatyrus Wallengren, 1858
* Oxeoschistus Butler, 1867
** = Dioriste Thieme, 1907
* Palmaris Herrera, 1965
* Pampasatyrus Hayward, 1953
** = Pseudocercyonis Miller & Emmel, 1971
* Pamperis Heimlich, 1959
* Panyapedaliodes Forster, 1964
** = Muscopedaliodes Forster, 1964
* Paramo Adams & Bernard, 1977
* Parapedaliodes Forster, 1964
* Pedaliodes Butler, 1867
* Pherepedaliodes Forster, 1964
* Physcopedaliodes Forster, 1964
* Praepedaliodes Forster, 1964
* Praepronophila Forster, 1964
* Proboscis Thieme, 1907
* Protopedaliodes Viloria & Pyrcz, 1994
* Pseudomaniola Weymer, 1890
** = Catargynnis Röber, 1892
* Punapedaliodes Forster, 1964
* Punargentus Heimlich, 1963
* Quilaphoetosus Herrera, 1966
* Redonda Adams & Bernard, 1981
* Sabatoga Staudinger, 1897
** = Zapatoca Apolinar, 1924
* Sierrasteroma Adams & Bernard, 1977
* Spinantenna Hayward, 1953
* Steremnia Thieme, 1905
** = Pseudosteroma Weymer, 1912
* Steroma Westwood, 1850
* Steromapedaliodes Forster, 1964
** = Steropoda Thieme, 1905
* Stuardosatyrus Herrera & Etcheverry, 1965
* Tamania Prycz, 1995
* Tetraphlebia C. & R. Felder, 1867
* Thiemeia Weymer, 1912

### Melanargiina Wheeler, 1903

* Melanargia Meigen, 1828
** = Agapetes Billberg, 1820
** = Arge Hübner, 1819 non Arge Schrank, 1802 (Hymenoptera)
** = Ledargia Houlbert, 1922
** = Epimede Houlbert, 1922
** = Parce Oberthür & Houlbert, 1922
** = Halimede Oberthür & Houlbert, 1922
** = Lachesis Oberthür & Houlbert, 1922
** = Argeformia Verity, 1953

### Satyrina

* Satyrus Latreille, 1810
* Arethusana de Lesse, 1951
* Argestina Riley, 1923
* Aulocera Butler, 1867
* Austroypthima Holloway, 1974
* Berberia de Lesse, 1951
* Boerebia Prout, 1901
** = Erebomorpha Elwes, 1899 non Erebomorpha Walker, 1860 (Geometridae)
* Brintesia Fruhstorfer, 1911
** = Oreas Hübner, 1806
** = Oreas Oken, 1815
* Callerebia Butler, 1867
* Cassionympha van Son, 1955
* Chazara Moore, 1893
** = Philareta Moore, 1893 non Philareta Moore, 1881 (Noctuidae)
* Coenyra Hewitson, 1865
* Coenyropsis van Son, 1958
* Davidina Oberthür, 1879
** = Leechia Röber, 1907 non Leechia South, 1901 (Crambidae), nec Leechia Tutt, 1907 (Lycaenidae)
** = Sinosatyrus Lee
* Hipparchia Fabricius, 1807
** = Eumenis Hübner, 1819
** = Nytha Billberg, 1820
** = Melania Sodoffsky, 1837
** = Pseudotergumia Agenjo, 1947
** = Neohipparchia de Lesse, 1951
** = Parahipparchia Kurdna, 1977
** = Euhipparchia Kudrna, 1977
* Kanetisa Moore, 1893
* Karanasa Moore, 1893
* Loxerebia Watkins, 1925
** = Hemadara Moore, 1893
* Mashuna van Son, 1955
** = Ypthimorpha Overlaet, 1955
* Melampias Hübner, 1819
* Minois Hübner, 1819
* Neita van Son, 1955
* Neocoenyra Butler, 1886
* Neominois Scudder, 1875
* Oeneis Hübner, 1819
** = Chionobas Boisduval, 1833
* Paralasa Moore, 1893
* Paroeneis Moore, 1893
* Physcaeneura Wallengren, 1857
** = Periplysia Gerstaecker, 1871
* Pseudochazara de Lesse, 1951
* Pseudonympha Wallengren, 1857
* Strabena Mabille, 1877
** = Callyphthima Butler, 1880
* Stygionympha van Son, 1955
* Ypthima Hübner, 1818
** = Xois Hewitson, 1865
** = Kolasa Moore, 1893
** = Thymipa Moore, 1893
** = Nadiria Moore, 1893
** = Pandima Moore, 1893
** = Lohana Moore, 1893
** = Dallacha Moore, 1893
** = Shania Evans, 1912
** = Sundaypthima Uemura, 1996
* Ypthimomorpha van Son, 1955

### incertae sedis

* Dyndirus Capronnier, 1874
* Setodocis Billberg, 1820 (genus dubium)

## Stamboom

Het onderstaande cladogram geeft de verwantschappen tussen de subtribus van deze geslachtengroep aan zoals die volgens Marin et al. (2011) moeten worden gezien:
| Satyrini | Parargina Lethina Mycalesina Eritina Ragadiina Coenonymphina + Hypocystina Euptychiina Erebiina Maniolina Yptimina Pronophilina Melanargiina Satyrina |
* ↑ Peña, C., N. Wahlberg, E. Weingartner, U. Kodandaramaiah, S. Nylin, A.V.L. Freitas & A.V.Z. Brower (2006). Higher level phylogeny of Satyrinae butterflies (Lepidoptera: Nymphalidae) based on DNA sequence data. Molecular Phylogenetics and Evolution 40: 29-49.
* ↑ The higher classification of Nymphalidae, The Nymphalidae Systematics Group.
* ↑ Een asterisk betekent dat dit geslacht geen deel uitmaakte van het fylogenetisch onderzoek door Pena et al. (2006).
* 1 2 Door Peña (2006) in Coenonymphina geplaatst
* ↑ Marin, M.A., Peña, C., Freitas, A.V.L., Wahlberg, N. & Uribe, S.I. (2011). From the phylogeny of the Satyrinae butterflies to the systematics of Euptychiina (Lepidoptera, Nymphalidae): history, progress and prospects. Neotropical Entomology 40: 1-13.