# Pterostichus

Pterostichus is een geslacht van kevers uit de familie van de loopkevers (Carabidae). De wetenschappelijke naam van het geslacht is voor het eerst geldig gepubliceerd in 1810 door Bonelli.

## Soorten

Deze lijst van 1174 stuks is mogelijk niet compleet.
* P. aanistschenkoi (O. Berlov & E. Berlov, 1999)
* P. aapsorum (Belousov, 1991)
* P. abaciformis (Straneo, 1955)
* P. abagonensis (Reitter, 1896)
* P. abasarukini (O. Berlov & E. Berlov, 1996)
* P. abasinus (Belousov, 1991)
* P. abaxoides (Dejean, 1828)
* P. abensis (Morita, 2010)
* P. abishirensis (Belousov, 1991)
* P. abnormis (J.R.Sahlberg, 1880)
* P. abudarini (O. Berlov & E. Berlov, 1997)
* P. acuspina (Tschitscherine, 1901)
* P. acuspinus (Tschitscherine, 1901)
* P. acutidens (Fairmaire, 1889)
* P. acutipes (Barr, 1971)
* P. adatarasanus (Sasakawa, 2005)
* P. ademidovae (O. Berlov, 1996)
* P. adoxus (Say, 1823)
* P. adstrictus (Eschscholtz, 1823)
* P. adulterinus (Wrase & J.Schmidt, 2006)
* P. aemiliae (Facchini & Sciaky, 2000)
* P. aeneocupreus (Fairmaire, 1887)
* P. aenescens (Chaudoir, 1850)
* P. aeneus (Fischer von Waldheim, 1829)
* P. aereipennis (Solsky, 1871)
* P. aethiops (Panzer, 1796)
* P. agilis (Allegro & Sciaky, 2010)
* P. agonus (G.horn, 1880)
* P. aibgensis (Starck, 1890)
* P. aimaki (Jedlička, 1968)
* P. ainus (Plutenko, 2005)
* P. akitai (Morita, 2004)
* P. akkusianus (Kirschenhofer, 1981)
* P. akozyrevi (O. Berlov & E. Berlov, 1999)
* P. aksekianus (Straneo, 1988)
* P. alacer (A. Morawitz, 1862)
* P. alamedae (Casey, 1918)
* P. alexandrovi (Lafer, 1979)
* P. alexeji (Zamotajlov & Kryzhanovskij, 1992)
* P. algidus (LeConte, 1852)
* P. altaianus (Jedlička, 1958)
* P. altaicus (Germar, 1824)
* P. altaiensis (Poppius, 1906)
* P. amagisanus (Tanaka & Ishida, 1972)
* P. amanoi (Nakane, 1968)
* P. amaroides (Dejean, 1828)
* P. ambigenus (Bates, 1883)
* P. ambiguus (Fairmaire, 1858)
* P. amblypterus (Chaudoir, 1868)
* P. amethystinus (Mannerheim, 1843)
* P. amoenus (Dejean, 1828)
* P. amurensis (Poppius, 1906)
* P. analis (Jedlička, 1963)
* P. anatolicus (Jedlička, 1963)
* P. andreae (Tschitscherine, 1897)
* P. andreinii (Dodero, 1922)
* P. andrewesi (Jedlička, 1931)
* P. angoarnigi (Morvan, 1994)
* P. angustus (Dejean, 1828)
* P. annosus (Casey, 1913)
* P. anomostriatus (Sciaky, 1995)
* P. anthracinus (Illiger, 1798)
       Oeverzwartschild
* P. apenninus (Dejean, 1831)
* P. apfelbecki (Csiki, 1908)
* P. apiculatiphallus (Nemoto, 1988)
* P. aralarensis (Espanol & Mateu, 1945)
* P. araraticus (Kirschenhofer, 1987)
* P. arator (Faldermann, 1836)
* P. arcanoides (Lorens, 1998)
* P. arcanus (Casey, 1913)
* P. arcticola (Chaudoir, 1868)
* P. arcuaticarinatus (Kasahara, 1986)
* P. archangaicus (Shilenkov, 2000)
* P. argutoriformis (Poppius, 1906)
* P. armenus (Faldermann, 1836)
* P. arnoldii (Erjiomin, 1996)
* P. arribasi (Ortuno, 1991)
* P. arrowi (Jedlička, 1936)
* P. arrowianus (Jedlička, 1938)
* P. arsenjevi (Lafer, 1979)
* P. arvernus (Jeannel, 1937)
* P. asahinus (Habu & Baba, 1960)
* P. asiaticus (Poppius, 1906)
* P. asturicus (Jeanne, 1965)
* P. astutus (Tschitscherine, 1903)
* P. asymmetricus (Bates, 1883)
* P. aterrimus (Herbst, 1784)
       Veenzwartschild
* P. atratus (Gistel, 1857)
* P. atrox (Andrewes, 1937)
* P. aubryi (Coiffait, 1952)
* P. audax (Tschitscherine, 1895)
* P. auratus (Heer, 1837)
* P. auriga (Ball, 1962)
* P. avaricus (Kryzhanovskij & Abdurakhmanov, 1983)
* P. baenningeri (Jedlička, 1931)
* P. balachowskyi (Morvan, 1972)
* P. baldwini (Casey, 1924)
* P. balikliensis (Hovorka & Skoupy, 2007)
* P. balthasari (Jedlička, 1937)
* P. balu (Schmidt, 2009)
* P. bandotaro (Tanaka, 1958)
* P. bannaticus (Dejean, 1828)
* P. barbarae (Sciaky & Facchini, 2003)
* P. barbarinus (Casey, 1913)
* P. barbatus (Fischer von Waldheim, 1829)
* P. bargusinicus (Shilenkov, 2000)
* P. barlensis (Straneo, 1935)
* P. barri (Bousquet, 2006)
* P. barryorum (Ball, 1962)
* P. basilobatus (Sasakawa, 2005)
* P. basipunctatus (Straneo, 1955)
* P. batesi (Tschitscherine, 1894)
* P. belizini (Lutshnik, 1933)
* P. belousovi (Kryzhanovskij, 1989)
* P. bellatrix (Tschitscherine, 1895)
* P. beneshi (Sciaky, 1996)
* P. berezowskii (Tschitscherine, 1898)
* P. besucheti (Straneo, 1988)
* P. beyeri (Vandyke, 1925)
* P. bhadravati (Jedlička, 1965)
* P. bhutanensis (Morvan, 1980)
* P. bicolor (Fischer von Waldheim, 1829)
* P. bielzii (Fuss, 1858)
* P. biexcisus (Straneo, 1955)
* P. bifidiphallus (Nemoto, 1988)
* P. bifoveolatus (Park; Kwon & Lafer, 1996)
* P. binotatus (Fischer von Waldheim, 1829)
* P. biocryus (Ball, 1962)
* P. bischoffianus (Jedlička, 1936)
* P. bisetosus (Straneo, 1938)
* P. bituberculatus (Tschitscherine, 1899)
* P. blandulus (L. Miller, 1859)
* P. blodgettensis (Will, 2007)
* P. blumenthali (Heinz, 1965)
* P. bogatshevi (Kryzhanovskij, 1983)
* P. borcka (Jedlička, 1963)
* P. bousqueti (Bergdahl, 2011)
* P. bowanus (Sciaky, 1997)
* P. brancuccii (Straneo, 1982)
* P. breuili (Jeannel, 1919)
* P. brevicornis (Kirby, 1837)
* P. brevilama (Straneo, 1989)
* P. brevipennis (Chevrolat, 1840)
* P. brevis (Duftschmid, 1812)
* P. breviusculus (R.F. Sahlberg, 1844)
* P. brittoni (Habu, 1958)
* P. bruckii (Schaum, 1859)
* P. brunneipennis (Straneo, 1955)
* P. bryanti (Vandyke, 1951)
* P. bryantoides (Ball, 1962)
* P. bucolicus (Casey, 1913)
* P. buglaniensis (Kirschenhofer, 1981)
* P. bullatus (Allegro & Sciaky, 2010)
* P. bungei (Tschitscherine, 1894)
* P. bureli (Morvan, 1994)
* P. burjaticus (Poppius, 1906)
* P. burkhan (O.Berlov & Anistschenko, 2005)
* P. burmeisteri (Heer, 1838)
* P. bythiniensis (Maran, 1944)
* P. cacumensis (Ball, 1966)
* P. californicus (Dejean, 1828)
* P. caligans (G. Horn, 1891)
* P. calvitarsis (Breit, 1912)
* P. campania (Andrewes, 1937)
* P. campbelli (Bousquet, 1985)
* P. canallatus (Casey, 1913)
* P. cancellatus (Motschulsky, 1860)
* P. cantabricus (Schaufuss, 1862)
* P. cantalicus (Chaudoir, 1868)
* P. capitatus (Chaudoir, 1850)
* P. capito (Tschitscherine, 1900)
* P. capitolinus (Kurnakov, 1962)
* P. caribou (Ball, 1962)
* P. carinatus (Sciaky & Facchini, 2003)
* P. carinthiacus (Meschnigg, 1946)
* P. carolinus (Darlington, 1931)
* P. carradei (Gautier des Cottes, 1866)
* P. carri (Angus, Gallan, Wrase & Chaladze, 2009)
* P. carsticus (Kasahara & Y.ito, 1989)
* P. casalei (Straneo, 1984)
* P. casaleianus (Kirschenhofer, 1991)
* P. caspius (Menetries, 1832)
* P. castaneus (Dejean, 1828)
* P. castanipes (Menetries, 1843)
* P. castor (Goulet & Bousquet, 1983)
* P. catalonicus (J.Daniel, 1906)
* P. catei (Sciaky & Wrase, 1997)
* P. cathaicus (Sciaky, 1994)
* P. caucasicola (Tschitscherine, 1893)
* P. caucasicus (Menetries, 1832)
* P. caudicalis (Say, 1823)
* P. cavazzutii (Allegro & Sciaky, 2010)
* P. cecchiniae (Jakobson, 1907)
* P. cervenkai (Sciaky, 1994)
* P. circulosus (Lindroth, 1966)
* P. clepsydra (Sciaky & Wrase, 1997)
* P. cochlearis (Hacker, 1968)
* P. cognatus (Dejean, 1831)
* P. colasi (Jeannel, 1937)
* P. colchicus (Chaudoir, 1850)
* P. colonus (Bates, 1883)
* P. collaris (Motschulsky, 1844)
* P. cometes (Andrewes, 1936)
* P. commixtiformis (Roubal, 1920)
* P. commutabilis (Motschulsky, 1866)
* P. comorus (Jedlička, 1932)
* P. compressus (Rost, 1892)
* P. comptorius (Gistel, 1857)
* P. confucius (Sciaky & Wrase, 1997)
* P. congestus (Menetries, 1843)
* P. consanguineus (Chaudoir, 1878)
* P. consors (Tschitscherine, 1893)
* P. convexus (Gebler, 1847)
* P. coracinus (Newman, 1838)
* P. cordaticollis (Heyden, 1884)
* P. cordatissimus (Straneo, 1937)
* P. cordifer (Reitter, 1896)
* P. coreicus (Jedlička, 1962)
* P. corrusculus (LeConte, 1873)
* P. coruscus (Tschitscherine, 1895)
* P. corvinus (Dejean, 1828)
* P. costatus (Menetries, 1851)
* P. crassiusculus (Chaudoir, 1868)
* P. craterensis (Hatch, 1949)
* P. cratocephalus (Tschitscherine, 1897)
* P. crebrpunctatus (Stranao, 1937)
* P. crenicollis (LeConte, 1873)
* P. crenulatopunctatus (R.F.Sahlberg, 1844)
* P. creper (Tschitscherine, 1902)
* P. cribratus (Dejean, 1828)
* P. cristatoides (Straneo, 1955)
* P. cristatus (L. Dufour, 1820)
       Bronboszwartschild
* P. cristicaudis (Kurnakov, 1962)
* P. cryobioides (Chaudoir, 1868)
* P. cucujinus (Reitter, 1892)
* P. cuneatulus (Casey, 1913)
* P. cursor (Dejean, 1828)
* P. curtatus (Fairmaire, 1886)
* P. curvatus (Sciaky & Facchini, 2003)
* P. cylindricus (Herbst in Füessly, 1784)
* P. cylindriformis (Reitter, 1887)
* P. chainapaani (J.Schmidt, 2005)
* P. chameleon (Motschulsky, 1866)
* P. champenoisi (Croissandeau, 1893)
* P. championi (Andrewes, 1926)
* P. chasautianus (Vysoky, 1984)
* P. chechcirensis (Lafer, 1979)
* P. chefsuricus (Reitter, 1896)
* P. chenpengi (Li, 1992)
* P. chinkiangensis (Kirschenhofer, 1991)
* P. chipewyan (Ball, 1962)
* P. chogyesanus (Park; Kwon & Lafer, 1996)
* P. chokaisanus (Sasakawa, 2009)
* P. chotanensis (Tschitscherine, 1893)
* P. chujoi (Habu, 1959)
* P. chujoiellus (Jedlička, 1962)
* P. chunghusorum (Lutshnik, 1922)
* P. chungkingi (Jedlička, 1932)
* P. chydaeus (Tschitscherine, 1897)
* P. daghestanus (Reitter, 1896)
* P. daihizanus (Ishida, 1968)
* P. daisenicus (Ishida, 1958)
* P. dandongensis (Kirschenhofer, 1997)
* P. dandonis (Kasahara, 1989)
* P. darvasicus (Kryzhanovskij, 1968)
* P. dauricus (Gebler, 1832)
* P. davidi (Tschitscherine, 1897)
* P. davshensis (Shilenkov, 2000)
* P. decastriensis (Lafer, 1979)
* P. deceptrix (Tschitscherine, 1898)
* P. defossus (Bates, 1883)
* P. deino (Will, 2007)
* P. dentellus (Facchini & Sciaky, 2003)
* P. denticaudis (Kurnakov, 1962)
* P. dentifer (Allegro & Sciaky, 2010)
* P. depilatus (Bates, 1889)
* P. depressidorsis (Reitter, 1896)
* P. deuvei (Lassalle, 1985)
* P. deuvesianus (Morvan, 1995)
* P. devillei (Puel, 1924)
* P. dhorpatanicus (Straneo, 1977)
* P. diabolus (Casey, 1913)
* P. diligendus (Chaudoir, 1868)
* P. diligens (Sturm, 1824)
       Gladde zwartschild
* P. dilutipes (Motschulsky, 1844)
* P. dipojranus (Straneo, 1988)
* P. discrepans (A.Morawitz, 1862)
* P. dissimilis (A. & G.B.Villa, 1833)
* P. distinctissimus (Jedlička, 1940)
* P. diversus (Fairmaire, 1886)
* P. dolens (Tschitscherine, 1900)
* P. doris (Jedlička, 1962)
* P. dorjulensis (Morvan, 1978)
* P. dostali (Kirschenhofer, 1981)
* P. drescoi (Negre, 1957)
* P. drescheri (Fischer, 1817)
* P. dufourii (Dejean, 1828)
* P. dulcis (Bates, 1883)
* P. dundai (Sciaky, 1994)
* P. duratii (A. & G.B.Villa, 1835)
* P. dux (L.Schaufuss, 1862)
* P. ebeninus (Dejean, 1828)
* P. ebenus (Quensel, 1806)
* P. eboshiyamanus (Sasakawa, 2009)
* P. ecarinatus (Hatch, 1936)
* P. ehlersi (Heyden, 1881)
* P. elmbergi (Poppius, 1908)
* P. elongatus (Duftschmid, 1812)
* P. emei (Sciaky, 1994)
* P. empetricola (Dejean, 1828)
* P. enasanus (Morita, 2007)
* P. enodis (Bousquet, 1992)
* P. enyo (Will, 2007)
* P. eobius (Tschitscherine, 1899)
* P. eoyoritomus (Sasakawa, 2009)
* P. epiroticus (Csiki, 1930)
* P. eriwanicus (Tschitscherine, 1897)
* P. esakii (Ishida, 1959)
* P. eschscholtzii (Germar, 1824)
* P. espanoli (J. Vives & E. Vives, 1977)
* P. esuriens (Casey, 1913)
* P. ethnicus (Gistel, 1857)
* P. euxinus (Straneo, 1935)
* P. exceptus (J.R. Sahlberg, 1887)
* P. exilis (Mateu, 2001)
* P. eximius (A. Morawitz, 1862)
* P. exochus (Andrewes, 1930)
* P. expeditus (Tschitscherine, 1898)
* P. externepunctatus (Dejean, 1828)
* P. falcispinus (Sasakawa, 2005)
* P. faldermanni (Schatzmayr, 1929)
* P. fallettii (Sciaky, 1996)
* P. falli (Vandyke, 1925)
* P. farkaci (Sciaky, 1997)
* P. fasciatopunctatus (Creutzer, 1799)
* P. faunus (Kurnakav, 1962)
* P. femoralis (Kirby, 1837)
* P. fencli (Dvorak, 1995)
* P. fenestratus (Kasahara & Ohtani, 1989)
* P. fenyesi (Csiki, 1930)
* P. ferghanicus (Kabak, 2001)
* P. ferreri (Espanol & Mateu, 1942)
* P. filipjevi (Lutshnik, 1928)
* P. filum (Tschitscherine, 189, 1900)
* P. findelii (Dejean, 1828)
* P. firmus (Lutshnik, 1933)
* P. flavofemoratus (Dejean, 1828)
* P. focarilei (Casale & Giachino, 1985)
* P. fornicatus (Kolenati, 1845)
* P. forticornis (Fairmaire, 1888)
* P. foveolatus (Duftschmid, 1812)
* P. franzi (Negre, 1955)
* P. fritzhiekei (J.Schmidt, 1994)
* P. fuchsi (Schaeffer, 1910)
* P. fujimurai (Habu, 1958)
* P. fujisanus (Tanaka & Suga, 1972)
* P. fujitai (Tanaka & Ishida, 1972)
* P. fuligineus (A. Morawitz, 1862)
* P. fulvescens (Motschulsky, 1844)
* P. funakoshii (Morita, 2007)
* P. funestes (Csiki, 1930)
* P. fuscicornis (Reiche & Saulcy, 1855)
* P. gagates (Hope, 1831)
* P. galae (Farkac & Plutenko, 1996)
* P. galaecianus (Lauffer, 1909)
* P. galinae (Kabak, 1992)
* P. gallega (Fairmaire, 1859)
* P. gallopavo (Sciaky & Wrase, 1997)
* P. ganesh (J.Schmidt, 2005)
* P. ganja (J. Schmidt, 1995)
* P. gaoligongensis (Wrase & J.Schmidt, 2006)
* P. gassanus (Sasakawa, 2009)
* P. geberti (Sciaky & Wrase, 1997)
* P. geojensis (Sasakawa; Kim & Kubota, 2006)
* P. gerdi (Schmidt, 2009)
* P. gerstlensis (Ball, 1962)
* P. ghilarovi (Kryzhanovskij, 1988)
* P. ghilianii (Putzeys, 1846)
* P. giacomazzai (Sciaky, 1994)
* P. gibbicollis (Motschulsky, 1844)
* P. giresuni (Jedlička, 1965)
* P. glabricollis (Jedlička, 1962)
* P. glabripennis (Jedlička, 1962)
* P. glacialis (Brisout de Barneville, 1863)
* P. glaferi (O.Berlov & E.Berlov, 1996)
* P. gliscans (Casey, 1913)
* P. gobettii (Straneo, 1983)
* P. gompanus (Straneo, 1983)
* P. gongga (Sciaky, 1997)
* P. goriensis (Tschitscherine, 1897)
* P. goschi (Jedlička, 1930)
* P. gotoensis (Kasahara & Matsumoto, 1990)
* P. gracilior (LeConte, 1873)
* P. gracilis (Dejean, 1828)
       Smalle zwartschild
* P. grajus (Dejean, 1828)
* P. gravis (Jedlička, 1939)
* P. gregalis (Casey, 1913)
* P. gromykoi (Sundukov, 2005)
* P. guizhouensis (Sciaky, 1997)
* P. gwervaenus (Morvan, 1995)
* P. gyrosus (Motschulsky, 1866)
* P. haesitatus (Fairmaire, 1888)
* P. haftorni (Lindroth, 1969)
* P. hagenbachii (Sturm, 1824)
* P. hakusanus (Kasahara, 1989)
* P. haptoderoides (Tschitscherine, 1889)
* P. harmandi (Tschitscherine, 1900)
* P. harponifer (Tanaka, 1987)
* P. hartmanni (J.Schmidt, 1995)
* P. hatchi (Hacker, 1968)
* P. hedkeinekus (Morvan, 1995)
* P. heilongjiangensis (Li, 1992)
* P. heinzi (Jedlička, 1965)
* P. heinzianus (Sciaky, 1996)
* P. heptapotamicus (Lutshnik, 1927)
* P. herculaneus (Mannerheim, 1843)
* P. herzi (Poppius, 1906)
* P. hidanus (Morita, 2010)
* P. hikosanus (Kasahara, 1994)
* P. himifuriho (Morita, 2007)
* P. hiramatsui (Morita, 2010)
* P. hirasawai (Morita, 2010)
* P. hirobane (Habu & Baba, 1958)
* P. hisamatsui (Ishida & Shibata, 1961)
* P. hoelli (Schmidt, 2009)
* P. homalonotus (Tschitscherine, 1894)
* P. honnoratii (Dejean, 1828)
* P. hoplites (Bates, 1883)
* P. hornii (LeConte, 1873)
* P. horvatovichi (Kirschenhofer, 1991)
* P. hozumii (Ishida, 1961)
* P. huashanus (Sciaky, 1994)
* P. hubeicus (Facchini & Sciaky, 2003)
* P. hudsonicus (LeConte, 1863)
* P. humboldti (Casey, 1913)
* P. humidulus (Vandyke, 1943)
* P. humilis (Casey, 1913)
* P. hummeli (Jedlička, 1935)
* P. hungaricus (Dejean, 1828)
* P. hypogeus (Barr, 1971)
* P. iberlovi (O.Berlov, 1996)
* P. idahoae (Csiki, 1930)
* P. ienishteai (Nitzu, 1988)
* P. ignavus (Tschitscherine, 1897)
* P. ikizderensis (Hovorka & Skoupy, 2007)
* P. ikukoae (Morita, 2007)
* P. illigei (Panzer, 1805)
* P. illigeri (Panzer, 1803)
* P. illustris (LeConte, 1851)
* P. imasakai (Kasahara & Ohtani, 1988)
* P. imitatus (Morvan, 1978)
* P. immarginatus (Straneo, 1977)
* P. impressicollis (Fairmaire & Laboulbene, 1854)
* P. impressus (Fairmaire & Laboulbene, 1854)
* P. imurai (Morita, 2007)
* P. inanis (G. Horn, 1891)
* P. incommodus (Schaum, 1858)
* P. inermis (Fall, 1901)
* P. infaustus (Gistel, 1857)
* P. infernalis (Hatch, 1936)
* P. infimus (Chaudoir, 1868)
* P. ingushus (Lutshnik, 1928)
* P. innshanensis (Jedlička, 1960)
* P. inopinatus (Lutshnik, 1933)
* P. inopinus (Casey, 1918)
* P. insidiatrix (Brulerie, 1872)
* P. insignicollis (Tschitscherine, 1893)
* P. intectus (Casey, 1918)
* P. interruptus (Dejean, 1828)
* P. irideus (Sciaky, 1994)
* P. iripennis (Chaudoir, 1868)
* P. isabellae (LeConte, 1851)
* P. ishiharai (Ishida & Shibata, 1961)
* P. ishikawai (Nemoto, 1988)
* P. ishikawaioides (Sasakawa; Kim; Kim & Kubota, 2008)
* P. ishizuchiensis (Kasahara, 1985)
* P. ishizukai (Kasahara, 1995)
* P. isolatus (Sasakawa; Kim; Kim & Kubota, 2008)
* P. isumiensis (Kasahara & Saito, 1997)
* P. itoi (Kasahara, 1986)
* P. iwakiensis (Sasakawa, 2009)
* P. ixion (Tschitscherine, 1902)
* P. jacobinus (Casey, 1913)
* P. jacobsoni (Poppius, 1906)
* P. jaechi (Kirschenhofer, 1997)
* P. jaechianus (Sciaky & Facchini, 2003)
* P. jakobsonianus (Lutshnik, 1928)
* P. janatai (Sciaky & Wrase, 1997)
* P. janbritoi (Morvan, 1994)
* P. jani (Sciaky & Facchini, 2003)
* P. jankowskyi (Tschitscherine, 1897)
* P. janoi (Jedlička, 1952)
* P. japonicus (Motschulsky, 1860)
* P. jarkendis (Jedlička, 1965)
* P. jelepus (Andrewes, 1932)
* P. jenisseicus (Csiki, 1930)
* P. jiricola (Sasakawa; Kim; Kim & Kubota, 2008)
* P. jogaesanensis (Lafer; Paik & Park, 1996)
* P. johnsoni (Ulke, 1889)
* P. josephi (Csiki, 1930)
* P. juga (J.schmidt, 2005)
* P. jugicola (Lutshnik, 1916)
* P. jugivagus (Tschitscherine, 1898)
* P. jungens (Tschitscherine, 1893)
* P. jureceki (Jedlička, 1936)
* P. jurecekianus (Maran, 1940)
* P. jurinei (Panzer, 1803)
* P. justusii (W.Redtenbacher, 1842)
* P. kadjaranze (Jedlička, 1947)
* P. kadleci (Dvorak, 1995)
* P. kadoudali (Morvan, 1981)
* P. kajimurai (Habu & Tanaka, 1957)
* P. kalabi (Sciaky, 1994)
* P. kalhys (Motschulsky, 1866)
* P. kalo (Schmidt, 2012)
* P. kamtschaticus (Motschulsky, 1860)
* P. kangchenjunga (Schmidt, 2012)
* P. kaniei (Morita, 2005)
* P. kaninensis (Poppius, 1906)
* P. kansanus (Casey, 1918)
* P. karasawai (Tanaka, 1958)
* P. karateghinicus (Mikhailov, 1972)
* P. kataevi (Kryzhanovskij, 1989)
* P. katashinensis (Habu, 1958)
* P. kavanaughi (Shilenkov, 2000)
* P. kaynashensis (Schweiger, 1967)
* P. keltiekus (Morvan, 1981)
* P. kerzhneri (Lafer, 1983)
* P. kiangsu (Jedlička, 1965)
* P. kiiensis (Morita & Ohkura, 1988)
* P. kimurai (Morita, 1994)
* P. kiritshenkoi (Lutshnik, 1928)
* P. kirschenblatti (Kryzhanovskij, 1988)
* P. kitakamisanus (Sasakawa, 2005)
* P. klapperichi (Jedlička, 1956)
* P. kleinfeldi (Straneo, 1982)
* P. kleinfeldianus (Sciaky & Wrase, 1997)
* P. koenigi (Reitter, 1887)
* P. koenigianus (Tschitscherine, 1890)
* P. koheii (Nakane, 1963)
* P. kokeilii (L.Miller, 1850)
* P. kolosovi (Lutshnik, 1928)
* P. kolymensis (Erjiomin, 1998)
* P. komalus (Jedlička, 1936)
* P. kongosanus (Nakane, 1963)
* P. kopetzi (J.Schmidt, 2005)
* P. korgei (Jedlička, 1964)
* P. korolkowi (Tschitscherine, 1901)
* P. kosakai (Morita, 1998)
* P. kotzebuei (Ball, 1962)
* P. krali (Sciaky, 1997)
* P. krasnopolensis (Kirschenhofer, 1987)
* P. krulikovskyi (Lutshnik, 1933)
* P. kubanicus (Kirschenhofer, 1987)
* P. kucerai (Sciaky, 1997)
* P. kultianus (Jedlička, 1946)
* P. kuraicus (Shilenkov, 2000)
* P. kurentzovi (Lafer, 1979)
* P. kurikomasanus (Sasakawa, 2005)
* P. kurosai (Tanaka, 1958)
* P. kurosawai (Tanaka, 1958)
* P. kvirensis (Belousov, 1991)
* P. kyushuensis (Habu, 1955)
* P. laborans (Casey, 1913)
* P. labzuki (Lafer, 1979)
* P. lacertus (Casey, 1913)
* P. lacunosus (Chaudoir, 1844)
* P. lachrymosus (Newman, 1838)
* P. laevipunctatus (Tschitscherine, 1889)
* P. laferi (O. Berlov & E. Berlov, 1996)
* P. lama (Menetries, 1843)
* P. lamuticus (Poppius, 1906)
* P. lanei (Vandyke, 1925)
* P. lanista (Tschitscherine, 1898)
* P. lasserrei (Dejean, 1828)
* P. lassulus (Casey, 1920)
* P. latemarginatus (Straneo, 1936)
* P. laticollis (Motschulsky, 1844)
* P. latifianus (Apfelbeck, 1906)
* P. latistylis (Tanaka, 1958)
* P. latitemporis (Sciaky & Wrase, 1997)
* P. latiusculus (Chaudoir, 1868)
* P. lattini (Labonte, 2006)
* P. ledenikensis (Knirsch, 1925)
* P. leica (Schmidt, 2012)
* P. leonisi (Apfelbeck, 1904)
* P. leptis (Bates, 1873)
* P. letensis (Habu, 1973)
* P. levadensis (Lafer, 1979)
* P. leviculus (Andrewes, 1933)
* P. licenti (Jedlička, 1939)
* P. liciniformis (Csiki, 1930)
* P. lidara (Andrewes, 1937)
* P. lineatopunctatus (L.Miller, 1850)
* P. lingshanus (Sciaky & Wrase, 1997)
* P. liodactylus (Tschitscherine, 1898)
* P. lobatus (Hacker, 1968)
* P. lodosi (Heinz, 1977)
* P. lolo (Bergdahl, 2011)
* P. lombardus (K.Daniel, 1901)
* P. longicollis (Duftschmid, 1812)
       Langnekzwartschild
* P. longinquus (Bates, 1873)
* P. longipennis (Straneo, 1942)
* P. longipes (Poppius, 1906)
* P. longiusculus (J.Sahlberg, 1880)
* P. lubricus (Casey, 1918)
* P. lucidus (Motschulsky, 1844)
* P. luctuosus (Dejean, 1828)
* P. lugi (Morvan, 1982)
* P. lukjanovitshi (Lutshnik, 1927)
* P. lumensis (Apfelbeck, 1905)
* P. lustrans (LeConte, 1851)
* P. lutschniki (Jedlička, 1962)
* P. lutshnikianus (Bogachev & Kurnakov, 1958)
* P. lyroderus (Chaudoir, 1846)
* P. macedonicus (Apfelbeck, 1918)
* P. macer (Marsham, 1802)
       Platte zwartschild
* P. macrocephalus (Habu, 1955)
* P. macrogenys (Bates, 1883)
* P. macrothorax (Poppius, 1906)
* P. machulkai (Jedlička, 1931)
* P. maderi (Jedlička, 1938)
* P. madidus (Fabricius, 1775)
       Rondhalszwartschild
* P. magoides (Straneo, 1937)
* P. magus (Mannerheim, 1825)
* P. maichensis (Lafer, 1979)
* P. makalu (Schmidt, 2012)
* P. makolskii (J.Muller, 1933)
* P. malissorum (Apfelbeck, 1905)
* P. malkini (Hatch, 1953)
* P. mancus (LeConte, 1852)
* P. mandibularoides (Ball, 1966)
* P. mandzhuricum (Lutshnik, 1916)
* P. mandzhuricus (Lutshnik, 1916)
* P. mannerheimii (Dejean, 1831)
* P. marani (Jedlička, 1967)
* P. marginatus (Matsumura, 1911)
* P. marginellus (Sasakawa; Kim & Kim, 2006)
* P. mariae (Lutshnik, 1921)
* P. marinensis (Hacker, 1968)
* P. masahiroi (Kasahara, 1988)
* P. masatakai (Morita, 2007)
* P. masidai (Ishida, 1959)
* P. masumotoi (Tanaka; Morita & Suga, 1987)
* P. matchai (Jedlička, 1930)
* P. matsumurai (Habu, 1973)
* P. mattolensis (Hacker, 1968)
* P. matveichuki (O. Berlov, 1996)
* P. maurusiacus (Mannerheim, 1825)
* P. maximus (Tschitscherine, 1889)
* P. mediocris (Dvorak, 1995)
* P. medvedevi (Kryzhanovskij & Mikhailov, 1972)
* P. megaloderus (Sciaky, 1994)
* P. meisteri (Reitter, 1885)
* P. melanarius (Illiger, 1798)
       Gewone zwartschild
* P. melanodes (Chaudoir, 1878)
* P. melas (Creutzer, 1799)
* P. mellyi (Gebler, 1843)
* P. menetriesii (LeConte, 1873)
* P. mercedianus (Casey, 1918)
* P. merklii (J.Frivaldszky, 1879)
* P. meskheticus (Belousov, 1991)
* P. metallifer (Gistel, 1857)
* P. mewakholensis (Schmidt, 2012)
* P. meyeri (Jedlička, 1934)
* P. micans (Heer, 1838)
* P. microbus (Sciaky & Facchini, 2003)
* P. microcephalus (Motschulsky, 1860)
* P. microphthalmus (Delarouzee, 1857)
* P. microps (Heyden, 1887)
* P. michailovi (Wrase, 1992)
* P. middendorffi (J.R. Sahlberg, 1875)
* P. miehei (J.Schmidt, 2005)
* P. migliaccioi (Straneo, 1982)
* P. militaris (Tschitscherine, 1893)
* P. mimus (Casey, 1913)
* P. ming (Sciaky & Wrase, 1997)
* P. minor (Gyllenhal, 1827)
       Moeraszwartschild
* P. mirabilis (Jedlička, 1939)
* P. mirificus (Bates, 1883)
* P. miroshnikovi (Zamotajlov, 1991)
* P. miroslavi (Sciaky & Wrase, 1997)
* P. mirus (Tschitscherine, 1894)
* P. miscellus (Casey, 1913)
* P. mitoyamanus (Tanaka, 1971)
* P. miyamai (Kasahara & Y. Ito, 1987)
* P. mizunoyai (Morita, 2007)
* P. modicellus (Tschitscherine, 1897)
* P. moestus (Say, 1823)
* P. molopsoides (Jedlička, 1934)
* P. monostigma (Tschitscherine, 1898)
* P. montanus (Motschulsky, 1844)
* P. monticoloides (Shilenkov, 1995)
* P. montigena (Tschitscherine, 1898)
* P. montisdeorum (Schweiger, 1967)
* P. morawitzianus (Lutshnik, 1922)
* P. morio (Duftschmid, 1812)
* P. morionides (Chaudoir, 1868)
* P. mosaicus (Sasakawa, 2005)
* P. mucronatus (Straneo, 1955)
* P. muehlfeldii (Duftschmid, 1812)
* P. muelleri (Straneo, 1936)
* P. muellermotzfeldi (Wrase & J.Schmidt, 2006)
* P. mujahedeeni (Savich, 1999)
* P. mukdenensis (Breit, 1933)
* P. mulensis (Sciaky, 1997)
* P. multinodosus (Sasakawa; Kim & Kim, 2006)
* P. multipunctatus (Dejean, 1828)
* P. multiseta (Straneo, 1984)
* P. mundatus (Jedlička, 1940)
* P. mundus (Jedlička, 1938)
* P. musashiensis (Kasahara, 1993)
* P. mutoides (Bousquet, 1992)
* P. mutus (Say, 1823)
* P. nadari (Vuillefroy, 1893)
* P. nakanei (Straneo, 1955)
* P. namrum (Jedlička, 1963)
* P. naokii (Morita, 2003)
* P. napaea (Kasahara, 1988)
* P. nasuensis (Morita, 2007)
* P. nasui (Kasahara, 1993)
* P. necessarius (Tschitscherine, 1894)
* P. neglectus (A.Morawitz, 1862)
* P. negligens (Sturm, 1824)
* P. nemoralis (Graells, 1851)
* P. neobrunneus (Lindroth, 1966)
* P. nepalensis (Straneo, 1977)
* P. nicaeensis (A. Villa & C.B. Villa, 1835)
* P. nigellatus (Kirschenhofer, 1991)
* P. nigellus (A. Morawitz, 1862)
* P. niger (Schaller, 1783)
       Grote zwartschild
* P. nigripalpis (Poppius, 1906)
* P. nigrita (Paykull, 1790)
       Moerasboszwartschild
* P. nigrocaeruleus (Vandyke, 1925)
* P. nikodymi (Dvorak, 1995)
* P. nimbatidius (Chaudoir, 1878)
* P. nishidai (Nakane, 1989)
* P. nishiyamai (Kasahara, 1986)
* P. nivalis (R.F. Sahlberg, 1844)
* P. nivicola (Menetries, 1832)
* P. noborui (Morita, 2008)
* P. nodicornis (Fairmaire & Laboulbene, 1854)
* P. noguchii (Bates, 1873)
* P. nordqvisti (J.R. Sahlberg, 1887)
* P. novotnyorum (Vysoky, 1981)
* P. novus (Straneo, 1944)
* P. nowitzkii (Tschitscherine, 1899)
* P. obesulus (LeConte, 1873)
* P. oblongiusculus (Motschulsky, 1850)
* P. oblongoparallelus (Maran, 1932)
* P. oblongopunctatus (Fabricius, 1787)
       Bronzen boszwartschild
* P. obsidianus (Casey, 1913)
* P. occultus (Casey, 1913)
* P. ochoticus (R.F. Sahlberg, 1844)
* P. ochsi (Fagniez, 1921)
* P. odontocnemis (G.Muller, 1931)
* P. odvarkai (Dvorak, 1995)
* P. oenotrius (Ravizza, 1975)
* P. ogaensis (Morita, 1995)
* P. ohbayashii (Ishida, 1968)
* P. ohdaisanus (Nakane, 1963)
* P. ohkawai (Morita, 2010)
* P. ohkurai (Morita, 1996)
* P. ohsawacavus (Sasakawa, 2005)
* P. okiensis (Nakane, 1989)
* P. okutamae (Tanaka, 1963)
* P. olafi (J.Schmidt, 2005)
* P. olegi (Belousov, 1991)
* P. omogoensis (Nakane, 1972)
* P. ompoensis (Jedlička, 1932)
* P. opacipennis (Jedlička, 1934)
* P. ophryoderus (Chaudoir, 1878)
* P. ordinarius (Casey, 1913)
* P. ordinatus (Fischer, 1823)
* P. orduensis (Straneo, 1988)
* P. oregonus (LeConte, 1861)
* P. oreophilus (Tschitscherine, 1898)
* P. orestes (Jedlička, 1936)
* P. orientalis (Motschulsky, 1844)
* P. orion (Tschitscherine, 1901)
* P. orionis (Jedlička, 1962)
* P. orman (Jedlička, 1963)
* P. osseticus (Kirschenhofer, 1987)
* P. ottomanus (Apfelbeck, 1908)
* P. ovalipennis (Casey, 1913)
* P. ovaliphallus (Sasakawa, 2005)
* P. ovoideus (Sturm, 1824)
       Platoogzwartschild
* P. pachinus (Bates, 1883)
* P. palekhai (E. Berlov & O. Berlov, 1996)
* P. palgongsanus (Nemoto, 1988)
* P. palmi (Schaeffer, 1910)
* P. panticulatus (Casey, 1913)
* P. pantomus (Jedlička, 1958)
* P. panzeri (Panzer, 1803)
* P. paralleloides (Kirschenhofer, 1987)
* P. parasimilis (G.E. Ball, 1962)
* P. parkwon (Davies, 2004)
* P. parnassius (Schaum, 1859)
* P. parviceps (Poppius, 1906)
* P. parvicollis (Sciaky & Wrase, 1997)
* P. pasanauricus (Kirschenhofer, 1987)
* P. patruelis (Dejean, 1831)
* P. paulinoi (Vuillefroy, 1868)
* P. pavani (Straneo, 1988)
* P. pedemontanus (Ganglbauer, 1891)
* P. peilingi (Jedlička, 1937)
* P. pemphredo (Will, 2007)
* P. peninsularis (Park & Kwon, 1996)
* P. pensylvanicus (LeConte, 1873)
* P. pentheri (Apfelbeck, 1918)
* P. percontator (Reitter, 1887)
* P. pergracilis (Casey, 1920)
* P. perisi (Novoa, 1979)
* P. perlamatus (Straneo, 1989)
* P. perlutus (Jedlička, 1938)
* P. pertinax (Tschitscherine, 1895)
* P. petulans (Jedlička, 1938)
* P. pfizenmayeri (Poppius, 1906)
* P. phaeopus (Sainte-Claire Deville, 1902)
* P. phaeus (Lutshnik, 1922)
* P. phungaraziensis (Wrase & J.Schmidt, 2006)
* P. piceolus (Chaudoir, 1850)
* P. picipes (Fischer von Waldheim, 1829)
* P. pilifer (Bates, 1878)
* P. pilosus (Host, 1789)
* P. pinguedineus (Eschscholtz, 1823)
* P. piniphilus (Gistel, 1857)
* P. pithyusicus (Lutshnik, 1933)
* P. placerensis (Casey, 1918)
* P. planaticollis (Kirschenhofer, 1987)
* P. planctus (LeConte, 1852)
* P. planicola (Tschitscherine, 1899)
* P. planiusculus (Chaudoir, 1859)
* P. planus (J.R. Sahlberg, 1887)
* P. platyderus (Chaudoir, 1850)
* P. platyops (Sciaky, 1997)
* P. plesiomorphus (Nemoto, 1989)
* P. plutonicus (Casey, 1913)
* P. poecilma (Andrewes, 1937)
* P. pohnerti (Jedlička, 1934)
* P. polygenus (Bates, 1883)
* P. ponticus (Kirschenhofer, 1987)
* P. poppiusi (Semenov, 1906)
* P. poppiusianus (Jakobson, 1907)
* P. porcellus (Kurnakov, 1962)
* P. potanini (Tschitscherine, 1889)
* P. praedo (Tschitscherine, 1901)
* P. prattii (Bates, 1890)
* P. primus (Darlington, 1931)
* P. probus (Park & Kwon, 1996)
* P. procax (A. Morawitz, 1862)
* P. procephalus (Bates, 1873)
* P. procerulus (Heyden, 1880)
* P. properans (Chaudoir, 1868)
* P. protensiformis (Casey, 1924)
* P. protensipennis (Casey, 1918)
* P. protractus (LeConte, 1860)
* P. przewalskyi (Tschitscherine, 1888)
* P. pseodostuxbergi (Poppius, 1906)
* P. pseudoastutus (Hovorka & Skoupy, 2007)
* P. pseudobarbarae (Sciaky & Facchini, 2003)
* P. pseudoharmandi (Morvan, 1981)
* P. pseudopachinus (Nakane, 1963)
* P. pseudopedius (Reitter, 1887)
* P. pseudopercus (Reitter, 1889)
* P. pseudopilifer (Straneo, 1957)
* P. pseudoplatyderus (Sciaky, 1994)
* P. pseudoplatysma (Straneo, 1982)
* P. pseudorotundus (Sciaky & Facchini, 2003)
* P. pseudosinensis (Sciaky & Facchini, 2003)
* P. pulchellus (Faldermann, 1836)
* P. pumilio (Dejean, 1828)
* P. pumilus (Casey, 1913)
* P. punctatissimus (Randall, 1838)
* P. punctatus (L. Redtenbacher, 1843)
* P. punctiger (J.R. Sahlberg, 1880)
* P. punctiventris (Chaudoir, 1878)
* P. pusillus (Dejean, 1828)
* P. putus (Casey, 1913)
* P. quadraticollis (Chaudoir, 1846)
* P. quadrifoveolatus (Letzner, 1852)
       Brandzwartschild
* P. quadriimpressus (Straneo, 1983)
* P. quelpartensis (Habu, 1985)
* P. raptor (Tschitscherine, 1901)
* P. raraensis (Morvan, 1980)
* P. rasilis (Park & Kwon, 1996)
* P. regeli (Tschitscherine, 1894)
* P. reiseri (Ganglbauer, 1889)
* P. reissi (G.Muller, 1931)
* P. relictus (Newman, 1838)
* P. rengensis (Morita, 2010)
* P. restrictus (Casey, 1918)
* P. reuteri (Wrase & Schmidt, 2006)
* P. rhaeticus (Heer, 1837)
       Heidezwartschild
* P. rhanis (Tschitscherine, 1902)
* P. rhilensis (Rottenberg, 1874)
* P. riffensis (Antoine, 1933)
* P. ripensis (Motschulsky, 1866)
* P. robustistylis (Sasakawa, 2005)
* P. rodionovi (Shilenkov, 2000)
* P. rolex (Morvan, 1995)
* P. rostratus (Newman, 1838)
* P. rothi (Hatch, 1951)
* P. rotundangulus (A.Morawitz, 1862)
* P. rotundus (Sciaky, 1994)
* P. rousi (Kirschenhofer, 1982)
* P. rousianus (Kirschenhofer, 1987)
* P. rousselli (Colas, 1963)
* P. rubripalpis (Csiki, 1930)
* P. rudestriatus (Reitter, 1883)
* P. ruffoi (Sciaky, 1984)
* P. rufipes (Dejean, 1828)
* P. rufitarsis (Dejean, 1828)
* P. rufonitens (Fairmaire, 1866)
* P. rufopiceus (Heyden, 1890)
* P. rugiceps (Straneo, 1984)
* P. rugosiceps (Schmidt, 2012)
* P. rugosipennis (Jedlička, 1932)
* P. rugosus (Gebler, 1823)
* P. rutilans (Dejean, 1828)
* P. ryomoensis (Morita & Suda, 2007)
* P. sacheri (I.Frivaldszky Von Frivald, 1865)
* P. sachtlebeni (Jedlička, 1962)
* P. safonovae (Budarin, 1995)
* P. sagarmatha (Schmidt, 2012)
* P. sahlbergi (Tschitscherine, 1894)
* P. sakuraii (Morita, 2008)
* P. santostamangi (J.Schmidt, 2006)
* P. sasajii (Morita, 2007)
* P. satoi (Ishida, 1961)
* P. satsumanus (Habu, 1958)
* P. satunini (Tschitscherine, 1903)
* P. satyrus (Kurnakov, 1962)
* P. saueri (Sciaky, 1994)
* P. saxicola (Tschitscherine, 1899)
* P. sayanus (Csiki, 1930)
* P. scalptus (Sciaky & Wrase, 1997)
* P. scitus (Maklin, 1878)
* P. scurra (Tschitscherine, 1901)
* P. scutellaris (LeConte, 1873)
* P. scuticollis (Fairmaire, 1888)
* P. schaschli (Marseul, 1880)
* P. schneideri (Sciaky & Facchini, 2003)
* P. schodaicus (Kirschenhofer, 1987)
* P. schoenherri (Faldermann, 1836)
* P. schoenmanni (Kirschenhofer, 1991)
* P. schrettenbrunneri (Schmidt, 2012)
* P. schuelkei (Sciaky & Wrase, 1997)
* P. sediorcus (Sasakawa & Kim, 2006)
* P. sejunctus (Bates, 1883)
* P. sejungendus (Chaudoir, 1868)
* P. selmanni (Duftschmid, 1812)
* P. semenovi (Tschitscherine, 1888)
* P. septemtrionalis (Sciaky & Facchini, 2003)
* P. septentrionis (Chaudoir, 1868)
* P. seriatus (Chaudoir, 1850)
* P. serripes (LeConte, 1875)
* P. seticeps (Straneo, 1984)
* P. setipes (Tschitscherine, 1898)
* P. setosus (Hatch, 1951)
* P. seunglaki (Park & Kwon, 1996)
* P. seungmoi (Park & Kwon, 1996)
* P. shaovuensis (Jedlička, 1956)
* P. shavrini (O.Berlov & Anistschenko, 1999)
* P. shennongjianus (Facchini & Sciaky, 2003)
* P. shibatai (Ishida, 1961)
* P. shiibanus (Habu, 1958)
* P. shilenkovi (Erjiomin & Kabak, 1991)
* P. shimizui (Morita, 2010)
* P. shingarevi (Lafer, 1979)
* P. shirakamisan (Sasakawa, 2009)
* P. shirakamisanus (Sasakawa, 2005)
* P. shiranus (Straneo, 1984)
* P. shirozui (Habu, 1985)
* P. shotaroi (Morita, 1987)
* P. shulli (Hatch, 1949)
* P. sierranus (Casey, 1913)
* P. similis (Mannerheim, 1852)
* P. simillimus (Fairmaire, 1886)
* P. sincerus (Park & Kwon, 1996)
* P. sinensis (Jedlička, 1962)
* P. singularis (Tschitscherine, 1889)
* P. sinicus (Fairmaire, 1897)
* P. sintanus (Andrewes, 1927)
* P. skoupyi (Hovorka & Skoupy, 2007)
* P. smetanai (Bousquet, 1985)
* P. sodalicius (Heyden, 1885)
* P. sojaki (Kirschenhofer, 1987)
* P. sojot (Shilenkov, 2000)
* P. solskyi (Chaudoir, 1878)
* P. songoricus (Motschulsky, 1845)
* P. soperi (Ball, 1966)
* P. sotkaensis (Jedlička, 1958)
* P. spathifer (Bousquet, 1992)
* P. sphodriformis (Bates, 1873)
* P. sphodrinus (LeConte, 1863)
* P. spiculifer (Bates, 1883)
* P. spinolae (Dejean, 1828)
* P. sponsor (Casey, 1913)
* P. spraguei (LeConte, 1873)
* P. stagnatilis (Gistel, 1857)
* P. stantonensis (Ball, 1966)
* P. stapedius (Hacker, 1968)
* P. starcki (Heyden, 1885)
* P. starckianus (Reitter, 1887)
* P. staveni (Hovorka & Skoupy, 2007)
* P. sterbai (Jedlička, 1934)
* P. stictopleurus (Fairmaire, 1888)
* P. stoeckleini (Straneo, 1941)
* P. stomoides (Chaudoir, 1868)
* P. straneellus (Jedlička, 1938)
* P. strasseri (Reitter, 1898)
* P. strenuus (Panzer, 1796)
       Gepuncteerde zwartschild
* P. stricticollis (Solsky, 1874)
* P. strigosus (Sciaky & Wrase, 1997)
* P. stygicus (Say, 1823)
* P. styx (Andrewes, 1937)
* P. subacutus (Casey, 1918)
* P. subaeneus (Chaudoir, 1850)
* P. subgibbus (Motschulsky, 1860)
* P. subiasi (Ortuno & Zaballos, 1992)
* P. subitus (Csiki, 1930)
* P. sublaevis (J.Sahlberg, 1880)
* P. subovatus (Motschulsky, 1860)
* P. subrugosus (Straneo, 1955)
* P. subsinuatus (Dejean, 1828)
* P. subtilissimus (Sciaky, 1996)
* P. sudai (Morita, 1991)
* P. sudrei (Deville, 1922)
* P. suffusus (Casey, 1913)
* P. sugimurai (Morita, 2007)
* P. sulcitarsis (A. Morawitz, 1862)
* P. sumondakensis (Sasakawa, 2005)
* P. sungariensis (Lafer, 1979)
* P. superciliosus (Say, 1823)
* P. surgens (LeConte, 1878)
* P. suruganus (Nakane & Straneo, 1979)
* P. sutschanensis (Jedlička, 1962)
* P. svanicus (Lutshnik, 1922)
* P. swaneticus (Reitter, 1883)
* P. syleus (Kirschenhofer, 1997)
* P. symmetricus (Straneo, 1955)
* P. szekessyianus (Sciaky, 1996)
* P. tachongi (Jedlička, 1936)
* P. taebaegsanus (Nemoto, 1988)
* P. tahirai (Kasahara, 1992)
* P. tahoensis (Casey, 1913)
* P. takadateyamanus (Sasakawa, 2009)
* P. takahashii (Ishida, 1958)
* P. takaosanus (Habu, 1958)
* P. taksonyis (Csiki, 1930)
* P. talibani (Savich, 1999)
* P. tamarae (Wrase & Kirschenhofer, 1991)
* P. tamsii (Dejean, 1831)
* P. tanakai (Ishida, 1964)
* P. tanakaorum (Morita & Ohkura, 1988)
* P. tanjae (Hovorka & Skoupy, 2007)
* P. tantillus (Fairmaire, 1888)
* P. taoi (Kasahara, 1993)
* P. taradakensis (Kasahara & Ohtani, 1988)
* P. tareumiut (G. E. Ball, 1962)
* P. tarsalis (LeConte, 1873)
* P. taskylensis (Shilenkov, 2000)
* P. tatianae (E. Berlov, 1996)
* P. tatricus (Kult, 1947)
* P. tenenbaumianus (Jedlička, 1930)
* P. tenuimarginatus (Chaudoir, 1868)
* P. tenuis (Casey, 1924)
* P. teretis (Park & Kwon, 1996)
* P. testaceus (Vandyke, 1926)
* P. thanglaensis (Schmidt, 2012)
* P. theeli (Maklin, 1878)
* P. thorectes (Bates, 1873)
* P. thorectoides (Jedlička, 1958)
* P. thunbergi (A. Morawitz, 1862)
* P. tichomirovi (Erjiomin, 1990)
* P. tienmushanus (Sciaky, 1997)
* P. tiliaceoradix (Ball, 1962)
* P. todai (Morita & Kanie, 1997)
* P. tokejii (Yoshida & Tanaka, 1960)
* P. tokui (Morita, 2007)
* P. toledanoi (Facchini & Sciaky, 2003)
* P. tomensis (Gebler, 1847)
* P. tosanus (Kasahara & Y.ito, 1997)
* P. tottoriensis (Morita, 2003)
* P. toyodai (Morita & Kurosa, 1998)
* P. transversalis (Duftschmid, 1812)
* P. triangularis (Sciaky & Facchini, 2003)
* P. trinarius (Casey, 1918)
* P. trinitensis (Hacker, 1968)
* P. triseriatus (Gebler, 1847)
* P. tristis (Dejean, 1828)
* P. troglophilus (Mateu, 1997)
* P. tropicalis (Bates, 1882)
* P. trubilini (Zamotajlov, 1997)
* P. truncatus (Dejean, 1828)
* P. tsagerkericus (Lutshnik, 1928)
* P. tschitscherinianus (Jakobson, 1907)
* P. tsukubasanus (Kasahara, 1988)
* P. tsurugaensis (Ishida, 1959)
* P. tsurugiyamanus (Habu, 1959)
* P. tuberculiger (Tschitscherine, 1897)
* P. tuberculofemoratus (Hatch, 1936)
* P. tuberifer (Sasakawa In Sasakawa & Kubota, 2006)
* P. tulobalu (Schmidt, 2009)
* P. tumescens (LeConte, 1863)
* P. tundrae (Tschitscherine, 1894)
* P. tunkinensis (Shilenkov, 2000)
* P. turanensis (Jedlička, 1959)
* P. turcomannicus (Motschulsky, 1850)
* P. uchiyamai (Morita, 1987)
* P. udokanensis (Shilenkov, 2000)
* P. uedaorum (Morita & Hirasawa, 1996)
* P. uenoi (Straneo, 1955)
* P. ulubeyensis (Kirschenhofer, 1981)
* P. umbralis (Gistel, 1857)
* P. umbricola (Gistel, 1857)
* P. unctulatus (Duftschmid, 1812)
* P. uralensis (Motschulsky, 1850)
* P. urengaicus (Jurecek, 1924)
* P. ussuriensis (Tschitscherine, 1897)
* P. vagepunctatus (Heer, 1837)
* P. validiceps (Reitter, 1887)
* P. validior (Tschitscherine, 1889)
* P. validus (Dejean, 1828)
* P. vandykei (Schaeffer, 1910)
* P. variabilis (Menetries, 1832)
* P. variolatus (Dejean, 1828)
* P. variseta (Straneo, 1984)
* P. vecors (Tschitscherine, 1897)
* P. ventralis (Say, 1823)
* P. ventricosus (Eschscholtz, 1823)
* P. vermiculosus (Menetries, 1851)
* P. vernalis (Panzer, 1796)
       Groeftarszwartschild
* P. vexatus (Bousquet, 1985)
* P. vicinus (Mannerheim, 1843)
* P. virescens (Gebler, 1833)
* P. vladivostokensis (Lafer, 1979)
* P. vseteckai (Maran, 1944)
* P. walteri (Reitter, 1883)
* P. watanabei (Tanaka, 1960)
* P. weigeli (J.Schmidt, 2005)
* P. weiperti (J.Schmidt, 2005)
* P. wenxianensis (Allegro & Sciaky, 2010)
* P. wittmeri (Morvan, 1978)
* P. woongbii (Park & Kwon, 1996)
* P. woronowi (Lutshnik, 1916)
* P. wrasei (Sciaky & Facchini, 2003)
* P. xatartii (Dejean, 1828)
* P. xilingensis (Allegro & Sciaky, 2010)
* P. yahikosanus (Sasakawa, 2009)
* P. yakushimanus (Nakane & Ishida, 1961)
* P. yamajii (Kasahara, 1993)
* P. yamauchii (Morita, 1992)
* P. yamizosanus (Sasakawa, 2005)
* P. yanoi (Jedlička, 1953)
* P. yatsuensis (Straneo, 1955)
* P. ybousqueti (O.Berlov, 1999)
* P. yokohamae (Nakane & Straneo In Nakane, 1979)
* P. yoritomus (Bates, 1873)
* P. yosemitensis (Hacker, 1968)
* P. yoshidai (Kasahara, 1985)
* P. yoshikawai (Ishida, 1959)
* P. yoshizawai (Morita, 2010)
* P. yulongshanensis (Sciaky, 1997)
* P. yunnanensis (Jedlička, 1934)
* P. yvanii (Dejean, 1828)
* P. zamotajlovi (Belousov, 1991)
* P. zhejiangensis (Kirschenhofer, 1997)
* P. ziegleri (Duftschmid, 1812)
* P. zoiai (Sciaky, 1996)
* P. zolotarewi (Reitter, 1911)
* P. zunianus (Casey, 1913)
* P. zyzzovi (O. Berlov & E. Berlov, 1996)
* ↑ Anichtchenko A. et al., (editors): Carabidae of the World. www.carabidae.org (bezocht: 20 tot 28 maart 2013)