# Tylopilus

Tylopilus is een geslacht van schimmels behorend tot de familie Boletaceae. Veel soorten uit dit geslacht zijn eetbaar maar hebben een bittere smaak. De typesoort is de Bittere boleet (Tylopilus felleus), die als een van de weinige soorten uit dit geslacht ook in Europa voorkomt. De meeste andere soorten komen voor in Noord-Amerika.

## Soorten

Volgens de Index Fungorum bevat het geslacht de volgende 146 soorten (peildatum januari 2023):
| Soortnaam | Auteur(s) | Publicatiejaar |
| Tylopilus acutesquamosus | Singer | 1983 |
| Tylopilus albirubens | (Corner) E. Horak | 2011 |
| Tylopilus alboater | (Schwein.) Murrill | 1909 |
| Tylopilus albofarinaceus | (W.F. Chiu) F.L. Tai | 1979 |
| Tylopilus albopurpureus | Yan C. Li & Zhu L. Yang | 2021 |
| Tylopilus alkalixanthus | Halling & Amtoft | 2003 |
| Tylopilus alpinus | Yan C. Li & Zhu L. Yang | 2016 |
| Tylopilus alutaceoumbrinus | Hongo | 1971 |
| Tylopilus ammiratii | Thiers | 1975 |
| Tylopilus appalachiensis | Singer | 1948 |
| Tylopilus aquarius | (Singer) Wartchow, Barb.-Silva, B. Ortiz & Ovrebo | 2017 |
| Tylopilus arenarius | Singer | 1978 |
| Tylopilus argentatae | (Corner) E. Horak | 2011 |
| Tylopilus argillaceus | Hongo | 1985 |
| Tylopilus ascendens | (Corner) E. Horak | 2011 |
| Tylopilus atratus | Both | 1998 |
| Tylopilus atripurpureus | (Corner) E. Horak | 2011 |
| Tylopilus atrobrunneus | (Lj.N. Vassiljeva) Wolfe | 1980 |
| Tylopilus atronicotianus | Both | 1998 |
| Tylopilus atroviolaceobrunneus | Yan C. Li & Zhu L. Yang | 2016 |
| Tylopilus aurantiacus | Yan C. Li & Zhu L. Yang | 2021 |
| Tylopilus aurantitubus | (Corner) E. Horak | 2011 |
| Tylopilus austrofelleus | (Cleland) Watling & N.M. Greg. | 1988 |
| Tylopilus badiceps | (Peck) A.H. Sm. & Thiers | 1971 |
| Tylopilus balloui | (Peck) Singer | 1947 |
| Tylopilus beelii | Heinem. & Gooss.-Font. | 1951 |
| Tylopilus brachypus | Singer, J. García & L.D. Gómez | 1991 |
| Tylopilus brunneirubens | (Corner) Watling & E. Turnbull | 1994 |
| Tylopilus bulbosus | Halling & G.M. Muell. | 2001 |
| Tylopilus callainus | N.K. Zeng, Zhi Q. Liang & M.S. Su | 2018 |
| Tylopilus castanoides | Har. Takah. | 2002 |
| Tylopilus cellulosus | Singer | 1947 |
| Tylopilus cervicolor | (Corner) E. Horak | 2011 |
| Tylopilus cervinicoccineus | (Corner) E. Horak | 2011 |
| Tylopilus chromoreticulatus | Wolfe & Bougher | 1993 |
| Tylopilus corneri | Singer, J. García & L.D. Gómez | 1991 |
| Tylopilus costaricensis | Singer & L.D. Gómez | 1991 |
| Tylopilus cremeus | (Corner) E. Horak | 2011 |
| Tylopilus cutifractus | (Corner) E. Horak | 2011 |
| Tylopilus cyanescens | T.H. Li & Watling | 1999 |
| Tylopilus cyanogranulifer | T.H. Li, Watling & N.M. Greg. | 1999 |
| Tylopilus dunensis | Magnago & M.A. Neves | 2017 |
| Tylopilus exiguus | T.W. Henkel | 1999 |
| Tylopilus felleus (Bittere boleet) | (Bull.) P. Karst. | 1881 |
| Tylopilus ferrugineus | (Kuntze) Singer | 1947 |
| Tylopilus fuligineoviolaceus | Har. Takah. | 2007 |
| Tylopilus funerarius | (Massee) Pegler & T.W.K. Young | 1981 |
| Tylopilus fuscatus | (Corner) Yan C. Li & Zhu L. Yang | 2021 |
| Tylopilus fuscescens | Watling | 1991 |
| Tylopilus glutinosus | Iqbal Hosen | 2021 |
| Tylopilus gomezii | Singer | 1991 |
| Tylopilus griseipurpureus | (Corner) E. Horak | 2011 |
| Tylopilus griseiviridus | Yan C. Li & Zhu L. Yang | 2021 |
| Tylopilus griseocarneus | Wolfe & Halling | 1989 |
| Tylopilus griseolivaceus | Angelini, Gelardi, Costanzo & Vizzini | 2019 |
| Tylopilus griseolus | Yan C. Li & Zhu L. Yang | 2021 |
| Tylopilus guanacastensis | Singer | 1983 |
| Tylopilus hayatae | Rodríguez-Ramírez & Luna-Vega | 2020 |
| Tylopilus himalayanus | D. Chakr., K. Das & Vizzini | 2018 |
| Tylopilus hondurensis | Singer & M.H. Ivory | 1983 |
| Tylopilus hongoi | Wolfe & Bougher | 1993 |
| Tylopilus humilis | Thiers | 1967 |
| Tylopilus intermedius | A.H. Sm. & Thiers | 1971 |
| Tylopilus isabellescens | Murrill | 1946 |
| Tylopilus jalapensis | Singer & J. García 1991 |
| Tylopilus javanicus | Henn. | 1898 |
| Tylopilus jiangxiensis | Kuan Zhao & Yan C. Li | 2020 |
| Tylopilus leucomycelinus | (Singer & M.H. Ivory) R. Flores & Simonini | 2000 |
| Tylopilus lividobrunneus | Singer | 1973 |
| Tylopilus louisii | Heinem. | 1964 |
| Tylopilus microsporus | S.Z. Fu, Q.B. Wang & Y.J. Yao | 2006 |
| Tylopilus minor | Singer | 1945 |
| Tylopilus mitissimus | Singer & L.D. Gómez | 1991 |
| Tylopilus montanus | Singer | 1989 |
| Tylopilus montoyae | Singer & J. García | 1991 |
| Tylopilus nebulosus | (Peck) Wolfe | 1980 |
| Tylopilus neofelleus | Hongo | 1967 |
| Tylopilus nicaraguensis | Singer & M.H. Ivory | 1983 |
| Tylopilus nigricans | Singer | 1947 |
| Tylopilus nigripes | A. Barbosa-Silva & Wartchow | 2019 |
| Tylopilus obscureviolaceus | Har. Takah. | 2004 |
| Tylopilus obscurus | Halling | 1989 |
| Tylopilus ochraceosquamosus | Watling | 1994 |
| Tylopilus olivaceirubens | (Corner) E. Horak | 2011 |
| Tylopilus olivaceobrunneus | Yan C. Li & Zhu L. Yang | 2021 |
| Tylopilus oradivensis | Osmundson & Halling | 2010 |
| Tylopilus orsonianus | Fulgenzi & T.W. Henkel | 2007 |
| Tylopilus otsuensis | Hongo | 1966 |
| Tylopilus pachycephalus | (Massee) Singer | 1983 |
| Tylopilus pakaraimensis | T.W. Henkel | 2001 |
| Tylopilus peralbidus | (Snell & Beardslee) Murrill | 1938 |
| Tylopilus pernanus | (Pat. & C.F. Baker) Watling | 2000 |
| Tylopilus perplexus | Watling | 1994 |
| Tylopilus phaeoruber | Yan C. Li & Zhu L. Yang | 2021 |
| Tylopilus phaseolisporus | (T.H. Li, R.N. Hilton & Watling) Osmundson, Bougher, R. Rob. & Halling | 2021 |
| Tylopilus piniphilus | Wolfe & Bougher | 1993 |
| Tylopilus pisciodorus | (Murrill) Murrill | 1948 |
| Tylopilus plumbeoviolaceoides | T.H. Li, B. Song & Y.H. Shen | 2002 |
| Tylopilus plumbeoviolaceus | (Snell & E.A. Dick) Snell & E.A. Dick | 1941 |
| Tylopilus potamogeton | Singer | 1978 |
| Tylopilus praeanisatus | (Murrill) Murrill | 1948 |
| Tylopilus primulinus | (Corner) E. Horak | 2011 |
| Tylopilus pseudoalpinus | Yan C. Li & Zhu L. Yang | 2021 |
| Tylopilus pseudoballoui | D. Chakr., K. Das & Vizzini | 2018 |
| Tylopilus pseudoscaber | Secr. ex A.H. Sm. & Thiers | 1968 |
| Tylopilus psittacinus | E. Horak | 2011 |
| Tylopilus punctatofumosus | (W.F. Chiu) F.L. Tai | 1979 |
| Tylopilus purpureorubens | Yan C. Li & Zhu L. Yang | 2021 |
| Tylopilus pygmaeus | Magnago & R.M.B. Silveira | 2017 |
| Tylopilus rhoadsiae | (Murrill) Murrill | 1944 |
| Tylopilus rhodoconius | (Singer) T.J. Baroni, Both & Bessette | 1998 |
| Tylopilus rigens | Hongo | 1979 |
| Tylopilus rubriporus | (Corner) E. Horak | 2011 |
| Tylopilus rubrobrunneus | Mazzer & A.H. Sm. | 1967 |
| Tylopilus rubrotinctus | Yan C. Li & Zhu L. Yang | 2021 |
| Tylopilus rufobrunneus | Yan C. Li & Zhu L. Yang | 2021 |
| Tylopilus rufonigricans | T.W. Henkel | 1999 |
| Tylopilus rugulosoreticulatus | Hongo | 1979 |
| Tylopilus sanctae-rosae | Singer | 1983 |
| Tylopilus snellii | Wolfe | 1983 |
| Tylopilus striatulus | Heinem. | 1951 |
| Tylopilus suavissimus | Heinem. & Gooss.-Font. | 1951 |
| Tylopilus subcellulosus | Singer & J. García | 1991 |
| Tylopilus subfusipes | A.H. Sm. | 1973 |
| Tylopilus subniger | Singer, J. García & L.D. Gómez | 1991 |
| Tylopilus subotsuensis | T.H.G. Pham, A.V. Alexandrova & O.V. Morozova | 2020 |
| Tylopilus subunicolor | E.A. Dick & Snell | 1961 |
| Tylopilus subvinaceipallidus | T.H. Li & Watling | 1999 |
| Tylopilus sultanii | S. Sarwar, Khalid & Niazi | 2014 |
| Tylopilus tabacinus | (Peck) Singer | 1944 |
| Tylopilus temucensis | Palfner | 2005 |
| Tylopilus tenuis | Heinem. | 1951 |
| Tylopilus tristiculus | (Massee) E. Horak | 2011 |
| Tylopilus tristior | (Corner) E. Horak | 2011 |
| Tylopilus variobrunneus | Roody, A.R. Bessette & Bessette | 1998 |
| Tylopilus veluticeps | (Pat. & C.F. Baker) Singer | 1947 |
| Tylopilus vinaceipallidus | (Corner) T.W. Henkel | 1999 |
| Tylopilus vinaceogriseus | Singer, J. García & L.D. Gómez | 1991 |
| Tylopilus vinosobrunneus | Hongo | 1979 |
| Tylopilus violaceobrunneus | Yan C. Li & Zhu L. Yang | 2016 |
| Tylopilus violaceorubrus | Yan C. Li & Zhu L. Yang | 2021 |
| Tylopilus violaceus | Heinem. | 1951 |
| Tylopilus violatinctus | T.J. Baroni & Both | 1998 |
| Tylopilus virescens | (Har. Takah. & Taneyama) N.K. Zeng, H. Chai & Zhi Q. Liang | 2019 |
| Tylopilus williamsii | Singer & J. García | 1991 |
| Tylopilus zambianus | Watling | 1994 |

## Externe link

* Tylopilus op MushroomExpert.com
* (en) Index Fungorum