# Chlaenius

Chlaenius is een geslacht van kevers uit de familie van de loopkevers (Carabidae). De wetenschappelijke naam van het geslacht is voor het eerst geldig gepubliceerd in 1810 door Franco Andrea Bonelli.
Chlaenius is een kosmopolitisch geslacht dat in alle zoögeografische provincies voorkomt. Het is ook een erg groot geslacht met meer dan 950 soorten en ondersoorten. Ongeveer driekwart daarvan komen voor in het Afrotropisch en Oriëntaals gebied.
De kevers hebben geheel of gedeeltelijk een glanzende metaalkleur.

## Soorten

* C. abacoides (Alluaud, 1933)
* C. aberanus (Sternberg, 1908)
* C. aberrans (Bates, 1882)
* C. abjectus (Andrewes, 1920)
* C. abstersus (Bates, 1873)
* C. accedens (Chaudoir, 1876)
* C. acroxanthus (Chaudoir, 1876)
* C. acutecostatus (Alluaud, 1933)
* C. adagidensis (Sternberg, 1908)
* C. adametzi (Kuntzen, 1913)
* C. adonis (Andrewes, 1923)
* C. aeneocephalus (Dejean, 1826)
* C. aeratus (Quensel, 1806)
* C. aestivus (Say, 1823)
* C. aethiopicus (Chaudoir, 1876)
* C. afganus (Jedlička, 1956)
* C. africanicus (Kirschenhofer, 2009)
* C. agilis (Chaudoir, 1856)
* C. agiloides (Jedlička, 1935)
* C. albissoni (Reitter, 1908)
* C. alesi (Jedlička, 1935)
* C. alexanderdostali (Kirschenhofer, 2008)
* C. alternans (Imhoff, 1843)
* C. alternatus (G. Horn, 1871)
* C. alutaceus (Gebler, 1829)
* C. allacteus (Alluaud, 1919)
* C. allardi (Basilewsky, 1967)
* C. allardianus (Lorenz, 1998)
* C. alluaudi (Fairmaire, 1901)
* C. alluaudianus (Jedlička, 1957)
* C. amabilis (Chaudoir, 1876)
* C. amarae (Andrewes, 1920)
* C. amauropterus (Chaudoir, 1856)
* C. amazonicus (Chaudoir, 1876)
* C. ammon (Fabricius, 1801)
* C. amoenus (Dejean, 1831)
* C. amplians (Bates, 1891)
* C. amplicollis (Basilewsky, 1953)
* C. amplipennis (Chaudoir, 1876)
* C. amydrus (Alluaud, 1934)
* C. analis (Olivier, 1795)
* C. analisimilis (Pomeroy, 1932)
* C. andamanensis (Andrewes, 1920)
* C. androyanus (Jeannel, 1949)
* C. angolanus (Basilewsky, 1953)
* C. angustatus (Dejean, 1831)
* C. anischenkoi (Kirschenhofer, 2010)
* C. annulipes (Bates, 1892)
* C. antennatus (Chaudoir, 1876)
* C. anthraceus (Alluaud, 1918)
* C. anthracoderus (Laferte-Senectere, 1851)
* C. apicalis (Wiedemann, 1819)
* C. apollo (Andrewes, 1919)
* C. argentinicus (Jedlička, 1946)
* C. armenus (Jedlička, 1950)
* C. arnoldi (Basilewsky, 1949)
* C. aruwimius (Bates, 1890)
* C. aspericollis (Bates, 1873)
* C. assamensis (Kirschenhofer, 2003)
* C. assecla (Laferte-Senectere, 1851)
* C. aterrimus (Basilewsky, 1956)
* C. athleta (Kryzhanovskij, 1976)
* C. atratus (Chaudoir, 1876)
* C. atripes (Chaudoir, 1876)
* C. atropos (Andrewes, 1941)
* C. attenuatus (Klug, 1833)
* C. augustus (Newman, 1838)
* C. aurifex (Basilewsky, 1949)
* C. auripilis (Andrewes, 1936)
* C. aurolimbatus (LaFerté-Sénectére, 1851)
* C. australis (Dejean, 1831)
* C. azurescens (Chaudoir, 1876)
* C. backanensis (Kirschenhofer, 2014)
* C. baehri (Kirschenhofer, 1998)
* C. baeticus (Rambur, 1837)
* C. balthasari (Jedlička, 1935)
* C. bandjermasinensis (Kirschenhofer, 2008)
* C. bangkokensis (Kirschenhofer, 2004)
* C. baoxingensis (Kirschenhofer, 2004)
* C. barkeri (Csiki, 1931)
* C. basilimbatus (Grundmann, 1956)
* C. baxii (Gory, 1833)
* C. beatus (Bates, 1891)
* C. bengalensis (Chaudoir, 1856)
* C. benyovszkyi (Csiki, 1931)
* C. berndjaegeri (Kirschenhofer, 2008)
* C. bicolor (Chaudoir, 1876)
* C. bifenestratus (Klug, 1832)
* C. bilyi (Kirschenhofer, 2009)
* C. bimaculatus (Dejean, 1826)
* C. binghami (Andrewes, 1919)
* C. bioculatus (Chaudoir, 1856)
* C. bipustulatus (Boheman, 1848)
* C. birmanicus (Chaudoir, 1876)
* C. bisignatus (Dejean, 1826)
* C. bivulnerus (Motschulsky, 1858)
* C. bocandei (Laferte-Senectere, 1852)
* C. bohemani (Chaudoir, 1856)
* C. boisduvalii (Dejean, 1831)
* C. bonelli (Mateu, 1947)
* C. borellyi (Burgeon, 1941)
* C. borgouensis (Kirschenhofer, 2007)
* C. bottegoi (Alluaud, 1933)
* C. boueti (Jeannel, 1949)
* C. boukali (Kirschenhofer, 2005)
* C. braminus (Chaudoir, 1876)
* C. brasiliensis (Dejean, 1831)
* C. breuningi (Jedlička, 1931)
* C. brevilabris (Le Conte, 1848)
* C. brevior (Fairmaire, 1901)
* C. breviusculus (Chaudoir, 1876)
* C. brunnescens (Kirschenhofer, 2009)
* C. bulirschi (Kirschenhofer, 2010)
* C. buquetii (Dejean & Boisduval, 1830)
* C. buriensis (Kirschenhofer, 1998)
* C. caecus (Dejean, 1831)
* C. caeruleiceps (Bates, 1892)
* C. caeruleicollis (Chaudoir, 1876)
* C. caesitius (Andrewes, 1923)
* C. caffer (Boheman, 1848)
* C. callichloris (Bates, 1873)
* C. cambodiensis (Bates, 1889)
* C. camerunensis (Kirschenhofer, 2003)
* C. canariensis (Dejean, 1831)
* C. carbonatus (Chaudoir, 1876)
* C. caurinus (G. Horn, 1885)
* C. cavilabrum (Barker, 1922)
* C. cebuensis (Kirschenhofer, 2008)
* C. cecrops (Kirschenhofer, 2009)
* C. celer (Chaudoir, 1876)
* C. ceramensis (Kirschenhofer, 2003)
* C. cinctus (Fabricius, 1781)
* C. circumcinctus (Say, 1830)
* C. circumdatus (Brulle, 1835)
* C. circumductus (A.Morawitz, 1862)
* C. circumscriptus (Duftschmid, 1812)
* C. clarksoni (Barker, 1922)
* C. clypeopatens (Basilewsky, 1949)
* C. coeruleus (Steven, 1809)
* C. colasi (Kirschenhofer, 2008)
* C. colmanti (Alluaud, 1933)
* C. colombensis (Jedlička, 1964)
* C. columbinus (Dejean, 1831)
* C. collarti (Alluaud, 1933)
* C. comans (Andrewes, 1919)
* C. comes (Péringuey, 1896)
* C. communimacula (Chaudoir in Oberthur, 1883)
* C. conformis (Dejean, 1831)
* C. congoanus (Alluaud, 1934)
* C. congoensis (Burgeon, 1935)
* C. conradsi (Kuntzen, 1913)
* C. consobrinus (Peringuey, 1896)
* C. consors (Péringuey, 1896)
* C. contractus (Chaudoir, 1876)
* C. controversus (Péringuey, 1926)
* C. convexus (Fairmaire, 1886)
* C. cookei (Andrewes, 1933)
* C. coquereli (Fairmaire, 1868)
* C. corbetti (Andrewes, 1919)
* C. cordicollis (Kirby, 1837)
* C. cordifer (Bates, 1891)
* C. corrosulus (Bates, 1892)
* C. coscinioderus (Chaudoir, 1856)
* C. cosciniophorus (Chaudoir, 1876)
* C. costiger (Chaudoir, 1856)
* C. costipennis (Boheman, 1848)
* C. costulatus (Motschulsky, 1859)
* C. coxalis (Fischer von Waldheim, 1844)
* C. crebrepunctatus (Chaudoir, 1856)
* C. crenistriatus (Chaudoir, 1876)
* C. cribellatus (Chaudoir, 1876)
* C. cribellicollis (Chaudoir, 1876)
* C. cribricollis (Dejean, 1831)
* C. croesus (Fabricius, 1801)
* C. croyi (Kirschenhofer, 2003)
* C. cruciatus (Dejean, 1831)
* C. crudelis (Péringuey, 1896)
* C. cruralis (Fischer von Waldheim, 1829)
* C. cubanus (Chaudoir, 1876)
* C. cumatilis (LeConte, 1851)
* C. cupreocinctus (Reiche, 1847)
* C. cupreolineatus (Chaudoir, 1876)
* C. cupreolus (Fairmaire, 1901)
* C. cupreopurpureus (Kirschenhofer, 2009)
* C. cupripennis (Chaudoir, 1876)
* C. cuprithorax (Quedenfeldt, 1883)
* C. cursor (Chevrolat, 1835)
* C. curtii (Lemaire, 2001)
* C. curtulus (Kirschenhofer, 2004)
* C. cyaneus (Brulle, 1835)
* C. cyanipennis (Boheman, 1848)
* C. cyanostolus (Andrewes, 1924)
* C. cylindricollis (Dejean, 1831)
* C. chagga (Alluaud, 1927)
* C. chainatensis (Kirschenhofer, 2011)
* C. chalcoderus (Chaudoir, 1876)
* C. chalcothorax (Wiedemann, 1823)
* C. cham (Chaudoir, 1876)
* C. championi (Andrewes, 1923)
* C. changwatensis (Kirschenhofer, 1998)
* C. chapanus (Andrewes, 1919)
* C. chaudoiri (G. Horn, 1876)
* C. chengduensis (Kirschenhofer, 2004)
* C. chiangmaiensis (Kirschenhofer, 2008)
* C. chitwanensis (Kirschenhofer, 2005)
* C. chlorochrous (Chaudoir, 1876)
* C. chlorodius (Dejean, 1826)
* C. chrysocephalus (P. Rossi, 1790)
* C. chrysoderus (Chaudoir, 1876)
* C. chuanqianensis (Liu & Liang, 2011)
* C. chuji (Jedlička, 1946)
* C. daer (Darlington, 1968)
* C. dajuensis (Kirschenhofer, 2002)
* C. dalibaiensis (Kirschenhofer, 2008)
* C. daressalaami (Jedlička, 1957)
* C. darfurensis (Kirschenhofer, 2007)
* C. darlingensis (Castelnau, 1867)
* C. darlingtoni (Basilewsky, 1951)
* C. davidsoni (Mandl, 1978)
* C. decellei (Basilewsky, 1968)
* C. decipiens (L. Dufour, 1820)
* C. decorsei (Alluaud, 1916)
* C. dejeanii (Dejean, 1831)
* C. delicatus (Bates, 1892)
* C. deliciolus (Bates, 1873)
* C. delkeskampi (Basilewsky, 1956)
* C. densaticollis (Fairmaire, 1901)
* C. denticulatus (Dejean, 1831)
* C. deplanatus (Laferte-Senectere, 1851)
* C. deserticola (Raffray, 1885)
* C. deuvei (Kirschenhofer, 2008)
* C. deyrollei (Laferte-Senectere, 1851)
* C. dhawalagiriensis (Kirschenhofer, 2002)
* C. diabolicus (Basilewsky, 1961)
* C. dianus (Jedlička, 1957)
* C. diephelus (Alluaud, 1934)
* C. differens (Peyron, 1858)
* C. difficilis (Sternberg, 1908)
* C. dilatatus (Motschulsky, 1864)
* C. dilutipes (Reitter, 1894)
* C. dimidiatus (Chaudoir, 1842)
* C. dinodoides (Chaudoir, 1876)
* C. discicollis (Laferte-Senectere, 1851)
* C. discopictus (Fairmaire, 1893)
* C. dispreticus (Basilewsky, 1968)
* C. dissidens (Péringuey, 1926)
* C. distigma (Chaudoir, 1876)
* C. ditulus (Péringuey, 1904)
* C. diversus (Facchini, 2011)
* C. dives (Dejean, 1826)
* C. djaina (Maindron, 1899)
* C. dohrnii (Bertoloni, 1857)
* C. dondoensis (Kirschenhofer, 2007)
* C. doriae (Chaudoir, 1876)
* C. dorsalis (Dejean, 1831)
* C. dostojevskji (Tschitscherine, 1895)
* C. drescheri (Louwerens, 1951)
* C. ducalis (Chaudoir, 1876)
* C. dureli (Maindron, 1899)
* C. dusaultii (L.Dufour, 1821)
* C. duvaucelii (Dejean, 1831)
* C. ecuadoricus (Jedlička, 1946)
* C. effugiens (Péringuey, 1908)
* C. elegans (Schmidt-Goebel, 1846)
* C. elgonensis (Alluaud, 1939)
* C. elisabethanus (Burgeon, 1935)
* C. elongatulus (Kirschenhofer, 2009)
* C. elongatus (Klug, 1833)
* C. emarginatus (Say, 1823)
* C. eneides (Bates, 1892)
* C. epicosmoides (Basilewsky, 1956)
* C. episcopalis (Dejean, 1831)
* C. epistrophus (Alluaud, 1927)
* C. eritreaensis (Kirschenhofer, 2007)
* C. ernesti (Gory, 1833)
* C. erythrocnemis (Chaudoir, 1876)
* C. erythropus (Germar, 1824)
* C. eugrammus (Basilewsky, 1947)
* C. eurybates (Bates, 1891)
* C. euryscopus (Bates, 1886)
* C. exaratus (Basilewsky, 1949)
* C. exilis (Andrewes, 1923)
* C. extensus (Mannerheim, 1825)
* C. eyeni (Basilewsky, 1949)
* C. fairmairei (Murray, 1858)
* C. fallax (Olivier, 1795)
* C. farai (Jedlička, 1949)
* C. farkaci (Kirschenhofer, 2005)
* C. fasciger (Chaudoir In Oberthur, 1883)
* C. fastigatus (Andrewes, 1921)
* C. feanus (Bates, 1892)
* C. femoratus (Dejean, 1826)
* C. fenestratus (Chaudoir, 1876)
* C. feronioides (Murray, 1858)
* C. festivus (Panzer, 1796)
* C. fimbriatus (Klug, 1833)
* C. finitimus (Péringuey, 1908)
* C. fizianus (Basilewsky, 1961)
* C. flaccidus (G. Horn, 1876)
* C. flavicornis (Fischer von Waldheim, 1842)
* C. flaviguttatus (W.S. Macleay, 1825)
* C. flavipes (Menetries, 1832)
* C. flavofemoratus (Laporte, 1834)
* C. flavomaculatus (Kolbe, 1889)
* C. flavoscriptus (Quedenfeldt, 1891)
* C. fletcheri (Andrewes, 1919)
* C. floridanus (G. Horn, 1876)
* C. formosensis (Lorenz, 1998)
* C. forreri (Bates, 1884)
* C. frater (Chaudoir, 1876)
* C. fraterculus (Maindron, 1899)
* C. fraternus (Kolbe, 1889)
* C. freyellus (Jedlička, 1959)
* C. freyi (Jedlička, 1960)
* C. freynei (Burgeon, 1937)
* C. fugax (Chaudoir, 1876)
* C. fulgidicollis (L. Dufour, 1820)
* C. fulvicollis (Chaudoir, 1876)
* C. fulvipes (Chaudoir, 1835)
* C. fuscicornis (Dejean, 1831)
* C. gabonensis (Kirschenhofer, 2009)
* C. gabonicus (Gemminger & Harold, 1868)
* C. gansuensis (Jedicka, 1935)
* C. geayi (Jeannel, 1949)
* C. geisthardti (Kirschenhofer, 2007)
* C. gemmingeri (Ballion, 1869)
* C. geniculatus (Basilewsky, 1956)
* C. germanus (Chaudoir, 1876)
* C. gestroi (Chaudoir, 1876)
* C. ghesquierei (Burgeon, 1935)
* C. gikondo (Kirschenhofer, 2009)
* C. glabratus (Dejean, 1831)
* C. glabricollis (Dejean, 1831)
* C. glaucinus (Kirschenhofer, 2007)
* C. glaucus (LeConte, 1856)
* C. gonioderus (Laferte-Senectere, 1851)
* C. gonospilus (Walker, 1871)
* C. goossensi (Alluaud, 1933)
* C. goryi (Gory, 1833)
* C. gotschii (Chaudoir, 1846)
* C. greensladei (Darlington, 1971)
* C. greyanus (White, 1841)
* C. grundmanni (Basilewsky, 1956)
* C. guatemalenus (Bates, 1882)
* C. guineensis (Kolbe, 1883)
* C. gundlachi (Chaudoir, 1876)
* C. guttula (Chaudoir, 1856)
* C. hainanensis (Kirschenhofer, 2004)
* C. hamatus (Eschscholtz, 1829)
* C. hamifer (Chaudoir, 1856)
* C. harpalinus (Eschscholtz, 1833)
* C. heidenfelderi (Kirschenhofer, 2008)
* C. helvetorum (Burgeon, 1935)
* C. hemichlorus (Fairmaire, 1888)
* C. henryi (Andrewes, 1919)
* C. herbaceus (Chevrolat, 1834)
* C. heyrovskyi (Jedlička, 1949)
* C. hildebrandti (Harold, 1880)
* C. himalayicus (Andrewes, 1923)
* C. holomelas (Alluaud, 1915)
* C. hornburgi (Kirschenhofer, 2008)
* C. horni (Sternberg, 1908)
* C. hostilis (Putzeys, 1880)
* C. howa (Kunckel dHerculais, 1891)
* C. huedepohli (Mandl, 1983)
* C. humeralis (Chaudoir, 1856)
* C. humphreyi (Alluaud, 1927)
* C. hunanensis (Kirschenhofer, 2003)
* C. hypocrita (Péringuey, 1896)
* C. imitatus (Reitter, 1895)
* C. immaculatus (Péringuey, 1885)
* C. immunitus (Murray, 1858)
* C. imperialis (Sternberg, 1908)
* C. impictus (Alluaud, 1933)
* C. impressicollis (Chaudoir, 1876)
* C. impunctifrons (Say, 1823)
* C. inaequalis (Fairmaire, 1901)
* C. inderiensis (Motschulsky, 1850)
* C. indicus (Jedlička, 1956)
* C. indutus (Klug, 1833)
* C. infantulus (Chaudoir, 1876)
* C. infersus (Péringuey, 1926)
* C. inops (Chaudoir, 1856)
* C. insignis (Chaudoir, 1876)
* C. instabilis (Raffray, 1886)
* C. insulanus (Louwerens, 1956)
* C. insularis (Uéno, 1964)
* C. intermedius (Chaudoir, 1856)
* C. interruptus (G. Horn, 1876)
* C. irakensis (Jedlička, 1959)
* C. iranensis (Kirschenhofer, 1998)
* C. itombwanus (Basilewsky, 1961)
* C. ivorensis (Facchini, 2011)
* C. jacobsoni (Andrewes, 1926)
* C. jactus (Kirschenhofer, 2004)
* C. jaechi (Kirschenhofer, 1991)
* C. jakeschi (Jedlička, 1967)
* C. jaleswarensis (Kirschenhofer, 2004)
* C. jamaicae (Darlington, 1935)
* C. janthinus (L. Redtenbacher, 1844)
* C. jeanneli (Basilewsky, 1949)
* C. jordani (Basilewsky, 1955)
* C. jucundulus (Basilewsky, 1970)
* C. judianensis (Kirschenhofer, 2004)
* C. junceus (Andrewes, 1923)
* C. jureceki (Jedlička, 1935)
* C. juvencus (Dejean, 1831)
* C. kafakumbae (Burgeon, 1935)
* C. kanarae (Andrewes, 1919)
* C. kapangae (Burgeon, 1935)
* C. kashmiricus (Grundmann, 1955)
* C. kaszabi (Jedlička, 1951)
* C. katanganus (Burgeon, 1935)
* C. keniaensis (Kirschenhofer, 2008)
* C. kenyerii (Kirschenhofer, 2003)
* C. kerkvoordeae (Burgeon, 1935)
* C. kilimanus (Basilewsky, 1956)
* C. kindermanni (Chaudoir, 1856)
* C. kira (Alluaud, 1919)
* C. kirki (Chaudoir, 1876)
* C. kivuanus (Basilewsky, 1949)
* C. kivuensis (Alluaud, 1933)
* C. koenigi (Semenov, 1888)
* C. kolariensis (Maindron, 1898)
* C. kolbei (Duvivier, 1892)
* C. koltzei (Grundmann, 1956)
* C. kotys (Kirschenhofer, 2004)
* C. kraatzi (Sternberg, 1908)
* C. kryzhanovskyi (Basilewsky, 1968)
* C. kulti (Basilewsky, 1949)
* C. kurosawai (Kasahara, 1986)
* C. labroexcisus (Basilewsky, 1948)
* C. lacunosus (Andrewes, 1920)
* C. ladon (Kirschenhofer, 1998)
* C. laetiusculus (Chaudoir, 1856)
* C. laetoides (Burgeon, 1935)
* C. laetus (Fabricius, 1794)
* C. laevicollis (Péringuey, 1926)
* C. laevipennis (Chaudoir, 1876)
* C. laeviplaga (Chaudoir, 1876)
* C. laeviusculus (Chaudoir, 1856)
* C. lafertei (Guérin-Méneville, 1843)
* C. lamottei (Basilewsky, 1951)
* C. langmusiensis (Kirschenhofer, 2008)
* C. langsonensis (Kirschenhofer, 2008)
* C. laosensis (Kirschenhofer, 2008)
* C. laotinus (Andrewes, 1919)
* C. lapillus (Basilewsky, 1949)
* C. lastii (Bates, 1886)
* C. lateralis (Brulle, 1838)
* C. latesternalis (Basilewsky, 1956)
* C. laticollis (Say, 1823)
* C. latipalpis (Mandl, 1992)
* C. latipennis (Sternberg, 1908)
* C. latithorax (Mannerheim, 1844)
* C. lativittis (Chaudoir, 1876)
* C. latreillei (Laferte-Senectere, 1852)
* C. lederi (Reitter, 1888)
* C. leigongshanensis (Kirschenhofer, 2004)
* C. leishanensis (Kirschenhofer, 2005)
* C. leleupi (Basilewsky, 1952)
* C. leopoldi (Burgeon, 1935)
* C. leprieuri (Gory, 1833)
* C. leptopus (Basilewsky, 1956)
* C. leucops (Wiedemann, 1823)
* C. leucoristus (Chaudoir, 1876)
* C. leucoscelis (Chevrolat, 1835)
* C. leytensis (Kirschenhofer, 2008)
* C. limbatus (Wiedemann, 1821)
* C. limbicollis (Chaudoir, 1876)
* C. limbipennis (Boheman, 1860)
* C. lineatus (Putzeys, 1880)
* C. lineellus (Motschulsky, 1859)
* C. lineicinctus (Fairmaire, 1901)
* C. linwensini (Liu & Liang, 2011)
* C. lioderus (Andrewes, 1923)
* C. liothorax (Alluaud, 1934)
* C. lirifer (Andrewes, 1941)
* C. lissoderus (Chaudoir, 1876)
* C. lithophilus (Say, 1823)
* C. litongaensis (Kirschenhofer, 2009)
* C. loango (Kirschenhofer, 2009)
* C. lobozianus (Alluaud, 1934)
* C. loeblianus (Kirschenhofer, 2004)
* C. lombokensis (Kirschenhofer, 2008)
* C. lomii (G.Muller, 1941)
* C. lomsakensis (Kirschenhofer, 1998)
* C. longeantennatus (Basilewsky, 1951)
* C. longicornis (Chaudoir, 1843)
* C. longulus (Jeannel, 1949)
* C. louwerensi (Andrewes, 1936)
* C. loveridgei (Basilewsky, 1951)
* C. loxias (Kirschenhofer, 2004)
* C. lucasi (Peyron, 1858)
* C. lucidicollis (Laferte-Senectere, 1851)
* C. lucidulus (Boheman, 1848)
* C. luculentus (Andrewes, 1920)
* C. lugens (Chaudoir, 1876)
* C. luisae (Gestro, 1895)
* C. lujai (Burgeon, 1935)
* C. lulengae (Burgeon, 1935)
* C. luluanus (Basilewsky, 1949)
* C. lunatus (Dejean, 1826)
* C. luteicauda (Chaudoir, 1876)
* C. luteoapicalis (Facchini, 2011)
* C. luzonicus (Chaudoir, 1856)
* C. lyperus (Jeannel, 1949)
* C. macropus (Chaudoir, 1876)
* C. maculatus (Dejean, 1826)
* C. maculiceps (Boheman, 1848)
* C. maculiger (Castelnau, 1867)
* C. madecassus (Csiki, 1931)
* C. madrasensis (Kirschenhofer, 2004)
* C. makalolo (Bates, 1886)
* C. malachinus (Motschulsky, 1865)
* C. malcheri (Emden, 1937)
* C. mandli (Grundmann, 1956)
* C. manowianus (Jedlička, 1957)
* C. manus (Louwerens, 1969)
* C. maowensis (Kirschenhofer, 2008)
* C. maputoensis (Kirschenhofer, 2007)
* C. marginellus (Dejean, 1831)
* C. marginicollis (Boheman, 1848)
* C. marginipennis (Gory, 1833)
* C. marianensis (Darlington, 1970)
* C. marleyi (Barker, 1922)
* C. martinbaehri (Kirschenhofer, 2008)
* C. masoni (Andrewes, 1923)
* C. maxi (Gory, 1833)
* C. maxillosus (G. Horn, 1876)
* C. medioguttatus (Chaudoir, 1876)
* C. mediornatus (Basilewsky, 1956)
* C. melampus (Menetries, 1849)
* C. melancholicus (Laferte-Senectere, 1851)
* C. melanocnemis (Basilewsky, 1949)
* C. melanopterus (Chaudoir, 1876)
* C. melanopus (Andrewes, 1923)
* C. menevillei (Chaudoir, 1876)
* C. merkli (Kirschenhofer, 2003)
* C. meteorus (Alluaud, 1939)
* C. meticulosus (Laferte-Senectere, 1851)
* C. micans (Fabricius, 1792)
* C. microspilus (Bates, 1892)
* C. milleti (Antoine, 1932)
* C. milloti (Jeannel, 1949)
* C. moaboensis (Kirschenhofer, 2009)
* C. moestus (Csiki, 1931)
* C. moheliensis (Kirschenhofer, 2009)
* C. moluccensis (Kirschenhofer, 2003)
* C. momboensis (Kirschenhofer, 2007)
* C. monardi (Alluaud, 1934)
* C. montanus (Lucas, 1859)
* C. montivagus (Andrewes, 1923)
* C. morettoi (Kirschenhofer, 2008)
* C. morio (Boheman, 1860)
* C. morosus (Laferte-Senectere, 1851)
* C. mpanga (Kirschenhofer, 2009)
* C. muehlei (Kirschenhofer, 2004)
* C. multicolor (Andrewes, 1919)
* C. mutatus (Gemminger & Harold, 1868)
* C. naeviger (A. Morawitz, 1862)
* C. nakuruensis (Kirschenhofer, 2012)
* C. nanlingensis (Deuve & Tian, 2005)
* C. nanpingensis (Kirschenhofer, 1999)
* C. natalensis (Chaudoir, 1876)
* C. nebraskensis (LeConte, 1856)
* C. nebrioides (Antoine, 1961)
* C. nemoralis (Say, 1823)
* C. neocaledonicus (Chaudoir In Oberthur, 1883)
* C. neochloodes (Lorenz, 1998)
* C. nepalensis (Hope, 1831)
* C. nepos (Chaudoir, 1876)
* C. newdehliensis (Kirschenhofer, 2002)
* C. niger (Randall, 1838)
* C. nigerrimus (Jedlička, 1958)
* C. nigrans (Kirschenhofer, 2003)
* C. nigratus (Kirschenhofer, 2007)
* C. nigricans (Wiedemann, 1821)
* C. nigricornis (Fabricius, 1787)
       Zwartsprietfluweelloper
* C. nigripennis (Chaudoir, 1856)
* C. nigrita (Dejean, 1826)
* C. nigroscelis (Chaudoir, 1856)
* C. nigrosuturatus (Mandl, 1978)
* C. nilgiricus (Andrewes, 1919)
* C. nimbanus (Basilewsky, 1951)
* C. nitidiceps (Dejean, 1826)
* C. nitidicollis (Dejean, 1826)
* C. nitidifrons (Fairmaire, 1901)
* C. nitidulus (Schrank, 1781)
       Groene fluweelloper
* C. noeli (Alluaud, 1933)
* C. noguchii (Bates, 1873)
* C. nossibianus (Facchini, 2011)
* C. notabilis (Laferte-Senectere, 1851)
* C. notula (Fabricius, 1801)
* C. nubicus (Chaudoir, 1876)
* C. nyassaensis (Jedlička, 1957)
* C. nyika (Kirschenhofer, 2009)
* C. obenbergeri (Jedlička, 1935)
* C. oberthuri (Sternberg, 1908)
* C. obesus (Laferte-Senectere, 1851)
* C. obliquatus (Barker, 1922)
* C. oblongus (Dejean, 1826)
* C. obscuripennis (Chevrolat, 1835)
* C. obscurus (Klug, 1832)
* C. obsidianoides (Alluaud, 1935)
* C. obsidianus (Alluaud, 1935)
* C. obsoletus (LeConte, 1851)
* C. obtusus (Dejean, 1831)
* C. occultus (Sloane, 1907)
* C. ocreatus (Bates, 1873)
* C. oculatus (Fabricius, 1801)
* C. ochroperas (Bates, 1892)
* C. oligochrysus (Alluaud, 1934)
* C. olivieri (Crotch, 1871)
* C. olthofi (Darlington, 1968)
* C. omochlorus (Andrewes, 1931)
* C. oneili (Barker, 1922)
* C. oodioides (Chaudoir, 1876)
* C. opacipennis (Chaudoir, 1876)
* C. ophonoides (Fairmaire, 1843)
* C. opisthographus (Alluaud, 1934)
* C. orbicollis (Chaudoir, 1876)
* C. orbiculicollis (Barker, 1922)
* C. orbus (G. Horn, 1871)
* C. orphanus (Péringuey, 1908)
* C. ostrinus (Andrewes, 1924)
* C. ovalipennis (Quedenfeldt, 1883)
* C. ovampo (Péringuey, 1892)
* C. overlaeti (Burgeon, 1935)
* C. oxygonus (Chaudoir, 1843)
* C. pacholatkoi (Kirschenhofer, 2003)
* C. pachys (Chaudoir, 1876)
* C. paenulatus (Erichson, 1843)
* C. palpalis (LaFerte-Senectere, 1851)
* C. paluensis (Kirschenhofer, 2011)
* C. pallidicornis (Ballion, 1871)
* C. pallipes (Gebler, 1823)
* C. pan (Darlington, 1968)
* C. panagaeoides (Chaudoir, 1876)
* C. panjabensis (Kirschenhofer, 1998)
* C. parallelus (Dejean, 1831)
* C. parepus (Jedlička, 1957)
* C. paromius (Basilewsky, 1949)
* C. patricius (Harold, 1879)
* C. patrizii (Basilewsky, 1953)
* C. patruelis (LeConte, 1844)
* C. pauxillus (Kirschenhofer, 2002)
* C. paykulli (Crotch, 1870)
* C. pectinipes (Bates, 1892)
* C. peltastes (Jedlička, 1935)
* C. pennsylvanicus (Say, 1823)
* C. pericallus (L.Redtenbacher, 1867)
* C. perinetanus (Facchini, 2011)
* C. peringueyi (Kuntzen, 1919)
* C. perpunctatus (Kuntzen, 1913)
* C. perrieri (Fairmaire, 1899)
* C. persimilis (Chaudoir, 1876)
* C. perspicillaris (Erichson, 1843)
* C. pertinax (Casey, 1920)
* C. peruanus (Erichson, 1847)
* C. peterseni (Louwerens, 1967)
* C. phaenoderus (Chaudoir, 1876)
* C. phakhaoensis (Kirschenhofer, 2012)
* C. pharaonis (Motschulsky, 1865)
* C. phenax (Basilewsky, 1949)
* C. philemon (Andrewes, 1936)
* C. piceus (Chaudoir, 1876)
* C. pictus (Chaudoir, 1856)
* C. piligenys (Liu & Liang, 2010)
* C. pilosicoerulans (Mandl, 1989)
* C. pimalicus (Casey, 1914)
* C. planipennis (Chaudoir, 1876)
* C. planulatus (Bates, 1884)
* C. platensis (G.R.Waterhouse, 1841)
* C. platyderus (Chaudoir, 1856)
* C. platynoides (Alluaud, 1934)
* C. plausibilis (Basilewsky, 1949)
* C. pleuroderus (Chaudoir In Oberthur, 1883)
* C. poecilinus (Bates, 1892)
* C. porinus (Alluaud, 1929)
* C. porphyrius (Bates, 1891)
* C. porrectus (Chaudoir, 1876)
* C. posticalis (Motschulsky, 1854)
* C. posticus (Fabricius, 1798)
* C. postmaculatus (Basilewsky, 1952)
* C. postscriptus (Bates, 1873)
* C. pradieri (Chaudoir, 1876)
* C. praefectus (Bates, 1873)
* C. prasinus (Dejean, 1826)
* C. pratensis (Chaudoir, 1876)
* C. pretiosus (Chaudoir, 1856)
* C. principalis (Sternberg, 1908)
* C. privatus (Bates, 1892)
* C. probsti (Kirschenhofer, 2004)
* C. prolixus (Erichson, 1843)
* C. propeagilis (Liu & Kavanaugh, 2011)
* C. propinquus (Csiki, 1931)
* C. prostenus (Bates, 1873)
* C. protensus (Chaudoir, 1876)
* C. pseudobipustulatus (Facchini, 2011)
* C. pseudocruciatus (Kirschenhofer, 2009)
* C. pseudocupreolus (Kirschenhofer, 2010)
* C. pseudoglabratus (Kirschenhofer, 2009)
* C. pseudolulengae (Kirschenhofer, 2009)
* C. pseudopalpalis (Basilewsky, 1949)
* C. pseudoraffrayi (Basilewsky, 1949)
* C. pterostichoides (Andrewes, 1941)
* C. ptuchodes (Andrewes, 1923)
* C. puberulus (Boheman, 1848)
* C. pubifer (Chaudoir, 1876)
* C. pubipennis (Chaudoir, 1856)
* C. puchneri (Kirschenhofer, 2007)
* C. pudicus (Fabricius, 1801)
* C. pugni (Camerano, 1879)
* C. pulchellus (Boheman, 1848)
* C. pulcher (Nietner, 1857)
* C. pulchriceps (Fairmaire, 1901)
* C. pumilio (Kolbe, 1889)
* C. punctatostriatus (Chaudoir, 1856)
* C. puncticephalis (Saha, 1984)
* C. puncticollis (Dejean, 1826)
* C. purpureus (Chaudoir, 1876)
* C. purpuricollis (Randall, 1838)
* C. pusillus (Say, 1823)
* C. putzeysi (Chaudoir, 1876)
* C. pyrrhopodus (Fairmaire, 1903)
* C. pyrrhos (Kirschenhofer, 2004)
* C. quadricolor (Olivier, 1790)
* C. quadrimaculatus (Boheman, 1848)
* C. quadrinotatus (Dejean, 1826)
* C. quadriornatus (Basilewsky, 1956)
* C. quadripustulatus (Dejean, 1831)
* C. quadrisignatus (Boheman, 1860)
* C. quadrisulcatus (Paykull, 1790)
* C. quadrivittatus (Jedlička, 1958)
* C. radama (Kunckel, 1891)
* C. raffrayi (Chaudoir, 1876)
* C. rafiki (Alluaud, 1929)
* C. rambouseki (Lutshnik, 1933)
* C. ranavalonae (Csiki, 1931)
* C. rebellis (Péringuey, 1926)
* C. reichei (Laferte-Senectere, 1851)
* C. reitteri (Jakobson, 1906)
* C. retropictus (Fairmaire, 1901)
* C. rhodesianus (Péringuey, 1899)
* C. richardsi (Ali, 1967)
* C. riparius (Lorenz, 1998)
* C. ripicola (Andrewes, 1937)
* C. robustus (Boheman, 1848)
* C. rodriguezi (Chaudoir, 1876)
* C. roeschkei (Sternberg, 1908)
* C. rotundatulus (Lorenz, 1998)
* C. rotundithorax (Liu & Kavanaugh, 2010)
* C. rotundus (Andrewes, 1920)
* C. rubenticollis (Kirschenhofer, 2009)
* C. rubricrus (Alluaud, 1916)
* C. rudesculptus (Chaudoir, 1876)
* C. rudicollis (Chaudoir, 1876)
* C. ruficauda (Chaudoir, 1856)
* C. rufifemoratus (MacLeay, 1825)
* C. rufithorax (Wiedemann, 1821)
* C. rufomarginatus (Dejean, 1831)
* C. rugulosus (Nietner, 1857)
* C. rukwaensis (Kirschenhofer, 2007)
* C. ruvumaensis (Kirschenhofer, 2008)
* C. rysonotus (Fairmaire, 1901)
* C. sabahensis (Kirschenhofer, 1998)
* C. sagaingensis (Kirschenhofer, 2005)
* C. saginatus (Laferte-Senectere, 1851)
* C. sagittarius (Dejean, 1831)
* C. salisburiensis (Barker, 1922)
* C. sallei (Chaudoir, 1876)
* C. samoensis (Csiki, 1915)
* C. sangaicus (Alluaud, 1929)
* C. sankuruensis (Burgeon, 1935)
* C. sassanus (Alluaud, 1934)
* C. savanicola (Basilewsky, 1968)
* C. scabricollis (Chevrolat, 1833)
* C. scapularis (Chaudoir, 1876)
* C. sciakyi (Kirschenhofer, 2004)
* C. scotti (Alluaud, 1937)
* C. sculptilis (Bates, 1886)
* C. scutellaris (Harold, 1880)
* C. scheerpeltzi (Basilewsky, 1956)
* C. schillhammeri (Kirschenhofer, 2003)
* C. schmidti (Gestro, 1895)
* C. schoenherri (Dejean, 1831)
* C. schoutedeni (Burgeon, 1935)
* C. schuelei (Kirschenhofer, 2007)
* C. sehnali (Kirschenhofer, 2008)
* C. seiferti (Kirschenhofer, 2005)
* C. sellatus (Dejean, 1831)
* C. semenowi (Tschitscherine, 1895)
* C. semicupreus (Basilewsky, 1949)
* C. semicyaneus (Solsky, 1874)
* C. semipurpureus (Motschulsky, 1865)
* C. semiviridis (Andrewes, 1920)
* C. senegalensis (Dejean, 1831)
* C. sennaariensis (Chaudoir, 1856)
* C. sericeus (Forster, 1771)
* C. sericimicans (Chaudoir, 1876)
* C. sexguttatus (Andrewes, 1919)
* C. sexmaculatus (Dejean, 1831)
* C. seyrigi (Alluaud, 1935)
* C. shaanxinensis (Kirschenhofer, 2004)
* C. siccus (Darlington, 1968)
* C. sichuanensis (Kirschenhofer, 2008)
* C. sierraleonensis (Kirschenhofer, 2008)
* C. siffointei (Basilewsky, 1968)
* C. signatus (Boheman, 1848)
* C. silvestrii (G.Muller, 1942)
* C. simba (Alluaud, 1929)
* C. simbabwensis (Kirschenhofer, 2008)
* C. similatus (Boheman, 1848)
* C. similis (Chaudoir, 1856)
* C. simillimus (Chaudoir, 1856)
* C. simplex (Wiedemann, 1821)
* C. simulatorius (Barker, 1922)
* C. sinensis (Chaudoir, 1856)
* C. sinuatus (Dejean, 1826)
* C. sivorii (Chaudoir, 1876)
* C. sobrinus (Dejean, 1826)
* C. soccatus (Say, 1830)
* C. soginoides (Chaudoir, 1876)
* C. sokotranus (Csiki, 1931)
* C. solitarius (Say, 1823)
* C. sollicitus (Laferte-Senectere, 1851)
* C. somaliae (Basilewsky, 1956)
* C. somereni (Alluaud, 1929)
* C. soricinus (Gerstaecker, 1867)
* C. sparsepunctatus (Chaudoir, 1876)
* C. spathulifer (Bates, 1873)
* C. specularis (Emden, 1937)
* C. splendidus (Dejean, 1831)
* C. spoliatus (P. Rossi, 1792)
* C. startellus (Basilewsky, 1949)
* C. stenoristus (Chaudoir, 1876)
* C. sterbai (Jedlička, 1935)
* C. steveni (Quensel, 1806)
* C. stichai (Jedlička, 1949)
* C. straneoi (Basilewsky, 1949)
* C. stschukini (Menetries, 1837)
* C. stungtrengensis (Kirschenhofer, 2004)
* C. stygius (Laferte-Senectere, 1851)
* C. suavis (Alluaud, 1939)
* C. subcostatus (W.J.Macleay, 1864)
* C. suberbiei (Jeannel, 1949)
* C. subferrugineus (Kirschenhofer, 2003)
* C. submarginatus (Chaudoir, 1876)
* C. subovatus (Chaudoir, 1876)
* C. subsulcatus (Dejean, 1831)
* C. suensoni (Mandl, 1992)
* C. sulcicollis (Paykull, 1798)
       Groefhalsfluweelloper
* C. sulcipennis (Dejean, 1826)
* C. sumptuosus (Alluaud, 1934)
* C. superbus (Sternberg, 1908)
* C. superstes (Péringuey, 1926)
* C. suppletor (Bates, 1891)
* C. surdipennis (Jeannel, 1949)
* C. suvorovi (Semenov, 1912)
* C. swahilius (Bates, 1886)
* C. syangyaensis (Kirschenhofer, 2004)
* C. sykesi (Hope, 1835)
* C. synaptus (Alluaud, 1918)
* C. syriacus (Chaudoir, 1876)
* C. systolocranioides (Alluaud, 1933)
* C. tamdaoensis (Kirschenhofer, 2003)
* C. tanahrataensis (Kirschenhofer, 2008)
* C. teani (Gestro, 1881)
* C. tecospilus (Alluaud, 1933)
* C. tenellus (Klug, 1832)
* C. tenuicollis (Fabricius, 1801)
* C. tenuilimbatus (Ballion, 1871)
* C. tenuis (Fairmaire, 1901)
* C. terminatus (Dejean, 1826)
* C. testaceicrus (Fairmaire, 1891)
* C. tetracelis (Alluaud, 1933)
* C. tetragonoderus (Chaudoir, 1876)
* C. tetraphacus (Alluaud, 1918)
* C. tetraspilus (Alluaud, 1918)
* C. texanus (G. Horn, 1876)
* C. thieleni (Kirschenhofer, 1998)
* C. thiesensis (Kirschenhofer, 2007)
* C. tibialis (Dejean, 1826)
       Rivierfluweelloper
* C. timorensis (Darlington, 1971)
* C. tinantae (Burgeon, 1935)
* C. titschacki (Jedlička, 1946)
* C. togatus (Klug, 1829)
* C. tomentosus (Say, 1823)
* C. topali (Kirschenhofer, 2004)
* C. toubaensis (Kirschenhofer, 2008)
* C. touzalini (Andrewes, 1920)
* C. trachys (Andrewes, 1923)
* C. transfuga (Chaudoir, 1876)
* C. transversalis (Dejean, 1831)
* C. trapezicollis (Chaudoir, 1856)
* C. treichi (Alluaud, 1916)
* C. tricolor (Dejean, 1826)
* C. trichrous (Alluaud, 1918)
* C. trigonotomoides (Emden, 1928)
* C. trinotatus (Laferte-Sinectere, 1851)
* C. tripustulatus (Dejean, 1831)
* C. tristis (Schaller, 1783)
       Zwarte fluweelloper
* C. tschitscherini (Jedlička, 1952)
* C. tshibindensis (Burgeon, 1935)
* C. tudicus (Andrewes, 1919)
* C. tuky (Basilewsky, 1949)
* C. uluguruanus (Basilewsky, 1956)
* C. umtalianus (Péringuey, 1904)
* C. unicolor (Chaudoir, 1856)
* C. uninotatus (Andrewes, 1919)
* C. unyorensis (Alluaud, 1934)
* C. uzungwensis (Basilewsky, 1951)
* C. vadoni (Basilewsky, 1950)
* C. vafer (LeConte, 1852)
* C. validicornis (Boheman, 1848)
* C. validus (Chevrolat, 1835)
* C. variabilipes (Eschscholtz, 1833)
* C. varians (Chaudoir, 1876)
* C. variicornis (A.Morawitz, 1863)
* C. variipes (Chaudoir, 1856)
* C. variolosus (Kirschenhofer, 2008)
* C. vartianorum (Mandl, 1989)
* C. velocipes (Chaudoir, 1876)
* C. venustulus (Dejean, 1831)
* C. vertagoides (Laferte-Senectere, 1851)
* C. veselyi (Kirschenhofer, 2004)
* C. vestitus (Paykull, 1790)
       Geelrandfluweelloper
* C. vethi (Bates, 1889)
* C. viangchanensis (Kirschenhofer, 1998)
* C. victor (Andrewes, 1928)
* C. victoriae (Kirschenhofer, 2009)
* C. viduus (G. Horn, 1871)
* C. vietnamensis (Kirschenhofer, 2008)
* C. vietnami (Jedlička, 1966)
* C. violaceipennis (Chaudoir, 1876)
* C. violatus (Gemminger & Harold, 1868)
* C. virens (Rambur, 1837)
* C. virescens (Chaudoir, 1835)
* C. virgulatus (Jeannel, 1949)
* C. virgulifer (Chaudoir, 1876)
* C. viridicollis (Reiche, 1843)
* C. viridis (Menetries, 1832)
* C. vitalisi (Andrewes, 1919)
* C. vitticollis (Boheman, 1848)
* C. vividus (Chaudoir, 1876)
* C. vulneratus (Dejean, 1831)
* C. waddellii (Murray, 1858)
* C. wallacei (Chaudoir, 1876)
* C. watsaensis (Burgeon, 1935)
* C. wegneri (Louwerens, 1953)
* C. wenchuanensis (Kirschenhofer, 2008)
* C. wittei (Burgeon, 1937)
* C. wittmerianus (Mandl, 1978)
* C. wrasei (Kirschenhofer, 1997)
* C. xanthacrus (Wiedemann, 1823)
* C. xanthomerus (Alluaud, 1918)
* C. xanthospilus (Wiedemann, 1821)
* C. yangonensis (Kirschenhofer, 2005)
* C. yunnanulus (Mandl, 1992)
* C. zanzibaricus (Chaudoir In Oberthur, 1883)
* C. ziloensis (Basilewsky, 1970)
* C. zygogrammus (Laferte-Senectere, 1851)
* ↑ Yves Bousquet. "Catalogue of Geadephaga (Coleoptra, Adephaga) of America, north of Mexico." ZooKeys 245 (201), blz. 963. DOI:10.3897/zookeys.245.3416
* ↑ Anichtchenko A. et al., (editors): Carabidae of the World (bezocht: 20 tot 28 maart 2013)
* ↑ Ye Liu, Hongliang Shi en Hongbin Liang. "Four new Chlaenius species (Coleoptera: Carabidae: Chlaeniini) from Asia and a key to the species of subgenus Chlaenioctenus." Zootaxa 3630(3):505-518 (2013). DOI:10.11646/zootaxa.3630.3.6