# Pterostichinae

Pterostichinae Bonelli, 1810 è una sottofamiglia di coleotteri carabidi diffusa in tutto il mondo.

## Descrizione

Gli Pterostichinae sono coleotteri di medie o piccole dimensioni, forma ovale e colore scuro, spesso metallico. Mancano quasi completamente specie dotate di decorazioni a colori contrastanti. I caratteri che permettono di distinguerli dagli altri carabidi sono: le tibie anteriori con una incavatura all'apice, le elitre a curvatura regolare, le antenne pubescenti dal 4° articolo e gli occhi sormontati da due setole. Quest'ultima caratteristica permette di riconoscerli facilmente dagli Harpalinae, con cui possono venire confusi.

## Sistematica

La sottofamiglia è suddivisa in otto tribù che comprendono i seguenti generi:
* Tribù Abacetini Chaudoir, 1872
** Abacetus Dejean, 1828
** Abacidus LeConte, 1873
** Aristopus Laferte-Senectere, 1853
** Celioinkosa Straneo, 1951
** Chlaeminus Motschulsky, 1865
** Colpodichius Straneo, 1952
** Cosmodiscus Sloane, 1907
** Cyrtomoscelis Chaudoir, 1874
** Distrigidius Jeannel, 1948
** Ecnomolaus Bates, 1892
** Haptoderidius Straneo, 1975
** Haptoderodes Straneo, 1986
** Holconotus Schmidt-Goebel, 1846
** Inkosa Peringuey, 1926
** Mateuellus Deuve, 1990
** Metabacetus Bates, 1892
** Metaxellus Straneo, 1960
** Metaxys Chaudoir, 1857
** Novillidius Straneo, 1941
** Oodinkosa Straneo, 1939
** Ophonichius Straneo, 1942
** Pediomorphus Chaudoir, 1878
** Pioprosopus Tschitscherine, 1899
** Pollicobius Vinson, 1939
** Prostalomus Basilewsky, 1950
** Pseudabacetus Burgeon, 1935
** Pterostillichus Straneo, 1949
** Rhagadillius Straneo, 1951
** Tiferonia Darlington, 1962
** Trachelocyphoides Alluaud, 1934
** Trachelocyphus Tschitscherine, 1900
* Tribù Chaetodactylini Tschitscherine, 1903
** Chaetodactyla Tschitscherine, 1897
* Tribù Cnemalobini Germain, 1911
** Cnemalobus Guerin-Meneville, 1838
* Tribù Cratocerini Lacordaire, 1854
** Sottotribù Catapieseina Bates, 1882
*** Catapiesis Solier, 1835
*** Homalomorpha Brulle, 1835
** Sottotribù Cratocerina sensu stricto
*** Brachidius Chaudoir, 1852
*** Cratocerus Dejean, 1829
*** Oxyglychus Straneo, 1938
** Sottotribù Drimostomatina Chaudoir, 1872
*** Abacaelostus Straneo, 1952
*** Andrewesinulus Straneo, 1938
*** Apsidocnemus Alluaud, 1936
*** Barylaus Liebherr, 1985
*** Caecocaelus Straneo, 1949
*** Caelostomus W.S. MacLeay, 1825
*** Camptogenys Tschitscherine, 1899
*** Capabatus Csiki, 1930
*** Crenulostrigus Straneo, 1942
*** Cyrtolaus Bates, 1882
*** Dactyleurys Tschitscherine, 1899
*** Dactylinius Straneo, 1941
*** Diachipteryx Alluaud, 1926
*** Diceromerus Chaudoir, 1872
*** Dromistomus Jeannel, 1948
*** Drymonaxus Straneo, 1941
*** Eurystomis Chaudoir, 1878
*** Feostoma Straneo, 1941
*** Hemitelestus Alluaud, 1895
*** Hoplizomenus Chaudoir, 1872
*** Leleuporites Straneo, 1960
*** Madapelmus Dajoz, 1985
*** Monodryxus Straneo, 1942
*** Pachycaecus Straneo, 1971
*** Pachyroxochus Straneo, 1942
*** Platyxythrius Straneo, 1942
*** Stegazopteryx Will, 2004
*** Stomonaxellus Tschitscherine, 1901
*** Strigomerodes Straneo, 1939
*** Strigomerus Chaudoir, 1872
*** Trichillinus Straneo, 1938
* Tribù Microcheilini Jeannel, 1948
** Microcheila Brulle, 1834
* Tribù Morionini Brulle, 1835
** Buderes Murray, 1857
** Hyperectenus Alluaud, 1935
** Hyperion Castelnau, 1834
** Megamorio Chaudoir, 1880
** Morion Latreille, 1810
** Morionidius Chaudoir, 1880
** Moriosomus Motschulsky, 1864
** Platynodes Westwood, 1846
** Stereostoma Murray, 1857
* Tribù Pterostichini Bonelli, 1810
** Sottotribù Euchroina Chaudoir, 1874
*** Abacillius Straneo, 1949
*** Abacillodes Straneo, 1988
*** Abaris Dejean, 1831
*** Apsaustodon Tschitscherine, 1901
*** Argutoridius Chaudoir, 1876
*** Blennidus Motschulsky, 1865
*** Blenniventer Straneo, 1986
*** Bothynoproctus Tschitscherine, 1900
*** Cephalostichus Straneo, 1977
*** Cynthidia Chaudoir, 1873
*** Euchroa Brulle, 1834
*** Eumara Tschitscherine, 1901
*** Gastrogmus Sloane, 1915
*** Haplobothynus Tschitscherine, 1901
*** Hybothecus Chaudoir, 1874
*** Litarthrum Sloane, 1915
*** Lobobrachus Sharp, 1885
*** Marsyas Putzeys, 1845
*** Meropalpus Tschitscherine, 1900
*** Microcephalus Dejean, 1828
*** Mireius Straneo, 1984
*** Nelidus Chaudoir, 1878
*** Neotalus Will, 2002
*** Oribazus Chaudoir, 1874
*** Pachythecus Chaudoir, 1874
*** Phaenaulax Tschitscherine, 1898
*** Prosopogmus Chaudoir, 1865
*** Pseudabarys Chaudoir, 1873
*** Setalimorphus Sloane, 1895
*** Setalis Castelnau, 1867
*** Simodontus Chaudoir, 1843
*** Trirammatus Chaudoir, 1838
** Sottotribù Loxandrina Erwin & Sims, 1984
*** Adrimus Bates, 1871
*** Cerabilia Castelnau, 1867
*** Feronista Moore, 1965
*** Haploferonia Darlington, 1962
*** Homalonesiota Maindron, 1908
*** Loxandrus LeConte, 1852
*** Metoncidus Bates, 1871
*** Nebrioferonia Straneo, 1939
*** Oxycrepis Reiche, 1843
*** Stolonis Motschulsky, 1866
*** Zeodera Castelnau, 1867
** Sottotribù Metiina Straneo, 1951
*** Abropus Waterhouse, 1842
*** Antarctiola Straneo, 1977
*** Chaetauchenium Tschitscherine, 1900
*** Feroniola Tschitscherine, 1900
*** Kuschelinus Straneo, 1963
*** Metius Curtis, 1839
*** Parhypates Motschulsky, 1866
** Sottotribù Molopina Bonelli, 1810
*** Abacops Tschitscherine, 1902
*** Abax Bonelli, 1810
*** Aristochroa Tschitscherine, 1898
*** Aristochroodes Marcillac, 1993
*** Camptoscelis Dejean, 1828
*** Cedrorum Borges & Serrano, 1993
*** Chalcochrous Chaudoir, 1838
*** Eucamptognathus Chaudoir, 1837
*** Eudromus Klug, 1835
*** Henrotius Jeannel, 1953
*** Lesticus Dejean, 1828
*** Molopidius Jeannel, 1942
*** Molopinus Jeannel, 1948
*** Molops Bonelli, 1810
*** Myas Sturm, 1826: 171
*** Nirmala Andrewes, 1930
*** Orthomus Chaudoir, 1838
*** Oscadytes Lagar Mascaro, 1975
*** Pedius Motschulsky, 1850
*** Percus Bonelli, 1810
*** Peyrieraselus Deuve, 1981
*** Sthenocranion Tschitscherine, 1899
*** Styracoderus Chaudoir, 1874
*** Teratotarsa Tschitscherine, 1893
*** Trigonognatha Motschulsky, 1858
*** Tropidocerus Chaudoir, 1878
*** Zariquieya Jeannel, 1924
** Sottotribù Pterostichina sensu stricto
*** Abacoleptus Fauvel, 1903
*** Abacomorphus Chaudoir, 1878
*** Abacophrastus Will, 2011
*** Acanthoferonia Moore, 1965
*** Allotriopus Bates, 1882
*** Analoma Darlington, 1971
*** Aulacopodus Britton, 1940
*** Basilewskya Straneo, 1948
*** Castelnaudia Tschitscherine, 1890
*** Catadromus W.S. Macleay, 1825
*** Conchitella Moore, 1962
*** Cophosomorpha Tschitscherine, 1890
*** Cratoferonia Tschitscherine, 1902
*** Cratogaster Blanchard, 1843
*** Cyclotrachelus Chaudoir, 1838
*** Cyrtoderus Hope, 1842
*** Darodilia Castelnau, 1867
*** Delinius Westwood, 1864
*** Euryabax Fauvel, 1903
*** Euryaptus Bates, 1892
*** Gastrellarius Casey, 1918
*** Gourlayia Britton, 1964
*** Harpostomus Chaudoir, 1856
*** Holcaspis Chaudoir, 1865
*** Leiradira Castelnau, 1867
*** Liopasa Tschitscherine, 1901
*** Lophoglossus LeConte, 1852
*** Loxodactylus Chaudoir, 1865
*** Loxogenius Sloane, 1907
*** Mecynognathus W.J.MacLeay, 1873
*** Megadromus Motschulsky, 1865
*** Neoferonia Britton, 1940
*** Nesites Andrewes, 1939
*** Notabax Moore, 1976
*** Notolestus Sloane, 1894
*** Notonomus Chaudoir, 1862
*** Nurus Motschulsky, 1866
*** Oberthueria Vuillet, 1911
*** Ogmophora Tschitscherine, 1896
*** Onawea Johns, 2007
*** Pachymorphus Chaudoir, 1838
*** Paniestichus Will, 2011
*** Paranurus Tschitscherine, 1901
*** Pareuryaptus Dubault, Lassalle & Roux, 2008
*** Percolaus Bates, 1882
*** Piesmus LeConte, 1852
*** Platycaelus Blanchard, 1843
*** Platysmodes Fauvel, 1903
*** Plocamostethus Britton, 1940
*** Poecilinus Jeannel, 1948
*** Poecilus Bonelli, 1810
*** Psegmatopterus Chaudoir, 1878
*** Pseudoceneus Tschitscherine, 1891
*** Pterostichus Bonelli, 1810
*** Rhabdotus Chaudoir, 1865
*** Rhytiferonia Darlington, 1962
*** Rhytisternus Chaudoir, 1865
*** Sarticus Motschulsky, 1865
*** Secatophus Castelnau, 1867
*** Setalidius Chaudoir, 1878
*** Speluncarius Reitter, 1886
*** Speomolops Patrizi, 1955
*** Sphodrosomus Perroud & Montrouzier, 1864
*** Stereocerus Kirby, 1837
*** Steropanus Fairmaire, 1888
*** Stomis Clairville, 1806
*** Straneostichus Sciaky, 1994
*** Taphoxomimus Straneo, 1975
*** Tapinopterus Schaum, 1858
*** Teropha Castelnau, 1867
*** Trichosternus Chaudoir, 1865
*** Trigonotoma Dejean, 1828
*** Wahlbergiana Bousquet, 2002
*** Xenion Tschitscherine, 1902
*** Zeopoecilus Sharp, 1886
* Tribù Zabrini Bonelli, 1810
** Sottotribù Amarina C. Zimmermann, 1831
*** Amara Bonelli, 1810
*** Pseudamara Lindroth, 1968
** Sottotribù Zabrina Bonelli, 1810
*** Zabrus Clairville, 1806