# Flora of New York/Polypodiales

| ← Pre-polypod ferns | Flora of New York — Polypodiales | Flora of New York — Polypodiales | Flora of New York — Polypodiales | Flora of New York — Polypodiales | → Pinales |
| ← Pre-polypod ferns | Table of contents | Genus index | Protected species index | Invasive species index | → Pinales |
| Subclass | Order | Suborder | Family | Subfamily | Genus | Subgenus | Common names | # |
| Equisetidae | Equisetales | — | Equisetaceae | — | Equisetum | Equisetum | horsetail | 6 |
| Equisetidae | Equisetales | — | Equisetaceae | — | Equisetum | Hippochaete | scouring rush | 7 |
| Ophioglossidae | Ophioglossales | — | Ophioglossaceae | Botrychioideae | Botrychium | Botrychium | moonwort, grape fern | 8 |
| Ophioglossidae | Ophioglossales | — | Ophioglossaceae | Botrychioideae | Botrychium | Osmundopteris | rattlesnake fern | 1 |
| Ophioglossidae | Ophioglossales | — | Ophioglossaceae | Botrychioideae | Botrychium | Sceptridium | evergreen grapefern | 4 |
| Ophioglossidae | Ophioglossales | — | Ophioglossaceae | Ophioglossoideae | Ophioglossum | Ophioglossum | adder's tongue | 1 |
| Polypodiidae (leptosporangiate ferns) | Osmundales | — | Osmundaceae | — | Osmundastrum | Osmundastrum | cinnamon fern | 1 |
| Polypodiidae (leptosporangiate ferns) | Osmundales | — | Osmundaceae | — | Osmunda | Claytosmunda | interrupted fern | 1 |
| Polypodiidae (leptosporangiate ferns) | Osmundales | — | Osmundaceae | — | Osmunda | Osmunda | royal fern | 1 |
| Polypodiidae (leptosporangiate ferns) | Hymenophyllales | — | Hymenophyllaceae | Trichomanoideae | Crepidomanes | Crepidomanes | grotto felt, weft fern | 1 |
| Polypodiidae (leptosporangiate ferns) | Schizaeales | — | Lygodiaceae | — | Lygodium | Lygodium | climbing fern | 1 |
| Polypodiidae (leptosporangiate ferns) | Schizaeales | — | Schizaeaceae | — | Schizaea | Schizaea | curlygrass fern | 1 |
| Polypodiidae (leptosporangiate ferns) | Salviniales | — | Salviniaceae | — | Azolla | Azolla | mosquitofern | 1 |
| Polypodiidae (leptosporangiate ferns) | Salviniales | — | Salviniaceae | — | Salvinia | Salvinia | watermoss, floatingfern | x |
| Polypodiidae (leptosporangiate ferns) | Salviniales | — | Marsileaceae | — | Marsilea | Marsilea | waterclover, waterfern | 1 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Dennstaedtiineae | Dennstaedtiaceae | — | Dennstaedtia | Dennstaedtia | hay-scented fern | 1 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Dennstaedtiineae | Dennstaedtiaceae | — | Pteridium | Pteridium | brackenfern | 2 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Pteridineae | Pteridaceae | Cryptogrammoideae | Cryptogramma | Cryptogramma | rock brake | 1 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Pteridineae | Pteridaceae | Pteridoideae | Pteris | Pteris | brake fern | x |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Pteridineae | Pteridaceae | Cheilanthoideae | Pellaea | Pellaea | cliff brake | 2 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Pteridineae | Pteridaceae | Cheilanthoideae | Cheilanthes | Cheilanthes | lip fern | 1 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Pteridineae | Pteridaceae | Vittarioideae | Adiantum | Adiantum | maidenhair-fern | 1 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Pteridineae | Pteridaceae | Vittarioideae | Vittaria | Vittaria | ribbon fern | 1 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Aspleniineae | Cystopteridaceae | — | Cystopteris | Cystopteris | bladder fern | 5 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Aspleniineae | Cystopteridaceae | — | Gymnocarpium | Gymnocarpium | oak fern | 2 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Aspleniineae | Diplaziopsidaceae | — | Homalosorus | Homalosorus | glade fern | 1 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Aspleniineae | Aspleniaceae | — | Asplenium | Asplenium | spleenwort | 10 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Aspleniineae | Thelypteridaceae | Phegopteridoideae | Phegopteris | Phegopteris | beech fern | 2 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Aspleniineae | Thelypteridaceae | Thelypteridoideae | Thelypteris | Thelypteris | marsh fern, New York fern | 3 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Aspleniineae | Woodsiaceae | — | Woodsia | Woodsia | cliff fern | 5 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Aspleniineae | Onocleaceae | — | Onoclea | Onoclea | sensitive fern | 1 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Aspleniineae | Onocleaceae | — | Matteuccia | Matteuccia | ostrich fern | 1 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Aspleniineae | Blechnaceae | Woodwardioideae | Anchistea | Anchistea | Virginia chainfern | 1 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Aspleniineae | Blechnaceae | Woodwardioideae | Lorinseria | Lorinseria | netted chainfern | 1 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Aspleniineae | Athyriaceae | — | Deparia | Deparia | false spleenwort | 1 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Aspleniineae | Athyriaceae | — | Athyrium | Athyrium | lady fern | 3 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Polypodiineae | Dryopteridaceae | Dryopteridoideae | Cyrtomium | Cyrtomium | netvein hollyfern | x |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Polypodiineae | Dryopteridaceae | Dryopteridoideae | Dryopteris | Dryopteris | woodfern | 30 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Polypodiineae | Dryopteridaceae | Dryopteridoideae | Polystichum | Polystichum | hollyfern | 5 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Polypodiineae | Polypodiaceae | Polypodioideae | Polypodium | Polypodium | polypody, rockcap fern | 3 |
| Polypodiidae (leptosporangiate ferns) | Polypodiales | Polypodiineae | Polypodiaceae | Polypodioideae | Pleopeltis | Pleopeltis | resurrection fern | x |

## Suborder Pteridineae
### Family Pteridaceae

The Pteridaceae E.D.M.Kirchn. (1831), (brakefern family)...

#### Subfamily Cryptogrammoideae
##### Cryptogramma

* Cryptogramma stelleri slender cliffbrake, Steller’s cliffbrake
Polypodiales — Pteridaceae — Cryptogrammoideae — Cryptogramma :
| Genus Cryptogramma | Rock-brake | N.Y. Status | Images | Distribution | NY NPT |
| Cryptogramma stelleri (S.G.Gmel.) Prantl1768. Pteris stelleri S.G.Gmel. 1882. Cryptogramma stelleri Prantl | Fragile rockbrake, Slender cliff brake, Slender rock-brake, Steller's rock brakeCryptogramme de steller | Native, CoC: 9, Vulnerable, DEC-D FACU Perennial, Herb-fern, 2n=60 | | NYFA: 3-4 | USDA-N Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos NatureServe IPN Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ||||||

#### Subfamily Pteridoideae
##### Pteris

* Pteris multifida spider brakePteris multifida, the spider brake was listed as present in New York by the 1955 American Fern Journal, which cited Willard Nelson Clute:
* New York: It is fond of growing on old walls as far north as New York City — Clute (1938, p. 102).
* USDA-NRCS Plants Database lists the 1958 American Fern Journal as the source for the plant being listed in New York, but it doesn't appear to be listed in that volume. Pteris multifida is now been listed as "excluded" in the New York Flora Atlas.
* ↑ G.N. Jones (1955) "On the Occurrence of Pteris multifida in the United States." American Fern Journal, 45:107-111.
* ↑ USDA-NRCS Plants Database for Pteris multifida (July 21, 2015)
Polypodiales — Pteridaceae — Pteridoideae — Pteris :
| Genus Pteris (excluded taxa) | Brake-fern | N.Y. Status | Images | Distribution | NY NPT |
| Pteris multifida Poir.1804. Pteris multifida Poir. 1938. Pycnodoria multifida (Poir.) Small  auct. Pteris serrulata non L.f. | Spider brake, Spider fern, Chinese brake, Huguenot fern, Saw-leaved bracken | Introduced from  temperate Asia,  tropical Asia, N.Y. excluded | | NYFA-Excluded | USDA-X0 ARS-GRIN ITIS FNA Tropicos IPN Images Wikispecies |

#### Subfamily Cheilanthoideae

* Subfamily Cheilanthoideae.

##### Pellaea

* Pellaea atropurpureaCliff-brake ferns
Polypodiales — Pteridaceae — Cheilanthoideae — Pellaea :
| Genus Pellaea | Cliff-brake | N.Y. Status | Distribution | NY NPT |
| Pellaea atropurpurea (L.) Link    Pteris atropurpurea (L.) Link | Purple cliffbrake, Purple-stemmed cliff-brake | Native, Vulnerable, DEC-D | NYFA: 4 | USDA-NN Go Botany Images Wikispecies |
| Pellaea glabella Mett. ex Kuhnssp. glabella1858. Pellaea rigida  var. rigida   1869. Pellaea glabella  var. glabella       Pellaea glabella  var. bushii  Mack.     Pellaea glabella  ssp. glabella   1902. Pellaea atropurpurea  var. bushii  Mack. & Bush 1938. Pellaea atropurpurea  f. glabella  Clute | Smooth cliff-brake ⓘ Slender cliff-brake ⓘ Dwarf cliffbrake Simple cliffbrake Pelléade glabre ⓘ Pelléade glabre typique ⓘ Pelléade glabre variété glabre ⓘ | Native, Threatened, DEC-B | NYFA: 2 | USDA-NN Go Botany VASCAN FNA Tropicos Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (b) Threatened. ||||||

##### Myriopteris

* Myriopteris lanosaHairy lip-fern or woolly lip fern (Myriopteris lanosa) can no longer be found in any of its known sites but it's possible that it's present somewhere in the state.
* ↑ 2023 New York Flora Atlas. New York Flora Association, Albany, New York.
Polypodiales — Pteridaceae — Cheilanthoideae — Myriopteris :
| Genus Myriopteris | Lip-fern | N.Y. Status | Images | Distribution | NY NPT |
| Myriopteris lanosa (Michx.) Grusz & Windham1803. Nephrodium lanosum Michx. 1804. Adiantum vestitum Spreng. 1806. Cheilanthes vestita Sw. 1859. Cheilanthes lanosa (Michx.) D.C.Eaton 2013. M. lanosa (Michx.) Grusz & Windham | Hairy lipfern, Wooly lip-fern | Native, CoC: 10, Endangered, DEC-A, S1, G5 | | Bronx, Dutchess, New York, Orange, Rockland, Ulster, Westchester NYFA: Z | USDA-N0 Go Botany ARS-GRIN ITIS FNA Tropicos BONAP Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered. ||||||

#### Subfamily Vittarioideae

* The Vittarioideae subfamily contains the former families Adiantaceae (maidenhair ferns) and Vittariaceae (shoestring ferns).

##### Adiantum

* Adiantum pedatum maidenhair fern
Polypodiales — Pteridaceae — Vittarioideae — Adiantum :
| Genus Adiantum | Maidenhair-fern | N.Y. Status | Distribution | NY NPT |
| Adiantum pedatum L.1753. Adiantum pedatum L. | Maidenhair fern, Northern maidenhair-fern, Northern maidenhair, Five-finger fernAdiante du Canada | Native, Likely secure, DEC-D | NYFA: 4 | USDA-NN Go Botany VASCAN ARS-GRIN FNA Tropicos Images Wikispecies |
| Adiantum capillus-veneris L.1753. Adiantum capillus-veneris | Venus’s hair fern ⓘ Common maidenhair ⓘ Southern maidenhair ⓘ Black maidenhair ⓘ | Introduced from  southern US, N. America native, Not naturalized | NYFA: Greene (1923) iNat: Seuben (2021), New York (2021) NYFA: Xm iNat | USDA-NN Go Botany-0 ARS-GRIN Tropicos Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ||||||

##### Vittaria

* Vittaria appalachiana Appalachian shoestring fernThe genus Vittaria is represented in New York by the endangered Appalachian shoestring fern, which has been found in only a few counties and only in its gametophyte stage.
* ↑ New York Flora Atlas: Vittaria appalachiana appears only as "minute moss-like gametophytes" in New York.
Polypodiales — Pteridaceae — Vittarioideae — Vittaria :
| Genus Vittaria | Shoestring-fern | N.Y. Status | Images | Distribution | NY NPT |
| Vittaria appalachiana Farrar & Mickel (1991) | Appalachian shoestring fern, Appalachian vittaria, Appalachian gametophyte fern | Native, Endangered, DEC-A | | NYFA: 3 counties iNat: Chautauqua (2021) NYFA: 1 | USDA-N0 Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered. ||||||

## Suborder Dennstaedtiineae
### Family Dennstaedtiaceae

* The Dennstaedtiaceae Lotsy (1909), (bracken-fern family)
* Although they are native to the Northeast, the hay-scented fern (Dennstaedtia punctilobula), and brackenfern (Pteridium aquilinum) are considered to be "interfering" plants in New York forests. These ferns produce dense stands that prevent the establishment of tree seedlings. While this has been found to be beneficial in limited situations, such as under power-lines and on ski-area trails, these ferns are generally seen as detrimental to forest regeneration.
* ↑ Heather M. Engelman & Ralph D. Nyland (2006). "Interference to Hardwood Regeneration in Northeastern North America: Assessing and Countering Ferns in Northern Hardwood Forests." Northern Journal of Applied Forestry (Society of American Foresters) 23(3):166–175.

##### Dennstaedtia

* Dennstaedtia punctilobula hay-scented fernHay-scented fern (Dennstaedtia punctilobula) is often considered to be an interfering species in New York forest understories, where large deer populations over-browse other native plants but avoid this fern, which prevents tree regeneration.
* ↑ Peter Smallidge (2015). "Native Invasive: Hay-scented Fern." The Overstory. New York Forest Owners Association, Southeatern Adirondack Chapter, 26:2.
* ↑ Fei, S., Gould, P., Kaeser, M., & Steiner, K. (2010). Distribution and Dynamics of the Invasive Native Hay-Scented Fern. Weed Science, 58(4), 408-412. doi:10.1614/WS-D-10-00040.1
Polypodiales — Dennstaedtiaceae — Dennstaedtia :
| Genus Dennstaedtia | Hay-scented fern | N.Y. Status | Images | Distribution | NY NPT |
| Dennstaedtia punctilobula (Michx.) T.Moore1803. Nephrodium punctilobulum Michx. 1810. Aspidium punctilobum Willd. 1827. Sitobolium punctilobum Desv. 1848. Dicksonia punctilobula A.Gray 1857. Dennstaedtia punctilobula T.Moore | Hay-scented fern, Eastern hay-scented fern, Hay-scent fern, Boulder fernDennstaedtie à   lobules ponctués | Native, CoC: 4, Secure, Interfering FACU-UPL Perennial, Herb-fern | | NYFA: 5 | USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos Images Wikispecies |

##### Pteridium

* Pteridium aquilinum eastern bracken fern
Polypodiales — Dennstaedtiaceae — Pteridium :
| Genus Pteridium | Brackenfern | N.Y. Status | Distribution | NY NPT |
| Pteridium latiusculum (Desv.) Hieron.1827. Pteris latiuscula Desv. in Mém. Soc. Linn. Paris 6: 303 1909. Pteridium aquilinum var. latiusculum (Desv.) Underw. ex A.Heller in Cat. N. Amer. Pl.:17 1914. Pteridium latiusculum (Desv.) Hieron. in R.E.Fries, Wiss. Erg. Schwed. Rhodesia-Kongo-Exp. 1941. Pteridium aquilinum ssp. latiusculum (Desv.) Hultén in Acta Univ. Lund., 2, 37(1):44 1943. Pteris aquilina subvar. latiuscula (Desv.) Farw. in Amer. Fern J. 33:9 1949. Pteridium aquilinum ssp. latiusculum (Desv.) Hultén ex R.T.Clausen in Cornell Univ. Ag. Mem. | Eastern brackenfern, Eastern bracken, BrackenfernFougère-aigle de l'Est | Native, CoC: 3, Secure, Interfering FACU Perennial, Fern, Heliophily: 6 | NYFA: 5 | USDA-NN POWO FSUS Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos BONAP Images Wikispecies |
| Pteridium pseudocaudatum (Clute) Christenh.1900. Pteris aquilina var. pseudocaudata Clute in Fern Bull. 8:39 1900. Pteridium aquilinum var. pseudocaudatum Clute in Fern Bull. 8:39 1901. Pteris pseudocaudata (Clute) Hay, T.S.Palmer & C.D.White in Proc. Biol. Soc. Washington 1919. Pteridium latiusculum var. pseudocaudatum (Clute) Maxon in Amer. Fern J. 9:44 1931. Filix-foemina aquilina var. pseudocaudata (Clute) Farw. in Amer. Midl. Naturalist 12:290 1935. Pteris latiuscula var. pseudocaudata (Clute) E.P.St.John & R.P.St.John in Amer. Fern J. 25:40 1941. Pteridium aquilinum ssp. pseudocaudatum (Clute) Hultén in Acta Univ. Lund., 2, 37(1):44 2018. Pteridium pseudocaudatum (Clute) Christenh. in Global Fl. 4:7 | Southeastern Bracken, Tailed bracken, Brackenfern, Western brackenfern | Native, CoC: 3, Endangered, DEC-A FACU Perennial, Fern, Heliophily: 5 | NYFA: Richmond (1889-1901), Suffolk (1932) NYFA: Z | USDA-N0 FSUS Go Botany ITIS FNA Tropicos BONAP Images Wikispecies |
| Pteridium aquilinum (L.) Kuhn1753. Pteris aquilina L. in Sp. Pl.:795 1799. Asplenium aquilinum (L.) Bernh. in J. Bot. (Schrader) 1799(1):310 1879. Pteridium aquilinum (L.) Kuhn in C.C.von der Decken, Reisen Ost Afr. 3(3):11 1941. Pteridium aquilinum ssp. typicum R.M. Tryon in Rhodora 43:15, not validly publ. | Western brackenfern | Introduced, N.Y. excluded | NYFA-Excluded | USDA-NN POWO ARS-GRIN FNA Tropicos BONAP Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered. ||||||

## Suborder Aspleniineae

* The classification used here follows PPG I (2016), which places all of eupolypods II in suborder Aspleniineae.
* Aspleniineae
* Cystopteridaceae
* Cystopteris (4 bladderferns)
* Gymnocarpium dryopteris (northern oak fern)
* Diplaziopsidaceae—Homalosorus pycnocarpos (narrow-leaved glade fern)
* Aspleniaceae—Asplenium (11 spleenworts)
* Thelypteridaceae
* Phegopteridoideae—Phegopteris (2 beech ferns)
* Thelypteridoideae—Thelypteris (3 marsh ferns)
* Woodsiaceae—Woodsia (5 cliff ferns)
* Athyriaceae
* Athyrium (2 lady ferns)
* Deparia acrostichoides (silvery spleenwort)
* Onocleaceae
* Onoclea sensibilis (sensitive fern)
* Matteuccia struthiopteris (ostrich fern)
* Blechnaceae : Woodwardioideae
* Anchistea virginica (Virginia chainfern)
* Lorinseria areolata (netted chainfern)
* ↑ PPG I (2016), A community‐derived classification for extant lycophytes and ferns. Jnl of Sytematics Evolution, 54: 563-603. https://doi.org/10.1111/jse.12229

### Family Cystopteridaceae

The Cystopteridaceae Ching & Z.R.Wang (1997) (bladder-ferns)...

#### Cystopteris

* Cystopteris bulbifera bulblet fern
Polypodiales — Cystopteridaceae — Cystopteris :
| Genus Cystopteris | Bladder-fern | N.Y. Status | Images | Distribution | NY NPT |
| Cystopteris bulbifera (L.) Bernh.1753. Polypodium bulbiferum L. 1806. Cystopteris bulbifera (L.) Bernh. 1900. Filix bulbifera (L.) Underw. | Bulblet fern, Bulblet bladder-fern, Bulblet fragile-fern, Berry fernCystoptère bulbifère, Cystoptèride bulbifère | Native, CoC: 8, Secure, DEC-D Perennial, Herb-fern | | NYFA: 5 | USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNA Images Wikispecies |
| Cystopteris fragilis (L.) Bernh.1753. Polypodium fragile L. 1806. Cystopteris fragilis (L.) Bernh. 1848. Cystopteris dickieana  R.Sim 1900. Filix fragilis (L.) Underw. | Brittle bladder-fern, Fragile fern | Native, CoC: 7, Likely secure, DEC-D Perennial, Herb-fern | | NYFA: 4 | USDA-NN Go Botany Images Wikispecies |
| Cystopteris tenuis (Michx.) Desv.  Cystopteris protrusa ×   "Cystopteris hemifragilis"†1803. Nephrodium tenue Michx. 1827. Cystopteris tenuis (Michx.) Desv. 1889. C. fragilis  var. mackayi  G.Lawson | Mackay's bladder-fern, Mackay's fragile fern, Upland brittle bladder-fernCystoptère ténue | Native, CoC: 7, Likely secure, DEC-D Perennial, Herb-fern | | NYFA: 4 | USDA-NN Go Botany Images Wikispecies |
| Cystopteris protrusa (Weath.) Blasdell1935. Cystopteris fragilis  var. protrusa  Weath. 1963. Cystopteris protrusa Blasdell | Lowland bladder-fern, Lowland brittle fern, Lowland fragile fern, Southern bladder-fern | Native, CoC: 10, Endangered, DEC-A Perennial, Herb-fern | | Broome, Erie, Richmond, Suffolk, Tompkins NYFA: 1 | USDA-NN Go Botany FNA Images Wikispecies |
| Genus Cystopteris (excluded taxa) | Bladder-fern | N.Y. Status | Images | Distribution | NY NPT |
| Cystopteris bulbifera × fragilis  Cystopteris bulbifera ×   Cystopteris fragilis | Hybrid of  bulblet bladder-fern &  fragile fern | Native, N.Y. excluded Perennial, Herb-fern | | NYFA-Excluded | Images Wikispecies |
| 1 2 3 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered. ||||||

#### Gymnocarpium

* Gymnocarpium dryopteris
Polypodiales — Cystopteridaceae — Gymnocarpium :
| Genus Gymnocarpium | Oak-fern | N.Y. Status | Images | Distribution | NY NPT |
| Gymnocarpium dryopteris (L.) Newman1753. Polypodium dryopteris L. 1851. G. dryopteris (L.) Newman 1852. Phegopteris dryopteris (L.) Fée 1905. Dryopteris linnaeana C. Chr. 1909. Dryopteris dryopteris (L.) Christ 1917. Thelypteris dryopteris (L.) Sloss. 1966. Carpogymnia dryopteris Á.& D.Löve | Northern oak fern, Common oak fern, Western oak fernGymnocarpe fougère-du-chêne | Native, CoC: 8, Likely secure, DEC-D | | NYFA: 4-5 | USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNA Images Wikispecies |
| Gymnocarpium × brittonianum (Sarvela) Pryer & Haufler  Gymnocarpium disjunctum ×   Gymnocarpium dryopteris1980. Gymnocarpium dryopteris  ssp. brittonianum  Sarvela 1993. Gymnocarpium × brittonianum Pryer & Haufler | Britton's fern, Britton's oak-fernGymnocarpe de Britton | Native, Unranked | | NYFA: U | USDA-NN VASCAN ITIS FNA Images Wikispecies |
| Genus Gymnocarpium (excluded) | Oak-fern | N.Y. Status | Images | Distribution | NY NPT |
| Gymnocarpium disjunctum (Rupr.) Ching1845. Polypodium dryopteris  var. disjunctum  Rupr. 1965. G. disjunctum (Rupr.) Ching | Pacific oakfern, Western oak fernGymnocarpe disjoint, Dryoptère disjointe, Dryoptèride du chêne | N. America native, N.Y. excluded | | NYFA-Excluded | USDA-NN VASCAN ITIS FNA Tropicos Images Wikispecies |
| Gymnocarpium robertianum (Hoffm.) Newman1795. Polypodium robertianum Hoffm. 1850. Phegopteris robertianum (Hoffm.) Fée 1851. Gymnocarpium robertianum (Hoffm.) Newman 1905. Dryopteris robertiana (Hoffm.) C.Chr. 1917. Thelypteris robertiana (Hoffm.) Sloss. ex Rydb. 1962. Gymnocarpium dryopteris  var. pumilum  B.Boivin | Scented oakfern, Limestone fern, Robert's oak fern, Northern oak fernGymnocarpe de Robert | N. America native, N.Y. excluded | | NYFA-Excluded | USDA-NN VASCAN ITIS FNA Tropicos Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ||||||

### Family Diplaziopsidaceae

The glade fern family includes the genera Diplaziopsis C. Chr. (2–4 spp.) and Homalosorus Pic. Serm. (1 sp.)

#### Homalosorus

* Homalosorus pycnocarpos narrow-leaved glade fern
Polypodiales — Aspleniaceae — Athyrioideae — Homalosorus :
| Genus Homalosorus Sw. | Twin-sorus fern | N.Y. Status | Images | Distribution | NY NPT |
| Homalosorus pycnocarpos Serm.1803. Asplenium angustifolium Michx. in… 1804. Asplenium pycnocarpon Spreng. 1906. Athyrium pycnocarpon Tidestr. 1938. Diplazium pycnocarpon M.Broun 1977. Homalosorus pycnocarpos Serm. | Glade fern, Narrow-leaved glade fern, Narrow-leaved-spleenwortAthyrie à sores denses, Diplazie à sores denses | Native, CoC: 9, Likely secure, DEC-D FAC Perennial, Fern | | NYFA: 4 | USDA-NN Go Botany VASCAN ITIS FNA Tropicos Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ||||||

### Family Aspleniaceae

The Aspleniaceae Link (1841) (spleenworts)...

#### Asplenium

* Asplenium platyneuron ebony spleenwortSpleenworts (Asplenium) have the largest number of species (over 700 worldwide) of any fern genus. However, only about 10 Asplenium species occur in New York State. Here Asplenium is divided into clades acording to Xu et al. (2020).
* Asplenium
* II.
* Asplenium scolopendrium (hart's-tongue fern)
* V.
* Asplenium platyneuron (ebony spleenwort)
* Asplenium rhizophyllum (walking fern)
* VI.
* Asplenium trichomanes (maidenhair spleenwort)
* Asplenium viride (green spleenwort)
* XI.
* Asplenium ruta-muraria (wall rue)
* Asplenium montanum (mountain spleenwort)
* Asplenium bradleyi (Bradley’s spleenwort)
* ↑ Xu, K.‐W., Zhang, L., Rothfels, C.J., Smith, A.R., Viane, R., Lorence, D., Wood, K.R., Chen, C.‐W., Knapp, R., Zhou, L., Lu, N.T., Zhou, X.‐M., Wei, H.‐J., Fan, Q., Chen, S.‐F., Cicuzza, D., Gao, X.‐F., Liao, W.‐B. and Zhang, L.‐B. (2020), A global plastid phylogeny of the fern genus Asplenium (Aspleniaceae). Cladistics, 36: 22-71. https://doi.org/10.1111/cla.12384

##### Asplenium II. Phyllitis clade

* Asplenium scolopendriumThe Phyllitis clade contains the Asplenium species previously assigned to genus Phyllitis, as well as species previously in genera Ceterach and Ceterachopsis. The sole New York species in this clade, Asplenium scolopendrium, was named as such by Linnaeus in 1753 but was segregated into Phyllitis by Newman in 1844.
Polypodiales — Aspleniaceae — Asplenium :
| Genus Asplenium | Spleenwort | N.Y. Status | Images | Distribution | NY NPT |
| Asplenium scolopendrium L.var. americanum (Fernald) Kartesz & Gandhi1793. Scolopendrium vulgare Small 1935. Phyllitis scolopendrium  var. americana  Fernald 1954. Phyllitis fernaldiana Á. Löve 1973. Phyllitis japonica  ssp. americana   1991. A. scolopendrium  var. americanum | American  hart's-tongue fern, Hart's-tongue fern, Hart's tonguefern | Native, CoC: 9, Threatened, DEC-B | | Genesee, Madison, Onondaga NYFA: 2 | USDA-N ARS-GRIN FNA Tropicos Images Wikispecies |
| Genus Asplenium (excluded taxa) | Spleenwort | N.Y. Status | Images | Distribution | NY NPT |
| Asplenium scolopendrium L.var. scolopendrium1753. Asplenium scolopendrium L. 1844. Phyllitis scolopendrium (L.) Newman 1881. Scolopendrium scolopendrium H.Karst. | Hart's-tongue fern | Introduced from  Eurasia,  northern Africa, N.Y. excluded, Cultivated | | NYFA-Excluded | ARS-GRIN Tropicos Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (b) Threatened. ||||||

##### Asplenium V. Schaffneria clade

Polypodiales — Aspleniaceae — Asplenium :
| Genus Asplenium | Spleenwort | N.Y. Status | Distribution | NY NPT |
| Asplenium platyneuronvar. platyneuron1753. Acrostichum platyneuros L. 1789. Asplenium ebeneum Aiton 1878. Asplenium platyneuron Oakes ex D.C. Eaton 1888. Asplenium platyneuron (L.) Britton, Sterns & Poggenb. 1931. Chamaefilix platyneuros (L.) Farw. | Ebony spleenwort | Native, CoC: 5, Secure, DEC-D | NYFA: 5 | USDA-NN Go Botany Tropicos Images Wikispecies |
| Asplenium rhizophyllum L.1753. Asplenium rhizophyllum L.1833. Camptosorus rhizophyllus (L.) Link | Walking fern, Walking-fern spleenwort, American walking fernDoradille ambulante, Camptosore à  feuilles radicantes | Native, CoC: 10, Likely secure, DEC-D | NYFA: 4-5 iNat | USDA-NN Go Botany VASCAN ARS-GRIN ITIS Images Wikispecies |
| Asplenium × ebenoides R.R.Scott  Asplenium rhizophyllum ×   Asplenium platyneuron1865. Asplenium ebenoides R.R.Scott1937. × Asplenosorus ebenoides Wherry | Scott's spleenwort, Dragon tail fern, Walking spleenwort | Native, No recent reports | Dutchess, Saratoga, Ulster, Washington NYFA: Z | USDA-N0 Go Botany Images Wikispecies |
| 1 2 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ||||||

##### Asplenium VI. A. trichomanes clade

Polypodiales — Aspleniaceae — Asplenium :
| Genus Asplenium | Spleenwort | N.Y. Status | Distribution | NY NPT |
| Asplenium trichomanes L. ssp. trichomanes1753. Asplenium trichomanes L. | Maidenhair spleenwort (diploid) | Native, CoC: 9, Likely secure, DEC-D | NYFA: 4 | USDA-NN Go Botany Images Wikispecies |
| Asplenium trichomanes L. ssp. quadrivalens D.E.Mey.1888. Asplenium melanolepis Colenso 1962. Asplenium trichomanes  ssp. quadrivalens  D.E.Mey. 2010. Asplenium quadrivalens (D.E. Mey.) Landolt | Maidenhair spleenwort (tetraploid) | Native, CoC: 9, Likely secure, S4?, G5, DEC-D | Onondaga (1924) NYFA: 4 | USDA-NN Go Botany Tropicos Images Wikispecies |
| Asplenium trichomanes L.nssp. lusaticum (D.E.Mey.) Lawalrèe  Asplenium trichomanes ssp. trichomanes ×   Asplenium trichomanes ssp. quadrivalens1958. Asplenium × lusaticum D.E.Mey. 1978. Asplenium trichomanes  ssp. lusaticum | Hybrid  Maidenhair spleenwort | Native, Unranked, No NY reports | | Go Botany Tropicos Images Wikispecies |
| Asplenium viride Huds.1753. Asplenium ramosum L. 1753. Asplenium trichomanes-ramosum L. 1762. Asplenium viride Huds. | Green spleenwort, Bright-green spleenwort | Native, CoC: 10, Endangered, DEC-A | NYFA: 1 | USDA-NN Go Botany Tropicos Images Wikispecies |
| 1 2 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered. ||||||

##### Asplenium XI. Pleurosorus clade

Polypodiales — Aspleniaceae — Asplenium :
| Genus Asplenium | Spleenwort | N.Y. Status | Distribution | NY NPT |
| Asplenium ruta-muraria L.1753. Asplenium ruta muraria L.1928. Asplenium cryptolepis Fernald1928. A. cryptolepisvar. ohionis Fernald1935. A. ruta-murariavar. cryptolepisC. Chr. ex Massey1939. A. ruta-murariassp. cryptolepisR.T. Clausen & Wahl | Wallrue, Wall-rue spleenwortDoradille des murailles, Mauerraute | Native, CoC: 10, Likely secure, DEC-D | NYFA: 4 | USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos Images Wikispecies |
| Asplenium montanum | Mountain spleenwort | Native, CoC: 10, Threatened, DEC-B | NYFA: 2-3 | USDA-N0 Go Botany Images Wikispecies |
| Asplenium bradleyi D.C.Eaton  Asplenium montanum ×   Asplenium platyneuron1873. Asplenium bradleyi D.C.Eaton1925. Asplenium stotleri Wherry1931. Chamaefilix bradleyi Farw. | Bradley's spleenwort | Native, CoC: 10, No recent reports, Endangered, DEC-A | NYFA: Z | USDA-N0 ITIS FNA Images Wikispecies |
| Asplenium ×‌ clermontae Syme  Asplenium ruta-muraria ×   Asplenium trichomanes1886. Asplenium clermontae Syme | Hybrid  Wallrue &  Maidenhair spleenwort | Native, Unranked, No NY reports | NYFA: D | USDA-N0 Go Botany Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (b) Threatened. ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered. ||||||

### Family Thelypteridaceae

Family Thelypteridaceae Ching ex Pic.Serm. (1970)

#### Subfamily Phegopteridoideae

The Phegopteridoideae Salino, A.R.Sm. & T.E.Almeida (2016)

##### Phegopteris

* Phegopteris connectilisTwo species of Beech ferns (Phegopteris) are found in New York.
Polypodiales — Thelypteridaceae — Phegopteridoideae — Phegopteris :
| Genus Phegopteris | Beech-fern | N.Y. Status | Distribution | NY NPT |
| Phegopteris connectilis (Michx.) Watt1753. Polypodium phegopteris L.1803. Polypodium connectile Michx.1826. Lastrea phegopteris (L.) Bory1852. Phegopteris polypodioides Fée1866. Phegopteris connectilis (Michx.) Watt1905. Dryopteris phegopteris (L.) C.Chr.1917. Thelypteris phegopteris (L.) Sloss. | Long beech fern, Northern beech fern, Narrow beech fernPhégoptère à     segments joint, Fougère du hêtre | Native, CoC: 7, Secure, DEC-D FACU Perennial, Fern | NYFA: 5 | USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos Images Wikispecies |
| Phegopteris hexagonoptera (Michx.) Fée1803. Polypodium hexagonopterum Michx.1852. Phegopteris hexagonoptera (Michx.) Fée1905. Dryopteris hexagonoptera (Michx.) C.Chr.1919. Thelypteris hexagonoptera (Michx.) Weath. | Broad beech fern, Broad beechfern, Southern beech fernPhégoptère à hexagones, Phégoptéride à hexagones, Thélyptère à hexagones | Native, Likely secure, DEC-D | NYFA: 4-5 | USDA-NN Go Botany VASCAN ARS-GRIN FNA Tropicos IPN LBJ Images Wikispecies |
| 1 2 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ||||||

#### Subfamily Thelypteridoideae
##### Thelypteris

* Thelypteris palustris marsh fernAs with the hay-scented fern (Dennstaedtia punctilobula) and bracken fern (Pteridium aquilinum), listed above, the New York fern (Thelypteris noveboracensis) is considered to be an "interfering" plant in New York forests. New York ferns produce dense stands that prevent the establishment of tree seedlings. Despite being considered weeds (or conceivably because they are), New York ferns, as well as the other two Thelypteris species, are listed as "exploitably vulnerable" by the New York State Department of Environmental Conservation.
* ↑ Heather M. Engelman & Ralph D. Nyland (2006). "Interference to Hardwood Regeneration in Northeastern North America: Assessing and Countering Ferns in Northern Hardwood Forests." Northern Journal of Applied Forestry (Society of American Foresters) 23(3):166–175.
* ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.
Polypodiales — Thelypteridaceae — Thelypteridoideae — Thelypteris :
| Genus Thelypteris | Maiden-fern | N.Y. Status | Images | Distribution | NY NPT |
| Thelypteris palustris Schottvar. pubescens (G.Lawson) Fernald1836. Dryopteris thelypterisvar. pubescens1864. Lastrea thelypterisvar. pubescens1864. T. confluensvar. pubescens1910. Dryopteris tremula Christ1929. T. palustrisvar. haleana1929. T. palustrisvar. pubescens | Marsh fern, Eastern marsh fern, Northern marsh fernThélyptère des marais     d'Amérique | Native, CoC: 4, Secure, DEC-D FACW Perennial, Herb-fern | | NYFA: 5 | USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos Images Wikispecies |
| Thelypteris noveboracensis (L.) Nieuwl.1753. Polypodium noveboracense L.1803. Nephrodium thelypterioides Michx.1827. Nephrodium novaeboracense Desv.1848. Dryopteris noveboracensis1910. Thelypteris noveboracensis1963. Parathelypteris noveboracensisChing1972. Thelypteris thelypterioides Holub | New York fernFougère de New York, Thélyptéride de New York | Native, CoC: 4, Secure, DEC-D FAC Perennial, Herb-fern | | NYFA: 5 | USDA-NN Go Botany ARS-GRIN FNA Images Wikispecies |
| Thelypteris simulata (Davenp.) Nieuwl.1894. Aspidium simulatum Davenp. 1894. Dryopteris simulata Davenp. 1904. Filix simulata (Davenp.) Farw. 1910. T. simulata (Davenp.) Nieuwl. 1976. Parathelypteris simulata Holttum | Massachusetts fern, Bog fernThélyptère simulatrice | Native, CoC: 8, Vulnerable, DEC-D FACW Perennial, Herb-fern | | NYFA: 3-4 | USDA-N Go Botany FNA Tropicos Images Wikispecies |
| Genus Thelypteris (excluded taxa) | Maiden-fern | N.Y. Status | Images | Distribution | NY NPT |
| Thelypteris palustris Schottvar. palustris1834. Thelypteris palustris Schott | Eurasian marsh fernThélyptère des marais | Introduced from  Eurasia, N.Y. excluded | | NYFA-Excluded | USDA- Images Wikispecies |
| 1 2 3 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ||||||

### Family Woodsiaceae

The Woodsiaceae Herter (1949)

#### Woodsia

* Woodsia ilvensisGenus Woodsia contains five species of cliff-ferns that are found in New York.
Polypodiales — Woodsiaceae — Woodsia :
| Genus Woodsia | Cliff-fern | N.Y. Status | Distribution | NY NPT |
| Woodsia ilvensis (L.) R.Br.    Acrostichum ilvense L. | Rusty cliff fern, Rusty woodsiaWoodsie de l'île d'Elbe | Native, Secure, DEC-D | NYFA: 5 | USDA-NN Go Botany Images Wikispecies |
| Woodsia obtusa (Spreng.) Torreyssp. obtusa | Bluntlobe cliff fern, Blunt-lobe woodsia | Native, Secure, DEC-D | NYFA: 5 | USDA-NN Go Botany Images Wikispecies |
| Woodsia alpina (Bolt.) S.Gray    Acrostichum alpinum    W. alpinavar. bellii    W. glabellavar. bellii | Alpine cliff fern, Alpine woodsia, Northern woodsiaWoodsie alpine | Native, Endangered, DEC-A | NYFA: 1 | USDA-NN Go Botany ITIS Images Wikispecies |
| Woodsia glabella R.Br. ex Richardson (1823)    W. alpinavar. glabella    W. hyperboreavar. glabella | Smooth cliff fern, Smooth woodsiaWoodsie glabre | Native, Endangered, DEC-A | NYFA: 1 | USDA-NN glabella/ Go Botany FNA Images Wikispecies |
| Woodsia oregana D.C.Eaton (1865)ssp. cathcartiana1908. W. cathcartiana B.L.Rob.    W. oreganavar. cathcartiana | Oregon cliff fern, Oregon woodsia, (tetraploid) | Native, Likely extirpated | NYFA: Z | USDA-NN Go Botany ITIS Images Wikispecies |
| 1 2 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. 1 2 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered. ||||||

### Family Athyriaceae

The Athyriaceae (lady-ferns)...

#### Athyrium

* Athyrium angustumAccording to Plants of the World, Athyrium angustum is now considered to be a variety of Athyrium asplenioides.
* ↑ Plants of the World (2021). Athyrium asplenioides var. angustum. Board of Trustees of the Royal Botanic Gardens, Kew, referencing Roskov Y. & al. (eds.) (2018). World Ferns: Checklist of Ferns and Lycophytes of the World Species 2000 & ITIS Catalogue of Life Naturalis, Leiden, the Netherlands.
* ↑ Hassler, Michael (2021). Athyrium Roth; Tent. Fl. Germ. 3: 58 (1790). World Ferns. Synonymic Checklist and Distribution of Ferns and Lycophytes of the World. Version 12.5; last update October 14th, 2021. - www.worldplants.de/ferns/.
Polypodiales — Aspleniaceae — Athyrioideae — Athyrium :
| Genus Athyrium Roth | Lady-fern | N.Y. Status | Images | Distribution | NY NPT |
| Athyrium angustum (Willd.) C.Presl1810. Aspidium angustum Willd. 1825. Athyrium angustum (Willd.) C.Presl 1860. Athyrium asplenioides  var. angustum  (Willd.) T.Moore 1864. Athyrium filix-femina  var. angustum  G.Lawson 1949. Athyrium filix-femina  ssp. angustum  R.T.Clausen | Northern lady fern ⓘ Common lady fern Narrow lady fern Subarctic lady fern Northeastern ladyfern ⓘ | Native, CoC: 5, Secure, DEC-D | | NYFA: 5 | USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos IPN Images Wikispecies |
| Athyrium asplenioides (Michx.) A.A.Eaton1803. Nephrodium asplenioides Michx. 1817. Athyrium asplenioides (Michx.) A.A.Eaton. 1923. Athyrium filix-femina  var. asplenioides  Farw, 1962. Athyrium filix-femina  ssp. asplenioides  Hultén | Southern lady fern, Lowland lady fern | Native, CoC: 7, Vulnerable, DEC-D | | NYFA: 3 | USDA-N0 Go Botany ITIS FNA Tropicos Images Wikispecies |
| Genus Athyrium (excluded taxa) | Lady-fern | N.Y. Status | Images | Distribution | NY NPT |
| Athyrium filix-femina (L.) Roth ex Mert. | Lady fern, Hybrid of  Northern lady fern &  Southern lady fern | Native, N.Y. excluded | | NYFA-Excluded | BONAP Images Wikispecies |
| Athyrium nipponicum (Mett.) Hance1865. Asplenium nipponicum Mett.1873. Athyrium nipponicum (Mett.) Hance2011. Anisocampium niponicumYea C.Liu, W.L.Chiou & M.Kato (Mett.) | Japanese painted lady fern, Japanese painted fern | Introduced from  eastern Asia, No specimens | | NYFA: 0 | Go Botany ARS-GRIN Tropicos Images Wikispecies |
| 1 2 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ↑ [ Liu & al. (2011) "Molecular phylogeny and taxonomy of the fern genus Anisocampium (Athyriaceae)". TAXON 60 (3), June 2011: 824–830.] ||||||

#### Deparia

* Deparia acrostichoides silvery spleenwort
Polypodiales — Aspleniaceae — Athyrioideae — Deparia :
| Genus Deparia | False-spleenwort | N.Y. Status | Images | Distribution | NY NPT |
| Deparia acrostichoides (Sw.) M.Kato    Athyrium thelypterioides     Diplazium acrostichoides | Silvery spleenwort, Silver false spleenwort, Silvery glade fern | Native, CoC: 8, Secure, DEC-D | | NYFA: 5 | USDA-NN Go Botany VASCAN Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ||||||

### Family Onocleaceae

Family Onocleaceae

#### Onoclea

The sensitive fern, so called due to its sensitivity to frost, is very common in areas that are wet for part of the year.
Polypodiales — Onocleaceae — Onoclea :
| Genus Onoclea | Sensitive fern | N.Y. Status | Images | Distribution | NY NPT |
| Onoclea sensibilis L.1753. Onoclea sensibilis L. | Sensitive fern, Bead fern, Polypod brake, Meadow brakeOnoclée sensible | Native, CoC: 2, Secure FACW Perennial, Fern | | NYFA: 5 | USDA-NN Go Botany ARS-GRIN ITIS FNA BONAP Images Wikispecies |

#### Matteuccia

* With the transfer of Onoclea (above) to Blechnoideae, Christenhusz & Chase (2014) recommended that Matteuccia be moved back to Onoclea.
Polypodiales — Onocleaceae — Matteuccia :
| Genus Matteuccia | Ostrich fern | N.Y. Status | Images | Distribution | NY NPT |
| Matteuccia struthiopteris (L.) Tod.var. pensylvanica (Willd.) C.V.Morton1810. Struthiopteris pensylvanica Willd.1819. Onoclea pensylvanica Sm.1945. Pteretis pensylvanica Fernald1950. Matteuccia pensylvanica Raymond1950. Matteuccia struthiopterisvar. pensylvanica C.V.Morton1961. Onoclea struthiopterisvar. pensylvanica B.Boivin | Ostrich fern, Shuttlecock fern, Fiddlehead fernFougère à l'autruche | Native, CoC: 5, Likely secure, DEC-D FACW-FAC Perennial, Herb-fern | | NYFA: 4 | USDA-NN Go Botany ARS-GRIN ITIS FNA BONAP Images Wikispecies |
| Genus Matteuccia (excluded taxa) | Ostrich fern | N.Y. Status | Images | Distribution | NY NPT |
| Matteuccia struthiopteris (L.) Tod.var. struthiopteris1753. Osmunda struthiopteris L.1866. Matteuccia struthiopteris (L.) Tod. | Ostrich fern,Fougère à l'autruche | Introduced from  Eurasia, N.Y. excluded | | NYFA-Excluded | USDA-NN Go Botany ARS-GRIN ITIS FNA BONAP Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ||||||

### Family Blechnaceae

* Family Blechnaceae

#### Subfamily Woodwardioideae
##### Anchistea

Polypodiales — Blechnaceae — Woodwardioideae — Anchistea :
| Genus Anchistea | Chainfern | N.Y. Status | Images | Distribution | NY NPT |
| Anchistea virginica (L.) C. Presl1771. Blechnum virginicum L. 1793. Woodwardia virginica (L.) Sm. 1851. Anchistea virginica (L.) C. Presl | Virginia chainfern | Native, CoC: 9, Secure, DEC-D OBL Perennial, Herb-fern | | NYFA: 5 | USDA-N Go Botany Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ||||||

##### Lorinseria

Polypodiales — Blechnaceae — Woodwardioideae — Lorinseria :
| Genus Lorinseria | Chainfern | N.Y. Status | Images | Distribution | NY NPT |
| Lorinseria areolata (L.) C. Presl1753. Acrostichum areolatum L. 1851. Lorinseria areolata (L.) C. Presl 1857. Woodwardia areolata (L.) T. Moore | Netted chainfern | Native, CoC: 10, Vulnerable, DEC-D OBL-FACW Perennial, Herb-fern | | NYFA: 3 | USDA-N Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ||||||

## Suborder Polypodiineae

Suborder Polypodiineae Dumort. (1829)

### Family Dryopteridaceae

Family Dryopteridaceae Herter. (1949)

#### Subfamily Dryopteridoideae

Subfamily Dryopteridoideae Link (1841)

##### Cyrtomium

Polypodiales — Dryopteridaceae — Dryopteridoideae — Cyrtomium :
| Genus Cyrtomium (excluded taxa) | Netvein-hollyfern | N.Y. Status | Images | Distribution | NY NPT |
| Cyrtomium falcatum (L.f.) C.Presl1782. Polypodium falcatum L.f.1783. Polypodium japonicum Houtt.1801. Aspidium falcatum (L.f.) Sw.1836. Cyrtomium falcatum (L.f.) C. Presl1891. Dryopteris falcata (L.f.) Kuntze1899. Polystichum falcatum (L.f.) Diels1947. Phanerophlebia falcata (L.f.) Copel. | Japanese holly fern, Japanese netvein hollyfern | Introduced from  Asia, N.Y. excluded | | NYFA-Excluded | USDA-X ARS-GRIN Tropicos Images Wikispecies |

##### Dryopteris

* Most Dryopteris (wood fern) species are characterized by sori located near the mid leaf vein. Dryopteris hybrids are very common.

###### Dryopteris diploid species

* Diploid wood fern species have 2n = 82 chromosmes (twice the Dryopteris base chromosome number of n = 41). D. intermedia and D. Goldiana have been shown to be part of a North American "reticulate complex", in which, along with three other Dryopteris species, they are parents to most of the allopolyploid Dryopteris species shown below.
* ↑ Emily B. Sessa, Elizabeth A. Zimmer and Thomas J. Givnish (2012). "Unraveling reticulate evolution in North American Dryopteris (Dryopteridaceae)." BioMed Central Evolutionary Biology 2012, 12:104 doi:10.1186/1471-2148-12-104
* ↑ Emily B. Sessa, Elizabeth A. Zimmer and Thomas J. Givnish (2012). "Reticulate evolution on a global scale: A nuclear phylogeny for New World Dryopteris (Dryopteridaceae)." Molecular Phylogenetics and Evolution Volume 64, Issue 3, September 2012, Pages 563–581.
Polypodiales — Dryopteridaceae — Dryopteridoideae — Dryopteris :
| Genus Dryopteris (diploid) | Wood fern | N.Y. Status | Distribution | NY NPT |
| Dryopteris marginalis (L.) A.Gray1753. Polypodium marginale L.1848. Dryopteris marginalis A.Gray | Marginal wood fern | Native, Secure, DEC-D Perennial, Herb-fern, Anemochore, 2n = 82 | NYFA: 5 | USDA-NN Go Botany FNA Images Wikispecies |
| Dryopteris intermedia (Muhl. ex Willd.) A. Gray1810. Aspidium intermedium Muhl. ex Willd. 1920. Thelypteris intermedia (Muhl. ex Willd.) House(?) 1825. Nephrodium intermedium (Muhl. ex Willd.) C. Presl 1848. Dryopteris intermedia (Muhl. ex Willd.) A. Gray 1893. D. spinulosa  var. intermedia  (Muhl. ex Willd.) Underw. 1950. D. austriaca  var. intermedia  (Muhl. ex Willd.) C.V.Morton | Evergreen woodfern, Fancy wood fern, Common wood fern | Native, Secure, DEC-D Perennial, Herb-fern, Anemochore, 2n = 82 | NYFA: 5 | USDA-NN Go Botany FNA Images Wikispecies |
| Dryopteris goldiana (Hook. ex Goldie) A.Gray1822. Aspidium goldieanum Hook. ex Goldie1848. Dryopteris goldieana A.Gray1910. Thelypteris goldieana Nieuwl.1916. Filix goldiana Farw.1931. Filix-mas goldiana Farw. | Goldie's woodfern, Goldie's shield fern, Giant woodfernDryoptère de Goldie, Dryoptéride de Goldie, Dryoptéris géant | Native, Likely secure, DEC-D Perennial, Herb-fern, Anemochore, 2n = 82 | NYFA: 4 | USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos NatureServe IPN Images Wikispecies |
| Dryopteris fragrans (L.) Schott | Fragrant cliff wood-fern, Fragrant woodfern | Native, Endangered, DEC-A Perennial, Herb-fern, Anemochore, 2n = 82 | NYFA: 1 | USDA-NN Go Botany FNA Images Wikispecies |
| 1 2 3 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered. ||||||

###### Dryopteris allopolyploid species

* These are fertile species of hybrid origin. Most are allotetraploid, derived from diploid Dryopteris parents, including two of the New York diploid species, D. intermedia and D. goldiana, shown above. The other diploid parents, D. expansa and D. ludoviciana, are North American, but not New York natives. A fifth, likely extinct, diploid parent "D. semicristata" has been hypothesized to form the well-supported model used here. The species Dryopteris clintoniana is allohexaploid, resulting from the diploid D. goldiana (above) and the allotetraploid D. Cristata, below. The parentage of the introduced non-NY-native, Dryopteris filix-mas has not been resolved, but it is probably not part of the same North American "reticulate complex" as the rest.
* ↑ Emily B. Sessa, Elizabeth A. Zimmer and Thomas J. Givnish (2012). "Unraveling reticulate evolution in North American Dryopteris (Dryopteridaceae)." BioMed Central Evolutionary Biology 2012, 12:104 doi:10.1186/1471-2148-12-104
Polypodiales — Dryopteridaceae — Dryopteridoideae — Dryopteris :
| Genus Dryopteris (allopolyploid) | Woodfern | N.Y. Status | Images | Distribution | NY NPT |
| Dryopteris cristata (L.) A.Gray  ♀ Dryopteris "semicristata" (SS) ×   ♂ Dryopteris ludoviciana (LL) → SSLL1753. Polypodium cristatum L. 1848. Dryopteris cristata A.Gray | Crested shield-fern, Crested woodfern | Native, Secure, DEC-D, 2n = 164 | | NYFA: 5 | USDA-NN Go Botany FNA Images Wikispecies |
| Dryopteris carthusiana (Vill.) H.P.Fuchs  ♀ Dryopteris "semicristata" (SS) ×   ♂ Dryopteris intermedia (II) → SSII1777. Polypodium spinulosum O.F.Muell. 1786. Polypodium carthusianum  Vill. in… 1867. Dryopteris spinulosa Watt 1959. Dryopteris carthusiana  (Vill.) H.P.Fuchs in… | Spinulose shield fern, Spinulose wood fern, Toothed wood fern, Narrow buckler fernDryoptère spinuleuse | Native, CoC: 6, Secure, DEC-D, S5, G5 FACW-FAC Perennial, Fern, 2n = 164 | | NYFA: 5 | USDA-N Go Botany VASCAN ARS-GRIN FNA Images Wikispecies |
| Dryopteris clintoniana (D.C.Eaton) Dowell  ♀ Dryopteris cristata (SSLL) ×   ♂ Dryopteris goldiana (GG) → SSLLGG1867. Aspidium cristatum  var. clintonianum  D.C.Eaton  in… 1893. Dryopteris cristata  var. clintoniana  (D.C.Eaton) Underw.  in… 1906. Dryopteris clintoniana  (D.C.Eaton) Dowell in… 1916. Filix cristata  var. clintoniana  (D.C.Eaton) Farw.  in… 1919. Thelypteris cristata  var. clintoniana  (D.C.Eaton) Weath.  in… 1922. Thelypteris clintoniana  (D.C.Eaton) House in… 1931. Filix-mas cristata  var. clintoniana  (D.C.Eaton) Farw.  in… 1960. Dryopteris poyseri  Wherry in… | Clinton's wood fernDryoptère de clinton | Native, Secure, DEC-D, 2n = 246 | | NYFA: 5 | USDA-NN Go Botany FNA Tropicos Images Wikispecies |
| Dryopteris campyloptera (Kunze) Clarkson  ♀ Dryopteris intermedia (II) ×   ♂ Dryopteris expansa (EE) → IIEE1848. Aspidium campylopterum  Kunze 1915. Dryopteris austriaca  Woyn. ex Schinz & Thell. 1915. Dryopteris spinulosa  var. americana  Fernald 1916. Dryopteris dilatata  var. americana  Benedict 1920. Thelypteris dilatata  (Hoffm.) House 1930. Dryopteris campyloptera  Clarkson | Mountain woodfern, Spreading wood fern, Eastern spreading wood fernDryoptère arquée | Native, Likely secure, DEC-D, 2n = 164 | | NYFA: 4 | USDA-N Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos Images Wikispecies |
| Dryopteris celsa (W.Palmer) Knowlt., Palmer & Pollard ex Small  ♀ Dryopteris ludoviciana (LL) ×   ♂ Dryopteris goldiana (GG) → LLGG1899. Dryopteris goldiana  ssp. celsa  W.Palmer 1900. Dryopteris celsa Knowlt. &al. 1938. Dryopteris atropalustris Small 1960. Dryopteris wherryi Crane | Log fern | Native, Endangered, DEC-A, 2n = 164 | | NYFA: 1 | USDA-N ARS-GRIN ITIS FNA Tropicos Images Wikispecies |
| Genus Dryopteris (excluded) | Woodfern | N.Y. Status | Images | Distribution | NY NPT |
| Dryopteris filix-mas (L.) Schott1753. Polypodium filix-mas L.1801. Nephrodium filix-mas Rich.1834. Dryopteris filix-mas Schott | Male fern, Male wood-fern | N. America native, N.Y. excluded, 2n = 164 | | NYFA-Excluded | USDA-NN Go Botany ARS-GRIN FNA Tropicos NatureServe BONAP Images Wikispecies |
| 1 2 3 4 New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (a) Endangered. ||||||

###### Dryopteris named hybrids

Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris subsect. named hybrids :
| Genus Dryopteris | Woodfern | N.Y. Status | Distribution | NY NPT |
| Dryopteris × boottii (Tuck.) Underw.  Dryopteris cristata (SSLL) ×   Dryopteris intermedia (II) → ILS 1843. Aspidium boottii Tuck.1893. Dryopteris boottii (Tuck.) Underw.1910. Thelypteris boottii (Tuck.) Nieuwl. | Boott's wood fern, Glandular swamp fern | Native, Vulnerable | NYFA: 3-5 | USDA-NN Go Botany ITIS Images Wikispecies |
| Dryopteris × triploidea Wherry  Dryopteris carthusiana (IISS) ×   Dryopteris intermedia (II) → IIS 1930. D. spinulosavar. fructuosa Trudell1950. D. austriacavar. fructuosa C.V.Morton1958. D. intermediavar. fructuosa Wherry1960. Dryopteris × triploidea Wherry | Triploid woodfern | Native, Vulnerable | NYFA: 3 | USDA-NN ITIS FNA Images Wikispecies |
| Dryopteris × uliginosa (A.Braun ex Döll) Kuntze ex Druce  Dryopteris carthusiana (IISS) ×   Dryopteris cristata (SSLL) → ILSS 1843. Aspidium spinulosum  var. uliginosum  A.Braun ex Döll  in… 1850. Lastrea spinulosa  var. uliginosa  (A.Braun ex Döll) H.C.Watson & Dennes  in… 1872. Lastrea cristata  ssp. uliginosa  (A.Braun ex Döll) Hayw.  in… 1908. Dryopteris × uliginosa  (A.Braun ex Döll) Druce in… 1884. Aspidium cristatum  ssp. uliginosum  (A.Braun ex Döll) Nyman  in… | Spinulose crested wood fern, Braun's wood fern | Native, Vulnerable | NYFA: 3 | USDA-NN Images Wikispecies |
| Dryopteris × slossoniae Wherry ex Lellinger  Dryopteris cristata (SSLL) ×   Dryopteris marginalis (MM) → LMS 1894. Dryopteris cristata × marginalis  Davenport 1894. Aspidium cristatum × marginale  Davenport 1904. Nephrodium slossonae  Hahne 1942. Dryopteris × slossonae  Wherry | Crested marginal fern, Miss Slosson's wood fern | Native, Threatened | NYFA: 1-3 | USDA-NN Images Wikispecies |
| Dryopteris × pittsfordensis Slossen  Dryopteris carthusiana (IISS) ×   Dryopteris marginalis (MM) → IMS1904. Dryopteris pittsfordensis Sloss.1904. Nephrodium pittsfordensis Woolson | Pittsford's wood fern | Native, Endangered | NYFA: 1 | USDA-NN ITIS Tropicos IPN Images Wikispecies |
| Dryopteris × burgessii Boivin  Dryopteris clintoniana (SSLLGG) ×   Dryopteris marginalis (MM) → GLMS | Burgess' shield fern | Native, No recent reports | NYFA: Z? | USDA-NN Images Wikispecies |
| Dryopteris × benedictii (Farw.) Wherry  Dryopteris carthusiana (IISS) ×   Dryopteris clintoniana (SSLLGG) → GILSS | Benedict's wood fern | Native, No recent reports | NYFA: Z? | USDA-NN Images Wikispecies |
| Dryopteris × dowellii (Farw.) Wherry  Dryopteris clintoniana (SSLLGG) ×   Dryopteris intermedia (II) → GILS | Dowell's wood fern | Native, No recent reports | NYFA: Z | USDA-NN Images Wikispecies |
| Dryopteris × mickelii  Dryopteris clintoniana (SSLLGG) ×   Dryopteris goldiana (GG) → GGLS | Wood fern | Native, Likely extirpated | NYFA: Z | USDA-N Images Wikispecies |
| Dryopteris × neowherryi W.H.Wagner  Dryopteris goldiana (GG) ×   Dryopteris marginalis (MM) → GM | Wood fern | Native, No recent reports | NYFA: Z | USDA-NN Images Wikispecies |
| Dryopteris × separabilis Small (pro sp.)  Dryopteris celsa (LLGG) ×   Dryopteris intermedia (II) → GIL | Glandular log fern | Native, No recent reports | NYFA: Z | USDA-N0 Images Wikispecies |
| Dryopteris × correllii W.H.Wagner  Dryopteris carthusiana (IISS) ×   Dryopteris goldiana (GG) → GIS | Correll's wood fern, Poyser's wood fern | Native, No recent reports | NYFA: ? | USDA-N0 Images Wikispecies |
| Dryopteris × leedsii Wherry  Dryopteris celsa (LLGG) ×   Dryopteris marginalis (MM) → GLM | Leeds' wood fern | Native, No recent reports | NYFA: Z | USDA-N0 Go Botany ITIS Images Wikispecies |

###### Dryopteris unnamed hybrids

Polypodiales — Polypodiaceae — Dryopteridoideae — Dryopteris subsect. unnamed hybrids :
| Genus Dryopteris | Woodfern | N.Y. Status | Distribution | NY NPT |
| Dryopteris filix-mas × marginalis | Marginal male fern | Native, Threatened | NYFA: 1-3 | Images Wikispecies |
| Dryopteris celsa × goldiana  D. celsa (LLGG) ×   D. goldiana (GG) → GGL | Hybrid wood-fern | Native, No recent reports | NYFA: Z | Images Wikispecies |
| Dryopteris intermedia × marginalis  D. intermedia (II) ×   D. marginalis (MM) → IM | Glandular marginal fern | Native, No recent reports | NYFA: Z | Images Wikispecies |
| Dryopteris celsa × cristata  D. celsa (LLGG) ×   D. cristata (SSLL) → GLLS | Hybrid wood-fern | Native, No recent reports | NYFA: Z | Images Wikispecies |
| Dryopteris carthusiana × celsa  D. carthusiana (IISS) ×   D. celsa (LLGG) → IGLS | Spinulose wood fern | Native, No recent reports | NYFA: Z | Images Wikispecies |
| Dryopteris goldiana × intermedia  D. goldiana (GG) ×   D. intermedia (II) → GI | Wood fern | Native, No recent reports | NYFA: Z | Images Wikispecies |
| Dryopteris cristata × goldiana  D. cristata (SSLL) ×   D. goldiana (GG) → GLS | Benedict's wood fern | Native, Unranked | NYFA: U | Images Wikispecies |

##### Polystichum

* Polystichum acrostichoides Christmas fernPolystichum Roth (1799) goes by the English vernacular names Christmas fern, holy fern, and sword fern.
Polypodiales — Polypodiaceae — Dryopteridoideae — Polystichum :
| Genus Polystichum | Hollyfern | N.Y. Status | Distribution | NY NPT |
| Polystichum acrostichoides (Michx.) Schott1803. Nephrodium acrostichoides Michx.1834. Polystichum acrostichoides(Michx.) Schott | Christmas fern, Dagger fernPolystic faux-acrostiche | Native, Secure | NYFA: 5 | USDA-NN Go Botany ARS-GRIN FNA Tropicos BONAP Images Wikispecies |
| Polystichum braunii (Spenner) Fée1825. Aspidium braunii Spenner1852. Polystichum braunii Fée | Braun's holly-fern | Native, Likely secure | NYFA: 4 | USDA-NN Go Botany Tropicos Images Wikispecies |
| Polystichum × potteri Barrington[ P. acrostichoides × P. braunii ]1986. Polystichum × potteri Barrington | Hybrid Christmas fern   & Braun's holly-fern | Native, Unranked | NYFA: U | USDA-NN Tropicos Images Wikispecies |
| Polystichum lonchitis (L.) Roth1753. Polypodium lonchitis L.1800. Polystichum lonchitis Roth1801. Aspidium lonchitis Sw.1924. Polystichum mohrioidesvar. lemmonii | Northern holly-fernPolystic faux-lonchitis | Native, Endangered | NYFA: Z | USDA-NN FNA Images Wikispecies |
| Polystichum × hagenahii Cody  Polystichum acrostichoides ×   Polystichum lonchitis1924. P. lonchitisfo. marginale McColl1966. P. marginale (McColl) Cody1968. P. × hagenahii Cody | Hagenah's holly fern, Hagenah's polystichum, Hybrid Christmas fern   & northern holly-fernPolystic de Hagenah | Native, Unranked, Questionable | | USDA-NN ITIS FNA Images Wikispecies |
| ↑ Flora of North America states that the rare hybrid Polystichum × hagenahii has only been found growing with both parents in Ontario. NYFA does not list it, but USDA-PLANTS does show it in NY as well as PA, listing the NY source: Lellinger, D.B. (1985) A field manual of the ferns and fern-allies of the U.S. and Canada, Smithsonian Institution, Washington, DC. ||||||

### Family Polypodiaceae
#### Subfamily Polypodioideae
##### Polypodium

* Polypodium virginianum
Polypodiales — Polypodiaceae — Polypodioideae — Polypodium :
| Genus Polypodium | Polypody | N.Y. Status | Distribution | NY NPT |
| Polypodium virginianum L.1753. Polypodium virginianum L.1818. P. vulgarevar. virginianum Eaton1840. P. vulgarevar. americanum Hook.1962. P. vulgaressp. virginianum Hultén1977. P. vinlandicum Á. Löve & D. Löve | Rock polypody, Rock-cap fern, Common polypodyTripes-de-roches, Polypode de virginie | Native, Secure, DEC-D | NYFA: 5 | USDA-NN Go Botany FNA Images Wikispecies |
| Polypodium appalachianum Haufler & Windham1900. P. vulgarevar. oreophilum Maxon1902. P. vulgarevar. acuminatum Gilbert1922. P. virginianumfo. acuminatum Fernald1991. P. appalachianum Haufler & Windham | Appalachian rockcap fern, Appalachian polypody, Appalachian rock polypodyPolypode des Appalaches | Native, Secure | NYFA: 5 | USDA-NN Go Botany ARS-GRIN ITIS FNA Tropicos Images Wikispecies |
| Polypodium × incognitum Cusick  Polypodium appalachianum ×   Polypodium virginianum2002. Polypodium × incognitum Cusick | Disguised rock polypody ⓘ Disguised polypody ⓘ Hybrid polypody ⓘ | Native, Unranked | Clinton (1995), Cortland (1891) NYFA: SNR | Go Botany Images Wikispecies |
| ↑ New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable. ||||||

##### Pleopeltis

* Pleopeltis polypodioides resurrection fernPleopeltis polypodioides (resurrection fern) has been excluded from the New York Flora Atlas. The most northern US iNaturalist observations of Pleopeltis polypodioides are in Key Largo in southern Florida. However Pleopeltis michauxiana, which has recently been split from P. polypodioides, has a coastal range as far north as southern Delaware.
* ↑ USDA-NRCS-PLANTS includes New York in the range of Pleopeltis polypodioides ssp. michauxiana, but the reference for this (House, H.D. 1924. Annotated list of the ferns and flowering plants of New York State. New York State Museum Bulletin No. 254) mentions only that the "gray polypody, or tree fern (Polypodium polypodioides) (L.) Watt, has been found on driftwoud on the Staten Island shore, although its natural range is from Virginia southward."
* ↑ Pleopeltis polypodioides (Tropical Resurrection Fern) iNaturalist species page.
Polypodiales — Polypodiaceae — Polypodioideae — Pleopeltis :
| Genus Pleopeltis (excluded taxa) | Scaly-polypody | N.Y. Status | Images | Distribution | NY NPT |
| Pleopeltis polypodioides (L.) E.G.Andrews & Windhamvar. michauxiana (Weath.) E.G.Andrews & Windham1939. Polypodium polypodioides  var. michauxianum  Weath. 1993. Pleopeltis polypodioides  var. michauxiana   in… 2014. Pleopeltis michauxiana (Weath.) Hickey & Sprunt | Resurrection fern, Gray's polypody | N. America native, N.Y. excluded | | NYFA-Excluded | USDA-N0 ITIS FNA Tropicos Images Wikispecies |