# Flora of New York/Amygdaloideae 2

| ← Rosales : Rosaceae : Amygdaloideae 1 | Flora of New York — Rosales : Rosaceae : Amygdaloideae 2 : Maleae | Flora of New York — Rosales : Rosaceae : Amygdaloideae 2 : Maleae | Flora of New York — Rosales : Rosaceae : Amygdaloideae 2 : Maleae | Flora of New York — Rosales : Rosaceae : Amygdaloideae 2 : Maleae | → Rosales : Rhamnaceae … Urticaceae |
| ← Rosales : Rosaceae : Amygdaloideae 1 | Table of contents | Genus index | Protected species index | Invasive species index | → Rosales : Rhamnaceae … Urticaceae |

##### Family Rosaceae (cont'd)

* The grouping of the Rosaceae (rose family) taxa used here is based on the Angiosperm Phylogeny Website, Eriksson (2003), Longhi (2014), Sun et al. (2018). X. Chen et al. (2020).
* The subfamilies Rosoideae and Amygdaloideae are present in New York.
* Rosaceae
* Dryadoideae
* Cercocarpus
* Dryas
* Rosoideae
* Ulmarieae
* Filipendula
* Rosodeae
* Colurieae
* Geum (avens)
* Waldsteinia (barren strawberry)
* Rubeae
* Rubus (brambles)
* Roseae
* Rosa (roses)
* Potentilleae
* Potentillineae
* Argentina (siverweed)
* Potentilla (cinquefoil)
* Fragariinae
* Fragaria (strawberries)
* Dasiphora (shrubby cinquefoil)
* Drymocallis (tall cinquefoil)
* Sibbaldia (three-toothed cinquefoil)
* Comarum (marsh cinquefoil)
* Agrimonieae
* Agrimoniinae
* Agrimonia (agrimony)
* Sanguisorbineae
* Poteridium (American burnet)
* Poterium (salad burnet)
* Sanguisorba (burnet)
* Amygdaloideae
* Niellieae
* Physocarpus
* Spiraeeae
* Spiraea
* Lyonothamneae
* Lyonothamnus floribundus
* Amygdaleae
* Prunus
* Sorbarieae
* Sorbaria (false spiraea)
* Osmaronieae
* Exochorda (pearlbush)
* Kerrieae
* Rhodotypos (jetbead)
* Kerria (Japanese rose)
* Gillenieae
* Gillenia
* Maleae
* Lindleyinae
* Kageneckia
* Vauquelinia
* Malinae
* Crataegus (hawthorn)
* Amelanchier (shadbush)
* Cydonia (quince)
* Pyracantha (firethorn)
* Sorbus (mountain ash)
* Pyrus (pear)
* Photinia
* Cotoneaster
* Chaenomeles (quince)
* Malus (apple)
* Aronia (chokeberry)
* ↑ Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 14, July 2017 (and more or less continuously updated since). Accessed 4 Jan. 2024
* ↑ Torsten Eriksson, Malin S. Hibbs, Anne D. Yoder, Charles F. Delwiche, and Michael J. Donoghue (2003). "The phylogeny of Rosoideae (Rocaceae) based on sequences of the internal transcribed spacers (ITS) of Nuclear Ribosomal DNA and the TRN/F region of Chloroplast DNA." Int. J. Plant Sci. 164(2): 197–211.
* ↑ Sara Longhi, Lara Giongo, Matteo Buti, Nada Surbanovski, Roberto Viola, Riccardo Velasco, Judson A. Ward & Daniel J. Sargent (2014). "Molecular genetics and genomics of the Rosoideae: state of the art and future perspectives." Horticulture Research 1, Article number: 1.
* ↑ Fengjie Sun, Jiahui Sun, Shuo Shi, Jinlu Li, Jing Yu, Ling Wang, Xueying Yang, Ling Guo, Shiliang Zhou 2018. Phylogeny of Maleae (Rosaceae) Based on Multiple Chloroplast Regions: Implications to Genera Circumscription. BioMed Research International. Hindawi.
* ↑ Chen, Xun, Li, J., Cheng, T., Zhang, W., Liu, Y., Wu, P., Yang, X., Wang, L., & Zhou, S. 2020. Molecular systematics of Rosoideae (Rosaceae). Plant Syst. Evol. 306:9.

###### Subfamily Amygdaloideae (cont'd)
## Tribe Gillenieae

* The tribe Gillenieae, with a base chromosome number of x = 9, shares a clade with sister tribe Maleae but does not have the high (x = 17) chromosome numbers of Maleae.
* ↑ Hodel Richard G. J., Zimmer Elizabeth A., Liu Bin-Bin, Wen Jun (2022). Synthesis of Nuclear and Chloroplast Data Combined With Network Analyses Supports the Polyploid Origin of the Apple Tribe and the Hybrid Origin of the Maleae—Gillenieae Clade. Frontiers in Plant Science.

### Gillenia

* Gillenia trifoliata (L.) Moench Bowman’s root, Indian physicThe genus name Gillenia was published in the early 1800s, but was changed to Porteranthus by Britton in 1893 to avoid confusion with a similar genus name. Recent interpretations of naming rules have caused the accepted name to be switched back to Gillenia.
* Sun et al. (2018) recommended moving Gillenia into Maleae for phylogenetic consistency.
* ↑ Fengjie Sun, Jiahui Sun, Shuo Shi, Jinlu Li, Jing Yu, Ling Wang, Xueying Yang, Ling Guo, Shiliang Zhou 2018. Phylogeny of Maleae (Rosaceae) Based on Multiple Chloroplast Regions: Implications to Genera Circumscription. BioMed Research International. Hindawi.
Rosales — Rosaceae — Amigdaloideae — Gillenieae — Gillenia :
| Genus Gillenia | Indian physic | N.Y. Status | Images | Distribution | NY NPT |
| Gillenia trifoliata (L.) Moench1753. Spiraea trifoliata L.1802. Gillenia trifoliata Moench1893. Porteranthus trifoliatus Britton | Bowman's root, Mountain Indian-physic, Fawn's breathGillénie trifoliée | Native, CoC: 8, Endangered, S1 Perennial, Herb | | NYFA: 4? | USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNA Images Wikispecies |
| Gillenia stipulata (Muhl. ex Willd.) Nutt.1809. Spiraea stipulata Muhl. ex Willd.1817. Gillenia stipulata Nutt.1893. Porteranthus stipulatus Britton | American ipecac, Indian physic | Native, Likely extirpated | | NYFA: Z | USDA-N0 ARS-GRIN ITIS Images Wikispecies |

## Tribe Maleae

The Maleae comprise the serviceberries, chokeberries, hawthorns, apples, pears, and quinces.

### Amelanchier

* Amelanchier arboreaThe species of Amelanchier are typically shrubs or small trees that go by the common names serviceberry, shadbush, or Juneberry.
* Amelanchier humilis Wiegand is here merged into Amelanchier alnifolia (Nutt.) Nutt. var. alnifolia (the
* older name) following morphological and genetic data from Burgess et al.(2015).
* For the ploidy levels shown here, 2n = 2x indicates diploidy, 3n = 3x triploidy, and 2n = 4x tetraploidy. As with most members of Maleae, Amelanchier species have a base chromosome number of x=17.
* ↑ L. Brouillet, F. Coursol, S.J. Meades, M. Favreau, M. Anions, P. Bélisle & P. Desmet. 2010+. "Amelanchier Medikus." VASCAN, the Database of Vascular Plants of Canada. http://data.canadensys.net/vascan/ (consulted on 2017-05-14)
* ↑ Kevin R. Cushman, Michael B. Burgess, Eric T. Doucette, Gretchen A. Nelson and Christopher S. Campbell (2017). "Species Delimitation in Tetraploid, Apomictic Amelanchier (Rosaceae)." Systematic Botany Vol. 42, No. 2 (April-June 2017), pp. 234-256.
* ↑ Michael B. Burgess, Kevin R. Cushman, Eric T. Doucette, Christopher S. Campbell (2015). "Understanding diploid diversity: A first step in unraveling polyploid, apomictic complexity in Amelanchier." American Journal of Botany, Vol. 102, No. 12 (December 2015), pp. 2041-2057.
* ↑ Bin-Bin Liu, Christopher S. Campbell, De-Yuan Hong, Jun Wen (2020). "Phylogenetic relationships and chloroplast capture in the Amelanchier-Malacomeles-Peraphyllum clade." Molecular Phylogenetics and Evolution, Volume 147, June 2020, 106784.
Rosales — Rosaceae — Amygdaloideae — Maleae — Malinae — Amelanchier :
| Genus Amelanchier Medik. | Serviceberry | N.Y. Status | Distribution | NY NPT |
| Amelanchier arborea (F.Michx.) Fernald1787. Pyrus botryapium  Wangenh. 1788. Mespilus canadensis  Walter (sensu auct.) 1803. Mespilus canadensis  var. cordata  Michx. 1810. Mespilus arborea  F.Michx. 1818. Aronia arborea  Barton 1818. Malus microcarpa  Raf. 1838. Pyrus wangenheimiana  Tausch 1908. Amelanchier alabamensis  Britton 1914. Amelancus canadensis  Medik. ex Vollm. 1941. Amelanchier arborea  (F.Michx.) Fernald 1946. Amelanchier arborea  var. alabamensis  (Britton) G.N.Jones 1946. Amelanchier arborea  var. arborea | Downy serviceberry, Downy shadbush, Common serviceberry, Apple shadbush, Shadblow, JuneberryAmélanchier arborescent, Amélanchier arbre | Native, CoC: 4, Secure, S5, G5 FAC-FACU Perennial, Tree-shrub, Sun - shade, 2n = 2x, 4x | NYFA: 5 GBIF | USDA-NN COL WFO POWO Go Botany VASCAN ARS-GRIN ITIS FNAFNA Tropicos LBJ Images Wikispecies |
| Amelanchier laevis Wiegand1824. Pyrus botryapium  Bigelow (sensu auct.) 1840. Amelanchier canadensis  Torr. & A.Gray (sensu auct.) 1846. Amelanchier botryapium  G.B.Emers. 1912. Amelanchier laevis  Wiegand 1912. Amelanchier laevis  f. nitida  Wiegand 1918. Amelanchier laevis  var. cordifolia  Ashe 1921. Amelanchier laevis  var. nitida  (Wiegand) Fernald 1949. Amelanchier laevis  f. villosa  Ter Pelkwijk 1964. Amelanchier arborea  var. laevis  (Wiegand) H.E.Ahles 1966. Amelanchier arborea  var. cordifolia  (Ashe) B.Boivin 1975. Amelanchier arborea  ssp. laevis  S.M.McKay ex P.Landry | Allegheny serviceberry, Smooth shadbush, Smooth serviceberryAmélanchier glabre, Petites poires | Native, CoC: 4, Secure Perennial, Tree-shrub, 2n = 2x, 4x | NYFA: 5 | USDA-N POWO Go Botany VASCAN ARS-GRIN FNAFNA Tropicos Images Wikispecies |
| Amelanchier canadensis (L.) Medik. var. canadensis1753. Mespilus canadensis  L. 1790. Mespilus amelanchier  var. canadensis  (L.) Castigl. 1793. Amelanchier canadensis (L.) Medik. 1796. Crataegus amoena  Salisb. 1806. Aronia botryapium  (L.f.) Pers. 1833. Pyrus canadensis  (L.) Billb. 1847. Amelanchier oblongifolia  (Torr. & A. Gray) M.Roem. 1930. Aronia canadensis  (L.) Farw. | Coastal shadbush, Eastern shadbush, Eastern serviceberry, Canadian serviceberry, Canada serviceberry, Oblongleaf serviceberry, Shadblow, Swamp shadbushAmélanchier du Canada, Amélanchier canadien | Native, CoC: 7, Secure FAC-FACU Perennial, Tree-shrub, 2n = 2x, 4x | NYFA: 5 | USDA-N POWO Go Botany VASCAN ARS-GRIN FNA Tropicos BONAP Images Wikispecies |
| Amelanchier intermedia Spach  Amelanchier canadensis ×   Amelanchier laevis1834. Amelanchier intermedia Spach in Hist. Nat. Vég. 2:85 1919. Amelanchier canadensis var. intermedia (Spach) Ashe in Bull. Torrey Bot. Club 46:221 | Intermediate shadbush, Intermediate serviceberry, Purple serviceberry, Swamp juneberryAmélanchier intermédiaire | Native, CoC: 7, Secure FACW Perennial, Tree, 2n = 4x | NYFA: 5 NYFA: 20 counties GBIF No RG opbservations | USDA-N COL POWO Go Botany VASCAN ARS-GRIN ITIS FNAFNA Tropicos Images Wikispecies |
| Amelanchier sanguinea (Pursh) DC.1803. Mespilus canadensis  var. rotundifolia  Michx. 1814. Pyrus sanguinea  Pursh 1818. Aronia sanguinea  (Pursh) Nutt. 1825. Amelanchier sanguinea (Pursh) DC. 1840. Amelanchier canadensis  var. rotundifolia  (Michx.) Torr.& Gray 1920. Amelanchier huronensis  Wiegand (orth. var.) 1975. Amelanchier sanguinea  var. arguta  (Greene) P.Landry | Roundleaf serviceberry, Round-leaved shadbush, Red-twigged shadbushAmélanchier sanguin, Petites poires | Native, CoC: 7, Secure, 2n = 2x, 3x, 4x | NYFA: 5 | USDA-NN POWO Go Botany ARS-GRIN FNA Images Wikispecies |
| Amelanchier spicata (Lam.) K.Koch.1783. Crataegus spicata Lam. in Encycl. 1:84 1912. Amelanchier stolonifera Wiegand in Rhodora 14: 144 1922. Amelanchier stolonifera var. lucida Fernald 1948. Amelanchier lucida Fernald 1969. Amelanchier spicata K.Koch. | Dwarf shadbush, Running serviceberry, Thicket shadbushAmélanchier en épis | Native, CoC: 7, Secure FACU Perennial, Shrub, 2n = 3x, 4x | NYFA: 5 | USDA-NN Go Botany ARS-GRIN ITIS FNA Images Wikispecies |
| Amelanchier alnifolia (Nutt.) Nutt. ex M. Roem.1818. Aronia alnifolia Nutt. in Gen. N. Amer. Pl. 1:306 1821. Pyrus alnifolia (Nutt.) Lindl. in Trans. Linn. Soc. London 13:98 1840. Amelanchier canadensis var. alnifolia (Nutt.) Torr. & A.Gray in Fl. N.Amer. 1:473 1843. Amelanchier diversifolia var. alnifolia (Nutt.) Torr. in J.C.Frémont, Rep. Exped. Rocky Mts. 1847. Amelanchier alnifolia (Nutt.) Nutt. ex M. Roem. in Fam. Nat. Syn. Monogr. 3:147 1891. Amelanchier glabra Greene in Fl. Francisc.: 52 1906. Amelanchier siskiyouensis C.K.Schneid. in Ill. Handb. Laubholzk. 1:735 1912. Amelanchier humilis Wiegand in Rhodora 14: 141 1913. Amelanchier macrocarpa Lunell in Amer. Midl. Naturalist 3:143 1918. Amelanchier leptodendron Lunell in Amer. Midl. Naturalist 5:237 1939. Amelanchier humilis var. typica E.L.Nielsen in Amer. Midl. Naturalist 22:173, not validly publ. 1939. Amelanchier humilis var. compacta E.L.Nielsen in Amer. Midl. Naturalist 22:174 1939. Amelanchier humilis var. campestris E.L.Nielsen in Amer. Midl. Naturalist 22:176 1939. Amelanchier humilis var. exserrata E.L.Nielsen in Amer. Midl. Naturalist 22:177 1939. Amelanchier mucronata E.L.Nielsen in Amer. Midl. Naturalist 22:178 1975. Amelanchier sanguinea var. alnifolia (Nutt.) P.Landry in Bull. Soc. Bot. France 122:249 1976. Amelanchier alnifolia var. compacta (E.L.Nielsen) McKay in Ontario Field Biol. 30:55 | Low shadbush, Low Juneberry, Low shadblow, Low serviceberry, Running serviceberryAmélanchier bas, Amélanchier nain | Native, CoC: 7, Likely secure Perennial, Shrub, Sun - shade, 2n = 2x, 3x, 4x | NYFA: 18 counties iNat: 4 counties GBIF: present in NY | USDA-N0 COL WFO POWO FSUS Go Botany VASCAN ARS-GRIN ITIS FNAFNA Tropicos BONAP LBJ Images Wikispecies |
| Amelanchier amabilis Wiegand1912. Amelanchier sanguinea  f. grandiflora  Wiegand  ... 1914. Amelanchier sanguinea  var. grandiflora  (Wiegand) Rehder in… 1920. Amelanchier grandiflora  (Wiegand) Wiegand in… 1921. Amelanchier amabilis  Wiegand in… | Lovely shadbush ⓘ Beautiful Serviceberry ⓘ Amélanchier gracieux ⓘ | Native, CoC: 8, Likely secure, S4? Perennial, Tree, shrub, 2n = 4x | NYFA: 4 iNat GBIF | USDA-NN COL WFO POWO FSUS Go Botany VASCAN ITIS FNAFNA Tropicos BONAP Images Wikispecies |
| Amelanchier bartramiana (Tausch) M.Roem.    Amelanchier oligocarpa (Michx.) M.Roem.1847. Amelanchier bartramiana (Tausch) M.Roem. | Bartram shadbush, Mountain shadbush, Oblongfruit serviceberryAmélanchier de Bartram | Native, CoC: 9, Likely secure, 2n = 2x, 3x, 4x | NYFA: 4 | USDA-NN Go Botany ARS-GRIN Images Wikispecies |
| Amelanchier nantucketensis E.P.Bicknell1908. Amelanchier oblongifoliavar. micropetala1911. Amelanchier nantucketense E.P.Bicknell1919. Amelanchier micropetala (B.L.Rob.) Ashe | Nantucket serviceberry, Nantucket shadbush | Native, CoC: 10, Endangered FACU Perennial, Shrub, 2n = 4x | NYFA: 1 | USDA-N0 Go Botany FNA Tropicos BONAP Images Wikispecies |

### Amelanchier hybrids

Rosales — Rosaceae — Amygdaloideae — Maleae — Malinae — Amelanchier :
| Genus Amelanchier Medik. | Serviceberry | N.Y. Status | Distribution | NY NPT |
| Amelanchier × lamarckii F.G.Schroed.  Amelanchier arborea ×   Amelanchier laevis1869. Amelanchier × canadensis  K.Koch (sensu auct.) 1906. Amelanchier canadensis  var. scharnkeana  Asch. & Graebn. 1920. Amelanchier × grandiflora Rehder (nom. illeg.) 1920. Amelanchier × grandiflora  f. rubescens  Rehder 1958. Amelanchier × confusa  Dandy 1968. Amelanchier × lamarckii  F.G.Schroed. 1968. Amelanchier lamarckii  var. scharnkeana  (Asch. & Graebn.) F.G.Schroed. 1976. Amelanchier arborea  ssp. grandiflora  P.Landry | Juneberry ⓘ Showy Mespilus ⓘ Showy Mespil ⓘ Large-flowered shadbush ⓘ | Native from  NY & VT, No specimens | NYFA: NYFA: 1 specimen iNat Kings (2023) | WFO POWO Images Wikispecies |
| Amelanchier × neglecta Eggl. ex K.R.Cushman et al.  Amelanchier bartramiana ×   Amelanchier laevis1916. Amelanchier canadensis  var. pauciflora  Farw. 2017. Amelanchier × neglecta Eggl. ex K.R.Cushman et al. | Hidden shadbush ⓘ Hybrid mountain & smooth shadbush ⓘ | Native, No specimens | NYFA: N NYFA: Franklin, Madison iNat St. Lawrence (?) | POWO Images Wikispecies |
| Amelanchier amabilis × A. arborea  Amelanchier amabilis ×   Amelanchier arborea | Hybrid lovely & downy shadbush ⓘ | Native, No specimens | NYFA: NYFA: Cayuga (2006) | Images Wikispecies |
| Amelanchier alnifolia × A. amabilis  Amelanchier alnifolia ×   Amelanchier amabilis | Hybrid low & lovely shadbush ⓘ | Native, No specimens, SNA, GNA | NYFA: U NYFA: Cayuga (2006) | Images Wikispecies |

### Aronia

* Aronia melanocarpaThe genus Aronia was included in Photinia by Robertson et al. (1991), but Kalkman (2004) determined that the name Aronia should have priority. Based on the molecular phylogeny, Potter et al. (2007) then found that the two genera should be classified as distinct.
Rosales — Rosaceae — Amygdaloideae — Maleae — Malinae — Aronia :
| Genus Aronia | Chokeberry | N.Y. Status | Distribution | NY NPT |
| Aronia melanocarpa (Michx.) Elliott1793. Hahnia arbutifolia  var. nigra  Medik. 1799. Pyrus arbutifolia  var. nigra  Willd. 1803. Mespilus arbutifolia  var. melanocarpa  Michx. 1809. Pyrus melanocarpa  (Michx.) Willd. 1821. Aronia melanocarpa (Michx.) Elliott 1841. Sorbus melanocarpa  Heynh. 1890. Pyrus nigra  Sarg. 1893. Aronia nigra  Koehne 1991. Photinia melanocarpa  (Michx.) K.R.Robertson & J.B.Phipps | Black chokeberryAronie à fruits noirs, Aronia gueules-noires, Aronia noir, Gueules noires | Native, CoC: 7, Secure FAC Perennial, Shrub, Sun - part shade | NYFA: 5 | USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNAFNA Tropicos BONAP IPN LBJ Images Wikispecies |
| Aronia arbutifolia (L.) Pers.1753. Mespilus arbutifolia L. 1782. Pyrus arbutifolia (L.) L.f. 1783. Crataegus pyrifolia Lam. 1806. Aronia arbutifolia (L.) Pers. 1841. Sorbus arbutifolia (L.) Heynh. 1991. Photinia pyrifolia (Lam.) K.R.Robertson & J.B.Phipps | Red chokeberryAronie à feuilles d'arbousier, Aronia à feuilles d'arbousier | Native, CoC: 7, Likely secure FACW Perennial, Shrub, Sun | NYFA: 4 | USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNA LBJ Images Wikispecies |
| Aronia × prunifolia (Marshall) Rehder (pro sp.)  Aronia arbutifolia ×   Aronia melanocarpa1785. Mespilus × prunifolia  Marshall 1790. Mespilus amelanchier  var. prunifolia  (Marshall) Castigl. 1790. Crataegus × prunifolia  (Marshall) Baumg. 1826. Pyrus floribunda  Lindl. 1826. Aronia × floribunda  (Lindl.) Sweet 1901. Aronia atropurpurea  Britton 1938. Aronia × prunifolia  (Marshall) Rehder 1991. Photinia × floribunda  (Lindl.) K.R.Robertson & J.B.Phipps 2018. Pyrus × hippomenes  M.F.Fay & Christenh. | Purple chokeberryAronie à feuilles de prunier, Arone noire | Native, CoC: 7, Likely secure FACW Perennial, Shrub | NYFA: 4 | USDA-NN POWO Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos BONAP LBJ Images Wikispecies |

### Pourthiaea

* Pourthiaea villosa
Rosales — Rosaceae — Amygdaloideae — Maleae — Malinae — Pourthiaea :
| Genus Pourthiaea Decne. | Chokeberry | N.Y. Status | Images | Distribution | NY NPT |
| Pourthiaea villosa (Thunb.) Decne.1784. Crataegus villosa  Thunb. 1784. Crataegus laevis  Thunb. 1798. Mespilus villosa  (Thunb.) Poir. 1825. Photinia villosa  (Thunb.) DC. 1873. Photinia arguta  var. villosa  (Thunb.) Wenz. 1874. Pourthiaea villosa  (Thunb.) Decne. 1899. Pourthiaea variabilis  Palib. 1915. Crataegus cavaleriei  H.Lév. & Vaniot 1925. Photinia laevis  var. villosa  (Thunb.) Koidz. 2018. Pyrus villosa  (Thunb.) M.F.Fay & Christenh. | Oriental redtip ⓘ Oriental photinia ⓘ Christmas Berry ⓘ | Introduced from  temperate Asia, Naturalized, SNA, GNR Perennial, Shrub, tree | | NYFA: X NYFA: 7 counties iNat 0 counties | USDA-X0 POWO ARS-GRIN FNA Images Wikispecies |

### Crataegus

* Crataegus submollisThere are about fifty hawthorn taxa listed for New York, and most are considered to be natives to New York State.
* The subgeneric system used here for hawthorns is from the Canadensys Database of Vascular Plants of Canada (VASCAN). Their data is based in part on the Flora of North America north of Mexico, Vol. 9, (in prep).

#### Crataegus sect. Coccineae

* Crataegus macrospermaSection Coccineae correspond to the primary North American Crataegus clade, containing 20 series.
* ↑ James B. Phipps (2021). Crataegus sect. Coccineae in Flora of North America. Volume 9. Flora of North America Association.
Rosales — Rosaceae — Amygdaloideae — Maleae — Crataegus subg.   sect. Coccineae :
| Genus Crataegus ser. Coccineae | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus coccinea L. var. coccinea1753. Crataegus coccinea  L. in… 1901. Crataegus pedicellata  Sarg. in… 1908. Crataegus aulica  Sarg. in… 1908. Crataegus letchworthiana  Sarg. in… | Scarlet hawthorn, Red-fruited hawthorn, Red hawAubépine écarlate, Épine écarlate | Native, CoC: 2, Likely secure, S4, G5 Perennial, Tree, shrub | | NYFA: 4 | USDA-NN COL Go Botany VASCAN ARS-GRIN ITIS Tropicos Images Wikispecies |
| Crataegus coccinea L.var. pringlei (Sarg.) J.A.Macklin & J.B.Phipps1901. Crataegus pringlei Sarg. 1901. Crataegus lobulata Sarg. 1903. Crataegus exclusa Sarg. 1908. Crataegus pringlei  var. lobulata  (Sarg.) Eggl. 1908. Crataegus pringlei  var. exclusa  (Sarg.) Eggl. 1920. Crataegus pedicellata  var. gloriosa  (Sarg.) Sarg. 2007. Crataegus coccinea  var. pringlei  J.A.Macklin & J.B.Phipps | Pringle's hawthorn ⓘ Aubépine de Pringle | Native, Threatened | | NYFA: 2? | USDA-NN COL POWO Go Botany VASCAN ARS-GRIN ITIS Tropicos Images Wikispecies |
| Crataegus coccinea L.var. fulleriana (Sarg.) J.A.Macklin & J.B.Phipps1903. Crataegus fulleriana Sarg. 1908. Crataegus chippewaensis Sarg. 1908. Crataegus confragosa Sarg. 1923. Crataegus illecebrosa Sarg. 2011. Crataegus coccinea  var. fulleriana  J.A.Macklin & J.B.Phipps | Fuller's hawthornAubépine de Fuller | Native, Unranked | | NYFA: U | USDA-NN COL VASCAN ARS-GRIN ITIS Tropicos BONAP Images Wikispecies |
| Crataegus holmesiana Ashe1900. Crataegus holmesiana Ashe1901. Crataegus (×) anomala Sarg.1901. Crataegus arnoldiana Sarg.1901. C. holmesianavar. villipes Ashe1903. Crataegus villipes Ashe1908. Crataegus tardipes Sarg.1920. C. holmesianavar. tardipes Sarg. | Holmes' hawthorn, Arnold hawthornAubépine de Holmes, Aubépine anormale | Native, Endangered | | NYFA: 1? | USDA-N0 Go Botany VASCAN ARS-GRIN ITIS Tropicos NatureServe BONAP IPN Images Wikispecies |
| Crataegus magniflora Sarg.1902. Crataegus corusca Sarg.1903. Crataegus magniflora Sarg.1903. Crataegus hillii Sarg.1908. Crataegus dayana Sarg.1908. Crataegus macounii Sarg.1969. Crataegus fullerianavar. magnifloraJ.S.Pringle | Shining-branch hawthorn, Gleaming hawthorn, Broad-leaved hawthornAubépine à grandes fleurs, Aubépine à rameaux luisants | Native, Unranked | | | USDA-NN VASCAN ARS-GRIN Images Wikispecies |
| Genus Crataegus ser. Rotundifoliae | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus chrysocarpa Ashevar. chrysocarpa1900. Crataegus chrysocarpa Ashe | Fireberry hawthorn, Fire hawthorn, Northern hawthornAubépine dorée | Native, CoC: 2, Likely secure Perennial, Tree, shrub | | NYFA: 4 | USDA-NN Go Botany VASCAN ARS-GRIN ITIS Images Wikispecies |
| Crataegus chrysocarpa Ashevar. virgintistamina    Crataegus chrysocarpavar. virgintistamina | Fireberry hawthorn, Fire hawthorn, Northern hawthornAubépine dorée | Native, CoC: 2, Unranked Perennial, Tree, shrub | | NYFA: U | USDA-NN Go Botany VASCAN ARS-GRIN ITIS Images Wikispecies |
| Crataegus irrasa Sarg.1903. Crataegus irrasa Sarg.1907. Crataegus noveboracensis Sarg.1907. Crataegus oakesiana Eggl.1923. Crataegus mansfieldensis Sarg. | Blanchard hawthorn, Blanchard's hawthornAubépine pubérulente | Native, CoC: 2, Endangered Perennial, Tree, shrub | | NYFA: 1? | USDA-NN Go Botany VASCAN ARS-GRIN ITIS Images Wikispecies |
| Genus Crataegus ser. Tenuifoliae | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus macrosperma Ashe1900. Crataegus macrosperma Ashe1900. Crataegus roanensis Ashe1901. Crataegus acutiloba Sarg.1901. Crataegus matura Sarg.1901. Crataegus pastorum Sarg.1902. Crataegus brumalis Ashe1903. Crataegus demissa Sarg. | Big-fruit hawthorn, Earlythorn, Variable hawthornAubépine à lobes aigus | Native, CoC: 2, Secure Perennial, Tree, shrub | | NYFA: 5 | USDA-NN Go Botany VASCAN ARS-GRIN ITIS Tropicos IPN Images Wikispecies |
| Crataegus flabellata (Bosc ex Spach) K.Koch1853. Crataegus flabellata K.Koch1901. C. densiflora Sarg.nom. illeg.1908. C. grayana Eggl.1946. C. flabellatavar. grayana E.J.Palmer | Fan-leaf hawthorn, Gray's hawthornAubépine flabelliforme | Native, CoC: 2, Endangered Perennial, Tree, shrub | | NYFA: 1? | USDA-NN Go Botany VASCAN ARS-GRIN ITIS Tropicos Images Wikispecies |
| Crataegus schuettei Ashevar. schuettei    Crataegus schuettei Ashe    Crataegus basilica Beadle | Schuette's hawthorn, Royal hawthorn | Native, CoC: 2, Endangered Aubépine de Schuette Perennial, Tree, shrub | | NYFA: 1? | USDA-NN Go Botany VASCAN ITIS Images Wikispecies |
| Crataegus prona Ashe = C. macrosperma (GRIN) 1903. Crataegus prona Ashe1904. Crataegus gravis Ashe | Illinois hawthorn | Native, CoC: 2, Unranked Perennial, Tree | | NYFA: U | USDA-N0 ARS-GRIN ITIS Images Wikispecies |
| Crataegus fluviatilis Sarg. = C. macrosperma (USDA) 1903. Crataegus fluviatilis Sarg.1903. Crataegus apiomorpha Sarg.1903. Crataegus cyanophylla Sarg.1965. C. roanensisvar. fluviatilis Kruschke1965. C. apiomorphavar. cyanophylla Kruschke | River hawthorn, Fort Sheridan hawthornAubépine fluviatile | Native, CoC: 2, Unranked Perennial, Tree | | NYFA: U | USDA-NN Go Botany VASCAN ITIS IPN Images Wikispecies |
| Genus Crataegus ser. Punctatae | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus punctata Jacq. (1770)C. fastosa Sarg. | Dotted hawthorn, Hillside hawthornAubépine ponctuée | Native, Likely secure | | NYFA: 4 | USDA-NN ARS-GRIN ITIS Images Wikispecies |
| Crataegus jonesiae Sarg. (1901 "jonesae")    C. harryi Sarg.    C. laurentiana Sarg. | Jones' hawthorn, Miss Jones' hawthornAubépine de Jones | Native, Unranked | | NYFA: U | USDA-NN ARS-GRIN ITIS Images Wikispecies |
| Genus Crataegus ser. Molles | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus submollis Sarg. (1901)    C. canadensis Sarg.    C. arnoldiana Sarg.    C. champlainensis Sarg. | Champlainthorn, Quebec hawthornAubépine subsoyeuse | Native, Vulnerable | | NYFA: 3? | USDA-NN ARS-GRIN Images Wikispecies |
| Crataegus mollis (Torr. & A. Gray) Scheele (1848) | Arnold's hawthorn, Downy hawthorn | Native, Endangered | | NYFA: 1 | USDA-NN ARS-GRIN ITIS Images Wikispecies |
| Genus Crataegus ser. Silvicolae | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus pruinosa (Wendl.f.) K.Koch (1853) | Frosted hawthorn, Frostythorn, Waxyfruit hawthorn, Waxy-fruited hawthorn | Native, Likely secure | | NYFA: 4 | USDA-NN ARS-GRIN ITIS Images Wikispecies |
| Crataegus jesupii Sarg. (1903)    C. filipes Ashe    C. dissonavar. bellula | Jesup's hawthorn, Jesup's-thorn | Native, Threatened | | NYFA: 2? | USDA-N0 ARS-GRIN ITIS Images Wikispecies |
| Crataegus compta Sarg. (1903)    Crataegus populnea Ashe    C. iracundavar. populnea | Adorned hawthornAubépine faux-peuplier | Native, Endangered | | NYFA: 1? | USDA-NN ITIS Images Wikispecies |
| Crataegus iracunda Beadle (1902)C. beckwithiae. Sarg.C. stolonifera. Sarg. | Stolonbearing hawthorn | Native, No recent reports | | NYFA: Z | USDA-NN ARS-GRIN ITIS Images Wikispecies |
| Crataegus suborbiculata Sarg. (1901) | Caughuawaga hawthorn | Native, Endangered | | NYFA: 1? | USDA-NN ARS-GRIN ITIS Images Wikispecies |
| Genus Crataegus interserial hybrids | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus × hudsonica Sarg.  C. pruinosa (ser. Silvicolae) ×   C. punctata (ser. Punctatae) | Hybrid of  Frosted hawthorn &  Dotted hawthorn | Native, Endangered | | NYFA: 1? | Images Wikispecies |
| Crataegus × immanis Ashe (pro sp.)  C. chrysocarpa (ser. Rotundifoliae) ×   C. pruinosa (ser. Silvicolae) | Hybrid of  Fireberry hawthorn &  Frosted hawthorn | Native, Endangered | | NYFA: 1? | USDA-N0 ITIS Images Wikispecies |
| Crataegus × maligna Sarg.  C. chrysocarpa (ser. Rotundifoliae) ×   C. macrosperma (ser. Tenuifoliae) | Hybrid of  Fireberry hawthorn &  Big-fruit hawthorn | Native, Unranked | | NYFA: U | USDA-N0 ITIS Images Wikispecies |
| Crataegus × puberis Sarg. (pro sp.)  C. flabellata (ser. Tenuifoliae) ×   C. punctata (ser. Punctatae) | Hybrid of  Fan-leaf hawthorn &  Dotted hawthorn | Native, Unranked | | NYFA: U | USDA-N0 ITIS Images Wikispecies |
| C. macrosperma × pruinosa  C. macrosperma (ser. Tenuifoliae) ×   C. pruinosa (ser. Silvicolae) | Hybrid of  Big-fruit hawthorn &  Frosted hawthorn | Native, Unranked | | NYFA: U | Images Wikispecies |
| Crataegus lucorum Sarg.  ser. Tenuifoliae ×   ser. Coccineae1901. Crataegus lucorum Sarg.1905. Crataegus insolens Sarg.1908. Crataegus lucorumvar. insolens Eggl.1913. Crataegus knieskerniana Sarg. | Grove hawthorn | Native, Unranked | | NYFA: U | USDA-N0 Go Botany ITIS Images Wikispecies |

#### Crataegus sect. Crataegus

Rosales — Rosaceae — Amygdaloideae — Maleae — Crataegus sect. Crataegus :
| Genus Crataegus ser. Dilatatae | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus coccinioides Ashe (1899) | Kansas hawthorn | Native, Endangered | | NYFA: 1? | USDA-NN ARS-GRIN ITIS Images Wikispecies |
| Genus Crataegus ser. Crus-galli | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus crus-galli L.1753. Crataegus crus-galli L.1784. Crataegus salicifolia Medik.1902. Crataegus regalis Beadle | Cockspur, Cockspurthorn, Cockspur hawthorn, Newcastle thorn, Bush hawthornAubépine ergot-de-coq | Native, Secure | | NYFA: 5 | USDA-NN ARS-GRIN ITIS Tropicos Images Wikispecies |
| Crataegus berberifolia Torr. & A.Gray (1840) | Barberry hawthorn | Native, Endangered | | NYFA: 1 | USDA-N Images Wikispecies |
| Genus Crataegus ser. Parvifoliae | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus uniflora Münchh. (1770) | Dwarf hawthorn, One-flower hawthorn | Native, Endangered, NYNHP: 1 | | NYFA: 1 | USDA-N0 ARS-GRIN ITIS Images Wikispecies |
| Genus Crataegus ser. Intricatae | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus intricata Lange1895. Crataegus intricata Lange1902. Crataegus foetida Ashe | Copenhagen hawthorn, Lange's-thorn, Thicket hawthorn, Biltmore hawthornAubépine intriquée | Native, Threatened | | NYFA: 2 | USDA-N Go Botany VASCAN ARS-GRIN ITIS Tropicos Images Wikispecies |
| Crataegus biltmoreana Beadle1899. Crataegus biltmoreana Beadle2011. Crataegus intricatavar. biltmoreanaR.W.Lance | Biltmore hawthorn | Native, Endangered | | NYFA: 1? | Go Botany ARS-GRIN ITIS Tropicos Images Wikispecies |
| Genus Crataegus ser. Montaninsulae | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus beata Sarg. (1903) | Dunbar's hawthorn | Native, Endangered | | NYFA: 1 | USDA-N Images Wikispecies |
| Crataegus disperma Ashe | Spreading hawthorn | Native, Endangered | | NYFA: 1 | USDA-NN Images Wikispecies |
| Crataegus laurentiana Sarg.    Crataegus brunetiana    C. laurentianavar. brunetiana    C. laurentianavar. dissimilifolia | Laurentian hawthorn | Native, Unranked | | NYFA: U | Images Wikispecies |
| Crataegus opulens Sarg.Crataegus beata. var. opulens | Rochester hawthorn | Native, Unranked | | NYFA: U | USDA-N0 ITIS Images Wikispecies |
| Genus Crataegus ser. Cordatae | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus phaenopyrum (L.f.) Medik.1757. Mespilus cordata Mill. nom. rej.1781. Mespilus phaenopyrum L.f.1788. Crataegus populifolia Walter1789. Crataegus cordata Aiton1793. Crataegus phaenopyrum Medik. | Washington hawthornAubépine petit-corail, Aubépine de Washington | Introduced, US South native | | NYFA: unk | USDA-NX Go Botany VASCAN Tropicos NatureServe BONAP Images Wikispecies |
| Genus Crataegus ser. Crataegus | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus monogyna Jacq.1775. Crataegus monogyna Jacq.1838. C. oxyacanthavar. praecox hort. | Common hawthorn, English hawthorn, Oneseed hawthorn, Single-seed hawthornAubépine monogyne | Introduced from  Eurasia,  northern Africa, Potentially invasive, CP-5 IPA | | NYFA: X | USDA-XX Go Botany ARS-GRIN ITIS Images Wikispecies |
| Crataegus laevigata (Poir.) DC.    Crataegus oxyacanthoides Thuillier    Crataegus palmstruchii Lindman    Mespilus laevigata Poiret | Smooth hawthorn, English hawthorn, Midland hawthornAubépine épineuse, Aubépine à deux styles | Introduced, from Eurasia | | | USDA-X VASCAN Images Wikispecies |
| ↑ New York Natural Heritage Program. 2024. Online Conservation Guide for Crataegus uniflora. Endangered, S1/G5. ↑ Appendix 5. Plant species of concern (Watch List) within the central Finger Lakes region. Policy on the use of non-native plants in Cornell Botanic Gardens' accessioned collections (2018) ||||||

#### Crataegus sect. Macracanthae

Rosales — Rosaceae — Amygdaloideae — Maleae — Crataegus sect. Macracanthae :
| Genus Crataegus ser. Macracanthae | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus calpodendron (Ehrh.) Medik.1788. Mespilus calpodendron Ehrh.1793. Crataegus calpodendron Medik.1908. Crataegus hispidula Sarg.1935. C. calpodendronvar. hispidulaE.J.Palmer | Pear hawthorn, Blackthorn hawthorn, Urn-fruit hawthornAubépine à fruits piriformes, Aubépine tomenteuse | Native, Likely secure | | NYFA: 4? | USDA-NN VASCAN ARS-GRIN ITIS Images Wikispecies |
| Crataegus succulenta Schrad. ex Link (1831) | Fleshy hawthorn, Long-spine hawthorn, Succulent hawthorn | Native, Likely secure | | NYFA: 4 | USDA-NN Go Botany ARS-GRIN ITIS Images Wikispecies |
| Crataegus macracantha Lodd. ex Loudonvar. macracantha1838. C. macracantha Lodd. ex Loudon1874. Mespilus macracantha Wenz.1886. C. coccineavar. macracantha Dudley1921. C. succulentavar. macracantha Eggl. | Long-thorned hawthornAubépine à épines longues | Native, Unranked | | NYFA: U | USDA-NN VASCAN ARS-GRIN ITIS Tropicos Images Wikispecies |
| Crataegus macracanthavar. occidentalis    C. succulentavar. divida    C. succulentavar. occidentalis    C. succulentavar. rhombifolia    C. succulenta occidentalis | Large-thorn hawthorn | Native, Unranked | | NYFA: U | USDA-NN ARS-GRIN Images Wikispecies |
| Crataegus menandiana Sarg. | Menands hawthorn | Native, No recent reports | | NYFA: Z | USDA-N0 ITIS Images Wikispecies |
| Genus Crataegus ser. Brainerdianae | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus brainerdii Sarg.1901. Crataegus brainerdii Sarg.1905. Crataegus seelyana Sarg.1913. Crataegus shirleyensis Sarg.1915. Oxyacantha brainerdii Nieuwl. | Brainerd hawthorn, Brainerd's hawthornAubépine de Brainerd | Native, Vulnerable | | NYFA: 3 | USDA-NN Go Botany ITIS Tropicos Images Wikispecies |
| Crataegus scabrida Sarg.1901. Crataegus scabrida Sarg.1908. C. brainerdiivar. scabrida Eggl. | Rough hawthorn, Scabrid hawthorn | Native, Vulnerable | | NYFA: 3? | USDA-NN Go Botany ITIS Images Wikispecies |
| Genus Crataegus interserial hybrids | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus × peckietta Sarg. (pro sp.)  Crataegus scabrida ×   Crataegus succulenta | Hybrid of  Rough hawthorn &  Fleshy hawthorn | Native, Endangered | | NYFA: 1? | USDA-N0 ITIS Images Wikispecies |
| Crataegus × laneyi Sarg. (pro sp.)  Crataegus brainerdii ×   Crataegus succulenta | Hybrid of  hawthorn &  hawthorn | Native, Unranked | | NYFA: U | USDA-N0 ITIS Images Wikispecies |

#### Crataegus sect. Virides

Rosales — Rosaceae — Amygdaloideae — Maleae — Crataegus sect. Virides :
| Genus Crataegus sect. Virides | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus viridis L.1753. Crataegus viridis L.1908. Crataegus dawsoniana Sarg. | Green hawthorn, Green haw, Southernthorn | Introduced, US South native, No NY reports, Cultivated | | NYFA: 0 | USDA-N0 ARS-GRIN ITIS Tropicos NatureServe BONAP Images Wikispecies |

#### Crataegus inter-sectional hybrids

Rosales — Rosaceae — Amygdaloideae — Maleae — Crataegus — inter-sectional hybrids :
| Genus Crataegus sect. Coccineae × Crataegus | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus × brevipes Peck (pro sp. 1910)  C. pruinosa (sect. Coccineae) ×   C. intricata (sect. Crataegus) | Hybrid of  Copenhagen hawthorn &  Frosted hawthorn | Native, Unranked | | NYFA: U | ITIS Images Wikispecies |
| C. monogyna × punctata  C. punctata (sect. Coccineae) ×   C. monogyna (sect. Crataegus) | Hybrid of  English hawthorn &  Dotted hawthorn | Native, Unranked | | NYFA: U | Images Wikispecies |
| Genus Crataegus sect. Coccineae × Macracanthae | Hawthorn | N.Y. Status | Images | Distribution | NY NPT |
| Crataegus × persimilis Sarg. (pro sp.)  C. crus-galli (sect. Crataegus) ×   C. succulenta (sect. Macracanthae)1798. Mespilus prunifolia Poir. illeg-hom.1806. C. prunifolia Pers, illeg-hom.1840. C. crus-gallivar. prunifolia Torr. & A.Gray1903. Crataegus persimilis Sarg.1908. Crataegus cerasina Sarg. | Plumleaf hawthorn, Plum-leaved hawthorn, Broadleaf cockspurthornAubépine à feuilles de prunier | Native, Unranked | | NYFA: 0 | USDA-NN VASCAN ARS-GRIN Tropicos Images Wikispecies |
| Crataegus × rubrocarnea Sarg. (pro sp.)  C. macrosperma (sect. Coccineae) ×   C. brainerdii (sect. Macracanthae) | Hybrid of  Brainerd hawthorn &  Big-fruit hawthorn | Native, Unranked | | NYFA: U | USDA-N0 ITIS Images Wikispecies |
| C. macrosperma × succulenta  C. macrosperma (sect. Coccineae) ×   C. succulenta (sect. Macracanthae) | Hybrid of  Big-fruit hawthorn &  Fleshy hawthorn | Native, Unranked | | NYFA: U | Images Wikispecies |

### Sorbus

* The mountain-ash trees in the genus Sorbus are not closely related to the ash trees of the genus Fraxinus and as such are not subject to the ravages of the emerald ash borer. The use of the term "ash" is probably due to the superfical similarity of the compound leaves. The mountain-ashes are more closely related to the hawthorns, apples and crabapples, which are also in the Maleae tribe.

#### Sorbus sect. Sorbus

* Sorbus americanaThere are two native mountain-ash trees in New York.
Rosales — Rosaceae — Amygdaloideae — Maleae — Sorbus sect. Commixtae :
| Genus Sorbus sect. Commixtae | Mountain-ash | N.Y. Status | Images | Distribution | NY NPT |
| Sorbus americana Marshall1785. Sorbus americana  Marshall in… 1806. Sorbus aucuparia  var. americana  (Marshall) Pers.  in… 1825. Pyrus microcarpa  (Pursh) DC. in… 1825. Pyrus americana  (Marshall) DC. in… 1825. Pyrus microcarpa  (Pursh) Spreng. in… 1840. Pyrus americana  var. microcarpa  (Pursh) Torr. & A.Gray  in… 1868. Crataegus americana  (Marshall) Chalon in… 1914. Pyrus americana  ssp. microcarpa  (Pursh) Juel, Cedergr. & Örtendahl  in… 1915. Aucuparia americana  (Marshall) Nieuwl. in… | American mountain-ash, Mountain-ashSorbier d'Amérique | Native, CoC: 8, Secure, S5 FAC-FACU Perennial, Tree, Shrub | | NYFA: 5 | USDA-NN POWO Go Botany VASCAN ARS-GRIN ITIS LBJ Images Wikispecies |
| Sorbus decora (Sarg.) C.K.Schneid.1825. Pyrus americana DC. 1902. Pyrus americana  var. decora  Sarg. 1906. Sorbus americana  var. groenlandica  C.K.Schneid. 1906. Sorbus decora C.K.Schneid. 1943. Pyrus decora (Sarg.) Hyl. 1974. Pyrus groenlandica K.R.Robertson | Northern mountain-ash, Showy mountain-ash | Native, CoC: 9, Secure, S5 FACU Perennial, Tree, Shrub | | NYFA: 5 | USDA-NN Go Botany ARS-GRIN ITIS LBJ Images Wikispecies |

#### Sorbus sect. Commixtae

* Sorbus aucupariaThe European mountain-ash, or rowan, is a commonly-planted ornamental tree that does naturalize in New York State. European mountain-ash trees can be distinguished from the American species by the long villous hairs on the undersides of the leaves as well as on the twigs and bud scales of the European species.
Rosales — Rosaceae — Amygdaloideae — Maleae — Sorbus sect. Commixtae :
| Genus Sorbus sect. Sorbus | Mountain-ash | N.Y. Status | Images | Distribution | NY NPT |
| Sorbus aucuparia L. ssp. aucuparia1753. Sorbus aucuparia  L. in… 1771. Mespilus aucuparia  (L.) Scop. in… 1790. Pyrus aucuparia  (L.) Gaertn. in… 1793. Aucuparia silvestris  Medik. in… 1796. Crataegus aucuparia  (L.) Salisb. in… 1811. Pyrenia aucuparia  (L.) Clairv. in… 1868. Aucuparia pinnata  Fourr. in… 1892. Sorbus aucuparia typica Beck in… 1906. Pyrus aucuparia  var. typica  (C.K.Schneid.) Asch. & Graebn.  in… 1913. Sorbus aucuparia  ssp. euaucuparia  Briq.  in… 2018. Pyrus altaica  (Koehne) M.F.Fay & Christenh. in… 2018. Pyrus anadyrensis  (Kom.) M.F.Fay & Christenh. in… 2018. Pyrus boissieri  (C.K.Schneid.) M.F.Fay & Christenh. in… | European mountain ash ⓘ Rowan Rowan-berry Sorbier des oiseleurs | Introduced from  Eurasia, Potentially invasive, IPA-US, WoW, Cultivated, Naturalized, SNA, G5 Perennial, Tree, shrub | | NYFA: X | USDA-XX POWO Go Botany ARS-GRIN ITIS Images Wikispecies |

### Malus

* Malus contains the apples and crabapples, one species of which (the sweet crabapple) is considered to be native to New York.

#### Malus sect. Chloromeles

* Malus coronariaThe sweet crabapple (Malus coronaria) has in the past been treated as a number of separate taxa, but most modern sources treat them as a single species. This is the only New York native crabapple and probably the only North American native crabapple found in the wild in New York. The native crabapple is not widely planted due to its susceptibility to cedar-apple rust (Gymnosporangium juniperivirginianae), cedar-quince rust (G. clavipes), and cedar-hawthorn rust (G. globosum).
Rosales — Rosaceae — Amygdaloideae — Maleae — Malus sect. Chloromeles :
| Genus Malus sect. Chloromeles | Sweet crabapple | N.Y. Status | Images | Distribution | NY NPT |
| Malus coronaria (L.) Mill.1753. Pyrus coronaria L. 1768. Malus coronaria (L.) Mill. 1892. Sorbus coronaria (L.) MacMill. 1911. M. glaucescens Rehder 1911. M. lancifolia Rehder 1913. M. bracteata Sarg. 1916. P. glaucescens L.H.Bailey 1918. M. carolinensis Ashe 1920. M. coronaria  var. dasycalyx  Rehder 1947. P. coronaria  var. dasycalyx  Fernald 1983. M. coronaria  var. glaucescens  Likhonos | Wild crab apple ⓘ Sweet crabapple ⓘ Sweet crab apple ⓘ Fragrant crabapple American crabapple American crab Narrow-leaved crabapple Garland crabapple Wild crabapple Golden crabapple Pommier odorant Pommier américain | Native, CoC: 2, Vulnerable Perennial, Strub-tree | | NYFA: 3? | USDA-NN Go Botany-0 VASCAN ARS-GRIN ITIS FNA Tropicos NatureServe BONAP IPN Images Wikispecies |

#### Malus sect. Gymnomeles

* Malus baccataSiberian crabapple (Malus baccata) is a popular ornamental tree and only occasionally escapes cultivation in New York.
* Tea crabapple (Malus hupehensis) wasn't collected in New York State until 2015. As of 2021, it has been reported from all New York City counties.
Rosales — Rosaceae — Amygdaloideae — Maleae — Malus sect. Gymnomeles :
| Genus Malus sect. Gymnomeles | Crabapple | N.Y. Status | Distribution | NY NPT |
| Malus baccata (L.) Borkh.1756. Pyrus baccata L. 1803. Malus baccata (L.) Borkh. 2018. Sinomalus baccata (L.) Rushforth | Siberian crabapple, Siberian crab-apple, Flowering crabapple, Chinese crabapplePommier de Sibérie, Pommier sibérien, Pommier à baies | Introduced from  eastern Asia, Highly invasive,  NYIS Tier: 2,  iMapInvasives,  Invasive.org: , Naturalized, CP-4 | NYFA: 7 counties iNat: 1 county iMap: 2 counties NYFA: X iNat | USDA-XX Go Botany VASCAN ARS-GRIN Images Wikispecies |
| Malus hupehensis (Pamp.) Rehder1915. Malus theifera Rehder 1916. Pyrus theifera (Rehder) L. H. Bailey 1933. Malus hupehensis (Pamp.) Rehder 2018. Sinomalus hupehensis (Pamp.) Rushforth | Tea crab apple ⓘ Tea crabapple ⓘ Hupeh crabapple ⓘ | Introduced from  China, Highly invasive,  NYIS Tier: 2,  iMapInvasives,  Invasive.org: , Naturalized, SNA | NYFA: 1 county iNat: 5 counties NYFA: Xn iNat GBIF | USDA-X0 ARS-GRIN Images Wikispecies |
| Malus toringo (Siebold) Siebold ex de Vriese1848. Sorbus toringo Siebold 1856. Malus toringo (Siebold) Siebold ex de Vriese 1858. Pyrus sieboldii Regel 1867. Pyrus toringo (Siebold) Miq. 1915. Malus sieboldii Rehder 1915. Malus sargentii Rehder 2018. Sinomalus sieboldii (Regel) Rushforth | Toringa crabapple, Toringo crabPommier toringo, Pommetier de Siebold   arborescent | Introduced from  China, Japan, Korea, Potentially invasive,  NYIS Tier: 3,  iMapInvasives,  Invasive.org: , NJIST Target | NYFA: X NYFA: 4 counties iNat 7 counties | USDA-X0 Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos NatureServe BONAP Images Wikispecies |
| ↑ Appendix 4. Plant species that are considered moderately invasive to natural areas in the central Finger Lakes region. Policy on the use of non-native plants in Cornell Botanic Gardens' accessioned collections (2018) ↑ New Jersey Invasive Species Strike Team (2015) "Do not Plant List." ↑ NJ Status: Emerging Stage 2 - Common and spreading rapidly. It is highly threatening to natural plant communities. All detected occurrences should be eradicated. New Jersey Invasive Species Strike Team (2012) "Target Species Fact Sheets." ||||||

#### Malus sect. Malus

* Isaac Newton's apple treeSection Malus contains the common domesticated apple (Malus pumila or Malus domestica) along with a number of Eurasian crabapples. Qian (2010) proposed conserving Malus domestica as the acceped name of the domesticated apple. It is important to note that apples are extremely heterozygous, resulting in naturalized apples that bear little resemblance to domestic apples, which are generally propagated by grafting. So the presence of apples in the wild that look and taste like cultivated apples is usually the result of long-living planted apple trees being subsumed by the wild and not by apple seeds being spread into a natural area.
* ↑ Guan-Ze Qian, Lian-Fen Liu and Geng-Guo Tang (2010). "(1933) Proposal to conserve the name Malus domestica against M. pumila, M. communis, M. frutescens, and Pyrus dioica (Rosaceae)." Taxon, Vol. 59, No. 2 (April 2010), pp. 650-652.
* ↑ Pollan, Michael (2001-06-12). The Botany of Desire: A Plant's-Eye View of the World (Kindle Location 315). Random House Publishing Group. Kindle Edition.
* ↑ "Malus pumila." Flora of North America, Vol. 9.
Rosales — Rosaceae — Amygdaloideae — Maleae — Malus sect. Malus :
| Genus Malus sect. Malus | Apple & crabapple | N.Y. Status | Images | Distribution | NY NPT |
| Malus domestica (Suckow) Borkh.1753. Pyrus malus L. 1753. Pyrus malus  var. sylvestris  L. 1768. Malus pumila Mill. 1768. Malus sylvestris (L.) Mill. 1798. Malus communis Desf. 1803. Malus domestica Borkh. 1821. Pyrus pumila K.Koch 1821. Pyrus sylvestris Gray | Cultivated apple, Orchard apple, Paradise apple, European crabapplePommier commun, Pommier cultivé, Pommier paradis, Pommier nain, Pommier sauvage | Introduced from  Europe, Highly invasive, Naturalized, CP-4 | | NYFA: X | USDA-XX Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos NatureServe BONAP IPN Images Wikispecies |
| Malus prunifolia (Willd.) Borkh.1794. Pyrus prunifolia Willd.1803. Malus prunifolia Borkh. | Pear-leaved crabapple, Plum-leaved crabapple, Plumleaf apple, Chinese apple, Chinese crabapplePommier à feuilles   de prunier | Introduced from  China, Potentially invasive, CP-5 | | Bronx, Oswego, Rensselaer NYFA: X | USDA-XX Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos NatureServe BONAP IPN Images Wikispecies |
| Malus floribunda Siebold ex Van Houtte1864. Pyrus floribunda G.Kirchn. 1865. Malus floribunda Siebold ex Van Houtte 1906. Pyrus pulcherrima Asch. & Graebn. 2018. Sinomalus sieboldii (Regel) Rushforth | Japanese flowering   crabapple, Japanese crab apple, Japanese crab | Introduced from  East Asia, Potentially invasive, CP-4 | | Bronx NYFA: X | USDA-X0 Go Botany VASCAN-0 ARS-GRIN ITIS NatureServe BONAP Images Wikispecies |
| Genus Malus sect. Malus (excluded taxa) | Apple & crabapple | N.Y. Status | Images | Distribution | NY NPT |
| Malus prunifolia × pumila  Malus prunifolia ×   Malus pumila | Hybrid of  cultivated apple &  plumleaf crabapple | Introduced, N.Y. excluded, CP-5 | | NYFA-Excluded | Go Botany Images Wikispecies |
| 1 2 Appendix 4. Plant species that are considered moderately invasive to natural areas in the central Finger Lakes region. Policy on the use of non-native plants in Cornell Botanic Gardens' accessioned collections (2018) 1 2 Appendix 5. Plant species of concern (Watch List) within the central Finger Lakes region. Policy on the use of non-native plants in Cornell Botanic Gardens' accessioned collections (2018) ||||||

### Pyrus

* Pyrus communis common pearPyrus contains two species of pears that have naturalized in New York. The common pear (Pyrus communis) is grown for its fruit and occasionally escapes or persists from cultivation. The Bradford or Callery pear is widely planted as an ornamental tree, but has been found to naturalize in some locations, particularly in Suffolk County.
Rosales — Rosaceae — Amygdaloideae — Maleae — Pyrus :
| Genus Pyrus | Pear | N.Y. Status | Distribution | NY NPT |
| Pyrus communis L.1753. Pyrus communis L.    Pyrus sativa DC.    Pyrus caucasica Fed.    Pyrus pyraster (L.) Du Roi | Pear, Common pear, Poirier | Introduced from  eastern Asia, Potentially invasive, Naturalized | NYFA: X NYFA: 15 counties 10 counties | USDA-XX ARS-GRIN ITIS Images Wikispecies |
| Pyrus calleryana1872. Pyrus calleryana Decne.1911. Pyrus kawakamii Hayata1920. P. calleryanafo. graciliflora Rehder1920. P. calleryanafo. tomentella Rehder | Bradford pear, Callery pear | Introduced from  Asia, Highly invasive,  NYIS: 65%, NYS Tier 4, CP-5, NPS | Orange, Suffolk NYFA: X | Images Wikispecies |
| ↑ New York non-native plant invasiveness assessment – Pyrus calleryana: Moderate (65). ↑ Appendix 5. Plant species of concern (Watch List) within the central Finger Lakes region. Policy on the use of non-native plants in Cornell Botanic Gardens' accessioned collections (2018) ||||||

### Cydonia

Rosales — Rosaceae — Amygdaloideae — Maleae — Malinae — Cydonia :
| Genus Cydonia | Quince | N.Y. Status | Images | Distribution | NY NPT |
| Cydonia oblonga Mill.1753. Pyrus cydonia L.1768. Cydonia oblonga Mill.1806. Cydonia vulgaris Pers. | Common quinceCognassier, Coing | Introduced from  temperate Asia | | Dutchess, Orange, Rockland, Tompkins NYFA: X | USDA-XX Go Botany VASCAN ARS-GRIN ITIS Images Wikispecies |

### Chaenomeles

Rosales — Rosaceae — Amygdaloideae — Maleae — Malinae — Chaenomeles :
| Genus Chaenomeles | Flowering quince | N.Y. Status | Images | Distribution | NY NPT |
| Chaenomeles japonica (Thunb.) Lindl. ex Spach1780. Pyrus japonica  Thunb. in… 1806. Cydonia japonica  (Thunb.) Pers. in… 1834. Chaenomeles japonica  (Thunb.) Lindl. ex Spach in… 1869. Aronia japonica  (Thunb.) K.Koch in… 1900. Cydonia sargentii  Lemoine in… | Japanese flowering quince, Maule's quinceCognassier du Japon | Introduced from  Japan, Korea | | Albany, Cattaraugus, Chemung, Ontario NYFA: X | USDA-XX ARS-GRIN ITIS Images Wikispecies |
| Chaenomeles speciosa (Sweet) Nakai (1929)    Chaenomeles lagenaria    Cydonia speciosa Sweet | Chinese quince, Flowering quince, Japanese quince | Introduced from  Asia | | Dutchess, Otsego NYFA: X | USDA-X0 ARS-GRIN ITIS Images Wikispecies |

### Pyracantha

Rosales — Rosaceae — Amygdaloideae — Maleae — Malinae — Pyracantha :
| Genus Pyracantha | Firethorn | N.Y. Status | Images | Distribution | NY NPT |
| Pyracantha coccinea M.Roem.1753. Mespilus pyracantha L.1793. Crataegus pyracantha Medik.1834. Cotoneaster pyracantha Spach1847. Pyracantha coccinea Roemer | Scarlet firethorn | Introduced, from Eurasia | | Otsego, Rockland NYFA: X | USDA-XN ARS-GRIN Images Wikispecies |

### Cotoneaster

Rosales — Rosaceae — Amygdaloideae — Maleae — Cotoneaster :
| Genus Cotoneaster | Cotoneaster | N.Y. Status | Images | Distribution | NY NPT |
| Cotoneaster atropurpureus Flinck & B. Hylmö    Cotoneaster atropurpureus in… | Purple-flowered cotoneaster ⓘ | Introduced, Not naturalized | | Ulster only NYFA: Xm | USDA- Images Wikispecies |
| Cotoneaster cochleatus (Franch.) G. Klotz    Cotoneaster cochleatus in… | Yunnan cotoneaster ⓘ | Introduced, Not naturalized | | Orange only NYFA: Xm | USDA- Images Wikispecies |
| Cotoneaster divaricatus Rehder & E.H. Wilson    Cotoneaster divaricatus in… | Spreading cotoneaster ⓘ | Introduced, Naturalization unknown | | Orange only NYFA: Xu | USDA-X0 Images Wikispecies |
| Genus Cotoneaster (excluded taxa) | Cotoneaster | N.Y. Status | Images | Distribution | NY NPT |
| Cotoneaster silvestrii Pamp.1910. Cotoneaster silvestrii Pamp.1912. C. hupehensis Rehder & E.H.Wilson1917. C. racemiflorusvar. veitchiiRehder & E.H.Wilson1957. C. veitchii G.Klotz1963. C. kangtinensis G.Klotz | Hupeh cotoneaster, Many-flowered cotoneaster, Redbead Cotoneaster | Introduced from  China, N.Y. excluded | | NYFA-Excluded | USDA-X0 ARS-GRIN ITIS Images Wikispecies |
| Cotoneaster adpressus Boiss.1904. Cotoneaster adpressa Bois | Creeping cotoneaster, Cotonéaster rampant | Introduced from  Asia, N.Y. excluded | | NYFA-Excluded | USDA-X0 ARS-GRIN ITIS Images Wikispecies |