# Flora of New York/Lycophytes

| ← Dipsacales | Flora of New York — Lycophytes | Flora of New York — Lycophytes | Flora of New York — Lycophytes | Flora of New York — Lycophytes | → Pre-polypod ferns |
| ← Dipsacales | Table of contents | Genus index | Protected species index | Invasive species index | → Pre-polypod ferns |
* Class Lycopodiopsida contains the lycophytes (clubmosses, firmosses and quillworts).
* tracheophytes
* lycophytes
* Lycopodiales : Lycopodiaceae
* Lycopodielloideae
* Lycopodiella spp. (5 bog cubmosses)
* Pseudolycopodiella caroliniana (slender bog clubmoss)
* Lycopodioideae
* Dendrolycopodium spp. (3 tree cubmosses)
* Diphasiastrum spp. (8 ground cedars)
* Spinulum spp. (3 bristly clubmosses)
* Lycopodium spp. (2 running clubmosses)
* Huperzioideae
* Huperzia spp. (4 firmosses)
* Isoetales : Isoetaceae
* Isoetes spp. (8 quillworts)
* Selaginellales : Selaginellaceae
* Selaginella spp. (3 spikemosses)
* euphyllophytes
* Polypodiopsida (ferns)
* spermatophytes (seed plants)
* ↑ P.F. Stevens, (2001-2015). "Ferns s.l." Angiosperm Phylogeny Website. Page last updated: 03/11/2021
* ↑ PPG I (2016), A community‐derived classification for extant lycophytes and ferns. Jnl of Sytematics Evolution, 54: 563-603. https://doi.org/10.1111/jse.12229
| Class | Order | Family | Subfamily | Genus | Common names | # |
| Lycopodiopsida (lycophytes) | Lycopodiales | Lycopodiaceae | Lycopodielloideae | Lycopodiella | bog clubmoss | 5 |
| Lycopodiopsida (lycophytes) | Lycopodiales | Lycopodiaceae | Lycopodielloideae | Pseudolycopodiella | false bog clubmoss | 1 |
| Lycopodiopsida (lycophytes) | Lycopodiales | Lycopodiaceae | Lycopodioideae | Lycopodium | clubmoss | 2 |
| Lycopodiopsida (lycophytes) | Lycopodiales | Lycopodiaceae | Lycopodioideae | Dendrolycopodium | tree clubmoss | 3 |
| Lycopodiopsida (lycophytes) | Lycopodiales | Lycopodiaceae | Lycopodioideae | Spinulum | interrupted clubmoss | 3 |
| Lycopodiopsida (lycophytes) | Lycopodiales | Lycopodiaceae | Lycopodioideae | Diphasiastrum | ground cedar | 7 |
| Lycopodiopsida (lycophytes) | Lycopodiales | Lycopodiaceae | Huperzioideae | Huperzia | firmoss | 5 |
| Lycopodiopsida (lycophytes) | Isoetales | Isoetaceae | — | Isoetes | quillwort | 8 |
| Lycopodiopsida (lycophytes) | Selaginellales | Selaginellaceae | — | Selaginella | spikemoss | 3 |

## Order Lycopodiales

The Lycopodiales ...

### Family Lycopodiaceae

| Lycopodiaceae   Huperzioideae (firmoss)   Huperzia lucidula (shining firmoss)     Huperzia appressa (mountain firmoss)     Huperzia selago (northern firmoss)     Huperzia × bartleyi (Bartley’s firmoss)         Lycopodielloideae bog clubmoss   Lycopodiella inundata (northern bog-clubmoss)     Lycopodiella appressa (southern bog-clubmoss)     Lycopodiella alopecuroides (foxtail bog-clubmoss)     slender bog-clubmoss Pseudolycopodiella caroliniana (Carolina clubmoss)     Lycopodioideae running clubmoss   Lycopodium clavatum (staghorn clubmoss)     Lycopodium lagopus (one-cone clubmoss)     tree clubmoss   Dendrolycopodium obscurum (flat-branched tree clubmoss)     Dendrolycopodium dendroideum (prickly tree clubmoss)     Dendrolycopodium hickeyi (Hickey’s tree clubmoss)     bristly clubmoss   Spinulum annotinum (common bristly clubmoss)     Spinulum canadense (northern bristly clubmoss)     groundcedar   Diphasiastrum digitatum (southern ground cedar)     Diphasiastrum tristachyum (blue ground cedar)     Diphasiastrum complanatum (northern ground cedar)     Diphasiastrum sitchense (Sitka ground cedar) | clumping (firmoss) creeping (clubmoss) |

#### Subfamily Huperzioideae

* The Huperzioideae, known as firmosses, have also been treated as a separate family Huperziaceae. Firmosses differ from other Lycopodiaceae in that they grow in clumps, instead of producing creeping, horizontal stems like other family members. Of the three genera in this group, Huperzia is a temperate and arctic genus and is the only one found naturally in New York. Of the other two genera, Phlegmariurus is native to tropical and subtropical regions, and Phylloglossum is native to Australia and New Zealand.
* ↑ "Family: Huperziaceae - firmoss family" Go Botany 3.5, Native Plant Trust, 2021

##### Huperzia

* Huperzia lucidula, shining firmossCommonly known as firmoss or fir-clubmoss, Huperzia (hoo-per-zee-uh) was named for Johann Peter Huperz, an early 19th century German botanist who specialized in ferns.
* The rare Appalachian firmoss was published as Huperzia appalachiana in 1992, but authorities, including Weakly (2023) consider it to be indistinct from the previously published Huperzia appressa.
* ↑ Weakley, A.S., and Southeastern Flora Team. 2023. Flora of the southeastern United States. University of North Carolina Herbarium, North Carolina Botanical Garden, Chapel Hill, U.S.A. Dec 23, 2023.
Lycopodiales — Lycopodiaceae — Huperzioideae — Huperzia :
| Genus Huperzia Bernh. (1801) | Firmoss | N.Y. Status | Distribution | NY NPT |
| Huperzia lucidula (Michx.) Trevis.1803. Lycopodium lucidulum  Michx. 1875. Huperzia lucidula  (Michx.) Trevis. 1903. Huperzia selago  ssp. lucidulum  Clute 1937. Lycopodium lucidulum  var. verum  Wherry 1939. Urostachys lucidulus  (Michx.) Herter ex Nessel 1939. Urostachys lucidulus  var. reclinatus  (Dup) Nessel 1965. Huperzia selago  ssp. lucidula  (Michx.) Á.Löve & D.Löve 1967. Lycopodium lucidulum  var. tryonii  Mohlenbr. 1999. Huperzia lucidula  var. tryonii  (Mohlenbr.) Mohlenbr. | Shining firmoss, Shining clubmoss, Shining fir-clubmossLycopode brillant, Huperzie brillant | Native, CoC: 5, Secure, S5, G5 FAC-FACW Perennial, Subshrub | NYFA: 5 iNat | USDA-NN POWO FSUS Go Botany VASCAN ARS-GRIN ITIS FNAFNA Tropicos NatureServe IPN Images Wikispecies |
| Huperzia appressa (Bach.Pyl. ex Desv.) Á.Löve & D.Löve1827. Lycopodium selago  var. appressum  Bach.Pyl. ex Desv. 1902. Lycopodium selago  f. appressum  (Bach.Pyl. ex Desv.) Gelert 1930. Lycopodium appressum  (Bach.Pyl. ex Desv.) Petrov in… 1939. Urostachys selago  var. appressus  (Bach.Pyl. ex Desv.) Herter ex Nessel  in… 1956. Lycopodium selago  ssp. appressa  (Bach.Pyl. ex Desv.) D.Löve 1961. Huperzia appressa  (Bach.Pyl. ex Desv.) Á.Löve & D.Löve in… 1968. Lycopodium selago  ssp. appressum  (Bach.Pyl. ex Desv.) Hultén 1981. Huperzia selago  var. appressa  (Bach.Pyl. ex Desv.) Ching 1992. Huperzia appalachiana Beitel & Mickel in… | Appalachian firmoss, Appalachian clubmoss, Appalachian fir-clubmoss, Mountain firmoss, Mountain fir-clubmossLycopode des Appalaches | Native, CoC: 9, Rare, S3, G4-G5, NYNHP: 3 FACU Perennial, Subshrub | NYFA: 3 NYFA: Essex, Greene iNat GBIF Essex County | USDA-NN COL WFO POWO FSUS Go Botany VASCAN ITIS FNAFNA NatureServe IPN Images Wikispecies |
| Huperzia selago (L.) Bernh. ex Schrank & Mart.1753. Lycopodium selago  L. 1779. Lycopodium densum  Lam. (nom. superfl.) 1805. Plananthus selago  (L.) P.Beauv. 1819. Plananthus fastigiatus  Opiz 1821. Lycopodium abietiforme  Gray 1829. Huperzia selago  (L.) Bernh. ex Schrank & Mart. 1831. Lycopodium selago  var. densum  Wallr. 1866. Selago vulgaris  Schur 1923. Urostachys selago  (L.) Herter 1942. Lycopodium selago  var. typicum  Wherry (not validly publ.) 1955. Mirmau selago  H.P.Fuchs 1997. Huperzia acuminata  (Sugim.) Satou | Northern fir-moss, Fir clubmossLycopode sélagine, Huperzie sélagine | Native, CoC: 10, Endangered, NYNHP: 1 | NYFA: 1 NYFA: 5 counties iNat 2 counties | USDA-NN FSUS Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos NatureServe IPN Images Wikispecies |
| Huperzia porophila (F.E.Lloyd & Underw.) Holub1900. Lycopodium porophilum  F.E.Lloyd & Underw. 1903. Lycopodium selago  var. porophilum  Clute 1905. Lycopodium lucidulum  var. porophilum  (F.E.Lloyd & Underw.) Clute 1908. Lycopodium lucidulum  f. porophilum  (F.E.Lloyd & Underw.) Clute 1923. Urostachys porophilus  (F.E.Lloyd & Underw.) Herter 1939. Urostachys lucidulus  var. porophilus  (F.E.Lloyd & Underw.) Nessel 1965. Huperzia selago  var. porophila  (F.E.Lloyd & Underw.) Á.Löve & D.Löve 1985. Huperzia porophila  (F.E.Lloyd & Underw.) Holub | Rock clubmoss, Rock firmoss, Lloyd's clubmossLycopode porophile | N. America native, N.Y. excluded | NYFA-Excluded NYFA: Excluded iNat Carbon County, PA | USDA-NN POWO FSUS VASCAN FNA Tropicos Images Wikispecies |
| Huperzia occidentalis (Clute) Kartesz & Gandhi1903. Lycopodium lucidulum  f. occidentale  Clute 1903. Lycopodium lucidulum  f. occidentalis  Clute 1932. Lycopodium lucidulum  var. occidentale  (Clute) L.R.Wilson 1966. Lycopodium selago  f. occidentale  (Clute) B.Boivin 1966. Lycopodium selago  var. occidentale  (Clute) B.Boivin 1967. Lycopodium lucidulum  var. tryonii  Mohlenbr. 1991. Huperzia occidentalis  (Clute) Kartesz & Gandhi 1992. Huperzia occidentalis  (Clute) Beitel (nom. illeg.) | Pacific Clubmoss ⓘ Western Clubmoss ⓘ Lycopode de l'Ouest ⓘ | N. America native  Northwest, N.Y. excluded Perennial, Herb | NYFA-Excluded NYFA: Excluded iNat GBIF Northwest N.A. only | USDA-NN POWO ITIS FNA Tropicos NatureServe BONAP Images Wikispecies |
| ↑ New York Natural Heritage Program. 2024. Online Conservation Guide for Huperzia appressa. Rare, S3/G5. ↑ New York Natural Heritage Program. 2024. Online Conservation Guide for Huperzia selago. Endangered, S1/G5. ||||||

##### Huperzia hybrids

Lycopodiales — Lycopodiaceae — Huperzioideae — Huperzia :
| Genus Huperzia Bernh. (1801) | Firmoss | N.Y. Status | Distribution | NY NPT |
| Huperzia × bartleyi (Cusick) Holub  Huperzia lucidula ×   Huperzia porophila1987. Lycopodium × bartleyi  Cusick 1991. Huperzia × bartleyi  (Cusick) Holub 1991. Huperzia × bartleyi  (Cusick) Kartesz & Gandhi | Bartley's clubmoss | Native, Unranked | NYFA: U NYFA: Ulster County iNat No NY obsevations | USDA-N0 POWO ITIS Tropicos Images Wikispecies |
| Huperzia × buttersii (Abbe) Kartesz & Gandhi  Huperzia lucidula ×   Huperzia selago1953. Lycopodium × buttersii  Abbe 1991. Huperzia × buttersii  Kartesz & Gandhi | Butters' club-moss, Butters' firmossLycopode de Butters | N. America native, N.Y. excluded | NYFA-Excluded iNat | USDA-NN POWO Go Botany VASCAN ITIS FNA Tropicos NatureServe Images Wikispecies |
| Huperzia × protoporophila A.Haines  Huperzia appressa ×   Huperzia lucidula2003. Huperzia × protoporophila  A.Haines | Hybrid rock firmoss | N. America native, No N.Y. reports | NYFA: D iNat | Go Botany VASCAN Tropicos IPN Images Wikispecies |
| Huperzia × josephbeitelii A.Haines  Huperzia appressa ×   Huperzia selago2003. Huperzia × beitelii  A.Haines 2003. Huperzia × josephbeitelii  A.Haines | Beitel's firmoss, Joseph Beitel's clubmoss | N. America native, No N.Y. reports | NYFA: D iNat | USDA-NN Go Botany VASCAN Tropicos IPN Images Wikispecies |

#### Subfamily Lycopodioideae
##### Lycopodium

* Running clubmoss The genus name Lycopodium (lī-kə-ˈpō-dē-əm) was derived from the Greek lukos (wolf) and podion (small foot). It is commonly known as running clubmoss.
* Also treated as Lycopodium sect. Lycopodium.
Lycopodiales — Lycopodiaceae — Lycopodioideae — Lycopodium :
| Genus Lycopodium | Running clubmoss | N.Y. Status | Images | Distribution | NY NPT |
| Lycopodium clavatum L.1753. Lycopodium clavatum  L. 1805. Lepidotis clavata  (L.) P.Beauv. 1805. Lepidotis ciliata  P.Beauv. 1805. Lepidotis inflexa  P.Beauv. 1810. Lepidotis aristatum  Humb. & Bonpl. ex Willd. 1925. Lycopodium clavatum  var. laurentianum  Vict. 1925. Lycopodium clavatum  var. subremotum  Vict. | Common clubmoss, Running clubmoss, Running pine, Running cedar, Staghorn clubmoss, Elk mossLycopode à massue | Native, CoC: 5, Secure | | NYFA: 5 iNat | USDA-NN POWO Go Botany ARS-GRIN ITIS FNAFNA Images Wikispecies |
| Lycopodium lagopus (Laest. ex Hartm.) Zinserl. ex Kuzen1831. Lycopodium clavatum  var. monostachyon  Hook. & Grev. 1838. Lycopodium clavatum  var. integrifolium  (Hook. ex Goldie) Spring 1842. Lycopodium clavatum  var. integerrimum  Spring 1858. Lycopodium clavatum  var. lagopus  Laest. ex C.Hartm. 1901. Lycopodium clavatum  var. brevispicatum  Peck 1910. Lycopodium clavatum  var. megastachyon  Fernald & Bissell 1925. Lycopodium clavatum  f. furcatum  Vict. 1950. Lycopodium clavatum  ssp. monostachyon  (Hook. & Grev.) Selander 1953. Lycopodium lagopus  (Laest. ex C.Hartm.) Zinserl. ex Kuzen 1958. Lycopodium clavatum  ssp. megastachyon  (Fernald & Bissell) Á.Löve & D.Löve | One-cone clubmoss, One-cone ground-pine, Running pine, Running cedar, Arctic stag's-horn clubmossLycopode patte-de-lapin | Native, CoC: 4, Likely secure | | NYFA: 4 iNat | USDA-NN POWO Go Botany ITIS FNAFNA Images Wikispecies |

##### Dendrolycopodium

* Tree clubmoss From Greek δενδρος (denros: tree), λυκος (lukos: wolf), and ποδιων (podion: legs), Dendrolycopodium (den-drō-lī-kə-ˈpō-dē-əm) contains tree-clubmoss species, which are fairly erect and have branched stems. This genus, also treated as Lycopodium sect. Obscura, is characterized by stalked terminal strobili, which contain their sporangia.
Lycopodiales — Lycopodiaceae — Lycopodioideae — Dendrolycopodium :
| Genus Dendrolycopodium | Tree clubmoss | N.Y. Status | Distribution | NY NPT |
| Dendrolycopodium obscurum (L.) A.Haines1753. Lycopodium obscurum  L. 1833. Lycopodium dendroideum  var. obscurum  (L.) Torr. ex Beck 1937. Lycopodium obscurum  var. genuinum  Wherry 2003. Dendrolycopodium obscurum  (L.) A.Haines | Ground pine, Princess-pine, Eastern tree clubmoss, Rare clubmoss, Flat-branch tree clubmossLycopode obscur | Native, CoC: 5, Secure FACU | NYFA: 5 iNat | USDA-NN Go Botany ARS-GRIN ITIS FNAFNA Images Wikispecies |
| Dendrolycopodium dendroideum (Michx.) A.Haines1803. Lycopodium dendroideum  Michx. 1890. Lycopodium obscurum  var. dendroideum  D.C.Eaton 1916. Lycopodium obscurum  var. hybridum  Farw. 2003. Dendrolycopodium dendroideum  (Michx.) A. Haines | Prickly tree clubmoss, Northern tree clubmoss, Treelike clubmoss, Prickle tree club-mossLycopode dendroïde | Native, CoC: 5, Secure FACU | NYFA: 5 iNat | USDA-NN Go Botany ARS-GRIN ITIS FNAFNA Images Wikispecies |
| Dendrolycopodium hickeyi (W.H.Wagner, Beitel, & Moran) A.Haines1977. Lycopodium obscurum  var. isophyllum  Hickey 1989. Lycopodium hickeyi  W.H.Wagner, Beitel & R.C.Moran 2003. Dendrolycopodium hickeyi  (W.H.Wagner, Beitel, & Moran) A.Haines | Pennsylvania clubmoss, Hickey's tree club-moss | Native, CoC: 5, Secure Perennial, Herb | NYFA: 5 iNat | USDA-NN Go Botany ITIS FNAFNA Images Wikispecies |

##### Spinulum

* Spinulum annotinumSpinulum (ˈspīn-yə-ləm) is also treated as Lycopodium sect. Annotina.
Lycopodiales — Lycopodiaceae — Lycopodioideae — Spinulum :
| Genus Spinulum | Bristly clubmoss | N.Y. Status | Distribution | NY NPT |
| Spinulum annotinum (L.) A.Haines1753. Lycopodium annotinum  L. 1772. Lycopodium dubium  Zoëga 2003. Spinulum annotinum  (L.) A.Haines | Common bristly clubmoss, Common stiff clubmoss, Interrupted clubmossLycopode interrompu | Native, CoC: 5, Secure | NYFA: 5 iNat | USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNA Images Wikispecies |
| Spinulum canadense (Nessel) A.Haines1772. Lycopodium dubium  auct. non Zoëga 1834. Lycopodium pungens  (Desv. ex Spring) Bach.Pyl. ex Iljin 1842. Lycopodium annotinum  var. pungens  Desv. 1842. Lycopodium pungens  La Pylaie ex Desv. 1843. Lycopodium annotinum  var. montanum  Tuck. 1939. Lycopodium canadense  Romhild ex Nessel 1968. Lycopodium annotinum  ssp. pungens  Hultén 1969. Lycopodium annotinum  ssp. dubium  (Zoëga) Kallio, Laine & Y.Mäkinen 2003. Spinulum canadense  (Romhild ex Nessel) A.Haines | Northern bristly clubmoss, Northern stiff clubmoss, Northern interrupted clubmoss | Native, CoC: 10, Endangered | NYFA: 1-2 iNat | Go Botany VASCAN Tropicos Images Wikispecies |
| Spinulum annotinum × canadense  Spinulum annotinum ×   Spinulum canadense | Hybrid bristly clubmoss | Native, Unranked | NYFA: U | Images Wikispecies |

##### Diphasiastrum

* Ground cedar (genus Diphasiastrum) is also treated as Lycopodium sect. Complanata.
Lycopodiales — Lycopodiaceae — Lycopodioideae — Diphasiastrum :
| Genus Diphasiastrum Holub (1975) | Ground-cedar | N.Y. Status | Distribution | NY NPT |
| Diphasiastrum digitatum (Dill. ex A.Braun) Holub1848. Lycopodium digitatum  Dill. ex A.Braun 1901. Lycopodium complanatum  var. flabelliforme  Fernald 1911. Lycopodium flabelliforme  (Fernald) Blanch. 1965. Diphasium complanatum  ssp. flabelliforme   in… 1975. Diphasiastrum digitatum (Dill. ex A.Braun) Holub | Southern groundcedar, Southern ground-cedar, Fan club-moss, Southern running-pineLycopode en éventail, Courants verts, Lycopode aplati variété en éventail | Native, CoC: 4, Secure, S5, G5 | NYFA: 5 iNat | USDA-NN Go Botany VASCAN ARS-GRIN ITIS FNAFNA Tropicos IPN Images Wikispecies |
| Diphasiastrum tristachyum (Pursh) Holub1814. Lycopodium tristachyum  Pursh in… 1840. Lycopodium clavatum  var. tristachyum  (Pursh) Hook.  in… 1848. Lycopodium complanatum  ssp. tristachyum  (Pursh) Dostál  in… 1859. Lycopodium complanatum  ssp. chamaecyparissus   in… 1905. Lycopodium complanatum  var. tristachyon  Clute  in… 1944. Diphasium tristachyum  (Pursh) Rothm. in… 1961. Diphasium chamaecyparissus  in… 1975. Diphasiastrum tristachyum (Pursh) Holub in… | Blue ground-cedar, Deep-root clubmoss | Native, CoC: 4, Secure, S5, G5 | NYFA: 5 iNat | USDA-NN POWO Go Botany ITIS FNAFNA Tropicos Images Wikispecies |
| Diphasiastrum complanatum (L.) Holub1753. Lycopodium complanatum  L. in… 1805. Lepidotis complanata  (L.) P.Beauv. in… 1840. Lycopodium anceps  Wallr. in… 1944. Diphasium complanatum  (L.) Rothm. in… 1958. Diphasium anceps  (Asch.) Á.Löve & D.Löve in… 1975. Diphasiastrum complanatum (L.) Holub in… | Northern groundcedar, Trailing clubmoss, Northern running-pine, Lycopode aplati | Native, CoC: 8, Endangered, S2, G5 | NYFA: 1 iNat | USDA-NN Go Botany ARS-GRIN ITIS FNAFNA Images Wikispecies |
| Diphasiastrum sitchense (Rupr.) Holub (1975)1845. Lycopodium sitchense  Rupr. in… 1916. Lycopodium complanatum  var. sitchense   in… 1923. Lycopodium × sabinifolium  var. sitchense   in… 1965. Lycopodium × sabinifolium  ssp. sitchense   in… 1965. Diphasium sitchense  (Rupr.) Á.Löve & D.Löve in… 1975. Diphasiastrum sitchense  (Rupr.) Holub in… | Sitka groundcedar, Alaskan clubmoss, Sitka club-moss, Lycopode de Sitka | Native, CoC: 10, Endangered, S1, G5 | NYFA: 1 iNat | USDA-NN Go Botany ARS-GRIN ITIS FNA Images Wikispecies |
| Diphasiastrum × sabinifolium (Willd.) Holub (1975)  Diphasiastrum sitchense ×   Diphasiastrum tristachyum    Lycopodium × sabinifolium Willd.     Lycopodium sabinifolium Willd.     Lycopodium armatum Desv. | Ground-fir, Savinleaf groundpine, Savin-leaved groundpine, Cypress clubmoss | Native, Unranked, SNA, GNA | NYFA: 3 iNat | USDA-NN ARS-GRIN ITIS FNA Images Wikispecies |
| Diphasiastrum × zeilleri (Rouy) Holub (1975)  Diphasiastrum complanatum ×   Diphasiastrum tristachyum    Lycopodium × zeilleri     Diphasium zeilleri (Rouy) Damboldt | Zeiller's running-pine, Zeiller's clubmoss | Native, Unranked, SNA, GNA | NYFA: 2 iNat | USDA-NN ITIS FNA Images Wikispecies |
| Diphasiastrum × habereri (House) Holub (1975)  Diphasiastrum digitatum ×   Diphasiastrum tristachyum    Lycopodium × habereri House     Lycopodium complanatum  var. habereri       Lycopodium tristachyum  var. habereri | Haberer's clubmoss | Native, Unranked, SNA, GNA | NYFA: U iNat | USDA-NN ITIS FNA Images Wikispecies |

#### Subfamily Lycopodielloideae
##### Lycopodiella

Bog clubmoss. (Lycopodiella in Latin means little wolf's-foot.)
Lycopodiales — Lycopodiaceae — Lycopodielloideae — Lycopodiella :
| Genus Lycopodiella Holub (1964) | Bog clubmoss | N.Y. Status | Images | Distribution | NY NPT |
| Lycopodiella inundata (L.) Holub1753. Lycopodium inundatum L. 1805. Plananthus inundatus (L.) P.Beauv. 1852. Lepidotis inundata (L.) Opiz 1912. Lepidotis inundata (L.) Börner 1964. Lycopodiella inundata (L.) Holub | Northern bog clubmoss, Marsh clubmoss, Inundated clubmossLycopodielle inondé | Native, CoC: 8, Secure OBL Perennial, Subshrub | | NYFA: 5 | USDA-NN Go Botany ARS-GRIN ITIS FNA Tropicos Images Wikispecies |
| Lycopodiella appressa (Chapm.) Cranfill1843. Lycopodium inundatum  var. bigelovii  Tuck. 1878. Lycopodium inundatum  var. appressum  Chapm. 1900. Lycopodium appressum F.E.Lloyd & Underw. 1981. Lycopodiella appressa (Chapm.) Cranfill 1985. Lycopodiella bigelowii (Tuck.) Holub | Southern bog clubmoss, Appressed bog-clubmoss, Chapman's club-moss | Native, CoC: 7, Likely secure FACW Perennial, Subshrub | | NYFA: 4 | USDA-NN Go Botany ITIS FNA Images Wikispecies |
| Lycopodiella alopecuroides (L.) Cranfill1753. Lycopodium alopecuroides L. 1944. Lepidotis alopecuroides (L.) Rothm. 1981. Lycopodiella alopecuroides (L.) Cranfill | Foxtail bog-clubmoss, Foxtail clubmoss | Native, CoC: 7, Vulnerable, S2?, G2 FACW Perennial, Subshrub | | Kings, Nassau, Richmond, Suffolk NYFA: 3 | USDA-N0 Go Botany ITIS FNA Images Wikispecies |
| Lycopodiella × copelandii (Eiger) Cranfill (1981)  L. alopecuroides ×   L. appressa | Copeland's bog clubmoss, Copeland's clubmoss | Native, Unranked | | Suffolk NYFA: U | USDA-N0 ITIS Images Wikispecies |
| Lycopodiella × gilmanii A.Haines (2003)  L. appressa ×   L. inundata | Gilman's bog clubmoss, Gilman's clubmoss | Native, Unranked | | NYFA: U | ITIS Images Wikispecies |
| Genus Lycopodiella (excluded taxa) | Bog clubmoss | N.Y. Status | Images | Distribution | NY NPT |
| Lycopodiella × robusta ( R.J.Eaton) A.Haines (2002)  L. alopecuroides ×   L. inundataLycopodium inundatum var. robustum | Bog clubmoss | Native, N.Y. excluded | | NYFA: U | Images Wikispecies |

##### Pseudolycopodiella

Lycopodiales — Lycopodiaceae — Lycopodielloideae — Pseudolycopodiella :
| Genus Pseudolycopodiella (L.) Holub | False bog-clubmoss | N.Y. Status | Images | Distribution | NY NPT |
| Pseudolycopodiella caroliniana (L.) Holub1753. Lycopodium carolinianum L. 1968. Lycopodiella caroliniana (L.) Pic.Serm. 1983. Pseudolycopodiella caroliniana Holub 1983. Pseudolycopodiella meridionalis Holub | Carolina clubmoss, Slender clubmoss, Slender bog club-moss | Native, CoC: 9, Endangered, NYNHP: 1 | | NYFA: 1 | USDA-N0 Go Botany ARS-GRIN ITIS FNA Tropicos Images Wikispecies |
| ↑ New York Natural Heritage Program. 2024. Online Conservation Guide for Pseudolycopodiella caroliniana. Endangered, S1/G5T4. ||||||

## Order Selaginellales
### Family Selaginellaceae
#### Selaginella

* Three spikemoss (Selaginella) species have been found growing wild in New York.
Selaginellales — Selaginellaceae — Selaginella :
| Genus Selaginella | Spikemoss | N.Y. Status | Images | Distribution | NY NPT |
| Selaginella rupestris (L.) Spring1753. Lycopodium rupestre  L. in… 1803. Stachygynandrum rupestre  (L.) P.Beauv. in… 1838. Selaginella rupestris  (L.) Spring in… 1849. Selaginella rupestris  var. borealis  Spring  in… 1891. Lycopodioides rupestris  (L.) Kuntze in… 1900. Selaginella bourgeaui  Hieron. in… 1908. Selaginella rupestris  f. borealis  (Spring) Clute  ... 1992. Bryodesma rupestre  (L.) Soják in… | Rock spikemoss, Dwarf spikemoss, Northern selaginellaSélaginelle des rochers, Sélaginelle rupestre | Native, CoC: 9, Likely secure, S4 UPL Perennial, Herb | | NYFA-4 iNat | NRCS-NN POWO Go Botany ITIS FNA Images Wikispecies |
| Selaginella apoda (L.) Spring1753. Lycopodium apodum L. 1840. Selaginella apus Spring 1840. Selaginella apoda (L.) Spring | Meadow spikemoss, Creeping spikemossSélaginelle apode | Native, CoC: 6, Unranked, Under review FACW | | NYFA-2? iNat | NRCS-N0 Go Botany ITIS FNA Images Wikispecies |
| Selaginella eclipes W.R.Buck1977. Selaginella eclipes W.R.Buck 1996. Selaginella apoda  ssp. eclipes  Skoda | Hidden spikemoss, Buck's meadow spikemoss, Creeping spikemoss | Native, CoC: 8, Endangered FACW | | NYFA-1 | NRCS-NN Go Botany ITIS FNA Tropicos Images Wikispecies |
| Genus Selaginella (excluded taxa) | Spikemoss | N.Y. Status | Images | Distribution | NY NPT |
| Selaginella selaginoides (L.) P. Beauv. ex Mart. & Schrank1753. Lycopodium selaginoides L. 1805. S. spinosa P.Beauv. illeg. 1841. S. selaginoides (L.) P.Beauv. ex Mart. & Schrank | Club spikemoss ⓘ Northern spikemoss ⓘ Northern spike-moss ⓘ Prickly mountain-moss ⓘ Lesser-club-moss ⓘ | N. America native, N.Y. excluded FACW Perennial, Herb | | NYFA-Excluded | USDA-NN Go Botany ARS-GRIN Images Wikispecies |
| ↑ 2019 Rare Plant Status Lists – Review List - New York Natural Heritage Program ||||||

## Order Isoetales
### Family Isoetaceae
#### Isoetes

* Isoetes echinosporaNorth American Isoetes lacustris plants have recently been segregated as Isoetes macrospora but can't be easily differentiated except by geography.
* ↑ W. Carl Taylor, Neil T. Luebke, Donald M. Britton, R. James Hickey, Daniel F. Brunton (2020). FNA Volume 2. Flora of North America Association
Isoetales — Isoetaceae — Isoetes :
| Genus Isoetes L. | Quillwort | N.Y. Status | Distribution | NY NPT |
| Isoetes engelmannii A.Braun1846. Isoetes engelmannii  A.Braun in… 1846. Isoetes lacustris  var. microspora  Engelm. in… 1867. Isoetes engelmannii  var. gracilis  Engelm. in… 1891. Calamaria engelmannii  (A.Braun) Kuntze in… 1905. Isoetes engelmannii  var. fontana  A.A.Eaton  in… 1994. Isoetes × fairbrothersii  J.D.Montgom. & W.C.Taylor in… | Engelmann's quillwort, Appalachian quillwortIsoète d'Engelmann | Native, CoC: 6, Secure, S5 OBL Perennial, Herb, Subshrub, diploid (2n = 22) | NYFA: 5 NYFA: 23 counties iNat GBIF Cortland (2019) | USDA-NN COL WFO POWO FSUS Go Botany VASCAN ITIS FNAFNA Tropicos NatureServe BONAP IPN Images Wikispecies |
| Isoetes echinospora Durieu ssp. muricata A.Löve & D.Löve1789. Isoetes setacea  Lam. p.p. 1827. Isoetes tenella  Léman ex Desv. POWO in… 1864. Isoetes braunii  Durieu in… 1867. Isoetes echinospora  var. braunii  Engelm. 1945. Isoetes muricata  var. braunii  (Engelm.) C.F.Reed  in… 1961. Isoetes echinospora  ssp. muricata  (Durieu) Á.Löve & D.Löve NYFA | Spiny-spore quillwort, Spring quillwort, Braun's quillwort, Bristlelike quillwortIsoète à spores épineuses, Isoète à spores spinuleuses | Native, CoC: 7, Likely secure, S4 OBL Perennial, Herb, diploid (2n = 22) | NYFA: 4 | USDA-? Go Botany VASCAN ARS-GRIN ITIS FNA Tropicos Images Wikispecies |
| Isoetes tuckermanii A.Braun ex Engelm.1867. I. tuckermanii A.Braun ex Engelm. 1900. I. tuckermanii  var. borealis  A.A.Eaton 1981. I. acadiensis Kott 1991. I. tuckermanii  ssp. acadiensis  Taylor & Luebke | Tuckerman's quillwortIsoète de Tuckerman | Native, CoC: 10, Likely secure, tetraploid (2n = 44) | NYFA: 4 | USDA-NN FSUS Go Botany ITIS FNA Images Wikispecies |
| Isoetes × eatonii R.Dodge  Isoetes echinospora ×   Isoetes engelmannii1896. Isoetes eatonii R.Dodge 1900. Isoetes gravesii A.A.Eaton     Isoetes eatonii  var. gravesii  A.A.Eaton | Eaton's quillwort | Native, Vulnerable | NYFA: 3 | USDA-NN ITIS Images Wikispecies |
| Isoetes macrospora Durieu1753. Isoetes lacustris  L. in… 1864. Isoetes macrospora  Durieu in… 1900. Isoetes hieroglyphica  A.A.Eaton 1922. Isoetes macrospora  f. hieroglyphica  N.Pfeiff.  ... | Big-spore quillwort ⓘ Lake quillwort Carved quillwort Warty Quillwort Isoète lacustre Isoète hiéroglyphique | Native, CoC: 10, Rare, S3 OBL Perennial, Herb, decaploid (2n = 110) | NYFA: 3 NYFA: 5 counties iNat GBIF St. Lawrence only | USDA-NN Go Botany ITIS FNAFNA Images Wikispecies |
| Isoetes septentrionalis D.F.Brunt.1846. Isoetes riparia  var. canadensis  Engelm.  in… 1903. Isoetes canadensis  var. robbinsii  A.A. Eaton 2015. Isoetes septentrionalis  D.F. Brunt. in… | Northern shore quillwort, Northern quillwortisoète du Nord | Native, CoC: 10, Endangered OBL Perennial, Herb, tetraploid (2n = 44) | NYFA: 1 | USDA-N0 FSUS Go Botany VASCAN ITIS FNA Images Wikispecies |
| Isoetes × robusta (Engelm.) D.F.Brunt.  Isoetes engelmannii ×   Isoetes septentrionalis1898. Isoetes × dodgei A.A.Eaton  auct. Isoetes × brittonii  auct. Isoetes engelmannii × riparia | Hybrid of  Engelmann's quillwort &  northern shore quillwort | Native, No recent reports | NYFA: Z? | USDA-N0 Go Botany ARS-GRIN ITIS Images Wikispecies |
| ↑ The New York Flora Atlas and New England Wild Flower Society list the spiny-spore quillwort as Isoëtes echinospora ssp. muricata because the leaves of the North American plants of bear stomata, while those of the European plants do not. The other sources used here treat ssp. muricata as a synonym. ||||||

#### Isoetes (excluded)

Isoetales — Isoetaceae — Isoetes :
| Genus Isoetes (excluded taxa) | Quillwort | N.Y. Status | Distribution | NY NPT |
| Isoetes riparia Engelm. ex A.Braun1846. Isoetes riparia Engelm. ex A.Braun 18??. Isoetes saccharata Engelm. in A.Gray 1891. Calamaria saccharata (Engelm.) Kuntze 1901. Isoetes canadensis (Engelm.) A.A.Eaton 1949. Isoetes riparia  f. saccharata  ( Engelm. ex A.Gray ) Proctor | Shore quillwort, Canada shore quillwort | Native, N.Y. excluded | NYFA-Excluded | USDA-NN Go Botany ITIS Tropicos BONAP Images Wikispecies |
| Isoetes × harveyi A.A.Eaton  Isoetes lacustris ×   Isoetes tuckermanii1900. Isoetes × heterospora A.A.Eaton     Isoetes tuckermanii  var. harveyi  Clute | Harvey's quillwort | Native, N.Y. excluded | NYFA-Excluded | USDA-NN Go Botany ITIS FNA Images Wikispecies |
| Isoetes × novae-angliae D.F. Brunt. & D.M. Britton  Isoetes septentrionalis ×   Isoetes tuckermanii | New England quillwort | Native, N.Y. excluded | NYFA-Excluded | USDA-N0 Go Botany ITIS FNA Images Wikispecies |
| ↑ Isoetes riparia was apparently misapplied to reports of Isoetes septentrionalis in New York. (NYFA (2016) ||||||