# Haplodrassus

Haplodrassus is a genus of ground spiders that was first described by R. V. Chamberlin in 1922. They range from 3 to 10 millimetres (0.12 to 0.39 in). H. signifer is the most widespread species, found across North America except for Alaska and northern Canada.

## Species

As of May 2019 it contains seventy-nine species:
* H. aenus Thaler, 1984 – Switzerland, Austria
* H. alexeevi Ponomarev & Shmatko, 2017 – Russia (Europe)
* H. ambalaensis Gajbe, 1992 – India
* H. atarot Levy, 2004 – Israel
* H. belgeri Ovtsharenko & Marusik, 1988 – Russia (South to north-east Siberia, Far East)
* H. bengalensis Gajbe, 1992 – India
* H. bicornis (Emerton, 1909) – USA, Canada
* H. bohemicus Miller & Buchar, 1977 – Czech Rep., Macedonia, Greece, Ukraine?, Russia (Europe, Caucasus)?
* H. canariensis Schmidt, 1977 – Canary Is.
* H. caspius Ponomarev & Belosludtsev, 2008 – Russia (Europe, Caucasus), Azerbaijan, Iran, Kazakhstan
* H. caucasius Ponomarev & Dvadnenko, 2013 – Caucasus (Russia, Georgia)
* H. chamberlini Platnick & Shadab, 1975 – North America
* H. chotanagpurensis Gajbe, 1987 – India
* H. cognatus (Westring, 1861) – Europe, Russia (Europe to Far East), Caucasus, Kazakhstan, Japan
** Haplodrassus c. ermolajewi Lohmander, 1942 – Russia (West Siberia)
* H. concertor (Simon, 1878) – France
* H. crassipes (Lucas, 1846) – Morocco, Algeria
* H. creticus (Roewer, 1928) – Greece (Crete)
* H. dalmatensis (L. Koch, 1866) – Europe, North Africa, Turkey, Middle East, Russia (Europe) to Central Asia
** Haplodrassus d. pictus (Thorell, 1875) – Spain, Madeira
* H. dentatus Xu & Song, 1987 – China
* H. dentifer Bosmans & Abrous, 2018 – Morocco, Algeria, Tunisia, Spain
* H. deserticola Schmidt & Krause, 1996 – Canary Is.
* H. dixiensis Chamberlin & Woodbury, 1929 – USA
* H. dumdumensis Tikader, 1982 – India
* H. eunis Chamberlin, 1922 – USA, Canada
* H. hatsushibai Kamura, 2007 – Japan
* H. hiemalis (Emerton, 1909) (type) – North America, Russia (Europe to Far East)
* H. huarong Yin & Bao, 2012 – China
* H. hunanensis Yin & Bao, 2012 – China
* H. ibericus Melic, Silva & Barrientos, 2016 – Portugal, Spain
* H. invalidus (O. Pickard-Cambridge, 1872) – Egypt, Cyprus, Turkey, Israel, Azerbaijan, Greece?
* H. ivlievi Ponomarev, 2015 – Russia (Europe)
* H. jacobi Gajbe, 1992 – India
* H. kanenoi Kamura, 1995 – Japan
* H. kulczynskii Lohmander, 1942 – Europe, Turkey, Russia (Europe to Far East), China, Korea
* H. lilliputanus Levy, 2004 – Israel
* H. longivulva Bosmans & Hervé, 2018 – Morocco, Algeria
* H. lyndae Abrous & Bosmans, 2018 – Morocco, Algeria, Spain
* H. macellinus (Thorell, 1871) – France, Italy, Portugal?, Spain?
* H. maculatus (Banks, 1904) – USA, Mexico
* H. mayumiae Kamura, 2007 – Korea, Japan
* H. mediterraneus Levy, 2004 – Turkey, Syria, Lebanon, Israel, Jordan
* H. mimus Chamberlin, 1922 – USA
* H. minor (O. Pickard-Cambridge, 1879) – Europe, Turkey
* H. moderatus (Kulczyński, 1897) – Europe, Russia (Europe to Far East), China
* H. montanus Paik & Sohn, 1984 – Russia (Far East), China, Korea
* H. morosus (O. Pickard-Cambridge, 1872) – Greece, Turkey, Israel
* H. nigroscriptus (Simon, 1909) – Morocco
* H. nojimai Kamura, 2007 – Japan
* H. omissus (O. Pickard-Cambridge, 1872) – Canary Is., Morocco, Mediterranean
* H. orientalis (L. Koch, 1866) – Bulgaria, Greece, Ukraine, Russia (Europe), Kazakhstan
* H. ovatus Bosmans & Hervé, 2018 – Algeria, Tunisia
* H. ovtchinnikovi Ponomarev, 2008 – Turkey, Iran, Kazakhstan
* H. paramecus Zhang, Song & Zhu, 2001 – China
* H. pargongsanensis Paik, 1992 – Korea
* H. ponomarevi Kovblyuk & Seyyar, 2009 – Greece, Turkey
* H. pseudosignifer Marusik, Hippa & Koponen, 1996 – Ukraine, Russia (Europe to Central Asia), Iran
* H. pugnans (Simon, 1880) – Israel, Russia (Europe to Far East), China, Japan
* H. reginae Schmidt & Krause, 1998 – Cape Verde Is.
* H. rhodanicus (Simon, 1914) – Portugal, Spain, France, Italy (Sardinia), Tunisia
* H. rufipes (Lucas, 1846) – Morocco, Algeria, Tunisia, Portugal, Spain, France, Italy
* H. rufus (Savelyeva, 1972) – Kazakhstan
* H. rugosus Tuneva, 2004 – Kazakhstan
* H. sataraensis Tikader & Gajbe, 1977 – India
* H. securifer Bosmans & Abrous, 2018 – Morocco, Algeria, Tunisia, Portugal, Spain, France, Italy, Belgium
* H. signifer (C. L. Koch, 1839) – North America, Europe, North Africa, Turkey, Israel, Caucasus, Russia (Europe to Far East), Central Asia, China, Korea
* H. silvestris (Blackwall, 1833) – Europe, Turkey, Caucasus
* H. soerenseni (Strand, 1900) – Europe, Turkey, Caucasus, Russia (Europe to Far East), Kazakhstan, China
* H. stuxbergi (L. Koch, 1879) – Russia (West to Middle Siberia)
* H. taepaikensis Paik, 1992 – Russia (South Siberia, Far East), Korea
* H. taibo (Chamberlin, 1919) – USA
* H. tegulatus (Schenkel, 1963) – Russia (South Siberia), China
* H. tehriensis Tikader & Gajbe, 1977 – India
* H. triangularis Bosmans, 2018 – Morocco, Tunisia
* H. typhon (Simon, 1878) – Algeria, Tunisia, Portugal, Spain, France, Italy (Sardinia)
* H. umbratilis (L. Koch, 1866) – Europe to Kazakhstan
** Haplodrassus u. gothicus Lohmander, 1942 – Sweden
* H. vastus (Hu, 1989) – China