# Aculus

Aculus is a genus of mites, including the following species:

## Species

* Aculus ablopurpurascus Huang, 2001
* Aculus acanthae (Mohanasundaram, 1982)
* Aculus acraspis (Nalepa, 1892)
* Aculus acutangulae Mohanasundaram, 1985
* Aculus advens (Keifer, 1938)
* Aculus aegerinus (Nalepa, 1892)
* Aculus aesculi Domes, 2003
* Aculus aflatunivagrans (Ponomareva, 1978)
* Aculus africanae (Meyer & Ueckermann, 1990)
* Aculus albopurpurascus Huang, 2001
* Aculus alfalfae (Roivainen, 1950)
* Aculus altus (Nalepa, 1909)
* Aculus amandae (Keifer, 1969)
* Aculus ambrosiae (Keifer, 1943)
* Aculus amygdalina (Banks, 1912)
* Aculus anthobius (Nalepa, 1892)
* Aculus aphanotrichus (Liro, 1943)
* Aculus aphelus Smith-Meyer & Ueckermann, 1990
* Aculus arbosti (Cotte, 1924)
* Aculus arceosae (Briones & McDaniel, 1976)
* Aculus argenteae (Farkas, 1963)
* Aculus arzakanensis (Bagdasarian, 1970)
* Aculus asclepiellus Keifer, 1965
* Aculus asperi (Chandrapatya & Boczek, 2002)
* Aculus asperus Channabasavanna, 1966
* Aculus asteri (Petanovic & Boczek, 2000)
* Aculus atlantazaleae (Keifer, 1940)
* Aculus atturensis Mohanasundaram, 1980
* Aculus aucupariae (Liro, 1943)
* Aculus baccaureae (Farkas, 1960)
* Aculus bacsetae Flechtmann & Davis, 1971
* Aculus balevskii Natcheff, 1966
* Aculus ballei (Nalepa, 1891)
* Aculus bambusae Kuang, 1991
* Aculus bangalorensis Mohanasundaram, 1985
* Aculus beeveri (Manson & Gerson, 1986)
* Aculus betuli (Nalepa, 1897)
* Aculus blagayanae Jocic, Petanovic & Vidovic, 2011
* Aculus brevisetus Carmona, 1972
* Aculus broussaisiae Keifer, 1964
* Aculus broussenetiae Lin-Fuping, Jin-Changle & Kuang-Haiyua, 1997
* Aculus bursifex (Nalepa, 1921)
* Aculus calcarifer (Liro, 1943)
* Aculus calystegiae (Lamb, 1952)
* Aculus campbelli Knihinicki & Boczek, 2003
* Aculus capellae Keifer, 1979
* Aculus capsibaccati (Keifer, 1979)
* Aculus carsonellus (Keifer, 1966)
* Aculus caryfoliae Keifer, 1961
* Aculus cassiae Mondal & Chakrabarti, 1981
* Aculus catappae Boczek & Davis, 1984
* Aculus cauliflorus Boczek & Davis, 1984
* Aculus cayratiae Kuang-Haiyua, 1987
* Aculus cercidii (Keifer, 1965)
* Aculus cercidis (Hall, 1967)
* Aculus chaitus Keifer, 1965
* Aculus chamaespartii Carmona, 1974
* Aculus charniae Smith-Meyer & Ueckermann, 1990
* Aculus chihuahuae (Keifer, 1979)
* Aculus clerodendri Smith-Meyer & Ueckermann, 1990
* Aculus cneorum (Smith-Meyer & Ueckermann, 1990)
* Aculus colei Channabasavanna, 1966
* Aculus comatus (Nalepa, 1892)
* Aculus conspicillatus Flechtmann in Flechtmann & Moraes, 2003
* Aculus convolvuli (Nalepa, 1891)
* Aculus coronillae (Canestrini & Massalongo, 1893)
* Aculus corynocarpi (Manson, 1984)
* Aculus cotyledonis (Keifer, 1939)
* Aculus craspedobius (Nalepa, 1925)
* Aculus crataegumplicans (Cotte, 1910)
* Aculus cribatus (Keifer, 1961)
* Aculus cysticola (Canestrini, 1892)
* Aculus cytisi Labanowski in Soika & Labanowski, 2000
* Aculus daibuensis Huang, 2001
* Aculus daneki Shi & Boczek, 2000
* Aculus daphnes (Roivainen, 1951)
* Aculus didymbotryae Abou-Awad & El-Banhawy, 1992
* Aculus dimidiatus (Hall, 1967)
* Aculus dipterocarpi Boczek, 1993
* Aculus dominguensis (Cromroy, 1958)
* Aculus epilobiorum (Liro, 1940)
* Aculus epiphyllus (Nalepa, 1892)
* Aculus eurynotus (Nalepa, 1894)
* Aculus excoecaria (Mondal & Chakrabarti, 1982)
* Aculus fabus Boczek & Knihinicki, 1998
* Aculus fascigrans Boczek & Shi in Boczek, Shi & Lewandowski, 2002
* Aculus ficivagrans Mohanasundaram, 1985
* Aculus flacourtiae Mohanasundaram, 1984
* Aculus fockeui (Nalepa & Trouessart, 1891)
* Aculus fraxini (Nalepa, 1894)
* Aculus gemmarum (Nalepa, 1892)
* Aculus glacialis (Thomas, 1885)
* Aculus glechomae (Liro, 1940)
* Aculus groenlandicus (Rostrup, 1900)
* Aculus gutierrezi (Keifer, 1973)
* Aculus haloragi (Lamb, 1953)
* Aculus hapsis Smith-Meyer & Ueckermann, 1990
* Aculus harpazi Smith-Meyer & Ueckermann, 1990
* Aculus heatherae (Manson, 1984)
* Aculus hedysari (Liro, 1941)
* Aculus helianthemi (Roivainen, 1950)
* Aculus helichrysi (Meyer & Ueckermann, 1990)
* Aculus hippocastani (Fockeu, 1890)
* Aculus huangzhongensis Kuang, 2000
* Aculus hygrophilus (Roivainen, 1953)
* Aculus hyperici (Liro, 1943)
* Aculus ichnocarpi (Ghosh & Chakrabarati, 1989)
* Aculus indicus Channabasavanna, 1966
* Aculus inquilinus Berezantsev, 1989
* Aculus jahandiezi (Cotte, 1924)
* Aculus jilinensis (Kuang, 1995)
* Aculus juglandis Natcheff, 1966
* Aculus khayae Smith-Meyer & Ueckermann, 1990
* Aculus kochi (Nalepa & Thomas, 1894)
* Aculus kolengii Mohanasundaram, 1981
* Aculus konoellus Keifer, 1965
* Aculus kumari (Mohanasundaram, 1982)
* Aculus lactucae (Canestrini, 1893)
* Aculus laevigatae (Hassan, 1928)
* Aculus laevis (Nalepa, 1892)
* Aculus lambi (Manson, 1965)
* Aculus latilobus (Keifer, 1955)
* Aculus latus (Nalepa, 1894)
* Aculus leguminae (Mohanasundaram, 1982)
* Aculus leionotus (Nalepa, 1891)
* Aculus leonotis Smith-Meyer & Ueckermann, 1990
* Aculus lepidii (Roivainen, 1953)
* Aculus leptadeniae Mohanasundaram, 1985
* Aculus ligustri (Keifer, 1938)
* Aculus liquidambaris (Keifer, 1940)
* Aculus longanensis Wei & Kuang, 1997
* Aculus longifilis (Canestrini, 1891)
* Aculus longinychus (Smith-Meyer & Ueckermann, 1990)
* Aculus longiseta Carmona, 1972
* Aculus magnirostris (Nalepa, 1892)
* Aculus malus (Zaher & Abou, Awad, 1979)
* Aculus malvae Boczek & Davis, 1984
* Aculus mansoni Amrine & Stasny, 1994
* Aculus marinkovici Petanovic, 1987
* Aculus martialis (Liro, 1941)
* Aculus masseei (Nalepa, 1925)
* Aculus mastigophorus (Nalepa, 1890)
* Aculus maximilianae (Briones & McDaniel, 1976)
* Aculus megacrinis Keifer, 1962
* Aculus meghriensis (Bagdasarian, 1970)
* Aculus meliae Kuang & Zhuo, 1989
* Aculus melicoccae (Boczek & Oleczek, 1988)
* Aculus menoni Channabasavanna, 1966
* Aculus merostichus (Nalepa, 1918)
* Aculus mespili (Bagdasarian, 1981)
* Aculus micheneri (Hall, 1967)
* Aculus microspinatus (Hall, 1967)
* Aculus minor (Nalepa, 1892)
* Aculus minutissimus (Hodgkiss, 1913)
* Aculus minutus (Nalepa, 1890)
* Aculus mogeri (Farkas, 1960)
* Aculus montanae Mohanasundaram, 1985
* Aculus moringae Channabasavanna, 1966
* Aculus morivagrans Boczek, 1968
* Aculus mumis (Kuang & Gong, 1996)
* Aculus myrsinites (Roivainen, 1947)
* Aculus nigrus Keifer, 1959
* Aculus niphlocladae (Keifer, 1966)
* Aculus ocimumae Mohanasundaram, 1991
* Aculus olearius Castagnoli, 1977
* Aculus oresterae (Keifer, 1978)
* Aculus ornatus (Roivainen, 1947)
* Aculus osteospermi Smith-Meyer & Ueckermann, 1990
* Aculus parafockeui Liu & Kuang, 1998
* Aculus parakarensis (Bagdasarian, 1972)
* Aculus parapycnanthemi Keifer, 1964
* Aculus parvensis (Manson, 1972)
* Aculus parvifoli (Smith-Meyer & Ueckermann, 1990)
* Aculus parvus (Nalepa, 1892)
* Aculus pavoniae Smith-Meyer & Ueckermann, 1990
* Aculus phaneris Amrine, 2002
* Aculus phyllocopteoides (Nalepa, 1891)
* Aculus physodesi (Keifer, 1979)
* Aculus pimpinellae (Liro, 1941)
* Aculus pistaciae Kuang-Haiyuan, 1998
* Aculus pitangae Boczek & Davis, 1984
* Aculus pittosporae (Mohanasundaram, 1982)
* Aculus pittosporacus Amrine & Stasny, 1994
* Aculus pittosporae (Mohanasundaram, 1982)
* Aculus podanthi (Nalepa, 1929)
* Aculus pretoriensis (Meyer & Ueckermann, 1990)
* Aculus pritchardi (Keifer, 1953)
* Aculus prunelli (Liro, 1942)
* Aculus pteleaevagrans Boczek, 1964
* Aculus pterygospermae Mohanasundaram, 1985
* Aculus pulaviensis Boczek, 1968
* Aculus pycnorhynchus (Nalepa, 1925)
* Aculus quinquilobus (Hodgkiss, 1913)
* Aculus reticulatus (Nalepa, 1890)
* Aculus retiolatus (Nalepa, 1892)
* Aculus rhamnivagrans (Keifer, 1939)
* Aculus rhododendronis (Keifer, 1940)
* Aculus rhopodacrus Smith-Meyer & Ueckermann, 1990
* Aculus ribis (Massee, 1929)
* Aculus rigidus (Nalepa, 1894)
* Aculus rubiae (Roivainen, 1953)
* Aculus rugosus (Sapozhnikova, 1980)
* Aculus salicis Kuang & Luo, 1997
* Aculus salicisalbae (Nalepa, 1925)
* Aculus salicisarbusculae (Nalepa, 1925)
* Aculus salicisforbyanae (Nalepa, 1925)
* Aculus salicisfragilis (Nalepa, 1925)
* Aculus salicisgrandifoliae (Nalepa, 1925)
* Aculus salicisincanae (Nalepa, 1925)
* Aculus salicisretusae (Nalepa, 1925)
* Aculus sarcococcae Mohanasundaram, 1985
* Aculus sargentodoxae Wei & Kuang, 1997
* Aculus sarothamni Boczek, 1961
* Aculus sayanicus Skoracka in Skoracka & Pacyna, 2003
* Aculus schlechtendali (Nalepa, 1890)
* Aculus schmardae (Nalepa, 1889)
* Aculus schubarti (Farkas, 1960)
* Aculus scutellariae (Canestrini & Massalongo, 1895)
* Aculus semenovi (Shevtchenko, Marikovski & Shamsutdinova, 1973)
* Aculus shanghaiensis Kuang, 1997
* Aculus shaoxingensis Kuang-Haiyuan, 1998
* Aculus shoreum (Ghosh, Mondal & Chakrabarti, 1986)
* Aculus smilacis Keifer, 1962
* Aculus solani Boczek & Davis, 1984
* Aculus stachysi (Petanovic & Boczek, 1991)
* Aculus staphyleae (Pantanelli, 1912)
* Aculus stigmatus (Nalepa, 1914)
* Aculus succiphagus (Meyer & Ueckermann, 1990)
* Aculus symphoricarpi (Keifer, 1939)
* Aculus syriacus (Fockeu, 1892)
* Aculus tamalpais (Keifer, 1939)
* Aculus tetanothrix (Nalepa, 1889)
* Aculus tetracanthus Smith-Meyer & Ueckermann, 1990
* Aculus tetraspermae (Chandrapatya, 1992)
* Aculus teucrii (Nalepa, 1892)
* Aculus thomasi (Nalepa, 1895)
* Aculus tibialis (Liro, 1943)
* Aculus tiliaevagrans Boczek, 1964
* Aculus tomentosi (Meyer & Ueckermann, 1990)
* Aculus triflorae (Di Stefano, 1966)
* Aculus trifolii Natcheff, 1979
* Aculus truncatus (Nalepa, 1892)
* Aculus ulae Boczek, 1961
* Aculus unctus Boczek, 1964
* Aculus undatae Smith-Meyer & Ueckermann, 1990
* Aculus vallis Keifer, 1966
* Aculus variabilis (Roivainen, 1953)
* Aculus verbenae (Keifer, 1977)
* Aculus vermicularis (Nalepa, 1914)
* Aculus verrucosus Flechtmann, 1996
* Aculus viburni Petanovic, Boczek & Shi, 2002
* Aculus viburnifoliae (Boczek & Shi, 1995)
* Aculus vitecicola (Kikuti, 1939)
* Aculus wagnoni (Keifer, 1959)
* Aculus wisterifoliae Keifer, 1963
* Aculus xylostei (Canestrini, 1891)
* Aculus yelagiriensis Mohanasundaram, 1980
* Aculus zaheri (Abou-Awad, 1979)
* Aculus zygophylli Smith-Meyer & Ueckermann, 1990