# List of Epedanidae species

This is a list of the described species of the harvestman family Epedanidae. The data is taken from Joel Hallan's Biology Catalog.

## Dibuninae

* Dibuninae Roewer, 1912
   * Dibunus Loman, 1906
      * Dibunus albitarsus (Roewer, 1927) — Philippines
      * Dibunus chapmani (Roewer, 1927) — Philippines
      * Dibunus dividuus Suzuki, 1977
      * Dibunus gracile (Roewer, 1912) — Philippines
      * Dibunus longipalpis Roewer, 1912 — Philippines
      * Dibunus maculatipes (Roewer, 1915) — Maluku Islands
      * Dibunus marianae Goodnight & Goodnight, 1957
      * Dibunus pseudobiantes Loman, 1906 — New Guinea
      * Dibunus similis Roewer, 1912 — Philippines
      * Dibunus transitorius (Roewer, 1927) — Philippines

## Epedaninae

* Epedaninae Sørensen, in L.Koch 1886
   * Alloepedanus Suzuki, 1985
      * Alloepedanus robustus Suzuki, 1985 — Thailand
   * Balabanus Suzuki, 1977
      * Balabanus quadrispinosus Suzuki, 1977
   * Caletor Loman 1892 (! contra Hallan)
      * Caletor bicolor (Sørensen, 1932)
      * Epedanus javanus Thorell, 1876 — Java
      * Caletor sumbawanus (Roewer, 1938) — Sumbawa
      * Caletor unguidens (Loman, 1892)
   * Caletorellus Roewer, 1938
      * Caletorellus siamensis (Hirst, 1912) — Thailand
   * Epedanellus Roewer, 1911
      * Epedanellus tuberculatus Roewer, 1911 — Japan
   * Epedanidus Roewer, 1945
      * Epedanidus globibunus Roewer, 1945 — Perak
   * Epedanulus Roewer, 1913
      * Epedanulus sarasinorum Roewer, 1913 — Sulawesi
   * Epedanus Thorell, 1876
      * Epedanus armatus Banks, 1930 (after Hallan)
      * Epedanus brevipalpus Banks, 1930 — Borneo
      * Epedanus cavicolus Banks, 1930 — Borneo
      * Epedanus pictus Thorell, 1876 — Sarawak
      * Epedanus praedo Sørensen, 1932 — Borneo
   * Euepedanus Roewer, 1915
      * Euepedanus chaiensis Suzuki, 1970
      * Euepedanus dividuus Suzuki, 1970
      * Euepedanus orientalis (Hirst, 1912) — Thailand
      * Euepedanus pentaspinulatus S. Suzuki, 1985 — Thailand
      * Euepedanus similis S. Suzuki, 1985 — Thailand
      * Euepedanus spinosus S. Suzuki, 1985 — Thailand
      * Euepedanus trispinosus Roewer, 1915 — Malacca
   * Funkikoa Roewer, 1927
      * Funkikoa maxima Roewer, 1927 — Funkiko (Taiwan)
   * Heteroepedanus Roewer, 1912
      * Heteroepedanus monacantha (Roewer, 1911)
      * Heteroepedanus triacantha (Roewer, 1911)
   * Aboriscus Roewer, 1940
      * Aboriscus singularis (Roewer, 1912) — India, Malaysia
      * Aboriscus longipes (Roewer, 1913) — India
      * Aboriscus aborensis (Roewer, 1913) — India
   * Lobonychium Roewer, 1938
      * Lobonychium palpiplus Roewer, 1938 — Borneo
   * Metathyreotus Roewer, 1913
      * Metathyreotus aborensis Roewer, 1913 — Assam
      * Metathyreotus kempi Roewer, 1913 — Assam
   * Metepedanulus Roewer, 1913
      * Metepedanulus sarasinorum Roewer, 1913 — Sulawesi
      * Metepedanulus flaveolus Banks, 1931 — Borneo
   * Metepedanus Roewer, 1912
      * Metepedanus venator (Roewer, 1911) — Borneo
      * Metepedanus accentuatus (Roewer, 1911) — Borneo
   * Mosfora Roewer, 1938
      * Mosfora silvestrii (Roewer, 1925) — Funkiko
   * Nanepedanus Roewer, 1938
      * Nanepedanus rufus Roewer, 1938 — Borneo
   * Neoepedanus Roewer, 1912
      * Neoepedanus fokiensis Roewer, 1912 — China
   * Paratakaoia S. Suzuki, 1985 — Thailand
      * Paratakaoia parva S. Suzuki, 1985
      * Paratakaoia minima (Suzuki, 1986) — Thailand
   * Parepedanulus Roewer, 1913
      * Parepedanulus sarasinorum Roewer, 1913 — Sulawesi
      * Parepedanulus bicmaculatus Roewer, 1915
   * Plistobunus Pocock, 1903
      * Plistobunus rapax Pocock, 1903 — Hong Kong
   * Pseudoepedanus Suzuki, 1969
      * Pseudoepedanus dolensis Suzuki, 1970
   * Pseudomarthana P. D. Hillyard, 1985
      * Pseudomarthana conspicua P. D. Hillyard, 1985 — Malaysia
   * Takaoia Roewer, 1911 — Takoia? (Solomon Islands)
      * Takaoia kubotai S. Suzuki, 1982 — Malaysia
      * Takaoia sauteri Roewer, 1911 — Taiwan
      * Takaoia similis Roewer, 1915 — Taiwan
   * Thyreotus Thorell, 1889
      * Thyreotus bifasciatus Thorell, 1889 — Burma
      * Thyreotus bimaculatus Roewer, 1912 — Burma
   * Toccolus Roewer, 1927
      * Toccolus minimus Roewer, 1927 — Tonking
      * Toccolus chibai Suzuki, 1976
      * Toccolus globitarsis Suzuki, 1970
   * Zepedanulus Roewer, 1927
      * Zepedanulus armatipalpus Roewer, 1927 — Malacca
      * Zepedanulus alter Roewer, 1963
      * Zepedanulus ishikawai Suzuki, 1971
      * Zepedanulus watanabei S. Suzuki, 1981 — Thailand

## Acrobuninae

* Acrobuninae Roewer, 1912
   * Acrobunus Thorell, 1891
      * Acrobunus nigropunctatus Thorell, 1891 — Sumatra
      * Acrobunus bifasciatus Thorell, 1911 — Sumatra
      * Acrobunus thorelli Banks, 1931 — Borneo
   * Anacrobunus Roewer, 1927
      * Anacrobunus filipes Roewer, 1927 — Riau Islands
   * Harpagonellus Roewer, 1927
      * Harpagonellus glaber Roewer, 1927 — Sumatra
   * Heterobiantes Roewer, 1912
      * Heterobiantes geniculatus (Pocock, 1903) — Hong Kong
   * Metacrobunus Roewer, 1915
      * Metacrobunus macrochelis Roewer, 1915 — Malacca
      * Metacrobunus frontalis Banks, 1931 — Borneo
   * Paracrobunus Suzuki, 1977
      * Paracrobunus bimaculatus Suzuki, 1977
      * Paracrobunus similis Suzuki, 1982

## Sarasinicinae

* Sarasinicinae Roewer, 1923 — should possibly be a family
   * Acanthepedanus Roewer, 1912
      * Acanthepedanus armatus Roewer, 1912 — Sumatra
   * Albertops Roewer, 1938
      * Albertops robustus Roewer, 1938 — Borneo
   * Asopella Kury, 2020 (not Sørensen, 1932, contra Hallan)
      * Asopella lutescens (Thorell, 1876) — Sarawak (contra Hallan)
      * Asopella robusta Suzuki, 1982 — Philippines
      * Asopella xanti Sørensen, 1932 — Java
   * Delicola Roewer, 1938
      * Delicola longipalpis Roewer, 1938 — Sumatra
   * Gintingius Roewer, 1938
      * Gintingius robustus Roewer, 1938 — Pahang
   * Kilungius Roewer, 1915
      * Kilungius bimaculatus Roewer, 1915 — Taiwan
      * Kilungius insulanus (Hirst, 1911) — Japan?
   * Koyanus Roewer, 1938
      * Koyanus clarus Roewer, 1938 — Borneo
   * Kuchingius Roewer, 1927
      * Kuchingius megalopalpus Roewer, 1927 — Borneo
   * Nobeoka Roewer, 1938 — Japan
      * Nobeoka laevis Roewer, 1938
   * Opelytus Roewer, 1927
      * Opelytus rugichelis Roewer, 1938 — Borneo
      * Opelytus simoni Roewer, 1927 — Malacca
      * Opelytus spinichelis Rower, 1938 — Tenasserim
      * Opelytus vepretum Roewer, 1927 — Pulu Pinang
   * Padangcola Roewer, 1963 — should possibly be in Epedaninae
      * Padangcola jacobsoni Roewer, 1963
   * Panticola Roewer, 1938 — placement is uncertain
      * Panticola elegans Roewer, 1938 — Malacca
   * Parepedanus Roewer, 1912
      * Parepedanus bispinosus Roewer, 1912 — Sumatra
      * Parepedanus bimaculatus Roewer, 1915 — Malacca
   * Pasohnus Suzuki, 1976 — was in Phalangodidae
      * Pasohnus bispinosus Suzuki, 1976
   * Pseudobiantes Hirst, 1911 — Japan
      * Pseudobiantes japonicus Hirst, 1911 — Japan
      * Pseudobiantes silvestrii (Mello-Leitão, 1944) — nomen novum for P. japonicus
   * Punanus Roewer, 1938
      * Punanus tenuis Roewer, 1938 — Borneo
   * Sarasinica Strand, 1914
      * Sarasinica atra Roewer, 1938 — Borneo
      * Sarasinica femoralis Roewer, 1938 — Borneo
      * Sarasinica tricommata (Roewer, 1913) — Sulawesi
          Sarasinica tricommata tricommata (Roewer, 1913)
          Sarasinica tricommata quadripunctata Roewer, 1913
          Sarasinica tricommata sexpunctata Roewer, 1913
      * Sarasinica henrikseni Mello-Leitão, 1944
   * Sembilanus Roewer, 1938
      * Sembilanus rugichelis Roewer, 1938 — Malacca
   * Sinistus Roewer, 1938
      * Sinistus maculatus Roewer, 1938 — Borneo
      * Sinistus fuscus Roewer, 1938 — Borneo
   * Siponnus Roewer, 1927
      * Siponnus stimulatus Roewer, 1927 — Pulu Pinang
   * Sungsotia Tsurusaki, 1995
      * Sungsotia uenoi Tsurusaki, 1995 — Vietnam
   * Tegestria Roewer, 1936
      * Tegestria borneensis Roewer, 1938 — Borneo
      * Tegestria coniata Roewer, 1938 — Malacca
      * Tegestria johorea Roewer, 1936 — Malacca
      * Tegestria montana Roewer, 1938 — Malaysia: Penang
      * Tegestria parva Suzuki, 1970
      * Tegestria pinangensis (Roewer 1938) - Malaysia: Penang (contra Hallan)
      * Tegestria seriata Roewer, 1938 — Malacca
      * Tegestria sumatrana Roewer, 1938 — Sumatra
   * Tonkinatus Roewer, 1938 — Tonking
      * Tonkinatus bimaculatus Roewer, 1938

## Incertae sedis

* Incertae sedis
   * Beloniscellus Roewer, 1912
      * Beloniscellus lombokiensis (Roewer, 1912) — Lombok
      * Beloniscellus floresianus Roewer, 1931 — Sunda Islands
      * Beloniscellus renschi Roewer, 1931 — Sunda Islands
      * Beloniscellus sumbawaensis Roewer, 1931 — Sunda Islands
      * Beloniscellus parvicalcar Roewer, 1931 — Sunda Islands
      * Beloniscellus narmadeus Roewer, 1949 — Indonesia
   * Beloniscops Roewer, 1949
      * Beloniscops flavicalcar Roewer, 1949 — Sumatra
      * Beloniscops lata Roewer, 1949 — Sumatra
   * Belonisculus Roewer, 1923
      * Belonisculus jacobsoni Roewer, 1923 — Simalur Island (Sumatra)
   * Beloniscus Thorell, 1891
      * Beloniscus albimarginatus Roewer, 1915 — Singapore
      * Beloniscus albiephippiatus Roewer, 1916 — Singapore
      * Beloniscus albipustulatus Roewer, 1949 — Sumatra
      * Beloniscus biconus Roewer, 1926 — Sumatra
      * Beloniscus bicornis Roewer, 1927 — Sumatra
      * Beloniscus malayanus Roewer, 1949 — Sumatra
      * Beloniscus morosus Thorell, 1891 — Sumatra
      * Beloniscus ochraceus Loman, in Weber 1892 — Sumatra
      * Beloniscus pustulosus Loman, in Weber 1892 — Sumatra
      * Beloniscus quinquespinosus Thorell, 1891 — Sumatra
      * Beloniscus simaluris Roewer, 1923 — Simalur Island
      * Beloniscus thienemanni Roewer, 1931 — Sunda Islands
      * Beloniscus tricalcaratus Roewer, 1949 — Java
      * Beloniscus tuberculatus Roewer, 1927 — Sumatra
   * Buparellus Roewer, 1949
      * Buparellus mitylus (Thorell, 1889) — Burma
      * Buparellus dibunichelis Roewer, 1949 — Burma
      * Buparellus patellaris Roewer, 1949 — Burma
      * Buparellus insolitus S. Suzuki, 1985 — Thailand
   * Bupares Thorell, 1889
      * Bupares armatus Roewer, 1949 — Burma
      * Bupares caper Thorell, 1889 — India, Burma
      * Bupares chelicornis Roewer, 1927 — Thailand
      * Bupares degerbolae S. Suzuki, 1985 — Thailand
      * Bupares granulatus Thorell, 1890 — Pinang
      * Bupares pachytarsus Roewer, 1934 — Perak, Singapore
      * Bupares stridulator Roewer, 1949 — Burma
      * Bupares trochanteralis Roewer, 1949 — Burma
   * Dhaulagirius Martens, 1977 — placement is uncertain
      * Dhaulagirius altitudinalis Marten, 1977
   * Dumaguetes Roewer, 1927
      * Dumaguetes chapmani Roewer, 1927 — Philippines
   * Parabeloniscus Suzuki, 1967
      * Parabeloniscus caudatus Suzuki, 1973
      * Parabeloniscus nipponicus Suzuki, 1967
      * Parabeloniscus shimojanai Suzuki, 1971
   * Parabupares S. Suzuki, 1982
      * Parabupares robustus S. Suzuki, 1982 — Indonesia
   * Sotekia S. Suzuki, 1982
      * Sotekia minima S. Suzuki, 1982 — Indonesia
   * Tithaeus Thorell, 1890
      * Tithaeus annandalei Roewer, 1927 — Singapore
      * Tithaeus birmanicus Roewer, 1949 — Burma
      * Tithaeus borneensis Roewer, 1949 — Borneo
      * Tithaeus cruciatus Roewer, 1927 — Sumatra
      * Tithaeus drac Lian, Zhu & Kury, 2008 — China
      * Tithaeus flavescens Banks, 1931 — Borneo
      * Tithaeus fraseri Suzuki, 1972 — Malaysia
      * Tithaeus fuscus Roewer, 1949 — Borneo
      * Tithaeus granulatus Banks, 1931 — Borneo
      * Tithaeus indochinensis Roewer, 1925 — Tonking
      * Tithaeus jacobsoni Roewer, 1923 — Sumatra
      * Tithaeus javanus Roewer, 1949 — Java
      * Tithaeus johorensis Roewer, 1949 — Malaya
      * Tithaeus kokutnus S. Suzuki, 1985 — Thailand
      * Tithaeus krakatauensis Roewer, 1949 — Indonesia
      * Tithaeus laevigatus Thorell, 1890 — Pinang
      * Tithaeus lesserti Roewer, 1949 — Borneo
      * Tithaeus longipes Banks, 1931 — Borneo
      * Tithaeus malakkanus Roewer, 1949 — Staits
      * Tithaeus metatarsalis Roewer, 1949 — Malaya
      * Tithaeus minor Roewer, 1949 — Singapore
      * Tithaeus nigripes Banks, 1931 — Borneo
      * Tithaeus pumilio Roewer, 1949 — Sarawak
      * Tithaeus rotundus Suzuki, 1970
      * Tithaeus rudispina Roewer, 1935 — Malaya
      * Tithaeus rudispina Roewer, 1949 — Johore (preoccupied?)
      * Tithaeus sarawakensis Roewer, 1912 — Sarawak
      * Tithaeus siamensis Roewer, 1949 — Thailand
      * Tithaeus similis S. Suzuki, 1985 — Thailand
      * Tithaeus tenuis Roewer, 1949 — Johore
      * Tithaeus timorensis Roewer, 1949 — Indonesia
      * Tithaeus trimaculatus Roewer, 1949 — Indonesia
      * Tithaeus vagus (Loman, 1892)
      * Tithaeus watanabei Suzuki, 1970b
   * Tokunosia Suzuki, 1964
      * Tokunosia tenuipes Suzuki, 1964
          Tokunosia tenuipes tenuipes Suzuki, 1964
          Tokunosia tenuipes tuberuculata Suzuki, 1973
          Tokunosia tenuipes taiwana Suzuki, 1977