# Sclerococcum

Sclerococcum is a genus of lichenicolous fungi in the family Dactylosporaceae.

## Taxonomy

The genus was circumscribed in 1825 by Elias Magnus Fries. The type species is Sclerococcum sphaerale, originally described in 1814 by Erik Acharius (as Spiloma sphaerale). This fungus is a lichenicolous hyphomycete – a mould that lives on a lichen. Most of the Sclerococcum species described since then are also lichenicolous, and most have a restricted host range. Molecular phylogenetics analysis published in 2012 showed that Sclerococcum sphaerale grouped together in a clade with species of Dactylospora in the class Eurotiomycetes.
In 2016, Réblová and colleagues proposed a new family Sclerococcaceae in a new order (Sclerococcales) to accommodate the type genus Sclerococcum, Dactylospora, Rhopalophora, three strains of beetle-associated fungi, and an isolate of Fusichalara minuta. This classification was not accepted in the 2017 Outline of the Ascomycota, which retained the family Dactylosporaceae.

## Species

* Sclerococcum acarosporae S.Y.Kondr. (2002)
* Sclerococcum acarosporicola Ertz & Diederich (2018)
* Sclerococcum aeruginosum (Holien & Ihlen) Ertz & Diederich (2018)
* Sclerococcum ahtii (Zhurb. & Pino-Bodas) Ertz & Diederich (2018)
* Sclerococcum allantoideum (Alstrup & D.Hawksw.) Ertz & Diederich (2018)
* Sclerococcum amygdalariae (Triebel) Ertz & Diederich (2018)
* Sclerococcum anziae (Zhurb., Ezhkin, Skirina & Y.Ohmura) Ertz & Diederich (2018)
* Sclerococcum aptrootii Diederich (2015)
* Sclerococcum areolatum Etayo (2008)
* Sclerococcum arthrobotryinum (Hafellner) Olariaga, Teres, J.M.Martín, M.Prieto & Baral (2019)
* Sclerococcum aspiciliicola (Alstrup & D.Hawksw.) Ertz & Diederich (2018)
* Sclerococcum athallinum (Müll.Arg.) Ertz & Diederich (2018)
* Sclerococcum attendendum (Nyl.) Ertz & Diederich (2018)
* Sclerococcum australe (Triebel & Hertel) Ertz & Diederich (2018)
* Sclerococcum bloxamii (Berk. & W.Phillips) Olariaga, Teres, J.M.Martín, M.Prieto & Baral (2019)
* Sclerococcum boreale (Holien & Ihlen) Ertz & Diederich (2018)
* Sclerococcum caledonicum (Hafellner) Olariaga, Teres, J.M.Martín, M.Prieto & Baral (2019)
* Sclerococcum canariense (Kohlm. & Volkm.-Kohlm.) Olariaga, Teres, J.M.Martín, M.Prieto & Baral (2019)
* Sclerococcum cladoniicola (Alstrup & Olech) Ertz & Diederich (2018)
* Sclerococcum crassitunicatum Zhurb., Diederich & U.Braun (2017)
* Sclerococcum crassum (Sarrión & Hafellner) Ertz & Diederich (2018)
* Sclerococcum davidii (Hafellner & H.Mayrhofer) Ertz & Diederich (2018)
* Sclerococcum deminutum (Th.Fr.) Ertz & Diederich (2018)
* Sclerococcum dendriscostictae Y.Joshi (2021) – India
* Sclerococcum dobrowolskii (Olech & Alstrup) Ertz & Diederich (2018)
* Sclerococcum epicladonia Zhurb. (2017)
* Sclerococcum epimyces (Tobisch) Olariaga, Teres, J.M.Martín, M.Prieto & Baral (2019)
* Sclerococcum epiphytorum Diederich (1990)
* Sclerococcum ewersii Elix, P.M.McCarthy & Hafellner (2019) – Australia
* Sclerococcum fissurinae Pérez-Ort. (2020) – Alaska
* Sclerococcum frigidum (Hafellner) Ertz & Diederich (2018)
* Sclerococcum gelidarium Etayo & F.Berger (2000)
* Sclerococcum glaucomarioides (Willey ex Tuck.) Ertz & Diederich (2018)
* Sclerococcum griseisporodochium Etayo (1995)
* Sclerococcum hafellnerianum (Sérus.) Ertz & Diederich (2018)
* Sclerococcum haliotrephum (Kohlm. & E.Kohlm.) Ertz & Diederich (2018)
* Sclerococcum heterodermiae (Etayo) Ertz & Diederich (2018)
* Sclerococcum homoclinellum (Nyl.) Ertz & Diederich (2018)
* Sclerococcum imperfectum (Ellis) Olariaga, Teres, J.M.Martín, M.Prieto & Baral (2019)
* Sclerococcum inconspicuum (Etayo) Ertz & Diederich (2018)
* Sclerococcum inopinum (Döbbeler & W.R.Buck) Olariaga, Teres, J.M.Martín, M.Prieto & Baral (2019)
* Sclerococcum inquilinum (Tuck.) Ertz & Diederich (2018)
* Sclerococcum leuckertii Diederich & P.Scholz (1995)
* Sclerococcum lobariellum (Nyl.) Ertz & Diederich (2018)
* Sclerococcum luridum (Hafellner) Olariaga, Teres, J.M.Martín, M.Prieto & Baral (2019)
* Sclerococcum mangrovei (E.B.G.Jones, Alias, Abdel-Wahab & S.Y.Hsieh) Ertz & Diederich (2018)
* Sclerococcum mediterraneum (Sarrión & Hafellner) Olariaga, Teres, J.M.Martín, M.Prieto & Baral (2019)
* Sclerococcum microsporum (Etayo) Ertz & Diederich (2018)
* Sclerococcum montagnei Hafellner (1996) – Macaronesia
* Sclerococcum olivaceum Etayo & F.Berger (2004)
* Sclerococcum ophthalmizae Coppins (2018)
* Sclerococcum orygmaeum (Nyl.) Ertz & Diederich (2018)
* Sclerococcum parasitaster (Nyl.) Ertz & Diederich (2018)
* Sclerococcum parasiticum (Flörke) Ertz & Diederich (2018)
* Sclerococcum parellarium (Nyl.) Ertz & Diederich (2018)
* Sclerococcum pertusariicola (Willey ex Tuck.) Ertz & Diederich (2018)
* Sclerococcum phaeophysciae Diederich & van den Boom (2017)
* Sclerococcum phyllobaeis Etayo (2017)
* Sclerococcum physciae Y.Joshi (2021) – India
* Sclerococcum pleiospermum (Triebel) Ertz & Diederich (2018)
* Sclerococcum polysporum (Triebel) Ertz & Diederich (2018)
* Sclerococcum porphyreum (Hafellner & Kalb) Ertz & Diederich (2018)
* Sclerococcum protothallinum (Anzi) Ertz & Diederich (2018)
* Sclerococcum pseudourceolatum (Sarrión & Hafellner) Olariaga, Teres, J.M.Martín, M.Prieto & Baral (2019)
* Sclerococcum purpurascens (Triebel) Ertz & Diederich (2018)
* Sclerococcum pyrenaicum (Etayo) Ertz & Diederich (2018)
* Sclerococcum rhyparizae (Arnold) Ertz & Diederich (2018)
* Sclerococcum ricasoliae (Vouaux) Flakus, Rodr.Flakus & Etayo (2019)
* Sclerococcum rimulicola (Müll.Arg.) Ertz & Diederich (2018)
* Sclerococcum rostrupii (Alstrup) Ertz & Diederich (2018)
* Sclerococcum rubiginosum (Hafellner) Olariaga, Teres, J.M.Martín, M.Prieto & Baral (2019)
* Sclerococcum saxatile (Schaer.) Ertz & Diederich (2018)
* Sclerococcum serusiauxii Boqueras & Diederich (1993)
* Sclerococcum simplex D.Hawksw. (1979)
* Sclerococcum sipmanii Diederich (2015)
* Sclerococcum sphaerale (Ach.) Fr. (1825)
* Sclerococcum stipitatum (Hafellner) Olariaga, Teres, J.M.Martín, M.Prieto & Baral (2019)
* Sclerococcum stygium (Berk. & M.A.Curtis) Olariaga, Teres, J.M.Martín, M.Prieto & Baral (2019)
* Sclerococcum suburceolatum (Coppins & Fryday) Ertz & Diederich (2018)
* Sclerococcum tegularum (Arnold) Ertz & Diederich (2018)
* Sclerococcum tephromelarum Etayo & Calat. (1998)
* Sclerococcum thelotrematicola (Elix) Fryday (2019)
* Sclerococcum toensbergii Diederich (2017)
* Sclerococcum urceolatum (Th.Fr.) Ertz & Diederich (2018)
* Sclerococcum verrucisporum Alstrup (1993)
* Sclerococcum verruculosum (Hafellner) Olariaga, Teres, J.M.Martín, M.Prieto & Baral (2019)
* Sclerococcum vrijmoediae (K.L.Pang, Sheng Y.Guo, Alias, Hafellner & E.B.G.Jones) Ertz & Diederich (2018)