# 2019 in paleobotany

This article records new taxa of fossil plants that are scheduled to be described during the year 2019, as well as other significant discoveries and events related to paleobotany that are scheduled to occur in the year 2019.

## Mosses

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Heinrichsiella | Gen. et sp. nov | Valid | Bippus et al. | Jurassic | | Argentina | A moss, possibly related to the family Polytrichaceae or Timmiellaceae. Genus includes new species H. patagonica. |
| Kulindobryum | Gen. et sp. nov | Valid | Ignatov in Mamontov & Ignatov | Middle or Late Jurassic | Ukureyskaya Formation | Russia ( Zabaykalsky Krai) | A form genus of dispersed moss capsules. Genus includes new species K. taylorioides. |
| Polycingulatisporites multiverrucata | Sp. nov | In press | Santamarina in Santamarina et al. | Late Cretaceous (Cenomanian) | Mata Amarilla Formation | Argentina | Spores of a member of Bryophyta of uncertain phylogenetic placement, possibly of sphagnaceous affinity. Announced in 2019; the final version was scheduled to be published in 2020. |
| Sphagnum heinrichsii | Sp. nov | Valid | Ignatov et al. | Late Eocene | Rovno amber | Ukraine | A moss, a species of Sphagnum. |
| Paleaethallus | Gen. et sp. nov | Valid | Mamontov, Katagiri & Borovich in Mamontov & Ignatov | Late Jurassic | Glushkovo Formation | Russia ( Zabaykalsky Krai) | A thalloid bryophyte. Genus includes new species P. squarrosus. |

## Liverworts

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Khasurtythallus | Gen. et sp. nov | Valid | Mamontov in Mamontov & Ignatov | Early Cretaceous | | Russia ( Buryatia) | A Marchantiidae liverwort. The type species is K. monosolenioides. |
| Ricciopsis sandaolingensis | Sp. nov | Valid | Li & Sun in Li et al. | Middle Jurassic | Xishanyao Formation | China | A Ricciaceae liverwort. |
| Thallites yangcaogouensis | Sp. nov | Valid | Wang et al. | Late Triassic | Yangcaogou Formation | China | A plant of uncertain phylogenetic placement, probably a liverwort. |

## Ferns and fern allies

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Alloiopteris loecsei | Sp. nov | Valid | Pšenička et al. | Carboniferous (Moscovian) | | Germany | A zygopterid fern. |
| Annularia noronhai | Sp. nov | Valid | Correia et al. | Carboniferous (Gzhelian) | Douro Basin | Portugal | A member of the family Calamitaceae. Announced in 2019; the final version of the article naming it was published in 2021. |
| Azolla keuja | Sp. nov | Valid | Jud, De Benedetti, Gandolfo & Hermsen | Paleocene (Danian) | Salamanca Formation | Argentina | A species of Azolla. |
| Berendtiopteris | Gen. et comb. nov | Valid | Sadowski et al. | Eocene | Baltic amber | Europe (Baltic Sea region) | A plant of uncertain phylogenetic placement, probably a fern; a new genus for "Pecopteris" humboldtiana. |
| Bifariusotheca | Gen. et sp. nov | Valid | X.H.Zhao ex Doweld | Late Permian | Longtan Formation | China | A marattialean fern. Genus includes new species B. notocathaysica Doweld. |
| Bowmanites yongchangensis | Sp. nov | Valid | Sun et al. | Permian (Cisuralian) | | China | A member of Sphenophyllales. |
| Clavatisporites cenomaniana | Sp. nov | In press | Santamarina in Santamarina et al. | Late Cretaceous (Cenomanian) | Mata Amarilla Formation | Argentina | Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020. |
| Collarisporites minor | Sp. nov | In press | Santamarina in Santamarina et al. | Late Cretaceous (Cenomanian) | Mata Amarilla Formation | Argentina | Spores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020. |
| Cyathocarpus yongchangensis | Sp. nov | Valid | Sun & Sun in Sun et al. | Permian (Cisuralian) | Shanxi Formation | China | A fern related to Psaronius. |
| Floratheca | Gen. et sp. nov | Valid | Lundgren et al. | Early Permian | Río Genoa Formation | Argentina | A member of Marattiales. Genus includes new species F. apokalyptika. |
| Germera brousmicheae | Sp. nov | Valid | Álvarez-Vázquez | Carboniferous (Westphalian) | | Canada | A member of Filicopsida. |
| Hausmannia olaensis | Sp. nov | Valid | Golovneva & Grabovskiy | Late Cretaceous (Santonian–early Campanian) | | Russia | A Dipteridaceae umbrella fern. |
| Heinrichsia | Gen. et sp. nov | Valid | Regalado et al. | Cretaceous | Burmese amber | Myanmar | A fern belonging to the family Pteridaceae. Genus includes new species H. cheilanthoides. |
| Kamatheca | Gen. et comb. nov | Valid | Doweld | Permian | | Russia | A marattialean fern; a new genus for "Acitheca" gigantea Esaulova. |
| Marsilea sprungerorum | Sp. nov | Valid | Hermsen | Eocene | Green River Formation Parachute Creek Member | United States Utah | A Marsilea species water fern. |
| Neolobatannularia | Gen. et sp. nov | Valid | Sun & Li in Wang et al. | Late Triassic | Yangcaogou Formation | China | A member of Equisetales. Genus includes new species N. liaoningensis. |
| Osmundastrum gvozdevae | Sp. nov | Valid | Bazhenova & Bazhenov | Middle Jurassic (Bathonian) | | Russia ( Kursk Oblast) | A species of Osmundastrum. |
| Palaeosorum waipiata | Sp. nov | Valid | Kaulfuss et al. | Early Miocene | | New Zealand | A member of the family Polypodiaceae. |
| Phlebopteris kirchneri | Sp. nov | Valid | Barbacka & Kustatscher in Barbacka, Kustatscher & Bodor | Early Jurassic (Hettangian) | Mecsek Coal Formation | Hungary | A fern belonging to the family Matoniaceae. |
| Plenasium xiei | Sp. nov | Valid | Cheng et al. | Cretaceous | | China | A member of Osmundaceae. Announced in 2019; the final version of the article naming it was published in 2021. |
| Polymorphopteris magdalenae | Sp. nov | Valid | R.H.Wagner ex Doweld | Late Carboniferous | | Spain | A marattialean fern. |
| Polymorphopteris wagneri | Sp. nov | Valid | Doweld | Late Carboniferous (Kasimovian) | | Spain | A marattialean fern. |
| Polypodiisporites serratus | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil spore of a member of the family Polypodiaceae. |
| Polypodiisporites timidus | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil spore of a member of the family Polypodiaceae. |
| Rinistachya | Gen. et sp. nov | Valid | Prestianni & Gess | Devonian (Famennian) | Witpoort Formation | South Africa | A member of Sphenophyllales. The type species is R. hilleri. |
| Rothwellopteris | Gen. et sp. nov | Valid | He et al. | Late Permian | Xuanwei Formation | China | A fern belonging to the group Marattiales. Genus includes new species R. pecopteroides. |
| Scolecopteris libera | Sp. nov | Valid | Li et al. | Permian (Asselian) | Taiyuan Formation | China | A marattialean fern |
| Scolecopteris renaultii | Sp. nov | Valid | Doweld | Permian (Cisuralian) | | France | A marattialean fern. |
| Tiania resinus | Sp. nov | Valid | He & Wang | Permian (Lopingian) | Xuanwei Formation | China | A member of Osmundales belonging to the extinct family Guaireaceae. |

## Lycophytes

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Bergeria wenquanensis | Sp. nov | Valid | Feng, D'Rozario & Zhang | Carboniferous (Viséan) | Akeshake Formation | China | A member of Lepidodendrales belonging to the family Flemingitaceae. |
| Guangdedendron | Gen. et sp. nov | | Wang et al. | Devonian (Famennian) | Wutong Formation | China | A member of Isoetales belonging to the group Dichostrobiles. Genus includes new species G. micrum. |
| Omphalophloios wagneri | Sp. nov | Valid | Opluštil, Pšenička & Bek | Carboniferous (Moscovian) | Illinois Basin | United States ( Indiana) | |
| Sawdonia hippotheca | Sp. nov | Valid | Berry & Gensel | Devonian (probably late Givetian) | Campo Chico Formation | Venezuela | A member of Zosterophyllopsida. |

## Conifers
### Araucariaceae

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Agathoxylon gilii | Sp. nov | Valid | Ríos-Santos & Cevallos-Ferriz | Late Jurassic | Todos Santos Formation | Mexico | |
| Agathoxylon hoodii | Comb nov | valid | (Tidwell & Medlyn) Gee et al | Late Jurassic | Morrison Formation | USA Utah | An araucariaceous petrified wood. Moved from Araucarioxylon hoodii (1993) |
| Agathoxylon jericonse | Sp. nov | Valid | Ríos-Santos & Cevallos-Ferriz | Late Jurassic | Todos Santos Formation | Mexico | |
| Agathoxylon kotaense | Sp. nov | In press | Chinnappa, Rajanikanth & Pauline Sabina | ?Late Jurassic – Early Cretaceous | Kota Formation | India | A member of the family Araucariaceae. |
| Agathoxylon parrensis | Sp. nov | Valid | Ríos-Santos & Cevallos-Ferriz | Paleocene | Las Encinas Formation | Mexico | |
| Araucaria balfourensis | Sp. nov | Valid | Hill et al. | Cenozoic | | Australia | A species of Araucaria. |
| Araucaria macrophylla | Sp. nov | Valid | Hill et al. | Cenozoic | | Australia | A species of Araucaria. |
| Araucaria mollifolia | Sp. nov | Valid | Hill et al. | Cenozoic | | Australia | A species of Araucaria. |
| Araucaria rothwellii | Sp. nov | Valid | Kvaček in Kvaček et al. | Late Cretaceous (Campanian-Maastrichtian) | Bozova Formation | Turkey | A species of Araucaria. |
| Brachyphyllum garciarum | Sp. nov | Valid | Carrizo et al. | Early Cretaceous (early Hauterivian/early Barremian) | Springhill Formation | Argentina | Probably a member of the family Araucariaceae. |

### Cupressaceae

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Austrocupressinoxylon | Gen. et sp. nov | Valid | Nunes et al. | Early Cretaceous | | Argentina | A member of Cupressaceae. Genus includes new species A. barcinense. |
| Austrohamia asfaltensis | Sp. nov | Valid | Contreras et al. | Early Jurassic | Cañadón Asfalto Formation | Argentina | A member of the family Cupressaceae. |
| Callitris blackburnii | Sp. nov | Valid | Paull et al. | Middle Miocene | | Australia | A species of Callitris. |
| Cupressinoxylon pliocenica | Sp. nov | Valid | Akkemik | Pliocene | Örencik Formation | Turkey | A member of the family Cupressaceae described on the basis of fossil wood. |
| Mesocyparis sinica | Sp. nov | Valid | Cui et al. | Paleocene (Danian) | Wuyun Formation | China | A member of the family Cupressaceae. |
| Protaxodioxylon sahnii | Sp. nov | Valid | Chinnappa, Kavali & Rajanikanth | Late Jurassic to Early Cretaceous | Kota Formation | India | A member of Cupressaceae, possibly related to Taxodium. |
| Protodammara reimatamoriori | Sp. nov | Valid | Mays & Cantrill | Late Cretaceous (Cenomanian) | Tupuangi Formation | New Zealand | A member of Cupressaceae. |
| Taxodioxylon cabullensis | Sp. nov | Valid | Ríos-Santos & Cevallos-Ferriz | Late Cretaceous | Packard Formation | Mexico | A cupressaceous fossil wood. |
| Taxodium viligense | Sp. nov | Valid | Golovneva | Late Cretaceous (Coniacian) | Chingandzha Formation | Russia | A species of Taxodium. |

### Pinceae

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Abies cuitlahuacii | Sp. nov | Valid | Cevallos-Ferriz, Ríos-Santos & Lozano-García | Pleistocene | | Mexico | A fir. |
| Pinus plioarmandii | Sp. nov | Valid | An et al. | Pliocene | | China | A pine. |
| Pinuxylon alonissianum | Sp. nov | Valid | Mantzouka & Sakala in Mantzouka et al. | Early Miocene | | Greece | A member of the family Pinaceae described on the basis of fossil wood. |
| Schizolepidopsis borealis | Sp. nov | Valid | Domogatskaya & Herman | Early Cretaceous (Albian) | Balyktakh Formation | Russia | A member of the family Pinaceae. |

### Podocarpaceae

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Dacrycarpus guipingensis | Sp. nov | Valid | Wu et al. | Miocene | Erzitang Formation | China | A species of Dacrycarpus. Announced in 2019; the final version of the article naming it was published in 2021. |
| Kirketapel | Gen. et sp. nov | Valid | Andruchow-Colombo et al. | Paleocene (early Danian) | Salamanca Formation | Argentina | A member of the family Podocarpaceae. The type species is K. salamanquensis. |
| Podocarpus pliomacrophyllus | Sp. nov | In press | Chen et al. | Early Pliocene | | China | A species of Podocarpus. Announced in 2019; the final version of the article naming it is not published yet. |

### Other conifers

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Cephalotaxus maguanensis | Sp. nov | Valid | Zhang et al. | Middle Miocene | | China | A species of Cephalotaxus. |
| Cupressinocladus shelikhovii | Sp. nov | Valid | Golovneva | Late Cretaceous (Coniacian) | Chingandzha Formation | Russia | A cheirolepidiaceous species |
| Frenelopsis justae | Sp. nov | Valid | Barral et al. | Early Cretaceous (Albian) | Escucha Formation | Spain | A member of the family Cheirolepidiaceae. |
| Ningxiaites shitanjingensis | Sp. nov | Valid | Wei et al. | Permian (Changhsingian) | Sunjiagou Formation | China | A conifer wood. |
| Protocupressinoxylon carrizalense | Sp. nov | Valid | Correa et al. | Late Triassic | Carrizal Formation | Argentina | |

## Other seed plants

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Amyelon bogdense | Sp. nov | Valid | Wan, Yang & Wang | Late Permian or Early Triassic | Guodikeng Formation | China | A silicified gymnospermous root. |
| Arazedispermum | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Aptian-early Albian) | Figueira da Foz Formation | Portugal | A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species A. lustanicum. |
| Axsmithia | Gen. et comb. nov | Valid | Anderson et al. | Triassic | | Antarctica | A seed fern. Genus includes "Umkomasia" uniramia Axsmith et al. (2000). |
| Bowenia johnsonii | Sp. nov | Valid | Hill et al. | Early Eocene | | Australia | A cycad, a species of Bowenia. |
| Brinkia | Gen. et 2 sp. nov | Valid | Kustatscher, Visscher & van Konijnenburg-van Cittert | Permian (Lopingian) | Bellerophon Formation Gröden/Val Gardena Sandstone | Italy | A possible member of Czekanowskiales. Genus includes new species B. kerpiana and B. cortianensis. |
| Cordaabaxicutis jacobii | Sp. nov | Valid | Šimůnek | Carboniferous (Pennsylvanian) | | Czech Republic | A member of Cordaitales. |
| Cordaadaxicutis detmarovicensis | Sp. nov | Valid | Šimůnek | Carboniferous (Pennsylvanian) | | Czech Republic | A member of Cordaitales. |
| Cordaadaxicutis doubravensis | Sp. nov | Valid | Šimůnek | Carboniferous (Pennsylvanian) | | Czech Republic | A member of Cordaitales. |
| Cordaadaxicutis jaroslavii | Sp. nov | Valid | Šimůnek | Carboniferous (Pennsylvanian) | | Czech Republic | A member of Cordaitales. |
| Cordaadaxicutis orlovensis | Sp. nov | Valid | Šimůnek | Carboniferous (Pennsylvanian) | | Czech Republic | A member of Cordaitales. |
| Cryptokerpia | Gen. et sp. nov | Valid | Blomenkemper, Abu Hamad & Bomfleur | Late Permian | Umm Irna Formation | Jordan | An enigmatic type of gymnosperm leaf. Genus includes new species C. sarlaccophora. |
| Douropteris | Gen. et sp. nov | Valid | Correia et al. | Carboniferous (Gzhelian) | Douro Basin | Portugal | A seed fern belonging to the group Medullosales. Genus includes new species D. alvarezii. |
| Ephedrispermum tenuicostatum | Sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (Aptian or early Albian) | | Portugal | A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. |
| Geminispermum | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (Albian) | Potomac Group | United States ( Virginia) | A seed plant belonging to the informal grouping Caytoniales-Umkomasiales-Petriellales. Genus includes new species G. virginiense. |
| Glossopteris thirroulensis | Sp. nov | Valid | McLoughlin & Mays in McLoughlin, Maksimenko & Mays | Permian (Wuchiapingian) | Wilton Formation | Australia | |
| Hirsutisperma | Gen. et sp. nov | Valid | Scott et al. | Carboniferous (Viséan) | | United Kingdom | An ovule adapted for wind dispersal and for deterring herbivory. Genus includes new species H. rothwellii. |
| Huncocladus | Gen. et sp. nov | Valid | Andruchow-Colombo, Wilf & Escapa | Early Eocene | La Huitrera Formation | Argentina | A seed plant of uncertain phylogenetic placement. Originally described as a member of the family Podocarpaceae related to the genus Phyllocladus; on the other hand, Dörken et al. (2021) rejected the podocarpaceous affinity of Huncocladus, and considered it to be more closely related to the cycad genera Bowenia or Eobowenia. Genus includes new species H. laubenfelsii. |
| Illawarraspermum | Gen. et sp. nov | Valid | McLoughlin & Mays in McLoughlin, Maksimenko & Mays | Permian (Wuchiapingian) | Wilton Formation | Australia | A glossopterid seed. Genus includes new species I. ovatum. |
| Kirchmuellia | Gen. et comb. nov | Valid | Anderson et al. | Early Jurassic | | Germany | A seed fern. Genus includes "Umkomasia" franconica Kirchner & Müller (1992). |
| Lepidopteris scassoi | Sp. nov | Valid | Elgorriaga, Escapa & Cúneo | Early Jurassic | Cañadón Asfalto Formation | Argentina | |
| Lignieriopsis | Gen. et 2 sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Aptian-Albian) | Figueira da Foz Formation Potomac Group | Portugal  United States ( Virginia) | A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species L. stenosperma and L. parva. |
| Mariopteris hexiensis | Sp. nov | Valid | Wang et al. | Permian (Cisuralian) | Shanxi Formation | China | Announced in 2019; the final version of the article naming it was published in 2021. |
| Mariopteris yongchangensis | Sp. nov | Valid | Wang et al. | Permian (Cisuralian) | Shanxi Formation | China | Announced in 2019; the final version of the article naming it was published in 2021. |
| Muelkirchium | Gen. et comb. nov | Valid | Anderson et al. | Early Jurassic | | Germany | A seed fern. Genus includes "Pteruchus" septentrionalis Kirchner & Müller (1992). |
| Mutoviaspermum | Gen. et sp. nov | Valid | Karasev et al. | Permian (Lopingian) | Poldarsa Formation | Russia ( Vologda Oblast) | A member of Voltziales. Genus includes new species M. krassilovii. |
| Noeggerathiopsis brasiliensis | Nom. nov | Valid | Degani-Schmidt & Guerra-Sommer | Early Permian | Rio Bonito Formation | Brazil | A member of Cordaitales; a replacement name for Rufloria gondwanensis Guerra-Sommer (1989). |
| Potoniea krisiae | Sp. nov | Valid | Pšenička, Zodrow & Bek | Carboniferous (Moscovian) | Sydney Coalfield | Canada ( Nova Scotia) | Reproductive male organ of a seed fern, possibly a member of the family Parispermaceae. |
| Protophyllocladoxylon zhaobishanensis | Sp. nov | Valid | Wan, Yang & Wang | Early Triassic (Induan) | Jiucaiyuan Formation | China | A silicified gymnospermous fossil wood. |
| Pseudotorellia yimaensis | Sp. nov | Valid | Dong et al. | Middle Jurassic | Yima Formation | China | |
| Ptilophyllum eminelidarum | Sp. nov | Valid | Carrizo, Lafuente Diaz & Del Fueyo | Early Cretaceous | Springhill Formation | Argentina | A member of Bennettitales. |
| Ptilophyllum micropapillosum | Sp. nov | Valid | Lafuente Diaz et al. | Early Cretaceous | Springhill Formation | Argentina | A member of Bennettitales. |
| Rothwellia | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (Albian) | Potomac Group | United States ( Virginia) | A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species R. foveata. |
| Sagenopteris trapialensis | Sp. nov | Valid | Elgorriaga, Escapa & Cúneo | Early Jurassic | Lonco Trapial Formation | Argentina | A member of Caytoniales. |
| Sclerospiroxylon xinjiangensis | Sp. nov | Valid | Wan, Yang & Wang | Permian (Kungurian) | Hongyanchi Formation | China | |
| Sueria laxinervis | Sp. nov | Valid | Yamada & Nishida in Yamada et al. | Late Cretaceous (Maastrichtian) | Quiriquina Formation | Chile | A cycad. |
| Thodaya | Gen. et sp. nov | Junior homonym | Friis, Crane & Pedersen | Early Cretaceous (Albian) | Potomac Group | United States ( Virginia) | A seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species T. sykesiae. The generic name is preoccupied by Thodaya Compton. |
| Umaltolepis yimaensis | Sp. nov | Valid | Dong et al. | Middle Jurassic | Yima Formation | China | |
| Umkomasia corniculata | Sp. nov | Valid | Shi et al. | Early Cretaceous (Aptian–Albian) | | Mongolia | |
| Umkomasia trilobata | Sp. nov | Valid | Shi et al. | Early Cretaceous (Aptian–Albian) | | Mongolia | |
| Wangjunia | Gen. et sp. nov | Valid | Backer, Bomfleur & Kerp | Permian (Guadalupian) | Lower Shihhotse Formation | China | A member of Cordaitales. Genus includes new species W. microphylla. |
| Xuanweioxylon damogouense | Sp. nov | Valid | Yang et al. | Permian (Lopingian) | Xuanwei Formation | China | A conifer stem. |
| Zhangwuia | Gen. et sp. nov | Valid | Liu, Hou & Wang | Middle Jurassic (Callovian) | Jiulongshan Formation | China | A reproductive organ of a seed plant of uncertain phylogenetic placement. Genus includes new species Z. mira. |

## Flowering plants
### Basal angiosperms
#### Nymphaeales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Nuphaea | Gen. et sp. nov | Valid | Gee & Taylor | Eocene | Messel pit | Germany | A member of Nymphaeaceae. Genus includes new species N. engelhardtii. | |

#### Other basal angiosperms

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Anaspermum | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | A flowering plant with affinities to Austrobaileyales or Nymphaeales. Genus includes new species A. operculatum. | |
| Gastonispermum antiquum | Sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | A flowering plant with affinities to Austrobaileyales or Nymphaeales. | |

### Monocots
#### Alismatales

| Name | Novelty | Status | Authors | Age | Location | Notes |
| Natantisphyllum | Gen. et sp. nov | Valid | Puebla, Vento & Prámparo | Late Cretaceous | Argentina | A member of the family Araceae. Genus includes new species N. crenae. Announced in 2019; the final version of the article naming it was published in 2021. |
| Orontiophyllum ferreri | Sp. nov | Valid | Sender et al. | Early Cretaceous (Albian) | Spain | A member or a relative of the family Araceae. |
| Turolospadix | Gen. et sp. nov | Valid | Sender et al. | Early Cretaceous (Albian) | Spain | A member or a relative of the family Araceae. Genus includes new species T. bogneri. |

#### Arecales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Palmoxylon deoriensis | Sp. nov | Valid | Khan, Mandal & Bera | Late Cretaceous (late Maastrichtian) – early Paleocene (Danian) | Deccan Intertrappean Beds | India | A permineralized palm stem. |
| Sabalites tibetensis | Sp. nov | Valid | Su & Zhou in Su et al. | Oligocene (Chattian) | Lunpola Basin | China | A member of the family Arecaceae belonging to the subfamily Coryphoideae. |
| Sclerosperma protoprofizianum | Sp. nov | Valid | Grímsson & Zetter in Grímsson et al. | Late Oligocene | | Ethiopia | A species of Sclerosperma. |
| Sclerosperma protomannii | Sp. nov | Valid | Grímsson & Zetter in Grímsson et al. | Late Oligocene | | Ethiopia | A species of Sclerosperma. |
| Spinopalmoxylon cicatricosum | Sp. nov | Valid | Winterscheid | Oligocene | Köln Formation | Germany | A member of the family Arecaceae belonging to the tribe Calameae. |
| Spinopalmoxylon parvifructum | Sp. nov | Valid | Winterscheid | Oligocene | Köln Formation | Germany | A member of the family Arecaceae belonging to the tribe Calameae. |
| Spinizonocolpites riochiquensis | Sp. nov | Valid | Vallati & De Sosa Tomas in Vallati, De Sosa Tomas & Casal | Late Cretaceous (Maastrichtian) | Lago Colhué Huapí Formation | Argentina | A member of Arecaceae described on the basis of fossil pollen grains. Announced in 2019; the final version of the article naming it was published in 2020. |

#### Dioscoreales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Dioscorea eocenicus | Sp. nov | Valid | Mehrotra & Shukla | Early Eocene | | India | A species of Dioscorea. | |
| Dioscorea manchesteri | Sp. nov | Valid | Kvaček | Miocene | Most Formation | Czech Republic | A species of Dioscorea. | |

#### Poales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Bambusiculmus makumensis | Sp. nov | Valid | Srivastava et al. | Late Oligocene | | India | A bamboo. |
| Bambusiculmus tirapensis | Sp. nov | Valid | Srivastava et al. | Late Oligocene | | India | A bamboo. |
| Bambusium arunachalense | Sp. nov | Valid | Srivastava et al. | Late Miocene to Pliocene | | India | A bamboo. |
| Bambusium deomarense | Sp. nov | Valid | Srivastava et al. | Late Miocene to Pliocene | | India | A bamboo. |
| Scirpus weichangensis | Sp. nov | Valid | Liang in Lu et al. | Early Miocene | Hannuoba Formation | China | A species of Scirpus. |

### Magnoliids
#### Laurales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Cinnamomum raptiensis | Sp. nov | Valid | Prasad et al. | Middle Miocene | Lower Churia Group | Nepal | A species of Cinnamomum. |
| Laurinoxylon acalensis | Sp. nov | Valid | Pérez-Lara, Estrada-Ruiz & Castañeda-Posadas | Eocene | El Bosque Formation | Mexico | A member of Lauraceae. |
| Laurinoxylon thomasii | Sp. nov | Valid | Akkemik in Akkemik, Akkılıç & Güngör | Early Miocene | | Turkey | |
| Laurophyllum alseodaphnoides | Sp. nov | Valid | Wang & Sun in Wang et al. | Miocene (Langhian) | Fotan Group | China | A member of Lauraceae described on the basis of fossil leaves. |
| Laurophyllum fotanensis | Sp. nov | Valid | Wang & Sun in Wang et al. | Miocene (Langhian) | Fotan Group | China | A member of Lauraceae described on the basis of fossil leaves. |
| Laurophyllum lindaiensis | Sp. nov | Valid | Wang & Sun in Wang et al. | Miocene (Langhian) | Fotan Group | China | A member of Lauraceae described on the basis of fossil leaves. |
| Laurophyllum triangulatum | Sp. nov | Valid | Wang & Sun in Wang et al. | Miocene (Langhian) | Fotan Group | China | A member of Lauraceae described on the basis of fossil leaves. |
| Laurophyllum zhangpuensis | Sp. nov | Valid | Wang & Sun in Wang et al. | Miocene (Langhian) | Fotan Group | China | A member of Lauraceae described on the basis of fossil leaves. |
| Mezilaurinoxylon oleiferum | Sp. nov | Valid | Ruiz, Brea & Pujana in Ruiz et al. | Paleocene (Danian) | Salamanca Formation | Argentina | A member of the family Lauraceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020. |
| Patagonoxylon | Gen. et sp. nov | Valid | Ruiz, Brea & Pujana in Ruiz et al. | Paleocene (Danian) | Salamanca Formation | Argentina | A Lauralean of uncertain phylogenetic placement. The type species is P. scalariforme. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020. |
| Persea masotkholaensis | Sp. nov | Valid | Prasad et al. | Middle Miocene | Lower Churia Group | Nepal | A species of Persea. |

#### Magnoliales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Anaxagorea mioluzonensis | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Anaxagorea. |
| Anonaspermum orientalis | Sp. nov | Valid | Li et al. | Late Oligocene | Yongning Formation | China | A member of the family Annonaceae. |
| Mitrephora mioreticulata | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Mitrephora. |
| Riaselis | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Aptian-early Albian or older) | | Portugal | Genus includes new species R. rugosa. |
| Serialis | Gen. et 9 sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Albian) | Almargem Formation Figueira da Foz Formation | Portugal | Genus includes new species S. antiqua, S. parva, S. elongata, S. tenuitesta, S. communis, S. crassitesta, S. grossa, S. undata and S. reticulata. |
| Uvaria miolucida | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Uvaria. |

#### Piperales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Appofructus | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | Genus includes new species A. nudus. |
| Appomattoxia minuta | Sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | |
| Burgeria | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | Genus includes new species B. striata. |
| Dejaxia | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | Genus includes new species D. brevicolpites. |
| Goczania | Gen. et 3 sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | Genus includes new species G. rugosa, G. inaequalis and G. punctata. |

### Unplaced non-eudicots
#### Chloranthales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Canrightia elongata | Sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | |
| Hedyflora | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Aptian–early Albian) | Figueira da Foz Formation | Portugal | A member of the family Chloranthaceae. Genus includes new species H. crystallifera. |
| Kvacekispermum costatum | Sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | |

### Basal eudicots
#### Proteales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Meliosma berryi | Sp. nov | Valid | Huegele & Manchester | Probably late Eocene | | United States ( Texas) | A species of Meliosma. |
| Platanus heilongjiangensis | Sp. nov | Valid | Sun et al. | Late Cretaceous (China) | Houshiigou Formation | China | A species of Platanus. |
| Scalarixylon romeroi | Sp. nov | Valid | Pujana & Ruiz | Eocene–Oligocene | Río Turbio Formation | Argentina | |

#### Ranunculales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Tinospora siwalika | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Tinospora. | |

### Superasterids
#### Aquifoliales

| Name | Novelty | Status | Authors | Age | Location | Notes |
| Ilex angustifolioides | Nom. nov | Valid | Doweld | Miocene | Germany | A holly; a replacement name for Ilex denticulata von Heer (1857). |
| Ilex aschutassica | Nom. nov | Valid | Doweld | Oligocene | Kazakhstan | A holly; a replacement name for Ilex integrifolia Baikovskaja (1956). |
| Ilex boulayi | Nom. nov | Valid | Doweld | Miocene | France | A holly; a replacement name for Ilex undulata Boulay (1887). |
| Ilex friedrichii | Nom. nov | Valid | Doweld | Oligocene | Germany | A holly; a replacement name for Ilex longifolia Friedrich (1884). |
| Ilex latifolioides | Nom. nov | Valid | Doweld | Oligocene | France | A holly; a replacement name for Ilex acuminata Saporta (1865). |
| Ilex mormonica | Nom. nov | Valid | Doweld | Oligocene | United States ( Montana) | A holly; a replacement name for Ilex acuminata Becker (1960). |
| Ilex opacina | Nom. nov | Valid | Doweld | Oligocene | France | A holly; a replacement name for Ilex microdonta Saporta (1865). |
| Ilex polarica | Nom. nov | Valid | Doweld | Paleocene | Greenland | A holly; a replacement name for Ilex macrophylla von Heer (1869). |
| Ilex subrotunda | Sp. nov | Valid | Doweld | Miocene | Japan | A holly; a replacement name for the previously invalidly published Ilex ohashii Huzioka (1963), lacking holotype designation when published. |

#### Asterales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Cichoreacidites? igapoensis | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil pollen of a member of the genus Pacourina or Vernonia. | |

#### Boraginales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Cordia siwalica | Sp. nov | Valid | Prasad et al. | Middle Miocene | Lower Churia Group | Nepal | A species of Cordia. | |

#### Caryophyllales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Basella keralensis | Sp. nov | Valid | Farooqui, Ray & Garg | Pleistocene | | India | A species of Basella. | |

#### Cornales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Eydeia jerseyensis | Sp. nov | Valid | Atkinson, Martínez & Crepet | Late Cretaceous (Turonian) | | United States ( New Jersey) | | |

#### Ericales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Juddicarpon | Gen. et sp. nov | Valid | Smith & Manchester | Miocene (Burdigalian-Langhian) | Clarkia fossil beds | United States ( Idaho) | A member of Vaccinioideae. Genus includes new species J. benewahensis. |
| Psilastephanocolporites brevissimus | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil pollen of a flowering plant, possibly a member of the genus Myrsine. |
| Sladenia zhengyii | Sp. nov | Valid | Jia & Zhou in Jia et al. | Early Miocene | Maguan Basin | China | A member of the family Sladeniaceae. Announced in 2019; the final version of the article naming it was published in 2021. |
| Symplocos amoena | Sp. nov | Valid | Huegele & Manchester | Probably late Eocene | | United States ( Texas) | A species of Symplocos. |
| Symplocos fritschii | Sp. nov | Valid | Huegele & Manchester | Probably late Eocene | | United States ( Texas) | A species of Symplocos. |
| Symplocos martinettoi | Sp. nov | Valid | Huegele & Manchester | Probably late Eocene | | United States ( Texas) | A species of Symplocos. |
| Symplocos platycarpa | Sp. nov | Valid | Huegele & Manchester | Probably late Eocene | | United States ( Texas) | A species of Symplocos. |
| Symplocos rothwelii | Sp. nov | Valid | Huegele & Manchester | Probably late Eocene | | United States ( Texas) | A species of Symplocos. |
| Symplocos trinitiensis | Sp. nov | Valid | Huegele & Manchester | Probably late Eocene | | United States ( Texas) | A species of Symplocos. |
| Symplocos trisulcata | Sp. nov | Valid | Huegele & Manchester | Probably late Eocene | | United States ( Texas) | A species of Symplocos. |

#### Gentianales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Calycophyllum plengei | Sp. nov | Valid | Woodcock, Meyer & Prado | Eocene | Piedra Chamana Fossil Forest | Peru | A species of Calycophyllum. |
| Psilatriporites aspidatus | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil pollen of a member of the genus Faramea. |
| Randia premacrophylla | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Randia. |

#### Icacinales

| Name | Novelty | Status | Authors | Age | Location | Notes |
| Iodes acuta | Sp. nov | Valid | Del Rio, Stull & De Franceschi | Early Eocene | France | A member of the family Icacinaceae. |
| Iodes parva | Sp. nov | Valid | Del Rio, Thomas & De Franceschi | Late Paleocene | France | A member of the family Icacinaceae. |
| Iodes reidii | Sp. nov | Valid | Del Rio, Thomas & De Franceschi | Late Paleocene | France | A member of the family Icacinaceae. |
| Iodes rigida | Sp. nov | Valid | Del Rio, Stull & De Franceschi | Early Eocene | France | A member of the family Icacinaceae. |
| Iodes rivecourtensis | Sp. nov | Valid | Del Rio, Thomas & De Franceschi | Late Paleocene | France | A member of the family Icacinaceae. |
| Iodes sinuosa | Sp. nov | Valid | Del Rio, Thomas & De Franceschi | Late Paleocene | France | A member of the family Icacinaceae. |
| Iodes tubulifera | Sp. nov | Valid | Del Rio, Thomas & De Franceschi | Late Paleocene | France | A member of the family Icacinaceae. |

### Superrosids
#### Malvids
##### Malvales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Anisoptera palaeoscaphula | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Anisoptera. |
| Ceiba archeopentandra | Sp. nov | Valid | Woodcock, Meyer & Prado | Eocene | Piedra Chamana Fossil Forest | Peru | A species of Ceiba. |
| Ceiba huancabambiana | Sp. nov | Valid | Woodcock, Meyer & Prado | Eocene | Piedra Chamana Fossil Forest | Peru | A species of Ceiba. |
| Dipterocarpus palaeoindicus | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Dipterocarpus. |
| Dryobalanoxylon neosumatrense | Sp. nov | Valid | Biswas, Khan & Bera | Late Miocene | | India | A member of the family Dipterocarpaceae. |
| Grewia americana | Sp. nov | Valid | Woodcock, Meyer & Prado | Eocene | Piedra Chamana Fossil Forest | Peru | A species of Grewia. |
| Grewia nepalensis | Sp. nov | Valid | Prasad et al. | Middle Miocene | Lower Churia Group | Nepal | A species of Grewia. |
| Grewia palaeodisperma | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Grewia. |
| Guazuma santacruzensis | Sp. nov | Valid | Woodcock, Meyer & Prado | Eocene | Piedra Chamana Fossil Forest | Peru | A member of the family Malvaceae. |
| Luehea stratificata | Sp. nov | Valid | Woodcock, Meyer & Prado | Eocene | Piedra Chamana Fossil Forest | Peru | A species of Luehea. |
| Muntingia solapora | Sp. nov | Valid | Woodcock, Meyer & Prado | Eocene | Piedra Chamana Fossil Forest | Peru | A species of Muntingia. |
| Ochroma pozoensis | Sp. nov | Valid | Woodcock, Meyer & Prado | Eocene | Piedra Chamana Fossil Forest | Peru | A species of Ochroma. |
| Sterculia arjunkholaensis | Sp. nov | Valid | Prasad et al. | Middle Miocene | Lower Churia Group | Nepal | A species of Sterculia. |
| Sterculia matrum | Sp. nov | Valid | Woodcock, Meyer & Prado | Eocene | Piedra Chamana Fossil Forest | Peru | A species of Sterculia. |
| Vasivaea weigendii | Sp. nov | Valid | Woodcock, Meyer & Prado | Eocene | Piedra Chamana Fossil Forest | Peru | |

##### Sapindales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Ailanthus maximus | Sp. nov | Valid | Liu, Su & Zhou in Liu et al. | Latest Paleocene to the Late Oligocene | Lunpola Basin Nima Basin | China | A species of Ailanthus. | |
| Antrocaryon panamaensis | Sp. nov | Valid | Herrera et al. | Early Miocene | Cucaracha Formation | Panama | A species of Antrocaryon. | |
| Arytera miolittoralis | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Arytera. | |
| Arytera nepalensis | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Arytera. | |
| Buchanania raptiensis | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Buchanania. | |
| Dodonaea piedra-chamana | Sp. nov | Valid | Woodcock, Meyer & Prado | Eocene | Piedra Chamana Fossil Forest | Peru | A species of Dodonaea. | |
| Dracontomelon montesii | Sp. nov | Valid | Herrera et al. | Early Miocene | Cucaracha Formation | Panama | A species of Dracontomelon. | |
| Erythrochiton masotkholaensis | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Erythrochiton. | |
| Euphoria churiaensis | Sp. nov | Valid | Prasad et al. | Middle Miocene | Lower Churia Group | Nepal | A member of the family Sapindaceae. | |
| Koelreuteria lunpolaensis | Sp. nov | Valid | Jiang et al. | Late Oligocene | Lunpola Basin | China | A species of Koelreuteria. | |
| Rhus asymmetrica | Sp. nov | Valid | Tosal, Sanjuan & Martín-Closas | Early Oligocene | | Spain | A sumac. | |
| Rhus boothillensis | Sp. nov | Valid | Flynn, DeVore & Pigg | Early Eocene | Klondike Mountain Formation | United States ( Washington) | A sumac. | Rhus boothillensi |
| Rhus garwellii | Sp. nov | Valid | Flynn, DeVore & Pigg | Early Eocene | Klondike Mountain Formation | United States ( Washington) | A sumac. | Rhus garwellii |
| Rhus republicensis | Sp. nov | Valid | Flynn, DeVore & Pigg | Early Eocene | Klondike Mountain Formation | United States ( Washington) | A sumac. | |
| Sapindus palaeomukorossi | Sp. nov | Valid | Prasad et al. | Middle Miocene | Lower Churia Group | Nepal | A species of Sapindus. | |
| Spondias rothwellii | Sp. nov | Valid | Herrera et al. | Early Miocene | Cucaracha Formation | Panama | A species of Spondias. | |
| Zanthoxylum reynelii | Sp. nov | Valid | Woodcock, Meyer & Prado | Eocene | Piedra Chamana Fossil Forest | Peru | A species of Zanthoxylum. | |

##### Other malvids

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Akania gibsonorum | Sp. nov | Valid | Conran et al. | Early Miocene | | New Zealand | A member of the family Akaniaceae. |
| Combretum siwalicum | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Combretum. |
| Eugenia nepalensis | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Eugenia. |
| Miconioidea | Gen. et sp. nov | Valid | Woodcock, Meyer & Prado | Eocene | Piedra Chamana Fossil Forest | Peru | A member of the family Melastomataceae. Genus includes new species M. eocenica. |
| Myrceugenellites grandiporosum | Sp. nov | Valid | Ruiz, Brea & Pujana in Ruiz et al. | Paleocene (Danian) | Salamanca Formation | Argentina | A member of the family Myrtaceae. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020. |
| Staphylea ochoterenae | Sp. nov | Valid | Hernández-Damián, Cevallos-Ferriz & Huerta-Vergara | Miocene | | Mexico | A species of Staphylea. |
| Terminalia arjunkholaensis | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Terminalia. |
| Turpinia tiffneyi | Sp. nov | Valid | Huegele & Manchester | Probably late Eocene | | United States ( Texas) | A species of Turpinia. |

#### Fabids
##### Fabales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Arcoa lindgreni | Sp. nov | Valid | Herendeen & Herrera | Eocene | Green River Formation | United States ( Wyoming) | A species of Arcoa. |
| Arcoa linearifolia | Comb. nov | Valid | (Lesquereux) Herendeen & Herrera | Eocene Priabonian | Florissant Formation | United States ( Wyoming) | A species of Arcoa. Moved from Mimosites linearifolius (1878) Originally named Caesalpinia (?) linearifolia (1873) |
| Bauhinia palaeomonandra | Sp. nov | Valid | Prasad et al. | Middle Miocene | Lower Churia Group | Nepal | A species of Bauhinia. |
| Butea nepalensis | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Butea. |
| Cassia arjunkholaensis | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Cassia. |
| Cercioxylon zeynepae | Sp. nov | Valid | Akkemik | Pliocene | Örencik Formation | Turkey | A relative of redbuds described on the basis of fossil wood. |
| Gleditsia europaea | Sp. nov | Valid | Worobiec in Worobiec & Worobiec | Miocene | | Poland | A species of Gleditsia. |
| Hopeoxylon umarsarensis | Sp. nov | Valid | Shukla, Singh & Mehrotra | Early Eocene | Naredi Formation | India | A member of the family Fabaceae belonging to the subfamily Detarioideae. |
| Leguminophyllum kvacekii | Sp. nov | Valid | Worobiec in Worobiec & Worobiec | Miocene | | Poland | Fossil leaflets resembling leaflets of extant and fossil members of Fabaceae. |
| Millettia arjunkholaensis | Sp. nov | Valid | Prasad et al. | Middle Miocene | Lower Churia Group | Nepal | A species of Millettia. |
| Mimosoxylon ceratonioides | Sp. nov | Valid | Akkemik in Akkemik, Akkılıç & Güngör | Early Miocene | | Turkey | |
| Ormosia zhangpuensis | Sp. nov | Valid | Wang et al. | Miocene | | China | A species of Ormosia. |
| Saraca palaeoindica | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Saraca. |
| Sindora eosiamensis | Sp. nov | Valid | Prasad et al. | Middle Miocene | Lower Churia Group | Nepal | A species of Sindora. |
| Sindora leguminocarpoides | Sp. nov | Valid | Prasad et al. | Middle Miocene | Lower Churia Group | Nepal | A species of Sindora. |
| Tzotziloxylon | Gen. et 2 sp. nov | Valid | Pérez-Lara & Estrada-Ruiz in Pérez-Lara, Estrada-Ruiz & Castañeda-Posadas | Eocene | El Bosque Formation | Mexico | A member of the family Fabaceae belonging to the subfamily Cercidoideae or Dialioideae. Genus includes new species T. cristalliferum and T. eocenica. |

##### Fagales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Castanopsis rothwellii | Sp. nov | Valid | Wilf et al. | Eocene | | Argentina | A species of Castanopsis. |
| Casuarinoxylon ildephonsi | Sp. nov | Valid | Vanner | Miocene | | New Zealand | A member of the family Casuarinaceae described on the basis of fossil wood. |
| Engelhardia trinitiensis | Sp. nov | Valid | Huegele & Manchester | Probably late Eocene | | United States ( Texas) | A species of Engelhardia. |
| Pterocaryoxylon tuncayi | Sp. nov | Valid | Akkemik in Akkemik, Akkılıç & Güngör | Early Miocene | | Turkey | |
| Quercus shangcunensis | Sp. nov | Valid | Liu et al. | Early Oligocene | Shangcun Formation | China | An oak |

##### Malpighiales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Calophyllum mioelatum | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Calophyllum. |
| Calophyllum zhangpuensis | Sp. nov | Valid | Wang et al. | Miocene | Fotan Group | China | A species of Calophyllum. |
| Elioxylon | Gen. et sp. nov | Valid | Srivastava, Miller & Baas | Late Cretaceous (Maastrichtian)–Paleocene (Danian) | Deccan Intertrappean Beds | India | A wood morphospecies with features of Achariaceae and Salicaceae. Type species includes new species E. seoniensis. |
| Garcinia zhangpuensis | Sp. nov | Valid | Wang et al. | Middle Miocene | Fotan Group | China | A species of Garcinia. |
| Mascogophyllum | Gen. et sp. nov | Valid | Centeno-González, Porras-Múzquiz & Estrada-Ruiz | Late Cretaceous (late Campanian) | Olmos Formation | Mexico | A possible member of Violaceae. Genus includes new species M. elizondoa. |
| Populus erratica | Nom. nov | Valid | Sachse | Late Oligocene and early Miocene | | Switzerland  France?  Germany?  Hungary? | A species of Populus; a replacement name for Juglans heerii Ettingshausen (1853). |
| Ryparia arjunkholaensis | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A member of the family Achariaceae. |

##### Oxalidales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Caldcluvioxylon torresiae | Sp. nov | Valid | Pujana & Ruiz | Eocene–Oligocene | Río Turbio Formation | Argentina | A member of the family Cunoniaceae. | |
| Tropidogyne lobodisca | Sp. nov | Valid | Poinar & Chambers | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | A probable member of Cunoniaceae. | |

##### Rosales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Artocarpus arjunkholaensis | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Artocarpus. |
| Cedrelospermum tibeticum | Sp. nov | Valid | Jia, Su & Zhou in Jia et al. | Late Oligocene | Dingqing Formation | China | A member of Ulmaceae. |
| Ficus preglobosa | Sp. nov | Valid | Prasad et al. | Middle Miocene | Lower Churia Group | Nepal | A species of Ficus. |
| Frangulops | Gen. et comb. nov | Valid | Doweld | Eocene | | United States ( Colorado) | A member of Rhamnaceae; a new genus for "Ilex" pseudostenophylla Lesquereux (1883). |
| Prunoidoxylon aytugii | Sp. nov | Valid | Akkemik in Akkemik, Akkılıç & Güngör | Early Miocene | | Turkey | |
| Pteroceltis shanwangensis | Sp. nov | Valid | Wong, Dilcher & Uemura | Miocene | Shanwang Formation | China | A species of Pteroceltis. |
| Pteroceltis taoae | Sp. nov | Valid | Wong, Dilcher & Uemura | Miocene | | China | A species of Pteroceltis. |
| Rubus eubaticus | Nom. nov | Valid | Doweld | Miocene | | Bulgaria | A species of Rubus; a replacement name for Rubus mucronatus Palamarev (1987). |
| Rubus primoricus | Nom. nov | Valid | Doweld | Miocene | | Russia ( Primorsky Krai) | A species of Rubus; a replacement name for Rubus ellipticus Pavlyutkin (2005). |
| Ulmus prestonia | Sp. nov | Valid | Lott, Manchester & Corbett | Miocene | | United States ( Florida) | An elm. |

#### Unplaced superrosid eudicots

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Cayratia palaeojaponica | Sp. nov | Valid | Prasad et al. | Late Miocene | Middle Churia Group | Nepal | A species of Cayratia. |
| Liquidambar bella | Sp. nov | Valid | Maslova et al. | Eocene | Huangniuling Formation | China | A Liquidambar species saxifragale. |
| Yua texana | Sp. nov | Valid | Huegele & Manchester | Probably late Eocene | | United States ( Texas) | A Yua species vitale. |

### Other angiosperms

| Name | Novelty | Status | Authors | Type locality | Type locality | Location | Notes |
| Baccatocarpon | Gen. et comb. nov | Valid | Bhowal & Sheikh ex Manchester, Ramteke, Kapgate & Smith | Late Cretaceous (Maastrichtian) | Deccan Intertrappean Beds | India | A fossil fruit of a flowering plant of uncertain affinities; a new genus for "Grewia" mohgaoensis Paradkar & Dixit (1984). |
| Battenipollis sabrinae | Sp. nov | Valid | Smith et al. | Early Paleogene | | Antarctica | An angiosperm pollen species. |
| Bonanzacarpum | Gen. et sp. nov | Valid | Manchester & Lott | Early to middle Eocene | Green River Formation | United States ( Utah) | A eudicot fruit of uncertain phylogenetic placement. The type species is B. sprungerorum. |
| Celastrilex | Gen. et comb. nov | Valid | Doweld | Paleocene | | United States ( Colorado) | A flowering plant of uncertain phylogenetic placement, described on the basis of fossil leaves; a new genus for "Celastrinites" artocarpidioides Lesquereux (1878). |
| Choffaticarpus | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | A flowering plant of uncertain phylogenetic placement. Genus includes new species C. compactus. |
| Cratolirion | Gen. et sp. nov | Valid | Coiffard, Kardjilov et Bernardes-de-Oliveira in Coiffard et al. | Early Cretaceous | Crato Formation | Brazil | A crown monocot of uncertain phylogenetic placement. Genus includes new species C. bognerianum. |
| Dalembia (?) gracilis | Sp. nov | Valid | Herman in Herman et al. | Late Cretaceous (Turonian-Coniacian) | Derevyannye Gory Formation | Russia ( Sakha Republic) | A flowering plant described on the basis of fossil leaves. |
| Dictyozonia | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | A non-eudicot flowering plant of uncertain affinity. Genus includes new species D. pusilla. |
| Dinisia | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | A non-eudicot flowering plant of uncertain affinity. Genus includes new species D. portugallica. |
| Dispariflora | Gen. et sp. nov | Valid | Poinar & Chambers | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | A flowering plant of uncertain phylogenetic placement, possibly a relative of members of Laurales, especially Southern Hemisphere families allied with the Monimiaceae. Genus includes new species D. robertae. |
| Eckhartia | Gen. et 3 sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | A non-eudicot flowering plant of uncertain affinity. Genus includes new species E. brevicolumella, E. longicolumella and E. intermedia. |
| Eckhartianthus | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | A non-eudicot flowering plant of uncertain affinity. Genus includes new species E. lusitanicus. |
| Eckhartiopsis | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | A non-eudicot flowering plant of uncertain affinity. Genus includes new species E. parva. |
| Exalloanthum | Nom. nov | Valid | Poinar | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | A flowering plant of uncertain phylogenetic placement; a replacement name for Diaphoranthus Poinar (2018). |
| Gambierina askiniae | Sp. nov | Valid | Smith et al. | Early Paleogene | | Antarctica | An angiosperm pollen species. |
| Herendeenoxylon | Gen. et sp. nov | Valid | Chin et al. | Late Cretaceous (Turonian) | Moreno Hill Formation | United States ( New Mexico) | A flowering plant of uncertain phylogenetic placement (possibly a member of Ericales), described on the basis of fossil wood. Genus includes new species H. zuniense. |
| Ibrahimia | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | An eudicot of uncertain phylogenetic placement, possibly related to Paisia. Genus includes new species I. vermiculata. |
| Juhaszia | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | A non-eudicot flowering plant of uncertain affinity. Genus includes new species J. portugallica. |
| Kempia | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | A non-eudicot flowering plant of uncertain affinity. Genus includes new species K. longicolpites. |
| Ladakhipollenites? densicolumellatus | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil pollen of a flowering plant. |
| Ladakhipollenites? lolongatus | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil pollen of Symmeria paniculata. |
| Ladakhipollenites? porolenticularis | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil pollen of a flowering plant (possibly a member of the family Marcgraviaceae). |
| Lagokarpos tibetensis | Sp. nov | Valid | Tang, Su & Zhou in Tang et al. | Paleogene | Niubao Formation | China | A fossil fruit with unknown modern affinities. |
| Mcdougallia | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | An eudicot of uncertain phylogenetic placement. Genus includes new species M. irregularis. |
| Nicholsia | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | An eudicot of uncertain phylogenetic placement. Genus includes new species N. brevicolpites. |
| Piercipollis | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | A non-eudicot flowering plant of uncertain affinity. Genus includes new species P. simplex. |
| Reyanthus | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | A flowering plant of uncertain phylogenetic placement, possibly related to Magnoliales. Genus includes new species R. lusitanicus. |
| Rhoipites? basicus | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil pollen of a flowering plant. |
| Rhoipites manausensis | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil pollen of a member of the genus Schefflera. |
| Rhoipites minuticirculus | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil pollen of a flowering plant. |
| Rhoipites negroensis | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil pollen of a flowering plant. |
| Samylinaea | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | An eudicot of uncertain phylogenetic placement. Genus includes new species S. punctata. |
| Sherwinoxylon | Gen. et sp. nov | Valid | Boura & Saulnier in Boura et al. | Late Cretaceous (Cenomanian) | | France | A vesselless angiosperm fossil wood of uncertain affinity. Genus includes new species S. winteroides. |
| Strombothelya | Gen. et 2 sp. nov | Valid | Poinar & Chambers | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | A flowering plant of uncertain phylogenetic placement. Genus includes new species S. monostyla and S. grammogyna. |
| Teebacia | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | A non-eudicot flowering plant of uncertain affinity. Genus includes new species T. hughesii. |
| Ubiquitoxylon | Gen. et sp. nov | Valid | Wheeler in Wheeler, Brown & Koch | Late Paleocene | Denver Formation | United States ( Colorado) | A dicotyledonous flowering plant of uncertain phylogenetic placement, described on the basis of fossil wood. Genus includes new species U. raynoldsii. |
| Vasunum | Gen. et sp. nov | Valid | Chin et al. | Late Cretaceous (Turonian) | Moreno Hill Formation | United States ( New Mexico) | A flowering plant of uncertain phylogenetic placement, described on the basis of fossil wood. Genus includes new species V. cretaceum. |
| Vedresia | Gen. et sp. nov | Valid | Friis, Crane & Pedersen | Early Cretaceous (late Barremian-early Aptian) | Almargem Formation | Portugal | A flowering plant of uncertain phylogenetic placement, possibly related to Chloranthales. Genus includes new species V. elliptica. |
| Zygadelphus | Gen. et sp. nov | Valid | Poinar & Chambers | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | A flowering plant of uncertain phylogenetic placement, possibly a member of Laurales. Genus includes new species Z. aetheus. |

## Other plants

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Acetabularia moldavica | Sp. nov | Valid | Barattolo, Ionesi & Ţibuleac | Middle Miocene | | Romania | A green alga belonging to the family Polyphysaceae, a species of Acetabularia. |
| Aloisalthella | Gen. et comb. nov | Valid | Granier in Granier & Lethiers | Late Jurassic and Early Cretaceous (Berriasian) | | Algeria  France  Spain  Ukraine  United Arab Emirates | A green alga belonging to the family Polyphysaceae; a new genus for "Actinoporella" sulcata von Alth (1882). |
| Aneurospora posongchongensis | Sp. nov | Valid | Cascales-Miñana et al. | Early Devonian | Posongchong Formation | China | A spore taxon. |
| Auerbachichara tataouinensis | Sp. nov | Valid | Tiss et al. | Middle Jurassic (Callovian) | Foum Tataouine Formation | Tunisia | A green alga belonging to the group Charophyta. |
| Bakalovaella deloffrei | Sp. nov | Valid | Granier & Bucur | Early Cretaceous (Hauterivian) | | France | A green alga belonging to the family Dasycladaceae. |
| Buthograptus gundersoni | Sp. nov | Valid | LoDuca | Ordovician (Sandbian) | Platteville Formation | United States ( Wisconsin) | A green alga belonging to the group Bryopsidales. |
| Buthograptus meyeri | Sp. nov | Valid | LoDuca | Ordovician (Sandbian) | Platteville Formation | United States ( Wisconsin) | A green alga belonging to the group Bryopsidales. |
| Calcipatera schoenlaubi | Sp. nov | Valid | Vachard in Krainer, Vachard & Schaffhauser | Permian | | Austria  Oman?  United States? ( New Mexico) | A green alga belonging to the group Bryopsidales and the family Anchicodiaceae. |
| Callixylon wendtii | Sp. nov | Valid | Tanrattana, Meyer-Berthaud & Decombeix | Devonian (Famennian) | | Morocco | An archaeopteridalean progymnosperm. |
| Cingulatisporites oligodistalis | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil spore. |
| Concavissimisporites varzeanus | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil spore. |
| Coniopteris moguqiensis | Sp. nov | Valid | Zhang, Liu & Liang | Middle Jurassic | Wanbao Formation | China | |
| Dissocladella? chahtorshiana | Sp. nov | Valid | Rashidi & Schlagintweit in Schlagintweit et al. | Paleocene | | Iran | A green alga belonging to the family Dasycladaceae. |
| Dissocladella compressa | Sp. nov | Valid | Rashidi & Schlagintweit | Late Cretaceous (Maastrichtian) | Tarbur Formation | Iran | A green alga belonging to the group Dasycladales. |
| Echinatisporis parviechinatus | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil spore. |
| Echinosporis conicus | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil spore. |
| Electrophycus | Gen. et sp. nov | Valid | Poinar & Brown | Late Cretaceous (Cenomanian) | Burmese amber | Myanmar | A green alga, possibly a member of the family Chaetophoraceae. Genus includes new species E. astroplethus. Announced in 2019; the final version of the article naming it was published in 2021. |
| Epiastopora | Gen. et comb. nov | Valid | Vachard in Krainer, Vachard & Schaffhauser | Carboniferous (Pennsylvanian) and Permian | | | A green alga belonging to the group Dasycladales and the family Seletonellaceae. A new genus for "Epimastopora" alpina Kochansky & Herak (1960) and several other species formerly assigned to the genera Epimastopora and Pseudoepimastopora. |
| Jowingera | Gen. et sp. nov | Valid | Bickner & Tomescu | Devonian (Emsian) | Battery Point Formation | Canada ( Quebec) | An early euphyllophyte. Genus includes new species J. triloba. |
| Leonophyllum | Gen. et sp. nov | Valid | Barbacka & Kustatscher in Barbacka et al. | Early Jurassic | | Hungary | A plant of uncertain phylogenetic placement, showing similarities to thalloid liverworts with raised vegetative bodies and to the fern family Hymenophyllaceae. Genus includes new species L. tenellum. |
| Leptocentroxyla | Gen. et sp. nov | Valid | Bickner & Tomescu | Devonian (Emsian) | Battery Point Formation | Canada ( Quebec) | An early euphyllophyte. Genus includes new species L. tetrarcha. |
| Maiaspora | Gen. et sp. nov | Valid | Mamontov et al. | Carboniferous (Viséan) | Moscow Syneclise | Russia | A miospore. Genus includes new species M. panopta. Announced in 2019; the final version of the article naming it was published in 2021. |
| Ninsaria | Gen. et sp. nov | Valid | Decombeix, Galtier, McLoughlin & Meyer-Berthaud in Decombeix et al. | Carboniferous (Viséan) | Rockhampton Group | Australia | A vascular plant belonging to the group Lignophytia, of uncertain phylogenetic placement within the latter group. Genus includes new species N. australiana. |
| Palambages pariunta | Sp. nov | Valid | Wainman et al. | Late Jurassic (late Kimmeridgian–early Tithonian) | Surat Basin | Australia | A colonial alga belonging to the group Chlorophyta. |
| Patruliuspora | Gen. et comb. nov | Valid | Barattolo, Ionesi & Ţibuleac | Late Triassic to Miocene | | Czech Republic  France  Slovakia | A green alga belonging to the family Polyphysaceae. Genus includes "Chalmasia" morelleti Pokorný (1948), "Halicoryne" carpatica Mišík (1987) and "Acicularia" valeti Segonzac (1970). |
| Porochara schudackii | Sp. nov | Valid | Tiss et al. | Middle Jurassic (Bajocian) | Krachoua Formation | Tunisia | A green alga belonging to the group Charophyta. |
| Pseudocymopolia acuta | Sp. nov | Valid | Rashidi & Schlagintweit | Late Cretaceous (Maastrichtian) | Tarbur Formation | Iran | A green alga belonging to the group Dasycladales and to the family Triploporellaceae. |
| Stenoloboxyla | Gen. et sp. nov | Valid | Bickner & Tomescu | Devonian (Emsian) | Battery Point Formation | Canada ( Quebec) | An early euphyllophyte. Genus includes new species S. ambigua. |
| Tainioxyla | Gen. et sp. nov | Valid | Bickner & Tomescu | Devonian (Emsian) | Battery Point Formation | Canada ( Quebec) | An early euphyllophyte. Genus includes new species T. quebecana. |
| Tichavekia | Gen. et sp. nov | Valid | Pšenička, Sakala & Kraft in Kraft et al. | Late Silurian | Prague Basin | Czech Republic | A large early land plant. Genus includes new species T. grandis. |
| Uteria naghanensis | Sp. nov | Valid | Rashidi & Schlagintweit | Late Cretaceous (Maastrichtian) | Tarbur Formation | Iran | A green alga belonging to the family Polyphysaceae. |
| Verrucatotriletes laesuraverrucatus | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil spore. |
| Verrucatotriletes tortus | Sp. nov | Valid | D'Apolito et al. | Pliocene–Pleistocene | | Brazil | Fossil spore. |

## General research

* Description of fossils of filamentous green algae from the Early Devonian Rhynie chert (Scotland) is published by Wellman, Graham & Lewis (2019).
* Cretaceous alga Falsolikanella campanensis, originally assigned to the tribe Diploporeae within the green alga order Dasycladales, is transferred to the genus Actinoporella within the tribe Acetabularieae, family Polyphysaceae by Barattolo et al. (2019).
* A study on the impact of the Cretaceous–Paleogene extinction event on European charophytes is published by Vicente, Csiki-Sava & Martín-Closas (2019).
* The oldest known trilete spore assemblages reported so far are described from the Sandbian successions from Motala (central Sweden) by Rubinstein & Vajda (2019).
* A study on the composition and distribution of dispersed spore assemblages from Middle Devonian deposits of northern Spain, and on their implications for inferring the nature of the Kačák Event, is published by Askew & Wellman (2019).
* A study on the morphology of the spore taxon Lagenoisporites magnus from the Carboniferous (Tournaisian) Toregua Formation (Bolivia) is published by Quetglas, Macluf & di Pasquo (2019).
* A review of research concerning early evolution of land plants during the Ordovician is published by Servais et al. (2019).
* A study on carbon isotope data from stratigraphic sections at Germany Valley (West Virginia) and Union Furnace (Pennsylvania) in the Central Appalachian Basin, evaluating its implications for the knowledge of change in atmospheric oxygen levels during the late Ordovician and its possible relationship with early diversification of land plants, is published by Adiatma et al. (2019).
* A study on the stable carbon isotopic composition of 190 fossil specimens belonging to 12 genera of Devonian and Early Carboniferous land plants is published by Wan et al. (2019).
* A study on the early evolution of vascular plants is published by Cascales-Miñana et al. (2019).
* A study on the evolution of early vascular plants is published by Crepet & Niklas (2019).
* A study on the fine-scale structure and the chemistry of the tracheids of the earliest known woody plant Armoricaphyton chateaupannense is published by Strullu-Derrien et al. (2019).
* A study on diversity and functions of lycopsid reproductive structures through time, based on data from extant and fossil taxa, is published by Bonacorsi & Leslie (2019).
* Redescription of the morphology of sterile and fertile structures of the Devonian lycopsid Kossoviella timanica is published by Orlova et al. (2019).
* A study on the ultrastructure of the spore wall in the Carboniferous lycopsid Oxroadia gracilis is published by Taylor (2019).
* A slab containing rooting systems which probably belonged to rhizomorphic lycopsids is reported from the Lower Permian Abo Formation (New Mexico, United States) by Hetherington et al. (2019).
* A study on the anatomy and affinities of Cheirostrobus pettycurensis is published by Neregato & Hilton (2019), who report the discovery of spores conforming to the species Retusotriletes incohatus associated with fossils of Cheirostrobus, representing the first discovery of Retusotriletes-type spores reported in situ within sphenophytes.
* A study on the anatomy and affinities of silicified stems of Sphenophyllum from the Tournaisian deposits in the Montagne Noire region of France and in the Saalfeld area in Germany is published by Terreaux de Felice, Decombeix & Galtier (2019).
* Fossils assigned to the genus Equisetum are reported from a new fossil plant assemblage of late Eocene or early Oligocene age from central Queensland (Australia) by Rozefelds et al. (2019), representing the first evidence of this genus from the Cenozoic of Australia and the most recent fossil record of this genus from Australia.
* A study on the evolutionary history of horsetails, based on genetic data and fossil record, is published by Clark, Puttick & Donoghue (2019), who report evidence indicative of two successive whole-genome duplication events occurring during the Carboniferous and Triassic rather than in association with the Cretaceous–Paleogene extinction event.
* A study aiming to determine links between volcanic activity in the Central Atlantic magmatic province, elevated concentrations of mercury in marine and terrestrial sediments and abnormalities of fossil fern spores across the Triassic-Jurassic boundary in southern Scandinavia and northern Germany is published by Lindström et al. (2019).
* A study on the fossil record of fern spores at the Cretaceous-Paleogene boundary, on the viability of fern spores, and on their implications for the knowledge of the duration of the impact winter at the Cretaceous-Paleogene boundary is published by Berry (2019).
* A study on the molecular structural characteristics of organic remains of a fern belonging to the family Osmundaceae from the Early Jurassic Korsaröd site in southern Sweden is published by Qu et al. (2019).
* A study on anatomy and growth of large specimens of the fossil fern species Weichselia reticulata from the Barremian La Huérguina Formation (Spain) is published by Blanco-Moreno et al. (2019).
* A study on the morphological characters of 42 fossil species of Dicksoniaceae from China, and on their implications for the taxonomy of the fossil members of this group, is published by Xin et al. (2019).
* Fossil occurrences of members of the genus Christella are reported from the late Paleocene of Liuqu, southern Tibet and middle Miocene of the Jinggu Basin in western Yunnan (China) by Xu et al. (2019), who transfer the species "Cyclosorus" nervosus Tao (1988) to the genus Christella.
* A study on the fossils of Glossopteris from the Permian succession of eastern India, aiming to identify the molecular signatures of solvent-extractable and non-extractable organic matter, will be published by Tewari et al. (2019).
* A study on the diversity trends of Glossopteris flora from the Barakar, Raniganj, and Panchet formations of Tatapani–Ramkola Coalfield (India) is published by Saxena et al. (2019).
* A study on the architecture of the ovuliferous reproductive organs of Permian glossopterids is published by Mcloughlin & Prevec (2019).
* A study on the pinnule and stomatal morphology of extant and fossil members of the genera Bowenia and Eobowenia, and on its implications for the knowledge of adaptations of fossil plants to different environments, is published by Hill, Hill & Watling (2019).
* Seed of the ginkgoalean Yimaia capituliformis with damage interpreted as likely oviposition lesions inflicted by a kalligrammatid lacewing is described from the Middle Jurassic Jiulongshan Formation (China) by Meng et al. (2019).
* A study on the phytogeographic history of ten conifer genera that are endemic to East Asia, based on fossil data from humid temperate forests in the Japanese Islands and Korean Peninsula, is published by Yabe et al. (2019).
* A study on the evolution of male and female cone sizes in members of the family Araucariaceae, as indicated by data from extant and fossil members of this family, is published by Gleiser et al. (2019).
* Five fossil foliage specimens of Calocedrus lantenoisi, representing one of the earliest records of the genus Calocedrus worldwide, are described from the Oligocene Shangcun Formation of the Maoming Basin (Guangdong Province, South China) by Wu et al. (2019).
* Leaves including cuticles and ovuliferous cones of members of the genus Metasequoia are described from the middle Miocene of Zhenyuan, Yunnan (Southwest China) by Wang et al. (2019), comprising the southernmost fossil record of this genus worldwide.
* A review of the fossil record of woods which might have affinities with Taxaceae, and a study on the palaeobiogeographical history of this family, is published by Philippe et al. (2019).
* Putative Cretaceous siliceous sponge Siphonia bovista is reinterpreted as an internal mould of the cone-like plant fossil Dammarites albens by Niebuhr (2019).
* A review of epidermal features of bennettites, comparing them with analogous features in living taxa and aiming to identify homologous character states, is published by Rudall & Bateman (2019).
* The first fossil record of a cycad seedling found in close association with a leaf flush of an adult cycad plant of the same species (Dioonopsis praespinulosa) is reported from the Palaeocene (Danian) Castle Rock flora in the Denver Basin (Colorado, United States) by Erdei et al. (2019).
* A review of the paleobotanical evidence of the age and early history of the flowering plants is published by Coiro, Doyle & Hilton (2019).
* A study aiming to establish when the flowering plants originated is published by Li et al. (2019).
* Presence of endothelium (a specialized seed tissue that develops from the inner epidermis of the inner integument) is reported in several different kinds of flowering plant seeds (including in the lineage leading to extant Chloranthaceae) from the Early Cretaceous of eastern North America and Portugal by Friis, Crane & Pedersen (2019).
* A study on the phylogenetic relationships of palm fruit fossils from the Cretaceous–Paleogene (Maastrichtian–Danian) Deccan Intertrappean Beds (India) is published by Matsunaga et al. (2019), who interpret these fossils as representing a crown group member of palm subtribe Hyphaeninae (tribe Borasseae, subfamily Coryphoideae) related to extant genera Satranala and Bismarckia.
* Fossil fruits of members of the genera Fragaria and Rubus are reported from the Pliocene outcrops in the Heqing Basin (China) by Huang et al. (2019).
* Description of alder leaf and infructescence fossils from the Upper Eocene Lawula Formation (Qinghai–Tibetan Plateau) is published by Xu, Su & Zhou (2019).
* A study on the morphology, paleoecology, historical biogeography and phylogenetic relationships of fossil pollen of members of Malvaceae belonging to the species Rhoipites guianensis and Malvacipolloides maristellae, and on its implications for inferring the impact of Cenozoic geological processes (including the uplift of the Andes) on members of Malvaceae living in northern South America, is published by Hoorn et al. (2019).
* A study aiming to determine the location of refugia of two North American species of hickories during the Last Glacial Maximum on the basis of genomic data is published by Bemmels, Knowles & Dick (2019).
* A study on functional leaf traits of the Eocene-Miocene taxa Rhodomyrtophyllum reticulosum (family Myrtaceae) and Platanus neptuni (family Platanaceae), evaluating whether leaf traits of these taxa reflect environmental conditions including climate, is published by Moraweck et al. (2019).
* A study on the morphology and phylogenetic relationships of Eocene fruits belonging to the species Mastixicarpum crassum and Eomastixia bilocularis is published by Manchester & Collinson (2019).
* Seeds of Eurya stigmosa are reported from the Early Pleistocene lacustrine and fluvial sediments of Porto da Cruz, Madeira by Góis-Marques et al. (2019).
* A study on the putative cycad "Zamia" australis from the Miocene Ñirihuau Formation (Argentina) is published by Passalia, Caviglia & Vera (2019), who reinterpret the fossil specimens as flowering plant leaves, and transfer this species to the genus Lithraea.
* New method for reconstructing water transport properties of fossil wood is proposed by Tanrattana et al. (2019).
* Signatures of Devonian (Famennian) forests and soils preserved in black shales in the southernmost Appalachian Basin (Chattanooga Shale; Alabama, United States) are presented by Lu et al. (2019).
* A study on reproductive structures of Devonian plants and on their implications for the knowledge of large-scale patterns of reproductive evolution over the Devonian is published by Bonacorsi & Leslie (2019).
* Revision of a fossil plant assemblage from the Carboniferous site in San Juan Province, Argentina known as Retamito or Río del Agua is published by Correa & Césari (2019).
* A study on the stratigraphic ranges and diversities of plant taxa from the upper Permian (Lopingian) to the Middle Triassic is published by Nowak, Schneebeli-Hermann & Kustatscher (2019), who interpret their findings as indicating that the extinction of land plants during the Permian–Triassic extinction event was much less severe than previously thought.
* A study on the timing of the collapse of the Permian Glossopteris flora from the Sydney Basin (Australia) is published by Fielding et al. (2019).
* New fossil flora dominated by cuticles of Dicroidium is reported from the Middle Triassic (Anisian) Mukheiris Formation (Jordan) by Abu Hamad et al. (2019).
* A study on changes of land vegetation resulting from the Toarcian oceanic anoxic event is published by Slater et al. (2019).
* Plant disseminules are documented from four Middle Jurassic to Lower Cretaceous lacustrine Lagerstätten in China and Australia by McLoughlin & Pott (2019).
* A study comparing the Jurassic floras of the Ayuquila Basin and the Otlaltepec Basin (Mexico) and evaluating their implications for the knowledge of the Jurassic environments of these basins is published by Velasco-de León et al. (2019).
* A study on phototropism in extant trees from Beijing and Jilin Provinces and fossil tree trunks from the Jurassic Tiaojishan and Tuchengzi formations in Liaoning and Beijing regions (China), and on its implications for inferring the history of the rotation of the North China Block, is published by Jiang et al. (2019).
* A study on the link between climatic changes and changes plant distribution in South America during the Early Cretaceous, as indicated by palynological data from the Aptian of the Sergipe Basin (Brazil), is published by Carvalho et al. (2019).
* A study on the frequency and diversity of damage types caused by insect oviposition in plants from the Upper Triassic Yangcaogou Formation, Middle Jurassic Jiulongshan Formation and Lower Cretaceous Yixian Formation (China), assessing the degree of plant host specificity, is published by Lin et al. (2019).
* A study on the plant specimens (ferns, gymnosperms and angiosperms) from the Lower Cretaceous Araripe Basin (Brazil) preserving evidence of plant–insect interactions and potentially of paleoecological relationships between plants and insects is published by Edilson Bezerra dos Santos Filho et al. (2019).
* Leaves of members of the family Nymphaeaceae preserving evidence of insect herbivory are reported from the Albian Utrillas Formation (Spain) by Estévez-Gallardo et al. (2019).
* A study on Cenomanian plants from the Redmond no.1 mine near Schefferville (Redmond Formation; Labrador Peninsula, Canada) and on their implications for the knowledge of paleoclimate of this site is published by Demers-Potvin & Larsson (2019).
* A study on the canopy structure of Late Cretaceous and Paleocene forests in South America, as indicated by the carbon isotope composition of fossil angiosperm leaves from two localities in the Paleocene Cerrejón Formation and one locality in the Maastrichtian Guaduas Formation (Colombia), is published by Graham et al. (2019).
* A quantitative analysis of an earliest Paleocene megaflora from the Ojo Alamo Sandstone in the San Juan Basin (New Mexico, United States) is published by Flynn & Peppe (2019).
* A study on the evolution of plant assemblages in the area of Primorye (Russia) throughout the Paleogene is published by Bondarenko, Blokhina & Utescher (2019).
* A study on changes in plant and insect communities across the Paleocene–Eocene boundary within the Hanna Basin (Wyoming, United States) is published by Azevedo Schmidt et al. (2019).
* A study on stomata of fossil specimens of members of the family Lauraceae from the Eocene of Australia and New Zealand, evaluating their implications for reconstructions of Eocene pCO$_2$ levels, is published by Steinthorsdottir et al. (2019).
* Description of early Eocene leaf fossils from the Dinmore locality (Redbank Plains Formation, Booval Basin; Australia) and a study on the implications of these fossils for reconstructions of paleoclimate is published by Pole (2019).
* A study on changes of plant communities from the Herren beds (Oregon, United States) during the Eocene and on the implications of plant fossils from this area for the reconstruction of Eocene climate is published by Jijina, Currano & Constenius (2019).
* Su et al. (2019) use radiometrically dated plant fossil assemblages to quantify when southeastern Tibet achieved its present elevation, and what kind of floras existed there at that time.
* Description of a plant megafossil assemblage from the Kailas Formation in western part of the southern Lhasa terrane, and a study on its implications for inferring the elevation history of the southern Tibetan Plateau, is published by Ai et al. (2019).
* A study on the dynamics and evolution of the flora of Turgai ecological type in Western Siberia during the early Oligocene to earliest Miocene is published by Popova et al. (2019).
* A study on the paleoclimate, vegetational type and ecological strategies adopted by fossil plants from the Oligocene Baigang Formation (China), as indicated by characteristics of fossil leaves from this formation, is published by Li et al. (2019).
* Description of a fossil plant assemblage from the Miocene Hattiesburg Formation (Mississippi, United States) is published by McNair et al. (2019).
* A study on changes of C$_4$ vegetation composition in southwestern Montana (United States) from the late Miocene through present is published by Hyland et al. (2019).
* A study aiming to test the hypothesis that fire contributed to the rise of C$_3$-dominated grasslands in Eurasia, based on data from core retrieved from the late Miocene to Pleistocene sediments from the Black Sea, is published by Feurdean & Vasiliev (2019).
* A study on the origin of the African C$_4$ savannah grasslands is published by Polissar et al. (2019).
* A study on vegetation changes in west African tropical montane forest over the past 90,000 years, as indicated by pollen data from the Lake Bambili site (Cameroon), is published by Lézine et al. (2019).
* A study on changes of vegetation in southern Borneo over the past 40,000  calibrated years BP, as indicated by data from Saleh Cave (South Kalimantan, Indonesia), is published by Wurster et al. (2019).
* A study on the role of past climate, extinct megafauna and guanaco in shaping the vegetation of the Patagonian steppe is published by Hernández, Ríos & Perotto-Baldivieso (2019).
* The discovery of ancient chestnut, hazelnut and flax DNA recovered from stalagmites from the Solkota cave (Georgia) is reported by Stahlschmidt et al. (2019).
* The discovery of oldest fossil trees, dating back 386 million years, in the Catskill region near Cairo, New York, is published online by Stein et al. (2019).