# Aphodius

Aphodius is a genus of beetles in the family Scarabaeidae. In most species both the adults and larvae are coprophagous (dung feeding) although some species have herbivorous or saprophagous larvae. Aphodius species typically dominate dung beetle communities in north temperate ecosystems. Most species are functionally classified as endocoprids, also known as dwellers, because the larvae live and feed within the dung pat itself.
With a discovery of an earlier published version by Hellwig, the correct citation for this genus is now: Aphodius Hellwig, 1798 (p. 101; 24 January). Type species by subsequent designation by Latreille (1810): Scarabaeus fimetarius Linnaeus, 1758.

## Species

These 44 species belong to the genus Aphodius, including 21 extinct species.
* Aphodius beloni Mulsant & Godart, 1879
* Aphodius calichromus Balthasar, 1932
* Aphodius cardinalis Reitter, 1892
* Aphodius clypeatus Fischer von Waldheim, 1821
* Aphodius coniugatus (Panzer, 1795)
* Aphodius corallifer Koshantschikov, 1913
* Aphodius crux Wiedemann, 1823
* Aphodius elegans Allibert, 1847
* Aphodius fasciger Harold, 1881
* Aphodius fimetarius (Linnaeus, 1758)
* Aphodius foetidus (Herbst, 1783)
* Aphodius guangdongensis Maté, 2008
* Aphodius irregularis Westwood, 1839
* Aphodius marginatus Fischer von Waldheim, 1842
* Aphodius micros Walker, 1871
* Aphodius minatorius Péringuey, 1908
* Aphodius pedellus (Degeer, 1774)
* Aphodius pedrosi Wollaston, 1854
* Aphodius plasoni Käufel, 1914
* Aphodius reginae Král, 1997
* Aphodius spadix Schmidt, 1916
* Aphodius swaneticus Reitter, 1892
* Aphodius thoracicus Fischer von Waldheim, 1842
* † Aphodius aboriginalis Wickham, 1912
* † Aphodius anteactus Krell, 2000
* † Aphodius bosniaskii Handlirsch, 1907
* † Aphodius brevipennis Heer, 1862
* † Aphodius charauxi Piton, 1940
* † Aphodius cretaceous Nikolajev, 2008
* † Aphodius florissantensis Wickham, 1911
* † Aphodius granarioides Wickham, 1913
* † Aphodius helvolus Statz, 1952
* † Aphodius inundatus Wickham, 1914
* † Aphodius krantzi Heyden & Heyden, 1866
* † Aphodius laminicola Wickham, 1910
* † Aphodius mediaevus Wickham, 1914
* † Aphodius meyeri Heer, 1847
* † Aphodius praeemptor Wickham, 1913
* † Aphodius precursor Horn, 1876
* † Aphodius schlickumi Statz, 1952
* † Aphodius senex Wickham, 1914
* † Aphodius shoshonis Wickham, 1912
* † Aphodius theobaldi Krell, 2000
* † Aphodius vectis Krell, 2019
The species of the following genera were formerly classified in Aphodius:
    Acrossus Mulsant, 1842
    Agoliinus Schmidt, 1913
    Alloblackburneus Bordat, 2009
    Ballucus Gordon & Skelley, 2007
    Blackburneus Schmidt, 1913
    Calamosternus Motschulsky, 1859
    Caligodorus Gordon & Skelley, 2007
    Cephalocyclus Dellacasa et al., 1998
    Chilothorax Motschulsky, 1859
    Cinacanthus Schmidt, 1913
    Coelotrachelus Schmidt, 1913
    Colobopterus Mulsant, 1842
    Cryptoscatomaseter Gordon & Skelley, 2007
    Dellacasiellus Gordon & Skelley, 2007
    Dialytodius Gordon & Skelley, 2007
    Drepanocanthoides Schmidt, 1913
    Eupleurus Mulsant, 1842
    Flaviellus Gordon & Skelley, 2007
    Geomyphilus Gordon & Skelley, 2007
    Haroldiellus Gordon & Skelley, 2007
    Hornosus Dellacasa, Dellacasa & Gordon, 2015
    Irrasinus Gordon & Skelley, 2007
    Labarrus Mulsant & Rey, 1869
    Lechorodius Gordon & Skelley, 2007
    Liothorax Motschulsky, 1859
    Luxolinus Gordon & Skelley, 2007
    Maculaphodius Gordon & Skelley, 2007
    Melinopterus Mulsant, 1842
    Mendidius Harold, 1868
    Merogyrus Gordon & Skelley, 2007
    Neotrichonotulus Dellacasa et al., 2004
    Nialaphodius Kolbe, 1908
    Orodaliscoides Schmidt, 1913
    Oscarinus Gordon & Skelley, 2007
    Otophorus Mulsant, 1842
    Oxyomoides Dellacasa et al., 2016
    Pardalosus Gordon & Skelley, 2007
    Phaeaphodius Reitter, 1892
    Planolinellus Dellacasa & Dellacasa, 2005
    Planolinoides Dellacasa & Dellacasa, 2005
    Planolinus Mulsant & Rey, 1869
    Pseudagolius Schmidt, 1913
    Rugaphodius Gordon & Skelley, 2007
    Scabrostomus Gordon & Skelley, 2007
    Schaefferellus Gordon & Skelley, 2007
    Setodius Gordon & Skelley, 2007
    Stenotothorax Schmidt, 1913
    Strigodius Gordon & Skelley, 2007
    Tetraclipeoides Schmidt, 1913
    Teuchestes Mulsant, 1842
    Trichaphodioides Paulian, 1942
    Trichonotulus Bedel, 1911