# List of Copris species

These species belong to Copris, a genus of dung beetles in the family Scarabaeidae.

## Copris species

* Copris acutidens Motschulsky, 1860
* Copris afgoi Cambefort, 1992
* Copris agnus Sharp, 1875
* Copris algol Nguyen-phung, 1989
* Copris amabilis Kolbe, 1914
* Copris amyntor Klug, 1855
* Copris anceus (Olivier, 1789)
* Copris andrewesi Waterhouse, 1891
* Copris angusticornis Arrow, 1933
* Copris angustus Nguyen-phung & Cambefort, 1986
* Copris anomiopseoides Boucomont, 1924
* Copris antares Ferreira, 1958
* Copris arcturus Gillet, 1907
* Copris ares Zelenka, 1993
* Copris arimotoi Ochi & Araya, 1992
* Copris arizonensis Schaeffer, 1906
* Copris armatus Harold, 1869
* Copris armeniacus Faldermann, 1835
* Copris armiger Gillet, 1910
* Copris arrowi Felsche, 1910
* Copris aspericollis Gillet, 1910
* Copris atropolitus Gillet, 1910
* Copris bacchus Thunberg, 1787
* Copris barclayi Ochi, Kon & Bai, 2009
* Copris basilewskyi Ferreira, 1962
* Copris basipunctatus Balthasar, 1942
* Copris bellator Chevrolat, 1844
* Copris bengalensis Gillet, 1911
* Copris bidens Kolbe, 1893
* Copris binodis
* Copris bootes Klug, 1855
* Copris boucardi Harold, 1869
* Copris bovinus Gillet, 1908
* Copris brachypterus Nomura, 1964
* Copris brittoni Ferreira, 1962
* Copris caelatus (Fabricius, 1794)
* Copris caliginosus Kohlmann, Cano & Delgado, 2003
* Copris cambeforti Nguyen-phung, 1988
* Copris cambodiensis Ochi, Kon & Kawahara, 2008
* Copris camerunus Felsche, 1904
* Copris caobangensis Bui, Dumack & Bonkowski, 2018
* Copris capensis Waterhouse, 1891
* Copris cariniceps Felsche, 1910
* Copris carinicus Gillet, 1910
* Copris carmelita Fabricius, 1801
* Copris cassius Péringuey, 1901
* Copris celebensis Lansberge, 1886
* Copris cheni Ochi, Kon & Bai, 2007
* Copris chimalapensis Mora-Aguilar & Delgado, 2015
* Copris ciliatus
* Copris colmanti Gillet, 1908
* Copris complexus Nguyen-phung, 1989
* Copris compressipennis Gillet, 1910
* Copris confucius Harold, 1877
* Copris coriarius Gillet, 1907
* Copris cornifrons Boheman, 1860
* Copris corniger Sahlberg, 1823
* Copris corpulentus Gillet, 1910
* Copris costaricensis Gahan, 1894
* Copris costariensis Matthews
* Copris crassus Deschodt & Davis, 2015
* Copris cribratus Gillet, 1927
* Copris cribricollis Gillet, 1910
* Copris curlettii Moretto, 2011
* Copris davidis (Deyrolle, 1878)
* Copris davisi Nguyen-phung & Cambefort, 1986
* Copris davisoni Waterhouse, 1891
* Copris decellei Cambefort, 1992
* Copris delicatus Arrow, 1931
* Copris denticulatus Nguyen-phung, 1988
* Copris diversus Waterhouse, 1891
* Copris draco Arrow, 1906
* Copris dracunculus Fereira, 1959
* Copris druidum Heer, 1862
* Copris dudleyi Cambefort, 1992
* Copris eburneus Cambefort, 1992
* Copris elpheneroides Felsche, 1910
* Copris elphenor Klug, 1855
* Copris erratus Lansberge, 1886
* Copris evanidus Klug, 1855
* Copris excisus Waterhouse, 1891
* Copris fallaciosus Gillet, 1907
* Copris fallax Felsche, 1910
* Copris felschei Reitter, 1892
* Copris fidius (Olivier, 1789)
* Copris frankenbergerianus Balthasar, 1958
* Copris freyi Ferreira, 1961
* Copris fricator (Fabricius, 1787)
* Copris fukiensis Balthasar, 1952
* Copris furciceps Felsche, 1910
* Copris garambae Cambefort, 1992
* Copris gazellarum Gillet, 1918
* Copris gibbulus Lansberge, 1886
* Copris gilleti Kolbe, 1907
* Copris gladiator Arrow, 1933
* Copris gopheri Hubbard, 1894
* Copris gracilis Waterhouse, 1891
* Copris graueri Kolbe, 1914
* Copris halffteri Matthews, 1959
* Copris harrisi Waterhouse, 1891
* Copris hartli Frey, 1975
* Copris hatayamai Ochi & Araya, 1992
* Copris hispanus (Linnaeus, 1764)
* Copris howdeni Matthews & Halffter, 1959
* Copris humilis Gillet, 1908
* Copris hybridus Gillet, 1910
* Copris igualensis Warner, 1990
* Copris illotus Balthasar, 1942
* Copris imitans Felsche, 1910
* Copris inaequabilis Zhang, 1997
* Copris incertus Say, 1835
* Copris inemarginatus Blatchley, 1918
* Copris inhalatus Quedenfeldt, 1884
* Copris insidiosus Péringuey, 1901
* Copris integer Reiche, 1847
* Copris interioris Kolbe, 1897
* Copris iris Sharp, 1875
* Copris jacchoides Nguyen-phung & Cambefort, 1987
* Copris jacchus (Fabricius, 1775)
* Copris jahi Nguyen-phung & Cambefort, 1986
* Copris johkii Ochi & Kon, 2014
* Copris juanmorai Mora-Aguilar & Delgado, 2015
* Copris jucundus Gillet, 1932
* Copris kachinensis Ochi, Kon & Kawahara, 2008
* Copris kartlinus Kabakov, 1988
* Copris kasagii Ochi & Kon, 1996
* Copris kasanka Josso & Prévost, 2016
* Copris katangae Cambefort & Nguyen-phung, 1996
* Copris keralensis Gill, 1986
* Copris kiuchii Masumoto, 1989
* Copris klapperichi Balthasar, 1942
* Copris klugi Harold, 1869
* Copris kusakabei Ochi, Kon & Kawahara, 2005
* Copris laeviceps Harold, 1862
* Copris laevigatus Gillet, 1927
* Copris laiiformis Nguyen-phung & Cambefort, 1987
* Copris laioides Boucomont, 1932
* Copris laius Harold, 1868
* Copris lannathai Hanboonsong & Masumoto & Ochi, 2003
* Copris leakeyorum Paulian, 1976
* Copris lecontei Matthews, 1962
* Copris leruei Josso & Prévost, 2017
* Copris lugubris Boheman, 1858
* Copris lunaris [fr] (Linnaeus, 1758)
* Copris macacus Lansberge, 1886
* Copris macclevei Warner, 1990
* Copris macer Péringuey, 1901
* Copris maesi Ratcliffe, 1998
* Copris magicus Harold, 1881
* Copris manni Ochi, Kon & Bai, 2009
* Copris marcus Gillet, 1921
* Copris martinae Nguyen-phung, 1988
* Copris martinezi Matthews and Halffter, 1968
* Copris matthewsi Delgado & Kohlmann, 2001
* Copris medogensis Zhang, 1988
* Copris megaceratoides Waterhouse, 1891
* Copris megasoma Matthews & Halffter, 1959
* Copris mesacanthus Harold, 1878
* Copris mexicanus Matthews & Halffter, 1959
* Copris minutus (Drury, 1773)
* Copris misellus Péringuey, 1901
* Copris moechus LeConte, 1854
* Copris moffartsi Gillet, 1907
* Copris mongkhoni Hanboonsong & Masumoto & Ochi, 2003
* Copris montivagus Gillet, 1908
* Copris morphaeus Gillet, 1932
* Copris mourgliai Zelenka, 1993
* Copris neglectus Moxey, 1963
* Copris neotridens Génier & Prévost, 2017
* Copris nepos Gillet, 1908
* Copris nevinsoni Waterhouse, 1891
* Copris novaki Zelenka, 1992
* Copris nubilosus Kohlmann & Cano & Delgado, 2003
* Copris numa Lansberge, 1886
* Copris obenbergeri Balthasar, 1933
* Copris obesus Boheman, 1857
* Copris ochus (Motschulsky, 1860)
* Copris orion Klug, 1835
* Copris orphanus Guérin-Méneville, 1847
* Copris parapecuarius Ochi, Kon & Kawahara, 2008
* Copris pauliani Nguyen-phung & Cambefort, 1986
* Copris pecuarius Lewis, 1884
* Copris pedarioides Lansberge, 1886
* Copris phungae Cambefort, 1992
* Copris phylax Gillet, 1908
* Copris poggii Ochi & Kon, 2005
* Copris potanini Semenov, 1891
* Copris prevosti Josso, 2011
* Copris pristinus Pierce, 1946
* Copris pseudobootes Ferreira, 1962
* Copris pseudochus Ochi & Kon, 2004
* Copris pseudomoffartsi Moretto, 2011
* Copris pseudosinicus Balthasar, 1958
* Copris pueli Mollandin De Boissy, 1905
* Copris punctatus Gillet, 1910
* Copris puncticollis Boheman, 1857
* Copris punctipennis Boucomont, 1914
* Copris punctulatus Wiedemann, 1823
* Copris punjabensis Gillet, 1921
* Copris quasilaevigatus Ochi, Kon & Bai, 2007
* Copris ramosiceps Gillet, 1921
* Copris rebouchei Harold, 1869
* Copris remotus Leconte, 1866
* Copris renwarti Nguyen-phung & Cambefort, 1986
* Copris repertus Walker, 1858
* Copris ribbei Lansberge, 1886
* Copris riekoae Ochi & Kon, 2003
* Copris ritsemae Harold, 1875
* Copris rugosus Gillet, 1908
* Copris ruricola Balthasar, 1933
* Copris sabinus Gillet, 1910
* Copris sacontala Redtenbacher, 1848
* Copris sallei Harold, 1869
* Copris sarpedon Harold, 1868
* Copris saundersi Harold, 1869
* Copris schoolmeestersi Ochi & Kon, 2010
* Copris scorpio Zelenka, 1993
* Copris serius Nguyen-phung, 1987
* Copris sexdentatus Thunberg, 1818
* Copris siamensis Gillet, 1910
* Copris siangensis Biswas, 1980
* Copris sierrensis Matthews, 1961
* Copris signatus Walker, 1858
* Copris simonettai Cambefort, 1992
* Copris simulator Nguyen-phung & Cambefort, 1986
* Copris singularis Nguyen-phung & Cambefort, 1986
* Copris sinicus Hope, 1842
* Copris sinon (Olivier, 1789)
* Copris sodalis Walker, 1858
* Copris sonensis Bui, Dumack & Bonkowski, 2018
* Copris sorex Balthasar, 1942
* Copris sphaeropterus Harold, 1877
* Copris spinator Harold, 1881
* Copris subdolus Balthasar, 1958
* Copris subpunctatus Gillet, 1910
* Copris subsidens Péringuey, 1901
* Copris subterraneus Heer, 1862
* Copris sumatrensis Gillet, 1921
* Copris surdus Arrow, 1931
* Copris szechouanicus Balthasar, 1958
* Copris tetraodon Gillet, 1910
* Copris tridentatus Solis & Kohlmann, 2003
* Copris tripartitus Waterhouse, 1875
* Copris truncatus Felsche, 1901
* Copris tsukamotoi Ochi, Kon & Kawahara, 2008
* Copris typhoeus Gerstaecker, 1884
* Copris uenoi Hanboonsong & Masumoto & Ochi, 2003
* Copris umbilicatus Abeille De Perrin, 1901
* Copris urus Boheman, 1857
* Copris usambaricus Gillet, 1908
* Copris vankhaii Nguyen-phung, 1988
* Copris victorini Boheman, 1857
* Copris vietnamicus Kabakov, 1994
* Copris vilhenai Ferreira, 1962
* Copris vrydaghi Ferreira, 1962
* Copris warneri Mccleve & Kohlmann, 2005
* Copris wiesei Kolbe, 1914
* Copris yangi Ochi, Kon & Bai, 2007
* Copris youngai Balthasar, 1967
* Copris zhangi Ochi, Kon & Bai, 2009