# Bulimulidae

Bulimulidae is a taxonomic family of medium-sized to large, air-breathing, tropical and sub-tropical land snails, terrestrial pulmonate gastropod molluscs in the superfamily Orthalicoidea.

## Distribution

Distribution of species in the family Bulimulidae includes Ecuador (9 genera) and other South American countries. Some species also occur in North America.

## Fossil record

The family's oldest fossil record dates from the late Cretaceous of Brazil (Itaboraí Basin).

## Anatomy

Members of this family have a haploid chromosome number between 26 and 30 (according to the values in this table).
Shells of species within Bostrycinae have a smooth protoconch and the genital organs feature a relatively long penis sheath (ca. 1/4–1/6 total phallus length).

## Taxonomy

Previously, the members of the Orthalicidae were also included in this family, as the subfamily Orthalicinae, and the taxa listed here were placed in their own subfamily, the Bulimulinae.

### 2005 taxonomy

Bulimulinae was placed in the family Orthalicidae according to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005).
The subfamily Bulimulinae included 3 tribes:
* Tribe Bulimulini Tryon, 1867 - synonyms: Bulimidae Guilding, 1828 (inv.); Berendtiinae P. Fischer & Crosse, 1872; Bothiembryontidae Iredale, 1937
* Tribe Odontostomini Pilsbry & Vanatta, 1898 - synonym: Tomogeridae Jousseaume, 1877
* Tribe Simpulopsini Schileyko, 1999

### 2010 taxonomy

Breure et al. (2010) moved Bothriembryon to Placostylidae (since 2012 as Bothriembryontidae), elevated Bulimulinae to Bulimulidae and Odontostomini to Odontostomidae.

### 2012 taxonomy

Breure & Romero (2012) confirmed previous results from 2010, additionally they elevated Simpulopsini to Simpulopsidae. There are three subfamilies within Bulimulidae:
* subfamily Bulimulinae Tryon, 1867
* subfamily Peltellinae Gray, 1855
* subfamily Bostrycinae Breure, 2012

## Genera

Genera in the family Bulimulidae include:

### subfamily Bulimulinae Tryon, 1887

    (synonym: Berendtiinae P. Fischer & Crosse, 1872)
* Anctus E. von Martens, 1860
* Auris Spix, 1827
* Berendtia Crosse & P. Fischer, 1869
* Bocourtia Rochebrune, 1882
* Bulimulus Leach, 1814 - type genus of the subfamily Bulimulinae
* Cochlorina Jan, 1830
* Graptostracus Pilsbry, 1939
* Kora Simone, 2012
* Lopesianus Weyrauch, 1958
* Naesiotus Albers, 1850
* Neopetraeus E. von Martens, 1885
* Newboldius Pilsbry, 1932
* Otostomus H. Beck, 1837
* Oxychona Mörch, 1852
* Protoglyptus Pilsbry, 1897
* Pseudoxychona Pilsbry, 1930
* Rabdotus Albers, 1850
* Scutalus Albers, 1850
* Spartocentrum Dall, 1895
* Sphaeroconcha Breure, 1978
* Stenostylus Pilsbry, 1898
* Suniellus Breure, 1978

### subfamily Peltellinae

* Antidrymaeus Germain, 1907
* Drymaeus Albers, 1850
* Peltella Gray, 1855 - type genus of the subfamily Peltellinae
* Mesembrinus Albers, 1850

### subfamily Bostrycinae

* Bostryx Troschel, 1847 sensu lato- type genus of the subfamily Bostrycinae

### Unassigned within Bulimulidae

* Bulimus Bruguière, 1789 (temporary name)
* †Itaborahia Maury, 1935
* † Oreoconus D. W. Taylor in McKenna et al., 1962
* † Palaeobulimulus Parodiz, 1949
* Stapafurdius Simone, 2021
* † Tocobaga Auffenberg, Slapcinsky & Portell, 2015

## Synonyms

* Adzharia P. Hesse, 1933: synonym of Bulimulus Leach, 1814 (junior synonym)
* Antidrymaeus Germain, 1907: synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
* Atahualpa Strebel, 1910: synonym of Thaumastus E. von Martens, 1860
* Ataxellus Dall, 1912: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented asBostryx Troschel, 1847
* Ataxus Albers, 1850: synonym of Bostryx Troschel, 1847
* Bilamelliferus Weyrauch, 1958: synonym of Bostryx Troschel, 1847 (junior synonymy)
* Cochlogena Férussac, 1821: synonym of Bulimulus Leach, 1814 (junior synonym)
* Diaphanomormus Weyrauch, 1964: synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
* Elatibostryx Weyrauch, 1958: synonym of Bostryx Troschel, 1847
* Floreziellus Weyrauch, 1967: synonym of Bostryx Troschel, 1847 (junior synonymy)
* Geoceras Pilsbry, 1896: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
* Globulinus Crosse & P. Fischer, 1875: synonym of Rabdotus Albers, 1850
* Goniognathmus Crosse & P. Fischer, 1875: synonym of Drymaeus Albers, 1850 (junior synonym)
* Hamadryas Albers, 1850: synonym of Drymaeus Albers, 1850
* Hannarabdotus Emerson & Jacobson, 1964: synonym of Rabdotus Albers, 1850
* Kionoptyx Haas, 1966: synonym of Bostryx Troschel, 1847 (junior synonymy)
* Kuschelenia Hylton Scott, 1951: synonym of Bocourtia (Kuschelenia) Hylton Scott, 1951 represented as Bocourtia Rochebrune, 1882
* Leptodrymaeus Pilsbry, 1946: synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
* Leptomerus Albers, 1850: synonym of Bulimulus Leach, 1814
* Leptomormus Weyrauch, 1958: synonym of Drymaeus (Mesembrinus) Albers, 1850 represented as Drymaeus Albers, 1850
* Liostracus [sic]: synonym of Leiostracus Albers, 1850 (misspelling of original genus, Leiostracus Albers, 1850)
* Loboa Ihering, 1917: synonym of Bulimulus Leach, 1814
* Mormus E. von Martens, 1860: synonym of Drymaeus Albers, 1850
* Multifasciatus Weyrauch, 1958: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
* Naesiotellus Weyrauch, 1967: synonym of Bostryx Troschel, 1847 (junior synonym)
* Navicula Spix, 1827: synonym of Cochlorina Jan, 1830
* Nesiotes E. von Martens, 1860: synonym of Naesiotus Albers, 1850
* Obstrussus Parodiz, 1946: synonym of Naesiotus Albers, 1850
* Olinodia Dall, 1920: synonym of Naesiotus (Reclasta) Dall, 1920 represented as Naesiotus Albers, 1850
* Orphaicus Schaufuss, 1869: synonym of Thaumastus E. von Martens, 1860
* Orphnus Albers, 1850: synonym of Thaumastus E. von Martens, 1860
* Orthotomium Crosse & P. Fischer, 1875: synonym of Rabdotus Albers, 1850
* Pachyotus H. Beck, 1837: synonym of Auris Spix, 1827
* Pachytholus Strebel, 1909: synonym of Thaumastus E. von Martens, 1860
* Pampasinus Weyrauch, 1958: synonym of Bostryx (Platybostryx) Pilsbry, 1896: synonym ofBostryx Troschel, 1847
* Peronaeus Albers, 1850: synonym of Bostryx Troschel, 1847
* Phenacotaxus Dall, 1912: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
* Plecochilus Agassiz, 1846: synonym of Plekocheilus Guilding, 1827 (unjustified emendation)
* Pseudoperonaeus Weyrauch, 1958: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
* Pseudorhodea Dall, 1895: synonym of Rabdotus (Plicolumna) J.G. Cooper, 1895 represented as Rabdotus Albers, 1850
* Pyrgus Albers, 1850: synonym of Bostryx (Geopyrgus) Pilsbry, 1896 represented as Bostryx Troschel, 1847
* Rhabdotus [sic]: synonym of Rabdotus Albers, 1850 (incorrect subsequent spelling)
* Rhaphiellus L. Pfeiffer, 1856: synonym of Naesiotus Albers, 1850
* Rimatula Parodiz, 1946: synonym of Protoglyptus Pilsbry, 1897
* Saeronia Dall, 1920: synonym of Naesiotus Albers, 1850
* Semiclausaria L. Pfeiffer, 1856: synonym of Drymaeus Albers, 1850 (junior synonym)
* Siphalomphix Rafinesque, 1833: synonym of Bulimulus Leach, 1814
* Sonorina Pilsbry, 1896: synonym of Rabdotus (Leptobyrsus) Crosse & P. Fischer, 1875 represented as Rabdotus Albers, 1850
* Spiroscutalus Pilsbry, 1932: synonym of Scutalus Albers, 1850
* Stenostoma Spix, 1827: synonym of Anctus E. von Martens, 1860
* Tatutor Jousseaume, 1887: synonym of Thaumastus E. von Martens, 1860
* Teneritia J. Mabille, 1898: synonym of Spartocentrum Dall, 1895
* Tholus Strebel, 1909: synonym of Thaumastus E. von Martens, 1860
* Xenothauma Fulton 1896: synonym of Scutalus Albers, 1850
* Zaplagius Pilsbry, 1896: synonym of Cochlorina Jan, 1830