# List of members of the National Academy of Sciences (Biochemistry)

This list is a subsection of theList of members of the National Academy of Sciences, which includes approximately 2,000 members and 350 foreign associates of theUnited StatesNational Academy of Sciences, each of whom is affiliated with one of 31 disciplinary sections. Each person's name, primary institution, and election year are given. The designation (d) after the name means the member is deceased.
| Name | Institution | Year |
| John Abelson | California Institute of Technology | 1985 |
| Julius Adler | University of Wisconsin–Madison | 1978 |
| Paul Ahlquist | University of Wisconsin–Madison | 1993 |
| Bruce Alberts | University of California, San Francisco | 1981 |
| Jorge Allende | University of Chile | 2001 |
| Sidney Altman | Yale University | 1990 |
| Bruce Ames | University of California, Berkeley | 1972 |
| Gilbert Ashwell | National Institutes of Health | 1979 |
| Tania Baker | Massachusetts Institute of Technology | 2007 |
| Clinton Ballou (d) | University of California, Berkeley | 1975 |
| David Bartel | Massachusetts Institute of Technology | 2011 |
| Jacqueline Barton | California Institute of Technology | 2002 |
| Brenda Bass | University of Utah | 2015 |
| Philip Beachy | Johns Hopkins University | 2002 |
| Lorena Beese | Duke University | 2009 |
| Marlene Belfort | New York State Department of Health | 1999 |
| James Berger | Johns Hopkins University | 2013 |
| Klaus Biemann | Massachusetts Institute of Technology | 1993 |
| Pamela Björkman | California Institute of Technology | 2001 |
| Elizabeth Blackburn | University of California, San Francisco | 1993 |
| Elkan Blout (d) | Harvard University | 1969 |
| Michael Botchan | UC Berkeley | 2008 |
| Herbert Boyer | University of California, San Francisco | 1985 |
| Paul D. Boyer (d) | University of California, Los Angeles | 1970 |
| Roscoe Brady (d) | National Institutes of Health | 1975 |
| Ronald Breaker | Yale University | 2013 |
| Patrick O. Brown | Stanford University | 2002 |
| John Buchanan (d) | Massachusetts Institute of Technology | 1962 |
| Giulio Cantoni (d) | National Institutes of Health | 1983 |
| John Carbon | University of California, Santa Barbara | 1986 |
| H. E. Carter (d) | University of Arizona | 1953 |
| Thomas R. Cech | Howard Hughes Medical Institute | 1987 |
| Michael Chamberlin | University of California, Berkeley | 1986 |
| Britton Chance (d) | University of Pennsylvania | 1954 |
| Aaron Ciechanover | Technion-Israel Institute of Technology | 2007 |
| W. Wallace Cleland (d) | University of Wisconsin–Madison | 1985 |
| Philip Cohen | University of Dundee | 2008 |
| Seymour Cohen (d) | Marine Biological Laboratory | 1967 |
| Mildred Cohn (d) | University of Pennsylvania | 1971 |
| Joan W. Conaway | Stowers Institute for Medical Research | 2020 |
| Minor Coon (d) | University of Michigan | 1983 |
| Nicholas R. Cozzarelli (d) | University of California, Berkeley | 1989 |
| Nancy Craig | Johns Hopkins University | 2010 |
| Benjamin Cravatt III | Scripps Research Institute | 2014 |
| James Dahlberg | University of Wisconsin–Madison | 1996 |
| Seth Darst | Rockefeller University | 2008 |
| Earl Davie (d) | University of Washington | 1980 |
| Christian de Duve (d) | Christian de Duve Institute of Cellular Pathology | 1975 |
| Hector DeLuca | University of Wisconsin–Madison | 1979 |
| Peter Dervan | California Institute of Technology | 1986 |
| Raymond Deshaies | Caltech | 2016 |
| Jack Dixon | Howard Hughes Medical Institute | 2000 |
| Russell Doolittle | University of California, San Diego | 1984 |
| Jennifer Doudna | University of California, Berkeley | 2002 |
| Gideon Dreyfuss | University of Pennsylvania | 2012 |
| Gerald Edelman (d) | The Scripps Research Institute | 1969 |
| Ronald Estabrook | University of Texas Southwestern Medical Center at Dallas | 1979 |
| Gary Felsenfeld | National Institutes of Health | 1976 |
| Edmond Fischer (d) | University of Washington | 1973 |
| Perry A. Frey | University of Wisconsin–Madison | 1998 |
| Irwin Fridovich | Duke University | 1981 |
| E. Peter Geiduschek | University of California, San Diego | 1974 |
| Quentin Gibson (d) | Rice University | 1982 |
| Alexander Glazer | University of California System | 2001 |
| Rachel Green | Johns Hopkins University | 2012 |
| Carol Greider | Johns Hopkins University | 2003 |
| Marianne Grunberg-Manago | Institut de Biologie Physico-Chimique | 1982 |
| Michael Grunstein | University of California, Los Angeles | 2008 |
| Irwin Gunsalus (d) | University of Illinois at Urbana–Champaign | 1965 |
| Lowell Hager | University of Illinois at Urbana–Champaign | 1995 |
| Sen-itiroh Hakomori | Pacific Northwest Research Institute | 2000 |
| Gordon Hammes (d) | Duke University | 1973 |
| Gregory Hannon | Cold Spring Harbor Laboratory | 2012 |
| F. Ulrich Hartl | Max Planck Institute of Biochemistry | 2011 |
| Ari Helenius | ETH Zurich | 2009 |
| Richard Henderson | Medical Research Council | 1998 |
| Wayne Hendrickson | Columbia University | 1993 |
| Leon Heppel (d) | Cornell University | 1970 |
| Avram Hershko | Technion Israel Institute of Technology | 2003 |
| George Hess | Cornell University | 1987 |
| Robert L. Hill | Duke University | 1975 |
| Mahlon Hoagland (d) | Worcester Foundation for Biomedical Research | 1984 |
| B. L. Horecker (d) | Cornell University | 1961 |
| Arthur Horwich | Yale University | 2003 |
| Robert Huber | Max Planck Institute for Biochemistry | 1995 |
| Jerard Hurwitz (d) | Memorial Sloan-Kettering Cancer Center | 1974 |
| Clyde Hutchison | University of North Carolina at Chapel Hill | 1995 |
| William Jencks (d) | Brandeis University | 1971 |
| Gerald Joyce | Scripps Research Institute | 2001 |
| Howard Ronald Kaback | University of California, Los Angeles | 1987 |
| Paul Kaesberg (d) | University of Wisconsin–Madison | 1991 |
| Seymour Kaufman (d) | National Institutes of Health | 1986 |
| Yoshito Kaziro (d) | Kyoto University | 2000 |
| Eugene P. Kennedy (d) | Harvard University | 1964 |
| Peter Kim | Merck & Co., Inc. | 1997 |
| Robert Kingston | Harvard University | 2016 |
| Judith Klinman | University of California, Berkeley | 1994 |
| Jeremy Knowles (d) | Harvard University | 1988 |
| Edward Korn | National Institutes of Health | 1990 |
| Arthur Kornberg (d) | Stanford University | 1957 |
| Roger Kornberg | Stanford University | 1993 |
| Alberto Kornblihtt | University of Buenos Aires | 2011 |
| Daniel E. Koshland, Jr. (d) | University of California, Berkeley | 1966 |
| Stephen Kowalczykowski | University of California, Davis | 2007 |
| Joseph Kraut (d) | University of California, San Diego | 1988 |
| John Kuriyan | University of California, Berkeley | 2001 |
| Alan Lambowitz | University of Texas at Austin | 2004 |
| Arthur Landy | Brown University | 1999 |
| M. Daniel Lane | Johns Hopkins University | 1987 |
| Henry Lardy (d) | University of Wisconsin–Madison | 1958 |
| I. Robert Lehman | Stanford University | 1977 |
| William Lennarz | State University of New York at Stony Brook | 1989 |
| Christopher D. Lima | Memorial Sloan Kettering Cancer Center | 2020 |
| Laszlo Lorand (d) | Northwestern University | 1987 |
| George Lorimer | University of Maryland, College Park | 1997 |
| James Manley | Columbia University | 2011 |
| Lynne E. Maquat | University of Rochester | 2011 |
| Michael Marletta | The Scripps Research Institute | 2006 |
| Emanuel Margoliash (d) | Northwestern University | 1975 |
| Brian Matthews | University of Oregon | 1986 |
| Rowena Green Matthews | University of Michigan | 2002 |
| Stephen L. Mayo | California Institute of Technology | 2004 |
| Robert Bruce Merrifield (d) | Rockefeller University | 1972 |
| Stanley Miller (d) | University of California, San Diego | 1973 |
| Kiyoshi Mizuuchi | National Institutes of Health | 1994 |
| Paul Modrich | Duke University | 1993 |
| Elizabeth Neufeld | University of California, Los Angeles | 1977 |
| Marshall Warren Nirenberg (d) | National Institutes of Health | 1967 |
| Harry Noller | University of California, Santa Cruz | 1992 |
| Masayasu Nomura (d) | University of California, Irvine | 1978 |
| Peter Novick | University of California, San Diego | 2013 |
| Michael E. O'Donnell | The Rockefeller University | 2006 |
| Erin O'Shea | University of California, San Francisco | 2004 |
| Leslie Orgel (d) | Salk Institute for Biological Studies | 1990 |
| Roy Parker | University of Colorado Boulder | 2012 |
| Armando Parodi | Fundacion Instituto Leloir | 2000 |
| Dinshaw Patel | Memorial Sloan-Kettering Cancer Center | 2009 |
| Nikola Pavletich | Memorial Sloan-Kettering Cancer Center | 2012 |
| David Phillips (d) | University of Oxford | 1985 |
| Frank Putnam (d) | Indiana University | 1976 |
| Charles Radding | Yale University | 1995 |
| Venkatraman Ramakrishnan | Medical Research Council | 2004 |
| Linda Randall | University of Missouri | 1997 |
| Tom Rapoport | Harvard University | 2005 |
| Lester Reed (d) | University of Texas at Austin | 1973 |
| Douglas C. Rees | California Institute of Technology | 2000 |
| Peter Reichard | Karolinska Institute | 1980 |
| Danny Reinberg | New York University | 2015 |
| Alexander Rich (d) | Massachusetts Institute of Technology | 1970 |
| Charles C. Richardson | Harvard University | 1983 |
| Phillips Robbins | Boston University | 1982 |
| John D. Roberts (d) | California Institute of Technology | 1956 |
| Ellen Rothenberg | California Institute of Technology | 2021 |
| Saul Roseman (d) | Johns Hopkins University | 1972 |
| William J. Rutter | Synergenics, LLC | 1984 |
| Bengt I. Samuelsson | Karolinska Institute | 1984 |
| Anthony San Pietro (d) | Indiana University | 1983 |
| Aziz Sancar | University of North Carolina at Chapel Hill | 2005 |
| Frederick Sanger (d) | Medical Research Council | 1967 |
| Robert Sauer | Massachusetts Institute of Technology | 1996 |
| Howard Schachman (d) | University of California, Berkeley | 1968 |
| Gottfried Schatz | University of Basel | 1989 |
| Robert Schimke (d) | Stanford University | 1976 |
| Paul Schimmel | Scripps Research Institute | 1990 |
| Vern Schramm | Albert Einstein College of Medicine | 2007 |
| Brenda Schulman | St. Jude Children's Research Hospital | 2014 |
| Kevan Shokat | University of California, San Francisco | 2009 |
| Maxine Singer | Carnegie Institution of Washington | 1979 |
| Robert Sinsheimer | University of California, Santa Barbara | 1967 |
| Emil Smith (d) | University of California, Los Angeles | 1962 |
| Hamilton O. Smith | J. Craig Venter Institute | 1980 |
| Dieter Söll | Yale University | 1997 |
| Nahum Sonenberg | McGill University | 2015 |
| David Sprinson (d) | Columbia University | 1990 |
| E. R. Stadtman (d) | National Institutes of Health | 1969 |
| Thressa Stadtman (d) | National Institutes of Health | 1981 |
| George Stark | Cleveland Clinic Foundation | 1987 |
| Donald Steiner (d) | University of Chicago | 1975 |
| Joan A. Steitz | Yale University | 1983 |
| Thomas Steitz (d) | Yale University | 1990 |
| Audrey Stevens (d) | Oak Ridge National Laboratory | 1998 |
| Bruce Stillman | Cold Spring Harbor Laboratory | 2000 |
| Jack Strominger | Harvard University | 1970 |
| JoAnne Stubbe | Massachusetts Institute of Technology | 1992 |
| Julian M. Sturtevant (d) | Yale University | 1973 |
| Wesley Sundquist | University of Utah | 2014 |
| Jack Szostak | Harvard University | 1998 |
| Herbert Tabor (d) | National Institutes of Health | 1977 |
| Paul Talalay (d) | Johns Hopkins University | 1987 |
| Glauco Tocchini-Valentini | National Research Council of Italy | 2009 |
| Susan S. Taylor | University of California, San Diego | 1996 |
| Jeremy Thorner | UC Berkeley | 2015 |
| Robert Tjian | University of California, Berkeley | 1991 |
| Olke Uhlenbeck | Northwestern University | 1993 |
| P. Roy Vagelos | Merck & Co., Inc. | 1972 |
| Bert Vallee (d) | Harvard University | 1974 |
| Alexander Varshavsky | California Institute of Technology | 1995 |
| Sidney Velick (d) | University of Utah | 1981 |
| Peter von Hippel | University of Oregon | 1978 |
| Salih Wakil (d) | Baylor College of Medicine | 1990 |
| Christopher T. Walsh | Harvard University | 1989 |
| Peter Walter | University of California, San Francisco | 2004 |
| James C. Wang | Harvard University | 1986 |
| Herbert Weissbach | Florida Atlantic University | 1982 |
| Jonathan Weissman | University of California, San Francisco | 2009 |
| James A. Wells | University of California, San Francisco | 1999 |
| Stephen C. West | Francis Crick Institute | 2016 |
| Reed Wickner | National Institutes of Health | 2000 |
| Sue Hengren Wickner | National Institutes of Health | 2004 |
| William T. Wickner | Dartmouth College | 1996 |
| Richard Wolfenden | University of North Carolina at Chapel Hill | 2002 |
| Carl Wu | National Institutes of Health | 2006 |
| Wei Yang | National Institutes of Health | 2013 |
| Paul Zamecnik (d) | Harvard University | 1968 |