# List of Stygnidae species

This is a list of the described species of the harvestman family Stygnidae. The data is taken from Joel Hallan's Biology Catalog.

## Heterostygninae

Heterostygninae Roewer, 1913
* Eutimesius Roewer, 1913
   * Eutimesius albicinctus (Roewer, 1915) — Venezuela
   * Eutimesius ephippiatus (Roewer, 1915) — Colombia
   * Eutimesius ornatus (Roewer, 1943) — Colombia, Venezuela
   * Eutimesius simoni Roewer, 1913 — Brazil, Colombia, Ecuador, Peru
* Imeri Pinto-da-Rocha & Tourinho, 2012 (! after Hallan)
   * Imeri ajuba Coronato-Ribeiro, Pinto-da-Rocha & Rheims, 2013 — Brazil (Amazonas)
   * Imeri lomanhungae Pinto-da-Rocha & Tourinho, 2012 — Brazil (Amazonas)
* Innoxius Pinto-da-Rocha, 1997
   * Innoxius magnus (Caporiacco, 1951) — Venezuela
* Minax Pinto-da-Rocha, 1997
   * Minax tetraspinosus Pinto-da-Rocha, 1997 — Venezuela
* Stenostygnellus Roewer, 1913
   * Stenostygnellus flavolimbatus Roewer, 1913 — Venezuela
   * Stenostygnellus macrochelis (Roewer, 1917) — Venezuela
* Stygnidius Simon, 1879
   * Stygnidius guerinii Sørensen, 1932 — French Guayana
   * Stygnidius inflatus (Guérin-Méneville, 1829-1843) — Brazil, Venezuela, French Guiana
* Stygnoplus Simon, 1879
   * Stygnoplus antiguanus (Roewer, 1943) — Leeward Islands, Antigua
   * Stygnoplus biguttatus Pinto-da-Rocha, 1997 — Venezuela
   * Stygnoplus clavotibialis (Goodnight & Goodnight, 1947) — Trinidad
   * Stygnoplus flavitarsis (Simon, 1879) — Leeward Islands, Guadeloupe
   * Stygnoplus forcipatus (C.L. Koch, 1845) — Colombia
   * Stygnoplus granulosus Mello-Leitão, 1940 — Venezuela
   * Stygnoplus longipalpus (Goodnight & Goodnight, 1942) — Guyana
   * Stygnoplus meinerti Sørensen, 1932 — Venezuela
   * Stygnoplus triacanthus (C.L. Koch, 1839) — South America
   * Stygnoplus tuberculatus (Goodnight & Goodnight, 1942) — Dominica (plus synonym Stygnoplus dominicanus (Roewer, 1943) per Pérez-González, 2000, after Hallan).
* Timesius Simon, 1879
   * Timesius vesicularis (Gervais, 1844) — Colombia
* Yapacana Pinto-da-Rocha, 1997
   * Yapacana tibialis (Pinto-da-Rocha, 1997) — Venezuela

## Stygninae

Stygninae Simon, 1879
* Actinostygnoides Goodnight & Goodnight, 1942
   * Actinostygnoides carus Goodnight & Goodnight, 1942 — Guyana
* Auranus Mello-Leitão, 1941
   * Auranus hoeferscovitorum Pinto-da-Rocha, 1997 — Brazil
   * Auranus parvus Mello-Leitão, 1941 — Brazil
* Fortia Villarreal et al., 2022 (! after Hallan)
   * Fortia jedi Villarreal et al., 2022 — Colombia
   * Fortia sith Villarreal et al., 2022 — Colombia
* Iguarassua Roewer, 1943
   * Iguarassua schubarti Roewer, 1943 — Brazil
* Jabbastygnus Kury & Villarreal, 2015 (! after Hallan)
   * Jabbastygnus huttorum Kury & Villarreal, 2015 — Colombia (Boyacá)
* Kaapora Pinto-da-Rocha, 1997
   * Kaapora minutissimus (Roewer, 1943) — Brazil
* Metaphareus Roewer, 1912
   * Metaphareus albimanus Roewer, 1912 — Colombia
   * Metaphareus punctatus Roewer, 1913 — Venezuela
* Niceforoiellus Mello-Leitão, 1941
   * Niceforoiellus assimilis Mello-Leitão, 1941 — Colombia
* Obidosus Roewer, 1931 (! after Hallan, genus restored per Villarreal et al. 2019)
   * Obidosus albilineatus Roewer, 1957 — Peru, Ecuador, Brazil (Amazonas)
   * Obidosus amigos (Ochoa & Pinto-da-Rocha, 2013) — Peru
   * Obidosus amplichelis Roewer, 1931 — Brazil (Amazonas)
   * Obidosus apiacas (Pinto-da-Rocha, 2000) — Brazil (Mato Grosso)
   * Obidosus bahiensis (Pinto-da-Rocha & Villarreal, 2009) — Brazil (Bahía)
   * Obidosus boibumba (Villarreal & Pinto-da-Rocha, 2006) — Brazil (Acre), Peru
   * Obidosus carnaval (Villarreal & Pinto-da-Rocha, 2006) — Brazil (Acre), Colombia (Amazonas)
   * Obidosus cirio (Villarreal & Pinto-da-Rocha, 2006) — Brazil (Pará)
   * Obidosus coxalis (Roewer, 1931) — Brazil (Amazonas)
   * Obidosus evelineae (Soares & Soares, 1978) — Brazil (Pará)
   * Obidosus foliadereis (Villarreal & Pinto-da-Rocha, 2006) — Brazil (Mato Grosso)
   * Obidosus jirau (Bragagnolo, 2013) — Brazil (Rondonia)
   * Obidosus junina (Villarreal & Pinto-da-Rocha, 2006) — Brazil (Bahía)
   * Obidosus kakinte (Ochoa & Pinto-da-Rocha, 2013) — Peru
   * Obidosus laevis (Sørensen, 1932) — Brazil (Pará, Matto Grosso)
   * Obidosus longipalpis (Roewer, 1943) — Brazil (Amazonas)
   * Obidosus machiguenga (Ochoa & Pinto-da-Rocha, 2013) — Peru
   * Obidosus mendopictus (Soares, 1978) — Brazil (Bahía)
   * Obidosus osvaldoi (Kury & Pinto-da-Rocha, 2008) — Brazil (Espírito Santo)
   * Obidosus palpalis (Roewer, 1931) — Brazil (Amazonas)
   * Obidosus regalo (Bragagnolo, 2013) — Brazil (Tocantins)
   * Obidosus trocaraincola (Pinto-da-Rocha, 1997) — Brazil (Pará)
* Ortonia Wood, 1869
   * Ortonia ferox Wood, 1869 — Ecuador
* Paraphareus Goodnight & Goodnight, 1943
   * Paraphareus tatei Goodnight & Goodnight, 1943 — Guyana/Venezuela (Bolívar: Monte Roraima)
* Phareus Simon, 1879 (! revised after Hallan)
   * Phareus antrophilus Villarreal & Rodríguez-T, 2006 — Colombia
   * Phareus raptator (Gervais, 1844) — Colombia, Venezuela
* Pickeliana Mello-Leitão, 1932 (! revised after Hallan)
   * Pickeliana albimaculata Hara & Pinto-da-Rocha, 2008
   * Pickeliana capito (Soares & Soares, 1974)
   * Pickeliana pickeli Mello-Leitão, 1932 — Brazil
* Planophareus Goodnight & Goodnight, 1943
   * Planophareus pallidus Goodnight & Goodnight, 1943 — Monte Roraima, Brazil
* Protimesius Roewer, 1913 (! revised after Hallan)
   * Protimesius gracilis Roewer, 1913 — Brazil (Amapá), Suriname, French Guiana
   * Protimesius lucifer Villarreal, Ázara & Kury, 2019 — Brazil (Pará)
   * Protimesius orcus Villarreal, Ázara & Kury, 2019 — Brazil (Ceará)
* Ricstygnus Kury, 2009 (! after Hallan)
   * Ricstygnus quineti Kury, 2009 — Brazil (Ceará)
* Sickesia Soares, 1979
   * Sickesia helmuti Soares, 1979 — Brazil
* Stenophareus Goodnight & Goodnight, 1943
   * Stenophareus roraimus Goodnight & Goodnight, 1943 — Roraima, Brazil
* Stenostygnoides Roewer, 1913
   * Stenostygnoides cosmetitarsus Roewer, 1913 — Surinam
* Stygnus Perty, 1833
   * Stygnus aggerum Sørensen, 1932 — Venezuela
   * Stygnus armatus Perty, 1833 — Brazil
   * Stygnus brevispinis Pinto-da-Rocha, 1997 — Bolivia
   * Stygnus ferrugineus (Perty, 1833) — Brazil, French Guiana
   * Stygnus gertschi (Roewer, 1963) — Colombia
   * Stygnus grasshoffi Pinto-da-Rocha, 1997 — Venezuela
   * Stygnus klugi (Goodnight & Goodnight, 1943) — Peru
   * Stygnus lesserti (Roewer, 1943) — Brazil
   * Stygnus luteus (Mello-Leitão, 1931) — Brazil
   * Stygnus marthae Pinto-da-Rocha, 1997 — Brazil
   * Stygnus mediocris (Roewer, 1931) — Ecuador
   * Stygnus multispinosus (Piza, 1938) — Brazil
   * Stygnus pectinipes (Roewer, 1943) — Surinam, Brazil, Colombia
   * Stygnus peruvianus (Roewer, 1957) — Peru
   * Stygnus polyacanthus (Mello-Leitão, 1923) — Brazil
   * Stygnus simonis Sørensen, 1932 — Peru
   * Stygnus simplex (Roewer, 1913) — Ecuador, Colombia, Peru
   * Stygnus weyrauchi (Roewer, 1963) — Peru
* Verrucastygnus Pinto-da-Rocha, 1997
   * Verrucastygnus caliginosus (R. Pinto-da-Rocha, 1990) — Brazil

## incertae sedis

incertae sedis
* Gaibulus Roewer, 1943
   * Gaibulus schubarti Roewer, 1943 — Brazil

### Since excluded

Subsequently transferred are former Stygnidae: Nomoclastinae Roewer, 1943
* Nomoclastes Sørensen, 1932
   * Nomoclastes quasimodo Pinto-da-Rocha, 1997 — Colombia
   * Nomoclastes taedifer Sørensen, 1932 — Colombia
Moved to => Nomoclastidae Roewer, 1943