# Sclerosomatidae

The Sclerosomatidae are a family of harvestmen with about 1,300 known species. One former subfamily has been recently removed to form a new family, Globipedidae.

## Name

The name is combined from Ancient Greek skleros "hard" and soma "body".

## Genera

* Gagrellinae Thorell, 1889
   * Abaetetuba Tourinho-Davis, 2004 (5 species)
   * Adungrella Roewer, 1955 (3 species)
   * Aguilaia González-Sponga, 2003 (1 species; Venezuela)
   * Akalpia Roewer, 1915 (2 species; India and Japan)
   * Altobunus Roewer, 1910 (3 species; Celebes and Philippines)
   * Amazonesia Soares, 1970 (2 species)
   * Antigrella Roewer, 1954 (1 species)
   * Aurivilliola Roewer, 1910 (17 species; South and Southeast Asia)
   * Azucarella Roewer, 1959 (1 species)
   * Bakerinulus Roewer, 1955 (1 species)
   * Bastia Roewer, 1910 (3 species; South and Southeast Asia)
   * Bastioides Mello-Leitão, 1931 (1 species; South America)
   * Baturitia Roewer, 1931 (1 species; Sunda Islands)
   * Bellonia González-Sponga, 2003 (1 species; Venezuela)
   * Biceropsis Roewer, 1935 (1 species; Burma)
   * Bonthainia Roewer, 1913 (5 species; South and Southeast Asia)
   * Bullobunus Roewer, 1910 (7 species; Philippines)
   * Caluga Roewer, 1959 (1 species)
   * Cardenalia Kury, 2017 (1 species; Venezuela)
   * Carinobius Roewer, 1955 (1 species)
   * Carmenia Roewer, 1915 (1 species; Colombia)
   * Carmichaelus Roewer, 1929 (1 species; India)
   * Ceratobunellus Roewer, 1911 (3 species; India)
   * Ceratobunoides Roewer, 1923 (2 species; Sumatra)
   * Ceratobunus Thorell, 1889 (13 species)
   * Cervibunus Roewer, 1912 (3 Species; South and Southeast Asia)
   * Chasenella Roewer, 1955 (2 species; Borneo)
   * Chebabius Roewer, 1935 (1 species; Burma)
   * Coonoora Roewer, 1929 (1 species; India)
   * Dentobunus Roewer, 1910 (33 species; Southeast Asia)
   * Diangathia Roewer, 1955 (1 species)
   * Echinobunus Roewer, 1912 (1 species; Java)
   * Euceratobunus Roewer, 1923 (1 species; India)
   * Eugagrella Roewer, 1910 (25 species; South and Southeast Asia)
   * Euzaleptus Roewer, 1911 (4 species; South and Southeast Asia)
   * Fesa Roewer, 1953 (1 species)
   * Gagrella Stoliczka, 1869 (226 species; Asia)
   * Gagrellenna Roewer, 1929 (1 species; India)
   * Gagrellina Roewer, 1913 (1 species; Celebes)
   * Gagrellissa Roewer, 1931 (1 species; Sunda Islands)
   * Gagrellopsis Sato & Suzuki, 1939 (1 species; Japan)
   * Gagrellula Roewer, 1910 (67 species; Asia)
   * Garleppa Roewer, 1912 (4 species; South America)
   * Geaya Roewer, 1910 (82 species; Latin America)
   * Globulosoma Martens, 1987 (2 species; Nepal)
   * Guaranobunus Ringuelet, 1954 (1 species; Argentina)
   * Hamitergum Crawford, 1992 (1 species; Russia)
   * Harmanda Roewer, 1910 (15 species; Asia)
   * Harmandina Schenkel, 1954 (1 species)
   * Hehoa Roewer, 1929 (1 species; India)
   * Heterogagrella Roewer, 1954 (2 species; Southeast Asia)
   * Hexazaleptus Suzuki, 1966 (1 species)
   * Hexomma Thorell, 1876 (9 species; Southeast Asia)
   * Himaldroma Martens, 1987 (2 species; Nepal)
   * Himalzaleptus Martens, 1987 (1 species; Nepal)
   * Holcobunus Roewer, 1910 (26 or 27 species; Latin America)
   * Holmbergiana Mello-Leitão, 1931 (3 species; South America)
   * Hologagrella Roewer, 1910 (7 species; Southeast Asia)
   * Hypogrella Roewer, 1955 (1 species)
   * Hypsibunus Thorell, 1891 (11 species; India & Southeast Asia)
   * Jussara Mello-Leitão, 1935 (10 species; South America)
   * Koyamaia Suzuki, 1972 (1 species)
   * Krusa Goodnight & Goodnight, 1947 (10 species; Mexico)
   * Krusella Roewer, 1953 (1 species; Venezuela)
   * Liopagus Chamberlin, 1916 (1 species)
   * Marthana Thorell, 1891 (24 species; Southeast Asia)
   * Melanopa Thorell, 1889 (30 species; India, Southeast Asia)
   * Melanopella Roewer, 1931 (3 species; Sunda Islands)
   * Melanopula Roewer, 1929 (5 species; South and Southeast Asia)
   * Metadentobunus Roewer, 1915 (3 species; Taiwan)
   * Metahehoa Suzuki, 1985 (1 species)
   * Metasyleus Roewer, 1955 (3 species)
   * Metaverpulus Roewer, 1912 (7 species; South Asia)
   * Metazaleptus Roewer, 1912 (8 species; South and Southeast Asia)
   * Microzaleptus Roewer, 1955 (1 species)
   * Munequita Mello-Leitão, 1941 (1 species; Brazil)
   * Neogagrella Roewer, 1913 (3 species; South and Southeast Asia)
   * Nepalgrella Martens, 1987 (2 species; Nepal)
   * Nepalkanchia Martens, 1990 (2 species; Nepal)
   * Obigrella Roewer, 1955 (1 species)
   * Octozaleptus Suzuki, 1966 (1 species)
   * Onostemma Mello-Leitão, 1938 (2 species; Brazil)
   * Oobunus Kishida, 1930 (1 species)
   * Orissula Roewer, 1955 (1 species)
   * Padangrella Roewer, 1954 (1 species)
   * Palniella Roewer, 1929 (1 species; India)
   * Paradentobunus Roewer, 1915 (1 species; India)
   * Paragagrella Roewer, 1912 (5 species; South and Southeast Asia)
   * Paragagrellina Schenkel, 1963 (1 species)
   * Parageaya Mello-Leitão, 1933 (6 species; Latin America)
   * Paratamboicus Mello-Leitão, 1940 (30 species; South America)
   * Paraumbogrella Suzuki, 1963 (1 species)
   * Paruleptes Soares, 1970 (1 species)
   * Pectenobunus Roewer, 1910 (4 species; South America)
   * Pergagrella Roewer, 1954 (1 species)
   * Pokhara Suzuki, 1970 (7 species; Nepal)
   * Prionostemma Pocock, 1903 (114 species; Mexico, Central America, and South America)
   * Prodentobunus Roewer, 1923 (4 species; Asia)
   * Psammogeaya Mello-Leitão, 1946 (1 species; Uruguay)
   * Psathyropus L. Koch, 1878 (40 species; Asia)
   * Pseudarthromerus Karsch, 1892 (1 species)
   * Pseudogagrella Redikortsev, 1936 (13 species; East Asia)
   * Pseudomelanopa Suzuki, 1974 (2 species; East Asia)
   * Pseudosystenocentrus Suzuki, 1985 (1 species)
   * Romerella Goodnight & Goodnight, 1943 (5 species; Mexico and South America)
   * Sarasinia Roewer, 1913 (1 species; Celebes)
   * Sataria Roewer, 1915 (3 species; India)
   * Scotomenia Thorell, 1889 (1 species; Burma)
   * Sericicorpus Martens, 1987 (1 species; Nepal)
   * Sinadroma Roewer, 1955 (1 species)
   * Syleus Thorell, 1876 (2 species; India)
   * Syngagrella Roewer, 1913 (1 species; Celebes)
   * Systenocentrus Simon, 1886 (6 species; Southeast and East Asia)
   * Tamboicus Roewer, 1912 (3 species; South America)
   * Taperina Roewer, 1953 (2 species)
   * Tetraceratobunus Roewer, 1915 (3 species; South Asia)
   * Thunbergia Martens, 2020 (4 species; South America)
   * Toragrella Roewer, 1955 (2 species)
   * Trachyrhinus Weed, 1892 (7 species; North America: Mexico and United States)
   * Umbogrella Roewer, 1955 (1 species)
   * Umbopilio Roewer, 1956 (2 species)
   * Vaho González-Sponga, 2003 (1 species)
   * Valle González-Sponga, 2003 (3 species)
   * Varinodulia Canals, 1935 (1 species)
   * Verpulus Simon, 1901 (13 species; South and Southeast Asia)
   * Verrucobunus Roewer, 1931 (3 species; East and Southeast Asia)
   * Xerogrella Martens, 1987 (1 species; Nepal)
   * Zaleptiolus Roewer, 1955 (4 species; Asia)
   * Zaleptulus Roewer, 1955 (3 species)
   * Zaleptus Thorell, 1876 (72 species; Asia and Australia)
* Gyantinae Silhavý, 1946
   * Gyas Simon, 1879 (1 species; Alps)
   * Gyoides Martens, 1982 (6 species; Nepal)
   * Rongsharia Roewer, 1957 (3 species; Nepal)
* Leiobuninae Banks, 1893
   * †Amauropilio Mello-Leitão, 1937 (fossil) (2 species; fossils found in Colorado)
   * Cosmobunus Simon, 1879 (5 species; Spain and Mexico)
   * Dilophiocara Redikorzev, 1931 (2 species)
   * Eumesosoma Cokendolpher, 1980 (6 species; United States)
   * Eusclera Roewer, 1910 (2 species; China and India)
   * Hadrobunus Banks, 1900 (2 species; United States)
   * Leiobunum C.L.Koch, 1839 (126 species - 2 extinct fossils; worldwide)
   * Leuronychus Banks, 1900 (2 species; North America)
   * Microliobunum Roewer, 1912 (2 species; Middle East)
   * Micronelima Schenkel, 1938 (1 species; Spain)
   * Nelima Roewer, 1910 (44 species; worldwide)
   * Paranelima Caporiacco, 1938 (6 species; Mexico and Central America)
   * Schenkeliobunum Starega, 1964 (1 species)
   * Togwoteeus Roewer, 1952 (1 species; United States)
* Sclerosomatinae Simon, 1879
   * Astrobunus Thorell, 1876 (12 species; Europe)
   * Granulosoma Martens, 1973 (1 species)
   * Homalenotus C.L.Koch, 1839 (13 species; Southern Europe and North Africa)
   * Mastobunus Simon, 1879 (2 species; Southern Europe and North Africa)
   * Metasclerosoma Roewer, 1912 (5 species; Italy)
   * Pseudastrobunus Martens, 1973 (1 species)
   * Pseudohomalenotus Caporiacco, 1935 (1 species; the Karakoram)
   * Pygobunus Roewer, 1957 (2 species; Japan)
   * Umbopilio Roewer, 1956 (1 species)

## Removed

   * Caiza Roewer, 1925. Subjective synonym of Pectenobunus Roewer, 1910) in Tourinho, 2004a
   * Pseudoarthromerus Karsch, 1891 (1 species; Japan, nomen nudum, therefore non-valid)