# Bryaxis (beetle)

Bryaxis is a genus of ant-loving beetles in the family Staphylinidae. There are more than 380 described species in Bryaxis.

## Bryaxis species

These 385 species are members of the genus Bryaxis.
* Bryaxis abastumanus (Reitter, 1881)
* Bryaxis abkhasicus Besuchet & Kurbatov, 2007
* Bryaxis acernus Castellini, 1978
* Bryaxis adjaricus Besuchet & Kurbatov, 2007
* Bryaxis adumbratus Besuchet & Kurbatov, 2007
* Bryaxis aelistae (Reitter, 1882)
* Bryaxis affinis (Sharp, 1883)
* Bryaxis alayai Löbl & Kurbatov, 1996
* Bryaxis albanicus (Apfelbeck, 1907)
* Bryaxis alesi Löbl & Kurbatov, 1996
* Bryaxis alishanus Löbl & Kurbatov, 1996
* Bryaxis alpestris (Dodero, 1900)
* Bryaxis altivagus Besuchet, 1962
* Bryaxis amasiae (Reitter, 1890)
* Bryaxis ami Löbl & Kurbatov, 1996
* Bryaxis ammon (Saulcy, 1878)
* Bryaxis amurensis Kurbatov, 1985
* Bryaxis anatolicus (Saulcy, 1878)
* Bryaxis andalusiacus (Jeannel, 1956)
* Bryaxis appendiculatus (Reitter, 1885)
* Bryaxis aragonensis (Franz, 1955)
* Bryaxis argiolus (Reitter, 1888)
* Bryaxis argodi (Croissandeau, 1885)
* Bryaxis argus (Kraatz, 1863)
* Bryaxis armeniacus Brachat, 2019
* Bryaxis armipes (Reitter, 1881)
* Bryaxis arnoldii Besuchet, 1961
* Bryaxis artvinensis Besuchet & Kurbatov, 2007
* Bryaxis asciicornis Kurbatov, 1985
* Bryaxis assingi Besuchet & Kurbatov, 2007
* Bryaxis asturiensis (Reitter, 1880)
* Bryaxis atilla (Saulcy, 1878)
* Bryaxis atlanticides Newton, 2015
* Bryaxis auritides Newton, 2015
* Bryaxis badius Besuchet, 1961
* Bryaxis bajulus (Hampe, C., 1863)
* Bryaxis balabanus Besuchet & Kurbatov, 2007
* Bryaxis balneator Besuchet & Kurbatov, 2007
* Bryaxis bedeki Bekchiev & Hlaváč, 2016
* Bryaxis bellax Löbl & Kurbatov, 1996
* Bryaxis bergamascus (Stolz, 1917)
* Bryaxis bericus Pace, 1974
* Bryaxis beroni Karaman, 1969
* Bryaxis biokovensis Bekchiev & Hlaváč, 2016
* Bryaxis blacensis (Karaman, 1954)
* Bryaxis bodemeyeri (Reitter, 1903)
* Bryaxis bojanensis (Apfelbeck, 1907)
* Bryaxis borckensis Besuchet & Kurbatov, 2007
* Bryaxis bosnicus (Ganglbauer, 1895)
* Bryaxis bothrophorus (Stolz, 1917)
* Bryaxis brachati Besuchet, 1980
* Bryaxis brescianus Daffner, 1986
* Bryaxis brusinae (Reitter, 1880)
* Bryaxis buddha Kurbatov & Löbl, 1995
* Bryaxis bulbifer (Reichenbach, 1816)
* Bryaxis bushido Löbl, Kurbatov & Nomura, 1998
* Bryaxis callipus (Apfelbeck, 1906)
* Bryaxis carinula (Rey, 1888)
* Bryaxis carpathicus (Saulcy, 1875)
* Bryaxis casalei Poggi, 1992
* Bryaxis castanoptera Motschulsky, 1851
* Bryaxis catalanus (Jeannel, 1950)
* Bryaxis cateniger (Krauss, 1899)
* Bryaxis cebennicus (Jeannel, 1950)
* Bryaxis cephalotes (Motschulsky, 1845)
* Bryaxis cepliki Hlaváč, 2009
* Bryaxis cetinjensis (Apfelbeck, 1906)
* Bryaxis chaudoirii (Chaudoir, 1845)
* Bryaxis chevrolati (Aubé, 1833)
* Bryaxis chiarae Monguzzi, 2016
* Bryaxis chilimontanus Nomura & Chang Eon Lee, 1993
* Bryaxis chobautianus Löbl, 1998
* Bryaxis claudioi Daffner, 1992
* Bryaxis clavicornis (Panzer, 1805)
* Bryaxis clavipes (Motschulsky, 1851)
* Bryaxis cocles (Saulcy, 1863)
* Bryaxis coelestis Löbl & Kurbatov, 1996
* Bryaxis coiffaiti Jeannel, 1958
* Bryaxis collaris (Baudi di Selve, 1859)
* Bryaxis colledanii Daffner, 1984
* Bryaxis comita (Rambousek, 1909)
* Bryaxis convexus (Kiesenwetter, 1858)
* Bryaxis corcyreus (Reitter, 1884)
* Bryaxis cornutus (Machulka, 1938)
* Bryaxis corsus Besuchet, 1999
* Bryaxis crassicornides Newton, 2017
* Bryaxis crassicornis (Motschulsky, 1835)
* Bryaxis credibilis Besuchet & Kurbatov, 2007
* Bryaxis crepsensis (Müller, G., 1947)
* Bryaxis crotchi (Sharp, 1874)
* Bryaxis curtisii (Leach, 1817)
* Bryaxis cythereias (Pic, 1903)
* Bryaxis dalmatinus (Reitter, 1881)
* Bryaxis dichrous (Reitter, 1882)
* Bryaxis difficilis (Reitter, 1884)
* Bryaxis disciger (Roubal, 1910)
* Bryaxis distinguendus Besuchet, 1961
* Bryaxis diversicornis (Raffray, 1873)
* Bryaxis dolosus Poggi & Sardu, 2013
* Bryaxis effeminatus Besuchet, 1983
* Bryaxis egens Besuchet & Kurbatov, 2007
* Bryaxis emeicus Kurbatov & Löbl, 1995
* Bryaxis emendatus Besuchet & Kurbatov, 2007
* Bryaxis emilianus (Stolz, 1926)
* Bryaxis erichsonii (Kiesenwetter, 1849)
* Bryaxis eurus Karaman, 1967
* Bryaxis euryscapus Besuchet & Kurbatov, 2007
* Bryaxis extremalis Kurbatov, 1990
* Bryaxis fagei (Jeannel, 1926)
* Bryaxis fauconneti (Fauvel, 1886)
* Bryaxis femoratus (Aubé, 1844)
* Bryaxis festivus Besuchet, 1964
* Bryaxis fictor Kurbatov & Löbl, 1998
* Bryaxis focarilei Besuchet, 1980
* Bryaxis fossicornis (Rambousek, 1916)
* Bryaxis frivaldszkyi (Reitter, 1887)
* Bryaxis fronticornis Bekchiev, 2013
* Bryaxis frustratus Besuchet, 1983
* Bryaxis gallicus (Reitter, 1887)
* Bryaxis ganglbaueri (Stolz, 1917)
* Bryaxis gemellus Besuchet & Kurbatov, 2007
* Bryaxis ghilarovi Besuchet, 1961
* Bryaxis giraffa (Reitter, 1880)
* Bryaxis goliath (Jeannel, 1922)
* Bryaxis gracilipalpis Jeannel, 1958
* Bryaxis grandicollis Jeannel, 1958
* Bryaxis grilati (Reitter, 1886)
* Bryaxis grouvellei (Reitter, 1882)
* Bryaxis hakka Löbl & Kurbatov, 1996
* Bryaxis halbherri (Reitter, 1885)
* Bryaxis hallamontanus Nomura & Chang Eon Lee, 1992
* Bryaxis harmandi Raffray, 1909
* Bryaxis heian Löbl, Kurbatov & Nomura, 1998
* Bryaxis heilongjiangensis Li, Jingke, 1993
* Bryaxis heliophobus (Apfelbeck, 1906)
* Bryaxis herculinus Besuchet, 1962
* Bryaxis heydeni (Reitter, 1880)
* Bryaxis hiranoi Taru & Nomura, 2021
* Bryaxis hisamatsui Löbl, Kurbatov & Nomura, 1998
* Bryaxis hokkiensis Löbl & Kurbatov, 1996
* Bryaxis hoko Löbl, Kurbatov & Nomura, 1998
* Bryaxis holciki Löbl, 1964
* Bryaxis humilis Raffray, 1909
* Bryaxis hypocritus Plavilstschikov, 1940
* Bryaxis ibericus (Saulcy, 1870)
* Bryaxis immodicus Besuchet & Kurbatov, 2007
* Bryaxis indistincta Motschulsky, 1851
* Bryaxis insularis (Holdhaus, 1905)
* Bryaxis ipsimus Besuchet & Kurbatov, 2007
* Bryaxis iriomotensis Löbl, Kurbatov & Nomura, 1998
* Bryaxis islamitus (Reitter, 1885)
* Bryaxis issensis (Müller, J., 1909)
* Bryaxis italicus (Baudi di Selve, 1870)
* Bryaxis japonicus (Sharp, 1874)
* Bryaxis joffrei (Sainte-Claire Deville, 1928)
* Bryaxis jomon Löbl, Kurbatov & Nomura, 1998
* Bryaxis jucundus Besuchet, 1961
* Bryaxis judaeus (Pic, 1900)
* Bryaxis judicariensis (Dodero, 1919)
* Bryaxis kahleni Pace, 1978
* Bryaxis kamakura Löbl, Kurbatov & Nomura, 1998
* Bryaxis karamanae Besuchet, 1958
* Bryaxis karaormani (Karaman, 1954)
* Bryaxis karate Löbl, Kurbatov & Nomura, 1998
* Bryaxis katana Löbl, Kurbatov & Nomura, 1998
* Bryaxis khnzoriani Besuchet, 1964
* Bryaxis kimi Kurbatov & Löbl, 1998
* Bryaxis kimjongkuki Nomura & Chang Eon Lee, 1993
* Bryaxis klimeschi (Blattný, W. & C. Blattný, 1914)
* Bryaxis kofun Löbl, Kurbatov & Nomura, 1998
* Bryaxis koltzei (Reitter, 1887)
* Bryaxis konecznii (Machulka, 1938)
* Bryaxis kovali Besuchet & Kurbatov, 2007
* Bryaxis krilei Hlaváč, 2008
* Bryaxis kruegeri (Machulka, 1932)
* Bryaxis kumensis Daffner, 1992
* Bryaxis kumgangsanensis Löbl, 2000
* Bryaxis kurnakovi Besuchet, 1960
* Bryaxis kuzmini Besuchet & Kurbatov, 2007
* Bryaxis laevipennis Besuchet & Kurbatov, 2007
* Bryaxis lagari (Halbherr, 1890)
* Bryaxis lanai Poggi, 2019
* Bryaxis lata Motschulsky, 1851
* Bryaxis lazistanicus Besuchet & Kurbatov, 2007
* Bryaxis lederi (Reitter, 1888)
* Bryaxis leechanyoungi Nomura & Chang Eon Lee, 1993
* Bryaxis leleupi (Jeannel, 1952)
* Bryaxis leonhardi (Reitter, 1902)
* Bryaxis lesbius Brachat, 2016
* Bryaxis lessinicus Pace, 1974
* Bryaxis libanicola (Sahlberg, J., 1908)
* Bryaxis liguricus (Dodero, 1919)
* Bryaxis litoralis Besuchet & Kurbatov, 2007
* Bryaxis lokayi (Machulka, 1927)
* Bryaxis longifrons Jeannel, 1952
* Bryaxis longifronsides Newton, 2015
* Bryaxis longipalpis (Motschulsky, 1835)
* Bryaxis longulus (Kiesenwetter, 1849)
* Bryaxis lurensis Besuchet, 2002
* Bryaxis lusitanicus (Saulcy, 1870)
* Bryaxis macedonicus (Karaman, 1953)
* Bryaxis macheonensis Nomura & Chang Eon Lee, 1993
* Bryaxis mahunkai Löbl, 1975
* Bryaxis mancinii (Dodero, 1919)
* Bryaxis mandarinus Löbl & Kurbatov, 1996
* Bryaxis marinae Sabella, 1989
* Bryaxis mayumi Löbl, Kurbatov & Nomura, 1998
* Bryaxis meghruicus Brachat, 2019
* Bryaxis meledanus Agazzi, 1964
* Bryaxis melinensis (Reitter, 1881)
* Bryaxis mendax Kurbatov & Löbl, 1998
* Bryaxis merkli Löbl, 2000
* Bryaxis meskischesianus Meggiolaro, 1966
* Bryaxis meybohmianus Brachat, 2019
* Bryaxis mirei (Jeannel, 1956)
* Bryaxis mirificus Besuchet, 1983
* Bryaxis moczarskii (Winkler, A., 1911)
* Bryaxis mohamedis (Reitter, 1903)
* Bryaxis monguzzii Besuchet, 1980
* Bryaxis monstrosetibialis (Stolz, 1923)
* Bryaxis monticola (Jeannel, 1950)
* Bryaxis montivagus (Reitter, 1885)
* Bryaxis mosorensis (Karaman, 1953)
* Bryaxis mulsanti Kiesenwetter, 1850
* Bryaxis multiplex Besuchet & Kurbatov, 2007
* Bryaxis muscorum (Kiesenwetter, 1849)
* Bryaxis myops Besuchet & Kurbatov, 2007
* Bryaxis naginata Löbl, Kurbatov & Nomura, 1998
* Bryaxis nakeralae (Reitter, 1885)
* Bryaxis nametkini Kurbatov & Löbl, 1998
* Bryaxis nebrodensis Besuchet, 1980
* Bryaxis nemilensis (Reitter, 1885)
* Bryaxis nigriceps (Leach, 1826)
* Bryaxis nigripennis (Aubé, 1844)
* Bryaxis nigritus Löbl, 1998
* Bryaxis nitidulus Besuchet, 1961
* Bryaxis nivarius Besuchet & Kurbatov, 2007
* Bryaxis nodicornis (Aubé, 1833)
* Bryaxis nogodanensis Nomura & Chang Eon Lee, 1993
* Bryaxis noricus (Schubert, F., 1958)
* Bryaxis normandi (Jeannel, 1950)
* Bryaxis obenbergeri (Machulka, 1938)
* Bryaxis obventicius Besuchet & Kurbatov, 2007
* Bryaxis odontogena (Dodero, 1919)
* Bryaxis oertzeni (Reitter, 1882)
* Bryaxis oraniensis (Pic, 1896)
* Bryaxis orcinus Besuchet & Kurbatov, 2007
* Bryaxis oreophilus (Meixner, 1912)
* Bryaxis orousseti Secq, B. & M. Secq, 1991
* Bryaxis osellai Besuchet & Kurbatov, 2007
* Bryaxis oseticus Besuchet & Kurbatov, 2007
* Bryaxis ossaeus Besuchet, 2008
* Bryaxis pachyscelis Besuchet & Kurbatov, 2007
* Bryaxis paganettii (Blattný, W. & C. Blattný, 1914)
* Bryaxis paiwan Löbl & Kurbatov, 1996
* Bryaxis pallidior (Pic, 1901)
* Bryaxis panda Kurbatov & Löbl, 1995
* Bryaxis pandellei (Saulcy, 1863)
* Bryaxis pangeonicus Brachat, 2019
* Bryaxis pastoralis (Peyerimhoff, 1901)
* Bryaxis pavani (Tamanini, 1940)
* Bryaxis pawlowskii Löbl, 1974
* Bryaxis peckorum Löbl, Kurbatov & Nomura, 1998
* Bryaxis pedator (Reitter, 1882)
* Bryaxis pedemontanus Besuchet, 1958
* Bryaxis pedestris (Motschulsky, 1845)
* Bryaxis peloponnesius (Reitter, 1884)
* Bryaxis peninsulanus Nomura & Chang Eon Lee, 1993
* Bryaxis peninsularis (Saulcy, 1870)
* Bryaxis pentagonoceras (Stolz, 1917)
* Bryaxis persicoi (Rasetti, F., 1939)
* Bryaxis pescaroloi Poggi, 1984
* Bryaxis petri Raffray, 1915
* Bryaxis pictetii (Tournier, 1859)
* Bryaxis pilifera Motschulsky, 1851
* Bryaxis pingtungensis Löbl & Kurbatov, 1996
* Bryaxis pinkeri (Stolz, 1917)
* Bryaxis planifer Löbl & Kurbatov, 1996
* Bryaxis platalea Löbl, Kurbatov & Nomura, 1998
* Bryaxis pletnevi Kurbatov & Löbl, 1998
* Bryaxis polemon Besuchet & Kurbatov, 2007
* Bryaxis ponticus Besuchet & Kurbatov, 2007
* Bryaxis porsenna Reitter, 1882
* Bryaxis portalegrensis (Schaufuss, L. W., 1882)
* Bryaxis procerus (Gredler, 1873)
* Bryaxis propinquus Besuchet & Kurbatov, 2007
* Bryaxis pulchrotibialis Besuchet & Kurbatov, 2007
* Bryaxis pumilus Bekchiev & Brachat, 2015
* Bryaxis puncticollis (Denny, 1825)
* Bryaxis pygmaeides Newton, 2015
* Bryaxis pyrenaeus (Saulcy, 1863)
* Bryaxis rambouseki (Matcha, 1916)
* Bryaxis raptor Löbl & Kurbatov, 1996
* Bryaxis redemptus (Dodero, 1919)
* Bryaxis reissi (Machulka, 1933)
* Bryaxis reitteri (Saulcy, 1875)
* Bryaxis reversus (Sharp, 1883)
* Bryaxis rhinophorus (Blattný, W. & C. Blattný, 1914)
* Bryaxis rifensis Besuchet, 1962
* Bryaxis rivularis Besuchet & Kurbatov, 2007
* Bryaxis rossii Castellini, 1986
* Bryaxis rosti (Reitter, 1894)
* Bryaxis rostratus (Motschulsky, 1845)
* Bryaxis roubalianus Löbl, 1961
* Bryaxis roumaniae Raffray, 1904
* Bryaxis rousi Besuchet & Kurbatov, 2007
* Bryaxis rugosicollis (Fiori, 1905)
* Bryaxis ruidus Kurbatov & Löbl, 1998
* Bryaxis ruthenus (Saulcy, 1877)
* Bryaxis sacer Kurbatov & Löbl, 1995
* Bryaxis sacrificus Kurbatov & Löbl, 1995
* Bryaxis samniticus (Blattný, W. & C. Blattný, 1914)
* Bryaxis samothracicus Brachat, 2019
* Bryaxis samurai Löbl, Kurbatov & Nomura, 1998
* Bryaxis sarplaninensis (Karaman, 1953)
* Bryaxis sauteri Raffray, 1909
* Bryaxis sawadai Löbl, Kurbatov & Nomura, 1998
* Bryaxis scapularis (Reitter, 1881)
* Bryaxis schuelkei Besuchet & Kurbatov, 2007
* Bryaxis schusteri Neuhäuser-Happe, 1999
* Bryaxis scrutandus Besuchet & Kurbatov, 2007
* Bryaxis sculpticornis (Guillebeau, 1891)
* Bryaxis sculptifrons (Reitter, 1880)
* Bryaxis seductus Besuchet & Kurbatov, 2007
* Bryaxis serripes (Fauvel, 1891)
* Bryaxis sichotensis Kurbatov, 1994
* Bryaxis sichuanus Kurbatov & Löbl, 1995
* Bryaxis siculus (Fiori, 1913)
* Bryaxis silvicolides Newton, 2015
* Bryaxis simoni (Reitter, 1880)
* Bryaxis smetanai Löbl, 1964
* Bryaxis solidus (Reitter, 1881)
* Bryaxis sparcepunctatus Jeannel, 1958
* Bryaxis splendidus (Croissandeau, 1891)
* Bryaxis steindachneri (Reitter, 1881)
* Bryaxis stolzi (Machulka, 1932)
* Bryaxis sturanyi (Apfelbeck, 1906)
* Bryaxis subdentatus (Dodero, 1919)
* Bryaxis subseriatus (Weise, J., 1877)
* Bryaxis subsolidus (Reitter, 1902)
* Bryaxis sumo Löbl, Kurbatov & Nomura, 1998
* Bryaxis swaneticus (Reitter, 1885)
* Bryaxis taiwanus Löbl & Kurbatov, 1996
* Bryaxis tanto Löbl, Kurbatov & Nomura, 1998
* Bryaxis taradakensis Löbl, Kurbatov & Nomura, 1998
* Bryaxis tarokanus Löbl & Kurbatov, 1996
* Bryaxis temporalis Besuchet & Kurbatov, 2007
* Bryaxis tendensis Besuchet, 2002
* Bryaxis tenuicornides Newton, 2015
* Bryaxis testatus Kurbatov, 1994
* Bryaxis tetralobus Löbl, Kurbatov & Nomura, 1998
* Bryaxis theanus (Reitter, 1894)
* Bryaxis tienmushanus Kurbatov & Löbl, 1995
* Bryaxis tingitanus Besuchet, 1962
* Bryaxis tithonus (Apfelbeck, 1906)
* Bryaxis transitorius Besuchet & Kurbatov, 2007
* Bryaxis transsilvanicus (Ganglbauer, 1897)
* Bryaxis trezzii Monguzzi, 2016
* Bryaxis trichosus Jeannel, 1958
* Bryaxis trigonoceras (Holdhaus, 1904)
* Bryaxis troglocerus (Saulcy, 1870)
* Bryaxis troglodytes (Fiori, 1900)
* Bryaxis troglophilus Agazzi, 1961
* Bryaxis tscherkessicus (Reitter, 1888)
* Bryaxis tuberculiceps Nonveiller, Pavićević & Besuchet, 2003
* Bryaxis tychobythinoides Löbl & Kurbatov, 1996
* Bryaxis ullrichii (Motschulsky, 1851)
* Bryaxis unicus Löbl & Kurbatov, 1996
* Bryaxis ursus (Reitter, 1882)
* Bryaxis ussuriensis Löbl, 1964
* Bryaxis valentulus Kurbatov & Löbl, 1998
* Bryaxis validicornides Newton, 2015
* Bryaxis verrucipalpis (Apfelbeck, 1907)
* Bryaxis vicinus (Dodero, 1919)
* Bryaxis viertli (Reitter, 1882)
* Bryaxis viti Besuchet & Kurbatov, 2007
* Bryaxis vlastae Roubal, 1913
* Bryaxis wankai (Reitter, 1915)
* Bryaxis weisei (Saulcy, 1875)
* Bryaxis witzgalli Daffner, 1982
* Bryaxis wolongensis Kurbatov & Löbl, 1995
* Bryaxis yari Löbl, Kurbatov & Nomura, 1998
* Bryaxis ypsilon Besuchet & Kurbatov, 2007
* Bryaxis yushanensis Löbl & Kurbatov, 1996
* Bryaxis zubroides Bekchiev, 2022