# Helicostyla

Helicostyla is a genus of small, air-breathing land snails, terrestrial pulmonate gastropod mollusks in the subfamily Helicostylinae of the family Camaenidae. Helicostyla is the type genus of the subfamily Helicostylinae.

## Distribution

This genus is endemic to the Philippines.

## Species

Species within the genus Helicostyla include:
* Helicostyla albina (Grateloup, 1840)
* Helicostyla amagaensis de Chavez, Kendrich, Fontanilla, Batomalaque & Chiba, 2015
* Helicostyla amaliae (Möllendorff, 1890)
* Helicostyla aplomorpha (Jonas, 1843)
* Helicostyla bicolorata (I. Lea, 1840)
* Helicostyla boettgeriana (Möllendorff, 1888)
* Helicostyla bullula (Broderip, 1841)
* Helicostyla buschi (L. Pfeiffer, 1846)
* Helicostyla bustoi (Hidalgo, 1887)
* Helicostyla caesar (L. Pfeiffer, 1855)
* Helicostyla calamianica (Quadras Moellendorff, 1894)
* Helicostyla canonizadoi (Bartsch, 1932)
* Helicostyla chocolatina (Poppe, Tagaro & Sarino, 2015)
* Helicostyla cinerosa (L. Pfeiffer, 1855)
* Helicostyla collodes (G. B. Sowerby I, 1841)
* Helicostyla cunctator (Reeve, 1849)
* Helicostyla curta (G. B. Sowerby I, 1841)
* Helicostyla daphnis (Broderip, 1841)
* Helicostyla decorata (Férussac, 1821)
* Helicostyla delicata Fulton, 1903
* Helicostyla dimera (Jonas, 1846)
* Helicostyla domingoi (Bartsch, 1919)
* Helicostyla dimera (Jonas, 1846)
* Helicostyla effusa (Pfeiffer, 1842)
* Helicostyla euconica (Bartsch, 1932)
* Helicostyla fastidiosa (Bartsch, 1932)
* Helicostyla faunus (Broderip, 1841)
* Helicostyla frater (Férussac, 1821)
* Helicostyla fulgens (G. B. Sowerby I, 1841)
* Helicostyla fulgetra (Broderip, 1841)
* Helicostyla gigas (Hidalgo, 1901)
* Helicostyla gilva (Pfeiffer, 1845)
* Helicostyla glaucophthalma (L. Pfeiffer, 1851)
* Helicostyla halichlora (Semper, 1866)
* Helicostyla hololeuca (L. Pfeiffer, 1855)
* Helicostyla huberorum Thach, 2021
* Helicostyla imperator (Pfeiffer, 1848)
* Helicostyla indusiata (L. Pfeiffer, 1871)
* Helicostyla jonasi (L. Pfeiffer, 1846)
* Helicostyla juglans (L. Pfeiffer, 1842)
* Helicostyla lacerata (Semper, 1877) (nomen nudum)
* Helicostyla leai (L. Pfeiffer, 1846)
* Helicostyla leopardus (L. Pfeiffer, 1845)
* Helicostyla leucophaea (G. B. Sowerby I, 1841)
* Helicostyla librosa (L. Pfeiffer, 1857)
* Helicostyla lignaria (L. Pfeiffer, 1842)
* Helicostyla macrostoma (L. Pfeiffer, 1843)
* Helicostyla marinduquensis (Hidalgo, 1887)
* Helicostyla mateoi (Bartsch, 1932)
* Helicostyla mearnsi Bartsch, 1905
* Helicostyla mirabilis (Férussac, 1821)
* Helicostyla montana (C. Semper, 1877)
* Helicostyla mus (Delessert, 1841)
* Helicostyla nobilis (Reeve, 1848)
* Helicostyla nux (C. Semper, 1877)
* Helicostyla okadai Kuroda, 1932
* Helicostyla orbitula (Sowerby, 1841)
* Helicostyla palavanensis (L. Pfeiffer, 1857)
* Helicostyla persimilis (Deshayes, 1850)
* Helicostyla philippinensis (L. Pfeiffer, 1846)
* Helicostyla pictor (Broderip, 1841)
* Helicostyla pithogaster (Ferussac, 1821)
* Helicostyla polita Reeve, 1848
* Helicostyla portii (L. Pfeiffer, 1861)
* Helicostyla rufogaster (Lesson, 1832)
* Helicostyla saranganica (Hidalgo, 1887)
* Helicostyla satyrus (Broderip, 1841)
* Helicostyla schadenbergi (Möllendorff, 1890)
* Helicostyla seckendorffiana (L. Pfeiffer, 1847)
* Helicostyla simonei Thach & F. Huber, 2021
* Helicostyla simplex (Jonas, 1843)
* Helicostyla smaragdina (Reeve, 1842)
* Helicostyla solida (L. Pfeiffer, 1843)
* Helicostyla solivaga (Reeve, 1849)
* Helicostyla speciosa (Jay, 1839)
* Helicostyla tephrodes (L. Pfeiffer, 1843)
* Helicostyla thomsoni (L. Pfeiffer, 1871)
* Helicostyla ticaonica (Broderip, 1841)
* Helicostyla turbo (Pfeiffer, 1845)
* Helicostyla ventricosa (Bruguière, 1792)
* Helicostyla vidali (Hidalgo, 1887)
* Helicostyla villari (Hidalgo, 1887)
* Helicostyla volubilis (Reeve, 1851)
* Helicostyla weberi (Bartsch, 1919)
* Helicostyla woodiana (I. Lea, 1840)
* Helicostyla worcesteri Bartsch, 1909

### Species brought into synonymy

* Helicostyla aegle (Broderip, 1841): synonym of Cochlostyla aegle (Broderip, 1841)
* Helicostyla annulata (G.B. Sowerby I, 1841): synonym of Pachysphaera annulata (G. B. Sowerby I, 1841) (unaccepted combination)
* Helicostyla balteata (G.B. Sowerby I, 1841): synonym of Pachysphaera balteata (G. B. Sowerby I, 1841) (unaccepted combination)
* Helicostyla bembicodes (Pfeiffer, 1851): synonym of Helicobulinus bembicodes (L. Pfeiffer, 1851) (unaccepted combination)
* Helicostyla butleri (Pfeiffer, 1842): synonym of Dryocochlias butleri (L. Pfeiffer, 1842) (unaccepted combination)
* Helicostyla camelopardalis (Broderip, 1841): synonym of Hypselostyla camelopardalis (Broderip, 1841) (unaccepted combination)
* Helicostyla carinata (Lea, 1840): synonym of Hypselostyla carinata (I. Lea, 1840) (superseded combination)
* Helicostyla cincinna (G.B. Sowerby I, 1840): synonym of Hypselostyla cincinna (G. B. Sowerby I, 1841) (unaccepted combination)
* Helicostyla cincinniformis (G.B. Sowerb Iy, 1841): synonym of Hypselostyla cincinniformis (G. B. Sowerby I, 1841) (superseded combination)
* Helicostyla collodes (G.B. Sowerby I, 1841): synonym of Cochlostyla collodes (G. B. Sowerby I, 1841) (superseded combination)
* Helicostyla concinna (G.B. Sowerby I, 1841): synonym of Hypselostyla concinna (G. B. Sowerby I, 1841) (unaccepted combination)
* Helicostyla curta (G.B. Sowerby I, 1841) : synonym of Cochlostyla curta (G. B. Sowerby I, 1841) (superseded combination)
* Helicostyla dactylus (Broderip, 1841): synonym of Hypselostyla dactylus (Broderip, 1841) (superseded combination)
* Helicostyla daphnis (Broderip, 1841): synonym of Cochlostyla daphnis (Broderip, 1841) (superseded combination)
* Helicostyla decora (A. Adams & Reeve, 1850): synonym of Cochlodryas decora (A. Adams & Reeve, 1850) (superseded combination)
* Helicostyla denticulata (Jay, 1839): synonym of Calocochlea denticulata (Jay, 1839) (superseded combination)
* Helicostyla dimera (Jonas, 1846): synonym of Cochlostyla dimera (Jonas, 1846) (superseded combination)
* Helicostyla diana (Broderip, 1841): synonym of Hypselostyla diana (Broderip, 1841) (unaccepted combination)
* Helicostyla dubiosa (Pfeiffer, 1846): synonym of Cochlostyla dubiosa (L. Pfeiffer, 1846) (superseded combination
* Helicostyla euconica (Bartsch, 1932): synonym of Cochlostyla effusa (L. Pfeiffer, 1843)
* Helicostyla evanescens (Broderip, 1841): synonym of Hypselostyla evanescens (Broderip, 1841) (unaccepted combination)
* Helicostyla fenestrata (Sowerby I, 1841): synonym of Pachysphaera fenestrata (G. B. Sowerby I, 1841) (superseded combination)
* Helicostyla florida (Sowerby I, 1841): synonym of Cochlodryas florida (G. B. Sowerby I, 1841) (superseded combination)
* Helicostyla fulgens (Broderip, 1841): synonym of Cochlostyla fulgens (G. B. Sowerby I, 1841) (superseded combination)
* Helicostyla fulgetra (Broderip, 1841): synonym of Cochlostyla fulgetra (Broderip, 1841)
* Helicostyla grandis (Pfeiffer, 1845): synonym of Helicobulinus grandis (L. Pfeiffer, 1845) (unaccepted combination)
* Helicostyla hydrophana G.B. Sowerby I, 1841: synonym of Dryocochlias metaformis (Férussac, 1821) (junior synonym)
* Helicostyla ignobilis Sowerby I, 1841: synonym of Cochlodryas ignobilis (G. B. Sowerby I, 1841) (superseded combination)
* Helicostyla iloconensis (Sowerby I, 1841): synonym of Pachysphaera iloconensis (G. B. Sowerby I, 1841) (misspelling of original genus name, Orustia, Mörch, 1852)
* Helicostyla jonasi (Pfeiffer, 1845): synonym of Cochlostyla jonasi (L. Pfeiffer, 1846) (superseded combination)
* Helicostyla juglans (Pfeiffer, 1842): synonym of Cochlostyla juglans (L. Pfeiffer, 1842) (superseded combination)
* Helicostyla leopardus (Pfeiffer, 1845): synonym of Cochlostyla leopardus (L. Pfeiffer, 1845) (superseded combination)
* Helicostyla leucophaea (Sowerby I, 1841): synonym of Cochlostyla leucophaea (G. B. Sowerby I, 1841) (superseded combination)
* Helicostyla lignaria (Pfeiffer, 1842): synonym of Cochlostyla lignaria (L. Pfeiffer, 1842) (suspected synonym)
* Helicostyla macrostoma (Pfeiffer, 1843): synonym of Cochlostyla macrostoma (L. Pfeiffer, 1843) (superseded combination)
* Helicostyla marinduquensis (Hidalgo, 1887): synonym of Cochlostyla marinduquensis Hidalgo, 1887 (superseded combination)
* Helicostyla metaformis (Ferussac, 1821): synonym of Dryocochlias metaformis (Férussac, 1821) (misspelling of original name, Corasia, Albers, 1850)
* Helicostyla mindoroensis W. J. Broderip, 1892 : synonym of Chrysallis mindoroensis (Broderip, 1841) (unaccepted generic combination)
* Helicostyla mirabilis (Ferussac, 1821): synonym of Cochlostyla mirabilis (Férussac, 1821) (superseded combination)
* Helicostyla montana Semper, 1891: synonym of Cochlostyla montana C. Semper, 1877 (superseded combination)
* Helicostyla monticula (G.B. Sowerby I, 1841): synonym of Orustia monticula (G. B. Sowerby I, 1841) (misspelling of original genus name, Orustia, Mörch, 1852)
* Helicostyla nimbosa (Broderip, 1841): synonym of Hypselostyla nimbosa (Broderip, 1841) (unaccepted combination)
* Helicostyla olanivanensis (Bartsch, 1913): synonym of Cochlostyla olanivanensis Bartsch, 1913
* Helicostyla opalina (G.B. Sowerby I, 1841): synonym of Phengus opalinus (G. B. Sowerby I, 1841)
* Helicostyla phaeostyla (Pfeiffer, 1856): synonym of Cochlostyla phaeostyla (L. Pfeiffer, 1857)
* Helicostyla pictor (Broderip, 1841): synonym of Cochlostyla pictor (Broderip, 1841) (superseded combination)
* Helicostyla polillensis (Pfeiffer, 1861): synonym of Cochlostyla polillensis (L. Pfeiffer, 1861) accepted as Calocochlea polillensis (L. Pfeiffer, 1861)
* Helicostyla pudibunda Semper, 1891: synonym of Pfeifferia pudibunda (C. Semper, 1877) (unaccepted combination)
* Helicostyla rehbeini (L. Pfeiffer, 1852): synonym of Dryocochlias rehbeini (L. Pfeiffer, 1852) (unaccepted combination)
* Helicostyla rufogaster (Lesson, 1831): synonym of Cochlostyla rufogaster (Lesson, 1832) (superseded combination)
* Helicostyla rustica (Mousson, 1849): synonym of Dryocochlias metaformis rustica (Mousson, 1849)
* Helicostyla sarcinosa (Ferussac, 1821): synonym of Helicobulinus sarcinosa (Férussac, 1821)
* Helicostyla satyra (Broderip, 1841): synonym of Cochlostyla satyrus (Broderip, 1841) (superseded combination)
* Helicostyla simplex (Jonas, 1843): synonym of Cochlostyla simplex (Jonas, 1843) (superseded combination)
* Helicostyla solivaga (Reeve, 1849): synonym of Cochlostyla solivaga (Reeve, 1849) (superseded combination)
* Helicostyla sphaerica (Sowerby I, 1841): synonym of Pachysphaera sphaerica (G. B. Sowerby I, 1841)
* Helicostyla submirabilis Möllendorff, 1897: synonym of Cochlostyla submirabilis Möllendorff, 1897 (unaccepted combination)
* * Helicostyla turbinoides (Broderip, 1841): synonym of Helicobulinus turbinoides (Broderip, 1841) (superseded combination)
* Helicostyla velata (Broderip, 1841): synonym of Hypselostyla velata (W.J. Broderip, 1841)
* Helicostyla ventricosa (Bruguière, 1792): synonym of Cochlostyla ventricosa (Bruguière, 1792) (superseded combination)
* Helicostyla viridostriata (Lea, 1840): synonym of Cochlodryas viridostriata (I. Lea, 1840) (superseded combination)
* Helicostyla woodiana (Lea, 1840): synonym of Cochlostyla woodiana (I. Lea, 1840) (superseded combination)