# Agraphydrus

Agraphydrus is a genus of water scavenger beetle in the family Hydrophilidae represented by 205 described species. It is distributed across the Afrotropical, Australasian, and Indomalayan realms.

## Taxonomy

The genus Agraphydrus was described for the first time by Régimbart in 1903.
It currently contains 205 described species, partly thanks to a series revisionary work developed by Albretch Komarek.

## Description

Small beetles (1.4–4.8 mm), pale/yellowish to dark brown in coloration, eyes not emarginate, labrum exposed, maxillary palps moderately long. Elytra without sutural striae, not laterally explanate; the elytral punctation ranges from very fine and subtle to coarse.
A complete diagnosis was presented by Girón and Short, but also see Komarek and collaborators.

## Habitat

According to Girón and Short, based on annotations by Komarek and collaborators:
Agraphydrus can be found in an extremely broad range of habitats, from rivers, streams and forest pools, to hygropetric environments around waterfalls or seepages over rocks; a few species have been collected in terrestrial habitats by sifting moss and leaves from near water bodies, or in the gravel along the bank of a river; in many cases specimens have been found associated with floating vegetation, mosses and algae.

## Species

* Agraphydrus abrasus Komarek and Freitag, 2020
* Agraphydrus activus Komarek and Hebauer, 2018
* Agraphydrus acutus Komarek, 2020
* Agraphydrus aethiopicus Komarek, 2020
* Agraphydrus agilis Komarek and Hebauer, 2018
* Agraphydrus albescens (Régimbart, 1903)
* Agraphydrus ampullatus Komarek and Freitag, 2020
* Agraphydrus anacaenoides Komarek, 2019
* Agraphydrus anatinus Komarek, 2018
* Agraphydrus andamanicus Komarek, 2018
* Agraphydrus andringitra Komarek, 2020
* Agraphydrus angulatus Komarek, 2019
* Agraphydrus angustatus Komarek, 2020
* Agraphydrus angustipenis Komarek, 2018
* Agraphydrus anhuianus (Hebauer, 2000)
* Agraphydrus annapurnensis Komarek, 2018
* Agraphydrus arduus Komarek and Hebauer, 2018
* Agraphydrus ater Komarek, 2018
* Agraphydrus atripalpis Komarek, 2020
* Agraphydrus attenuatus (Hansen, 1999)
* Agraphydrus audax Komarek and Hebauer, 2018
* Agraphydrus avita (Hansen, 1997)
* Agraphydrus bacchusi Komarek, 2019
* Agraphydrus balkeorum Komarek, 2019
* Agraphydrus batak Komarek and Freitag, 2020
* Agraphydrus bhutanensis Komarek, 2018
* Agraphydrus bicoloratus Komarek, 2020
* Agraphydrus bilardoi Komarek, 2020
* Agraphydrus biltoni Komarek, 2020
* Agraphydrus biprojectus Minoshima, Komarek, and Ôhara, 2015
* Agraphydrus borneensis Komarek, 2019
* Agraphydrus boukali Komarek, 2018
* Agraphydrus brevilobatus Komarek and Freitag, 2020
* Agraphydrus brevipenis Komarek, 2019
* Agraphydrus burmensis Komarek, 2019
* Agraphydrus calvus Komarek and Hebauer, 2018
* Agraphydrus camerunensis Komarek, 2020
* Agraphydrus cantonensis Komarek and Hebauer, 2018
* Agraphydrus carinatulus Komarek, 2019
* Agraphydrus cervus Komarek, 2019
* Agraphydrus ceylonensis Komarek, 2018
* Agraphydrus chinensis Komarek and Hebauer, 2018
* Agraphydrus cinnamum Komarek, 2018
* Agraphydrus clarus Komarek, 2019
* Agraphydrus comes Komarek and Hebauer, 2018
* Agraphydrus communis Komarek, 2018
* Agraphydrus confusus Komarek and Hebauer, 2018
* Agraphydrus congolensis Komarek, 2020
* Agraphydrus conicus Komarek and Hebauer, 2018
* Agraphydrus connexus Komarek and Hebauer, 2018
* Agraphydrus constrictus Komarek, 2018
* Agraphydrus contractus Komarek and Hebauer, 2018
* Agraphydrus coomani (d'Orchymont, 1927)
* Agraphydrus coronarius Minoshima, Komarek, and Ôhara, 2015
* Agraphydrus crassipenis Komarek, 2018
* Agraphydrus dapengensis Yang & Jia, 2021
* Agraphydrus decipiens Minoshima, Komarek, and Ôhara, 2015
* Agraphydrus delineatus Komarek, 2019
* Agraphydrus elongatus Ribera, Hernando, and Cieslak, 2019
* Agraphydrus engkari Komarek, 2019
* Agraphydrus excisus Komarek, 2019
* Agraphydrus exedis (d'Orchymont, 1937)
* Agraphydrus exiguus Komarek, 2019
* Agraphydrus falcatus Komarek, 2018
* Agraphydrus fasciatus Komarek and Hebauer, 2018
* Agraphydrus fikaceki Komarek and Hebauer, 2018
* Agraphydrus flavescens Komarek, 2020
* Agraphydrus flavipes Komarek, 2020
* Agraphydrus flavonotus Komarek, 2018
* Agraphydrus floresinus Komarek, 2019
* Agraphydrus fontis Komarek, 2020
* Agraphydrus forcipatus Komarek and Hebauer, 2018
* Agraphydrus fortis Komarek, 2018
* Agraphydrus fujianensis Komarek and Hebauer, 2018
* Agraphydrus geminus (d'Orchymont, 1932)
* Agraphydrus gereckei Komarek, 2020
* Agraphydrus gilvus Komarek, 2018
* Agraphydrus glaber Komarek, 2018
* Agraphydrus globipenis Komarek and Hebauer, 2018
* Agraphydrus goldschmidti Komarek, 2020
* Agraphydrus gracilipalpis Komarek, 2018
* Agraphydrus hamatus Komarek, 2019
* Agraphydrus hanseni (Satô and Yoshitomi, 2004)
* Agraphydrus heinrichi Komarek, 2018
* Agraphydrus helicopter Komarek, 2019
* Agraphydrus hendrichi Komarek, 2019
* Agraphydrus heterochromatus Komarek, 2019
* Agraphydrus hortensis Komarek, 2019
* Agraphydrus hygropetricus Komarek, 2018
* Agraphydrus igneus Komarek and Hebauer, 2018
* Agraphydrus imitans Komarek, 2019
* Agraphydrus indicus (d'Orchymont, 1932)
* Agraphydrus inflatus Komarek, 2018
* Agraphydrus infuscatus Komarek, 2019
* Agraphydrus insidiator Minoshima, Komarek, and Ôhara, 2015
* Agraphydrus ishiharai (Matsui, 1994)
* Agraphydrus jaechi (Hansen, 1999)
* Agraphydrus jankodadai Komarek, 2019
* Agraphydrus jilanzhui Komarek and Hebauer, 2018
* Agraphydrus kallar Komarek, 2018
* Agraphydrus kathapa Komarek, 2019
* Agraphydrus kempi (d'Orchymont, 1922)
* Agraphydrus khasiensis Komarek, 2018
* Agraphydrus kodaguensis Komarek, 2018
* Agraphydrus komareki Yang & Jia, 2021
* Agraphydrus laocaiensis Komarek, 2019
* Agraphydrus latus Komarek, 2019
* Agraphydrus longipalpus (Jia, 1998)
* Agraphydrus longipenis Komarek and Hebauer, 2018
* Agraphydrus lunaris Komarek, 2019
* Agraphydrus luteilateralis (Minoshima and Fujiwara, 2009)
* Agraphydrus madagascarensis Komarek, 2020
* Agraphydrus maehongsonensis Komarek, 2019
* Agraphydrus malayanus (Hebauer, 2000)
* Agraphydrus malkini Komarek, 2020
* Agraphydrus manfredjaechi Komarek, 2019
* Agraphydrus masatakai Minoshima, Komarek, and Ôhara, 2015
* Agraphydrus matoposensis Komarek, 2020
* Agraphydrus mazzoldii Komarek, 2019
* Agraphydrus meghalayanus Komarek, 2018
* Agraphydrus microphthalmus Komarek, 2019
* Agraphydrus minutissimus (Kuwert, 1890)
* Agraphydrus mirabilis Komarek, 2019
* Agraphydrus montanus Minoshima, Komarek, and Ôhara, 2015
* Agraphydrus muluensis Komarek, 2019
* Agraphydrus musculus Komarek, 2019
* Agraphydrus namthaensis Komarek, 2019
* Agraphydrus nanus Komarek, 2018
* Agraphydrus narusei (Satô, 1960)
* Agraphydrus nemorosus Komarek, 2019
* Agraphydrus nepalensis Komarek, 2018
* Agraphydrus niger Komarek, 2018
* Agraphydrus nigroflavus Komarek, 2019
* Agraphydrus obesus Komarek, 2019
* Agraphydrus obscuratus Komarek, 2018
* Agraphydrus obsoletus Komarek, 2018
* Agraphydrus occultus Komarek and Freitag, 2020
* Agraphydrus ogatai Minoshima, 2016
* Agraphydrus orbicularis Komarek, 2019
* Agraphydrus orientalis (d'Orchymont, 1932)
* Agraphydrus palawanensis Komarek and Freitag, 2020
* Agraphydrus pallidus Komarek, 2019
* Agraphydrus papuanus Komarek, 2019
* Agraphydrus pauculus (Knisch, 1924)
* Agraphydrus pauper Komarek, 2020
* Agraphydrus pelingeni Komarek and Freitag, 2020
* Agraphydrus penangensis Komarek, 2019
* Agraphydrus piceus Komarek, 2019
* Agraphydrus politus (Hansen, 1999)
* Agraphydrus praecipuus (d'Orchymont, 1937)
* Agraphydrus protentus Komarek, 2018
* Agraphydrus pseudoniger Yang & Jia, 2021
* Agraphydrus pullus Komarek, 2018
* Agraphydrus punctatellus Régimbart, 1903
* Agraphydrus punctulatus Komarek, 2018
* Agraphydrus puzhelongi (Jia, 2010)
* Agraphydrus pygmaeus (Knisch, 1924)
* Agraphydrus raucus Komarek, 2019
* Agraphydrus reductus Komarek and Hebauer, 2018
* Agraphydrus regularis (Hansen, 1999)
* Agraphydrus reticulatus Komarek, 2019
* Agraphydrus reticuliceps Komarek and Hebauer, 2018
* Agraphydrus rhodesiensis Komarek, 2020
* Agraphydrus rhomboideus Komarek, 2019
* Agraphydrus rivalis Komarek, 2020
* Agraphydrus robustus Komarek and Hebauer, 2018
* Agraphydrus rostratus Komarek, 2018
* Agraphydrus rugosus Komarek, 2018
* Agraphydrus sabulosus Yang & Jia, 2021
* Agraphydrus sarawakensis Komarek, 2019
* Agraphydrus schoedli Komarek, 2019
* Agraphydrus schoenmanni Komarek and Hebauer, 2018
* Agraphydrus scintillans Komarek, 2019
* Agraphydrus scutifer Komarek, 2020
* Agraphydrus setifer Komarek and Hebauer, 2018
* Agraphydrus shaverdoae Komarek, 2019
* Agraphydrus siamensis (Hansen, 1999)
* Agraphydrus sipekorum Komarek, 2018
* Agraphydrus skalei Komarek, 2019
* Agraphydrus spadix Komarek, 2019
* Agraphydrus spinosus Komarek, 2019
* Agraphydrus splendens Komarek and Hebauer, 2018
* Agraphydrus stagnalis (d'Orchymont, 1937)
* Agraphydrus stramineus Komarek, 2019
* Agraphydrus sucineus Komarek, 2019
* Agraphydrus sundaicus Komarek, 2019
* Agraphydrus tamdao Komarek, 2019
* Agraphydrus taprobanensis Komarek, 2018
* Agraphydrus tenuipalpis Komarek and Freitag, 2020
* Agraphydrus thaiensis Minoshima, Komarek, and Ôhara, 2015
* Agraphydrus tristis Komarek, 2019
* Agraphydrus tulipa Komarek, 2019
* Agraphydrus tumidus Komarek, 2020
* Agraphydrus tumulosus Komarek, 2018
* Agraphydrus umbrosus Komarek and Hebauer, 2018
* Agraphydrus uncinatus Komarek and Hebauer, 2018
* Agraphydrus usambaraensis Komarek, 2020
* Agraphydrus uvaensis (Hebauer, 2000)
* Agraphydrus vadoni Komarek, 2020
* Agraphydrus variabilis Komarek and Hebauer, 2018
* Agraphydrus vietnamensis Komarek, 2019
* Agraphydrus villiersi (Balfour-Browne, 1958)
* Agraphydrus wangmiaoi Komarek and Hebauer, 2018
* Agraphydrus yunnanensis Komarek and Hebauer, 2018
* Agraphydrus zetteli Komarek and Freitag, 2020