# 2012 in paleobotany

This paleobotany list records new fossil plant taxa that were to be described during the year 2012, as well as notes other significant paleobotany discoveries and events which occurred during 2012.

## Bryophytes

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Bryokhutuliinia crassimarginata | Sp nov | | Ignatov et al | Late Jurassic | Ukurey formation | Russia | A moss of uncertain placement. | |

## Lycophytes

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Winslowia | Gen. et sp. nov | Valid | Dunn et al. | Carboniferous (Serpukhovian) | Pride Mountain Formation | United States | A member of Isoetales belonging to the group Chaloneriaceae. Genus includes new species W. tuscumbiana. | |

## Marchantiophytes

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Riccardiothallus | Gen. et sp. nov | Valid | Guo et al. | Devonian (Pragian) | Posongchong Formation | China | A liverwort belonging to the family Aneuraceae. Genus includes new species R. devonicus. | |

## Ferns and fern allies

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Korallipteris | Gen. et comb. nov | Valid | Vera & Passalia | Triassic to Miocene | | Antarctica  Argentina  Chile  New Zealand | A fern of uncertain phylogenetic placement. A new genus for "Gleichenia" argentinica (1962); Additional species were originally placed in Gleichenia, Gleichenites and Microphyllopteris. |
| Millerocaulis tekelili | Sp. nov | Valid | Vera | Early Cretaceous (Aptian) | Cerro Negro Formation | Antarctica (Livingston Island) | An Osmundaceae fern. Originally placed in Millerocaulis; Moved to Claytosmunda tekelili in (2017). |
| Osmundopsis rafaelii | Sp. nov | Valid | Escapa & Cúneo | Early Jurassic | | Argentina | A member of Osmundaceae. |
| Todites cacereii | Sp. nov | Valid | Escapa & Cúneo | Early Jurassic | | Argentina | A member of Osmundaceae. |

## Bennettitales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Anomozamites harrisi | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Anomozamites heeri | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Anomozamites villosus | Sp. nov | Valid | Pott et al. | Probably late Middle Jurassic | Daohugou Beds | China | A member of Bennettitales. Originally described as a species of Anomozamites Moved to the Wielandiella villosus (2015). |
| Cycadolepis barnardii | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Cycadolepis karatauensis | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Cycadolepis kirillii | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Cycadolepis kokalensis | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Cycadolepis lata | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Cycadolepis maija | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Cycadolepis multistomata | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Cycadolepis pachydermatica | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Cycadolepis perfecta | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Cycadolepis prynadае | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Cycadolepis schairensis | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Nilssoniopteris chirchilica | Sp. nov | Valid | Kiritchkova & Nosova | Middle Jurassic (Aalenian) | Кokala Suite | Kazakhstan | |
| Nilssoniopteris localis | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Nilssoniopteris saveljevii | Sp. nov | Valid | Kiritchkova & Nosova | Middle Jurassic (Aalenian) | Tonasha Formation | Kazakhstan | |
| Pterophyllum chagaense | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Pterophyllum doludenkoae | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Pterophyllum originalum | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Pterophyllum pаrvisegmentatum | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Pterophyllum pinniforme | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Pterophyllum valentinaeа | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Pterophyllum vsevolodii | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Williamsonia antonia | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Williamsonia baranovaeа | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Williamsonia chagaensе | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Williamsonia kokalensis | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Williamsonia vialovaea | Sp. nov | Valid | Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |

## "Pteridospermatophytes"

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Ctenozamites schweitzerii | Sp. nov | Valid | Kiritchkova & Nosova | Middle Jurassic (Bajocian) | Karadiirmen' Formation | Kazakhstan | A member of Pteridospermatophyta of uncertain affinities. |
| Eretmonia macloughlinii | Sp. nov | Valid | Ryberg, Taylor & Taylor | Late Permian | Upper Buckley Formation | Antarctica | A member of Glossopteridales. |
| Kuvakospermum | Gen. et sp. nov | Valid | Naugolnykh & Sidorov | Permian (Wordian) | | Russia | A seed fern belonging to the family Peltaspermaceae. Genus includes new species K. pedatum. |
| Lidgettoniopsis | Gen. et sp. nov | Valid | Ryberg, Taylor & Taylor | Late Permian | Beacon Supergroup (upper Buckley Formation) | Antarctica | A seed fern belonging to the group Dictyopteridiales and the family Eretmoniaceae. Genus includes new species L. ramulus. |
| Scutum leiophyllum | Sp. nov | Valid | Ryberg, Taylor & Taylor | Late Permian | Beacon Supergroup (upper Buckley Formation) | Antarctica | A member of Glossopteridales belonging to the family Dictyopteridiaceae. |
| Vetlugospermum | Gen. et sp. nov | Valid | Naugolnykh | Early Triassic | | Russia | A seed fern belonging to the group Peltaspermales. Genus includes new species V. rombicum. |

## Ginkgoales

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes |
| Baiera kazachstanica | Sp. nov | Valid | Nosova & Kiritchkova in Kiritchkova & Nosova | Early Jurassic (Toarcian) | Кokala Suite | Kazakhstan | |
| Baieroxylon rocablanquense | Sp. nov | Valid | Gnaedinger | Early Jurassic | Roca Blanca Formation | Argentina | |
| Ginkgo cranei | Sp. nov | Valid | Zhou, Quan, & Liu | upper Paleocene | Sentinel Butte Formation | USA North Dakota | An extinct species in the modern genus Ginkgo |
| Ginkgo gomolitzkyana | Sp. nov | Valid | Nosova | Middle Jurassic | Angren Formation | Uzbekistan | A member of Ginkgoales. |
| Ginkgo huolinhensis | Sp. nov | Valid | Dong & Sun | Early Cretaceous | Huolinhe Formation | China | |
| Ginkgoites barnardianus | Sp. nov | Valid | Nosova & Kiritchkova in Kiritchkova & Nosova | Middle Jurassic (Aalenian) | Кokala Suite | Kazakhstan | |
| Ginkgoites irdalicus | Sp. nov | Valid | Nosova & Kiritchkova in Kiritchkova & Nosova | Early Jurassic | Ergozi Suite | Kazakhstan | |
| Ginkgoites kelendensis | Sp. nov | Valid | Nosova & Kiritchkova in Kiritchkova & Nosova | Middle Jurassic (Aalenian) | Tonasha Formation | Kazakhstan | |
| Ginkgoites ketovianus | Sp. nov | Valid | Nosova & Kiritchkova in Kiritchkova & Nosova | Middle Jurassic (Bathonian) | Sarydiirmeny Suite | Kazakhstan | |
| Leptotoma asiatica | Sp. nov | Valid | Nosova & Kiritchkova in Kiritchkova & Nosova | Middle Jurassic (Bajocian) | Karadiirmen' Formation | Kazakhstan | |
| Pseudotorellia karadiirmenica | Sp. nov | Valid | Nosova & Kiritchkova in Kiritchkova & Nosova | Middle Jurassic (Bajocian) | Karadiirmen' Formation | Kazakhstan | |

## Conifers
### Araucariaceae

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Emwadea | Gen. et sp. nov | Valid | Dettmann, Clifford & Peters | Early Cretaceous (Albian) | Winton Formation | Australia | A member of the family Araucariaceae. Genus includes new species E. microcarpa. | |

### Cheirolepidiaceae

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Duartenia | Gen. et sp. nov | Valid | Mohr et al. | Early Cretaceous | Crato Formation | Brazil | A possible member of Cheirolepidiaceae. The type species is Duartenia araripensis. | |
| Glenrosa falcata | Sp. nov | Valid | Gomez, Ewin & Daviero-Gomez | Early Cretaceous (Barremian) | Calizas de La Huergina Formation | Spain | A conifer. | |

### Cupressaceae

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Calocedrus huashanensis | Sp. nov | Valid | Shi, Zhou, & Xie | Oligocene | Ningming Formation | China | A Calocedrus species | |
| Fokienia shengxianensis | Sp. nov | Valid | He, Sun & Liu | Late Miocene | Shengxian Formation Xiaolongtan Formation | China | Originally placed in Fokienia. Moved to Calocedrus shengxianensis in (2015). | |

### Pinaceae

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes |
| Eathiestrobus | Gen. et sp. nov | Valid | Rothwell et al. | Late Jurassic (Kimmeridgian) | Kimmeridge Clay Formation | United Kingdom | A member of Pinaceae. The type species is Eathiestrobus mackenziei. |
| Picea burtonii | Sp. nov | Valid | Klymiuk & Stockey | Early Cretaceous (Valanginian) | | Canada | A spruce species seed cone. |
| Pinus yorkshirensis | Sp. nov | Valid | Ryberg et al. | Early Cretaceous (Hauterivian-Barremian) | Speeton Clay Formation | United Kingdom | A pine. |

### Podocarpaceae

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Dacrycarpus puertae | Sp. nov | Valid | Wilf | Eocene | | Argentina | A species of Dacrycarpus | |

### Taxaceae

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Marskea kazachstanica | Sp. nov | Valid | Nosova & Kiritchkova in Kiritchkova & Nosova | Early Jurassic (Toarcian) | Kokala Suite | Kazakhstan | | |
| Torreya kazachstanica | Sp. nov | Valid | Nosova & Kiritchkova in Kiritchkova & Nosova | Early Jurassic (Toarcian) | Kokala Suite | Kazakhstan | | |

### Other conifers

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
| Elatocladus bulakus | Sp. nov | Valid | Nosova & Kiritchkova in Kiritchkova & Nosova | Early Jurassic (Toarcian) | Kokala Suite | Kazakhstan | A conifer of uncertain affinities. | |
| Plyophyllioxylon | Gen. et sp. nov | Valid | Feng et al. | Permian (Roadian/Wordian) | Lower Shihhotse Formation | China | A conifer of uncertain phylogenetic placement. Genus includes new species P. hulstaiense. | |

## Other gymnosperms

| Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
| Nilssonia prosvirjakovaea | Sp. nov | Valid | Kiritchkova & Nosova | Middle Jurassic (Bajocian) | Karadiirmen' Formation | Kazakhstan | A cycad. | |
| Rehezamites | Gen. et sp. nov | Valid | Pott et al. | Early Cretaceous (Aptian) | Yixian Formation | China | A seed plant of uncertain phylogenetic placement. Genus includes new species R. anisolobus. | |

## Angiosperms

| Name | Novelty | Status | Authors | Age | Unit | Location | Notes |
| Archaeampelos senonica | Sp. nov | Valid | Alekseev | Late Cretaceous | | Russia | |
| Baasia | Gen. et sp. nov | Valid | Estrada-Ruiz et al. | Late Cretaceous (probably early to middle Maastrichtian) | McRae Formation | United States | A member of the family Celastraceae. Genus includes new species B. armendarisense. |
| Bertilanthus | Gen. et sp. nov | Valid | Friis & Pedersen | Late Cretaceous | | Sweden | An asterid. Genus includes new species B. scanicus. |
| Camptodromites sibiricus | Sp. nov | Valid | Alekseev | Late Cretaceous | | Russia | |
| Celastrinites lanceolatus | Sp. nov | Valid | Alekseev | Late Cretaceous | | Russia | |
| Chaneya hainanensis | Sp. nov | Valid | Feng & Jin | Eocene | Changchang Basin | China | A fossil fruit. |
| Cissites basicordatus | Sp. nov | Valid | Alekseev | Late Cretaceous | | Russia | |
| Cissus lombardii | Sp. nov | Valid | Manchester, Chen & Lott | Early Oligocene | Mancora Formation | Peru | A species of Cissus. |
| Eucalyptus caldericola | Sp. nov | Valid | Hermsen, Gandolfo & Zamaloa | Eocene (Ypresian) | | Argentina | A species of Eucalyptus. |
| Eucalyptus frenguelliana | Sp. nov | Valid | Gandolfo & Zamaloa in Hermsen, Gandolfo & Zamaloa | Eocene (Ypresian) | | Argentina | A species of Eucalyptus. |
| Eucalyptus lynchiae | Sp. nov | Valid | Gandolfo & Hermsen in Hermsen, Gandolfo & Zamaloa | Eocene (Ypresian) | | Argentina | A species of Eucalyptus. |
| Fulleroxylon | Gen. et sp. nov | Valid | Estrada-Ruiz et al. | Late Cretaceous (probably early to middle Maastrichtian) | McRae Formation | United States | A member of the family Myrtaceae. Genus includes new species F. armendarisense. |
| Juglandiphyllites microdentatus | Sp. nov | Valid | Alekseev | Late Cretaceous | | Russia | |
| Laurinoxylon stichkai | Sp. nov | Valid | Boonchai & Manchester | Early Eocene | Bridger Formation | United States | A member of Lauraceae described on the basis of fossil wood. |
| Laurophyllum calicarioides | Sp. nov | Valid | Bannister, Conran & Lee | Early Miocene | | New Zealand | A member of Lauraceae with affinities to members of the genus Litsea. |
| Laurophyllum lacustris | Sp. nov | Valid | Bannister, Conran & Lee | Early Miocene | | New Zealand | A member of Lauraceae with affinities to members of the genus Beilschmiedia. |
| Laurophyllum maarensis | Sp. nov | Valid | Bannister, Conran & Lee | Early Miocene | | New Zealand | A member of Lauraceae with affinities to members of the genus Cryptocarya. |
| Laurophyllum microphyllum | Sp. nov | Valid | Bannister, Conran & Lee | Early Miocene | | New Zealand | A member of Lauraceae with affinities to members of the genus Cryptocarya. |
| Laurophyllum otagoensis | Sp. nov | Valid | Bannister, Conran & Lee | Early Miocene | | New Zealand | A member of Lauraceae with affinities to members of the genus Beilschmiedia. |
| Laurophyllum sylvestris | Sp. nov | Valid | Bannister, Conran & Lee | Early Miocene | | New Zealand | A member of Lauraceae with affinities to members of the genus Beilschmiedia. |
| Laurophyllum taieriensis | Sp. nov | Valid | Bannister, Conran & Lee | Early Miocene | | New Zealand | A member of Lauraceae with affinities to members of the genus Cryptocarya. |
| Laurophyllum vulcanicola | Sp. nov | Valid | Bannister, Conran & Lee | Early Miocene | | New Zealand | A member of Lauraceae with affinities to members of the genus Beilschmiedia. |
| Laurophyllum waipiata | Sp. nov | Valid | Bannister, Conran & Lee | Early Miocene | | New Zealand | A member of Lauraceae with affinities to members of the genus Cryptocarya. |
| Litseopsis nova-zelandiae | Sp. nov | Valid | Bannister, Conran & Lee | Early Miocene | | New Zealand | A member of Lauraceae. |
| Mahonia grimmii | Sp. nov | Valid | Güner & Denk | Miocene | Eskihisar Formation | Turkey | A species of Mahonia. |
| Mahonia somaensis | Sp. nov | Valid | Güner & Denk | Miocene | Soma Formation | Turkey | A species of Mahonia. |
| Morinda chinensis | Sp. nov | Valid | Shi et al. | Eocene | Changchang Formation | China | A species of Morinda. |
| Newtonia mushensis | Sp. nov | Valid | Pan et al. | Early Miocene | | Ethiopia | A species of Newtonia. |
| Palaeophytocrene hammenii | Sp. nov | Valid | Stull in Stull et al. | Middle-late Paleocene | Bogotá Formation | Colombia | A member of Icacinaceae belonging to the tribe Phytocreneae. |
| Palaeophytocrene piggae | Sp. nov | Valid | Stull in Stull et al. | Late Paleocene | Sentinel Butte Formation | United States | A member of Icacinaceae belonging to the tribe Phytocreneae. |
| Pygmaeoxylon | Gen. et sp. nov | Valid | Estrada-Ruiz et al. | Late Cretaceous (probably early to middle Maastrichtian) | McRae Formation | United States | A magnoliid of uncertain phylogenetic placement. Genus includes new species P. paucipora. |
| Pyrenacantha austroamericana | Sp. nov | Valid | Stull in Stull et al. | Late early Oligocene | | Peru | A species of Pyrenacantha. |
| Schenkeriphyllum glanduliferum | Gen. et sp. nov | Valid | Mohr, Coiffard & Bernardes-de-Oliveira | Early Cretaceous (Aptian) | Crato Formation | Brazil | Fossil magnolialean angiosperm. |
| Wilsonoxylon | Gen. et sp. nov | Valid | Boonchai & Manchester | Early Eocene | Bridger Formation | United States | A member of Canellaceae described on the basis of fossil wood. Genus includes new species W. edenense. |