# List of butterflies of Sudan and South Sudan

This is a list of butterflies of Sudan and South Sudan. About 341 species are known from the Sudans, 6 of which are endemic.

## Papilionidae
### Papilioninae
#### Papilionini

* Papilio antimachus Drury, 1782
* Papilio zalmoxis Hewitson, 1864
* Papilio nireus lyaeus Doubleday, 1845
* Papilio nireus pseudonireus Felder & Felder, 1865
* Papilio plagiatus Aurivillius, 1898
* Papilio phorcas sudanicola Storace, 1965
* Papilio rex franciscae Carpenter, 1928
* Papilio mechowi Dewitz, 1881
* Papilio mechowianus Dewitz, 1885
* Papilio echerioides joiceyi Gabriel, 1945
* Papilio jacksoni imatonga Clifton & Collins, 1997
* Papilio nobilis didingensis Carpenter, 1928
* Papilio hesperus sudana Gabriel, 1945
* Papilio lormieri Distant, 1874
* Papilio mackinnoni Sharpe, 1891

#### Leptocercini

* Graphium antheus (Cramer, 1779)
* Graphium policenes policenes (Cramer, 1775)
* Graphium policenes telloi Hecq, 1999
* Graphium colonna (Ward, 1873)
* Graphium angolanus baronis (Ungemach, 1932)
* Graphium ridleyanus (White, 1843)
* Graphium leonidas (Fabricius, 1793)
* Graphium philonoe whalleyi (Talbot, 1929)
* Graphium almansor escherichi (Gaede, 1915)

## Pieridae
### Coliadinae

* Eurema brigitta (Stoll, [1780])
* Eurema hecabe solifera (Butler, 1875)
* Catopsilia florella (Fabricius, 1775)
* Colias electo pseudohecate Berger, 1940
* Colias marnoana Rogenhofer, 1884

### Pierinae

* Colotis antevippe zera (Lucas, 1852)
* Colotis aurigineus (Butler, 1883)
* Colotis aurora evarne (Klug, 1829)
* Colotis celimene angusi Rothschild, 1921
* Colotis celimene sudanicus (Aurivillius, 1905)
* Colotis chrysonome (Klug, 1829)
* Colotis daira stygia (Felder & Felder, 1865)
* Colotis danae eupompe (Klug, 1829)
* Colotis elgonensis basilewskyi Berger, 1956
* Colotis ephyia (Klug, 1829)
* Colotis euippe exole (Reiche, 1850)
* Colotis halimede (Klug, 1829)
* Colotis hetaera aspasia (Ungemach, 1932)
* Colotis liagore (Klug, 1829)
* Colotis phisadia (Godart, 1819)
* Colotis pleione nilus Talbot, 1942
* Colotis protomedia (Klug, 1829)
* Colotis rogersi (Dixey, 1915)
* Colotis ungemachi (Le Cerf, 1922)
* Colotis vesta princeps Talbot, 1939
* Colotis vesta velleda (Lucas, 1852)
* Colotis vestalis castalis (Staudinger, 1884)
* Colotis eris (Klug, 1829)
* Calopieris eulimene (Klug, 1829)
* Eronia cleodora Hübner, 1823
* Eronia leda (Boisduval, 1847)
* Pinacopterix eriphia melanarge (Butler, 1886)
* Pinacopterix eriphia tritogenia (Klug, 1829)
* Nepheronia argia (Fabricius, 1775)
* Nepheronia buquetii (Boisduval, 1836)
* Nepheronia thalassina sinalata (Suffert, 1904)
* Euchloe falloui (Allard, 1867)
* Leptosia alcesta inalcesta Bernardi, 1959

#### Pierini

* Appias epaphia contracta (Butler, 1888)
* Appias phaola intermedia Dufrane, 1948
* Appias sabina (Felder & Felder, [1865])
* Appias sylvia sudanensis Talbot, 1932
* Pontia glauconome Klug, 1829
* Mylothris agathina (Cramer, 1779)
* Mylothris chloris (Fabricius, 1775)
* Mylothris jacksoni nagichota Talbot, 1944
* Mylothris rubricosta (Mabille, 1890)
* Mylothris rueppellii septentrionalis Carpenter, 1928
* Mylothris schumanni uniformis Talbot, 1944
* Dixeia dixeyi (Neave, 1904)
* Dixeia doxo doxo (Godart, 1819)
* Dixeia doxo venatus (Butler, 1871)
* Dixeia orbona vidua (Butler, 1900)
* Belenois aurota (Fabricius, 1793)
* Belenois creona (Cramer, [1776])
* Belenois raffrayi (Oberthür, 1878)
* Belenois solilucis loveni (Aurivillius, 1921)
* Belenois subeida (Felder & Felder, 1865)
* Belenois sudanensis (Talbot, 1929)
* Belenois theora laeta (Weymer, 1903)
* Belenois thysa meldolae Butler, 1872

## Lycaenidae
### Miletinae
#### Miletini

* Megalopalpus zymna (Westwood, 1851)
* Lachnocnema abyssinica Libert, 1996
* Lachnocnema emperamus (Snellen, 1872)
* Lachnocnema divergens Gaede, 1915
* Lachnocnema reutlingeri perspicua Libert, 1996

### Poritiinae
#### Liptenini

* Alaena subrubra Bethune-Baker, 1915
* Pentila pauli multiplagata Bethune-Baker, 1908
* Pentila umangiana connectens Hulstaert, 1924
* Telipna sulpitia Hulstaert, 1924
* Telipna albofasciata laplumei Devos, 1917
* Ornipholidotos overlaeti intermedia Libert, 2005
* Ornipholidotos latimargo (Hawker-Smith, 1933)
* Mimacraea krausei karschioides Carpenter & Jackson, 1950
* Liptena ferrymani (Grose-Smith & Kirby, 1891)
* Liptena xanthostola xantha (Grose-Smith, 1901)
* Tetrarhanis ilala (Riley, 1929)
* Larinopoda lircaea (Hewitson, 1866)

#### Epitolini

* Stempfferia cercenoides (Holland, 1890)
* Cephetola dolorosa (Roche, 1954)
* Phytala elais ugandae Jackson, 1964

### Aphnaeinae

* Pseudaletis agrippina Druce, 1888
* Chloroselas tamaniba (Walker, 1870)
* Chloroselas taposana Riley, 1932
* Cigaritis acamas bellatrix (Butler, 1886)
* Cigaritis nilus (Hewitson, 1865)
* Zeritis neriene Boisduval, 1836
* Zeritis pulcherrima Aurivillius, 1923
* Axiocerses harpax kadugli Talbot, 1935
* Axiocerses amanga (Westwood, 1881)
* Aphnaeus coronae Talbot, 1935

### Theclinae

* Myrina silenus (Fabricius, 1775)
* Myrina subornata nuba Talbot, 1935
* Dapidodigma demeter sudsudana d'Abrera, 1980
* Hypolycaena liara Druce, 1890
* Hypolycaena obscura Stempffer, 1947
* Leptomyrina sudanica Stempffer, 1964
* Iolaus alienus ugandae Stempffer, 1953
* Iolaus scintillans Aurivillius, 1905
* Iolaus sudanicus Aurivillius, 1905
* Iolaus pallene (Wallengren, 1857)
* Iolaus menas Druce, 1890
* Iolaus ismenias (Klug, 1834)
* Stugeta marmoreus (Butler, 1866)
* Paradeudorix ituri ugandae (Talbot, 1935)
* Hypomyrina mimetica Libert, 2004
* Deudorix dinochares Grose-Smith, 1887
* Deudorix dinomenes diomedes Jackson, 1966
* Deudorix livia (Klug, 1834)
* Deudorix lorisona baronica Ungemach, 1932
* Capys bamptoni Henning & Henning, 1988

### Polyommatinae
#### Lycaenesthini

* Anthene butleri (Oberthür, 1880)
* Anthene crawshayi minuta (Bethune-Baker, 1916)
* Anthene hodsoni (Talbot, 1935)
* Anthene lunulata (Trimen, 1894)
* Anthene wilsoni (Talbot, 1935)
* Anthene lusones (Hewitson, 1874)
* Anthene nigeriae (Aurivillius, 1905)

#### Polyommatini

* Cupidopsis jobates mauritanica Riley, 1932
* Uranothauma delatorum Heron, 1909
* Uranothauma heritsia intermedia (Tite, 1958)
* Leptotes marginalis (Stempffer, 1944)
* Tuxentius margaritaceus (Sharpe, 1892)
* Tarucus balkanicus (Freyer, 1843)
* Tarucus legrasi Stempffer, 1948
* Tarucus rosacea (Austaut, 1885)
* Tarucus theophrastus (Fabricius, 1793)
* Tarucus ungemachi Stempffer, 1942
* Zizeeria karsandra (Moore, 1865)
* Actizera stellata (Trimen, 1883)
* Euchrysops alberta (Butler, 1901)
* Euchrysops albistriata (Capronnier, 1889)
* Euchrysops crawshayi (Butler, 1899)
* Euchrysops cyclopteris (Butler, 1876)
* Euchrysops nilotica (Aurivillius, 1904)
* Chilades eleusis (Demaison, 1888)
* Lepidochrysops albilinea Tite, 1959
* Lepidochrysops negus wau (Wichgraf, 1921)
* Lepidochrysops nigrita Tite, 1959
* Lepidochrysops polydialecta (Bethune-Baker, [1923])
* Lepidochrysops victoriae occidentalis Libert & Collins, 2001

## Nymphalidae
### Danainae
#### Danaini

* Danaus chrysippus chrysippus (Linnaeus, 1758)
* Danaus chrysippus alcippus (Cramer, 1777)
* Danaus dorippus (Klug, 1845)
* Tirumala formosa neumanni (Rothschild, 1902)
* Tirumala petiverana (Doubleday, 1847)
* Amauris niavius aethiops Rothschild & Jordan, 1903
* Amauris tartarea Mabille, 1876
* Amauris albimaculata sudanica Talbot, 1940
* Amauris echeria mongallensis Carpenter, 1928
* Amauris echeria steckeri Kheil, 1890
* Amauris hecate (Butler, 1866)

### Satyrinae
#### Melanitini

* Gnophodes betsimena parmeno Doubleday, 1849
* Melanitis libya Distant, 1882

#### Satyrini

* Bicyclus angulosa (Butler, 1868)
* Bicyclus campus (Karsch, 1893)
* Bicyclus istaris (Plötz, 1880)
* Bicyclus kenia (Rogenhofer, 1891)
* Bicyclus milyas (Hewitson, 1864)
* Bicyclus pavonis (Butler, 1876)
* Bicyclus martius sanaos (Hewitson, 1866)
* Bicyclus saussurei angustus Condamin, 1970
* Heteropsis perspicua (Trimen, 1873)
* Ypthima albida Butler, 1888
* Ypthima antennata van Son, 1955
* Ypthima condamini Kielland, 1982
* Ypthima doleta Kirby, 1880
* Ypthima pupillaris Butler, 1888
* Ypthima simplicia Butler, 1876
* Ypthimomorpha itonia (Hewitson, 1865)

### Charaxinae
#### Charaxini

* Charaxes varanes vologeses (Mabille, 1876)
* Charaxes fulvescens monitor Rothschild, 1900
* Charaxes boueti rectans Rothschild & Jordan, 1903
* Charaxes cynthia kinduana Le Cerf, 1923
* Charaxes lactetinctus ungemachi Le Cerf, 1927
* Charaxes jasius Poulton, 1926
* Charaxes epijasius Reiche, 1850
* Charaxes hansali baringana Rothschild, 1905
* Charaxes brutus angustus Rothschild, 1900
* Charaxes julius somalicus Rothschild, 1900
* Charaxes ansorgei kinyeti Plantrou, 1989
* Charaxes pollux (Cramer, 1775)
* Charaxes druceanus vivianae Plantrou, 1982
* Charaxes eudoxus imatongensis Plantrou, 1982
* Charaxes eudoxus mechowi Rothschild, 1900
* Charaxes numenes aequatorialis van Someren, 1972
* Charaxes tiridates tiridatinus Röber, 1936
* Charaxes bipunctatus ugandensis van Someren, 1972
* Charaxes smaragdalis gobyae Plantrou, 1989
* Charaxes etesipe (Godart, 1824)
* Charaxes achaemenes monticola van Someren, 1970
* Charaxes jahlusa ganalensis Carpenter, 1937
* Charaxes eupale latimargo Joicey & Talbot, 1921
* Charaxes dilutus ngonga van Someren, 1974
* Charaxes baumanni didingensis van Someren, 1971
* Charaxes etheocles carpenteri van Someren & Jackson, 1957
* Charaxes cedreatis Hewitson, 1874
* Charaxes viola picta van Someren & Jackson, 1952
* Charaxes paphianus subpallida Joicey & Talbot, 1925
* Charaxes kahldeni Homeyer & Dewitz, 1882
* Charaxes zoolina (Westwood, [1850])
* Charaxes lycurgus bernardiana Plantrou, 1978
* Charaxes zelica depuncta Joicey & Talbot, 1921
* Charaxes amandae Rydon, 1989

#### Euxanthini

* Charaxes crossleyi ansorgei (Rothschild, 1903)

### Apaturinae

* Apaturopsis cleochares (Hewitson, 1873)

### Nymphalinae
#### Nymphalini

* Vanessa dimorphica (Howarth, 1966)
* Junonia chorimene (Guérin-Méneville, 1844)
* Junonia terea tereoides (Butler, 1901)
* Junonia westermanni Westwood, 1870
* Protogoniomorpha parhassus (Drury, 1782)
* Protogoniomorpha temora (Felder & Felder, 1867)
* Precis archesia ugandensis (McLeod, 1980)
* Precis coelestina Dewitz, 1879
* Precis octavia (Cramer, 1777)
* Precis tugela aurorina (Butler, 1894)
* Hypolimnas anthedon (Doubleday, 1845)
* Hypolimnas misippus (Linnaeus, 1764)
* Hypolimnas salmacis platydema Rothschild & Jordan, 1903
* Mallika jacksoni (Sharpe, 1896)

### Biblidinae
#### Biblidini

* Byblia anvatara acheloia (Wallengren, 1857)
* Mesoxantha ethosea reducta Rothschild, 1918
* Neptidopsis ophione nucleata Grünberg, 1911

### Limenitinae
#### Limenitidini

* Pseudoneptis bugandensis Stoneham, 1935
* Pseudacraea acholica Riley, 1932
* Pseudacraea boisduvalii (Doubleday, 1845)
* Pseudacraea eurytus (Linnaeus, 1758)
* Pseudacraea lucretia protracta (Butler, 1874)

#### Neptidini

* Neptis metella (Doubleday, 1848)
* Neptis nemetes nemetes Hewitson, 1868
* Neptis nemetes margueriteae Fox, 1968
* Neptis occidentalis Rothschild, 1918

#### Adoliadini

* Catuna crithea (Drury, 1773)
* Pseudargynnis hegemone (Godart, 1819)
* Euphaedra medon fraudata van Someren, 1935
* Euphaedra neumanni Rothschild, 1902
* Euphaedra harpalyce sudanensis Talbot, 1929

### Heliconiinae
#### Acraeini

* Acraea chilo Godman, 1880
* Acraea insignis Distant, 1880
* Acraea neobule Doubleday, 1847
* Acraea pseudolycia astrigera Butler, 1899
* Acraea quirina (Fabricius, 1781)
* Acraea zetes (Linnaeus, 1758)
* Acraea abdera Hewitson, 1852
* Acraea cepheus (Linnaeus, 1758)
* Acraea egina (Cramer, 1775)
* Acraea guluensis Le Doux, 1932
* Acraea caecilia (Fabricius, 1781)
* Acraea caldarena Hewitson, 1877
* Acraea doubledayi Guérin-Méneville, 1849
* Acraea marnois Rogenhofer, 1890
* Acraea pseudegina Westwood, 1852
* Acraea sykesi Sharpe, 1902
* Acraea aganice orientalis (Ungemach, 1932)
* Acraea epaea paragea (Grose-Smith, 1900)
* Acraea epiprotea (Butler, 1874)
* Acraea macarista (Sharpe, 1906)
* Acraea poggei nelsoni Grose-Smith & Kirby, 1892
* Acraea pseuderyta Godman & Salvin, 1890
* Acraea acerata Hewitson, 1874
* Acraea bonasia (Fabricius, 1775)
* Acraea encedana Pierre, 1976
* Acraea serena (Fabricius, 1775)
* Acraea esebria Hewitson, 1861
* Acraea jodutta (Fabricius, 1793)
* Acraea johnstoni Godman, 1885
* Acraea lycoa Godart, 1819
* Acraea pharsalus pharsalus Ward, 1871
* Acraea pharsalus rhodina Rothschild, 1902
* Acraea oreas Sharpe, 1891

#### Argynnini

* Issoria hanningtoni imatonga (Riley, 1932)

#### Vagrantini

* Lachnoptera anticlia (Hübner, 1819)
* Phalanta eurytis eurytis (Doubleday, 1847)
* Phalanta eurytis microps (Rothschild & Jordan, 1903)
* Phalanta phalantha aethiopica (Rothschild & Jordan, 1903)

## Hesperiidae
### Coeliadinae

* Coeliades forestan (Stoll, [1782])
* Coeliades keithloa menelik (Ungemach, 1932)
* Coeliades pisistratus (Fabricius, 1793)

### Pyrginae
#### Celaenorrhinini

* Celaenorrhinus proxima (Mabille, 1877)
* Eretis lugens (Rogenhofer, 1891)
* Eretis melania Mabille, 1891
* Eretis mixta Evans, 1937
* Sarangesa aza Evans, 1951
* Sarangesa brigida sanaga Miller, 1964
* Sarangesa laelius (Mabille, 1877)
* Sarangesa lucidella (Mabille, 1891)
* Sarangesa phidyle (Walker, 1870)

#### Tagiadini

* Tagiades flesus (Fabricius, 1781)
* Eagris decastigma purpura Evans, 1937
* Eagris denuba obliterata Carpenter, 1928
* Eagris lucetia (Hewitson, 1875)
* Eagris tigris liberti Collins & Larsen, 2005
* Caprona adelica Karsch, 1892
* Netrobalane canopus (Trimen, 1864)
* Abantis bismarcki Karsch, 1892

#### Carcharodini

* Spialia doris (Walker, 1870)
* Spialia zebra bifida (Higgins, 1924)

### Hesperiinae
#### Aeromachini

* Prosopalpus styla Evans, 1937
* Kedestes protensa Butler, 1901
* Kedestes rogersi Druce, 1907
* Gorgyra aretina (Hewitson, 1878)
* Pardaleodes incerta (Snellen, 1872)
* Pardaleodes sator pusiella Mabille, 1877
* Pardaleodes tibullus (Fabricius, 1793)
* Osmodes thora (Plötz, 1884)
* Acleros mackenii olaus (Plötz, 1884)
* Andronymus caesar philander (Hopffer, 1855)
* Chondrolepis niveicornis (Plötz, 1883)
* Zophopetes dysmephila (Trimen, 1868)
* Gretna cylinda (Hewitson, 1876)
* Caenides dacena (Hewitson, 1876)
* Fresna nyassae (Hewitson, 1878)

#### Baorini

* Borbo fallax (Gaede, 1916)
* Borbo perobscura (Druce, 1912)
* Gegenes nostrodamus (Fabricius, 1793)

### Heteropterinae

* Metisella quadrisignatus (Butler, 1894)
* Metisella tsadicus (Aurivillius, 1905)
* Lepella lepeletier (Latreille, 1824)