# Anapidae

Anapidae is a family of rather small spiders with 233 described extant species in 59 genera. It includes the former family Micropholcommatidae as the subfamily Micropholcommatinae, and the former family Holarchaeidae. Most species are less than 2 millimetres (0.079 in) long.
They generally live in leaf litter and moss on the floor of rain forest. Many build orb webs with a diameter less than 3 centimetres (1.2 in). In some species, such as Pseudanapis parocula, the pedipalps of the female are reduced to coxal stumps.

## Description

Spiders of this family are very small, usually less than two millimeters long, and lack a cribellum. They can have zero, six or eight eyes, the rear median eyes either reduced or missing. One species: 'Epigastrina typhlops' (Rix & Harvey, 2010) has no eyes, an adaption to life underground. In some genera the carapace is modified so that the eyes are raised higher than usual. Color can range from reddish brown, yellowish brown to pale, pigment reduced, creams. Generally both margins of chelicerae have teeth, one exception being 'Acrobleps hygrophilus' who lacks retromarginal teeth. The legs are short and spineless, though there are some with reduced spination such as the genus: 'Teutoniella' or with only one or two long spines such as 'Borneanapis'). The labium has a spur that extends between the chelicerae and can be seen when the chelicerae are spread.

## Distribution

Anapidae are found worldwide, particularly in South America, Africa, Asia, Australia and New Zealand. Few genera occur in North America or Europe. Only Comaroma simoni and the three species of Zangherella are found in Europe; Gertschanapis shantzi and Comaroma mendocino are found in the United States.

## Systematics

The family Micropholcommatidae was synonymized with this family by Schütt in 2003 and by Lopa et al. in 2011, a change that has been accepted by the World Spider Catalog. Similarly, the family Holarchaeidae was synonymized by Dimitrov et al. in 2017, and likewise accepted by the World Spider Catalog.

## Genera

As of December 2024, the World Spider Catalog accepts the following extant genera:
* Acrobleps Hickman, 1979 — Australia
* Algidiella Rix & Harvey, 2010 — New Zealand
* Anapis Simon, 1895 — South America, Central America, Mexico, Jamaica
* Anapisona Gertsch, 1941 — South America, Central America, Saint Vincent and the Grenadines, Mexico
* Austropholcomma Rix & Harvey, 2010 — Australia
* Borneanapis Snazell, 2009 — Indonesia
* Caledanapis Platnick & Forster, 1989 — New Caledonia
* Chasmocephalon O. Pickard-Cambridge, 1889 — Australia
* Comaroma Bertkau, 1889 — Asia, United States
* Conculus Kishida, 1940 — Papua New Guinea, Asia
* Crassanapis Platnick & Forster, 1989 — Chile, Argentina
* Crozetulus Hickman, 1939 — Africa
* Dippenaaria Wunderlich, 1995 — South Africa
* Elanapis Platnick & Forster, 1989 — Chile
* Enielkenie Ono, 2007 — Taiwan
* Eperiella Rix & Harvey, 2010 — Chile, Australia
* Epigastrina Rix & Harvey, 2010 — Australia
* Eterosonycha Butler, 1932 — Australia
* Forsteriola Brignoli, 1981 — Burundi, Rwanda, Congo
* Gaiziapis Miller, Griswold & Yin, 2009 — China
* Gertschanapis Platnick & Forster, 1990 — United States
* Gigiella Rix & Harvey, 2010 — Australia, Chile
* Guiniella Rix & Harvey, 2010 — Papua New Guinea
* Hickmanapis Platnick & Forster, 1989 — Australia
* Holarchaea Forster, 1955 — Australia, New Zealand
* Mandanapis Platnick & Forster, 1989 — New Caledonia
* Maxanapis Platnick & Forster, 1989 — Australia
* Metanapis Brignoli, 1981 — Africa, Nepal
* Micropholcomma Crosby & Bishop, 1927 — Australia
* Minanapis Platnick & Forster, 1989 — Chile, China, Argentina
* Montanapis Platnick & Forster, 1989 — New Caledonia
* Normplatnicka Rix & Harvey, 2010 — Australia, Chile
* Nortanapis Platnick & Forster, 1989 — Australia
* Novanapis Platnick & Forster, 1989 — New Zealand
* Octanapis Platnick & Forster, 1989 — Australia
* Olgania Hickman, 1979 — Australia
* Paranapis Platnick & Forster, 1989 — New Zealand
* Patelliella Rix & Harvey, 2010 — Australia
* Pecanapis Platnick & Forster, 1989 — Chile
* Plectochetos Butler, 1932 — Australia
* Pseudanapis Simon, 1905 — Asia, Germany, South America, Africa, Mexico, Central America, Papua New Guinea
* Pua Forster, 1959 — New Zealand
* Queenslanapis Platnick & Forster, 1989 — Australia
* Raveniella Rix & Harvey, 2010 — Australia
* Rayforstia Rix & Harvey, 2010 — New Zealand, Australia
* Risdonius Hickman, 1939 — Australia
* Sheranapis Platnick & Forster, 1989 — Chile
* Sinanapis Wunderlich & Song, 1995 — China, Laos, Vietnam
* Sofanapis Platnick & Forster, 1989 — Chile
* Spinanapis Platnick & Forster, 1989 — Australia
* Taliniella Rix & Harvey, 2010 — New Zealand
* Taphiassa Simon, 1880 — Australia, New Zealand, Sri Lanka
* Tasmanapis Platnick & Forster, 1989 — Australia
* Teutoniella Brignoli, 1981 — Chile, Brazil
* Tinytrella Rix & Harvey, 2010 — New Zealand
* Tricellina Forster & Platnick, 1989 — Chile
* Victanapis Platnick & Forster, 1989 — Australia
* Zangherella Caporiacco, 1949 — Europe, Algeria, Turkey
* Zealanapis Platnick & Forster, 1989 — New Zealand
There are also genera only known as fossils, typically in Baltic amber:
* †Balticoroma Wunderlich, 2004
* †Balticonopsis Wunderlich, 2004
* †Cenotextricella Penney in Penney et al., 2007
* †Deanoorapsis Penney, 2020
* †Dubianapis Wunderlich, 2004
* †Flagellanapis Wunderlich, 2004
* †Fossilanapis Wunderlich, 2004
* †Palaeoanapis Wunderlich, 1988
* †Ruganapis Wunderlich, 2004
* †Saxonanapis Wunderlich, 2004
* †Tuberanapis Wunderlich, 2004