# Pacullidae

Pacullidae is a family of araneomorph spiders first described by Eugène Simon in 1894. It was merged into Tetrablemmidae in 1958, then raised back to family status after a large phylogenetic study in 2017.

## Description

The family Pacullidae contains three-clawed spiders with six eyes, lacking a cribellum. They resemble spiders from the family Tetrablemmidae in some respects but are much larger, always exceeding 5 mm long, have a very wrinkled (rugose) cuticle, and females do not have large membranous receptacles.

## Phylogeny

Pacullidae falls within the Synspermiata clade, a clade of former haplogyne spiders with "synsperm" – encapsulated groups of 2–4 fused sperm cells. Within this clade, it groups with four other families, including Tetrablemmidae, but is distinct from the latter, being most closely related to Diguetidae. Together with Pholcidae, these four families are placed in the "lost trachea clade", a group of families that have lost their posterior respiratory system.

## Genera and species

As of April 2019, the World Spider Catalog accepts the following genera and species:
* Lamania Lehtinen, 1981
** Lamania bernhardi (Deeleman-Reinhold, 1980) — Borneo
** Lamania bokor Schwendinger & Košulič, 2015 — Cambodia
** Lamania gracilis Schwendinger, 1989 — Bali
** Lamania inornata (Deeleman-Reinhold, 1980) — Borneo
** Lamania kraui (Shear, 1978) — Thailand, Malaysia
** Lamania lipsae Dierkens, 2011 — Borneo
** Lamania nirmala Lehtinen, 1981 — Borneo
** Lamania sheari (Brignoli, 1980) — Indonesia (Sulawesi)
* Paculla Simon, 1887
** Paculla bukittimahensis Lin & Li, 2017 — Singapore
** Paculla cameronensis Shear, 1978 — Malaysia
** Paculla globosa Lin & Li, 2017 — Singapore
** Paculla granulosa (Thorell, 1881) — New Guinea
** Paculla mului Bourne, 1981 — Borneo
** Paculla negara Shear, 1978 — Malaysia
** Paculla sulaimani Lehtinen, 1981 — Malaysia
** Paculla wanlessi Bourne, 1981 — Borneo
* Perania Thorell, 1890
** Perania annam Schwendinger & Košulič, 2015 — Vietnam
** Perania armata (Thorell, 1890) — Indonesia (Sumatra)
** Perania birmanica (Thorell, 1898) — Myanmar
** Perania cerastes Schwendinger, 1994 — Malaysia
** Perania coryne Schwendinger, 1994 — Malaysia
** Perania deelemanae Schwendinger, 2013 — Indonesia (Sumatra)
** Perania egregia Schwendinger, 2013 — Thailand
** Perania ferox Schwendinger, 2013 — Thailand
** Perania harau Schwendinger, 2013 — Indonesia (Sumatra)
** Perania korinchica Hogg, 1919 — Indonesia (Sumatra)
** Perania nasicornis Schwendinger, 1994 — Thailand
** Perania nasuta Schwendinger, 1989 — Thailand
** Perania nigra (Thorell, 1890) — Indonesia (Sumatra)
** Perania picea (Thorell, 1890) — Indonesia (Sumatra)
** Perania quadrifurcata Schwendinger, 2013 — Thailand
** Perania robusta Schwendinger, 1989 — China, Thailand
** Perania selatan Schwendinger, 2013 — Indonesia (Sumatra)
** Perania siamensis Schwendinger, 1994 — Thailand
** Perania tumida Schwendinger, 2013 — Thailand
** Perania utara Schwendinger, 2013 — Indonesia (Sumatra)
* Sabahya Deeleman-Reinhold, 1980
** Sabahya bispinosa Deeleman-Reinhold, 1980 — Borneo
** Sabahya kinabaluana Deeleman-Reinhold, 1980 — Borneo
* †Furcembolus Wunderlich 2008 9 spp, Burmese amber, Myanmar, Cenomanian