# Paleobiota of the Maotianshan Shales

This is a list of fossils found at Maotianshan Shales, whose most famous assemblage of organisms are referred to as the Chengjiang biota.
The Maotianshan Shales are a series of Early Cambrian sedimentary deposits in the Chiungchussu Formation, famous for their Konservat Lagerstätten, deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales form one of some forty Cambrian fossil locations worldwide exhibiting exquisite preservation of rarely preserved, non-mineralized soft tissue, comparable to the fossils of the Burgess Shale of British Columbia, Canada.

## Ecdysozoa

| Genus | Species | Material | Notes | Images |
| Acosmia | A. maotiania | 9 nearly complete specimens | A small ecdysozoan worm that was originally thought to have been a priapulid worm. | Fossil of Acosmia |
| Laojieella | L. thecata | One complete and one incomplete specimen | An ecdysozoan worm considered close to the priapulids, it has a spiny proboscis, an elliptical theca on the posterior part of the trunk, and a caudal appendage. | |

### Scalidophora
#### Palaeoscolecidomorpha

| Genus | Species | Notes | Images |
| Cricocosmia | C. jinningensis | A genus of palaeoscolecid worm. Fossils have been found with smaller worms seemingly attached to them. | Reconstruction of Cricocosmia with parasitic Inquicus |
| Mafangscolex | M. sinensis M. yunnanensis | | |
| Maotianshania | M. cylindrica | An extinct palaeoscolecid worm closely related to Cricocosmia. | Fossil of Maotianshania |
| Tabelliscolex | T. chengjiangensis T. hexagonus T. maanshanensis | A genus of palaeoscolecid worm | |
| Tylotites | T. petiolaris | | |

#### Priapulida

| Genus | Species | Material | Notes | Images |
| Corynetis | C. brevis C. fortis | | An archaeopriapulid worm known from the biota. | Reconstruction of Corynetis |
| Ercaivermis | E. sparios | A single specimen. | A stem-group priapulid. | |
| Eximipriapulus | E. globocaudatus | | A carnivorous priapulid-like worm that has been found living in the discarded shells of hyoliths. | Reconstruction of Eximipriapulus |
| Paraselkirkia | P. sinica | | An archaeopriapulid worm that resembles another worm, Selkirkia, from the Burgess shale | Reconstruction of Paraselkirkia |
| Paratubiluchus | P. bicaudatus | Two specimens | | Reconstruction of Paratubiluchus |
| Palaeopriapulites | P. parvus | | | Reconstruction of Palaeopriapulites |
| Selkirkia | S. elongata | | A burrowing archaeopriapulid worm also known from the burgess shale, alongside the Fezouata Shale. | Fossil of the relatedSelkirkia Columbia |
| Sicyophorus | S. rarus | | A genus of archaeopriapulid worm. | Reconstruction of Sicyophorus |
| Xiaoheiqingella | X. peculiaris | Fourteen specimens, five previously included in “Yunnanpriapulus” | A priapulid worm thought to belong to the crown group. | Reconstruction of Xiaoheiqingella |

### Panarthropoda
#### Lobopodia

| Genus | Species | Material | Notes | Images |
| Antennacanthopodia | A. gracilis | Known from two specimens, making the species very rare in the Chengjiang. | An unarmoured lobopodian, apparently closely related to Onychophora. The long, paired, cirriform structures at the end of the body could have been sensory, or perhaps represent viscera not externally present in life. | Reconstruction of Antennacanthopodia |
| Cardiodictyon | C. catenulum | | A highly elongate armoured lobopodian, with saddle-shaped sclerites and a lightly sclerotized head. | Reconstruction of Cardiodictyon |
| Diania | D. cactiformis | | A heavily armoured lobopodian. | Reconstruction of Diania |
| Lenisambulatrix | L. humboldti | A single specimen. | A simple, unarmoured lobopodian. | Reconstruction of Lenisambulatrix |
| Onychodictyon | O. ferox O. gracilis | Several complete specimens. | An enigmatic genus of lobopodian, potentially related to either the Luolishaniidae or to Tardigrada. Not always monophyletic. | Reconstruction of Onychodictyon ferox |
| Paucipodia | P. inermis | Known from well-preserved body fossils. | A large unarmoured lobopodian, commonly found in association with mass death-beds of legless worms and Eldonia. Thought to have been a scavenger. | Reconstruction of Paucipodia |

##### Eoconchariidae

| Genus | Species | Material | Notes | Images |
| Microdictyon | M. sinicum | Roughly 100 specimens | Microdictyon is known from sclerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy. | Fossil of Microdictyon sinicum |

##### Hallucishaniids

| Genus | Species | Material | Notes | Images |
| Luolishania | L. longicruris | Around forty specimens | Type genus of Luolishaniidae. The species Miraluolishania haikouensis is considered a juvenile synonym of Luolishania. | Reconstruction of Luolishania |
| Facivermis | F. yunnanicus | Thirty specimens, one formerly assigned to “Xishania” | Derived Luolishaniid lobopodian with five setiferous sieving appendages, and lacking any walking limbs. Sometimes discovered living in constructed tubes, buried into the sediment. | Reconstruction of Facivermis |
| Hallucigenia | H. fortis | Two specimens | A hallucigeniid lobopodian with a bulbous sclerotized head and two pairs of small feeding appendages, borne on a short neck. | Reconstruction of Hallucigenia fortis |

##### Siberiidae

| Genus | Species | Material | Notes | Images |
| Megadictyon | M. haikouensis | Posterior-most section of the trunk is unknown. | A poorly known siberiid. | Reconstruction of Megadictyon cf. haikouensis |
| Jianshanopodia | J. decorata | Known from multiple large individuals, all from a singular exposure of the Chengjiang. | A giant siberiid lobopodian related to Megadictyon. | Reconstruction of Jianshanopodia |

##### Gilled Lobopodians

| Genus | Species | Material | Notes | Images |
| Omnidens | O. amplus O. qiongqii | Numerous well preserved oral cones and portions of the frontal appendage. | A giant gilled lobopodian, closely related to Pambdelurion. Also present in the Xiaoshiba Biota. | Diagrammatic reconstruction of Omnidens |

#### Radiodonta

Radiodonta are a group of highly successful panarthropods. The Chengjiang Fauna includes a large number of Radiodont species, primarily from the clade Amplectobeluidae. While hurdiids are poorly known from the formation and surrounding region, several specimens attributable to the family have been discovered, but remain unnamed. Some radiodonts from the Chengjiang defy easy classification, or are otherwise debated in their exact affinity.
| Genus | Species | Material | Notes | Images |
| Houcaris | H. saron H. consimilis | Isolated appendages. | Previously described as "Anomalocaris" saron. | Appendage of Houcaris saron |
| Innovatiocaris | I. maotianshanensis I. multispiniformis I. sp. | Complete fossil and isolated appendages | Previously assigned to the Anomalocaris genus, and one of the most completely known radiodonts. | Reconstruction of I. maotianshanensis |
| Laminacaris | L. chimera | Giant isolated appendages, up to nearly 30 centimetres long. | A giant radiodont combining features from different clades. | Appendage of Laminacaris |

##### Amplectobeluidae

| Genus | Species | Material | Notes | Images |
| Amplectobelua | A. trispinata | Dozens of isolated appendages, and at least one complete body of a juvenile. | A large Amplectobeluid radiodont with long tail furcae. The frontal appendages formed a distinctive claw-like shape. | Reconstruction of Amplectobelua |
| Lyrarapax | L. trilobus L. unguispinus | Known from appendages and complete body fossils, including neural anatomy. | Small Amplectobeluid radiodont. Some specimens of this genus represent some of the smallest known radiodonts. | Reconstruction of Lyrarapax unguispinus (left) and Lyrarapax trilobus (right) |
| Ramskoeldia | R. platyacantha R. consimilis? | Known from appendages, often associated with GLS plates, sclerites, and swimming flaps. | Large Amplectobeluid radiodonts, with crushing gnathobases. The species R. consimilis may belong to Houcaris instead. | Appendages of Ramskoeldia |

##### Anomalocarididae

| Genus | Species | Material | Notes | Images |
| Anomalocaris | A. cf. canadensis | Isolated appendages. | | Appendage of A. cf. canadensis |
| Lenisicaris | L. lupata | Isolated appendages. | A second species is included in the genus, Lenisicaris pennsylvanica, from the Kinzers Formation. | Appendage of Lenisicaris |
| Shucaris | S. ankylosskelos | Isolated appendages. Oral cone Partial body | An unusual radiodont seemingly transitional between anomalocaridids and amplectobeluids, with an oral cone similar to the former but gnathobase-like structures similar to the latter. | Appendage model of Shucaris |

##### Hurdiidae (Peytoiidae)

Hurdiids (peytoiids) have historically been considered absent, or at least rare, from the Chengjiang Biota, an observation that researchers have found confusing given their regularity at other sites. In recent years, a number of fragmentary hurdiid fossils have been described, including 3 partial appendages belonging to a new species of Stanleycaris, an isolated sclerite assigned to Cambroraster, Zhenghecaris (the affinities of which, even its identity as a radiodont, remains questionable), and isolated lateral scelrites (also questionable).
| Genus | Species | Material | Notes | Images |
| Cambroraster | C. sp. | Head sclerites. | A large nektobenthic hurdiid, first known from the Burgess Shale. Fossils of Cambroraster are also known from Mantou Formation of North China, making this a very widespread genus. This genus was the first definitive peytoiid known from China. | Fossil of Cambroraster from the Burgess Shale. |
| Zhenghecaris | Z. shankouensis | Sclerites | Large-sized arthropod carapace, originally described as bivalved arthropod close to Thylacocephala, later reinterpreted as hurdiid radiodont close to Cambroraster, its classification is still discussed. Some fossils, apparently the lateral scelrites of a radiodont, connected anteriorly by a "beak" and featuring two-pronged, wing-like processes posteriorly (similar to the lateral processes of Zhenghecaris's central head sclerite), have also been attributed to Zhenghecaris. | Fossil of Zhenghecaris |

#### Arthropoda

| Genus | Species | Material | Notes | Images |
| Chuandianella | C. ovata | Several body fossils and carapaces | An enigmatc bivalved arthropod originally considered close to Waptia. It was then discovered that it lacked mandibles and therefore does not belong to Hymenocarina, the Waptiidae family, and even the larger mandibulata clade. | Reconstruction of Chuandianella |
| Combinivalvula | C. chengjiangensis | | An enigmatic bivalved arthropod. | |
| Cucumericrus | C. decorata | Multiple fragmentary specimens. | An enigmatic arthropod that has been tentatively classified as a possible radiodont, although a position closely related to Erratus as the most primitive deuteropod is more likely. | Reconstructed trunk appendage of Cucumericrus |
| Cyathocephalus | C. bispinosus | | | |
| Diplopyge | D. forcipatus D. minutus | | | |
| Dongshanocaris | D. foliiformis | | A small arthropod that is currently "too poorly preserved to verify their identity as a valid taxa". | |
| Erratus | E. sperare | Several carapaces Eyes and other body parts | A bivalved arthropod currently placed as the most basal deuteropod, that shows the origins of lungs and legs in early arthropods. | Reconstruction of Erratus |
| Ercaia | E. miniscula | | An arthropod that has been speculated to be an early crustacean. | |
| Forfexicaris | F. valida | | A small bivalved arthropod that somewhat resembles ostracods, but also has large, upward facing appendages with spines. | Reconstruction of Forfexicaris |
| Glossocaris | G. occulatus | | | |
| Isoxys | I. auritus I. curvirostratus I. paradoxus | | A basal nektonic arthropod that possessed a bivalved carapace, and large circular eyes, related to Surusicaris from the Burgess Shale. Also found in various other Cambrian aged sites. | Reconstruction of Isoxys curvirostratus |
| Jianshania | J. furcatus | | A small, somewhat enigmatic arthropod known from the biota | Fossil of Jianshania |
| Kunmingocaris | K. bispinosus | | | |
| Kylinxia | K. zhangi | Almost complete fossil Isolated frontal appendages | An early arthropod that may show the "missing link" between early stem-arthropods, and more advanced true arthropods. It possessed frontal appendages similar to radiodonts. | Reconstruction of Kylinxia |
| Mafangia | M. subscalaria | | | |
| Mafangocaris | M. multinodus | | | |
| Occacaris | O. oviformis | | A predatory arthropod that had spiked frontal appendages similar to those of the megacheirans, and was previously thought to have belonged to that grouping | Reconstruction of Occacaris |
| Ovalicephalis | O. mirabilis | | | |
| Parapaleomerus | P. sinensis | | An arachnomorph arthropod of the group strabopida. Unlike the other members of strabopida, Parapaleomerus lacks dorsal eyes and only possesses ten trunk tergites. The telson has been described as trapezoidal in shape. | Reconstructed carapace of Parapaleomerus |
| Pisinnocaris | P. subconigera | | | |
| Pseudoiulia | P. cambriensis | | A poorly known arthropod that, in 2013, was suggested to be a member of the family Kootenichelidae, alongside Kootenichela and Worthenella. | |
| Pterotum | P. triacanthus | | | |
| Rhombicalvaria | R. acanthi | | | |
| Sunella | S. grandis | | | |
| Syrrhaptis | S. intes | | | |
| Tanglangia | T. caudata | | | |
| Tauricornicaris | T. latizonae T. oxygonae | | A giant arthropod once misidentified as a peytoiid radiodont. | |
| Wutingella | W. bindosa | | | |
| Yiliangocaris | Y. ellipticus | | | |
| Youti | Y. yuanshi | | | Fossil of Youti |
| Yunnanocaris | Y. megista | | | |

##### Hymenocarina

| Genus | Species | Material | Notes | Images |
| Branchiocaris | B. yunnanensis | Several isolated carapaces | Also known in the Burgess Shale by B. pretiosa, B. yunnanensis is only known from isolated carapaces, although it is thought that this may be because in this species the carapace covered the entire body. It also has two morphotypes. | Reconstruction of the related Branchiocaris pretiosa |
| Canadaspis | C. laevigata | | A hymenocarine arthropod, also known from the Burgess Shale by C. perfecta, although some authors believe that both species should be separated into different genera. They were mainly benthic animals. | Reconstruction of Canadaspis laevigata |
| Clypecaris | C. pteroidea C. serrata | | A hymenocarine arthropod. The species in the genus are primarily distinguished by the presence of a serrated edge on the front of the carapace of C. serrata. C. serrata is noted for the modification of an anterior pair of limbs into spined grasping appendages, indicating a predatory lifestyle. | Reconstruction of Clypecaris serrata |
| Ercaicunia | E. multinodosa | | A hymenocarine arthropod that had a bivalved carapace which covered about a third of its total body-length, and had up to six serrations on its forward edge. The head had a pair of large uniramous antennae, as well as a smaller pair of secondary antennae, and a pair of mandibles and maxillae. | Reconstruction of Ercaicunia |
| Erjiecaris | E. minusculo | | A small euarthropod, and unlike other hymenocarines, it possibly had eyes directly on top of its carapace. It was most likely a nektobenthic animal. | Reconstruction of Erjiecaris |
| Pectocaris | P. spatiosa P. eurypetala P. inopinata | | A bivalved arthropod of the hymenocarina grouping. The species P. spatiosa is regarded as a filter feeder, using the setae on the endites of their limbs to filter out matter from the water column | Reconstruction of Pectocaris spatiosa |
| Synophalos | S. xynos | | A hymenocarine notable for fossils showing a bizarre chain of multiple individuals. Although the purpose for this behavior is unknown, it may have served a reproductive, migratory, or defensive purpose, with the authors of the describing paper of this genus considering migration as its most likely function. | Reconstruction of a Synophalos chain |
| Tuzoia | T. sinensis | several carapaces and soft body parts | A large hymenocarine that possessed a semicircular bivalved carapace, and was one of the largest arthropods from the Cambrian | Reconstruction of Tuzoia sp. |
| Jugatacaris | A. agillis | several carapaces and soft body parts | A hymenocarine arthropod. The carapace of the animal was around 28 to 37 millimetres (1.1 to 1.5 in) in length, with a pronounced ridge at the top of the carapace separating the two valves, which formed a fin-like structure raised above the carapace. The head had a pair of stalked eyes, as well as a dumbbell shaped medial eye between them. The head also bore a pair of mandibles as well as at least one and possibly two pairs of antennules. | Reconstruction of Jugatacaris |
| Xiazhuangocaris | X. chenggongensis | a single specimen (NIGP 172765), which only preserves the carapace and trunk regions | A hymenocarine with no known head fossils. The material known show that the carapace had a pronounced pair of notches at its front, as well as a posterior notch at its rear. The body had at least 13 tergite-pleurite rings, which terminate in a pair of rounded caudal rami, which are fringed with setae. | Reconstruction of Xiazhuangocaris |

##### Fuxianhuiida

| Genus | Species | Notes | Images |
| Chengjiangocaris | C. longiformis C. kunmingensis | A Fuxianhuiid arthropod with numerous appendages. It is one of the most complete arthropods from the lower Cambrian. | Reconstruction of Chengjiangocaris |
| Fuxianhuia | F. protensa | A large arthropod named after Fuxian Lake. Some adult specimens are found closely associated with numerous juveniles, indicating a level of parental care. | Reconstruction of Fuxianhuia with associated juveniles |
| Liangwangshania | L. biloba | A roughly 7 cm long fuxianhuiid, with which Shankouia is potentially synonymous through sexual dimorphism. | |

##### Bradoriida

| Genus | Species | Images |
| Comptaluta | C. inflata C. kailiensis | |
| Indiana | I. sp. | CT-scan of a fossil of Indiana sp. |
| Jiucunella | J. paulula | |
| Kunmingella | K. angustacostata K. douvillei K. guanshanensis | Reconstruction of Kunmingella douvillei |
| Kunyangella | K. cheni | |
| Liangshanella | L. liangshanensis | |
| Malongella | M. bituerculata | |
| Parakunmingella | P. malongensis | |
| Spinokunmingella | S. typica | |
| Tsunyiella | T. daindongensis | |

##### Megacheira

| Genus | Species | Material | Notes | Images |
| Haikoucaris | H. ercaiensis | | | Reconstruction of Haikoucaris |

###### Leanchoiliidae

| Genus | Species | Material | Notes | Images |
| Alalcomenaeus | A. cambricus | | A small arthropod related to Leanchoilia, also known from the Burgess Shale. | Reconstruction of Alalcomenaeus |
| Leanchoilia | D. asiatica L. illecebrosa | | One of the several species of Leanchoilia, it has known juvenile specimens and the labrum morphology known. | Fossil of Leanchoilia illecebrosa |

###### Jianfengiidae

| Genus | Species | Notes | Images |
| Fortiforceps | F. foliosa | | Reconstruction of Fortiforceps |
| Sklerolibyon | *S. maomima | | Reconstruction of Sklerolibyon |
| Jianfengia | J. multisegmentalis | | Reconstruction of Jianfengia |
| Parapeytoia | P. yunnanensis | A jianfengiid once misidentified as a radiodont | Reconstruction of known elements from Parapeytoia |

##### Marrellomorpha

| Genus | Species | Material | Notes | Images |
| Primicaris | P. larvaformis | | | Reconstruction of Primicaris |

##### Artiopoda

| Genus | Species | Material | Notes | Images |
| Acanthomeridion | A. serratum | Multiple Specimens. | | Reconstruction of a fully grown Acanthomeridion |
| Cindarella | C. eucalla | | | Reconstruction of Cindarella |
| Kuamaia | K. lata K. muricata | | | Reconstruction of Kuamaia lata |
| Panlongia | P. spinosa P. tetranudosa | | | Fossil of Panlognia |
| Pygmaclypeatus | P. daiensis | | | CT-scan of a specimen of Pygmaclypeatus |
| Retifacies | R. abnormalis | | | Reconstruction of Retifacies |
| Saperion | S. glumaceum | | | |
| Sidneyia | S. malongensis S. minor S. sinica | | | Reconstruction of Sidneyia minor |
| Sinoburius | S. lunaris | | | Reconstruction of Sinoburius lunaris |
| Skioldia | S. aldna | | | |
| Squamacula | S. clypeata | | | Reconstruction of Squamacula |
| Tonglaiia | T. bispinosa | Single specimen. | | Holotype of Tonglaiia bispinosa |
| Urokodia | U. aequalis | | An arthropod that had segmentation which resembled that of a millipede, as well as head and tail shields with thorny spikes. Recently, it was recovered as the most basal artiopod. | Reconstruction of Urokodia |
| Xandarella | X. spectaculum | | | Reconstruction of Xandarella |
| Zhugeia | Z. acuticaudata | | | Holotype of Zhugeia acuticaudata |

###### Nektaspida

| Genus | Species | Images |
| Misszhouia | M. longicaudata | Reconstruction of Misszhouia |
| Naraoia | N. spinosa | Reconstruction of Naraoia spinosa |
| Kangacaris | K. shui | Reconstruction of Kangacaris zhangi, the type species from Cambrian Australia |

##### Trilobita

| Genus | Species | Images |
| Eoredlichia | E. intermedia | Reconstruction of Eoredlichia |
| Kuanyangia | K. pustulosa | |
| Palaeolenus | E. miniscula | Fossil pair of Palaeolenus |
| Tsunyidiscus | T. aclis | Line drawing of Tsunyidiscus niutitangensis |
| Wutingapsis | W. tingi | |
| Yunnanocephalus | Y. yunnanensis | Fossil of Yunnanocephalus |

## Spiralia
### Clade Gnathifera

| Genus | Species | Material | Notes | Images |
| Amiskwia | A. sinica | | | |
| Inquicus | I. fellatus | | | Reconstruction of Inquicus |

#### Chaetognatha

| Genus | Species | Images |
| Ankalodous | A. ercainella | |
| Eognathacantha | E. ercainella | Reconstruction of Eognathacantha |
| Protosagitta | P. spinosa | Reconstruction of Protosagitta |

### Lophotrochozoa
#### Annelida

| Genus | Species | Material | Notes | Images |
| Iotuba | I. chengjiangensis | | | Fossil of Iotuba |

#### Mollusca

| Genus | Species | Material | Notes | Images |
| Orthrozanclus | O. elongata | | | Reconstruction of Orthrozanclus elongata |
| Wiwaxia | W. papillo | | | Reconstruction of Wiwaxia papilio |

#### Hyolitha

| Genus | Species |
| Ambrolinevitus | A. maximus A. platypluteus A. ventricosus |
| Burithes | B. yunnanensis |
| Glossolithes | G. magnus |
| Linevitus | L. billingsi L. flabellaris L. opimus |

#### Phoronida

| Genus | Species | Material | Notes | Images |
| Eophonoris | E. chengjiangensis | | | |

#### Entoprocta

| Genus | Species | Material | Notes | Images |
| Cotyledion | C. tylodes | | | Reconstruction of Cotyledion |

#### Cornulitida

| Genus | Species | Material | Notes | Images |
| Cambrocornulitus | C. rarus | | | |

#### Brachiopoda

| Genus | Species | Images |
| Diandongia | D. pista | |
| Heliomedusa | H. orienta | Fossil of Heliomedusa orienta |
| Lingulella | L. orienta | Fossil of the related Lingulella caelata |
| Lingulellotreta | L. malongensis | |
| Longtancunella | L. chengjiangensis | |
| Yuganotheca | Y. elegans | |

## Deuterostomia

| Genus | Species | Material | Notes | Images |
| Yunnanozoon | Y. lividium | | A species of deuterostome animal with controversial taxonomy, with some authors placing it as an chordate, stem-vertebrate, hemichordate, etc. | Fossil of Yunnanozoon |

### Chordata

| Genus | Species | Notes | Images |
| Cathaymyrus | C. diadexus C. haikouensis | A chordate known from the Yunnan Province with potential relations to cephalochordates like lancelets. | Reconstruction of C. diadexus and Vetulocystis |
| Haikouichthys | H. ercaicunensis | An early craniate (an animal with a distinct notochord and head) of the Myllokunmingiida order. Some authors consider it potentially synonymous with the contemporary Myllokunmingia. | Reconstruction of Haikouichthys |
| Myllokunmingia | M. fengjiaoa | A myllokunmingiid known for its wide body plan, sail-like fin on its dorsal plane, and a non-biomineralized skeleton. | Reconstruction of Myllokunmingia |
| Shankouclava | S. anningense S. shankouense | Likely one of the earliest known tunicates, alongside Cheungkongella and possibly “Palaeoikopleuria”. | Reconstruction of Shankouclava anningense |
| Zhongjianichthys | Z. rostratus | Another myllokunmingiid known for lacking a dorsal fin, and having a more slim body compared to its two contemporary relatives. | Reconstruction of Zhongjianichthys |
| Zhongxiniscus | Z. intermedius | An early chordate thought to represent a transitional form between earlier chordates, and more derived craniates. | Reconstruction of Zhongxiniscus |

#### Vetulicolia

| Genus | Species | Images |
| Beidazoon | B. venustum | Reconstruction of Beidazoon |
| Didazoon | D. haoae | Reconstruction of Beidazoon |
| Pomatrum | P. ventralis | Reconstruction of Beidazoon |
| Vetulicola | V. cuneata V. gantoucunensis V. rectangulata | Reconstruction of Vetulicola rectangulata |
| Yuyuanozoon | Y. magnificissimi | Reconstruction of Yuyuanozoon |

### Ambulacraria
#### Cambroernida

| Genus | Species | Notes | Images |
| Phlogites | P. brevis P. longus | Senior synonym of Cheungkongella ancestralis. | Fossil of Phlogites |
| Rotadiscus | R. grandis | | Reconstruction of Rotadiscus |
| Stellostomites | S. eumorphus | Senior synonym of Eldonia eumorpha. | Diagram of Stellostomites |

#### Echinodermata

| Genus | Species | Material | Notes | Images |
| Dianchicystis | D. jianshanensis | | | |
| Vetulocystis | V. catenata | | | Reconstruction of Vetulocystis (bottom) |

### Phylum Hemichordata

| Genus | Species |
| Cambrobranchus | C. pelagobenthos |
| Galeaplumosus | G. abilus |
| Yuknessia | Y. simplex |

### Phylum Ctenophora

| Genus | Species | Images |
| Maotianoascus | M. octanarius | Reconstruction of Maotianoascus octanarius |
| Sinoascus | S. paillatus | Reconstruction of Sinoascus papillatus |
| Stromatoveris | S. psygmoglena | |
| Trigoides | T.aclis |

### Phylum Porifera

| Genus | Species | Images |
| Allantospongia | A. mica | |
| Choia | C. xiaolantianensis | |
| Choiaella | C. radiata | |
| Hazelia | H. | |
| Leptomitella | L. confusa L. conica L. metta | |
| Leptomitus | L. teretiusculus | |
| Paraleptomitella | Paraleptomitella dictyodroma Paraleptomitella globula |
| Quadrolaminiella | Quadrolaminiella crassa Quadrolaminiella diagonalis | |
| Saetaspongia | S. densa | |
| Sinfoflabrum | S. antiquum | Fossil of Sinfoflabrum |
| Triticispongia | T. diagonata |

### Phylum Cnidaria

| Genus | Species | Images |
| Archisaccophyllia | A. kunmingensis | Fossil of Archisaccophyllia |
| Nailiana | N. elegans | |
| Priscapennamarina | P. angusta | |
| Xianguangia | X. sinica | Fossil of Xianguangia |

## Enigmatic

| Genus | Species | Notes | Images |
| Allonnia | A. phrixothrix | | Reconstruction of Allonnia sp. |
| Anthrotum | A. robustus | | |
| Batofasciculus | B. ramificans | | |
| Calathites | C. spinalis | | |
| Cambrotentacus | C. sanwuia | | Reconstruction of a pair of Cambrotentacus |
| Conicula | C. straita | | |
| Daihua | D. sanqiong | | Fossil of Daihua |
| Dinomischus | D. venustus | | Fossil of Dinomischus venustus |
| Discoides | D. abnormalis | | |
| Hippotrum | H. spinatus | | |
| Jiucunia | J. petalina | | |
| Maanshania | M. crusticeps | | |
| Macrocephalus | M. elongates | | |
| Petalilium | P. latus | Possible synonym of Nectocaris pteryx | Fossil of Petalilium |
| Parvulonoda | P. dubia | | |
| Phacatrum | P. tubifer | | |
| Phasangula | P. striata | | |
| Pristitoites | P. bifarius | | |
| Rhipitrus | R. calvifer | | |

## Miscellaneous macroalgae

Macroalgae rank fourth in species-level diversity behind arthropods, priapulids, and sponges, and account for 71.5% of total abundance from the biota, particularly dominated by unattached (floating, planktonic, and drifting) species.
| Genus | Species | Material | Notes | Images |
| Enteromophites | E. intestinalis | | Sessile, attached to the substrate. | |
| Fuxianospira | F. gyrata | The most common species of macroalgae present, accounting for 52% of all macroalgae fossils reported from the Chengjiang. | Unattached (planktonic, drifting, floating) | |
| Liulingjitaenia | L. alloplecta | | Unattached (planktonic, drifting, floating) | |
| Longfengshania | L. stipitata L. spheria L. cordata | | Sessile, attached to the substrate. | |
| Morania | M. fragmenta | | Unattached (planktonic, drifting, floating) | Fossil of Morania fragment from the Cambrian Wheeler Shale of Utah, USA. |
| Paralongfengshania | P. sicyoides | | Sessile, attached to the substrate. | |
| Paradelesseria | P. sanguinea | | Unattached (planktonic, drifting, floating) | |
| Plantulaformis | P. sinensis | | Sessile, attached to the substrate. | |
| Punctariopsis | P. latifolia P. simplexis | | Sessile, attached to the substrate. | |
| Sinocylindra | S. yunnanensis S. linearis | | Unattached (planktonic, drifting, floating) | Fossil of Sinocylindra yunnanensis |
| Tawuia | T. sp. | | Unattached (planktonic, drifting, floating) | Fossil of Tawuia sp. |
| Yunnanospirellus | Y. typica Y. elegans | | Unattached (planktonic, drifting, floating) | |