# Portunidae

Portunidae is a family of crabs which contains the swimming crabs. Its members include well-known shoreline crabs such as the blue crab (Callinectes sapidus) and velvet crab (Necora puber).

## Description

Portunid crabs are characterised by the flattening of the fifth pair of legs into broad paddles, which are used for swimming. This ability, together with their strong, sharp claws, allows many species to be fast and aggressive predators.

## Taxonomy

Swimming crabs reach their greatest species diversity in the Pacific and Indian Oceans. Portunidae consists of the following subfamilies and genera:

### Achelouinae Spiridonov, 2020

* Achelous De Haan, 1833

### Caphyrinae Guérin, 1832

* Caphyra Guérin, 1832
* Coelocarcinus Edmondson, 1930
* Lissocarcinus Adams & White, 1849
* †Mioxaiva Müller, 1978

### Carupinae Paulson, 1875

* Carupa Dana, 1851
* Catoptrus A. Milne-Edwards, 1870
* †Euronectes Karasawa, Schweitzer & Feldmann, 2008
* Kume Naruse & Ng, 2012
* Laleonectes Manning & Chace, 1990
* Libystes A. Milne-Edwards, 1867
* Pele Ng, 2011
* †Rakosia Müller, 1984
* Richerellus Crosnier, 2003

### Coelocarcininae Števćić, 1991

* Coelocarcinus Edmondson, 1930

### Lupocyclinae Alcock, 1899

* Lupocycloporus Alcock, 1899
* Lupocyclus Adams & White, 1849

### Necronectinae Glaessner, 1928

* †Necronectes A. Milne-Edwards, 1881
* Scylla De Haan, 1833

### Podophthalminae Dana, 1851

* Euphylax Stimpson, 1860
* †Paraeuphylax Varela & Schweitzer, 2011
* †Phenophthalmus Feldmann, Schweitzer & Encinas, 2010
* Podophthalmus Lamarck, 1801
* †Psygmophthalmus Schweitzer, Iturralde-Vinent, Hetler & Velez-Juarbe, 2006
* †Sandomingia Rathbun, 1919
* †Saratunus Collins, Lee & Noad, 2003
* †Viaophthalmus Karasawa, Schweitzer & Feldmann, 2008
* Vojmirophthalmus Števčić, 2011

### Portuninae Rafinesque, 1815

* †Acanthoportunus Schweitzer & Feldmann, 2002
* Alionectes Koch, Spiridonov & Ďuriš, 2022
* Allomonomia Koch, Spiridonov & Ďuriš, 2022
* †Archaeoportunus Artal, Ossó & Domínguez, 2021
* Arenaeus Dana, 1851
* Atoportunus Ng & Takeda, 2003
* Callinectes Stimpson, 1860
* Carupella Lenz in Lenz & Strunck, 1914
* Cavoportunus Nguyen & Ng, 2010
* †Colneptunus Lőrenthey in Lőrenthey & Beurlen, 1929
* Cycloachelous Ward, 1942
* Eodemus Koch, Spiridonov & Ďuriš, 2022
* Incultus Koch, Spiridonov & Ďuriš, 2022
* Monomia Gistel, 1848
* †Persianus Garassino, Pasini, Mirzaie Attabadi & Nyborg in Garassino, Pasini, Mirzaie Ataabadi, Hadi, Parsazad, Nyborg & Vega, 2024
* Portunus Weber, 1795
* †Rathbunites Schweitzer, Dworschak & Martin, 2011
* Sanquerus Manning, 1989
* Trionectes Koch, Spiridonov & Ďuriš, 2022
* Xiphonectes A. Milne-Edwards, 1873

### Thalamitinae Paulson, 1875

* Caphyra Guérin, 1832
* Charybdis De Haan, 1833
* Cronius Stimpson, 1860
* †Eocharybdis Beschin, Busulini, De Angeli & Tessier, 2002
* Gonioinfradens Leene, 1938
* Goniosupradens Davie, 2002
* †Lessinithalamita De Angeli & Ceccon, 2015
* Lissocarcinus Adams & White, 1849
* †Mioxaiva Müller, 1979
* Thalamita Latreille, 1829
* Thalamitoides A. Milne-Edwards, 1869
* Zygita Evans, 2018

### incertae sedis

* †Neptocarcinus Lörenthey, 1897
* †Pseudoachelous Portell & Collins, 2004