# Alpheus (crustacean)

Alpheus is a genus of snapping shrimp of the family Alpheidae. This genus contains in excess of 330 species, making this the most species-rich genus of shrimp. Like other snapping shrimp, the claws of Alpheus are asymmetrical, with one of the claws enlarged for making a popping noise. Some species in the genus enter into symbiotic relationships with gobiid fishes, and others associate with sea anemones.

## Species

Alpheus contains the following species:
* Alpheus abumusa Dehghani, Sari & Naderloo, 2019
* Alpheus acutocarinatus De Man, 1909
* Alpheus acutofemoratus Dana, 1852
* Alpheus adamastor Coutière, 1908
* Alpheus aequus Kim & Abele, 1988
* Alpheus africanus Balss, 1916
* Alpheus agilis Anker, Hurt & Knowlton, 2009
* Alpheus agrogon Ramos, 1997
* Alpheus alaincrosnieri Anker, 2020
* Alpheus albatrossae (AH Banner, 1953)
* Alpheus alcyone De Man, 1902
* Alpheus alpheopsides Coutière, 1905
* Alpheus amarillo Anker, 2012
* Alpheus amblyonyx Chace, 1972
* Alpheus amirantei Coutière, 1908
* Alpheus anchistus De Man, 1920
* Alpheus angulosus McClure, 2002
* Alpheus angustilineatus Nomura & Anker, 2005
* Alpheus ankeri Dehghani, Sari & Naderloo, 2019
* Alpheus antepaenultimus Kim & Abele, 1988
* Alpheus architectus De Man, 1897
* Alpheus arenensis (Chace, 1937)
* Alpheus arenicolus Banner & Banner, 1983
* Alpheus arethusa De Man, 1909
* Alpheus armatus Rathbun, 1901
* Alpheus armillatus H. Milne Edwards, 1837
* Alpheus arnoa Banner, 1957
* Alpheus astrinx Banner & Banner, 1982
* Alpheus australiensis Banner & Banner, 1982
* Alpheus australosulcatus Banner & Banner, 1982
* Alpheus baccheti Anker, 2010
* Alpheus bahamensis Rankin, 1898
* Alpheus balaenodigitus Banner & Banner, 1982
* Alpheus bannerorum Bruce, 1987
* Alpheus barbadensis (Schmitt, 1924)
* Alpheus barbatus Coutière, 1897
* Alpheus batesi Banner & Banner, 1964
* Alpheus bellimanus Lockington, 1877
* Alpheus bellulus Miya & Miyake, 1969
* Alpheus berumeni Anker, 2024
* Alpheus bicostatus De Man, 1908
* Alpheus bidens (Olivier, 1811)
* Alpheus bisincisus De Haan, 1849
* Alpheus blachei Crosnier & Forest, 1965
* Alpheus bouvieri A. Milne-Edwards, 1878
* Alpheus brachymerus (AH Banner, 1953)
* Alpheus bradypus Coutière, 1905
* Alpheus brasileiro Anker, 2012
* Alpheus brevicristatus De Haan, 1844
* Alpheus brevipes Stimpson, 1860
* Alpheus brevirostris (Olivier, 1811)
* Alpheus brucei Banner & Banner, 1982
* Alpheus bucephaloides Nobili, 1905
* Alpheus bucephalus Coutière, 1905
* Alpheus buchanorum Banner & Banner, 1983
* Alpheus buckupi Almeida, Terossi, Araújo-Silva & Mantelatto, 2013
* Alpheus bunburius Banner & Banner, 1982
* Alpheus burukovskyi Anker & Pachelle, 2015
* Alpheus caigeri Anker, 2024
* Alpheus californiensis Holmes, 1900
* Alpheus canaliculatus Banner & Banner, 1968
* Alpheus candei Guérin-Méneville, 1855
* Alpheus cangiopalus Marin & Thinh, 2024
* Alpheus carlae Anker, 2012
* Alpheus cedrici Anker & De Grave, 2012
* Alpheus chacei Carvacho, 1979
* Alpheus chamorro Banner, 1956
* Alpheus chilensis Coutière in Lenz, 1902
* Alpheus chiragricus H. Milne Edwards, 1837
* Alpheus christofferseni Anker, Hurt & Knowlton, 2007
* Alpheus clamator Lockington, 1877
* Alpheus clypeatus Coutière, 1905
* Alpheus coetivensis Coutière, 1908
* Alpheus collumianus Stimpson, 1860
* Alpheus colombiensis Wicksten, 1988
* Alpheus compressus Banner & Banner, 1981
* Alpheus confusus Carvacho, 1989
* Alpheus coralvivo Santos, Terossi, Mantelatto, Torres & Almeida, 2024
* Alpheus coutierei De Man, 1909
* Alpheus crinitus Dana, 1852
* Alpheus cristatus Coutière, 1897
* Alpheus cristulifrons Rathbun, 1900
* Alpheus crockeri (Armstrong, 1941)
* Alpheus cyanoteles Yeo & Ng, 1996
* Alpheus cylindricus Kingsley, 1878
* Alpheus cythereus Banner & Banner, 1966
* Alpheus dasycheles Coutière, 1908
* Alpheus davaoensis Chace, 1988
* Alpheus dentipes Guérin, 1832
* Alpheus deuteropus Hilgendorf, 1879
* Alpheus diadema Dana, 1852
* Alpheus digitalis De Haan, 1844
* Alpheus dingabadi Anker, 2023
* Alpheus dispar Randall, 1840
* Alpheus distinctus Kim & Abele, 1988
* Alpheus djeddensis Coutière, 1897
* Alpheus djiboutensis De Man, 1909
* Alpheus dolerus Banner, 1956
* Alpheus echiurophilus Anker, Komai & Marin, 2015
* Alpheus edamensis De Man, 1888
* Alpheus edwardsii (Audouin, 1826)
* Alpheus ehlersii De Man, 1909
* Alpheus estuariensis Christoffersen, 1984
* Alpheus euchirus Dana, 1852
* Alpheus eulimene De Man, 1909
* Alpheus euphrosyne De Man, 1897
* Alpheus eurydactylus De Man, 1920
* Alpheus exilis Kim & Abele, 1988
* Alpheus explorator Boone, 1935
* Alpheus facetus De Man, 1908
* Alpheus fagei Crosnier & Forest, 1965
* Alpheus fasciatus Lockington, 1878
* Alpheus fasqueli Anker, 2001
* Alpheus felgenhaueri Kim & Abele, 1988
* Alpheus fenneri Bruce, 1994
* Alpheus firmus Kim & Abele, 1988
* Alpheus floridanus Kingsley, 1878
* Alpheus foresti Banner & Banner, 1981
* Alpheus formosus Gibbes, 1850
* Alpheus frontalis H. Milne Edwards, 1837
* Alpheus fujitai Nomura & Anker, 2005
* Alpheus funafutensis Borradaile, 1899
* Alpheus fushima Nomura, 2009
* Alpheus galapagensis Sivertsen, 1933
* Alpheus gallicus Scioli & Anker, 2020
* Alpheus georgei Banner & Banner, 1982
* Alpheus glaber (Olivi, 1792)
* Alpheus gracilipes Stimpson, 1860
* Alpheus gracilis Heller, 1861
* Alpheus grahami Abele, 1975
* Alpheus haanii Ortmann, 1890
* Alpheus hailstonei Coutière, 1905
* Alpheus hebes Kim & Abele, 1988
* Alpheus heeia Banner & Banner, 1975
* Alpheus hephaestus Bracken-Grissom & Felder, 2014
* Alpheus heronicus Banner & Banner, 1982
* Alpheus heterocarpus (Yu, 1935)
* Alpheus heterochaelis Say, 1818
* Alpheus heurteli Coutière, 1897
* Alpheus hippothoe De Man, 1888
* Alpheus holthuisi Ribeiro, 1964
* Alpheus homochirus (Yu, 1935)
* Alpheus hoonsooi Kim & Abele, 1988
* Alpheus hoplocheles Coutière, 1897
* Alpheus hortensis Wicksten & McClure, 2003
* Alpheus hululensis Coutière, 1905
* Alpheus hutchingsae Banner & Banner, 1982
* Alpheus hyeyoungae Kim & Abele, 1988
* Alpheus hyphalus Chace, 1988
* Alpheus idiocheles Coutière, 1905
* Alpheus ikedosoma Komai, 2015
* Alpheus imitatrix De Man, 1909
* Alpheus immaculatus Knowlton & Keller, 1983
* Alpheus inca Wicksten & Méndez G., 1981
* Alpheus inopinatus Holthuis & Gottlieb, 1958
* Alpheus intrinsecus Spence Bate, 1888
* Alpheus japonicus Miers, 1879
* Alpheus javieri Anker, Hurt & Knowlton, 2009
* Alpheus kagoshimanus Hayashi & Nagata, 2000
* Alpheus karplusi Anker, 2022
* Alpheus kuroshimensis Nomura & Anker, 2005
* Alpheus labis Banner & Banner, 1982
* Alpheus lacertosus Kim & Abele, 1988
* Alpheus ladronis Banner, 1956
* Alpheus lanceloti Coutière, 1905
* Alpheus lanceostylus Banner, 1959
* Alpheus lancirostris Rankin, 1900
* Alpheus latus Kim & Abele, 1988
* Alpheus lentiginosus Anker & Nizinski, 2011
* Alpheus lepidus De Man, 1908
* Alpheus leptocheles Banner & Banner, 1975
* Alpheus leptochiroides De Man, 1909
* Alpheus leptochirus Coutière, 1905
* Alpheus leviusculus Dana, 1852
* Alpheus lobidens De Haan, 1849
* Alpheus longecarinatus Hilgendorf, 1879
* Alpheus longichaelis Carvacho, 1979
* Alpheus longiforceps Hayashi & Nagata, 2002
* Alpheus longinquus Kim & Abele, 1988
* Alpheus longipalma Komai & Ohtomi, 2018
* Alpheus lottini Guérin, 1829
* Alpheus luiszapatai Ramos-Tafur, 2018
* Alpheus lutosus Anker & De Grave, 2009
* Alpheus macellarius Chace, 1988
* Alpheus macrocheles (Hailstone, 1835)
* Alpheus macrodactylus Ortmann, 1890
* Alpheus macroskeles Alcock & Anderson, 1899
* Alpheus maindroni Coutière, 1898
* Alpheus malabaricus (Fabricius, 1775)
* Alpheus malleator Dana, 1852
* Alpheus malleodigitus (Spence Bate, 1888)
* Alpheus mangalis Anker, 2023
* Alpheus mannarensis Purushothaman, Abhilash, Ajith Kumar & Lal, 2021
* Alpheus margaritae Salgado-Barragán, Ayón-Parente & Zamora-Tavares, 2017
* Alpheus martini Kim & Abele, 1988
* Alpheus mathewsae Anker, 2012
* Alpheus mazatlanicus Wicksten, 1983
* Alpheus microrhynchus De Man, 1897
* Alpheus microscaphis (Banner, 1959)
* Alpheus microstylus (Spence Bate, 1888)
* Alpheus midas Pachelle & Anker, 2024
* Alpheus miersi Coutière, 1898
* Alpheus migrans Lewinsohn & Holthuis, 1978
* Alpheus millsae Anker, Hurt & Knowlton, 2007
* Alpheus mitis Dana, 1852
* Alpheus mohammadpouri Dehghani, Sari & Naderloo, 2019
* Alpheus moretensis Banner & Banner, 1982
* Alpheus naos Anker, Hurt & Knowlton, 2007
* Alpheus naranjo Anker, 2018
* Alpheus nipa Banner & Banner, 1985
* Alpheus nobili Banner & Banner, 1966
* Alpheus nomurai Anker, 2023
* Alpheus nonalter Kensley, 1969
* Alpheus normanni Kingsley, 1878
* Alpheus notabilis Stebbing, 1915
* Alpheus novaezealandiae Miers, 1876
* Alpheus nuno Anker, 2012
* Alpheus nuttingi (Schmitt, 1924)
* Alpheus oahuensis (AH Banner, 1953)
* Alpheus obesomanus Dana, 1852
* Alpheus octocellatus Anker & Benzoni, 2023
* Alpheus ovaliceps Coutière, 1905
* Alpheus pachychirus Stimpson, 1860
* Alpheus pacificus Dana, 1852
* Alpheus packardii Kingsley, 1880
* Alpheus paludicola Kemp, 1915
* Alpheus panamensis Kingsley, 1878
* Alpheus papillosus Banner & Banner, 1982
* Alpheus paracrinitus Miers, 1881
* Alpheus paradentipes Coutière, 1905
* Alpheus paraformosus Anker, Hurt & Knowlton, 2008
* Alpheus paralcyone Coutière, 1905
* Alpheus paralpheopsides Coutière, 1905
* Alpheus parasocialis Banner & Banner, 1982
* Alpheus pareuchirus Coutière, 1905
* Alpheus parvimaculatus Nomura & Anker, 2005
* Alpheus parvirostris Dana, 1852
* Alpheus parvus De Man, 1909
* Alpheus peasei (Armstrong, 1940)
* Alpheus percyi Coutière, 1908
* Alpheus perezi Coutière, 1908
* Alpheus perlas Anker & Pachelle, 2019
* Alpheus perplexus Banner, 1956
* Alpheus petronioi Almeida, Terrosi & Mantelatto, 2014
* Alpheus philoctetes De Man, 1909
* Alpheus platycheirus Boone, 1927
* Alpheus platydactylus Coutière, 1897
* Alpheus platyunguiculatus (AH Banner, 1953)
* Alpheus polystictus Knowlton & Keller, 1985
* Alpheus polyxo De Man, 1909
* Alpheus pontederiae de Rochebrune, 1883
* Alpheus pouang Christoffersen, 1979
* Alpheus praedator De Man, 1908
* Alpheus proseuchirus De Man, 1908
* Alpheus pseudopugnax (AH Banner, 1953)
* Alpheus puapeba Christoffersen, 1979
* Alpheus pubescens De Man, 1908
* Alpheus pugnax Dana, 1852
* Alpheus punctatus Anker, 2012
* Alpheus pustulosus Banner & Banner, 1968
* Alpheus quasirapacida Chace, 1988
* Alpheus ramosportoae Soledade, Terossi, Scioli, Mantelatto & Almeida, 2019
* Alpheus randalli Banner & Banner, 1980
* Alpheus rapacida De Man, 1908
* Alpheus rapax Fabricius, 1798
* Alpheus rectus Kim & Abele, 1988
* Alpheus ribeiroae Anker & Dworschak, 2004
* Alpheus richardsoni Yaldwyn, 1971
* Alpheus richpalmeri Anker, 2020
* Alpheus roblesi Bracken-Grissom & Felder, 2014
* Alpheus romensky Burukovsky, 1990
* Alpheus roquensis Knowlton & Keller, 1985
* Alpheus roseodigitalis Nomura & Anker, 2005
* Alpheus rostratus Kim & Abele, 1988
* Alpheus rudolphi Almeida & Anker, 2011
* Alpheus rugimanus A. Milne-Edwards, 1878
* Alpheus samoa Banner & Banner, 1966
* Alpheus samudra De Grave, Krishnan, Kumar K.P. & Christodoulou, 2020
* Alpheus sarapis Anker, 2025
* Alpheus savuensis De Man, 1908
* Alpheus saxidomus Holthuis, 1980
* Alpheus schmitti Chace, 1972
* Alpheus schubarti Anker, 2024
* Alpheus sciolii Anker, 2022
* Alpheus scopulus Kim & Abele, 1988
* Alpheus serenei Tiwari, 1964
* Alpheus shukran Anker, 2024
* Alpheus sibogae De Man, 1908
* Alpheus simus Guérin-Méneville, 1855
* Alpheus sizou Banner & Banner, 1967
* Alpheus socialis Heller, 1862
* Alpheus soelae Banner & Banner, 1986
* Alpheus songkla Banner & Banner, 1966
* Alpheus soror Bruce, 1999
* Alpheus spatulatus Banner & Banner, 1968
* Alpheus splendidus Coutière, 1897
* Alpheus spongiarum Coutière, 1897
* Alpheus stanleyi Coutière, 1908
* Alpheus stantoni Banner & Banner, 1986
* Alpheus staphylinus Coutière, 1908
* Alpheus stephensoni Banner & Smalley, 1969
* Alpheus strenuus Dana, 1852
* Alpheus styliceps Coutière, 1905
* Alpheus sudara Banner & Banner, 1966
* Alpheus sulcatus Kingsley, 1878
* Alpheus sulcipalma Purushothaman, Bharathi, Damodhar, Kumar & Lal, 2023
* Alpheus suluensis Chace, 1988
* Alpheus supachai Banner & Banner, 1966
* Alpheus superciliaris Coutière, 1905
* Alpheus takla Anker, 2023
* Alpheus talismani Coutière, 1898
* Alpheus tampensis Anker, 2012
* Alpheus tasmanicus Banner & Banner, 1982
* Alpheus tenuicarpus De Man, 1908
* Alpheus tenuipes De Man, 1910
* Alpheus tenuis Kim & Abele, 1988
* Alpheus thomasi Hendrix & Gore, 1973
* Alpheus thompsoni Anker, 2022
* Alpheus tigrinus Anker, 2024
* Alpheus tirmiziae Kazmi, 1974
* Alpheus tricolor Anker, 2001
* Alpheus triphopus Nobili, 1906
* Alpheus tungii Banner & Banner, 1966
* Alpheus ulalae Bracken-Grissom & Felder, 2014
* Alpheus umbo Kim & Abele, 1988
* Alpheus utriensis Ramos & von Prahl, 1989
* Alpheus vanderbilti Boone, 1930
* Alpheus vanuatu Anker, 2020
* Alpheus verrilli (Schmitt, 1924)
* Alpheus villosus (Olivier, 1811)
* Alpheus villus Kim & Abele, 1988
* Alpheus viridari (Armstrong, 1949)
* Alpheus viserion Anker, Leray & Pachelle, 2021
* Alpheus vladivostokiensis (Vinogradov, 1950)
* Alpheus waltervadi Kensley, 1969
* Alpheus websteri Kingsley, 1880
* Alpheus wickstenae Christoffersen & Ramos, 1987
* Alpheus williamsi Bruce, 1994
* Alpheus wonkimi Anker & Pachelle, 2013
* Alpheus xanthocarpus Anker, Hurt & Knowlton, 2008
* Alpheus xishaensis Liu & Lan, 1980
* Alpheus zarenkovi Anker & Pachelle, 2015
* Alpheus zimmermani Anker, 2007
* Alpheus zulfaquiri Kazmi, 1982