# Cochlostoma

Cochlostoma is a genus of land snails with an operculum, terrestrial gastropod mollusks in the family Cochlostomatidae.

## Distribution

Calcareous terrains of South Europe and North Africa.

## Description and Anatomy

Mantle caves serve as pulmones. Operculum corneous and multispiral. Penis is oriented above the right tentacle. Sexual dimorphism is often recognisable in external morphology.

## Ecology

All species are confined to limestone habitats, feeding on dead plants and microflora. Females lays large eggs, hatching after three to six weeks.

## Species

The genus Cochlostoma contains at least 140 species and subspecies, including:
subgenus Auritus
* Cochlostoma achaicum (O. Boettger, 1885)
* Cochlostoma alleryanum (Paulucci, 1879)
* Cochlostoma auritum (Rossmässler, 1837)
* Cochlostoma fuchsi Feher, 2004
* Cochlostoma georgi (A.J. Wagner, 1906)
* Cochlostoma hellenicum (Saint-Simon, 1869)
* Cochlostoma hoyeri (Polinski, 1922)
* Cochlostoma paladilhianum (Saint-Simon, 1869)
* Cochlostoma parnonis Schütt, 1981
* Cochlostoma pinteri Feher, 2004
* Cochlostoma roseoli (A.J. Wagner, 1901)
* Cochlostoma tessellatum (Rossmässler, 1837)
* Cochlostoma westerlundi (Paulucci, 1879)
subgenus Cochlostoma
* Cochlostoma septemspirale (Razoumowsky, 1789)
subgenus Turritus
* Cochlostoma acutum (Caziot, 1908)
* Cochlostoma adamii (Paulucci, 1879)
* Cochlostoma affine (Benoit, 1882)
* Cochlostoma braueri (A.J. Wagner, 1897)
* Cochlostoma crosseanum (Paulucci, 1879)
* Cochlostoma elegans (Clessin, 1879)
* Cochlostoma euboicum (Westerlund, 1885)
* Cochlostoma gracile (L. Pfeiffer, 1849)
* Cochlostoma kleciaki (Braun, 1887)
* Cochlostoma macei (Bourguignat, 1869)
* Cochlostoma montanum (Issel, 1866)
* Cochlostoma mostarensis (A.J. Wagner, 1906)
* Cochlostoma nanum (Westerlund, 1879)
* Cochlostoma pageti Klemm, 1962
* Cochlostoma patulum (Draparnaud, 1801)
* Cochlostoma sardoum (Westerlund, 1890)
* Cochlostoma simrothi (Caziot, 1908)
* Cochlostoma stossichi (Hirc, 1881)
* Cochlostoma sturanii (A.J. Wagner, 1897)
subgenus Dalfreddia
* Cochlostoma porroi (Strobel, 1850)
* Cochlostoma subalpinum (Pini, 1884)
subgenus Clessiniella
* Cochlostoma anomphale Boeckel, 1939
* Cochlostoma stelucarum Zallot et al., 2021
* Cochlostoma tergestinum (Westerlund, 1878)
* Cochlostoma villae (Strobel, 1851)
* Cochlostoma waldemari (A.J. Wagner, 1897)
subgenus Eupomatias
* Cochlostoma henricae (Strobel, 1851)
* Cochlostoma philippianum (Gredler, 1853)
subgenus Wagneriola
* Cochlostoma scalarinum (A. & J.B. Villa, 1841)
subgenus Lovcenia
* Cochlostoma dalmatinum (L. Pfeiffer, 1863)
* Cochlostoma erika (A.J. Wagner, 1906)
* Cochlostoma jakschae Zallot, Fehér, Bamberger, Gittenberger, 2015
* Cochlostoma lanatum Zallot, Fehér, Bamberger, Gittenberger, 2015
* Cochlostoma tropojanum Zallot, Fehér, Bamberger, Gittenberger, 2015
unknown subgenus
* Cochlostoma cinerascens (Rossmässler, 1837)
* Cochlostoma cretense (Maltzan, 1887)

## Synonyms of Cochlostoma

* Auritus Westerlund, 1883
* Auritus (Auritus) Westerlund, 1883
* Auritus (Holcopoma) Kobelt & Möllendorff, 1899
* Auritus (Pleuropoma) A. J. Wagner, 1897
* Auritus (Titanopoma) A. J. Wagner, 1897
* Cochlostoma (Auritus) Westerlund, 1883· accepted, alternate representation
* Cochlostoma (Cinerascens) A. J. Wagner, 1897· accepted, alternate representation
* Cochlostoma (Clessiniella) Zallot, Groenenberg, De Mattia, Fehér & E. Gittenberger, 2015 (junior objective synonym)
* Cochlostoma (Cochlostoma) Jan, 1830· accepted, alternate representation
* Cochlostoma (Dalfreddia) Zallot, Groenenberg, De Mattia, Fehér & E. Gittenberger, 2015· accepted, alternate representation
* Cochlostoma (Eupomatias) A. J. Wagner, 1897· accepted, alternate representation
* Cochlostoma (Holcopoma) Kobelt & Möllendorff, 1899
* Cochlostoma (Lovcenia) Zallot, Groenenberg, De Mattia, Fehér & E. Gittenberger, 2015· accepted, alternate representation
* Cochlostoma (Neglecta) A. J. Wagner, 1897· accepted, alternate representation
* Cochlostoma (Scalarina) A. J. Wagner, 1897· accepted, alternate representation
* Cochlostoma (Titanopoma) A. J. Wagner, 1897 (junior synonym)
* Cochlostoma (Turritus) Westerlund, 1883· accepted, alternate representation
* Cyclostoma (Cochlostoma) Jan, 1830 (original rank)
* Diversicolor A. J. Wagner, 1905 (a junior synonym)
* Hartmannia Newton, 1891
* Holcopoma Kobelt & Möllendorff, 1899
* Patuliana Caziot, 1908
* Personatus Westerlund, 1883
* Pleuropoma A. J. Wagner, 1897
* Pleuropomatia Tomlin, 1929
* Pomatias (Atlantica) A. J. Wagner, 1897 (invalid: junior homonym of Atlantica Ancey, 1887)
* Pomatias (Auritus) Westerlund, 1883
* Pomatias (Cinerascens) A. J. Wagner, 1897
* Pomatias (Cisalpina) A. J. Wagner, 1897
* Pomatias (Difficilis) A. J. Wagner, 1897 (invalid:junior objective synonym of Hartmannia)
* Pomatias (Eupomatias) A. J. Wagner, 1897
* Pomatias (Maculatus) Westerlund, 1883
* Pomatias (Nana) A. J. Wagner, 1897 (invalid: junior homonym of Nana Schumacher, 1817)
* Pomatias (Neglecta) A. J. Wagner, 1897
* Pomatias (Personatus) Westerlund, 1883
* Pomatias (Philippiana) A. J. Wagner, 1897
* Pomatias (Pleuropoma) A. J. Wagner, 1897 ( invalid: junior homonym of Pleuropoma Möllendorff, 1893; * * Holcopoma and Pleuropomatia are replacement names)
* Pomatias (Pomatiella) Clessin, 1889
* Pomatias (Sardoa) A. J. Wagner, 1897
* Pomatias (Scalarina) A. J. Wagner, 1897
* Pomatias (Septemspiraliana) Caziot, 1910
* Pomatias (Septemspiralis) A. J. Wagner, 1897
* Pomatias (Strobelia) Clessin, 1889 (invalid: junior homonym of Strobelia Rondani, 1868 [Diptera])
* Pomatias (Tesselata) A. J. Wagner, 1897
* Pomatias (Titanopoma) A. J. Wagner, 1897 (junior objective synonym of Auritus)
* Pomatias (Turritus) Westerlund, 1883
* Stereopoma A. J. Wagner, 1897 (junior synonym)