# Calcaires du Bou Dahar

The Calcaires du Bou Dahar (also known as Djebel Bou Dahar Paleoshoal, Djebel Bou Dahar, Calcaires du Bou Dahar Formation, or Bou Dahar Formation) is a geological formation or a sequence of formations of Late Sinemurian to Pliensbachian-Toarcian boundary (Early Jurassic) age in Bni Tadjite, the Central High Atlas, Morocco. This unit represents an excepcional record of an evolving reef complex (mountain laterals), platform slopes (Steep slopes between 20° and 35° on various sides) and a emerged shoal (nearly horizontal limestone layers on the top flat Bou Dahar plateau) developed inside a carbonate platform, recording the evolutionary cycles of this environment with notorious precision, also yielding what is considered one of the greatest/most diverse marine biotas of the entire Jurassic Tethys Ocean. The Bou Dahar carbonate platform shoal stands prominently and structurally above surrounding plains, spanning 35-40 km in length and 4-15 km in width, with a relief of 100-450 m. This carbonate formation originated on metamorphosed Silurian to Ordovician siliciclastic rocks and tholeiitic volcanic layers tied to Central Atlantic Magmatic Province basalts, forming a corridor oriented WSW to ENE. Surrounding alluvial plains expose green marls, shales, and dark lime-mudstones representing basinal deposits contemporaneous or subsequent to the platform. It has been considered to be a sequence of different coeval inner geological formations, including the Foum Zidet Formation, the Aganane Formation and Ouchbis Formation, but is usually interpreted as a single major unit due to it´s unique preservation.

## Economic importance

Lead, Zinc, and Baryte mining in the Bou Dahar district relies solely on artisanal methods managed by the public "CADETAF" cooperative. The cooperative’s claim area, spanning 60,000 km$^2$ across the central and eastern High Atlas, along with exploration licenses held by ManagemGroup, together constitute North Africa’s largest calamine region. Data from CADETAF and ManagemGroup reveal over 1 million tons of zinc ore with more than 30% ZnO content extracted to date, including 559,403 tons of 16% Zn ore produced by ManagemGroup from 2012 to 2019.

## Sequence evolution

The Bou Dahar platform is a well-preserved example of carbonate platform development within a tectonically active rift-basin environment. This region formed as part of the Jurassic High Atlas seaway, where tectonic activity shaped the basin, with influences from rifting, faulting, and transtensional forces. Early studies mapped the area’s Jurassic marine environments, with carbonate platform margins situated alongside deeper basin limestones. This platform records two primary periods of shallow-water carbonate growth: one during the Lower to Middle Liassic (Sinemurian to Domerian) and a subsequent phase spanning the Upper Liassic to Lower Dogger (Aalenian to Bajocian). However, only the earlier phase is preserved in Djebel Bou Dahar. This phase was abruptly interrupted at the Domerian-Toarcian transition, marked by the onlap of Toarcian shales and Aalenian lime mudstones over the platform, signaling a shift to deeper marine conditions.
The seismic modeling of Bou Dahar platform highlighted large-scale structures, such as the platform's form and the boundary marking its submergence, across various frequencies and modeling approaches. Lower-frequency wavelets emphasized truncation of layers and outward shifts in onlap points. In contrast, internal platform facies zones were less distinct at frequencies below 50 Hz, but with a geometry still discernible through disturbances in reflections, especially near the lower boundaries, with image-ray migration models proving more responsive to these subtle variations even at lower frequencies. Seismic contrast, driven by differences in acoustic impedance, tends to decline with deeper burial and subsequent diagenesis, with peak resolution typically present at initial deposition stages before significant burial effects.
Sequences I-II overlie CAMP basalts, spanning over 126 & 150 meters, marked by a dolomite-rich boundary featuring fenestral and stromatolitic structures, which indicate transgressive-regressive (T-R) cycles. Dominance of subtidal carbonates suggests extensive shallow marine conditions across the platform. Early dolomitization preserved the fine microstructures within these sequences, aligning with the Hettangian-Lower Sinemurian Idikel Formation.
Sequence III developed as a low-relief carbonate platform and varies in slope angle between northern and southern margins, with thickness peaking in northeastern synclines at around 100 meters. Thinning toward the center, it comprises laterally continuous skeletal and packstone deposits interspersed with sponge mounds, mirroring the Foum Zidet Formation.
Sequence IV with a wedge-shaped geometry, reveals steep slopes and retrogradational stacking, indicating subaerial exposure at its base with ammonite-rich wackestone beds. This sequence thickens at the southern margin, where coral and sponge boundstones, extending to 140 meters, transition into grainstone and packstone accumulations along the slope. The eastern platform edge hosts cross-bedded grainstone layers, lacking intertidal deposits, characteristic of open-marine conditions.
Sequence V, along Sequences IV to VI correlate with the Aganane Formation and Ouchbis Formations, being a progradational wedge, is thickest along the platform's southern and eastern edges, forming a concave-up clinoform structure, reaching 50-70 meters in height and indicating alternating subtidal and intertidal lithofacies and evidence of exposure events. Moving basinward, finer basinal sediments with green marls and peloidal wackestones emerge, suggesting periodic sediment starvation during the Early Pliensbachian.
The final sequence, Sequence VI, fills the last stage of platform development with angular unconformity over older layers and CAMP basalts. This sequence encompasses varied facies belts surrounding a central paleo-high with subtidal and intertidal deposits, coated-grain bars, and coastal-plain sediments like red shales and calcretes. Clinoforms along the lower slope show a concave-up profile that flattens basinward, with deposits consolidating into a fan along the southern margin, evidence of sediment redistribution likely from mass wasting.

## Paleoenvironment

The paleoenvironment of Bou Dahar is characterized by a dynamic evolution through distinct stages: pre-drowning, drowning, and post-drowning, each marked by specific sedimentological and ecological changes that reflect shifts in energy conditions and depositional environments.
During the pre-drowning phase, the carbonate platform was dominated by tranquil lagoonal settings, indicative of low-energy environments. These conditions fostered the accumulation of microsparite carbonates, which are essential in maintaining the stable conditions found in lagoons. The presence of bioturbation, mud cracks, and tidal flats illustrates the periodic exposure of the substrate, suggesting a complex interplay of biogenic and abiogenic processes. The platform margin transitioned into coarser zones where higher-energy environments were characterized by sorted carbonate sands and natural barriers. This transitional buffer from the lagoon to slope facilitated the formation of intertidal mounds, which played a crucial role in the sediment dynamics of the region.
As the platform underwent drowning, it shifted towards high-energy conditions. Sediments from this phase exhibited dense biosparites with significant skeletal deposits, indicating heightened wave action and the erosion of previously accumulated lagoonal deposits. The transition to these high-energy environments underscores a significant change in the depositional dynamics, where sediment transport was dominated by stronger currents. The emergence of tepee structures in the platform carbonates reflects this period of increased energy. Formed from early lithification and sediment infill in intertidal zones, these structures were characterized by cracks filled with fine sediments and stromatolitic algae, highlighting rapid infilling processes shortly after deposition. Unlike their recent counterparts in the Persian Gulf, the Moroccan tepees developed in fully lithified rocks, shaped by thermal expansion and contraction driven by temperature variations in intertidal settings.
Microscopic examination of the sediments reveals a rich diversity of biogenic and non-biogenic components, categorized into nine distinct microfacies types. These include laminated pelmicrites, biopelsparites, and various intraclasts and bioclasts, showcasing the complex interplay of environmental factors influencing sediment composition. Foraminifers along with corals, indicate thriving marine ecosystems during the deposition of Bou Dahar. The various microfacies illustrate shifts in sedimentation dynamics, with microfacies types reflecting different depositional environments-from tranquil lagoonal settings to more energetic reef environments.
Local carbonate factories present a unique contrast with other coeval units in the Tethys, with siliceous sponge microbial boundstone mounds, which were more prevalent in Middle and Upper Jurassic systems. Despite their earlier emergence, these mounds show similarities to younger Jurassic formations in texture and biological components. They thrived in deep-water environments, particularly in areas that experienced low detrital input. Siliceous sponge mounds record distinct rare earth element and yttrium patterns among carbonate components, indicating that Early Jurassic seawater resembled modern distributions. Microbialites contained higher rare earth element concentrations, likely influenced by organic matter and diagenetic processes. These mounds thrived in well-oxygenated environments, emphasizing the need to examine individual carbonate types to understand their unique geochemical signatures. The coral calcareous sponge microbial boundstone aligns with the coral-microbolite-debris reef classification, indicating a high-energy, shallow-water habitat.

## Fossil content

| Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon | Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited. |

### Foraminifera

| Genus | Species | Location | Material | Notes |
| Agerina | A. martana | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Cornuspiridae family |
| Ammobaculites | A. coprolithiformis A. spp. | Bou Dahar | Isolated Tests/Shells | A foraminifer of the family Ammomarginulininae. |
| Bosniella | B. oenensis | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Biokovinidae family |
| Bullopora | B. spp. | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Webbinellinae family |
| Everticyclammina | E. praevirguliana | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Everticyclamminidae family. |
| Kurnubia | K. sp. | Bou Dahar | Isolated Tests/Shells | A foraminifer of the family Kurbuniinae |
| Glomospira | G. sp. | Bou Dahar | Isolated Tests/Shells | A foraminifer of the family Ammodiscidae. |
| Haurania | H. amiji H. deserta H. ssp. | Bou Dahar | Isolated Tests/Shells | A foraminifer of the family Hauraniinae. |
| Involutina | I. farinacciae I. liassica | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Involutinidae family |
| Lituosepta | L. recoarensis L. compressa L. ssp. | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Mesoendothyridae family. |
| Mesoendothyra | M. sp. | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Mesoendothyridae family. |
| Mayncina | M. termieri | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Mayncinidae family |
| Nodosaria | N. sexcostata N. sp. | Bou Dahar | Isolated Tests/Shells | A foraminifer of the family Nodosariinae. |
| Orbitopsella | O. primaeva O. preacursor O. dubari O. spp. | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Mesoendothyridae family. |
| Ophtalmidium | O. concentricum O. sp. | Bou Dahar | Isolated Tests/Shells | A foraminifer of the family Ophthalmidiidae. |
| Pseudocyclammina | P. liasica P. spp. | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Pfenderinidae family. |
| Planisepta | P. compressa | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Planiseptinae family. |
| Reophax | R. sp. | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Reophacinae family |
| Siphovalvulina | S. gibraltarensis S. variabilis S. sp.A | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Pfenderinidae family. |
| Tetrataxis | T. spp. | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Tetrataxidae family. |
| Textularopsis | T. sinemuriensis | Bou Dahar | Isolated Tests/Shells | A foraminifer of the family Nodosariinae. |
| Trochammina | T. sp | Bou Dahar | Isolated Tests/Shells | A foraminifer of the Trochamminidae family |

### Sponges

| Genus | Species | Stratigraphic position | Material | Notes | Images |
| Arhoussia | A. calyx | Jebel Bou Arhous Srhir | Colonial Imprints | A Glass sponge, member of the family Euplectellidae | |
| Chaetetes | C. (Pseudoseptifer) zignoi C. (Pseudoseptifer) murciana C. (Pseudoseptifer) liasica | Bou Dahar | Colonial Imprints | A Demosponge, member of the family Chaetetinae. Monospecific assamblages with encrusting and symbiont forms | Specimen of the same genus |
| Palaeopora | P. ramulosa P. liasica | Bou Dahar | Colonial Imprints | A Calcareous sponge, Incertade sedis affinities | |
| Stromatomorpha | S. californica S. cf. californica S. liassica S. (Cylicopsis) atlantis | Bou Dahar IttoFezzou Ait-Ani | Colonial Imprints | A Demosponge, member of the family Anthaspidellidae | |
| Tillichtia | T. aedificator T. spp. | Kadoussa | Colonial Imprints | A Glass sponge, member of the family Euplectellidae | |

### Hydrozoa

| Genus | Species | Stratigraphic position | Material | Notes | Images |
| Spongiomorpha | S. ramosa S. gibbosa S. (Heptastylopsis) sp. 1 | Bou Dahar | Colonial Imprints | A Hydrozoan, member of the family Spongiomorphidae | |
| Tubiphytes | T spp. | Bou Dahar | Colonial Imprints | A Hydrozoan, member of the family Nigriporellidae | Tubiphytes specimens |

### Anthozoa

| Genus | Species | Stratigraphic position | Material | Notes | Images |
| Actinastrea | A. minima | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Actinastreidae | |
| Allocoeniopsis | A. dendroidea | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Actinastreidae | |
| Alloiteausmilia | A. boudaharensis | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Zardinophyllidae | |
| Araiophyllum | A. liasicum | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Comoseridae | |
| Archaeosmilia | A. menchikoffi A. beata A. sp. | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Archaeosmiliidae | |
| Brachyseris | B. walfordi | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Latomeandridae | |
| Chomatoseris | C. orbulites | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Comoseridae | |
| Cladophyllia | C. dresnayi | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Cladophylliidae | |
| Coenastraea | C. cf.maccoyi C. sp. | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Actinastreidae | |
| Cyclophyllopsis | C. cornutiformis | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Cyclophyllopsiidae | |
| Dimorphastraea | D. menchikoffi | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Latomeandridae | |
| Eocomoseris | E. minima | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Comoseridae | |
| Epismilia | E. mauretaniensis E.? sp. | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Zardinophyllidae | |
| Epismiliopsis | E. liasica E. sp. | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Zardinophyllidae | |
| Epistreptophyllum | E. vetusum | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Dermosmiliidae | Epistreptophyllum specimen (from Germany) |
| Funginella | F. domeriensis | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Funginellidae | |
| Goniocora | G. concinna | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Stylinidae | Goniocora specimens |
| Haplaraea | H. spp. | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Astraraeidae | |
| Margarosmilia | M. gemminata | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Margarophylliidae | Margarosmilia specimens |
| Mesomorpha | M. gracilis M. bussoni | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Thamnasteriidae | |
| Neorylstonia | N. pseudocolumellata N. spp. | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Volzeioidea | |
| Omphalophylliopsis | O. lobatus | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Conophyllidae | |
| Plesiophyllum | P. cylindricum | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Zardinophyllidae | |
| Proleptophyllia | P. multiradiata P. rejuvenescens P. subcylindrica P. bilobata | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Acrosmiliidae | |
| Rodinosmilia | R. elegantula | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Pachycoeniidae | |
| Stylosmilia | S. dresnayi | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Stylinidae | |
| Stylophyllopsis | S. vetusum S. cf. rudis | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Stylophyllidae | |
| Trigerastrea | T. minima | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Latomeandridae | |
| Volzeia | V. badiotica | Bou Dahar | Imprints & isolated specimens | A Stony coral member of the family Volzeiidae | |

### Brachiopoda

| Genus | Species | Stratigraphic position | Material | Notes |
| Aulacothyris | A. semiplana | Bou Dahar | Isolated Shells | A Terebratulidan, member of the family Zeilleriidae |
| Dictyothyris | D. drepanensis | Bou Dahar | Isolated Shells | A Terebratulidan, member of the family Dictyothyrididae |
| Fimbriothyris | F. itoensis F. tenuiplicata F. brevicostata | Bou Dahar | Isolated Shells | A Terebratulidan, member of the family Zeilleriidae |
| Linguithyris | L. aspasia L. adnethensis | Bou Dahar | Isolated Shells | A Terebratulidan, member of the family Nucleatidae |
| Rhynchonella | R. quadrata R. spp. | Bou Dahar | Isolated Shells | A Rhynchonellidan, member of the family Rhynchonellidae |
| Prionorhynchia | P. msougari P. subcynocephala P. rubriaxensis | Bou Dahar | Isolated Shells | A Rhynchonellidan, member of the family Wellerellidae |
| Prototegulithyris | P. khadijae | Bou Dahar | Isolated Shells | A Terebratulidan, member of the family Dictyothyrididae |
| Pseudogibbirhynchia | P. jurensis P. moorei P. waehneri | Bou Dahar | Isolated Shells | A Rhynchonellidan, member of the family Basiliolidae |
| Stolmorhynchia | S. limata | Bou Dahar | Isolated Shells | A Rhynchonellidan, member of the family Wellerellidae |
| Suessia | S. liasina | Bou Dahar | Isolated Shells | A Terebratulidan, member of the family Zeilleriidae |

### Gastropoda

| Genus | Species | Stratigraphic position | Material | Notes | Images |
| Actaeonina | A. radius | Bou Dahar N7 Bou Dahar "Carriere" | Isolated Shells | An Architectibranchian, member of the family Acteonidae | |
| Amphitrochilia | A. (Aristerella) undata A. (Aristerella) rorata | Bou Dahar N2 Bou Dahar N3 Bou Dahar N5 Bou Dahar N6 Bou Dahar N7 Bou Dahar N12 Bou Dahar "Carriere" | Isolated Shells | A Trochoidean, member of the family Proconulidae | |
| Ampullina | A. (Ampullospira) dominans A. sp. nov A. (Pseudamura) sp. nov | Bou Dahar N1 Bou Dahar N2 Bou Dahar N6 Bou Dahar N12 Bou Dahar "Carriere" | Isolated Shells | A Caenogastropodan, member of the family Ampullinidae | Ampullina specimens |
| Aphanoptyxis | A. cossmanni | Bou Dahar N7 Bou Dahar N9 | Isolated Shells | A Nerineoidean, member of the family Ceritellidae | |
| Aptyxiella | A. praevia A. fimbriata A. subgradata | Bou Dahar N1 Bou Dahar N2 Bou Dahar N3 Bou Dahar N5 Bou Dahar N6 Bou Dahar N7 Bou Dahar N12 Bou Dahar "Carriere" | Isolated Shells | A Nerineoidean, Incertade sedis | |
| Ataphrus | A. cf.piettei A. (Pleuratella) cf.normaniensis | Bou Dahar N3 Bou Dahar N7 Bou Dahar "Carriere" Tizi N´Oumzour | Isolated Shells | A Trochoidean, member of the family Ataphridae | |
| Boehmia | B. speciosa B. costata | Bou Dahar N3 Bou Dahar N6 | Isolated Shells | A Architectibranchian, member of the family Acteonidae | |
| Brachytrema | B. despujolsi | Bou Dahar N2 Bou Dahar N3 Bou Dahar N5 Bou Dahar N7 Bou Dahar N12 | Isolated Shells | A Caenogastropodan, member of the family Brachytrematidae | |
| Buvignieria | B. sp. | Bou Dahar | Isolated Shells | A Littorinimorphan, member of the family Palaeorissoinidae | |
| Cerithiella | C. africana C. sarrazini C. aff.nudae | Bou Dahar N1 Bou Dahar N3 Bou Dahar N7 Bou Dahar N8 Bou Dahar "Carriere" | Isolated Shells | A Triphoroidean, member of the family Newtoniellidae | Cerithiella specimen |
| Cerithinella | C. termieri C. cf.italica C. gratiosa C. miliare C. gemmifera C. margaritata C. baccata C. cf.baccata C. aff.cerithiiformis C. (Laevibaculus) sp. | Bou Dahar N1 Bou Dahar N2 Bou Dahar N3 Bou Dahar N5 Bou Dahar N6 Bou Dahar N7 Bou Dahar N11 Bou Dahar N12 | Isolated Shells | A Caenogastropodan, member of the family Procerithiidae | |
| Climacina | C. daharensis C. cf.mariae C. sp. | Bou Dahar N2 Bou Dahar N3 Bou Dahar N7 | Isolated Shells | A Caenogastropodan, member of the family Procerithiidae | |
| Coelostylina | C. chartroni C. choffati C. rigida C. (Cosmostylina) darestei C. (Cosmostylina) lima | Bou Dahar N2 Bou Dahar N3 Bou Dahar N4 Bou Dahar N6 Bou Dahar N7 Tizi N´Oumzour Aguelmane-Azougar | Isolated Shells | A Caenogastropodan, member of the family Coelostylinidae | |
| Crossostoma | C. atlantis | Bou Dahar N7 Tizi N´Oumzour | Isolated Shells | A Trochoidean, member of the family Colloniidae | |
| Cryptaulax | C. angulodentatum C. album C. cf.contortum C. cf.densestritum C. aff.tetrataeniatum C. cf.tetrataeniatum | Bou Dahar N2 Bou Dahar N3 Bou Dahar N4 Bou Dahar N6 Bou Dahar N7 Bou Dahar N11 | Isolated Shells | A Cerithiimorphan, member of the family Cryptaulacinae | |
| Diarthema | D. sp. | Bou Dahar N3 | Isolated Shells | A Littorinimorphan, member of the family Aporrhaidae | |
| Diatinostoma | D. sp. | Tizi N´Oumzour | Isolated Shells | A Caenogastropodan, member of the family Eustomatidae | |
| Dicroloma | D. sp. | Bou Dahar N7 | Isolated Shells | A Littorinimorphan, member of the family Aporrhaidae | |
| Discocirrus | D. crispus D. armatus | Bou Dahar N1 Bou Dahar N2 Bou Dahar N3 Bou Dahar N5 Bou Dahar N12 Bou Dahar "Carriere" | Isolated Shells | A Seguenziidan, member of the family Cirridae | |
| Discohelix | D. (Colpomphalus) spledens D. (Colpomphalus) cf.glauca D. (Colpomphalus) sp. A | Bou Dahar N3 Bou Dahar N6 Bou Dahar N7 Bou Dahar N12 Bou Dahar "Carriere" | Isolated Shells | A Seguenziidan, member of the family Eucyclidae | Discohelix specimen |
| Exelissa | E. pusilla E. douxamii E. spissa E. galenae E. menchikoffi E. cf.prealpina | Bou Dahar N1 Bou Dahar N2 Bou Dahar N3 Bou Dahar N4 Bou Dahar N6 Bou Dahar N7 Bou Dahar N8 Bou Dahar N9 Bou Dahar N12 Ait Daoud-ou-Azzi | Isolated Shells | A Cerithiimorphan, member of the family Cryptaulacinae | |
| Emarginula | E. henrici E. cf.orthogonia E. (Rimula) pileata E. (Rimula) sp. A E. (Rimula) sp. B | Bou Dahar N3 Bou Dahar N7 Bou Dahar N12 Bou Dahar "Carriere" Aït Daoud | Isolated Shells | A Lepetellidan, member of the family Fissurellidae | Emarginula specimens |
| Eucyclus | E. lateclathratus | Bou Dahar N3 | Isolated Shells | A Seguenziidan, member of the family Eucyclidae | |
| Fibulella | F. afra F. nicolai F. sp. | Bou Dahar, all the levels | Isolated Shells | A Nerineoidean, member of the family Ceritellidae | |
| Gymnocerithium | G. sp. | Bou Dahar "Carriere" | Isolated Shells | A Caenogastropodan, member of the family Campanilidae | |
| Hesperidina | H. striolata | Bou Dahar N7 Tizi N´Oumzour Imi n´Ikis | Isolated Shells | A Caenogastropodan, member of the family Purpurinidae | |
| Nerinella | N. marucchiensis N. cayeuxi N. cf.atava N. convoluta N. sp. | Bou Dahar N1 Bou Dahar N2 Bou Dahar N3 Bou Dahar N6 Bou Dahar N7 Bou Dahar N8 Bou Dahar N11 Bou Dahar N12 Bou Dahar "Carriere" | Isolated Shells | A Nerineoidean, member of the family Nerinellidae | |
| Neritodomus | N. calaminae N. (Sphaerochilus) daharensis N. cf.tethys | Bou Dahar N1 Bou Dahar N2 Bou Dahar N3 Bou Dahar N7 Bou Dahar "Carriere" Bou Dahar Plateau Tizi N´Oumzour | Isolated Shells | A Cycloneritida, member of the family Neridomidae | |
| Neritopsis | N. falloti N. sp. | Bou Dahar N2 Bou Dahar "Carriere" | Isolated Shells | A Cycloneritida, member of the family Neritopsidae | Neritopsis specimens |
| Ovacteonina | O. sp. | Bou Dahar N7 | Isolated Shells | An Architectibranchian, member of the family Acteonidae | |
| Paracerithium | P. lacostei P. echinatum P. liasinum P. merciaii P. ingens P. angulosum P. cf.strueveri P. cf.multiforme | Bou Dahar N1 Bou Dahar N2 Bou Dahar N3 Bou Dahar N6 Bou Dahar N7 Bou Dahar N11 Bou Dahar N12 Bou Dahar "Carriere" | Isolated Shells | A Caenogastropodan, member of the family Batillariidae | |
| Pileolus | P. laevis | Bou Dahar, all the levels Tizi N´Oumzour | Isolated Shells | A Cycloneritida, member of the family Pileolidae | |
| Platyacra | P. (Asperilla) russoi | Bou Dahar N1 Bou Dahar N2 Bou Dahar N12 Amellago | Isolated Shells | A Trochoidean, member of the family Angariidae | |
| Pleurotomaria | P. cf.pisana | Bou Dahar | Isolated Shells | A Pleurotomariidan, member of the family Pleurotomariidae | Pleurotomaria specimens |
| Procerithium | P. canescens P. curtum P. sp. | Bou Dahar N2 Bou Dahar N3 Bou Dahar N7 Bou Dahar N12 | Isolated Shells | A Caenogastropodan, member of the family Procerithiidae | |
| Proconulus | P. spinatus P. sp. A P. sp. B | Bou Dahar N3 Bou Dahar N6 Bou Dahar N7 Bou Dahar N12 Bou Dahar "Carriere" | Isolated Shells | A Trochoidean, member of the family Proconulidae | |
| Protorcula | P. fischeri | Bou Dahar N2 Bou Dahar N7 Bou Dahar N8 | Isolated Shells | A Pseudozygopleuroidea, member of the family Protorculidae | |
| Pseudomelania | P. aristomacha P. dealbata P. chrysallis P. (Gardetia) chouberti P. (Oonia) marucchiensis P. (Oonia) cf.turgidula P. (Oonia) aff.pupoideae P. (Oonia) sp. P. (Rhabdoconcha) tenuestriata P. cf.procera P. cf.eulimoides P. cf.rhea P. cf.globosa P. sp. | Bou Dahar N1 Bou Dahar N2 Bou Dahar N3 Bou Dahar N6 Bou Dahar N7 Bou Dahar N8 Bou Dahar N12 Bou Dahar "Carriere" Tizi N´Oumzour | Isolated Shells | A Caenogastropodan, member of the family Pseudomelaniidae | Pseudomelania specimen |
| Pseudonerinea | P. terebra P. liasica P. sp. A P. sp. | Bou Dahar N1 Bou Dahar N2 Bou Dahar N3 Bou Dahar N5 Bou Dahar N6 Bou Dahar "Carriere" | Isolated Shells | A Nerineoidean, member of the family Pseudonerineidae | |
| Purpurina | P. rudis P. moghrabiensis P. patroclus P. berberica | Bou Dahar N1 Bou Dahar N2 Bou Dahar N3 Bou Dahar N4 Bou Dahar N6 Bou Dahar N7 Bou Dahar N12 Bou Dahar "Carriere" Kasba d´Amellago Tizi N´Oumzour | Isolated Shells | A Caenogastropodan, member of the family Purpurinidae | Example of Purpurina specimen |
| Rigauxia | R. baculus | Bou Dahar N2 Bou Dahar N7 Bou Dahar "Carriere" | Isolated Shells | A Streptacidoidea, Incertade sedis | |
| Scurriopsis | S. altissima S. parma S. (Scurria?) cf.nana S. (Scurria?) sp. | Bou Dahar N3 Bou Dahar N7 Bou Dahar N12 Bou Dahar "Carriere" Aït Daoud El-Kansera | Isolated Shells | A Lottioidean, member of the family Acmaeidae | Scurria specimens |
| Teliochilus | T. cf.deslongchampsi T. clathratus | Bou Dahar N2 Bou Dahar N3 Bou Dahar N4 Bou Dahar N6 Bou Dahar N7 Bou Dahar N12 | Isolated Shells | A Cerithiimorphan, member of the family Cryptaulacinae | |
| Tectus | T. (Dimorphotectus) chouberti | Bou Dahar N3 Bou Dahar N7 Bou Dahar "Carriere" | Isolated Shells | A Trochoidean, member of the family Tegulidae | Tectus specimens |
| Teinostoma | T. sp. | Bou Dahar N2 Bou Dahar "Carriere" | Isolated Shells | A Littorinimorphan, member of the family Teinostomatidae | Teinostoma specimens |
| Trochotoma | T. (Ditremaria) secans T. (Ditremaria) pachyspira | Bou Dahar N3 Bou Dahar N7 Bou Dahar N12 Bou Dahar "Carriere" | Isolated Shells | A Pleurotomariidan, member of the family Trochotomidae | |
| Tylostoma | T. maurum | Bou Dahar N7 Bou Dahar "Carriere" | Isolated Shells | A Caenogastropodan, member of the family Tylostomatidae | Tylostoma specimen |
| Zygopleura | Z. pectita Z. appenninica Z. paradisi Z. tatia Z. menchikoffi Z. sp. | Bou Dahar N1 Bou Dahar N2 Bou Dahar N3 Bou Dahar N4 Bou Dahar N6 Bou Dahar N7 Bou Dahar N12 Bou Dahar "Carriere" | Isolated Shells | A Pseudozygopleuroidea, member of the family Zygopleuridae | Example of Zygopleura specimen |

### Polyplacophora

| Genus | Species | Stratigraphic position | Material | Notes | Images |
| Allochiton | A. sp. nov? | Bou Dahar | Isolated Shells | A Chitonidan, member of the family Mopaliidae | Example of modern Mopalia, relative of the same family |
| Heterochiton | H. cf.giganteus H. cf.zitteli H. sp. nov? | Bou Dahar | Isolated Shells | A Chitonidan, member of the family Mopaliidae | Example of modern Mopalia, relative of the same family |

### Bivalvia

| Genus | Species | Stratigraphic position | Material | Notes | Images |
| Anisocardia | A. quadrata | Bou Dahar N4 | Isolated Shells | An Veneridan, member of the family Arcticidae | |
| Astarte | A. vendaeensis A. marcaisi A. (Theveninia) aff. tetragona A. aff.interlineata A. sp. | Bou Dahar N3 Bou Dahar N5 Bou Dahar N6 Bou Dahar N12 Bou Dahar Sup | Isolated Shells | A Carditidan, member of the family Astartidae | Astarte specimens |
| Barbatia | B. metellaria | Bou Dahar N3 Bou Dahar N4 Bou Dahar N5 Bou Dahar N7 | Isolated Shells | An Arcidan, member of the family Arcidae | Barbatia specimens |
| Cardium | C. pustulatum C. galenae C. (Protocardia) praecalva C. sp. | Bou Dahar N2 Bou Dahar N3 Bou Dahar N7 Tizi N´Oumzour | Isolated Shells | A Pectinidan, member of the family Pectinidae | Cardium specimens |
| Camptonectes | C. sp. | Bou Dahar "Carriere" | Isolated Shells | A Cardiidan, member of the family Cardiidae | |
| Chlamys | C. aff.uhligi | Bou Dahar Sup | Isolated Shells | A Pectinidan, member of the family Pectinidae | |
| Cochlearites | C. loppianus | Bou Dahar "Carriere" SE Azou N Aït Oufella Foum Kheneg Tizi N´Oumzour | Isolated Shells | An Ostreidan, member of the family Plicatostylidae | Cochlearites |
| Fimbria | F. (Corbis) alveolus F. (Corbis) cochlearis F. (Corbis) trulla F. (Corbis) sp. | Bou Dahar N1 Bou Dahar N3 Bou Dahar N4 Bou Dahar N12 Bou Dahar "Carriere" | Isolated Shells | A Lucinidan, member of the family Lucinidae. | Fimbria specimens (from Hawaii) |
| Gervillia | G. (Gervilleia) spinicosta G. (Gervilleia) sp. | Bou Dahar "Carriere" Toutia Foum Kheneg | Isolated Shells | An Ostreidan, member of the family Bakevelliidae | |
| Gervilleioperna | G. atlantis G. buchi G. termieri G. timorensis G. sp. | Bou Dahar "Carriere" SE Azou N Aït Oufella Foum Kheneg Tizi N´Oumzour | Isolated Shells | An Ostreidan, member of the family Plicatostylidae | Gervillioperna |
| Lima | L. (Plagiostoma) eucharis Lima (Limea) aff.juliana L. (Ctenostreon) cf.raricostata L. (Ctenostreon) sp. | Bou Dahar N3 Bou Dahar "Carriere" | Isolated Shells | A Limidan, member of the family Limidae | Lima specimens |
| Lithioperna | L. (Lithiopedalion) kuehni L. scutata L. spp. | Bou Dahar "Carriere" SE Azou N Aït Oufella Foum Kheneg Tizi N´Oumzour | Isolated Shells | An Ostreidan, member of the family Plicatostylidae | Lithioperna |
| Lucina | L. (Phacoides) lunulicava L. (Phacoides) laevis L. (Phacoides) cf.zonaria L. (Phacoides) sp. nov | Bou Dahar N2 Bou Dahar N4 Bou Dahar N7 Bou Dahar "Carriere" Aguelmane-Azougar | Isolated Shells | A Lucinidan, member of the family Lucinidae. | Lucina specimens |
| Isoarca | I. ssp. | Bou Dahar | Isolated Shells | A Nuculanidan, member of the family Isoarcidae | |
| Isocyprina | I. pulla | Bou Dahar N3 Bou Dahar N7 Bou Dahar "Carriere" | Isolated Shells | A Veneridan, member of the family Isocyprinidae | |
| Myoconcha | M. linguaeformis M. (Daharina) gentili M. aff.corniculun | Bou Dahar N3 Bou Dahar N7 Bou Dahar "Carriere" | Isolated Shells | An Imparidentian, member of the family Kalenteridae | |
| Mytilus | M. maurus M. aff.furcatus M. (Pachymytilus) superbus | Bou Dahar N6 Bou Dahar N7 Bou Dahar N12 Bou Dahar "Carriere" | Isolated Shells | A Mytilidan, member of the family Mytilidae | Mytilus specimens |
| Opis | O. alata O. caudata O. bicristata | Bou Dahar Aguelmane-Azougar | Isolated Shells | A Carditidan, member of the family Astartidae. | |
| Opisoma | O. excavatum O. menchikoffi O. curvidens O. scalprum O. rex O. bourcarti O. spp. | Bou Dahar N2 Bou Dahar N3 Bou Dahar N6 Bou Dahar N7 Bou Dahar N8 Bou Dahar N12 Bou Dahar "Carriere" Khorchef | Isolated Shells | A Carditidan, member of the family Astartidae. Is considered a genus that evolved from shallow burrowing ancestors, becoming a secondarily semi-infaunal edgewise recliner adapted to photosymbiosis. | |
| Pachyerisma | P. opimum | Bou Dahar N3 Bou Dahar N4 Bou Dahar N7 | Isolated Shells | A Megalodontidan, member of the family Megalodontidae | |
| Pachymegalodon | P. timorense | Bou Dahar "Carriere" | Isolated Shells | A Megalodontidan, member of the family Megalodontidae | |
| Parallelodon | P. (Beushausenia) daharensis P. (Beushausenia) sp. | Bou Dahar N1 Bou Dahar N3 Bou Dahar N4 Bou Dahar "Carriere" | Isolated Shells | An Arcidan, member of the family Parallelodontidae | |
| Perna | P. scutata P. sp. nov P. sp. | Bou Dahar N6 Bou Dahar N7 Bou Dahar N12 Bou Dahar "Carriere" | Isolated Shells | An Ostreidan, member of the family Pteriidae | Perna specimens |
| Pecten | P. (Aequipecten) julianus P. (Aequipecten) norigliensis P. aff.bersaskensis P. spp. | Bou Dahar N1 Bou Dahar N3 Bou Dahar N12 Bou Dahar "Carriere" Foum Kheneg | Isolated Shells | A Pectinidan, member of the family Pectinidae | Pecten specimens |
| Plicatula | P. daharensis | Bou Dahar N3 Bou Dahar N6 Bou Dahar N7 Bou Dahar "Carriere" | Isolated Shells | A Pectinidan, member of the family Plicatulidae | Plicatula specimens |
| Pinna | P. (Pinnigena) sp. | Bou Dahar | Isolated Shells | An Ostreidan, member of the family Pinnidae | Pinna specimens |
| Pseudotrapezium | P. spp. | Bou Dahar N7 | Isolated Shells | A Veneridan, member of the family Trapezidae | |
| Spondylus | S. numidus | Bou Dahar Sup | Isolated Shells | A Pectinidan, member of the family Spondylidae | Spondylus specimens |

### Ammonites

| Genus | Species | Stratigraphic position | Material | Notes | Images |
| Apoderoceras | A. davoei A. sp. | Bou Dahar N1 | Isolated Shells | An Ammonite, member of the family Coeloceratidae | Apoderoceras specimen |
| Arieticeras | A. algovianum A. bertrandi | Jebel Bou Rharraf | Isolated Shells | An Ammonite, member of the family Hildoceratidae | |
| Calaiceras | C. calais | Jebel Bou Rharraf | Isolated Shells | An Ammonite, member of the family Phylloceratidae | |
| Dactylioceras | D. tenuicostatum | Bou Dahar N1 | Isolated Shells | An Ammonite, member of the family Dactylioceratidae | Dactylioceras reconstruction |
| Dubariceras | D. dubari D. spp. | Jebel Bou Rharraf | Isolated Shells | An Ammonite, member of the family Eoderoceratidae | |
| Emaciaticeras | E. archimedis E. emaciatum E. fervidum | Jebel Bou Rharraf | Isolated Shells | An Ammonite, member of the family Hildoceratidae | |
| Fuciniceras | F. (Paltarpites) aequiondulatum F. balatonense F. bonarellii F. costicillatum F. cornacaldense F. dilectum F. isseli F. lavinianum F. mellahense F. volubile F. wiedenmayeri F. sp. | Jebel Bou Rharraf Bou Dahar | Isolated Shells | An Ammonite, member of the family Hildoceratidae | |
| Galaticeras | G. aegoceroides G. subtriangulare G. harpoceratoides G. aff. harpoceratoides G. spp. | Jebel Bou Rharraf Bou Dahar | Isolated Shells | An Ammonite, member of the family Lytoceratidae | |
| Hildoceratoides | H. pectinatum | Bou Dahar N7 | Isolated Shells | An Ammonite, member of the family Hildoceratidae | |
| Juraphyllites | J. libertus J. limatus J. (Harpophylloceras) eximius J. sp. | Jebel Bou Rharraf Bou Dahar | Isolated Shells | An Ammonite, member of the family Juraphyllitidae | |
| Leptaleoceras | L. accuratum L. insigne | Jebel Bou Rharraf | Isolated Shells | An Ammonite, member of the family Hildoceratidae | |
| Liparoceras | L. (Becheiceras) bechei | Jebel Bou Rharraf | Isolated Shells | An Ammonite, member of the family Liparoceratidae | Liparoceras specimen |
| Lytoceras | L. baconicum L. fimbriatum L. fimbriatoides L. furcicrenatum L. ovimontanum L. villae L. spp. | Jebel Bou Rharraf Bou Dahar | Isolated Shells | An Ammonite, member of the family Lytoceratidae | Lytoceras fimbriatum specimen |
| Mauretaniaceras | M. elmii | Jebel Bou Rharraf | Isolated Shells | An Ammonite, member of the family Hildoceratidae | |
| Metaderoceras | M. apertum M. brutum M. gemmellaroi M. cf.gemmellaroi M. pseudomuticum M. spp. | Jebel Bou Rharraf | Isolated Shells | An Ammonite, member of the family Eoderoceratidae | |
| Microderoceras | M.(Eoderoceras) birchiades | Jebel Bou Rharraf | Isolated Shells | An Ammonite, member of the family Eoderoceratidae | Microderoceras specimen |
| Miltoceras | M. bettonii M. involutum M. roseum M. sellae M. taguendoufi M. spp. | Jebel Bou Rharraf Bou Dahar | Isolated Shells | An Ammonite, member of the family Coeloceratidae | |
| Paltechioceras | P. boehmi | Bou Dahar | Isolated Shells | An Ammonite, member of the family Echioceratidae | |
| Partschiceras | P. sp. | Jebel Bou Rharraf | Isolated Shells | An Ammonite, member of the family Phylloceratidae | |
| Phricodoceras | P. bettonii P. cantaluppii | Jebel Bou Rharraf | Isolated Shells | An Ammonite, member of the family Phricodoceratidae | |
| Phylloceras | P. hebertinum | Jebel Bou Rharraf Bou Dahar | Isolated Shells | An Ammonite, member of the family Phylloceratidae | Phylloceras reconstruction |
| Prodactylioceras | P. colubriforme P. italicum | Jebel Bou Rharraf Bou Dahar | Isolated Shells | An Ammonite, member of the family Dactylioceratidae | Prodactylioceras specimen |
| Reynesoceras | R. mortilleti R. ragazzoni | Jebel Bou Rharraf Bou Dahar | Isolated Shells | An Ammonite, member of the family Dactylioceratidae | |
| Reynesocoeloceras | R. fallax R. indunense R. praeincertum R. simulans | Jebel Bou Rharraf Bou Dahar | Isolated Shells | An Ammonite, member of the family Dactylioceratidae | |
| Radstockiceras | R. spp. | Jebel Bou Rharraf | Isolated Shells | An Ammonite, member of the family Oxynoticeratidae | |
| Tauromeniceras | T. mazetieri T. nerinea T. spp. | Jebel Bou Rharraf Bou Dahar | Isolated Shells | An Ammonite, member of the family Hildoceratidae | |
| Tropidoceras | T. demonense T. flandrini T. heterogeneum T. zitteli | Jebel Bou Rharraf Bou Dahar N4 | Isolated Shells | An Ammonite, member of the family Polymorphitidae | |
| Zetoceras | Z. zetes Z. ssp. | Jebel Bou Rharraf | Isolated Shells | An Ammonite, member of the family Phylloceratidae | |

### Annelida

| Genus | Species | Stratigraphic position | Material | Notes | Images |
| Serpula | S. spp. | Bou Dahar | Tubiform Structures | A Serpulidan, member of the family Serpulidae | Serpula specimen |
| Terebella | T. lapilloides T. spp. | Bou Dahar | Tubiform Structures | A Terebellidan, member of the family Terebellidae | Terebella specimen |

### Echinodermata

| Genus | Species | Stratigraphic position | Material | Notes | Images |
| Apiocrinus | A. amalthei | Bou Dahar | Stems | A Crinoid, member of the family Apiocrinidae | |
| Atlasaster | A. termieri | Bou Dahar | Specimens | An Echinoidean, member of the group Irregularia | |
| Diademopsis | D. behtensis | Bou Dahar | Specimens & Isolated spines | An Echinoidean, member of the family Pedinidae | Diademopsis specimen |
| Diplechinus | D. hebbriensis | Bou Dahar | Specimens & Isolated spines | An Echinoidean, member of the family Stomechinidae | |
| Diplocidaris | D. menchikoffi | Bou Dahar | Specimens & Isolated spines | An Echinoidean, member of the family Diplocidaridae | Diplocidaris specimen |
| Firmacidaris | F. precincta | Bou Dahar | Specimens & Isolated spines | An Echinoidean, incertae sedis | |
| Hemicidaris | H. termieri | Bou Dahar | Specimens & Isolated spines | An Echinoidean, member of the family Hemicidaridae | Hemicidaris specimen |
| Microdiadema | M. richeri | Bou Dahar | Specimens & Isolated spines | An Echinoidean, member of the family Pseudodiadematidae | |
| Miocidaris | M. dubari | Bou Dahar | Specimens & Isolated spines | An Echinoidean, member of the family Miocidaridae | |
| Pseudopedina | P. atlantis | Bou Dahar | Specimens & Isolated spines | An Echinoidean, member of the family Pedinidae | |
| Pygaster | P. daguini | Bou Dahar | Specimens & Isolated spines | An Echinoidean, member of the family Pygasteridae | |

### Microproblematic/Algae

| Genus | Species | Location | Material | Notes | Images |
| Aeolissacus | A. spp. | Bou Dahar | Calcified Thalli | A Possible Green Algae of the family Dasycladaceae. | |
| Apophoretella | A. spp. | Bou Dahar | Calcified specimens | A Cyanobacterial Alga of the family Cyanophyceae | |
| Baccanella | B. floriformis | Bou Dahar | Calcified specimens | Incertade sedis | |
| Bacinella | B. irregularis B. ordinata | Bou Dahar | Calcified specimens | A Cyanobacterial Alga of the family Garwoodiaceae | |
| Boueina | B. hochstetteri | Bou Dahar IttoFezzou Ait-Ani | Calcified specimens | A Green Algae of the family Halimedaceae. | |
| Cayeuxia | C. liasica C. spp. | Bou Dahar | Calcified Thalli | A green algae of the Halimedaceae or Udoteaceae family. | Modern Udotea |
| Lithocodium | L. aggregatum | Bou Dahar | Calcified specimens | Incertade sedis | |
| Muranella | M. sphaerica | Bou Dahar | Calcified specimens | Incertade sedis | |
| Neomizzia | N. elongata | Bou Dahar | Calcified specimens | A Green Algae of the family Dasycladaceae. | Considered to be convergent with modern Cymopolia |
| Thaumatoporella | T. parvovesiculifera | Bou Dahar | Calcified Thalli | A Green alga of the Thaumatoporellales group | |
| Palaeodasycladus | P. fragilis P. mediterraneus P. dolomiticus P. anae P. spp. | Bou Dahar | Calcified Thalli | A Green Algae of the family Dasycladaceae. A reefal algae usually found in carbonate settings along all the Mediterranean | |
| Radiomura | R. cautica | Bou Dahar | Calcified specimens | Incertade sedis | |
| Rivularia | R. moesica | Bou Dahar | Calcified specimens | A Cyanobacterial Alga of the family Rivulariaceae | Rivularia specimens |
| Solenopora | S. liasica | Bou Dahar | Calcified specimens | A Red Alga of the family Solenoporaceae | |
| Tersella | T. genotii T. alpina | Bou Dahar | Calcified specimens | A Green Algae of the family Dasycladaceae. | |
| Toulaina | T. liassica | Bou Dahar IttoFezzou Ait-Ani | Calcified specimens | A Green Algae of the family Halimedaceae. | |