# List of Amphidromus species

There are over 200 species within the land snail genus Amphidromus. 95 = 75 by Laidlaw and Solem (1961) + 16 new + 2 new species by Sutcharit et al. 2015 + 2 new species by Inkhavilay et al. 2017
These species used to be divided into three subgenera:, They are all accepted as an alternate representation
* subgenus Amphidromus Albers, 1850
* subgenus Goniodromus Bülow, 1905
* subgenus Syndromus Pilsbry, 1900
* Amphidromus abbasi Chan & Tan, 2010
* Amphidromus abbasorum Thach, 2017
* Amphidromus abbotthuberorum Thach, 2017
* Amphidromus adamsii (Reeve, 1848)
* Amphidromus agnieszkae Thach, 2020
* Amphidromus alfi Thach, 2020
* Amphidromus alicetandiasae J. Parsons, 2016
* Amphidromus alicjaboronae Thach, 2021
* Amphidromus alticola Fulton, 1896
* Amphidromus ameliae Dharma, 2007
* Amphidromus andamanicus (Hanley & Theobald, 1876)
* Amphidromus angulatus Fulton, 1896
* Amphidromus anhdaoorum
* Amphidromus anhduongae Thach, 2020
* Amphidromus annae E. von Martens, 1891
* Amphidromus anthonyabbotti Thach & F. Huber, 2017
* Amphidromus areolatus (L. Pfeiffer, 1861)
* Amphidromus asperoides Jirapatrasilp & C.-T. Lee, 2024
* Amphidromus atricallosus (Gould, 1843)
** Amphidromus atricallosus classiarius Sutcharit & Panha, 2006
** Amphidromus atricallosus leucoxanthus (von Martens, 1864)
** Amphidromus atricallosus perakensis Fulton, 1901
* Amphidromus aureocinctus Fulton, 1896
* Amphidromus backeljaui Thach, 2020
* Amphidromus baili Thach, 2020
* Amphidromus banksi Butot, 1955
* Amphidromus baoi Thach, 2017
* Amphidromus baolocensis Thach & F. Huber, 2016
* Amphidromus basilanensis Bartsch, 1917
* Amphidromus bataviae (Grateloup, 1840)
* Amphidromus beccarii (Tapparone Canefri, 1883)
* Amphidromus begini (Morlet, 1886)
* Amphidromus bernardfamyi Thach, 2017
* Amphidromus berschaueri Thach, 2018 (taxon inquirendum, debated synonym)
** Amphidromus berschaueri mingi Thach, 2019
* Amphidromus binhduongensis Thach & F. Huber, 2021
* Amphidromus binhphuocensis Thach, 2019
* Amphidromus boroni Thach, 2020
* Amphidromus bozhii Y.C. Wang, 2019
* Amphidromus bramvanderbijli Thach, 2019
* Amphidromus buelowi Fruhstorfer, 1905
* Amphidromus buluanensis Bartsch, 1917
* Amphidromus calista Pilsbry, 1900: synonym of Amphidromus chloris (Reeve, 1848) accepted as Amphidromus sulphuratus (Hombron & Jacquinot, 1847) (junior subjective synonym)
* Amphidromus cambojiensis (Reeve, 1860)
* Amphidromus capistratus E. von Martens, 1903
* Amphidromus centrocelebensis Bollinger, 1918
* Amphidromus cochinchinensis (L. Pfeiffer, 1857)
* Amphidromus coeruleus Clench & Archer, 1932
* Amphidromus cognatus Fulton, 1907
* Amphidromus columellaris Möllendorff, 1892
* Amphidromus comes (L. Pfeiffer, 1861)
* Amphidromus consobrinus Fulton, 1897
* Amphidromus contrarius (O. F. Müller, 1774)
* Amphidromus cossignanii Thach, 2021
* Amphidromus costiferE. A. Smith, 1893
* Amphidromus crassus Fulton, 1899
* Amphidromus cruentatus
* Amphidromus dancei Dharma, 2021
* Amphidromus daoae Thach, 2016
* Amphidromus dautzenbergi Fulton, 1899
* Amphidromus davidberschaueri Thach, 2021
* Amphidromus dekkeri Thach, 2021
* Amphidromus delsaerdti Thach, 2016
* Amphidromus djajasasmitai Dharma, 1993
* Amphidromus dohrni (L. Pfeiffer, 1864)
* Amphidromus donchani Thach, 2019
* Amphidromus dubius Fulton, 1896
* Amphidromus duboismercesylviae Thach & F. Huber, 2020
* Amphidromus dungae Thach, 2021
* Amphidromus eichhorsti Thach, 2020: synonym of Amphidromus cruentatus (Morelet, 1875) (junior subjective synonym)
* Amphidromus elviae Dharma, 2007
* Amphidromus elvinae Dharma, 2007
* Amphidromus enganoensis Fulton, 1896
* Amphidromus entobaptus Dohrn, 1889
* Amphidromus epidemiae Y.-C. Wang, 2021
* Amphidromus escondidus Poppe, 2020
* Amphidromus everetti Fulton, 1896
* Amphidromus feliciae Thach & Abbas, 2020
* Amphidromus felixi Dharma, 2021
* Amphidromus fengae Y.C. Wang, 2019
* Amphidromus filozonatus (E. von Martens, 1867)
* Amphidromus flavus (L. Pfeiffer, 1861)
* Amphidromus floresianus Fulton, 1897
* Amphidromus frednaggsi Thach & F. Huber, 2018
* Amphidromus friedahuberae Thach & F. Huber, 2017
* Amphidromus fultoni Ancey, 1897
* Amphidromus furcillatus (Mousson, 1849)
* Amphidromus gattingeri Thach & F. Huber, 2020
* Amphidromus gisellelieae Thach & Abbas, 2022
* Amphidromus givenchyi Geret, 1912
* Amphidromus glaucolarynx (Dohrn, 1861)
* Amphidromus globonevilli Sutcharit & Panha, 2015
* Amphidromus grohi Thach, 2021
* Amphidromus gustafi Thach & F. Huber, 2020
* Amphidromus haematostoma Möllendorff, 1898
* Amphidromus hamatus Fulton, 1896
* Amphidromus harryleei Thach, 2020
* Amphidromus haszprunari Thach, 2020
* Amphidromus heerianus (L. Pfeiffer, 1871)
* Amphidromus heinrichhuberi Thach & F. Huber, 2016
* Amphidromus hejingi Thach, 2019
* Amphidromus herosae Thach, 2021
* Amphidromus hildagoi Bartsch, 1917
* Amphidromus hongdaoae Thach, 2017
* Amphidromus huanganhi Thach, 2020
* Amphidromus huberi Thach, 2014
* Amphidromus huynhanhi Thach, 2019
* Amphidromus huynhi Thach, 2019
* Amphidromus ilsa B. Rensch, 1933
* Amphidromus inconstans Fulton, 1898
* Amphidromus inflatus Fulton, 1896
* Amphidromus ingens Möllendorff, 1900
* Amphidromus ingensoides Jirapatrasilp & C.-T. Lee, 2024
* Amphidromus inversus (Müller, 1774)
** Amphidromus inversus albulus Sutcharit & Panha, 2006
** Amphidromus inversus annamiticus (Crosse & Fischer, 1863)
** Amphidromus inversus inversus Müller, 1774
* Amphidromus iunior Cilia, 2013
* Amphidromus jacobsoni Laidlaw, 1954
* Amphidromus javanicus
* Amphidromus jadeni Thach, 2021
* Amphidromus janus (L. Pfeiffer, 1854)
* Amphidromus javanicus (G. B. Sowerby I, 1833)
* Amphidromus jeffabbasorum Thach, 2016
* Amphidromus johnstanisici Thach & Huber, 2017
* Amphidromus jomi Dumrongrojwattana, Wongkamhaeng & Tanamai, 2019
* Amphidromus jonabletti Thach, 2019
* Amphidromus juliekeppensae Thach, 2021
* Amphidromus kalaoensis Fulton, 1896
* Amphidromus kantori Thach & F. Huber, 2020
* Amphidromus keppensdhondtorum Thach, 2018
* Amphidromus khammouanensis Thach & F. Huber, 2017
* Amphidromus khoatuanorum Thach, 2021
* Amphidromus kiati Thach, 2020
* Amphidromus koenigi Thach & F. Huber, 2018
* Amphidromus kiati Thach, 2020
* Amphidromus koonpoi
* Amphidromus laii Thach, 2019
* Amphidromus laosianus Bavay, 1898
* Amphidromus latestrigatus M.M. Schepman, 1892
* Amphidromus ledaoae Thach, 2016
* Amphidromus leeanus Thach, 2021
* Amphidromus leucoxanthus (E. von Martens, 1864)
* Amphidromus liei Thach, 2017
** Amphidromus liei joshuathami Thach, 2018
* Amphidromus lilianaboronae Thach, 2021
* Amphidromus loanphungae Thach, 2020
* Amphidromus lucsegersi Thach & Abbas, 2017
* Amphidromus madelineae Thach, 2020
* Amphidromus marekboroni Thach, 2021
* Amphidromus mariaeThach & F. Huber, 2017
* Amphidromus mariasendersae Thach & F. Huber, 2017
* Amphidromus marki Thach, 2021
* Amphidromus markpankowskii Thach, 2020
* Amphidromus martensi Boettger, 1894
* Amphidromus maryae Thach, 2021
* Amphidromus masoni (Godwin-Austen, 1876)
* Amphidromus maxi Thach, 2020
* Amphidromus michaeli Thach, 2021
* Amphidromus mingmini Thach, 2019
* Amphidromus minhthaoae Thach, 2020
* Amphidromus mirandus Bavay & Dautzenberg, 1912
* Amphidromus monsecourorum Thach & F. Huber, 2017
* Amphidromus ngai Thach, 2019
* Amphidromus nganguyeni Thach, 2021
* Amphidromus ngocae Thach, 2021
* Amphidromus ngocanhi Thach, 2017
* Amphidromus nguyenkhoai Thach, 2020
* Amphidromus nguyetminhae Thach, 2020
* Amphidromus niasensis Fulton, 1907
* Amphidromus nicoasiarum Thach, 2021
* Amphidromus nicoi Thach, 2017
* Amphidromus noriokowasoei Thach & F. Huber, 2017
* Amphidromus palaceus (Mousson, 1848)
* Amphidromus pankowskiae Thach, 2020
* Amphidromus pankowskianus Thach, 2020
* Amphidromus pankowskiorum Thach, 2021
* Amphidromus patbaili Thach & F. Huber, 2021
* Amphidromus pattinsonae Iredale, 1943
* Amphidromus perrieri A. de Rochebrune, 1882
* Amphidromus persimilis J. Parsons, 2019
* Amphidromus pervariabilis Bavay & Dautzenberg, 1909
* Amphidromus perversus (Linnaeus, 1758) - the type species of the genus Amphidromus
** Amphidromus perversus butoti
** Amphidromus perversus emaciatus
** Amphidromus perversus natunensis Fulton, 1896
* Amphidromus petestimpsoni Thach, 2021
* Amphidromus phamanhi Thach, 2016
* Amphidromus phamanhorum Thach, 2021
* Amphidromus phamtuanhae Thach, 2021
* Amphidromus phamvutuanhae Thach, 2022
* Amphidromus philippeboucheti Thach, 2019
* Amphidromus poppei Thach, 2020
* Amphidromus protania Lehmann & Maassen, 2004
* Amphidromus qiongensis J. He & Q.-H. Zhou, 2017
* * Amphidromus reflexilabris Schepman, 1892
* Amphidromus quangtrungi Thach, 2021
* Amphidromus reflexilabris Schepman, 1892
* Amphidromus reuselaarsi Thach, 2018
* Amphidromus roseolabiatus Fulton, 1896
* Amphidromus schomburgki (Pfeiffer, 1860)
** Amphidromus schomburgki dextrochlorus Sutcharit & Panha, 2006
* Amphidromus sandersae Thach & F. Huber, 2020
* Amphidromus sekincauensis
* Amphidromus similis Pilsbry, 1900
* Amphidromus simonei Thach, 2020
* Amphidromus siongkiati Thach, 2019
* Amphidromus sowyani Thach, 2019
* Amphidromus sriabbasae Thach, 2017
* Amphidromus stanyi Dharma, 2021
* Amphidromus stevenabbasorum Thach, 2021
* Amphidromus sylviae Stark, 2017
* Amphidromus syndromoideus Inkhavilay & Panha, 2017
* Amphidromus szekeresi Thach, 2020
* Amphidromus thachi
* Amphidromus thachorum F. Huber, 2020
* Amphidromus thaitieni Thach & F. Huber, 2020
* Amphidromus thomasi Thach, 2021
* Amphidromus thuthaoae Thach, 2021
* Amphidromus timorensis Parsons & Abbas, 2020
* Amphidromus trianensis Thach & F. Huber, 2018
* Amphidromus truongi Thach, 2021
* Amphidromus tuanhae Thach, 2021
* Amphidromus tuani Thach, 2021
* Amphidromus tureki Thach, 2021
* † Amphidromus ubaldii Dharma, 2021
* Amphidromus yauyeejiae Thach & Abbas, 2017
* Amphidromus verbinneni Segers, 2020
* Amphidromus vincekessneri Thach, 2020
* Amphidromus walleri Thach, 2020
* Amphidromus yauyeejiae Thach & Abbas, 2017
* Amphidromus zebrinus (L. Pfeiffer, 1861)
* Amphidromus zelosus Y.-C. Wang & Z.-Y. Chen, 2021
* Amphidromus atricallosus perakensis
* Amphidromus givenchyi
* Amphidromus inversus
* Amphidromus roseolabiatus
* Amphidromus syndromoideus

## Subgenus Syndromus

subgenus Syndromus Pilsbry, 1900
All species of the subgenus Syndromus are sinistral with two exceptions: amphidromine Amphidromus glaucolarynx and dextral Amphidromus kuehni.
* Amphidromus adamsii (Reeve, 1848)
** Amphidromus adamsii articulata Fulton, 1896
** Amphidromus adamsii duplocincta Fulton, 1896
** Amphidromus adamsii inornata Fulton, 1896
** Amphidromus adamsii luteofasciata Fulton, 1896
** Amphidromus adamsii ornata Fulton, 1896
** Amphidromus adamsii rubiginosa Fulton, 1896
** Amphidromus adamsii rufocincta Fulton, 1896
** Amphidromus adamsii simplex Fulton, 1896
** Amphidromus adamsii subunicolor Fulton, 1896
** Amphidromus adamsii superba Fulton, 1896
* Amphidromus alicetandiasae Parsons, 2016
* Amphidromus areolatus (Pfeiffer, 1861)
* Amphidromus coeruleus Clench & Archer, 1932
* Amphidromus contrarius (Müller, 1774) - the type species of the subgenus Syndromus
** Amphidromus contrarius maculata Fulton, 1896
** Amphidromus contrarius multifasciata Fulton, 1896
** Amphidromus contrarius rolfei Thach, 2018
* Amphidromus flavus (Pfeiffer, 1861)
** Amphidromus flavus proxima Fulton, 1896
* Amphidromus fuscolabris Möllendorff, 1898
* Amphidromus floresianus Fulton, 1897
* Amphidromus glaucolarynx (Dohrn, 1861) - This is the only one amphidromine (left-handed and right-handed snails occur in the population) species in the subgenus Syndromus.
* Amphidromus globonevilli Sutcharit & Panha, 2015
* Amphidromus haematostoma Möllendorff, 1898
* Amphidromus iunior Cilia, 2013
* Amphidromus kuehni - This is the only dextral species in the subgenus Syndromus.
* Amphidromus laevus (Müller, 1774)
** Amphidromus laevus romaensis Rolle, 1903
* Amphidromus pictus Fulton, 1896
* Amphidromus poecilochrous Fulton, 1896 / Amphidromus poecilochroa Fulton, 1896
* Amphidromus porcellanus (Mousson, 1848)
* Amphidromus principalis Sutcharit & Panha, 2015
* Amphidromus richardi Severns, 2006
** Amphidromus richardi manacocoensis Severns, 2006
* Amphidromus roemeri (Pfeiffer, 1863)
* Amphidromus rottiensis Chan & Tan, 2010
* Amphidromus semitessellatus (Morlet, 1884)
* Amphidromus stevenliei Parsons, 2016
* Amphidromus sumatranus (Martens, 1864)
* Amphidromus xiengensis Morlet, 1891
* Amphidromus xiengkhaungensis Inkhavilay & Panha, 2017
* Amphidromus sp. - an undescribed species from Lampung, Indonesia
* Amphidromus adamsii
* Amphidromus areolatus
* Amphidromus flavus
* Amphidromus fuscolabris
* Amphidromus globonevilli
* Amphidromus principalis
* Amphidromus xiengensis

## Subgenus Goniodromus

* Amphidromus buelowi Fruhstorfer, 1905
** Amphidromus buelowi malalakensis Parsons & Abbas, 2016
subgenus Incertae sedis
* Amphidromus cognatus Fulton, 1907
subgenus ?
* Amphidromus alticola Fulton, 1896
* Amphidromus andamanicus
** Amphidromus andamanicus nicobarica Godwin-Austen, 1895
* Amphidromus angulatus Fulton, 1896
* Amphidromus annamiticus
** Amphidromus annamiticus roseotincta Möllendorff, 1894
* Amphidromus asper Haas, 1934
* Amphidromus aureocincta Fulton, 1896
* Amphidromus banksi Butot, 1955
* Amphidromus baolocensis Thach & F. Huber, 2016
* Amphidromus bataviae (Grateloup, 1840)
* Amphidromus begini (Morlet, 1886)
* Amphidromus cambojiensis (Reeve, 1860)
* Amphidromus candidus
* Amphidromus chloris Reeve, 1848
* Amphidromus cochinchinensis (Pfeiffer, 1857)
* Amphidromus columellaris
** Amphidromus columellaris gloriosa Fulton, 1896
* Amphidromus comes (Pfeiffer, 1861)
* Amphidromus concinna Fulton, 1896
* Amphidromus consobrinus Fulton, 1897
* Amphidromus contusus (Reeve, 1848)
* Amphidromus costifer Smith, 1893
* Amphidromus crassa Fulton, 1899
* Amphidromus cruentatus (Morelet, 1875)
* Amphidromus dambriensis Thach & F. Huber, 2016
* Amphidromus daoae Thach, 2016
* Amphidromus dautzenbergi Fulton, 1899
* Amphidromus delsaerdti Thach, 2016
* Amphidromus djajasasmitai
* Amphidromus dohrni (Pfeiffer, 1864)
* Amphidromus dubius Fulton, 1896
* Amphidromus enganoensis Fulton, 1896
* Amphidromus entobaptus
* Amphidromus eques (Pfeiffer, 1857)
* Amphidromus everetti Fulton, 1896
** Amphidromus everetti connectens Fulton, 1896
* Amphidromus filozonatus
** Amphidromus filozonatus jucunda Fulton, 1896
* Amphidromus friedae Thach & F. Huber, 2016
* Amphidromus hamatus Fulton, 1896
* Amphidromus heerianus
** Amphidromus heerianus poecillus
* Amphidromus heinrichhuberi Thach & F. Huber, 2016
* Amphidromus hongdaoae Thach, 2017
* Amphidromus hosei Smith, 1895
* Amphidromus hueae Thach & F. Huber, 2016
* Amphidromus inconstans Fulton, 1898
* Amphidromus janus (Pfeiffer, 1854)
* Amphidromus kalaoensis Fulton, 1896
* Amphidromus kruijti
* Amphidromus lepidus (Gould, 1856)
* Amphidromus lindstedti (Pfeiffer, 1857)
* Amphidromus loricatus (Pfeiffer, 1855)
* Amphidromus lamdongensis Thach & F. Huber, 2016
* Amphidromus ledaoae Thach, 2016
* Amphidromus luangensis Gra-tes, 2015
* Amphidromus maculatus
* Amphidromus maculiferus
** Amphidromus maculiferus gracilior Fulton, 1896
** Amphidromus maculiferus inflata Fulton, 1896
** Amphidromus maculiferus obscura Fulton, 1896
* Amphidromus masoni (Godwin-Austen, 1876)
* Amphidromus melanomma (Pfeiffer, 1852)
* Amphidromus metabletus
* Amphidromus moniliferus (Gould, 1846)
* Amphidromus mouhoti (Pfeiffer, 1861)
* Amphidromus niasensis Fulton, 1907
** Amphidromus niasensis f. sowerbyi
* Amphidromus perakensis Fulton, 1901
** Amphidromus perakensis globosus Fulton, 1903
* Amphidromus phamanhi Thach, 2016
* Amphidromus placidus Fulton, 1896
* Amphidromus psephos Vermeulen, Liew & Schilthuizen, 2015
* Amphidromus quadrasi
** Amphidromus quadrasi solida Fulton, 1896
* Amphidromus reflexilabris
* Amphidromus robustus Fulton, 1896
* Amphidromus schileykoi Thach, 2016
* Amphidromus sinensis (Benson, 1851)
** Amphidromus sinensis gracilis Fulton, 1896
** Amphidromus sinensis vicaria Fulton, 1896
* Amphidromus singalangensis Rolle, 1908
* Amphidromus sinistralis (Reeve, 1849)
** Amphidromus sinistralis lutea Fulton, 1896
** Amphidromus sinistralis rosea Fulton, 1896
* Amphidromus smithii Fulton, 1896 or Amphidromus smithi Fulton, 1896
* Amphidromus sowerbyi Fulton, 1907
* Amphidromus sumbaensis Fulton, 1896
* Amphidromus suspectus
** Amphidromus suspectus albolabiata Fulton, 1896
* Amphidromus taluensis Gra-tes, 2015
** Amphidromus taluensis borealis Gra-tes, 2015
* Amphidromus thanhhoaensis Thach & F. Huber, 2016
* Amphidromus theobaldianus (Benson, 1857)
* Amphidromus ventrosulus Möllendorff, 1900
* Amphidromus versicolor Fulton, 1896
* Amphidromus webbi Fulton, 1907
** Amphidromus webbi babiensis Laidlaw, 1954
** Amphidromus webbi simalurensis Laidlaw, 1954
* Amphidromus winteri
** Amphidromus winteri inauris Fulton, 1896
* Amphidromus zebrinus (Pfeiffer, 1861)
Other
* Amphidromus sinensis globosa Nevill, 1878 - nomen nudum