# Gnaphalieae

The Gnaphalieae, commonly known as paper daisies, are a tribe of flowering plants in the family Asteraceae. It is most closely related to the tribes Anthemideae, Astereae, and Calenduleae.

## Characteristics

This group is most diverse in South America, Southern Africa and Australia. There are only a few genera with species native to temperate regions: Anaphalis, Antennaria, Gamochaeta, Helichrysum, Leontopodium (Edelweiss), Phagnalon, Diaperia, and Pseudognaphalium.
The classification of the tribe into subtribes is unclear, with a number of past classifications not being supported by late 20th-century evidence.

## Genera

Gnaphalieae genera recognized by the Global Compositae Database as April 2022:
* × Filfia Holub
* Acanthocladium F.Muell.
* Achyrocline (Less.) DC.
* Acomis F.Muell.
* Actinobole Fenzl ex Endl.
* Alatoseta Compton
* Ammobium R.Br.
* Amphiglossa DC.
* Anaphalioides (Benth.) Kirp.
* Anaphalis DC.
* Anaxeton Gaertn.
* Ancistrocarphus A.Gray
* Anderbergia B.Nord.
* Anemocarpa Paul G.Wilson
* Angianthus J.C.Wendl.
* Antennaria Gaertn.
* Apalochlamys Cass.
* Argentipallium Paul G.Wilson
* Argyroglottis Turcz.
* Argyrotegium J.M.Ward & Breitw.
* Arrowsmithia DC.
* Asteridea Lindl.
* Athrixia Ker Gawl.
* Atrichantha Hilliard & B.L.Burtt
* Basedowia E.Pritz.
* Bellida Ewart
* Belloa J.Rémy
* Berroa Beauverd
* Blennospora A.Gray
* Bombycilaena (DC.) Smoljan.
* Bryomorphe Harv.
* Calocephalus R.Br.
* Calomeria Vent.
* Calotesta P.O.Karis
* Cassinia R.Br.
* Castroviejoa Galbany, L.Sáez & Benedí
* Catatia Humbert
* Cephalipterum A.Gray
* Cephalosorus A.Gray
* Chamaepus Wagenitz
* Chevreulia Cass.
* Chiliocephalum Benth.
* Chionolaena DC.
* Chondropyxis D.A.Cooke
* Chryselium Urtubey & S. E. Freire
* Chrysocephalum Walp.
* Chthonocephalus Steetz
* Cladochaeta DC.
* Coronidium Paul G. Wilson
* Craspedia G.Forst.
* Cremnothamnus Puttock
* Cuatrecasasiella H.Rob.
* Dasyanthus Bubani
* Decazesia F.Muell.
* Diaperia Nutt.
* Dicerothamnus Koek.
* Dielitzia P.S.Short
* Disparago Gaertn.
* Dithyrostegia A.Gray
* Dolichothrix Hilliard & B.L.Burtt
* Edmondia Cass.
* Elytropappus Cass.
* Epitriche Turcz.
* Eriochlamys Sond. & F.Muell.
* Erymophyllum Paul G.Wilson
* Euchiton Cass.
* Ewartia Beauverd
* Ewartiothamnus Anderb.
* Facelis Cass.
* Feldstonia P.S.Short
* Filago Loefl.
* Fitzwillia P.S.Short
* Fluminaria N.G.Bergh
* Galeomma Rauschert
* Gamochaeta Wedd.
* Gilberta Turcz.
* Gilruthia Ewart
* Gnaphaliothamnus Kirp.
* Gnaphalium L.
* Gnephosis Cass.
* Gnomophalium Greuter
* Gratwickia F.Muell.
* Haeckeria F.Muell.
* Haegiela P.S.Short & Paul G.Wilson
* Haptotrichion Paul G.Wilson
* Helichrysopsis Kirp.
* Helichrysum Mill.
* Hesperevax (A.Gray) A.Gray
* Humeocline Anderb.
* Hyalochlamys A.Gray
* Hyalosperma Steetz
* Hydroidea P.O.Karis
* Ifloga Cass.
* Ixiolaena Benth.
* Ixodia R.Br.
* Jalcophila M.O.Dillon & Sagást.
* Lachnospermum Willd.
* Langebergia Anderb.
* Laphangium (Hilliard & B.L.Burtt) Tzvelev
* Lasiopogon Cass.
* Lawrencella Lindl.
* Leiocarpa Paul G.Wilson
* Lemooria P.S.Short
* Leontopodium (Pers.) R.Br. ex Cass.
* Lepidostephium Oliv.
* Leptorhynchos Less.
* Leucochrysum (A.Cunn. ex DC.) Paul G.Wilson
* Leucogenes Beauverd
* Leucophyta R.Br.
* Leysera L.
* Logfia Cass.
* Loricaria Wedd.
* Lucilia Cass.
* Metalasia R.Br.
* Mexerion G.L.Nesom
* Micropsis DC.
* Micropus L.
* Millotia Cass.
* Mniodes (A.Gray) Benth. & Hook.f.
* Myriocephalus Benth.
* Myrovernix Koek.
* Neotysonia Dalla Torre & Harms
* Nestlera Spreng.
* Odixia Orchard
* Oedera L.
* Ozothamnus R.Br.
* Paenula Orchard
* Parantennaria Beauverd
* Pentatrichia Klatt
* Petalacte D.Don
* Phaenocoma D.Don
* Phagnalon Cass.
* Pithocarpa Lindl.
* Plecostachys Hilliard & B.L.Burtt
* Podolepis Labill.
* Podotheca Cass.
* Pogonolepis Steetz
* Polycalymma F.Muell. & Sond.
* Pseudognaphalium Kirp.
* Psilocarphus Nutt.
* Pterochaeta Steetz
* Pterygopappus Hook.f.
* Pycnosorus Benth.
* Quinetia Cass.
* Quinqueremulus Paul G.Wilson
* Rachelia J.M.Ward & Breitw.
* Raoulia Hook.f. ex Raoul
* Raouliopsis S.F.Blake
* Rhetinocarpha Paul G.Wilson & M.A.Wilson
* Rhodanthe Lindl.
* Rhynchopsidium DC.
* Rutidosis DC.
* Schoenia Steetz
* Scyphocoronis A.Gray
* Seriphium L.
* Siemssenia Steetz
* Siloxerus Labill.
* Sinoleontopodium Y.L.Chen
* Sondottia P.S.Short
* Stenocline DC.
* Stenophalium Anderb.
* Stoebe L.
* Stuartina Sond.
* Stylocline Nutt.
* Swammerdamia DC.
* Syncarpha DC.
* Syncephalum DC.
* Taplinia Lander
* Tenrhynea Hilliard & B.L.Burtt
* Thiseltonia Hemsl.
* Tietkensia P.S.Short
* Toxanthes Turcz.
* Trichanthodium Sond. & F.Muell.
* Triptilodiscus Turcz.
* Troglophyton Hilliard & B.L.Burtt
* Vellereophyton Hilliard & B.L.Burtt
* Waitzia J.C.Wendl.
* Xerochrysum Tzvelev

### Other genera

* Callilepis (may not belong in this tribe)
* Libinhania
* Omalotheca
* Oxylaena
* Planea