# Drzewica Formation

The Drzewica Formation (also called Drzewica Series, Polish: seria drzewicka gen. serii drzewickiej and in older literature Bronów Series, Polish: seria bronowska or Komorowo Beds, Polish: warstwy komorowskie) is a geologic formation in Szydłowiec, Poland. It is Pliensbachian in age. Vertebrate fossils have been uncovered from this formation, including dinosaur tracks. The Drzewica Formation is part of the Depositional sequence IV-VII of the late lower Jurassic Polish Basin, with the IV showing the presence of local Alluvial deposits, with possible meandriform deposition origin, dominated in Jagodne and Szydłowiec, while delta system occurred through the zone of the modern Budki. The sequence V shows a reduction of the erosion in the Zychorzyn borehole of the Drzewica Formation, showing changes on the extension of the marine facies, where upper deposits change from Alluvial to Deltaic-Seashore depositional settings. VI-VII facies were recovered on the Brody-Lubienia borehole, with a lower part exposed on the village of Śmiłów that shows a small fall of the Sea level. The stathigraphic setting of the dinosaur tracks reported from the formation suggest a Seashore or Deltaic barrier. Body fossils reported include bivalves, palynology, fossil trunks, roots. Trunks of coniferous wood, especially Cheirolepidiaceae and Araucariaceae trees show the occurrence of vast coniferous forests around the tracksite. The association of forests and dinosaur megafauna on the Pliensbachian suggests also a colder and specially dry ecosystem. Drzewica deposits where in part to be a gigantic shore barrel, setting at the time where the Polish basin sea was at its lowest point. Other related units are Fjerritslev or Gassum Formation (Danish Basin), Hasle & Sorthat Formation (Bornholm), upper Neringa Formation (Lithuania) and abandoned informal units in other regions of Poland: upper Sławęcin Beds (Polish: seria sławęcińska), Wieluń series (Polish: seria wieluńska) or Bronów series.

## Description

The Drzewica Formation between Gowarczów and Rozwady, forms sandstone complexes of +10 or so meters, usually uniformly formed, finely less often medium-grained, separated by Silt-sandstone blocks, sandstone-like places or Siltstones. In these sediments there are worm tubules (Ichnofossils), Rhizoids and the remains of Charred plants, sometimes thin carbon inserts. This section on the area of Gowarczów-Rozwady wasn't completely covered, but connects the appearance of the Drzewica Formation strata with several coeval Boreholes: Gowarczów (GW-1 and GW-2), Kuraszków (KP-1 and KP-4), Sielec (A2), Adamów (Al), Kraszków (MG-2), Gielniów (GG-1) and in a few exposures (quarries) from the vicinity of Kraszków and Gielniów. On this region the most complete profile of this formation was obtained in the Sielec Borehole (A2), which was pierced by the geological structure of the outcrop zone, covering the middle and upper section of the Drzewica Formation with a thickness of 146.7 m. Locally, the thickness of this series is estimated at 175–180 m.

### Profile

Stratigraphy on the Bodzanów IG 1 Borehole :
| Unit | Lithology | Thickness (metres) | Fossil Palynology/Flora | Fossil Fauna |
| Youngest | sandstones | 2630.0–2632.0 m depth on the General Lithological Profile | Non Reported | Non Reported |
| N1 | 2.6 m of core, including: 0.7 m-fine grained sandstone, light gray, almost white, solid/compact. 0.8 m-fine grained sandstone, light gray, almost white, with a tabular diagonal layering, emphasized by accumulations of carbonated organic matter and Muscovite. 1.1 m -fine grained sandstone, light gray, almost white, with blurry layering. | 2632,0–2638,0 m depth on the General Lithological Profile | Cupressacites subgranulatus (Cupressaceae Conifers) Ischyosporites variegatus (Dicksoniaceae Ferns) Carnisporites granulatus (Selaginellaceae Lycopsids) Taurocusporites verrucatus (Notothyladaceae Bryophytes) Classopollis classoides (Cheirolepidiaceae Conifers) Trachysporites sp. (Pteridophyta Ferns) Tsugaepollenites macroverrucosus (Pinaceae Conifers) Corrugatisporites scanicus (Lygodiaceae Ferns) Deltoidospora minor (Cyatheaceae Ferns) Carbonized vegetable chaff | Non Reported |
| N$_2$ | Downwards lutite turning into mudstones. At a depth of 2638.0–2794.0 m, interpretation based on geophysical measurements and crumb samples | 2638,0–2642,0 m depth on the General Lithological Profile | Non Reported | Non Reported |
| Oldest | sandstones and inverted sandstones, with individual inserts of sandy mudstones | 2638,0–2642,0 m depth on the General Lithological Profile | Non Reported | Non Reported |

## Paleogeography

At the Late Pliensbachian (Margaritatus-Spinatum) several changes occur on the Polish Basin. The last section, latest Raricostatum subzone, there were a series marine transgressive surfaces, linked with the called Pli1 transgressive phase, known thanks to molluscs and ooids, along with an increase of marine macrofauna and microfauna. A marine Ingression filled the Polish basin, depositing marine dark Clay, Mudstone and lenticular Heteroliths, that are abundant on the lower-Middle Pliensbachian Łobez Formation and Poland, where dark-grey marine Claystones appear on the Kaszewy 1 borehole. There was a visible shoreline retreat that ends on the beginning of the Late Pliensbachian Margaritatus Chronozone. After the Pli1, a series of local subordinate retrogradational stratal-pattern architectures indicate the transgressive phase of the sequence Pli2. Pli2 The transgressive phase led to a maximum flooding level at the middle Spinatum. That was known thanks to Ammonites found on an 85 m basin at NW Poland, on the Wolin IG-1 borehole. This transgression also impacted deposition, with 265 m sequences of Mudstones and sandstones in a syndepositional tectonic graben at the Wolin-Recław Zone. After the maximum peak of the flooding basin it followed a regressive phase, characterized by progradational fluvio-deltaic sandstones, like the series of sandstones that recover the major tracksites of the Drzewica Formation, the Śmiłów Quarry Tracksite and the Wólka Karwicka Tracksite, both know from a regression nearshore/barrier lagoonal derived sandstone deposit. The Coeval deposits of Usedom and East Prignitz on Germany contain numerous Glendonites (Calcite pseudomorphs after Ikaite), what lends support to the possibility of a Late Pliensbachian cooling event and polar Glaciation at that time. Mostly of the authors, based on the current scientific quorum and the wide distribution of Glendonites in Upper Pliensbachian strata, tend to support a glacial-derived hypothesis. The Spinatum regression collides with the worldwide Toarcian Oceanic Anoxic event. The oldest deposits of the sequence (Latest Pliensbachian-Lowermost Toarcian), partly of an age resting on a regional erosional surface, comprise alluvial and deltaic sediments assigned to a number of lithostratigraphic units of local significance. The sequence recovers significant sea-level falls, that are related with the recurrent Late Pliensbachian glaciation, that would be followed at the start of the Toarcian with Global warming and a large scale transgression, recovered locally on the Ciechocinek Formation.
Thanks to data from the Kaszewy 1 borehole levels of 1200 pCO$_2$ were recovered, thanks to Plant stomatal index, being the highest amount recovered on the whole Lower Jurassic interval. The studied strata also led to know a great variability in Total Organic Content, interpreted as result of a greater terrestrial biodegradation. This led to knowing that the lower temperatures recovered from the Polish Basin in this interval occurred in the latest Rhaetian/earliest Hettangian, the late Hettangian, early Pliensbachian and especially in the Late Pliensbachian (Stokesi, Gibbosus, Spinatum), being this last the coolest period in the whole Early Jurassic.

## Sedimentological evolution

The Formation strata starts on the Polish Jurassic Depositional sequence V, that besides Drzewica area and Budki–Szydłowiec–Jagodne area, is also known from the Brody-Lubienia borehole (with a massive exposure of the sequence V in an operating quarry near Bielowice, the major one along with Skrzynno near Przysucha), that is located on the eastern part of the Świętokrzyskie Mountains region, giving insight into development of Pliensbachian deposits in the marginal parts of this basin. The initial part of this sequence is associated with a local fall of the sea level, where the lower IV level transition exposes with erosion, on locations such as the Zychorzyn borehole (at depth of 96.1 m) on Drzewica. The latter sedimentation is marked with thin deposits from delta-distributary depositional subsystem, that become thicker around the Budki, where in the zone of the Brody-Lubienia borehole an alluvial-meandering channel deposition system developed. After this initial regression there was a local transgression that led to abundance of local trace fossils and the formation of barrier-lagoonal deposits on Budki and in Brody-Lubienia, along with the increased presence of marine bivalves. In the Brody-Lubienia borehole were developed in embayment-lagoon facies. After the major marine transgression there was a visible change on Budki–Szydłowiec–Jagodne, shifting to fluvial discharge continuously moving along the Nowe Miasto–Iłża fault to the NE direction during the Pliensbachian times. At this time on the Jagodne 1 borehole it was deposited the development of a barrier-lagoon linked with a delta.
The Depositional sequence VI starts on the Jagodne 1 borehole with very large Mud clasts, pointing to intensive erosion of the underlying lagoonal sediments, while on the basin center, on the Zychorzyn borehole the erosion on lower sediments was lesser aggressive. The sediments that were deposited at the start of this sequence are uniformly represented by coarse alluvial deposits, interpreted as meandering river with possible conspicuous share of bed load transport. On the Budki 1 borehole series of coarse-grained alluvial package (especially large scale tabular cross-bedding sets dominate and quartz pebbles association), can represent low-sinuosity braided channels. Another transgressive phase on this sequence led to the formation of foreshore/shoreface/barrier deposits, and a local event on the Polish Basin, where there was a rapid drowning of the palEvelina Bertoli (born 8 July 1986 in Rome, Italy) is an Olympic Italian eventing rider. She competed at the 2021 and 2023 European Championships, and at the 2022 World Championships in Pratoni del Vivaro. Her sister Sara Bertoli took part in the 2008 Olympic Games in Modern Pentathlon.
In 2024 she competed at the 2024 Summer Olympics in Paris. paleorelief with alluvial valleys and coastal plains, resulting in embayed coastline with detached beach andor barrier ridges. There was equilibrium with the sea level, with periodical little regressions and transgressions. The maximum flooding range of this phase is represented by the Szydłowiec sandstones, developed on a regressive period where on Śmiłów was developed an eolian dune depositional subsystem linked with a barrier-lagoon, know due to the presence of barrier crest eolian deposits with plants buried. In this environment, the major Tracksites of the formation were developed. Linked with the Śmiłów Quarry sandstone appear deltaic deposits on the Brody-Lubienia borehole and on Zychorzyn.
The Depositional sequence VII is the last Pliensbachian major sequence, know especially from the Brody-Lubienia borehole and the continuation of an alluvial portion exposed also at Śmiłów, where an exposure exhibits sequence VI/VII characters, showing that the sea level fall was not so significant, as it was in the base of the previous sequence. This sequence starts with a rise of sea level and quick drowning of the whole area, destroying the previous delta system/marsh subsystem development measured at the Brody-Lubienia borehole, that shows that the amplitude of this transgression was quite high, as is represented by a thickness of 16 m.

### Śmiłów outcrop

There are several Types off Seashore Stratification seen on the Drzewica Formation. Starting with the post-Spinatum regression derived shore prograding cycle, where the nearshore deposits show detailed Hummocky cross-stratification, at the time are covered by thin beach-welded facies showing irregular “massive” bedding and numerous Plant roots penetrating down from the undulated bounding surface. Superimposed are fine grained, very well sorted sandstones with giant tabular Cross-bedding set interpreted as beach (Backshore depositional subsystem) Eolian Dune with complete Plant remains buried in the wind-transported Sand (Mostly, Herbaceous-Built Plants). The Direction of the Wind sedimentation is clearly visible on these deposits, where the stratification of the sand is well oriented. Those deposits are abundant on the Antecki quarry. The most related to the shores, probably barrier-Lagoon and Delta depositional systems are clearly visible on the Smilów Quarry, where the exposure shows different lithofacies and strata, including fine-grained, white and light-grey sandstones with Hummocky cross-stratification, with a series of tabular and trough Cross-bedding sets, probably related to near marine depositional subsystem. A more fine-medium-grained, grey to brownish sandstones also with flat-tabular-horizontal bedding with abundant drifted Plant Fossils and plant remains in situ is considered to be part of a clear foreshore barrier–Eolian depositional subsystem, probably related to a series of Dune fields, related to the Sea Barriers. Kaolinized grey Mudstones with abundant Dispersed plant roots are related to more lagoonal depositional subsystems. The slightly smaller and fine built, brownish and yellow sandstones that show details Cross-bedding are related to a more alluvial deposition, probably from the influence of a local Channel. On the Main Smilów Quarry the dominant shore face depositional subsystem show detailed marks of clear Palaeocurrent directions, dispersed trough Cross-bedding inclinations that point to the West. There have been measurements of the Wave-formed ripple that approximately reflects a shoreline orientation. Thanks to the wave ripple crests orientation and the orientation of fossil tree logs it was indicated an inclination of the inclined tabular Foreset bed continuing across the whole outcrop. These Foreset bed show a more pronounced inclination thanks to seaward-dipping Clinoforms, what reflects the original inclination of a barrier slope, with the sum of palaeocurrent pointing to south-west.

## Economic Resources
### CO$_2$ Storage

Zaosie Anticline-Jeżów Anticline are two of the three structures selected as having the best conditions for geological CO$_2$ storage in the Bełchatów region, with several wells drilled. Both the structures are characterized by similar geological feature and petrophysical parameters of rocks, being potential reservoirs. Significant thickness, good reservoir properties and proper depth give sufficient grounds to select the Lower Jurassic sandstone levels of the Drzewica Formation.
Budziszewice-Zaosie structure is one of the potential CO$_2$ storage sites for the biggest emitter of that gas in Europe, the Bełchatów Power Plant, with the major and most perspective reservoir located on the Drzewica Formation Strata locally. Archival seismic sections subject to secondary interpretation made it possible for a more accurate mapping of the deep structure of the anticline and its definition as present state of recognition as tectonically continuous. The Major drilling hole was stablished on the top of the upper Pliensbachian, JPl3 (the top of the Drzewica formation of the first potential reservoir), with the top of the Ostrowiec formation (Polish: seria ostrowiecka) as the second potential reservoir. Parametric models of variability were made for the resulting geometrical matrix lithology, contamination, effective porosity and total permeability of the complexes. Several works based on static 3D model recovered effective storage capacity c.a. 10 Mt (during 6 or 30 years of injection depending on hydrodynamic conditions). Other major CO$_2$ storage is found on Sierpc anticline ("Upper Sławęcin Beds"), specially on the Sierpc 2 borehole, with measured capacity of 260 tonnes of dihydrogen.

### Sandstone Exploitation

The sandstones of the Drzewica Formation have been object of mining since the Middle Ages, used as key material for the local construction. Two kinds are distinctive, the main Szydłowiec sandstones and the Kunów sandstones, mined on Śmiłów (13th century–recent), Kunów (17th–20th century). It was used by members of the Polish renaissance such as Jan Michałowicz, for some of their most well known pieces, such as the Tomb of Wolski Brothers. Kunów sandstones have also been studied as potential geotourism attractions.

## Fungi

| Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon | Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited. |
| Genus | Species | Location | Material | Notes | Images |
| Xylophagous Fungi | Morphotype A (Aseptae) Morphotype B (Sphaerical) Morphotype C (Sack-shaped) Morphotype D (Transverse septa) Morphotype E (Amerospore) Morphotype F (Phragmospores) Morphotype G (Dictyospores) | Brody-Lubienia borehole. | Fungal Spores Hyphae-like attachments | Saprophyte fungal Spores from Marine and Deltaic Settings associated with wood and Litter. The frequency of fungal spores on the Polish basin on the Pliensbachian-Toarcian transition is correlated with negative C isotope peaks and enhanced cuticular plant litter accumulation, pointing to climate-driven enhanced decomposition of wood and rapid destruction of terrestrial carbon, which may have played an important role in the aggravation of the Jurassic Greenhouse disaster. Associated with a high rate of organic burial, the presence of Fungal Matter increased on the Uppermost layers of the Drzewica Formation, with a continued deposition between the T-OAE extinction, and several ups and downs on the Ciechocinek Formation, related with local climate and humidity changes. This is rather a reflection of the efficiency of terrestrial biodegradation. Measured increasing of temperature favoured local fungal-mediated decomposition of plant litter, specifically of normally resistant wood. Observed fungal spores represent various morphotypes and resemble superficially other palynogenic detritus, such as sphaerical Prasinophyceae. In some cases spherical fungal spores show structures related to Hyphae attachments. | Spores found on the Polish are mostly from Saprophyte (wood decomposers) fungi, resembling the extant genera like Scutellinia |

## Paleofauna

Along the Early Jurassic, there was a mostly marginal-marine (mainly Mesohaline), deltaic and continental sedimentation that prevailed in the area of Poland. A marine basin of Polyhaline character conditions occurred mainly in the Pliensbachian, due to a flooding of the Polish Basin. Due to that, Pliensbachian sedimentation in Pomerania developed in a shelf basin that attained depths of about 100 m, below the storm wave base. As the latest Pliensbachian began, the climate changes to a more cold environment, as prove the presence of ice sheets on northern latitude facies of the same age. That led to changes in salinity. The Drzewica Formation was influenced by an ancient swampy-lagoonal sub-system, with Deltaic currents depositing fluvial sediments. The Main Boreholes show eustatic changes on the marine conditions, with a retreat of the sea level, leading to the appearance of vegetation and abundant pollen on the uppermost parts of the formation. The regressive phase of sea level is represented by the Komorowo Beds and Upper Sławęcin Beds along with the own Drzewica Formation. As the level of the sea recovered on the Toarcian, most of the setting turn out to be a bathyal marine bottom. The major outcrop of the formation, found on the Śmiłów Quarry represents a nearshore, deltaic, barrier-lagoonal ecosystem, where the biota left a high report of footprints (+200), with different sizes, that range from small to large animals of various groups. The Smilow Quarry Tracksite is the main outcrop and setting for the Footprints. Along with the coeval (also belonging to the formation) Wólka Karwicka tracksite represent the only ones recovered from the Late Liassic of Europe. The footprints are related to mostly lagoon-derived facies, unlike the vegetation, that appears on Deltaic facies. Brackish deposits are relates to Brachiopodans and other molluscs and Limuloid animals. The presence of Late Pliensbachian common ammonoids and several vertebrate clades connects the marine fauna with similar environments of the same age across Europe. The Drzewica Fauna is also, one of the few from the Pliensbachian-Toarcian Anoxic event recorded worldwide, specially referred to the vertebrate fauna reported.

### Invertebrates
#### Ichnofossils

| Genus | Species | Location | Material | Type | Abundance | Notes |
| Cochlichnus | C. isp. | Szydłówek quarry Śmiłów quarry Podkowiński quarry Antecki quarry | Traces | Fodichnia | Rare | Brackish or freshwater trace fossils. Locomotion and feeding trace of aquatic annelids, insect larvae and nematodes |
| Conichnus | C. isp. | Szydłówek quarry Śmiłów quarry Podkowiński quarry Antecki quarry | Subcylindrical burrows | Fugichnia Domichnia | Common | Marine, brackish or freshwater tubular traces of annelids or crustaceans |
| Diplocraterion | D. parallelum | Kaszewy 1 | U-Shaped Burrows | Fodichnia | Common | Marine trace fossils. Is often associated with successions characterised by changes of sea level. Linked with crustaceans, specially shrimps |
| Imbrichnus | I. isp. | Szydłówek quarry Śmiłów quarry Podkowiński quarry Antecki quarry | Burrows | Fugichnia | Rare | Brackish or freshwater tubular burrows. Probably made by annelids, bivalves or crustaceans |
| Isopodichnus | I. isp. | Szydłówek quarry Śmiłów quarry Podkowiński quarry Antecki quarry | Traces | Fodichnia | Common | Brackish or freshwater trace fossils. Locomotion and feeding trace of phyllopod and notostracan crustaceans in nonmarine environments. It represents traces with great similarity of palaeoecology, population size-frequency, morphology and behaviour of Triops cancriformis |
| Lockeia | L. siliquaria L. amygdaloides L. czarnockii | Szydłówek quarry Śmiłów Quarry Podkowiński quarry Antecki quarry Wólka Karwicka Brody-Lubienia borehole | Dwelling traces | Cubichnia Domichnia | Very Abundant | Marine, brackish or freshwater resting traces of Bivalves. The Lockeia traces trend to accumulate in brackish waters on the case of the Drzewica Formation. They can occur in large aggregates, where occasionally traces present on agglomerations show a clear orientation, probably parallel to the palaeocurrent direction, which is associated with locomotion of the burrowing animal. |
| Scalichnus | S. phiale | Szydłówek quarry Śmiłów quarry Podkowiński quarry Antecki quarry | Sac/Bottle shaped burrows | Fugichnia Domichnia | Common | Marine, brackish or freshwater tubular traces of annelids, aquatic insect larvae or crustaceans |
| Scolicia | S. isp. | Szydłówek quarry Śmiłów quarry Podkowiński quarry Antecki quarry | Traces | Fodichnia | Common | Brackish or freshwater trace fossils. Locomotion and feeding trace of gastropods |
| Skolithos | S. isp. | Szydłówek quarry Śmiłów quarry Podkowiński quarry Antecki quarry | Sac/Bottle shaped burrows | Domichnia | Common | Brackish tubular traces of annelids, crustaceans or insects |
| Spongeliomorpha | S. isp. | Szydłówek quarry Śmiłów quarry Podkowiński quarry Antecki quarry Brody-Lubienia borehole Kaszewy 1 | Sac/Bottle shaped burrows | Domichnia | Common | Brackish tubular traces of annelids, crustaceans or insects |

#### Bivalvia

Unidentified blackish bivalves were reported from the formation on the first report of the Tracksites. Are related with shallow lagoonal facies.
| Genus | Species | Location | Material | Notes |
| Anodonta | A. liasokeuperina | Śmiłów Szydłowiec Podkowiński Quarry | Cunchs | A freshwater mussel, type member of the family Unionidae inside Unionida. Diverse and correlated with freshwater deposits on the Hettangian-Sinemurian of Bavaria |
| Cardinia | C. philea C. sp. | Szydłowiec | Shells | A Saltwater/blackish cockle, type member of the family Cardiniidae inside Carditida. On the Pliensbachian strata there is not a clear consensus if the Polish basin reached stenohaline conditions, but the presence of this genus on the Parkoszowice 58 BN borehole along with foraminiferans and marine phytoplankton tends to support more polyhaline-mesohaline conditions. |
| Nuculana | N. zieteni | Szydłowiec | Shells | A saltwater/brackish Pointed Nut clam, type member of the family Nuculanidae inside Protobranchia. Originally described as Dacryomya zieteni. |
| Pleuromya | P. forchammeri | Szydłowiec | Shells | A saltwater Clam, type member of the family Pleuromyidae inside Pholadida. A species are heretofore known solely from the Scania and Bornholm region, derived there from the Doshult and Katslosa series. |
| Pronoella | P. cf. elongata | Szydłowiec | Cunchs | A saltwater/brackish Clam, member of the family Arcticidae inside Cardiida. |

#### Gastropoda

| Genus | Species | Location | Material | Notes | Images |
| Viviparus | V. spp. | Szydłowiec sandstones | Shells | A freshwater snail, type member of the family Viviparidae inside Viviparoidea. In the area of the northern and north-western edge of the Swiętokrzyskie Mountains, the sea fauna has been found sporadically, but there are several finds of freshwater forms. Viviparus main identified snail on the Formation, and a typical form associated with Deltaic-Fluvial Deposits. | Example of extant specimen of Viviparus |

#### Ammonites

| Genus | Species | Location | Material | Notes | Images |
| Tragophylloceras | T. loscombi | Kaszewy 1 borehole | MUZ.PGI.80.VI.149, single 1,7 cm Shell. | An Ammonite, member of the family Juraphyllitidae inside Ammonoidea. Latest Pliensbachian index fossil, Tragophylloceras is considered to be part of the Mediterranean-Submediterranean fossil region, but it has exceptions, with specimens coming from southern England and northwestern Germany. In this case, the rocks that hosted the specimen also have poorly preserved dinoflagellate cysts, belonging to the genus Mendicodinium, that cannot expose its age beyond Late Pliensbachian. This is supported by the presence of the fossil 20 m below the Lower Toarcian deposits. It looks also to be related to a large flooding surface, with the specimen being probably washed to nearshore or epicontinental waters. | Example of Tragophylloceras Fossils |

#### Crustacea

| Genus | Species | Location | Material | Notes | Images |
| Euestheria | E. brodieana E. minuta | Podkowiński quarry | Valves | A Brackish/Freshwater Clam shrimp (Phyllopodan) of the family Lioestheriidae. The presence of Euestheria marks the appearance of less saline conditions, as this is a mostly freshwater genus. The local Phyllopods are related with a great amount of freshwater debris (especially plants), and suggest seasonal changes in the rivers on the Pliensbachian-Toarcian Polish Basin. | |
| Liasina | L. lanceolata L. sp. | Kaszewy 1 borehole Gutwin borehole Brody-Lubienia borehole | Docens of Specimens | A Brackish Ostracodan of the family Pontocyprididae. | |

#### Limuloidea

| Genus | Species | Location | Material | Notes | Images |
| Limulidae | Indeterminate | Śmiłów Quarry | Unknown, only Cited | A marine/brackish Horseshoe crab, member of the family Limulidae inside Xiphosurida. Body fossils reported from the Śmiłów Quarry tracksite include limuloid animals. Due to the environment representing a barrier-lagoonal setting, the specimens may have died during mating season. The specimens haven't been described. | Limulus, example of Limuloid |
| Limulitella | L. cf. liasokeuperinus | Skarżysko-Kamienna | Head and partial upper body | A marine/brackish Horseshoe crab, member of the family Limulidae inside Xiphosurida. Only representative Limuloid from the Lower Jurassic, Limulitella cf. liasokeuperinus was found associated with brackish and deltatic facies, related to nearshore deposits. Its affinities with the genus Limulitella has not been proved. It has been suggested as Limulitella incertae sedis, on the basis that there is not enough material to prove its affinities. The status of the species remain controversial, since the original fossil has been lost or in an unknown collection, more probably a private one. | |

### Vertebrates
#### Chondrichthyes

| Genus | Species | Location | Material | Notes | Images |
| Hybodontoidea | Indeterminate | Szydłowiec sandstones | Teeth | A Freshwater/Marine Shark, member of the family Hybodontidae inside Hybodontiformes. Contemporaneous Hybodus fossils are known from Rumania. Due to the abundance of Hybodus on Lower Jurassic Strata this has been a temporal assignation. Beyond that, the specimen hasn't been compared. The genus resembles the freshwater sharks from coeval units of North America. | Restoration of Hybodus |
| Paleoxyris | P. muensteri | Szydłowiec sandstones | Egg Capsules | A Freshwater/Brackish Shark Egg Capsule, probably from a member of the family of Lonchidiidae or Hybodontidae inside Hybodontiformes. With the study of the contemporaneous flora from Chmielow (Gielniów Formation), Paleoxyris egg capsules were identified. On the same paper are claimed to be recovered also from the Szydłowiec sandstones. This species is also known from the Rhaetian of Tübingen, Germany. The best candidate for this capsules can be the genus Lonchidion. | |

#### Actinopteri

Unidentified fin spines are known from this formation.
| Genus | Species | Location | Material | Notes | Images |
| Semionotidae | Indeterminate | Szydłowiec sandstones Podkowiński quarry | Right Dentary Dorsal body with Scales Referred Teeth Isolated scales | A Freshwater Bony Fish, member of the family Semionotiformes inside Neopterygii. Fossils of Semionotus cf. bergeri are known from the Hettangian of Poland. The found teeth can be from a relative. Represents a genus similar to coeval fishes from inland deposits of North America. | Example of Semionotiform, Semionotus |

#### Synapsida

| Genus | Species | Location | Material | Notes | Images |
| Brasilichnium | Cf. B. isp. | Śmiłów quarry Szydłówek quarry | Tracks | Represent tracks from Mammaliformes, member of the ichnofamily Chelichnopodidae, incertae sedis inside Synapsida. Resemble creatures such as Sinoconodon, Morganucodonta such as Erythrotherium, Megazostrodon or Dinnetherium; more advanced mammaliformes, such as members of Haramiyida, including arborear forms such as Shenshou. As show by Middle-Late Jurassic Finds, this group was ecologically diverse. | A genus similar to Morganucodon is most probably the best candidate for the local Brasilichnium footprints |

#### Crocodylomorpha

| Genus | Species | Location | Material | Notes | Images |
| Batrachopus | B. isp. Cf.B. isp. | Śmiłów quarry Szydłówek quarry Wólka Karwicka | Tracks | Terrestrial/Semiacuatic crocodylomorph tracks, type member of the ichnofamily Batrachopodidae, incertae sedis inside Pseudosuchia. The only ones know from the Late liassic of Europe. It resemble Protosuchus-like small terrestrial crocodylomorphs, and is probably related with the genera from the Kayenta Formation, such as Kayentasuchus or Eopneumatosuchus, or the Chinese genera Strigosuchus or Platyognathus. Seashore dwelling animals, with Carnivorous and Herbivorous representatives. Another series of larger footprints Cf.Batrachopus can come from primitive Amphicotylus-like Goniopholididae or even a member of Teleosauridae | Terrestrial crocodylomorphs such as Protosuchus, were most likely the Batrachopus trackmakers. |

#### Theropoda

| Genus | Species | Location | Material | Notes | Images |
| Anchisauripus | A. isp. A A. isp. B Cf.A. isp. | Szydłówek quarry Śmiłów quarry Wólka Karwicka | Footprints | Theropod tracks, type member of the ichnofamily Anchisauripodidae, incertae sedis inside Neotheropoda. Assigned to Coelophysidae-alike dinosaurs. Small to medium slender primitive predatory dinosaurs, related with genera such as Liliensternus, Tachiraptor, Zupaysaurus, Procompsognathus or contemporaneous taxa such as the North American Segisaurus. | Anchisauripus footprints belong to a genus similar to Procompsognathus |
| Eubrontes | E. isp. Cf.E. isp. | Szydłówek quarry Śmiłów quarry | Footprints | Theropod tracks, type member of the ichnofamily Eubrontidae, incertae sedis inside Theropoda. Eubrontes' is related to the Genus Dilophosaurus, representing a basal Neotheropods. Include some large tracks, suggestings specimens up to 7 m, using Dilophosaurus as reference. Dilophosaurus is found or North America Pliensbachian strata, although the assignation of these tracks to the genus is controversial. Similar tracks have been described from Hettangian deposits from Poland and Sweden. There is a distinctive crouching theropod trace assigned to Eubrontes plexus, suggested a feather ingument on Dilophosauroids or relatives | Model nicknamed "Dyzio", who represents a feathered reconstruction based on Drzewica and North American finds |
| Grallator | G. ispp. Cf.G. isp. | Śmiłów quarry Szydłówek quarry Wólka Karwicka | Footprints | Theropod tracks, member of the ichnofamily Eubrontidae, incertae sedis inside Neotheropoda. The footprints from Smilow are small-sized,tridactyl and relatively narrow. forms Assigned to Coelophysidae-alike dinosaurs. Small to medium slender primitive predatory dinosaurs, related with neotheropods such as Camposaurus or Dracoraptor. | Grallator footprints belong to a genus similar to Camposaurus |
| Kayentapus | K. ispp. Cf.K. isp. | Szydłówek quarry Śmiłów quarry | Footprints | Theropod tracks, member of the ichnofamily Eubrontidae, incertae sedis inside Neotheropoda. The tracks resemble Sinosaurus and are related with genera such as Notatesseraeraptor, Tachiraptor, Sarcosaurus and other basal Neotheropoda related to Tetanureae and Ceratosauriade. Small to medium-sized theropods. | |
| Megalosauripus | M. isp. | Szydłówek quarry | Footprints | Theropod tracks, member of the ichnofamily Eubrontidae, incertae sedis inside Theropoda. Largest Theropod Footprints from the Late Lower Jurassic. They have resemblance with the non-Tetanureae Sinosaurus, but match with Late Jurassic Orionides trackmakers. The trackmaker is expected to reach a length of 8.7-9.1 m long, with a weight over 1.9-2.05 tonnes, making it among the largest Early Jurassic know theropod Trackmakers. The footprints found on Drzewica seem even larger because of their large metatarsophalangeal area, which is often observed in the large theropod footprints from the post-Liassic strata. | Megalosauripus footprints can belong to a large relative of Chuandongocoelurus |
| Plesiornis | cf. P. isp. | Szydłówek quarry | Footprints | Theropod Tracks, member of the ichnofamily Anchisauripodidae, incertae sedis inside Theropoda. | |
| Stenonyx | S. isp. | Szydłówek quarry | Footprints | Theropod Tracks, member of the ichnofamily Anchisauripodidae, incertae sedis inside Theropoda. Considered a very Small Dinosaur, probably a juvenile. The Foot resembles those of young Coelophysis from Ghost Ranch Quarries. Stenonyx footprints had been described from the Early Jurassic Soltykow (Hettangian) and this ones Szydlowek, what matches with specimens found on the also Pliensbachian Hasle Formation, linking Southern Sweden, Bornholm and Poland, contiguous during the Early Jurassic and dinosaurs could thus freely roam this large area. | |

#### Sauropodomorpha

| Genus | Species | Location | Material | Notes | Images |
| Parabrontopodus | P. ispp. | Szydłówek quarry Śmiłów quarry | Footprints | Sauropod tracks, type member of the ichnofamily Parabrontopodidae, incertae sedis inside Sauropodomorpha. Two different casts of footprints of primitive Sauropods. Parabrontopodus isp. A ranges from a foot model identical to the genus Barapasaurus and the invalid "Damalasaurus", with an estimated body size of 6.8 m. Parabrontopodus isp. B represents footprints that match with the genus Gongxianosaurus. The small size of the footprints can suggest juvenile individuals travelled along the deltaic facies of the formation. | Local Parabrontopodus resemble the feet of the genus Barapasaurus |

#### Ornithischia

| Genus | Species | Location | Material | Notes | Images |
| Anomoepus | A. ispp. Cf.A. isp. | Szydłówek quarry Śmiłów quarry | Footprints | Ornithischian tracks, member of the ichnofamily Anomoepodidae, incertae sedis inside Neornithischia. A foot similar to Lesothosaurus, implying 2.2 m long specimens. On the other casts there are tracks that resemble the feet of Stormbergia and various Genasauria of different sizes, between 90 and 130 cm. Fast-running small herbivorous ornithischians, probably gregarious. | Stormbergia matches with the bodyplan assigned to the Anomoepus trackmaker |
| Moyenisauropus | M. isp. Cf.M. isp. | Szydłówek quarry Śmiłów quarry Wólka Karwicka | Footprints | Thyreophoran tracks, type member of the ichnofamily Moyenisauropodidae, incertae sedis inside Neornithischia. Is considered synonymous with the ichnogenus Anomoepus. The tracks adscribed resemble basal Thyreorphora feet. The tracks related to the genus are vinculated with genera such as Scelidosaurus, whose fossils have been found on Pliensbachian strata of England. In this case, implicates a ~4 m long animal. | Scelidosaurus matches with the bodyplan assigned to the Moyenisauropus trackmaker |

## Plants

There are traces of root plants with numerous plant remains, some of them vertically preserved, and on this basis he derived a possible dune-related origin of these sediments. The plants in the growth position were present at the bases of aeolian sandstone bodies, up to 2 m thick, which wedge out laterally and are covered by argillaceous strata of coastal lagoon origin with drifted plant remains. Mostly of the plant remains are composed by rhizoids and undeterminable fragments, recovered at the Szydłowiec sandstones, Śmiłów and Mirzec, near Starachowice. More recent studies suggest an environment of shallow seafront and beach, which of course does not exclude the presence of Aeolian zones there. Lithofacies on the Smilów Quarry shows slightly inclined beach forests and abundant plant roots in a series of barrier foreshore, backshore and/or aeolian dune facies, that include even plants buried in whole dunes, while drifted plant fossils and tree logs occur on the main depositional surface. This inclined surface exposes most likely a long period of non deposition and washed plants, recovering at the same time the original tilt of the local beach.

### Chmielów Flora

This flora comes from layers of the also Pliensbachian Gielniów Formation (Polish: seria gielniowska) (Marginal Brackish), yet is very likely derived from the coeval Drzewica Formation deposits (Coeval coastline) of the Subsequence V. This flora includes at least 30 species: Neocalamites hoerensis (Calamitaceae Equisetales, also found on the Podkowiński Quarry); Laccopteris angustiloba & Phlebopteris muensteri (Matoniaceae ferns); Dictyophyllum acutilobum, Hausmannia crenata, Clathropteris meniscoides & Thaumatopteris schenkii (Dipteridaceae Ferns); Todites williamsonii & Cladophlebis denticulata (Osmundaceae ferns); Pterophyllum subaequale & Nilssonia orientalis/acuminata/simplex (Bennettitales, with Nilssonia being dominant like in the Lublin Floras); Ctenis nilssoni (Cycads); Sagenopteris nilssoniana (Caytoniaceae Seed Ferns); Ginkgoites marginatus (Ginkgos); Ixostrobus siemiradzkii & Czekanowskia nathorstii/setacea (Czekanowskiales Ginkgoopsids); Hirmeriella muensteri (Cheirolepidiaceae conifers); Schizolepis follini/moelleri/braunii & Palissya sp. (Palissyaceae conifers); Podozamites angustifolius/gramineus/stobieckii/distans (Krassiloviaceae conifers).

### Coals

The Kaszewy Coals found mostly on the Kaszewy-1 borehole and Niekłan PIG-1 borehole are the major coals of the Drzewica Formation and Ciechocinek Formation, representing the Pliensbachian-Toarcian boundary. The Pliensbachian section of the Kaszewy-1 borehole comprises ~220 m of terrestrial and shallow marine siliciclastic sediments. This Section was located on a nearshore-deltaic influenced setting, with an increased amount of both terrestrial and marine organic matter, reflecting increased weathering and increased transport of terrestrial organic matter, with ups and downs of the amount of both marine and terrestrial matter. Here were collected fossil Charcoal in abundance and polycyclic aromatic hydrocarbon. Total fossil charcoal abundance is generally low in the Pliensbachian part of the study section, before peaking immediately above the Pl-To Boundary. The abundance of coarse fossil charcoal particles (>125 μm) in the Pliensbachian-Toarcian sections of the Kaszewy-1 core is very low (0-15 particles/10 g sediment), while fine fossil charcoal abundance is generally low in the Pliensbachian level. There is also a greater abundance of non-charcoal particles. At the start of the Toarcian locally, there is a measured increase on the amount of fine charcoal particles reflecting changes on the environment. Inside the Polycyclic Aromatic Hydrocarbons, the Pyrolytics (Benz-Anthracene, Benzo(k)Fluoranthene, Fluoranthene, Indeno[1,2,3-cd]Pyrene, Phenanthrene and Pyrene) were detected on a high variety of samples, with the Phenanthrene as the most abundant component, while Coronene is the lowest, suggesting an origin for this last one on the pyrolysis of organic matter. Petrogenics are more abundant on the local coal samples than the Pyrolytic, what can suggest that these last ones may not reflect particularly local high levels of wildfire activity. Apparently, the Kaszewy-1 borehole did not experience increased wildfire activity, but instead that the fine fossil charcoal abundance and Pyrolytics concentration records have shown a more clear regional wildfire signal. The fossil charcoal abundance and geochemical data from Pliensbachian and Toarcian samples show evidence of at least four, and possible another two, levels of increased wildfire activity, and that these can be grouped into two larger periods of increased wildfire activity.

### Palynology

The Drzewica Formation belongs to the Horstisporites planatus (Pl) Zone (Upper Sinemurian – Pliensbachian). This level starts on the older upper part of the Ostrowiec Formation, the Łobez Formation and Komorowo Formation of Pomerania and in other equivalent units from all Poland. This Megaspore section of relatively poor compared with the lower and upper sections, with the lower boundary of the level is determined by the appearance of the indicator species Horisporites planatus (Selaginellaceae) and The upper limit of the level is marked by the appearance of continuous occurrences of Paxillitriletes phyllicus (Isöetaceae), indicating more humid conditions, and Erlansonisporites sparassis (Selaginellaceae). This level has a series of characteristic megaspores, such as Minerisporites institus (Isöetaceae) and Hughesisporites pustulatus (Lycophyta) and on a minor ratio, rare occurrences of Hughesisporites planatus recorded even above the level range. Is important to recover that the limit of the Pliensbachian-Toarcian in Poland was the first precisely defined on the basis of a detailed stratigraphic-sequential correlation, fully later confirmed by a high-resolution chemostratigraphic correlation, based on isotope analysis, with the ranges of individual megaspores fitting well into this limit. This level can be correlated with the non-megasporous level of Kuqaia quadrata, found in the sediments of the Sinemurian- Lower Pliensbachian Yangxia Formation in the Xinjiang Province, China. In the Polish profiles, the Pl level correlates with the upper part of Rogalska's 1st phase as well as the 2nd and 3rd microflora phase, where the following indicator miospores appear: Lycopodiumsporites gristhorpensis (Lycopodiaceae), Undulatisporites undulapolus (Filicopsida) and Applanopsipollenites segmentatus (Gymnospermopsida), and in central Poland represents the works of the Upper Sinemurian and the whole Pliensbachian, being probably correlated with the middle part of the miospore level Cerebropollenites macroverrucosus (Coniferophyta), distinguished in the Pliensbachian and Toarcian deposits in northern Europe.
The Drzewica Formation palynoflora is recovered mostly on the Gutwin and Brody-Lubienia boreholes, and includes the next plant groups: Bryophyta, Lycopodiaceae, Equisetaceae, Selaginellaceae, Cyatheaceae, Dipteridaceae, Dicksoniaceae, Gleicheniaceae, Matoniaceae, Osmundaceae, Schizeaceae, Peltaspermaceae, Taxodiaceae, Cheirolepidiaceae, Araucariaceae, Podocarpaceae, Pinaceae, Gnetales, Cycadidae, Bennettitales and Ginkgoales. In between all the taxa, Pollen from Seed Ferns and Conifers dominate over miospores, indicating a relatively dry ecosystem. Freshwater algae miospores are known (Botryococcus sp., Cymatiosphaera sp.), as well Paleozoic palynoremains washed from uplands (Walchia sp., Pseudowalchia biangulina, Ullmannia frumentaria, Florinites antiquus, etc.), what indicate a clear active river influx, yet has also evidence of marine ingressions due to the presence of saltwater acritarchs (Psophosphaera coniferoides, Leiosphaeridia sp.).

### Biomass

Beyond proper palynogy, biomass associated has been recovered, specially on the Brody-Lubenia borehole, with abundance of C29 diasterenes (>70%), that proves a great contribution of land plants and thus terrestrial deposits nearby. There is also in some levels abundance of algae-derived C27 and C28 diasterenes, coeval with acritarchs, prasinophytes and dinoflagellates, being this last ones important primary producers in the Polish Basin. The presence of C27 sterols points to the importance of the dinoflagellates, but also other groups, such as Bangiophyceae and Eustigmatophyceae algae or marine protists like the Thraustochytriaceae. At least in some facies there is evidence of proliferation of freshwater-tolerant algae in the brackish environments of the Polish Basin.

### Megaflora

The Lublin Upland fluvial sandstones contain diverse types of fossil flora, associated genera and species only with Lower Jurassic sediments. Originally, while studying the Carboniferous flora from the boreholes in the area of the planned Bogdanka Coal Mine, appeared typical flora in similar to Jurassic formations. The age of the plant material was not determined concretely until 2020, where was recovered as being Pliensbachian-Toarcian in age, with the flora representing an arid environment, more probably Pliensbachian, that is covered locally by the Drzewica Formation. The Brody-Lubienia borehole is abundant on terrestrial Palynomorphs (know due to the presence of C29 Diasterenes, >70%), but also has high abundances of aquatic derived biomass. Even with that, there is a clear consensus with the more terrestrial character of the sediments from Brody-Lubienia, expressed by the frequent occurrence of plant roots and paleosol horizons. This also is recovered on the rate of MTTCs (mono-, di- and tri-methyl-trimethyltridecyl-chromanes) where higher indices at Brody-Lubienia indicate lower salinities and a stronger influx of riverine freshwater. The environment was probably Dry, developing flora on the near Freshwater influx settings. The Lublin Flora is linked with the flowing waters from the East. Lublin lias is dominated by cycads and Bennetites Ginkgoales and seed ferns occur sporadically, all on a conglomerate with numerous species occurs in the bottom, where the deposits are filled with of coal, mudstone, sandstone and clay siderite (reworked from the Carboniferous), as well as pebbles from Devonian limestones. Vegetation mostly grew outside the sedimentation area, as well as on shores and shallows.
At the Śmiłów Quarry, plant remains are known, and are interpreted as derived from an aeolian rework, with the plants buried in growth position by barrier crest aeolian dunes. Szydłówek quarry host also a nearshore-foreshore-backshore/eolian setting, where the flora is dominated by woody trunks, plant root moulds and impressions of drifted woody trunks, that suggest the presence of large coniferous forests around the Szydłówek tracksite. Plants here thrive on the foredune are exposed to salt spray, strong wind, and burial by blowing and accumulating sand. Ferns like Matonia braunii maybe worked as dune stabilisers, like the extant plant Leymus arenarius.
| Genus | Species | Stratigraphic position | Material | Notes |
| Anomozamites | A. cf. gracilis | Borehole L-95 near Łęczna | Pinnae | Affinities with the Bennettitales inside Bennettitopsida. Anomozamites is characterised by slender, (almost) completely and regularly segmented leaves whose leaflets are generally as long as broad or, at maximum, two times as long as broad. This genus is related with more arboreal Bennetitalean flora. Shows coriaceous leaves and is a genus linked more with dry climates. |
| Baiera | B. digitata B. sp. | Borehole L-95 near Łęczna Kaszewy 1 | Leaves Cuticles | Affinities with the Ginkgoaceae inside Ginkgoidae. Linked to the Hettangian-Sinemurian flora from Greenland and Skane, but also with coeval flora from the Sorthat Formation. Is the main Tree flora recovered locally. Is related with relatively warmer and humid conditions. It is the main recovered Floral cuticles on the Kasazewy 1 Borehole. |
| Caytonia | C. sp | Lublin Coal Basin | Ovulate Structure | Affinities with the Caytoniaceae inside Caytoniales. Represents the Ovulate organs or large Tree ferns, and is related with the middle Jurassic flora of United Kingdom. |
| Czekanowskia | C. sp. | Kaszewy 1 | Cuticles | Affinities with the Czekanowskiales inside Ginkgoopsida. This Genus is related with relatively drier-cooler conditions. |
| Ginkgoites | G. sp. | Kaszewy 1 | Cuticles | Affinities with the Ginkgoaceae inside Ginkgoidae. Is related with relatively warmer and humid conditions. |
| Hirmeriella | H. sp. | Borehole L-95 near Łęczna (Lublin Coal Basin) | Leaves | Affinities with the Cheirolepidiaceae inside Pinales. The type Leave from the Cheirolepidaceae family (as Cheirolepis, the genus that give name to the family, is a junior synonym), appears to be linked with Hot climates, able to survive in dry, extreme conditions, and been fire tolerant. |
| Matonia | M. braunii | Podkowiński Quarry | PC IB UJ 109/1 (isolated stipes, one frond, isolated pinnules) 109/2 (groups of stipes, one pinna). | Affinities with Matoniaceae inside Pteridophyta. Matonia braunii represents the second major plant fossil identified on the Drzewica formation Dunar settings, it probably formed monotypic stands on coastal dunes, suggesting a wide ecological amplitude that extant related ferns. |
| Neocalamites | N. hoernensis | Podkowiński Quarry | Stems | Affinities with Calamitaceae inside Equisetopsida. A common horsetail on the Liassic of Europe. |
| Nilssonia | N. compta N. orientalis N. brevis N. polymorpha N. sp. | Borehole L-95 near Łęczna | Pinnae | Affinities with the Cycadeoidaceae inside Bennettitales. The most abundant plant macrofossil locally, and the most diverse found on the Lublin coals. It represents a Cycad Like plant with Coriaceous leaves |
| Nilssoniopteris | N. major N. vittata | Borehole L-95 near Łęczna | Pinnae | Affinities with the Bennettitales inside Bennettitopsida. This Bennetite has a leaf similar to Nilssonia, large, lanceolate in outline with coriaceous structure, like the modern angiosperm Philodendron. Is related with rather Dry-hot environments, with less Fern flora. |
| Otozamites | O. beani O. graphicus | Borehole L-95 near Łęczna | Pinnae | Affinities with the Bennettitales inside Bennettitopsida. A more arbustive type of Bennetite, abundant on the Pliensbachian-Toarcian strata along Eurasia, related to rather dry climates. |
| Pachypteris | P. lanceolata P. rhomboidalis | Lublin Coal Basin | Pinnae | Affinities with the Peltaspermales inside Pteridospermatophyta. Bipinnate leaves, rachis longitudinally striated, with a long petiole and secondary rachises inserted oppositely to suboppositely on the upper side of the primary rachis. This Leaves belong to large Aboreal Ferns related with dry environments. |
| Pseudotorellia | P. sp. | Kaszewy 1 | Cuticles | Affinities with the Pseudotorelliaceae inside Ginkgoopsida. This Genus is related with relatively drier-cooler conditions. |
| Swedenborgia? | S.? sp. | Śmiłów Quarry | MUZ PIG 76.III.1, Complete Plant | Affinities with Krassiloviaceae inside Voltziales. A problematical plant that has been reported only on the Formation. It is plant buried in whole in an eolian dune strata, composed mostly by sandstone, that was deposited on a barrier-lagoon depositional system (possible backshore/backbarrier depositional subsystem). With leaves of 5–25 cm long and about 2–5 mm wide, and an angle between them and the short main axis ranges within 20-30°, This taxon has a morphology that resembles Schizeacean ferns (Specially Schizaea pectinata), but also that of the Voltziales Aethophyllum stipulare from Anisian of France (A Herbaceous Conifer From the Triassic). It was proposed as a specimen of Swedenborgia on a revision of the specimens of the Hettangian from also Poland. This suggests a herbaceous appearance of these plants, against traditional suppositions about their mainly tree-like character. Finally, can be leaves similar to Podozamites (also Krassiloviaceae). If not herbaceous plants, it can be leaves from trees or shrubs, but all probably coming from an Aeolian-Dunar Environment, growing like the extant Pinus halepensis in mediterranean dunes. |
| Pterophyllum | P. sp. | Borehole L-95 near Łęczna | Pinnae | Affinities with the Williamsoniaceae inside Bennettitopsida. Leaves from Arboreal Bennetites, similar to the modern Cyca Encephalartos woodii, with robust Trunks, built for Dry and hot climates. |
| Sagenopteris | S. sp | Borehole L-95 near Łęczna | Pinnae | Affinities with the Caytoniaceae inside Caytoniales. Sagenopteris is most likely the Pinnae of the Tree Fern that also contain Caytonia. |
| Sphenobaiera | S. sp. | Kaszewy 1 | Cuticles | Affinities with the Ginkgoaceae inside Ginkgoidae. Is related with relatively warmer and humid conditions. |
| Stenopteris | S. (Rhaphidopteris?) duetschii | Lublin Coal Basin | Pinnae | Affinities with the Umkomasiaceae inside Pteridospermatophyta. Is very similar to Rhaphidopteris, characterized by usually bi-tripinnate, coriaceous leaves with narrow segments. It belongs to Large (up to 25 m tall) arboreal Fern-like plants. |
| Weltrichia | W. spectabilis | Lublin Coal Basin | Bennetite "Flower" | Affinities with the Cycadeoidaceae inside Bennettitales. The Bennetite flowers are the main organ that links this relatives with the Cycas with modern Angiosperms. Probably the Pollen assigned to Angiosperms come from plants with this "Flowers". |