# List of kerivoulines

Kerivoulinae is one of the four subfamilies of Vespertilionidae, itself one of twenty families of bats in the mammalian order Chiroptera and part of the microbat suborder. A member of this subfamily is called a kerivouline, or a woolly bat. They are found in Africa and Asia, primarily in forests and caves, though some species can also be found in grasslands, savannas, or wetlands. They range in size from the least woolly bat, at 2 cm (1 in) plus a 2 cm (1 in) tail, to the Kachin woolly bat, at 6 cm (2 in) plus a 7 cm (3 in) tail. Like all bats, kerivoulines are capable of true and sustained flight, and have forearm lengths ranging from 2 cm (1 in) to 5 cm (2 in). They are all insectivorous and eat a variety of insects and spiders. Almost no kerivoulines have population estimates, though two species—the St. Aignan's trumpet-eared bat and the Tanzanian woolly bat—are categorized as endangered species with population sizes as low as 600.
The 30 extant species of Kerivoulinae are divided between two genera: Kerivoula with 26 species, and Phoniscus with the other four. A few extinct prehistoric kerivouline species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.

## Conventions

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the kerivouline's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted. Population figures are rounded to the nearest hundred.

## Classification

Kerivoulinae, one of the four subfamilies of the family Vespertilionidae, contains 30 extant species divided into 2 genera.
Subfamily Kerivoulinae
* Genus Kerivoula (woolly bats): 26 species
* Genus Phoniscus (trumpet-eared bats): four species

## Kerivoulines

The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.
Genus Kerivoula – Gray, 1842 – 26 species :
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
| Bismarck trumpet-eared bat | K. myrella Thomas, 1914 | Manus Island in Papua New Guinea | Size: 3–4 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Forest | VU  1,500–2,000 |
| Clear-winged woolly bat | K. pellucida (Waterhouse, 1845) | Southeastern Asia | Size: 3–5 cm (1–2 in), plus 4–6 cm (2 in) tail 2–4 cm (1–2 in) forearm length Habitat: Forest | NT  Unknown |
| Copper woolly bat | K. cuprosa Thomas, 1861 | Central and western Africa | Size: 3–4 cm (1–2 in), plus 4–5 cm (2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Forest | DD  Unknown |
| Cryptic woolly bat | K. crypta Wroughton & Ryley, 1913 | Southern India and Sri Lanka | Size: About 4 cm (2 in), plus about 4 cm (2 in) tail about 3 cm (1 in) forearm length Habitat: Forest and caves | LC  Unknown |
| Damara woolly bat | K. argentata Tomes, 1861 Three subspecies K. a. argentata K. a. nidicola K. a. zuluensis | Southeastern Africa | Size: 3–5 cm (1–2 in), plus 4–6 cm (2 in) tail 2–4 cm (1–2 in) forearm length Habitat: Savanna | LC  Unknown |
| Dark woolly bat | K. furva Kuo, Soisook, Ho, Csorba, Wang, & Rossiter, 2017 | Eastern and southeastern Asia | Size: Unknown length, plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Forest and caves | LC  Unknown |
| Ethiopian woolly bat | K. eriophora (Heuglin, 1877) | Ethiopia | Size: About 5 cm (2 in), plus about 3 cm (1 in) tail 2–4 cm (1–2 in) forearm length Habitat: Unknown | DD  Unknown |
| Flat-skulled woolly bat | K. depressa Miller, 1906 | Southeastern Asia | Size: About 3 cm (1 in), plus about 4 cm (2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Forest and caves | LC  Unknown |
| Flores woolly bat | K. flora Thomas, 1914 | Indonesia and Malaysia | Size: About 4 cm (2 in), plus about 5 cm (2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Forest | VU  Unknown |
| Fly River trumpet-eared bat | K. muscina Tate, 1941 | Papua New Guinea | Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Forest | LC  Unknown |
| Hardwicke's woolly bat | K. hardwickii (Horsfield, 1824) | Southeastern Asia | Size: 3–5 cm (1–2 in), plus 3–5 cm (1–2 in) tail 2–4 cm (1–2 in) forearm length Habitat: Forest and caves | LC  Unknown |
| Indochinese woolly bat | K. dongduongana Tu, Hassanin, Furey, Son, & Csorba, 2018 | Southeastern Asia | Size: Unknown length, plus about 4 cm (2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Forest and caves | LC  Unknown |
| Kachin woolly bat | K. kachinensis Bates, Struebig, Rossiter, Kingston, Oo, & Mya, 2004 | Southeastern Asia | Size: 4–6 cm (2 in), plus 5–7 cm (2–3 in) tail 4–5 cm (2 in) forearm length Habitat: Forest | LC  Unknown |
| Krau woolly bat | K. krauensis Francis, Kingston, & Zubaid, 2007 | Southeastern Asia | Size: 3–4 cm (1–2 in), plus 3–5 cm (1–2 in) tail 2–4 cm (1–2 in) forearm length Habitat: Forest | NT  Unknown |
| Least woolly bat | K. minuta Miller, 1898 | Indonesia and Malaysia | Size: 2–3 cm (1 in), plus 2–5 cm (1–2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Forest | NT  Unknown |
| Lenis woolly bat | K. lenis Thomas, 1916 | Southern and southeastern Asia | Size: 4–5 cm (2 in), plus 4–6 cm (2 in) tail 3–5 cm (1–2 in) forearm length Habitat: Forest | LC  Unknown |
| Lesser woolly bat | K. lanosa (A. Smith, 1847) Four subspecies K. l. harrisoni K. l. lanosa K. l. lucia K. l. muscilla | Sub-Saharan Africa | Size: 4–5 cm (2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Savanna and forest | LC  Unknown |
| Painted bat | K. picta (Pallas, 1767) Two subspecies K. p. bellissima K. p. picta | Southern and southeastern Asia | Size: 4–5 cm (2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Forest and grassland | NT  Unknown |
| Papillose woolly bat | K. papillosa Temminck, 1840 Two subspecies K. p. malayana K. p. papillosa | Southeastern Asia | Size: 4–6 cm (2 in), plus 4–6 cm (2 in) tail 3–5 cm (1–2 in) forearm length Habitat: Forest | LC  Unknown |
| Small woolly bat | K. intermedia Hill & Francis, 1984 | Malaysia | Size: 3–4 cm (1–2 in), plus 3–5 cm (1–2 in) tail 2–4 cm (1–2 in) forearm length Habitat: Forest | NT  Unknown |
| Smith's woolly bat | K. smithii Thomas, 1880 | Central Africa | Size: 3–5 cm (1–2 in), plus 4–5 cm (2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Forest | LC  Unknown |
| Spurrell's woolly bat | K. phalaena Thomas, 1912 | Central Africa | Size: 3–4 cm (1–2 in), plus 3–5 cm (1–2 in) tail 2–3 cm (1 in) forearm length Habitat: Forest | LC  Unknown |
| St. Aignan's trumpet-eared bat | K. agnella Thomas, 1908 | Papua New Guinea | Size: 4–5 cm (2 in), plus about 5 cm (2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Forest | EN  600–750 |
| Tanzanian woolly bat | K. africana Dobson, 1878 | Tanzania | Size: About 4 cm (2 in), plus about 3 cm (1 in) tail about 3 cm (1 in) forearm length Habitat: Forest | EN  Unknown |
| Titania's woolly bat | K. titania Bates, Struebig, Hayes, Furey, Mya, Thong, Tien, Son, Harrison, Francis, & Csorba, 2007 | Southeastern Asia | Size: 3–5 cm (1–2 in), plus 4–6 cm (2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Forest | LC  Unknown |
| Whitehead's woolly bat | K. whiteheadi Thomas, 1894 Three subspecies K. w. bicolor K. w. pusilla K. w. whiteheadi | Southeastern Asia | Size: 3–5 cm (1–2 in), plus 2–5 cm (1–2 in) tail 2–4 cm (1–2 in) forearm length Habitat: Grassland and forest | LC  Unknown |
Genus Phoniscus – Miller, 1905 – four species :
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
| Dubious trumpet-eared bat | P. aerosa (Tomes, 1858) | Possibly southeastern Africa | Size: 4–6 cm (2 in), plus 3–5 cm (1–2 in) tail about 4 cm (2 in) forearm length Habitat: Forest | DD  Unknown |
| Golden-tipped bat | P. papuensis (Dobson, 1878) | Papua New Guinea and eastern Australia | Size: 4–6 cm (2 in), plus 3–5 cm (1–2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Forest | VU  Unknown |
| Groove-toothed bat | P. atrox Miller, 1905 | Southeastern Asia | Size: About 4 cm (2 in), plus 2–4 cm (1–2 in) tail 3–4 cm (1–2 in) forearm length Habitat: Forest | NT  Unknown |
| Peters's trumpet-eared bat | P. jagorii (Peters, 1866) | Southeastern Asia | Size: 4–5 cm (2 in), plus 3–5 cm (1–2 in) tail 3–5 cm (1–2 in) forearm length Habitat: Forest and inland wetlands | LC  Unknown |