# Frullania

Frullania is the only genus of liverworts in family Frullaniaceae. It contains the following species:
A
* Frullania aculeata Taylor, 1846
* Frullania acutata Caspary, 1887
* Frullania acutiloba Gerola, 1947
* Frullania akiyamae Hattori, 1986
* Frullania albertii Stephani, 1916
* Frullania allanii Hodgson, 1949
* Frullania allionii Stephani, 1910
* Frullania alpina Stephani, 1911
* Frullania alstonii Verdoorn, 1930
* Frullania alstonii var. pfleidereri Hattori, 1972
* Frullania alternans Nees In G., L. & N., 1845
* Frullania amamiensis Kamimura, 1968
* Frullania ambronnii Stephani, 1916
* Frullania amplicrania Stephani, 1910
* Frullania ampullifera Jack & Stephani In Stephani, 1894
* Frullania anderssonii Ångström, 1873
* Frullania angulata Mitten, 1863
* Frullania angulata F. Serratoides Vanden Berghen, 1983
* Frullania angulata var. laciniata Demaret & Vanden Berghen, 1950
* Frullania angustistipa Stephani, 1908
* Frullania anomala Hodgson, 1949
* Frullania antaresensis Hattori, 1980
* Frullania aoshimensis Horikawa, 1929
* Frullania apertilobula Gerola, 1947
* Frullania apicalis Mitten, 1879
* Frullania apicalis var. camerunensis Vanden Berghen, 1976
* Frullania apiculata (R., B. & N.) Nees In G., L. & N., 1845
* Frullania apiculata var. goebelii Schiffner, 1893
* Frullania apiculata var. guianensis Lindenberg & Gottsche, 1851
* Frullania apiculata var. laxa Nees In G., L. & N., 1845
* Frullania apollinarii Stephani, 1911
* Frullania aposinensis Hattori & Lin, 1985
* Frullania appalachiana Schuster, 1983
* Frullania appendistipula Hattori, 1972
* Frullania appendistipula var. spinifera Hattori, 1974
* Frullania arecae (Sprengel) Gottsche, 1863
* Frullania arecae var. spiniloba (Stephani) Yuzawa, 1991
* Frullania armata Herzog & Clark In Clark, 1954
* Frullania armatifolia Verdoorn, 1932
* Frullania armitiana Stephani, 1911
* Frullania armitiana var. inflexula Hattori, 1988
* Frullania armitiana var. longe-Attenuata (Hattori) Hattori, 1980
* Frullania arsenii Stephani, 1924
* Frullania asagrayana Montagne, 1842
* Frullania astrolabea Stephani, 1910
* Frullania aterrima (Hooker & Taylor) Hooker & Taylor In G., L. & N., 1845
* Frullania aterrima var. lepida Hodgson, 1949
* Frullania atrata (Swartz) Montagne, 1839
* Frullania atrata var. flaccida (Weber) Nees In G., L. & N., 1845
* Frullania atrata var. mexicana Nees In G., L. & N., 1845
* Frullania atrosanguinea Taylor Ex Spruce, 1884
* Frullania attenuata Stephani, 1911
* Frullania auriculata Hattori, 1985
* Frullania azorica Sim-Sim, Sergio, Mues & Kraut, 1995
B
* Frullania baileyana Stephani, 1910
* Frullania bakeri Stephani, 1916
* Frullania baladina Gottsche Ex Stephani, 1894
* Frullania baladina var. edentata Hattori, 1984
* Frullania baltica Grolle, 1985
* Frullania baumannii Hattori, 1977
* Frullania beauverdii Stephani, 1916
* Frullania bella Stephani, 1911
* Frullania belmorensis Stephani In Stephani & Watts, 1914
* Frullania benjaminiana Inoue In Hattori, 1975
* Frullania bergmanii Hattori, 1974
* Frullania berthoumieui Stephani, 1894
* Frullania beyrichiana (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
* Frullania bhutanensis Hattori, 1971
* Frullania bicornistipula Spruce, 1884
* Frullania blastopetala Hattori, 1984
* Frullania blepharozia Spruce, 1884
* Frullania bogotensis Stephani, 1910
* Frullania bolanderi Austin, 1869
* Frullania bonariensis Reiner, 1988
* Frullania bonincola Hattori, 1978
* Frullania borbonica Lindenberg In G., L. & N., 1845
* Frullania boveana Massalongo, 1885
* Frullania brachycarpa Spruce, 1889
* Frullania brasiliensis Raddi, 1822
* Frullania brasiliensis var. cylindrica (Gottsche In Lehmann) Spruce, 1884
* Frullania brasiliensis var. cylindrica F. Lindigii (Gottsche) Spruce, 1884
* Frullania breuteliana Gottsche In G., L. & N., 1845
* Frullania brevicalycina Stephani, 1894
* Frullania brittoniae Evans, 1897
* Frullania brotheri Stephani, 1894
* Frullania brunnea (Sprengel) Drège, 1843
* Frullania buchtienii Herzog, 1942
* Frullania bullata Stephani, 1910
C
* Frullania caduca Hattori, 1980
* Frullania caespitans Beauverd In Stephani, 1924
* Frullania caffraria Stephani, 1894
* Frullania caffraria F. Anomala Vanden Berghen, 1976
* Frullania calcarata Ångström, 1873
* Frullania caldensis Ångström, 1876
* Frullania caledonica Gottsche Ex Stephani, 1894
* Frullania californica (Austin Ex Underwood) Evans, 1897
* Frullania campanulata Sande Lacoste, 1853
* Frullania campanulata var. caduca Verdoorn, 1930
* Frullania campanulata var. malesiaca (Verdoorn) Hattori, 1975
* Frullania canaliculata Gottsche Ex Stephani, 1910
* Frullania capensis Gottsche In G., L. & N., 1845
* Frullania capillaris Stephani, 1911
* Frullania carrii Kamimura & Hattori In Hattori & Kamimura, 1973
* Frullania casparyi Grolle, 1985
* Frullania cataractarum Stephani, 1911
* Frullania caulisequa (Nees In Martius) Montagne, 1839
* Frullania cavallii Gola, 1907
* Frullania cesatiana De Notaris, 1865
* Frullania changii Hattori & Gao, 1985
* Frullania chenii Hattori & Lin, 1985
* Frullania chevalieri (Schuster) Schuster, 1992
* Frullania chiapasana Stephani, 1910
* Frullania chilcootiensis Stephani, 1887
* Frullania chilensis Stephani, 1894
* Frullania chiovendae Gola, 1914
* Frullania chodatii Beauverd In Stephani, 1924
* Frullania ciliata Lindenberg & Gottsche In G., L. & N., 1847
* Frullania cinchonae Gottsche In G., L. & N., 1845
* Frullania clandestina (Nees & Montagne) Nees In G., L. & N., 1845
* Frullania clavata (Hooker & Taylor) Taylor In G., L. & N., 1845
* Frullania claviloba Stephani, 1911
* Frullania clemensiana Verdoorn, 1932
* Frullania cobrensis Gottsche Ex Stephani, 1894
* Frullania colliculosa von Konrat, Braggins, Hentschel & Heinrichs, 2010
* Frullania compacta Gottsche Ex Stephani, 1911
* Frullania complicata Stephani, 1911
* Frullania confertiloba Stephani, 1910
* Frullania consociata Stephani, 1910
* Frullania contracta Stephani, 1911
* Frullania controversa Beauverd In Stephani, 1924
* Frullania convoluta Lindenberg & Hampe In Hampe, 1851
* Frullania convoluta var. ampliata Herzog, 1953
* Frullania cordaeana Lindenberg In G., L. & N., 1845
* Frullania cordistipula (Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
* Frullania cordistipula var. (Beta) mutica Nees In G., L. & N., 1845
* Frullania cordistipula var. (Gamma) regularis Nees In G., L. & N., 1845
* Frullania cordistipula var. dentistipula Hattori, 1986
* Frullania cornuta Stephani, 1911
* Frullania crassitexta Stephani, 1910
* Frullania crawfordii Stephani, 1894
* Frullania crenulifolia Jack & Stephani, 1892
* Frullania crinoidea Spruce Ex Stephani In Stephani, 1911
* Frullania crispiloba Stephani, 1894
* Frullania crispiplicata Yuzawa & Hattori Ex Konstantinova, Potemkin & Schijakov, 1992
* Frullania cristata Hattori, 1981
* Frullania cuencensis Taylor, 1846
* Frullania cuneatistipula Stephani, 1924
* Frullania cuneiloba Nees In G., L. & N., 1845
* Frullania curviramea Stephani, 1911
* Frullania curvirostris Colenso, 1889
* Frullania curvistipula Stephani, 1911
* Frullania curvistipula var. falcatidenta Hattori, 1982
* Frullania curvistipula var. lamii Verdoorn, 1930
* Frullania curvistipula var. latistipula Hattori, 1978
* Frullania cuspidifolia Stephani, 1911
* Frullania cuspiloba Stephani, 1910
* Frullania cyparioides (Schwägrichen) Schwaegrichen In G., L. & N., 1845
D
* Frullania darwinii Gradstein & Uribe In Uribe, 2004
* Frullania davurica Hampe In G., L. & N., 1845
* Frullania davurica F. Dorsoblastos (Hattori) Hattori & Lin, 1985
* Frullania davurica F. Microphylla (Massalongo) Hattori & Lin, 1985
* Frullania davurica var. chichibuensis (Kamimura) Hattori, 1976
* Frullania davurica var. concava Chang In Gao & Chang, 1981
* Frullania debilis Stephani Ex Hattori, 1974
* Frullania decidua Spruce, 1884
* Frullania deflexa Mitten, 1862
* Frullania degelii Arnell, 1959
* Frullania densifolia Subsp. Andamana Hattori Ex Singh In Nath & Asthana, 2001
* Frullania densiloba Stephani Ex Evans, 1906
* Frullania dentata Hattori, 1974
* Frullania dentata var. secernens Hattori, 1988
* Frullania dentifera Hattori & Streimann, 1985
* Frullania dentiloba Hattori, 1975
* Frullania deplanata Mitten In Hooker, 1855
* Frullania deppii Gottsche Ex Lehmann, 1844
* Frullania depressa Mitten, 1863
* Frullania dilatata Subsp. (Dilatata 1982) (L.) Dumortier, 1835
* Frullania dilatata Subsp. (Dilatata 1982) F. Fuscovirens Jorgensen, 1934
* Frullania dilatata Subsp. (Dilatata 1982) var. anomala Corbiere, 1889
* Frullania dilatata Subsp. (Dilatata 1982) var. macrotus Nees, 1838
* Frullania dilatata Subsp. (Dilatata 1982) var. subtilissima Nees, 1838
* Frullania dilatata Subsp. Asiatica Hattori, 1982
* Frullania diptera (Lehmann & Lindenberg) Drège, 1843
* Frullania dispar Nees In G., L. & N., 1845
* Frullania diversitexta Stephani, 1897
* Frullania donnellii Austin, 1879
* Frullania dulimensis Uribe, 2006
* Frullania durifolia Stephani, 1894
* Frullania dusenii Stephani In Dusen, 1905
* Frullania duthiana Stephani, 1910
* Frullania duthiana var. laevis Hattori In Ohashi, 1975
* Frullania duthiana var. szechuanensis Hattori & Gao, 1985
E
* Frullania eboracensis (Subsp. Eboracensis 1992) Gottsche Ex Lehmann, 1844
* Frullania eboracensis Subsp. Virginica (Gottsche In Lehmann) Schuster, 1992
* Frullania echinantha Hattori, 1974
* Frullania echinatella Hattori, 1988
* Frullania ecklonii (Sprengel) Sprengel In G., L. & N., 1845
* Frullania ecklonii F. Robustior Gottsche, 1863
* Frullania ecklonii F. Tenerior Sprengel, 1847
* Frullania ecklonii var. huitamalcensis Gottsche, 1863
* Frullania ecklonii var. rufescens Sprengel, 1847
* Frullania ecuadoriensis Stephani, 1911
* Frullania elegans Lehmann, 1857
* Frullania elephantum Hattori, 1977
* Frullania engelii Hattori, 1983
* Frullania epiphylla Subsp. Fijiensis Hattori, 1985
* Frullania eplicata Stephani, 1911
* Frullania ericoides (Nees In Martius) Montagne, 1839
* Frullania ericoides F. Ericoides (Nees) Verdoorn Ex Piippo, 1990
* Frullania ericoides var. planescens (Verdoorn) Hattori, 1984
* Frullania erostrata Hattori, 1974
* Frullania errans Verdoorn, 1930
* Frullania errans var. angulistipula Hattori, 1972
* Frullania esenbeckiana Beauverd Ex Stephani, 1924
* Frullania evelynae Hattori & Thaithong, 1978
* Frullania evoluta Mitten, 1861
* Frullania expansa Stephani, 1897
* Frullania eymae Hattori, 1975
* Frullania eymae var. crispidentata Hattori & Streimann, 1985
F
* Frullania falciloba Taylor Ex Lehmann, 1844
* Frullania fallax Gottsche In G., L. & N., 1845
* Frullania falsicornuta Hattori, 1986
* Frullania falsisinuata Hattori & Piippo, 1986
* Frullania falsisinuata F. Parvistylata (Hattori) Hattori & Piippo, 1986
* Frullania falsisinuata var. crispidentata Hattori & Piippo, 1986
* Frullania fauriana Stephani, 1894
* Frullania fauriana F. Emarginata Kamimura, 1952
* Frullania fengyangshanensis Zhu & So, 1997
* Frullania ferdinandi-Muelleri Stephani, 1910
* Frullania fertilis var. major Massalongo, 1885
* Frullania flammea Taylor Ex Spruce, 1884
* Frullania flexicaulis Spruce, 1884
* Frullania flexuosa Hattori, 1983
* Frullania fragilifolia (Taylor) Taylor In G., L. & N., 1845
* Frullania franciscana Howe, 1894
* Frullania fuegiana Stephani, 1910
* Frullania fugax (Hooker & Taylor) Taylor In G., L. & N., 1845
* Frullania fulfordiae Hattori, 1987
* Frullania fusco-Virens Stephani, 1910
* Frullania fusco-Virens var. gemmipara (Schuster & Hattori In Hattori) Hattori & Lin, 1985
G
* Frullania gabonensis Vanden Berghen, 1976
* Frullania gaoligongensis Bai & Gao, 1999
* Frullania gaudichaudii (Nees & Montagne) Nees & Montagne In G., L. & N., 1845
* Frullania gaudichaudii F. Hasseltii (Sande Lacoste) Verdoorn, 1930
* Frullania gemmulosa Hattori & Thaithong In Hattori, Thaithong & Kitagawa, 1977
* Frullania gibbosa (Nees) Nees In Montagne, 1840
* Frullania gibbosa var. major Gottsche, 1863
* Frullania gigantea Stephani, 1911
* Frullania giraldiana Massalongo, 1897
* Frullania giraldiana var. handelii (Verdoorn) Hattori, 1972
* Frullania globosa Hattori & Streimann, 1985
* Frullania glomerata (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg Ex Montagne, 1838
* Frullania gracilicaulis Hattori, 1977
* Frullania gracilis Subsp. (Gracilis 1978) (Reinwardt, Blume & Nees) Gottsche In G., L. & N., 1845
* Frullania gracilis Subsp. (Gracilis 1978) var. brevior Nees In G., L. & N., 1845
* Frullania gracilis Subsp. (Gracilis 1978) var. vittata Hattori, 1986
* Frullania gracilis Subsp. Zennoskei Hattori & Thaithong, 1978
* Frullania gradsteinii Yuzawa, Mues & Hattori, 1987
* Frullania granatensis Gottsche, 1864
* Frullania grandifolia Stephani, 1911
* Frullania grandilobula Hattori & Piippo, 1986
* Frullania grandistipula Lindenberg In G., L. & N., 1845
* Frullania griffithsiana Gottsche In G., L. & N., 1846
* Frullania grolleana Hattori, 1972
* Frullania grossiclava Stephani, 1910
* Frullania grossifolia Stephani, 1911
* Frullania guadalupensis Gottsche Ex Stephani, 1911
* Frullania gualaquizana Stephani, 1911
* Frullania guatemalensis Stephani, 1911
H
* Frullania haeckeriana Lindenberg In G., L. & N., 1846
* Frullania haematocysta Spruce, 1884
* Frullania hainanensis Hattori & Lin, 1986
* Frullania hamata Stephani, 1911
* Frullania hamaticoma Stephani, 1889
* Frullania hamatiloba Stephani, 1910
* Frullania hamatiloba F. Grosse-Appendiculata Hattori, 1944
* Frullania hamatiloba var. latistipula Hattori, 1975
* Frullania hamatosetacea Grolle In Grolle & Meister, 2004
* Frullania hamiflora Herzog & Clark In Clark & Schultz, 1953
* Frullania handelii Verdoorn In Handel-Mazzetti, 1930
* Frullania handel-Mazzettii Hattori, 1981
* Frullania hariotana Stephani, 1911
* Frullania harpantha Herzog, 1942
* Frullania hasskarliana Lindenberg In G., L. & N., 1845
* Frullania hasskarliana var. gracilis Hattori, 1986
* Frullania hasskarliana var. integribractea (Verdoorn) Hattori, 1975
* Frullania hasskarliana var. parvidentata Hattori, 1986
* Frullania hattoriana Godfrey & Godfrey, 1980
* Frullania hattoriantha Udar & Nath, 1981
* Frullania hattorii Konrat & Braggins, 2003
* Frullania hawaiiensis Miller, 1953
* Frullania hebridensis Stephani, 1911
* Frullania hedrantha Hattori & Kamimura, 1973
* Frullania helleri Stephani, 1910
* Frullania herzogiana Stephani, 1911
* Frullania herzogii Hattori, 1955
* Frullania heteromorpha Schiffner, 1889
* Frullania hicksiae Hattori, 1984
* Frullania hicksiae F. Litoralis Hattori, 1988
* Frullania higuchii Yuzawa & Hattori In Yuzawa & Koike, 1994
* Frullania himalayensis Stephani, 1910
* Frullania hinoi Kamimura, 1982
* Frullania hiroshii Hattori, 1980
* Frullania hirtiflora Spruce, 1884
* Frullania holostipula Hattori & Griffin Iii, 1978
* Frullania hottana Hattori, 1976
* Frullania howeana Stephani In Stephani & Watts, 1914
* Frullania huerlimannii Hattori, 1976
* Frullania huerlimannii var. dioica Hattori, 1984
* Frullania humbertii Vanden Berghen, 1976
* Frullania humilis Spruce, 1890
* Frullania hypoleuca Nees In G., L. & N., 1843
* Frullania hypoleucula Hattori, 1984
I
* Frullania imerinensis Stephani, 1911
* Frullania immersa Stephani, 1896
* Frullania incisoduthiana Hattori In Mizutani, 1979
* Frullania incisoduthiana var. parva Hattori In Mizutani, 1979
* Frullania incisostipula Stephani, 1924
* Frullania inconstans Verdoorn, 1930
* Frullania inconstans F. Integrior Verdoorn, 1930
* Frullania inconstans var. grossedentata Kamimura & Hattori, 1973
* Frullania incumbens Mitten In Hooker, 1855
* Frullania incurva Hattori, 1988
* Frullania inflata Gottsche In G., L. & N., 1845
* Frullania inflata var. communis Schuster, 1985
* Frullania inflata var. dioica Hattori & Thaithong, 1978
* Frullania inflexa Mitten, 1861
* Frullania inflexiloba Hattori, 1984
* Frullania inouei Hattori, 1980
* Frullania integristipula (Nees) Nees In G., L. & N., 1845
* Frullania integristipula var. emarginata Verdoorn, 1929
* Frullania intermedia Subsp. (Intermedia 1980) F. Billardieriana (Nees & Montagne In Montagne) Verdoorn, 1930
* Frullania intermedia Subsp. (Intermedia 1980) var. non-Apiculata Hattori, 1975
* Frullania intermedia Subsp. (Intermedia) (Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
* Frullania intermedia Subsp. Morokensis (Stephani) Hattori, 1980
* Frullania intermixta Colenso, 1889
* Frullania intumescens (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
* Frullania involuta Hampe Ex Stephani, 1911
* Frullania involvens Hattori & Kamimura, 1973
* Frullania iriomotensis Hattori, 1980
* Frullania irregularis Hattori & Piippo, 1986
* Frullania iwatsukii Hattori, 1972
J
* Frullania jackii Subsp. (Jackii 1959) Gottsche In Gottsche & Rabenhorst, 1863
* Frullania jackii Subsp. (Jackii 1959) F. Depauperata Grolle, 1970
* Frullania jacobsii Hattori, 1986
* Frullania jacquinotii Gottsche Ex Stephani, 1910
* Frullania jelskii Loitlesberger, 1894
* Frullania johnsonii Stephani, 1894
* Frullania junghuhniana Gottsche In G., L. & N., 1845
* Frullania junghuhniana var. bisexualis Hattori, 1976
* Frullania junghuhniana var. tenella (Sande Lacoste) Grolle & Hattori In Hattori, 1982
* Frullania junghuhniana var. tenella F. Monoica (Hattori) Hattori, 1986
K
* Frullania kagoshimensis Subsp. (Kagoshimensis 1985) Stephani, 1910
* Frullania kagoshimensis Subsp. (Kagoshimensis 1985) var. minor Kamimura, 1961
* Frullania kagoshimensis Subsp. Hunanensis (Hattori) Hattori, 1985
* Frullania kalimantanensis Hattori, 1986
* Frullania kalimantanensis Piippo & Hattori In Piippo & Tan, 1992
* Frullania kashyapii Verdoorn, 1932
* Frullania kitagawana Hattori, 1984
* Frullania klotzschii Nees Ex Stephani, 1911
* Frullania koponenii Hattori, 1978
* Frullania kunzei (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
* Frullania kunzei var. maritima Schuster, 1991
L
* Frullania laetevirens Hampe In G., L. & N., 1845
* Frullania laevi-Periantha Bai & Gao, 2000
* Frullania lancistyla Stephani, 1910
* Frullania larjiana Singh & Singh, 2005
* Frullania laticaulis Spruce, 1890
* Frullania latiflora Spruce, 1884
* Frullania latogaleata Herzog, 1948
* Frullania laxiflora Spruce, 1884
* Frullania laxiflora var. crossii Spruce, 1884
* Frullania leana Austin, 1869
* Frullania leeuwenii Verdoorn, 1930
* Frullania lepida Hattori & Piippo, 1986
* Frullania letestui Vanden Berghen, 1976
* Frullania levieri Stephani, 1910
* Frullania libera Nees In G., L. & N., 1845
* Frullania lindbergiana Gottsche, 1863
* Frullania lindenbergii Gottsche Ex Lehmann, 1844
* Frullania lindenbergii var. (Beta ?) Fusca Gottsche In G., L. & N., 1847
* Frullania lindeniana Stephani, 1911
* Frullania lindmanii Stephani, 1897
* Frullania linii Hattori, 1981
* Frullania lobato-Hastata Stephani, 1911
* Frullania lobulata (Hooker) Hooker & Nees In G., L. & N., 1845
* Frullania longipinna Stephani, 1910
* Frullania longistipula Stephani In Renauld & Cardot, 1891
* Frullania longistipula var. apiculata Demaret & Vanden Berghen, 1950
* Frullania longistyla Yuzawa & Hattori, 1988
* Frullania loricata Pearson, 1891
* Frullania loricata var. laxa Pearson, 1891
* Frullania ludoviciae Stephani, 1908
* Frullania lushanensis Hattori & Lin, 1985
M
* Frullania macgregorii Stephani, 1894
* Frullania macgregorii var. rostellula (Hattori) Hattori, 1982
* Frullania macrocephala (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
* Frullania macrophylla Hattori, 1980
* Frullania macularis Taylor, 1846
* Frullania madagascariensis Gottsche, 1882
* Frullania madens Stephani, 1924
* Frullania madothecoides Spruce, 1884
* Frullania magellanica (Sprengel) Weber & Nees In G., L. & N., 1845
* Frullania magellanica var. diminutiva Herzog, 1938
* Frullania mammilligera Grolle, 2003
* Frullania mammillosa Hattori, 1977
* Frullania matafaoica Miller, 1981
* Frullania mathanii Stephani, 1911
* Frullania mauritiana Austin, 1869
* Frullania maymyoensis Svihla, 1958
* Frullania mcveanii Hattori, 1973
* Frullania media (Hodgson) Hattori, 1983
* Frullania megalostipa Spruce, 1884
* Frullania meijeri Hattori, 1974
* Frullania meridana Stephani, 1911
* Frullania meyeniana Lindenberg In G., L. & N., 1845
* Frullania meyeniana var. dioica Hattori, 1977
* Frullania microauriculata Verdoorn, 1929
* Frullania microauriculata var. rotundior Verdoorn, 1929
* Frullania microcaulis Gola, 1923
* Frullania microcephala Gottsche, 1863
* Frullania microphylla (Gottsche In Gottsche & Rabenhorst) Pearson, 1894
* Frullania microphylla F. Corticicola Schiffner, 1936
* Frullania microphylla var. deciduifolia Grolle, 1970
* Frullania microrhyncha Clark & Svihla, 1950
* Frullania microscopica Pearson, 1922
* Frullania minor Subsp. (Minor 1975) var. integribracteola Hattori, 1975
* Frullania minor Subsp. Recurviloba Hattori, 1975
* Frullania mirabilis Jack & Stephani In Stephani, 1892
* Frullania miradorensis Lindenberg & Gottsche In G., L. & N., 1847
* Frullania mizutanii Kamimura & Hattori, 1973
* Frullania moniliata Subsp. Breviramea (Stephani) Verdoorn, 1930
* Frullania moniliata Subsp. Moniliata Crandall-Stotler, Geissler & Stotler, 1987
* Frullania moniliata Subsp. Moniliata F. Minshanensis (Hattori) Crandall-Stotler Et Alii, 1987
* Frullania moniliata Subsp. Moniliata var. elongatistipula (Verdoorn) Crandall-Stotler Et Alii, 1987
* Frullania moniliata Subsp. Moniliata var. vietnamica (Hattori) Crandall-Stotler Et Alii, 1987
* Frullania monocera (Hooker & Taylor) Taylor In G., L. & N., 1845
* Frullania monocera var. depauperata Hattori, 1984
* Frullania monocera var. schiffneri (Verdoorn) Hattori, 1979
* Frullania monoica Stephani, 1900
* Frullania montagnei Gottsche In G., L. & N., 1845
* Frullania montana Stephani, 1910
* Frullania moritziana Lindenberg & Gottsche In G., L. & N, 1847
* Frullania moritziana var. (Beta) Mexicana Gottsche, 1863
* Frullania morobensis Hattori & Streimann, 1985
* Frullania motoyana Stephani, 1911
* Frullania mucronata (Lehmann & Lindenberg In Lehmann) Lehmann & Linde,nberg
* Frullania mucronata var. submutica Lehmann & Lindenberg In G., L. & N. 1845 In G., L. & N., 1845
* Frullania multilacera Subsp. (Multilacera 1987) Stephani, 1911
* Frullania multilacera Subsp. (Multilacera 1987) var. lacerissima Hattori, 1975
* Frullania multilacera Subsp. Gracilior Hattori, 1987
* Frullania multilaceroides Hattori, 1987
* Frullania muscicola Stephani, 1894
* Frullania mutilata Stephani, 1911
N
* Frullania nadeaudii Stephani, 1911
* Frullania neocaledonica Engel In Engel & Merrill, 1999
* Frullania neosheana Hattori, 1979
* Frullania nepalensis (Sprengel) Lehmann & Lindenberg In G., L. & N., 1845
* Frullania neurota Taylor, 1846
* Frullania nicholsonii Hodgson, 1949
* Frullania nigricaulis (R., B. & N.) Nees In G., L. & N., 1845
* Frullania nigricaulis var. elongata Verdoorn, 1930
* Frullania nisquallensis Sullivant, 1849
* Frullania nivimontana Hattori, 1982
* Frullania nobilis Stephani, 1894
* Frullania nobilis var. cochleata (Stephani) Hattori In Hattori & Streimann, 1985
* Frullania nodulosa (Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
* Frullania notarsii Stephani, 1911
* Frullania novocurvirostris Hattori, 1981
* Frullania novoguineensis Schiffner, 1890
O
* Frullania oahuensis Hampe In G., L. & N., 1843
* Frullania oakesiana Subsp. (Oakesiana 1992) Austin, 1870
* Frullania oakesiana Subsp. Takayuensis (Stephani) Schuster, 1992
* Frullania obovata Hattori, 1982
* Frullania obscurifolia Mitten, 1879
* Frullania ocanniensis Stephani, 1924
* Frullania ocellata Kamimura & Hattori, 1973
* Frullania odontostipa Spruce, 1890
* Frullania okinawensis Kamimura, 1982
* Frullania onraedtii Vanden Berghen, 1976
* Frullania orbicularis Austin, 1869
* Frullania orientalis Sande Lacoste, 1855
* Frullania orinocensis Spruce, 1883
* Frullania ornithocephala (Reinwardt, Blume & Nees) Nees In G., L. & N., 1845
* Frullania ornithocephala F. Magnilobula Hattori, 1976
* Frullania ornithocephala F. Retusa Hattori, 1976
* Frullania ornithocephala F. Teres (Sande Lacoste) Verdoorn, 1929
* Frullania ornithocephala var. maior Nees, 1845
* Frullania ornithocephala var. minor Nees, 1845
* Frullania ornithocephala var. pilosa Verdoorn, 1929
* Frullania ornithocephala var. tuberculosa Hattori, 1974
* Frullania osculatiana Notaris, 1855
* Frullania osumiensis (Hattori) Hattori In Iwatsuki & Hattori, 1956
* Frullania osumiensis var. dentata Kamimura, 1970
P
* Frullania pachyderma Hattori, 1978
* Frullania pallide-Virens Stephani, 1911
* Frullania pallidula Hattori, 1988
* Frullania pancheri Gottsche Ex Stephani, 1894
* Frullania papillata Stephani, 1911
* Frullania papillilobula Hattori, 1975
* Frullania papuana Verdoorn, 1930
* Frullania papulosa Stephani, 1911
* Frullania paradoxa Lehmann & Lindenberg In G., L. & N., 1845
* Frullania paradoxa F. Cornistipula Herzog, 1952
* Frullania parhami (Schuster) Schuster, 1992
* Frullania pariharii Hattori & Thaithong, 1978
* Frullania pariharii F. Intermedia Hattori & Thaithong, 1978
* Frullania parvifolia Stephani, 1910
* Frullania parvistipula Stephani, 1910
* Frullania patagonica Stephani, 1910
* Frullania patula Mitten In Hooker, 1855
* Frullania pauciramea Stephani, 1911
* Frullania pauciramea var. pauciramella Hattori & Piippo, 1986
* Frullania paucirameoides Hattori & Piippo, 1986
* Frullania pearceana Stephani, 1911
* Frullania pedicellata Stephani, 1897
* Frullania pentapleura Subsp. (Pentapleura 1988) Taylor, 1846
* Frullania peruviana Gottsche In G., L. & N., 1846
* Frullania phalangiflora Stephani, 1916
* Frullania philippinensis Stephani, 1911
* Frullania physantha Mitten Mitten, 1861
* Frullania pilibracteola Hattori, 1977
* Frullania piliflora Stephani, 1911
* Frullania piliflora var. appendiculata Herzog, 1951
* Frullania pilistipula Stephani, 1911
* Frullania piptophylla Hattori, 1980
* Frullania piptophylla var. minor Hattori, 1986
* Frullania piptophylloides Hattori, 1980
* Frullania pittier I Stephani, 1892
* Frullania plana Sullivant, 1849
* Frullania planifolia Stephani, 1910
* Frullania platycalyx Herzog, 1952
* Frullania platyphylla Colenso, 1889
* Frullania pluricarinata Gottsche, 1864
* Frullania pocsantha Thaithong & Hattori, 1977
* Frullania polyclada Colenso, 1889
* Frullania polyptera Taylor, 1846
* Frullania polyptera var. angustata (Mitten) Hattori, 1975
* Frullania polysticta Lindenberg In G., L. & N., 1845
* Frullania ponapensis Hattori & Koike In Koike, 1994
* Frullania pringlei Fulford & Sharp, 1990
* Frullania probosciphora Taylor, 1846
* Frullania prominula Hattori & Streimann, 1985
* Frullania propaginea Hattori & Streimann, 1985
* Frullania pseudericoides Hattori, 1982
* Frullania pseudericoides Hattori, 1986
* Frullania pseudomeyeniana Hattori, 1986
* Frullania pseudomonocera Hattori, 1986
* Frullania pseudoschensiana Hattori, 1980
* Frullania pseudoschensiana var. darjeelingensis Hattori, 1981
* Frullania ptychantha Montagne, 1843
* Frullania pulchella Herzog, 1954
* Frullania pullei Verdoorn, 1930
* Frullania pulogensis Stephani, 1911
* Frullania punctata Reimers, 1931
* Frullania purpurea Stephani, 1911
* Frullania pusilla Mitten, 1871
* Frullania pycnantha (Hooker & Taylor) Taylor In G., L. & N., 1845
* Frullania pycnoclada Grolle In Grolle & Meister, 2004
* Frullania pyricalycina Stephani, 1910
Q
* Frullania queenslandica Stephani, 1910
* Frullania quillotensis (Nees & Montagne) Nees & Montagne In Montagne In Alcide D'orbigny, 1839
* Frullania quillotensis F. Flaccida Herzog, 1954
* Frullania quillotensis F. Leiantha Herzog, 1954
R
* Frullania rabenhorstii Stephani, 1911
* Frullania ramuligera (Nees) Montagne, 1842
* Frullania recurvistipula Hattori, 1975
* Frullania reflexistipula Sande Lacoste, 1853
* Frullania reflexistipula var. novocaledonica Hattori, 1977
* Frullania reflexistipula var. squarrosa Hattori & Piippo, 1986
* Frullania regularis Schiffner, 1890
* Frullania reicheana Stephani, 1910
* Frullania reimersii Verdoorn, 1930
* Frullania remotidens Hattori, 1872
* Frullania remotifolia Stephani, 1911
* Frullania remotiloba Stephani, 1894
* Frullania repanda Gottsche, 1864
* Frullania repandistipula Subsp. (Repandistipula 1975) Sande Lacoste, 1853
* Frullania repandistipula Subsp. Queenslandica Hattori, 1987
* Frullania repandistipula Subsp. Spinibractea Hattori, 1975
* Frullania reptans var. integristipula Nicholson, 1925
* Frullania retusa Mitten, 1861
* Frullania retusa var. gymnantha Hattori & Thaithong, 1978
* Frullania retusa var. hirsuta Hattori & Thaithong, 1978
* Frullania rhystocolea Herzog Ex Verdoorn In Handel-Mazzetti, 1930
* Frullania rhytidantha Hattori, 1980
* Frullania rigescens Spruce, 1884
* Frullania rigida Stephani, 1910
* Frullania ringens Spruce, 1884
* Frullania riojaneirensis (Raddi) Ångström, 1876
* Frullania riparia Hampe Ex Lehmann, 1838
* Frullania rizalli Piippo & Hattori, 1992
* Frullania rostellata Mitten In Hooker, 1867
* Frullania rostrata (Hooker & Taylor) Hooker & Taylor In G., L. & N., 1845
* Frullania rubella Gottsche Ex Stephani, 1889
* Frullania rubella var. elongata (Stephani) Hattori, 1983
* Frullania rudolfiana Hattori, 1972
* Frullania rupicola Stephani, 1924
S
* Frullania sabahana Hattori, 1976
* Frullania sabaliana Schuster, 1983
* Frullania sabanetica Gottsche, 1864
* Frullania sachapatensis Stephani, 1911
* Frullania sackawana Stephani, 1897
* Frullania sackawana var. minor Kamimura, 1982
* Frullania saipanensis Hattori & Koike In Koike, 1994
* Frullania sarawakensis Hattori, 1976
* Frullania scalaris Hattori, 1977
* Frullania scandens Montagne, 1843
* Frullania schaefer-Verwimpii Yuzawa & Hattori, 1989
* Frullania schensiana Massalongo, 1897
* Frullania schiffneri Verdoorn, 1929
* Frullania schimperi Nees In G., L. & N., 1845
* Frullania schimperi var. laciniata Vanden Berghen, 1976
* Frullania schumannii (Caspary) Grolle, 1981
* Frullania schusteri Hattori, 1988
* Frullania schusteriana Hattori, 1972
* Frullania scottiana Hattori, 1987
* Frullania sebastianopolitana var. (Delta) Gottsche, 1864
* Frullania selwyniana Pearson, 1890
* Frullania semienana Gola, 1914
* Frullania semivillosa Lindenberg & Gottsche In G., L. & N., 1847
* Frullania semivillosa var. (Beta) Glabra Lindenberg & Gottsche In G., L. & N., 1847
* Frullania sergiae Sim-Sim, Fontihna, Mues & Lion, 2000
* Frullania seriata Gottsche Ex Stephani, 1889
* Frullania seriatifolia Stephani, 1894
* Frullania serrata Subsp. (Serrata 1982) Gottsche In G., L. & N., 1845
* Frullania serrata Subsp. (Serrata 1982) F. Crispulodentata Verdoorn, 1929
* Frullania serrata Subsp. (Serrata 1982) F. Vogelkopensis Hattori, 1982
* Frullania serrata Subsp. (Serrata 1982) var. ceramensis Hattori, 1986
* Frullania serrata Subsp. (Serrata 1982) var. hamatispina (Hattori) Hattori, 1982
* Frullania serrata Subsp. (Serrata 1982) var. oceanica Verdoorn, 1937
* Frullania serrata Subsp. (Serrata 1982) var. pertenuis (Nees) Gottsche In G., L. & N., 1845
* Frullania serrata Subsp. Grolleana Grolle & Piippo, 1984
* Frullania serrata Subsp. Spinistipula (Hattori) Hattori, 1982
* Frullania setacea Hattori, 1988
* Frullania setigera Stephani, 1894
* Frullania shanensis Svihla, 1957
* Frullania sharpantha Udar & Kumar, 1983
* Frullania sharpii Hattori, 1974
* Frullania sheana Hattori, 1979
* Frullania simmondsii Stephani In Stephani & Watts, 1914
* Frullania sinensis Stephani In Levier, 1906
* Frullania sinosphaerantha Hattori & Lin, 1985
* Frullania sinuata Subsp. Sinuata 1975 Sande Lacoste, 1853
* Frullania sinuata Subsp. Sinuata F. Parvistylata Hattori In Grolle & Piippo, 1984
* Frullania socotrana Mitten In Balfour, 1888
* Frullania solanderiana Colenso, 1889
* Frullania spathulistipa Stephani, 1910
* Frullania speciosa Herzog, 1949
* Frullania spegazzinii Reiner, 1988
* Frullania sphaerantha Hattori, 1980
* Frullania sphaerocephala Spruce, 1884
* Frullania sphaerolobulata Lin In Lin & Chen, 1997
* Frullania spicata (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
* Frullania spinifera Taylor, 1846
* Frullania spinigastria Hattori, 1979
* Frullania spiniplica Hattori, 1972
* Frullania spinistipula Stephani, 1911
* Frullania spongiosa Stephani, 1894
* Frullania squamuligera Spruce, 1884
* Frullania squarrosula (Hooker & Taylor) Taylor In G., L. & N., 1845
* Frullania standaerti Stephani, 1910
* Frullania steereana Hattori, 1987
* Frullania stenostipa Spruce, 1884
* Frullania streimannii Hattori, 1983
* Frullania stylifera (Schuster) Schuster, 1992
* Frullania subcaduca Hattori, 1974
* Frullania subclavata Stephani, 1910
* Frullania subdentata Stephani, 1911
* Frullania subdentata F. Longistipula Hattori, 1978
* Frullania subdilatata Massalongo In Levier, 1906
* Frullania subhampeana Hodgson, 1949
* Frullania subincumbens Hattori, 1987
* Frullania sublignosa Stephani, 1894
* Frullania submultilacera Hattori & Koike In Koike, 1994
* Frullania subnigricaulis Hattori, 1973
* Frullania subnigricaulis var. subtruncata Hattori, 1975
* Frullania subocellata Hattori, 1986
* Frullania subpedicellata Hattori, 1980
* Frullania subpilibracteola Hattori, 1977
* Frullania subpyricalycina Herzog, 1954
* Frullania subrostrata Hattori, 1972
* Frullania subsquarrosa Hattori, 1972
* Frullania subtilissima Lindenberg In G., L. & N., 1845
* Frullania subtropica Stephani, 1910
* Frullania subtruncata Stephani, 1910
* Frullania subvalida Hattori & Thaithong, 1978
* Frullania supradecomposita (Lehmann & Lindenberg In Lehmann) Lehmann & Lindenberg In G., L. & N., 1845
* Frullania svihlana Hattori, 1983
T
* Frullania tagawana (Hattori, Thaithong & Kitawana) Hattori In Hattori & Lin, 1985
* Frullania taiheizana Horikawa, 1934
* Frullania tamarisci (L.) Dumortier, 1835
* Frullania tamarisci var. atlantica Schiffner, 1936
* Frullania tamarisci var. atrovirens Carrington, 1863
* Frullania tamarisci var. commutata Nees In G., L. & N., 1845
* Frullania tamarisci var. cornubica Carrington In Carrington & Pearson, 1878
* Frullania tamarisci var. laxa Nees In G., L. & N., 1845
* Frullania tamarisci var. mediterranea (De Notaris) Nees In G., L. & N., 1845
* Frullania tamarisci var. robusta Lindberg, 1875
* Frullania tamarisci var. sardoa (De Notaris) De Notaris, 1865
* Frullania tamarisci var. schiffneri Nicholson, 1933
* Frullania tamsuina Stephani, 1910
* Frullania taradakenis Stephani, 1910
* Frullania taxodiocola Schuster, 1983
* Frullania teneriffae (Weber) Nees, 1838
* Frullania teneriffae F. Explanata (Macvicar) Bisang, Schumacker, Sérgio & Grolle, 1988
* Frullania teneriffae var. germana (Taylor) Bisang, Schumacker, Sérgio & Grolle, 1988
* Frullania tenuirostris Stephani, 1911
* Frullania ternatensis Gottsche In G., L. & N., 1846
* Frullania ternatensis var. non-Appendiculata Hattori, 1974
* Frullania tetraptera Nees & Montagne In Montagne, 1838
* Frullania thiersiae Hattori, 1988
* Frullania tixieri Hattori, 1976
* Frullania tjibodensis Hattori & Thaithong, 1977
* Frullania togashiana Hattori, 1975
* Frullania tongariroense Colenso, 1889
* Frullania trianae Gottsche, 1864
* Frullania tricarinata Sande Lacoste, 1856
* Frullania trichodes Mitten, 1862
* Frullania trigona Clark, Jovet-Ast & Frye, 1947
* Frullania trinervis (Lehmann & Lindenberg) Drège, 1843
* Frullania triquetra Lindenberg & Gottsche In G., L. & N., 1847
* Frullania trollii Herzog, 1942
* Frullania truncata Caspary, 1887
* Frullania truncatistyla von Konrat et al., 2011
* Frullania tubercularis Hattori & Lin, 1985
* Frullania tunguraguana Clark & Frye, 1952
* Frullania turfosa Lindenberg & Gottsche In G., L. & N., 1847
* Frullania tuyamae Hattori & Thaithong, 1978
U
* Frullania udarii Nath & Singh, 2006
* Frullania umbonata Mitten Ex Stephani, 1911
* Frullania undulata Kamimura, 1961
* Frullania usambarana Schiffner Ex Stephani, 1894
* Frullania usambarana var. reducta Vanden Berghen, 1976
* Frullania usamiensis Stephani, 1897
* Frullania utriculata Stephani, 1894
V
* Frullania vaga Mitten In Seemann, 1871
* Frullania vaginata (Swartz) Nees In G., L. & N., 1846
* Frullania valdiviensis Jack & Stephani In Stephani, 1894
* Frullania valida Stephani, 1910
* Frullania valparaisiana Lehmann, 1857
* Frullania vandenberghenii Pocs In Bizot & Pocs, 1979
* Frullania van-Zantenii Kamimura & Hattori, 1973
* Frullania variabilis Stephani, 1910
* Frullania varians Caspary, 1887
* Frullania variegata Stephani, 1894
* Frullania venusta Hattori, 1974
* Frullania verdoorniana Hattori, 1973
* Frullania victoriensis Stephani, 1910
* Frullania vitalii Yuzawa & Hattori, 1988
* Frullania vittata Hattori, 1974
* Frullania vittata F. Denticulata Hattori, 1980
* Frullania vittiana Hattori, 1987
W
* Frullania wagneri Stephani, 1910
* Frullania wairua Konrat & Braggins, 2005
* Frullania wangii Hattori & Lin, 1985
* Frullania warnckeana Hattori, 1974
* Frullania warnckeana var. dentosa Hattori, 1977
* Frullania weberbaueri Stephani, 1911
* Frullania winteri Stephani, 1910
* Frullania winteri var. vanderhammenii (Haarbrink) Yuzawa, 1991
Y
* Frullania yorkiana Stephani, 1911
* Frullania yuennanensis Stephani, 1894
* Frullania yuennanensis var. siamensis (Hatt. et al.) Hattori & Lin, 1985
* Frullania yusawae Kamimura, 1970
* Frullania yuzawana Hattori, 1981
Z
* Frullania zangii Hattori & Lin, 1985
* Frullania zennoskeana Hattori, 1984

## Extinct species

* †Frullania baerlocheri Heinrichs et al., 2012 Burmese amber, Myanmar, Cenomanian
* †Frullania cretacea Hentschel et al., 2009 Burmese amber, Myanmar, Cenomanian
* †Frullania partita Li et al., 2020 Burmese amber, Myanmar, Cenomanian
* †Frullania pinnata Heinrichs et al., 2017 Burmese amber, Myanmar, Cenomanian