# Phylinae

Phylinae is a subfamily of the plant bug family Miridae. Species of this family are found worldwide.
In research published in 2013, the subfamily Phylinae was reorganized. The tribe Auricillocorini is now considered a synonym of Hallodapini, and the tribe Pronotocrepini a synonym of the subtribe Cremnorrhina. New tribes were created under the new or resurrected names Decomiini, Exocarpocorini, Cremnorrhini, Nasocorini, and Semiini. The revised subfamily consists of nine tribes containing around 400 genera and over 2000 species.

## Subfamily Phylinae Douglas & Scott, 1865

    Tribe Cremnorrhinini Reuter, 1883 (sometimes spelled "Cremnorrhini")
       Subtribe Coatonocapsina Schuh & Menard, 2013
       Subtribe Cremnorrhina Reuter, 1883
    Tribe Decomiini Schuh & Menard, 2013
    Tribe Exaeretini Futon, 1875
    Tribe Hallodapini Van Duzee, 1916
    Tribe Leucophoropterini Schuh, 1974
       Subtribe Leucophoropterina Schuh, 1974
       Subtribe Tuxedoina Schuh & Menard, 2013
    Tribe Nasocorini Reuter, 1883
    Tribe Phylini Douglas & Scott, 1865
       Subtribe Keltoniina Schuh & Menard, 2013
       Subtribe Oncotylina Douglas & Scott, 1865
       Subtribe Phylina Douglas & Scott, 1865
    Tribe Pilophorini Douglas & Scott, 1876
    Tribe Semiini Knight, 1923
       Subtribe Exocarpocorina Schuh & Menard, 2013
       Subtribe Semiina Knight, 1923

## Genera

Phylinae contains the following genera:

### Subfamily Phylinae Douglas & Scott, 1865

    Tribe Cremnorrhinini Reuter, 1883
       Subtribe Coatonocapsina Schuh & Menard, 2013
          Genus Austropsallus Schuh, 1974
          Genus Capecapsus Schuh, 1974
          Genus Coatonocapsus Schuh, 1974
          Genus Heterocapillus Wagner, 1960
          Genus Parasciodema Poppius, 1914
       Subtribe Cremnorrhinina Reuter, 1883
          Genus Amblytylus Fieber, 1858
          Genus Brachyceratocoris Knight, 1968
          Genus Calidroides Schwartz, 2005
          Genus Coquillettia Uhler, 1890
          Genus Cremnorrhinus Reuter, 1880
          Genus Dacota Uhler, 1872
          Genus Denticulophallus Schuh, 1974
          Genus Ethelastia Reuter, 1876
          Genus Euderon Puton, 1888
          Genus Excentricoris Carvalho, 1955
          Genus Guentherocoris Schuh & Schwartz, 2004
          Genus Harpocera Curtis, 1838
          Genus Leutiola Wyniger, 2012
          Genus Lopidodenus V. Putshkov, 1974
          Genus Lopus Hahn, 1833
          Genus Macrotylus Fieber, 1858
          Genus Orectoderus Uhler, 1876
          Genus Pachyxyphus Fieber, 1858
          Genus Paralopus Wagner, 1957
          Genus Pronotocrepis Knight, 1929
          Genus Shendina Linnavuori, 1975
          Genus Strophopoda Van Duzee, 1916
          Genus Teleorhinus Uhler, 1890
          Genus Ticua Wyniger, 2012
          Genus Utopnia Reuter, 1881
          Genus Zinjolopus Linnavuori, 1975
    Tribe Decomiini Schuh & Menard, 2013
       Genus Aurantiocoris Schuh & Schwartz, 2004
       Genus Decomia Poppius, 1915
       Genus Decomioides Schuh, 1984
       Genus Malaysiamiris Schuh, 1984
       Genus Malaysiamiroides Schuh, 1984
       Genus Rubrocuneocoris Schuh, 1984
    Tribe Exaeretini Futon, 1875
       Genus Anonychiella Reuter, 1912
       Genus Aphaenophyes Reuter, 1899
       Genus Atractotomimus Kiritshenko, 1952
       Genus Auchenocrepis Fieber, 1858
       Genus Brendaphylus Yasunaga, 2013
       Genus Camptotylus Fieber, 1860
       Genus Camptozorus Kerzhner, 1996
       Genus Chrysochnoodes Reuter, 1901
       Genus Compsonannus Reuter, 1902
       Genus Eumecotarsus Kerzhner, 1962
       Genus Eurycranella Reuter, 1904
       Genus Frotaphylus Carvalho, 1984
       Genus Gonoporomiris Henry & Schuh, 2002
       Genus Hadrophyes Puton, 1874
       Genus Hyalopsallus Carvalho & Schaffner, 1973
       Genus Megalodactylus Fieber, 1858
       Genus Moissonia Reuter, 1894
       Genus Opuna Kirkaldy, 1902
       Genus Pastocoris Reuter, 1879
       Genus Psallopsis Reuter, 1901
       Genus Randallopsallus Yasunaga, 2013
       Genus Tuponia Reuter, 1875
       Genus Voruchiella Poppius, 1912
       Genus Yotvata Linnavuori, 1964
    Tribe Hallodapini Van Duzee, 1916
       Genus Acrorrhinium Noualhier, 1895
       Genus Aeolocoris Reuter, 1903
       Genus Alloeomimus Reuter, 1910
       Genus Artchawakomius Yasunaga, 2012
       Genus Aspidacanthus Reuter, 1901
       Genus Auricillocoris Schuh, 1984
       Genus Azizus Distant, 1910
       Genus Bibundiella Poppius, 1914
       Genus Boopidella Reuter, 1907
       Genus Carinogulus Schuh, 1974
       Genus Chaetocapsus Poppius, 1914
       Genus Clapmarius Distant, 1904
       Genus Cleotomiris Schuh, 1984
       Genus Cleotomiroides Schuh, 1984
       Genus Cremnocephalus Eieber, 1860
       Genus Cyrtopeltocoris Reuter, 1876
       Genus Diocoris Kirkaldy, 1902
       Genus Eminoculus Schuh, 1974
       Genus Formicopsella Poppius, 1914
       Genus Gampsodema Odhiambo, 1960
       Genus Glaphyrocoris Reuter, 1903
       Genus Hadrodapus Linnavuori, 1996
       Genus Hallodapomimus Herczek, 2000
       Genus Hallodapus Eieber, 1858
       Genus Ifephylus Linnavuori, 1993
       Genus Kapoetius Schmitz, 1969
       Genus Laemocoris Reuter, 1879
       Genus Leaina Linnavuori, 1974
       Genus Leptomimus Herczek & Popov, 2010
       Genus Lestonisca Carvalho, 1988
       Genus Linacoris Carvalho, 1983
       Genus Lissocapsus Bergroth, 1903
       Genus Malgacheocoris Carvalho, 1952
       Genus Mimocoris J. Scott, 1872
       Genus Myombea China & Carvalho, 1951
       Genus Myrmicomimus Reuter, 1881
       Genus Neolaemocoris Wagner, 1975
       Genus Omphalonotus Reuter, 1876
       Genus Pangania Poppius, 1914
       Genus Paralaemocoris Linnavuori, 1964
       Genus Phoradendrepulus Polhemus & Polhemus, 1985
       Genus Podullahas Schuh, 1984
       Genus Pongocoris Linnavuori, 1975
       Genus Ribautocapsus Wagner, 1962
       Genus Ruwaba Linnavuori, 1975
       Genus Skukuza Schuh, 1974
       Genus Sohenus Distant, 1910
       Genus Syngonus Bergroth, 1926
       Genus Systellonotidea Poppius, 1914
       Genus Systellonotopsis Poppius, 1914
       Genus Systellonotus Pieber, 1858
       Genus Trichophorella Reuter, 1905
       Genus Trichophthalmocapsus Poppius, 1914
       Genus Vitsikamiris Polhemus, 1994
       Genus Wygomiris Schuh, 1984
       Genus Zaratus Distant, 1909
    Tribe Leucophoropterini Schuh, 1974
       Subtribe Leucophoropterina Schuh, 1974
          Genus Abuyogocoris Schuh, 1984
          Genus Aitkenia Carvalho & Gross, 1982
          Genus Arafuramiris Schuh, 1984
          Genus Ausejanus Menard & Schuh, 2011
          Genus Austrodapus Menard & Schuh, 2011
          Genus Biromiris Schuh, 1984
          Genus Blesingia Carvalho & Gross, 1982
          Genus Collessicoris Carvalho & Gross, 1982
          Genus Ctypomiris Schuh, 1984
          Genus Gulacapsus Schuh, 1984
          Genus Johnstonsonius Menard & Schuh, 2011
          Genus Leucophoroptera Poppius, 1921
          Genus Missanos Menard & Schuh, 2011
          Genus Neaitkenia Menard & Schuh, 2011
          Genus Neoleucophoroptera Menard & Schuh, 2011
          Genus Papuamimus Schuh, 1984
          Genus Papuamiroides Menard & Schuh, 2011
          Genus Pseudohallodapocoris Schuh, 1984
          Genus Sejanus Distant, 1910
          Genus Solomonomimus Schuh, 1984
          Genus Transleucophoroptera Menard & Schuh, 2011
          Genus Trichocephalocapsus Schuh, 1984
          Genus Waterhouseana Carvalho, 1973
       Subtribe Tuxedoina Schuh & Menard, 2013
          Genus Ephippiocoris Poppius, 1912
          Genus Pseudophylus Yasunaga, 1999
          Genus Tuxedo Schuh, 2001
    Tribe Nasocorini Reuter, 1883
       Genus Adenostomocoris Schuh & Schwartz, 2004
       Genus Agrametra Buchanan-White, 1878
       Genus Arctostaphylocoris Schuh & Schwartz, 2004
       Genus Atomophora Reuter, 1879
       Genus Atomoscelis Reuter, 1875
       Genus Atractotomoidea Yasunaga, 1999
       Genus Atractotomus Fieber, 1858
       Genus Badezorus Distant, 1910
       Genus Beckocoris Knight, 1968
       Genus Bergmiris Carvalho, 1984
       Genus Boopidocoris Reuter, 1879
       Genus Caiganga Carvalho & Becker, 1957
       Genus Camptotylidea Wagner, 1957
       Genus Campylomma Reuter, 1878
       Genus Chinacapsus Wagner, 1961
       Genus Chlamydatus Curtis, 1833
       Genus Chlamyopsallus Schwartz, 2005
       Genus Hambletoniola Carvalho, 1954
       Genus Helenocoris Schmitz, 1976
       Genus Hirtopsallus Schmitz, 1976
       Genus Insulopus Schmitz, 1976
       Genus Karocris V. Putshkov, 1975
       Genus Kasumiphylus Schwartz & Stonedahl, 2004
       Genus Knightomiroides Stonedahl & Schwartz, 1996
       Genus Lamprosthenarus Poppius, 1914
       Genus Larinocerus
       Genus Lattinophylus Schuh, 2008
       Genus Lindbergopsallus Wagner, 1962
       Genus Lopsallus Schmitz, 1976
       Genus Maurodactylus Reuter, 1878
       Genus Megalopsallus Knight, 1927
       Genus Monosynamma J. Scott, 1864
       Genus Naresthus Schmitz, 1976
       Genus Nasocoris Reuter, 1879
       Genus Neisopsallus Schmitz, 1976
       Genus Neophylus Carvalho & Costa, 1992
       Genus Nevadocoris Knight, 1968
       Genus Nigrimiris Carvalho & Schaffner, 1973
       Genus Nigrocapillocoris Wagner, 1973
       Genus Oligobiella Reuter, 1885
       Genus Orthopidea Reuter, 1899
       Genus Phaxia Kerzhner, 1984
       Genus Phoenicocoris Reuter, 1875
       Genus Pinomiris Stonedahl & Schwartz, 1996
       Genus Pruneocoris Schuh & Schwartz, 2004
       Genus Psallomimus Wagner, 1951
       Genus Rhinacloa Reuter, 1876
       Genus Salicarus Kerzhner, 1962
       Genus Solenoxyphus Reuter, 1875
       Genus Spanagonicus Berg, 1883
       Genus Squamophylus Carvalho & Costa, 1992
       Genus Sthenaropsis Poppius, 1912
       Genus Taeniophorus Linnavuori, 1952
       Genus Tannerocoris Knight, 1970
       Genus Tapuruyunus Carvalho, 1946
       Genus Thymopsallus Linnavuori, 1975
       Genus Tijucaphylus Carvalho & Costa, 1992
       Genus Tunisiella Carapezza, 1997
       Genus Voruchia Reuter, 1879
    Tribe Phylini Douglas & Scott, 1865
       Subtribe Keltoniina Schuh & Menard, 2013
          Genus Keltonia Knight, 1966
          Genus Pseudatomoscelis Poppius, 1911
          Genus Reuteroscopus Kirkaldy, 1905
          Genus Waupsallus Linnavuori, 1975
       Subtribe Oncotylina Douglas & Scott, 1865
          Genus Acrotelus Reuter, 1885
          Genus Alloeotarsus Reuter, 1885
          Genus Americodema T. Henry, 1999
          Genus Angelopsallus Schuh, 2006
          Genus Antepia Seidenstiicker, 1962
          Genus Arizonapsallus Schuh, 2006
          Genus Asciodema Reuter, 1878
          Genus Bisulcopsallus Schuh, 2006
          Genus Brachyarthrum Fieber, 1858
          Genus Cariniocoris T. Henry, 1989
          Genus Ceratopsallus Schuh, 2006
          Genus Cercocarpopsallus Schuh, 2006
          Genus Chlorillus Kerzhner, 1962
          Genus Compsidolon Reuter, 1899
          Genus Crassomiris Weirauch, 2006
          Genus Damioscea Reuter, 1883
          Genus Dasycapsus Poppius, 1912
          Genus Europiella Reuter, 1909
          Genus Eurycolpus Reuter, 1875
          Genus Galbinocoris Weirauch, 2006
          Genus Glaucopterum Wagner, 1963
          Genus Hamatophylus Weirauch, 2006
          Genus Hoplomachus Fieber, 1858
          Genus Ihermocoris Puton, 1875
          Genus Insulaphylus Weirauch, 2006
          Genus Josifovius Konstantinov, 2008
          Genus Kmentophylus Duwal, Yasunaga, & Lee, 2010
          Genus Knightophylinia Schaffner, 1978
          Genus Knightopiella Schuh, 2004
          Genus Knightopsallus Schuh, 2006
          Genus Leucodellus Reuter, 1906
          Genus Lineatopsallus T. Henry, 1991
          Genus Litoxenus Reuter, 1885
          Genus Maculamiris Weirauch, 2006
          Genus Malacotes Reuter, 1878
          Genus Marrubiocoris Wagner, 1970
          Genus Megalocoleus Reuter, 1890
          Genus Moiseevichia Schuh, 2006
          Genus Nanopsallus Wagner, 1952
          Genus Neopsallus Schuh & Schwartz, 2004
          Genus Occidentodema T. Henry, 1999
          Genus Oligotylus Van Duzee, 1916
          Genus Omocoris Lindberg, 1930
          Genus Oncotylidea Wagner, 1965
          Genus Oncotylus Fieber, 1858
          Genus Opisthotaenia Reuter, 1901
          Genus Oreocapsus Linnavuori, 1975
          Genus Parachlorillus Wagner, 1963
          Genus Parapsallus Wagner, 1952
          Genus Paredrocoris Reuter, 1878
          Genus Phaeochiton Kerzhner, 1964
          Genus Phallospinophylus Weirauch, 2006
          Genus Phyllopidea Knight, 1919
          Genus Phymatopsallus Knight
          Genus Piceophylus Schwartz & Schuh, 1999
          Genus Pinophylus Schwartz & Schuh, 1999
          Genus Placochilus Fieber, 1858
          Genus Plagiognathus Fieber, 1858
          Genus Plesiodema Reuter, 1875
          Genus Pleuroxonotus Reuter, 1903
          Genus Pronototropis Reuter, 1879
          Genus Psallodema V. Putshkov, 1970
          Genus Psallomorpha Duwal Yasunaga, & Lee, 2010
          Genus Psallovius T. Henry
          Genus Pygovepres Weirauch, 2006
          Genus Quercophylus Weirauch, 2006
          Genus Quernocoris Weirauch, 2006
          Genus Ranzovius Distant, 1893
          Genus Rhinocapsus Uhler, 1890
          Genus Roburocoris Weirauch, 2009
          Genus Rubellomiris Weirauch, 2006
          Genus Rubeospineus Weirauch, 2006
          Genus Sacculifer Kerzhner, 1959
          Genus Salicopsallus Schuh, 2006
          Genus Schaffneropsallus Schuh, 2006
          Genus Stenoparia Fieber, 1870
          Genus Sthenaropsidea Henry & Schuh, 2002
          Genus Stictopsallus Schuh, 2006
          Genus Stirophylus Eckerlein & Wagner, 1965
          Genus Stoebea Schuh, 1974
          Genus Tapirula Carapezza, 1997
          Genus Tinicephalus Fieber, 1858
          Genus Tragiscocoris Fieber, 1861
          Genus Vanduzeephylus Schuh & Schwartz, 2004
          Genus Vesperocoris Weirauch, 2006
          Genus Viscacoris Weirauch, 2009
          Genus Zophocnemis Kerzhner, 1962
       Subtribe Phylina Douglas & Scott, 1865
          Genus Adelphophylus Wagner, 1959
          Genus Agraptocoris Reuter, 1903
          Genus Alnopsallus Duwal, Yasunaga, & Lee, 2010
          Genus Alvarengamiris Carvalho, 1991
          Genus Amazonophilus Carvalho & Costa, 1993
          Genus Anapsallus Odhiambo, 1960
          Genus Anomalocornis Carvalho & Wygodzinsky, 1945
          Genus Arlemiris Carvalho, 1984
          Genus Bicurvicoris Carvalho & Schaffner, 1973
          Genus Botocudomiris Carvalho, 1979
          Genus Brachycranella Reuter, 1905
          Genus Conostethus Fieber, 1858
          Genus Crassicornus Carvalho, 1945
          Genus Darectagela Linnavuori, 1975
          Genus Darfuromma Linnavuori, 1975
          Genus Dignaia Linnavuori, 1975
          Genus Dominiquella Linnavuori, 1983
          Genus Ectagela Schmidt, K., 1939
          Genus Ellacapsus Yasunaga, 2013
          Genus Eremophylus Yasunaga, 2001
          Genus Farsiana Linnavuori, 1998
          Genus Gediocoris Wagner, 1964
          Genus Ghazalocoris Linnavuori, 1975
          Genus Gressittocapsus Schuh, 1984
          Genus Icodema Reuter, 1875
          Genus Indatractus Linnavuori, 1975
          Genus Izyaius Schwartz, 2006
          Genus Juniperia Linnavuori, 1965
          Genus Knightensis Schaffner, 1978
          Genus Lalyocoris Linnavuori, 1993
          Genus Lasiolabopella Schuh, 1974
          Genus Lepidargyrus Muminov, 1962
          Genus Lepidocapsus Poppius, 1914
          Genus Leptoxanthus Reuter, 1905
          Genus Liviopsallus Carapezza, 1982
          Genus Mendozaphylus Carvalho & Carpintero, 1991
          Genus Millerimiris Carvalho, 1951
          Genus Mixtecamiris Carvalho & Schaffner, 1973
          Genus Natalophylus Schuh, 1974
          Genus Nubaia Linnavuori, 1975
          Genus Orthonotus Stephens, 1829
          Genus Parafulvius Carvalho, 1954
          Genus Paravoruchia Wagner, 1959
          Genus Phylus Hahn, 1831
          Genus Plagiognathidea Poppius, 1914
          Genus Platyscytisca Costa & Henry, 1999
          Genus Platyscytus Reuter, 1907
          Genus Porophoroptera Carvalho & Gross, 1982
          Genus Psallus Fieber, 1858
          Genus Roudairea Puton & Reuter, 1886
          Genus Sasajiophylus Yasunaga, 2001
          Genus Somalocoris Linnavuori, 1975
          Genus Sthenarus Fieber, 1858
          Genus Stibaromma Odhiambo, 1961
          Genus Tibiopilus Carvalho & Costa, 1993
          Genus Trevessa China, 1924
          Genus Villaverdea Carvalho, 1990
          Genus Widdringtoniola Schuh, 1974
          Genus Zakanocoris V. Putshkov, 1970
          Genus Zanchiophylus Duwal, Yasunaga, & Lee, 2010
    Tribe Pilophorini Douglas & Scott, 1876
       Genus Alepidiella Poppius, 1914
       Genus Aloea Linnavuori, 1975
       Genus Dilatops Weirauch, 2006
       Genus Druthmarus Distant, 1909
       Genus Ethatractus Linnavuori, 1975
       Genus Hypseloecus Reuter, 1891
       Genus Lasiolabops Poppius, 1914
       Genus Neoambonea Schuh, 1974
       Genus Parambonea Schuh, 1974
       Genus Parasthenaridea Miller, 1937
       Genus Pherolepis Kulik, 1968
       Genus Pilophorus Hahn, 1826
       Genus Pseudambonea Schuh, 1974
       Genus Randallophorus Henry, 2013
       Genus Spinolosus Zou, 1985
       Genus Sthenaridea Reuter, 1885
    Tribe Semiini Knight, 1923
       Subtribe Exocarpocorina Schuh & Menard, 2013
          Genus Ampimpacoris Weirauch & Schuh, 2011
          Genus Ancoraphylus Weirauch, 2007
          Genus Araucanophylus Carvalho, 1984
          Genus Basileobius Eyles & Schuh, 2003
          Genus Chiloephylus Carvalho, 1984
          Genus Cyrtodiridius Eyles & Schuh, 2003
          Genus Exocarpocoris Weirauch, 2007
          Genus Gonzalezinus Carvalho, 1981
          Genus Halormus Eyles & Schuh, 2003
          Genus Harpagophylus Schuh & Weirauch, 2010
          Genus Jiwarli Soto & Weirauch, 2007
          Genus Leptidolon Reuter, 1904
          Genus Mecenopa Eyles & Schuh, 2003
          Genus Melaleucoides Schuh & Weirauch, 2010
          Genus Pimeleocoris Eyles & Schuh, 2003
          Genus Polyozus Eyles & Schuh, 2003
          Genus Thryptomenomiris Schuh & Weirauch, 2010
          Genus Wallabicoris Schuh & Pedraza, 2010
          Genus Xiphoidellus Weirauch & Schuh, 2011
          Genus Xiphoides Eyles & Schuh, 2003
       Subtribe Semiina Knight, 1923
          Genus Criocoris Fieber, 1858
          Genus Hoplomachidea Reuter, 1909
          Genus Karoocapsus Schuh, 1974
          Genus Monocris V. Putshkov, 1974
          Genus Monospatha Eyles & Schuh 2003
          Genus Myrmicopsella Poppius, 1914
          Genus Nicholia Knight, 1929
          Genus Parapseudosthenarus Schuh, 1974
          Genus Pseudosthenarus Poppius, 1914
          Genus Rakula Odhiambo, 1967
          Genus Semium Reuter, 1876
          Genus Thoth Linnavuori, 1993
          Genus Tytthus Fieber, 1864